treatments-xml/data/1A/3B/9C/1A3B9C28246CFFC8FF488BBC738678FB.xml
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<document id="6DC6A92A4B099CE65F9DB429A39CBEE7" ID-CLB-Dataset="24041" ID-DOI="10.11646/zootaxa.4732.3.2" ID-GBIF-Dataset="fda8a42c-e690-401b-b536-861cf6b13d61" ID-ISSN="1175-5326" ID-Zenodo-Dep="3667209" ID-ZooBank="A4B6127B-2C87-4720-B643-29DAE34E849D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1581669909765" checkinUser="plazi" docAuthor="Purkayastha, Jayaditya, Das, Madhurima, Bohra, Sanath Chandra, Bauer, Aaron M. &amp; Agarwal, Ishan" docDate="2020" docId="1A3B9C28246CFFC8FF488BBC738678FB" docLanguage="en" docName="zootaxa.4732.3.2.pdf" docOrigin="Zootaxa 4732 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cyrtodactylus urbanus Purkayastha &amp; Das &amp; Bohra &amp; Bauer &amp; Agarwal 2020, sp. nov." docType="treatment" docVersion="8" lastPageNumber="387" masterDocId="E602E4502465FFC4FFDF8B2571057F79" masterDocTitle="Another new Cyrtodactylus (Squamata: Gekkonidae) from Guwahati, Assam, India" masterLastPageNumber="392" masterPageNumber="375" pageNumber="384" updateTime="1698783054916" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="BB93709CD032DE3186F054017BE2A426">Another new Cyrtodactylus (Squamata: Gekkonidae) from Guwahati, Assam, India</mods:title>
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<mods:namePart id="3C7419CB50059D6B6CAA5552C1E72CF7">Purkayastha, Jayaditya</mods:namePart>
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<mods:namePart id="FDDEF2D21C7D52D3DF088E748FCA9E43">Das, Madhurima</mods:namePart>
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<mods:namePart id="8867AF179E1D376F8BC7BBDA7D650DDB">Bohra, Sanath Chandra</mods:namePart>
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<mods:namePart id="55E7629D21D6C6D2CEBBF8DECEFE1BE8">Bauer, Aaron M.</mods:namePart>
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<mods:date id="7F90A0802C71002DAC35EDC7EB1638D5">2020</mods:date>
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<treatment id="1A3B9C28246CFFC8FF488BBC738678FB" ID-DOI="http://doi.org/10.5281/zenodo.3671684" ID-GBIF-Taxon="162142656" ID-Zenodo-Dep="3671684" LSID="urn:lsid:plazi:treatment:1A3B9C28246CFFC8FF488BBC738678FB" httpUri="http://treatment.plazi.org/id/1A3B9C28246CFFC8FF488BBC738678FB" lastPageId="12" lastPageNumber="387" pageId="9" pageNumber="384">
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<emphasis id="A0E6F12C246CFFCDFF488BBC730A7FCD" bold="true" box="[151,527,153,180]" pageId="9" pageNumber="384">
<taxonomicName id="559256BD246CFFCDFF488BBC70AF7FCD" authority="Purkayastha &amp; Das &amp; Bohra &amp; Bauer &amp; Agarwal, 2020" authorityName="Purkayastha &amp; Das &amp; Bohra &amp; Bauer &amp; Agarwal" authorityYear="2020" box="[151,426,153,180]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="384" phylum="Chordata" rank="species" species="urbanus" status="sp. nov.">
<emphasis id="A0E6F12C246CFFCDFF488BBC70AF7FCD" bold="true" box="[151,426,153,180]" italics="true" pageId="9" pageNumber="384">Cyrtodactylus urbanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD54C57246CFFCDFE6E8BBF730A7FCD" box="[433,527,154,180]" pageId="9" pageNumber="384" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="DA887EB5246CFFCDFF488B9870657FAE" box="[151,352,189,216]" pageId="9" pageNumber="384" type="description">
<paragraph id="922D2D3E246CFFCDFF488B9870657FAE" blockId="9.[151,527,153,216]" box="[151,352,189,216]" pageId="9" pageNumber="384">
<figureCitation id="0AA931BB246CFFCDFF488B9870077FA1" box="[151,258,189,216]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStart-2="FIGURE 7" captionStart-3="FIGURE 8" captionStart-4="FIGURE 9" captionStartId-0="10.[151,250,1962,1987]" captionStartId-1="11.[151,250,1396,1421]" captionStartId-2="12.[151,250,1285,1310]" captionStartId-3="13.[151,250,2001,2026]" captionStartId-4="15.[151,250,1536,1561]" captionTargetBox-0="[168,1419,174,1943]" captionTargetBox-1="[151,1436,641,1369]" captionTargetBox-2="[151,1436,454,1249]" captionTargetBox-3="[210,1377,171,1978]" captionTargetBox-4="[151,1436,553,1517]" captionTargetId-0="figure@10.[168,1419,174,1943]" captionTargetId-1="figure@11.[151,1436,641,1369]" captionTargetId-2="figure@12.[151,1436,454,1249]" captionTargetId-3="figure@13.[210,1377,171,1978]" captionTargetId-4="figure@15.[151,1436,553,1517]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionTargetPageId-2="12" captionTargetPageId-3="13" captionTargetPageId-4="15" captionText-0="FIGURE 5. A: Dorsal view, B: ventral view and C: details of dorsal pholidosis of Cyrtodactylus urbanus sp. nov. (male holotype, VR/ERS/ZSI/683) in life." captionText-1="FIGURE 6. Head of Cyrtodactylus urbanus sp. nov. (male holotype, VR/ERS/ZSI/683), A: dorsal view B: lateral view and C: ventral view." captionText-2="FIGURE 7. Fully everted hemipenis of Cyrtodactylus urbanus sp. nov. (male holotype, VR/ERS/ZSI/683), A: sulcal view and B: asulcal view." captionText-3="FIGURE 8. Type series of Cyrtodactylus urbanus sp. nov. Individuals are labeled by VR/ERS/ZSI specimen number." captionText-4="FIGURE 9. Habitat of Cyrtodactylus urbanus sp. nov., Basistha, Guwahati, Assam" figureDoi-0="http://doi.org/10.5281/zenodo.3667219" figureDoi-1="http://doi.org/10.5281/zenodo.3667221" figureDoi-2="http://doi.org/10.5281/zenodo.3667223" figureDoi-3="http://doi.org/10.5281/zenodo.3667225" figureDoi-4="http://doi.org/10.5281/zenodo.3667227" httpUri-0="https://zenodo.org/record/3667219/files/figure.png" httpUri-1="https://zenodo.org/record/3667221/files/figure.png" httpUri-2="https://zenodo.org/record/3667223/files/figure.png" httpUri-3="https://zenodo.org/record/3667225/files/figure.png" httpUri-4="https://zenodo.org/record/3667227/files/figure.png" pageId="9" pageNumber="384">Figs. 59</figureCitation>
,
<tableCitation id="DF101885246CFFCDFED28B9870657FAE" box="[269,352,189,215]" captionStart="TABLE 4" captionStartId="14.[151,239,154,178]" captionTargetPageId="14" captionText="TABLE 4. Morphometric and meristic data for Cyrtodactylus urbanus sp. nov. from Basishta, Guwahati, Assam, India." httpUri="http://table.plazi.org/id/C6ED7DB6246BFFCAFF488BBF74917FCB" pageId="9" pageNumber="384" tableUuid="C6ED7DB6246BFFCAFF488BBF74917FCB">Table 4</tableCitation>
</paragraph>
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<paragraph id="922D2D3E246CFFCDFF488A2574117E47" blockId="9.[151,1437,256,2041]" pageId="9" pageNumber="384">
<materialsCitation id="22FA2763246CFFCDFF488A2574117E47" ID-GBIF-Occurrence="2565852133" collectingDate="2018-06-13" collectionCode="ERS, SVL" country="India" elevation="106" latitude="26.1063" location="Basistha" longLatPrecision="1" longitude="91.7872" municipality="Guwahati" pageId="9" pageNumber="384" specimenCount="1" stateProvince="Assam" typeStatus="holotype">
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<typeStatus id="4D29939C246CFFCDFF488A2570037E63" box="[151,262,256,282]" pageId="9" pageNumber="384" type="holotype">Holotype</typeStatus>
.
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VR/
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/ZSI/683, adult male,
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.
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,
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,
<collectingRegion id="5056E3DC246CFFCDFBCB8A2575A07E63" box="[1044,1189,256,282]" country="India" name="Assam" pageId="9" pageNumber="384">Assam State</collectingRegion>
,
<collectingCountry id="EA856DAE246CFFCDFB6D8A2575E97E63" box="[1202,1260,256,282]" name="India" pageId="9" pageNumber="384">India</collectingCountry>
(
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,
<geoCoordinate id="F7A64BF9246CFFCDFF488A0170317E47" box="[151,308,292,318]" degrees="91.787199" direction="east" orientation="longitude" pageId="9" pageNumber="384" precision="1" value="91.7872">91.787199° E</geoCoordinate>
),
<quantity id="556A80DB246CFFCDFE968A0170957E47" box="[329,400,292,318]" metricMagnitude="2" metricUnit="m" metricValue="1.06" pageId="9" pageNumber="384" unit="m" value="106.0">
<elevation id="19BFCA0D246CFFCDFE968A0170957E47" box="[329,400,292,318]" metricMagnitude="2" metricUnit="m" metricValue="1.06" pageId="9" pageNumber="384" unit="m" value="106.0">106 m</elevation>
</quantity>
elevation above sea level, collected on
<date id="E62C0BFE246CFFCDFC938A0172E07E47" box="[844,997,292,318]" pageId="9" pageNumber="384" value="2018-06-13">
<collectingDate id="F668F216246CFFCDFC938A0172E07E47" box="[844,997,292,318]" pageId="9" pageNumber="384" value="2018-06-13">13 June 2018</collectingDate>
</date>
, by Jayaditya Purkayastha.
</materialsCitation>
</paragraph>
<paragraph id="922D2D3E246CFFCDFF188A6D72847ED0" blockId="9.[151,1437,256,2041]" pageId="9" pageNumber="384">
<materialsCitation id="22FA2763246CFFCDFF188A6D72787ED0" ID-GBIF-Occurrence="2565852134" collectionCode="ERS, SVL" pageId="9" pageNumber="384" specimenCount="2" typeStatus="paratype">
<emphasis id="A0E6F12C246CFFCDFF188A6D70437E1B" bold="true" box="[199,326,328,354]" pageId="9" pageNumber="384">
<typeStatus id="4D29939C246CFFCDFF188A6D70447E1B" box="[199,321,328,354]" pageId="9" pageNumber="384" type="paratype">Paratypes</typeStatus>
.
</emphasis>
Adult females (VR/
<collectionCode id="F483B5FB246CFFCDFDF18A6D73647E1B" box="[558,609,328,354]" pageId="9" pageNumber="384">ERS</collectionCode>
/ZSI/684,
<collectionCode id="F483B5FB246CFFCDFD0B8A6D720D7E1B" box="[724,776,328,354]" pageId="9" pageNumber="384">SVL</collectionCode>
<quantity id="556A80DB246CFFCDFCCF8A6D727D7E1B" box="[784,888,328,354]" metricMagnitude="-2" metricUnit="m" metricValue="7.43" pageId="9" pageNumber="384" unit="mm" value="74.3">74.3 mm</quantity>
; VR/
<collectionCode id="F483B5FB246CFFCDFC6B8A6D72E27E1B" box="[948,999,328,354]" pageId="9" pageNumber="384">ERS</collectionCode>
/ZSI/688,
<collectionCode id="F483B5FB246CFFCDFB868A6D75887E1B" box="[1113,1165,328,354]" pageId="9" pageNumber="384">SVL</collectionCode>
<quantity id="556A80DB246CFFCDFB4A8A6275FF7E1B" box="[1173,1274,327,354]" metricMagnitude="-2" metricUnit="m" metricValue="7.56" pageId="9" pageNumber="384" unit="mm" value="75.6">75.6 mm</quantity>
and VR/
<collectionCode id="F483B5FB246CFFCDFABD8A6D74917E1B" box="[1378,1428,328,354]" pageId="9" pageNumber="384">ERS</collectionCode>
/ ZSI/689,
<collectionCode id="F483B5FB246CFFCDFEDC8A4970327EFF" box="[259,311,364,390]" pageId="9" pageNumber="384">SVL</collectionCode>
<quantity id="556A80DB246CFFCDFE9F8A4970AE7EFF" box="[320,427,364,390]" metricMagnitude="-2" metricUnit="m" metricValue="7.32" pageId="9" pageNumber="384" unit="mm" value="73.2">73.2 mm</quantity>
), Adult males (VR/
<collectionCode id="F483B5FB246CFFCDFD558A4E73BB7EFF" box="[650,702,363,390]" pageId="9" pageNumber="384">ERS</collectionCode>
/ZSI/685,
<collectionCode id="F483B5FB246CFFCDFCEE8A4972607EFF" box="[817,869,364,390]" pageId="9" pageNumber="384">SVL</collectionCode>
<quantity id="556A80DB246CFFCDFCB18A4E72D27EFF" box="[878,983,363,390]" metricMagnitude="-2" metricUnit="m" metricValue="6.54" pageId="9" pageNumber="384" unit="mm" value="65.4">65.4 mm</quantity>
; VR/
<collectionCode id="F483B5FB246CFFCDFBCB8A4E75427EFC" box="[1044,1095,363,389]" pageId="9" pageNumber="384">ERS</collectionCode>
/ZSI/686,
<collectionCode id="F483B5FB246CFFCDFB658A4975EB7EFF" box="[1210,1262,364,390]" pageId="9" pageNumber="384">SVL</collectionCode>
<quantity id="556A80DB246CFFCDFB288A4E74657EFF" box="[1271,1376,363,390]" metricMagnitude="-2" metricUnit="m" metricValue="6.859999999999999" pageId="9" pageNumber="384" unit="mm" value="68.6">68.6 mm</quantity>
; VR/
<collectionCode id="F483B5FB246CFFCDFF488AAA71CE7ED0" box="[151,203,399,425]" pageId="9" pageNumber="384">ERS</collectionCode>
/ZSI/687,
<collectionCode id="F483B5FB246CFFCDFEE38AB570757ED3" box="[316,368,400,426]" pageId="9" pageNumber="384">SVL</collectionCode>
<quantity id="556A80DB246CFFCDFEA88AB570DA7ED3" box="[375,479,400,426]" metricMagnitude="-2" metricUnit="m" metricValue="7.029999999999999" pageId="9" pageNumber="384" unit="mm" value="70.3">70.3 mm</quantity>
); same collection details as
<typeStatus id="4D29939C246CFFCDFCC88AAA72787ED0" box="[791,893,399,425]" pageId="9" pageNumber="384" type="holotype">holotype</typeStatus>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DA887EB5246CFFCDFF188A96722F7A7F" pageId="9" pageNumber="384" type="diagnosis">
<paragraph id="922D2D3E246CFFCDFF188A9673357C69" blockId="9.[151,1437,256,2041]" pageId="9" pageNumber="384">
<emphasis id="A0E6F12C246CFFCDFF188A96723D7EB7" bold="true" box="[199,824,435,462]" pageId="9" pageNumber="384">Diagnosis and comparison with regional congeners.</emphasis>
<taxonomicName id="559256BD246CFFCDFC9E8A9675417EB4" authority="Purkayastha &amp; Das &amp; Bohra &amp; Bauer &amp; Agarwal, 2020" authorityName="Purkayastha &amp; Das &amp; Bohra &amp; Bauer &amp; Agarwal" authorityYear="2020" box="[833,1092,435,461]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="384" phylum="Chordata" rank="species" species="urbanus" status="sp. nov.">
<emphasis id="A0E6F12C246CFFCDFC9E8A9675417EB4" box="[833,1092,435,461]" italics="true" pageId="9" pageNumber="384">Cyrtodactylus urbanus</emphasis>
</taxonomicName>
<emphasis id="A0E6F12C246CFFCDFB928A9175AC7EB7" bold="true" box="[1101,1193,436,462]" pageId="9" pageNumber="384">
<taxonomicNameLabel id="BBD54C57246CFFCDFB928A9175AC7EB7" box="[1101,1193,436,462]" pageId="9" pageNumber="384" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished from regional congeners by its moderate body size (SVL to 74.0 mm); 911 supralabials; 810 infralabials; 2124 longitudinal rows of bluntly conical, feebly keeled dorsal tubercles; tubercles extending beyond base of tail; 3845 paravertebral tubercles; 3034 mid-ventral scales; no precloacal groove; 912 precloacal pores in a contiguous series, no femoral pores or pits; a row of enlarged scales above and below precloacal pore-bearing scales, slightly larger than pore-bearing scales;. There are 56 basal lamellae, 14 broken lamellae and 810 distal lamellae under 4
<superScript id="65E78076246CFFCDFF7A89AC71B77DEE" attach="left" box="[165,178,649,663]" fontSize="6" pageId="9" pageNumber="384">th</superScript>
finger and 67 basal lamellae, 23 broken lamellae and 1012 distal lamellae under 4
<superScript id="65E78076246CFFCDFBA689AC75837DEE" attach="left" box="[1145,1158,649,663]" fontSize="6" pageId="9" pageNumber="384">th</superScript>
toe. Subcaudal scalation of original tail without enlarged plates. Dorsal pattern of 67 longitudinal rows of two indistinct transverse pale buff blotches outlined by light edges, light mid-dorsal line. Tail with alternating dark and lighter bands. Iris in life silver-grey with brown reticulations.
</paragraph>
<paragraph id="922D2D3E246CFFCDFF18883F722F7A7F" blockId="9.[151,1437,256,2041]" pageId="9" pageNumber="384">
<taxonomicName id="559256BD246CFFCDFF18883F70CF7C4D" authority="Purkayastha &amp; Das &amp; Bohra &amp; Bauer &amp; Agarwal, 2020" authorityName="Purkayastha &amp; Das &amp; Bohra &amp; Bauer &amp; Agarwal" authorityYear="2020" box="[199,458,794,820]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="384" phylum="Chordata" rank="species" species="urbanus" status="sp. nov.">
<emphasis id="A0E6F12C246CFFCDFF18883F70CF7C4D" box="[199,458,794,820]" italics="true" pageId="9" pageNumber="384">Cyrtodactylus urbanus</emphasis>
</taxonomicName>
<emphasis id="A0E6F12C246CFFCDFE0C883F732A7C4D" bold="true" box="[467,559,794,820]" pageId="9" pageNumber="384">
<taxonomicNameLabel id="BBD54C57246CFFCDFE0C883F732A7C4D" box="[467,559,794,820]" pageId="9" pageNumber="384" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be differentiated from other members of the
<taxonomicName id="559256BD246CFFCDFBB5883E74047C4D" box="[1130,1281,794,821]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="384" phylum="Chordata" rank="species" species="khasiensis">
<emphasis id="A0E6F12C246CFFCDFBB5883E74047C4D" box="[1130,1281,794,821]" italics="true" pageId="9" pageNumber="384">C. khasiensis</emphasis>
</taxonomicName>
group by the following characters (see
<tableCitation id="DF101885246CFFCDFE1F881B731D7C21" box="[448,536,829,856]" captionStart="TABLE 5" captionStartId="16.[151,239,154,178]" captionTargetBox="[166,1419,189,509]" captionText="TABLE 5. Diagnostic characters for members of the Cyrtodactylus khasiensis complex. Abbreviations as in methods. PcP/ PcFP MVSR PVT DTR Transverse series of Middorsal stripe blotches on the dorsum" httpUri="http://table.plazi.org/id/C6ED7DB62475FFD4FF488BBF74857FCB" pageId="9" pageNumber="384" tableUuid="C6ED7DB62475FFD4FF488BBF74857FCB">Table 5</tableCitation>
): 912 precloacal pores in males (
<emphasis id="A0E6F12C246CFFCDFC73881A72F17C21" box="[940,1012,831,856]" italics="true" pageId="9" pageNumber="384">versus</emphasis>
1028 precloacofemoral pores in
<taxonomicName id="559256BD246CFFCDFA5C881B70537C05" authorityName="Bauer" authorityYear="2003" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="384" phylum="Chordata" rank="species" species="ayeyarwadyensis">
<emphasis id="A0E6F12C246CFFCDFA5C881B70537C05" italics="true" pageId="9" pageNumber="384">C. ayeyarwadyensis</emphasis>
</taxonomicName>
; 2639 precloacofemoral pores which may be interrupted by up to 11 unpored scales in
<taxonomicName id="559256BD246CFFCDFAEA884771FE7CD9" authorityName="Purkayastha &amp; Das &amp; Bohra &amp; Bauer &amp; Agarwal" authorityYear="2020" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="384" phylum="Chordata" rank="species" species="guwahatiensis">
<emphasis id="A0E6F12C246CFFCDFAEA884771FE7CD9" italics="true" pageId="9" pageNumber="384">C. guwahatiensis</emphasis>
</taxonomicName>
; 2937 precloacofemoral pores in
<taxonomicName id="559256BD246CFFCDFDA188A372207CE6" box="[638,805,902,928]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="384" phylum="Chordata" rank="species" species="tripuraensis">
<emphasis id="A0E6F12C246CFFCDFDA188A372207CE6" box="[638,805,902,928]" italics="true" pageId="9" pageNumber="384">C. tripuraensis</emphasis>
</taxonomicName>
) and 3034 MVSR (
<emphasis id="A0E6F12C246CFFCDFBD188A375537CE6" box="[1038,1110,902,927]" italics="true" pageId="9" pageNumber="384">versus</emphasis>
<quantity id="556A80DB246CFFCDFB8388A375BB7CD9" box="[1116,1214,902,928]" metricMagnitude="0" metricUnit="m" metricValue="1.016" metricValueMax="1.0922" metricValueMin="0.9398" pageId="9" pageNumber="384" unit="in" value="40.0" valueMax="43.0" valueMin="37.0">3743 in</quantity>
<taxonomicName id="559256BD246CFFCDFB1B88A374897CD9" authorityName="Agarwal, Mahony, Giri, Chaitanya &amp; Bauer" authorityYear="2018" box="[1220,1420,902,928]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="384" phylum="Chordata" rank="species" species="kazirangaensis">
<emphasis id="A0E6F12C246CFFCDFB1B88A374897CD9" box="[1220,1420,902,928]" italics="true" pageId="9" pageNumber="384">C. kazirangaensis</emphasis>
</taxonomicName>
). The new species may be distinguished from
<taxonomicName id="559256BD246CFFCDFD4C888F725E7CBA" authorityName="Agarwal, Mahony, Giri, Chaitanya &amp; Bauer" authorityYear="2018" box="[659,859,937,964]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="384" phylum="Chordata" rank="species" species="septentrionalis">
<emphasis id="A0E6F12C246CFFCDFD4C888F725E7CBA" box="[659,859,937,964]" italics="true" pageId="9" pageNumber="384">C. septentrionalis</emphasis>
</taxonomicName>
by a slightly lower precloacal pore count in males (912
<emphasis id="A0E6F12C246CFFCDFF3E88EB702C7C9E" box="[225,297,974,999]" italics="true" pageId="9" pageNumber="384">versus</emphasis>
1314), more subdigital lamellae beneath toe IV of the pes (1921
<emphasis id="A0E6F12C246CFFCDFBE988EB757B7C9E" box="[1078,1150,974,999]" italics="true" pageId="9" pageNumber="384">versus</emphasis>
1520) and fewer scales across the belly (3034
<emphasis id="A0E6F12C246CFFCDFE7888D770EA7B72" box="[423,495,1010,1035]" italics="true" pageId="9" pageNumber="384">versus</emphasis>
3538). Furthermore,
<taxonomicName id="559256BD246CFFCDFD2F88D472F47B72" authority="Purkayastha &amp; Das &amp; Bohra &amp; Bauer &amp; Agarwal, 2020" authorityName="Purkayastha &amp; Das &amp; Bohra &amp; Bauer &amp; Agarwal" authorityYear="2020" box="[752,1009,1009,1035]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="384" phylum="Chordata" rank="species" species="urbanus" status="sp. nov.">
<emphasis id="A0E6F12C246CFFCDFD2F88D472F47B72" box="[752,1009,1009,1035]" italics="true" pageId="9" pageNumber="384">Cyrtodactylus urbanus</emphasis>
</taxonomicName>
<emphasis id="A0E6F12C246CFFCDFC2788D475567B72" bold="true" box="[1016,1107,1009,1035]" pageId="9" pageNumber="384">
<taxonomicNameLabel id="BBD54C57246CFFCDFC2788D475567B72" box="[1016,1107,1009,1035]" pageId="9" pageNumber="384" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="559256BD246CFFCDFB3588D771CA7B56" authorityName="Bauer" authorityYear="2003" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="384" phylum="Chordata" rank="species" species="ayeyarwadyensis">
<emphasis id="A0E6F12C246CFFCDFB3588D771CA7B56" italics="true" pageId="9" pageNumber="384">C. ayeyarwadyensis</emphasis>
</taxonomicName>
in having a prominent mid-dorsal stripe (absent in
<taxonomicName id="559256BD246CFFCDFCCE8F3372F47B56" authorityName="Bauer" authorityYear="2003" box="[785,1009,1045,1072]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="384" phylum="Chordata" rank="species" species="ayeyarwadyensis">
<emphasis id="A0E6F12C246CFFCDFCCE8F3372F47B56" box="[785,1009,1045,1072]" italics="true" pageId="9" pageNumber="384">C. ayeyarwadyensis</emphasis>
</taxonomicName>
) and fewer transverse blotches on the dorsum (68
<emphasis id="A0E6F12C246CFFCDFEF28F1F70707B2A" box="[301,373,1082,1107]" italics="true" pageId="9" pageNumber="384">versus</emphasis>
911)
<taxonomicName id="559256BD246CFFCDFE1B8F1C73C07B2A" authority="Purkayastha &amp; Das &amp; Bohra &amp; Bauer &amp; Agarwal, 2020" authorityName="Purkayastha &amp; Das &amp; Bohra &amp; Bauer &amp; Agarwal" authorityYear="2020" box="[452,709,1081,1107]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="384" phylum="Chordata" rank="species" species="urbanus" status="sp. nov.">
<emphasis id="A0E6F12C246CFFCDFE1B8F1C73C07B2A" box="[452,709,1081,1107]" italics="true" pageId="9" pageNumber="384">Cyrtodactylus urbanus</emphasis>
</taxonomicName>
<emphasis id="A0E6F12C246CFFCDFD138F1C72237B2A" bold="true" box="[716,806,1081,1107]" pageId="9" pageNumber="384">
<taxonomicNameLabel id="BBD54C57246CFFCDFD138F1C72237B2A" box="[716,806,1081,1107]" pageId="9" pageNumber="384" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is very similar to
<taxonomicName id="559256BD246CFFCDFC2C8F1C75827B2A" box="[1011,1159,1081,1107]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="384" phylum="Chordata" rank="species" species="khasiensis">
<emphasis id="A0E6F12C246CFFCDFC2C8F1C75827B2A" box="[1011,1159,1081,1107]" italics="true" pageId="9" pageNumber="384">C. khasiensis</emphasis>
</taxonomicName>
, from which it differs in mean counts across a number of characters as follows:
<taxonomicName id="559256BD246CFFCDFCDB8F7875017B0E" authority="Purkayastha &amp; Das &amp; Bohra &amp; Bauer &amp; Agarwal, 2020" authorityName="Purkayastha &amp; Das &amp; Bohra &amp; Bauer &amp; Agarwal" authorityYear="2020" box="[772,1028,1117,1143]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="384" phylum="Chordata" rank="species" species="urbanus" status="sp. nov.">
<emphasis id="A0E6F12C246CFFCDFCDB8F7875017B0E" box="[772,1028,1117,1143]" italics="true" pageId="9" pageNumber="384">Cyrtodactylus urbanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD54C57246CFFCDFBD48F7875627B0E" box="[1035,1127,1117,1143]" pageId="9" pageNumber="384" rank="species">
<emphasis id="A0E6F12C246CFFCDFBD48F7875657B0E" bold="true" box="[1035,1120,1117,1143]" pageId="9" pageNumber="384">sp. nov</emphasis>
.
</taxonomicNameLabel>
has fewer paravertebral tubercles (3840
<emphasis id="A0E6F12C246CFFCDFE9E8FA7708C7BE2" box="[321,393,1154,1179]" italics="true" pageId="9" pageNumber="384">versus</emphasis>
<quantity id="556A80DB246CFFCDFE518FA570F57BE2" box="[398,496,1152,1179]" metricMagnitude="0" metricUnit="m" metricValue="1.2065" metricValueMax="1.3462" metricValueMin="1.0668" pageId="9" pageNumber="384" unit="in" value="47.5" valueMax="53.0" valueMin="42.0">4253 in</quantity>
<taxonomicName id="559256BD246CFFCDFE298FA4738D7BE2" box="[502,648,1153,1179]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="384" phylum="Chordata" rank="species" species="khasiensis">
<emphasis id="A0E6F12C246CFFCDFE298FA4738D7BE2" box="[502,648,1153,1179]" italics="true" pageId="9" pageNumber="384">C. khasiensis</emphasis>
</taxonomicName>
); fewer mid-ventral scale rows (3034
<emphasis id="A0E6F12C246CFFCDFBE38FA775817BE2" box="[1084,1156,1154,1179]" italics="true" pageId="9" pageNumber="384">versus</emphasis>
3442); and fewer apical subdigital lamellae (810
<emphasis id="A0E6F12C246CFFCDFE678F8073057BC7" box="[440,512,1189,1214]" italics="true" pageId="9" pageNumber="384">versus</emphasis>
1315). The tail pattern in the original tail of the new species has approximately 14 dark bands, the first six similar to spots on the dorsum, whereas the original tail of
<taxonomicName id="559256BD246CFFCDFBEB8FEC75CD7B9B" box="[1076,1224,1224,1251]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="384" phylum="Chordata" rank="species" species="khasiensis">
<emphasis id="A0E6F12C246CFFCDFBEB8FEC75CD7B9B" box="[1076,1224,1224,1251]" italics="true" pageId="9" pageNumber="384">C. khasiensis</emphasis>
</taxonomicName>
has approximately 11 dark bands, the first two similar to spots on the dorsum.
</paragraph>
</subSubSection>
<subSubSection id="DA887EB5246CFFCDFF188E3573BD7A0B" pageId="9" pageNumber="384" type="etymology">
<paragraph id="922D2D3E246CFFCDFF188E3573BD7A0B" blockId="9.[151,1437,256,2041]" pageId="9" pageNumber="384">
<emphasis id="A0E6F12C246CFFCDFF188E35704B7A53" bold="true" box="[199,334,1296,1322]" pageId="9" pageNumber="384">Etymology.</emphasis>
The specific epithet is an adjective meaning of or belonging to a city and reflects the urban habitat of the species, through which we also wish to highlight the importance of urban biodiversity. We propose Urban bent-toed gecko as common name for the gecko.
</paragraph>
</subSubSection>
<subSubSection id="DA887EB5246CFFC8FF188E5E750F7947" lastPageId="12" lastPageNumber="387" pageId="9" pageNumber="384" type="description">
<paragraph id="922D2D3E246CFFCFFF188E5E73727FAE" blockId="9.[151,1437,256,2041]" lastBlockId="11.[151,1437,153,611]" lastPageId="11" lastPageNumber="386" pageId="9" pageNumber="384">
<emphasis id="A0E6F12C246CFFCDFF188E5E70E37AEF" bold="true" box="[199,486,1403,1430]" pageId="9" pageNumber="384">
Description of
<typeStatus id="4D29939C246CFFCDFEA68E5970E77AEF" box="[377,482,1404,1430]" pageId="9" pageNumber="384" type="holotype">holotype</typeStatus>
.
</emphasis>
Adult male, SVL
<quantity id="556A80DB246CFFCDFD688E5972187AEF" box="[695,797,1404,1430]" metricMagnitude="-2" metricUnit="m" metricValue="7.19" pageId="9" pageNumber="384" unit="mm" value="71.9">71.9 mm</quantity>
, with both hemipenes everted. Head just more than onefifth of SVL (HL/SVL 0.23), wider than long (HW/HL 0.68), somewhat depressed (HD/HL 0.52), distinct from neck; loreal region somewhat inflated, interorbital area flat, canthus rostralis not prominent; snout slightly less than half of the head length (SE/HL 0.46), twice as long as orbit diameter (OD/SE 0.53); scales on forehead, canthus rostralis and snout homogeneous. Scales from posterior margin of eyes to nape smaller than those of forehead, roughly hexagonal, juxtaposed and weakly pointed; scales on interorbital and occipital regions heterogeneous, without distinct tubercles. Eye one quarter of head length (OD/HL 0.25); pupil vertical with crenulate margins; supraciliaries small, spiny, decreasing in size towards posterior end of orbit, those on the anterior end of orbit largest; ear opening oval, obliquely oriented, one-tenth of the head length (EL/HL 0.10); eye to ear distance about half eye diameter (OD/SE 0.54). Rostral wider than deep (RL/RW 0.60), partially divided dorsally by weakly developed rostral groove; single much enlarged supranasal on either side, separated by two small internasal, about the same size as enlarged scales on snout; rostral in contact with first supralabials, nasals, supranasals and an internasal; nostrils semicircular, laterally oriented, posterior half covered by nasal pad, each in broad contact with rostral and surrounded by supranasal, first supralabial, and three postnasals; two rows of scales separate orbit from supralabials; mental wider (
<quantity id="556A80DB246CFFCDFADB8C6B74597810" box="[1284,1372,1870,1897]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="9" pageNumber="384" unit="mm" value="2.1">2.1 mm</quantity>
) than long (
<quantity id="556A80DB246CFFCDFF068C54703078F5" box="[217,309,1905,1932]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="9" pageNumber="384" unit="mm" value="1.5">1.5 mm</quantity>
), triangular; two well developed postmentals on either side, inner pair more than twice the size (
<quantity id="556A80DB246CFFCDFAA78C5771C178C9" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="9" pageNumber="384" unit="mm" value="2.6">2.6 mm</quantity>
) of and separating outer pair (1.0 mm), in moderate contact (
<quantity id="556A80DB246CFFCDFC5D8CB372D978C9" box="[898,988,1942,1968]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="9" pageNumber="384" unit="mm" value="1.2">1.2 mm</quantity>
) behind mental; inner postmentals bordered by mental, infralabial I, outer postmental and six gular scales; outer postmental bordered by inner postmental, infralabial I and II, and four gular scales on either side; supralabials (to midorbital position) 10 (right) 9 (left), bordered by a row of large, flat, somewhat elongate scales; infralabials 9, infralabials I to III bordered ventrally by a row of enlarged scales, largest anteriorly.
</paragraph>
<caption id="C6ED7DB6246FFFCEFF488C8F70D1789E" ID-DOI="http://doi.org/10.5281/zenodo.3667219" ID-Zenodo-Dep="3667219" httpUri="https://zenodo.org/record/3667219/files/figure.png" pageId="10" pageNumber="385" startId="10.[151,250,1962,1987]" targetBox="[168,1419,174,1943]" targetPageId="10">
<paragraph id="922D2D3E246FFFCEFF488C8F70D1789E" blockId="10.[151,1436,1962,2023]" pageId="10" pageNumber="385">
<emphasis id="A0E6F12C246FFFCEFF488C8F701678BA" bold="true" box="[151,275,1962,1987]" pageId="10" pageNumber="385">FIGURE 5.</emphasis>
A: Dorsal view, B: ventral view and C: details of dorsal pholidosis of
<taxonomicName id="559256BD246FFFCEFC3F8C8E75CD78BA" authority="Purkayastha &amp; Das &amp; Bohra &amp; Bauer &amp; Agarwal, 2020" authorityName="Purkayastha &amp; Das &amp; Bohra &amp; Bauer &amp; Agarwal" authorityYear="2020" box="[992,1224,1963,1987]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="385" phylum="Chordata" rank="species" species="urbanus" status="sp. nov.">
<emphasis id="A0E6F12C246FFFCEFC3F8C8E75CD78BA" box="[992,1224,1963,1987]" italics="true" pageId="10" pageNumber="385">Cyrtodactylus urbanus</emphasis>
</taxonomicName>
<emphasis id="A0E6F12C246FFFCEFB108C8E741A78BA" bold="true" box="[1231,1311,1963,1987]" pageId="10" pageNumber="385">
<taxonomicNameLabel id="BBD54C57246FFFCEFB108C8E741A78BA" box="[1231,1311,1963,1987]" pageId="10" pageNumber="385" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(male holotype, VR/ERS/ZSI/683) in life.
</paragraph>
</caption>
<paragraph id="922D2D3E246EFFCFFF188BC471F47D1A" blockId="11.[151,1437,153,611]" pageId="11" pageNumber="386">Body moderately slender, trunk slightly less than half of SVL (TRL/SVL 0.43) with weakly developed ventrolateral folds that are not denticulate; dorsal scales heterogeneous, mostly rounded granules, intermixed with irregularly arranged, enlarged tubercles (24 times size of granules), bluntly conical and feebly keeled throughout, becoming more conical and slightly smaller towards flank, largest on the sacral region, ventrolateral folds with single row of scattered enlarged, smooth tubercles; tubercles extend from occipital region beyond the tail base; tubercles on nape smaller than those of dorsum; tail flat, weakly pointed and feebly keeled; 22 rows of dorsal tubercles; 45 paravertebral tubercles; ventral scales much larger than dorsals, smooth, cycloid, imbricate to subimbricate; slightly smaller in size under thighs, no enlarged scales between precloacal pores and vent; scales across the belly 31; gular region with small granular scales throughout except a few rows bordering mental, postmentals and infralabials larger, flat and juxtaposed. Nine distinct precloacal pores in a continuous series, precloacal grooves absent, hemipenal bulge distinct.</paragraph>
<caption id="C6ED7DB6246EFFCFFF488E51701E7AC8" ID-DOI="http://doi.org/10.5281/zenodo.3667221" ID-Zenodo-Dep="3667221" httpUri="https://zenodo.org/record/3667221/files/figure.png" pageId="11" pageNumber="386" startId="11.[151,250,1396,1421]" targetBox="[151,1436,641,1369]" targetPageId="11">
<paragraph id="922D2D3E246EFFCFFF488E51701E7AC8" blockId="11.[151,1437,1396,1457]" pageId="11" pageNumber="386">
<emphasis id="A0E6F12C246EFFCFFF488E5170167AF4" bold="true" box="[151,275,1396,1421]" pageId="11" pageNumber="386">FIGURE 6.</emphasis>
Head of
<taxonomicName id="559256BD246EFFCFFEAE8E50735D7AF4" authority="Purkayastha &amp; Das &amp; Bohra &amp; Bauer &amp; Agarwal, 2020" authorityName="Purkayastha &amp; Das &amp; Bohra &amp; Bauer &amp; Agarwal" authorityYear="2020" box="[369,600,1397,1421]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="386" phylum="Chordata" rank="species" species="urbanus" status="sp. nov.">
<emphasis id="A0E6F12C246EFFCFFEAE8E50735D7AF4" box="[369,600,1397,1421]" italics="true" pageId="11" pageNumber="386">Cyrtodactylus urbanus</emphasis>
</taxonomicName>
<emphasis id="A0E6F12C246EFFCFFD808E5073AA7AF4" bold="true" box="[607,687,1397,1421]" pageId="11" pageNumber="386">
<taxonomicNameLabel id="BBD54C57246EFFCFFD808E5073AA7AF4" box="[607,687,1397,1421]" pageId="11" pageNumber="386" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(male holotype, VR/ERS/ZSI/683), A: dorsal view B: lateral view and C: ventral view.
</paragraph>
</caption>
<paragraph id="922D2D3E246EFFCFFF188EC5736B78B4" blockId="11.[151,1437,1502,1998]" pageId="11" pageNumber="386">Forearm (FL/SVL 0.15) and tibia (CL/SVL 0.17) short; digits relatively short, strongly inflected at each joint, all bearing robust, recurved claws; subdigital lamellae widened beneath basal phalanx; basal lamellae 45664 (right manus) and 45575 (right pes); distal lamellae (intervening rows of nonlamellar granules between basal and distal lamellae series in parentheses): 6(1)8(2)10(3)9(4)9(2) (right manus) and 7(2)9(2)10(2)11(2) 10(2) (right pes); interdigital webbing absent from both manus and pes; relative length of digits: I &lt;V &lt;II &lt;III &lt;IV (right manus) and I &lt;II &lt;V &lt;III &lt;VI (right pes); scales on the palm and sole smooth, weakly raised, subimbricate; scales on forelimb heterogeneous, comprising flat, subimbricate scales on upper arm, those on forearm heterogeneous, ventral portion covered heterogenous imbricate scales; scales on hindlimbs heterogeneous, dorsal part of thigh and shank with larger scales, intermixed with scattered, enlarged, conical, feebly keeled tubercles; anterior portion of thighs and ventral aspect of hindlimbs with much enlarged, smooth, imbricate scales. Tail original, complete, rounded in cross-section, slender, tapering; Two rows of enlarged, flat, weakly pointed, smooth, tubercles positioned paravertebrally on tail base only, remaining dorsal caudal scales smooth, rounded, subimbricate, similar in size dorsally but becoming larger on lateral aspect; subcaudal scales larger, smooth, imbricate with midventral series of paired scales; no enlarged plates.</paragraph>
<paragraph id="922D2D3E2469FFC8FF188BBD73037ED6" blockId="12.[151,1437,152,431]" pageId="12" pageNumber="387">
<emphasis id="A0E6F12C2469FFC8FF188BBD730D7FCA" bold="true" box="[199,520,152,179]" pageId="12" pageNumber="387">
Colouration in life (
<figureCitation id="0AA931BB2469FFC8FE6C8BBD70FC7FCA" box="[435,505,152,179]" captionStart="FIGURE 5" captionStartId="10.[151,250,1962,1987]" captionTargetBox="[168,1419,174,1943]" captionTargetId="figure@10.[168,1419,174,1943]" captionTargetPageId="10" captionText="FIGURE 5. A: Dorsal view, B: ventral view and C: details of dorsal pholidosis of Cyrtodactylus urbanus sp. nov. (male holotype, VR/ERS/ZSI/683) in life." figureDoi="http://doi.org/10.5281/zenodo.3667219" httpUri="https://zenodo.org/record/3667219/files/figure.png" pageId="12" pageNumber="387">Fig. 5</figureCitation>
).
</emphasis>
Dorsal ground colouration dark brown. Head has dark brown ground colour with faint yellow patches behind the nasal scales as well as in the supraciliary region. Nape has few cream coloured spots on brown ground colour. The dorsal part of trunk region shows alternating dark and light pattern, a mid-dorsal cream coloured line divides the trunk and extends posteriorly to the level of the cloaca. The dorsal markings are “w” shaped. The same markings continue onto the tail extending to the tail tip, reducing, however, to simple transverse bars on the distal portion of the tail. The caudal region has 14 dark bands alternating with lighter ones. The thigh and hind limbs show indistinct cream coloured blotches or crossbars alternating with dark brown ground colour. Similar pattern observed in the forelimb.
</paragraph>
<caption id="C6ED7DB62469FFC8FF488E20703C7A3B" ID-DOI="http://doi.org/10.5281/zenodo.3667223" ID-Zenodo-Dep="3667223" httpUri="https://zenodo.org/record/3667223/files/figure.png" pageId="12" pageNumber="387" startId="12.[151,250,1285,1310]" targetBox="[151,1436,454,1249]" targetPageId="12">
<paragraph id="922D2D3E2469FFC8FF488E20703C7A3B" blockId="12.[151,1437,1285,1346]" pageId="12" pageNumber="387">
<emphasis id="A0E6F12C2469FFC8FF488E2070167A67" bold="true" box="[151,275,1285,1310]" pageId="12" pageNumber="387">FIGURE 7.</emphasis>
Fully everted hemipenis of
<taxonomicName id="559256BD2469FFC8FDF08E2372127A67" authority="Purkayastha &amp; Das &amp; Bohra &amp; Bauer &amp; Agarwal, 2020" authorityName="Purkayastha &amp; Das &amp; Bohra &amp; Bauer &amp; Agarwal" authorityYear="2020" box="[559,791,1286,1310]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="387" phylum="Chordata" rank="species" species="urbanus" status="sp. nov.">
<emphasis id="A0E6F12C2469FFC8FDF08E2372127A67" box="[559,791,1286,1310]" italics="true" pageId="12" pageNumber="387">Cyrtodactylus urbanus</emphasis>
</taxonomicName>
<emphasis id="A0E6F12C2469FFC8FCC28E23726B7A67" bold="true" box="[797,878,1286,1310]" pageId="12" pageNumber="387">
<taxonomicNameLabel id="BBD54C572469FFC8FCC28E23726B7A67" box="[797,878,1286,1310]" pageId="12" pageNumber="387" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(male holotype, VR/ERS/ZSI/683), A: sulcal view and B: asulcal view.
</paragraph>
</caption>
<paragraph id="922D2D3E2469FFC8FF188E55750F7947" blockId="12.[151,1436,1392,1922]" pageId="12" pageNumber="387">
<emphasis id="A0E6F12C2469FFC8FF188E55704B7AF2" bold="true" box="[199,334,1392,1419]" pageId="12" pageNumber="387">Hemipenis.</emphasis>
The hemipenis of the
<typeStatus id="4D29939C2469FFC8FD948E5573A87AF3" box="[587,685,1392,1418]" pageId="12" pageNumber="387" type="holotype">holotype</typeStatus>
of
<taxonomicName id="559256BD2469FFC8FD0C8E5472547AF3" authority="Purkayastha &amp; Das &amp; Bohra &amp; Bauer &amp; Agarwal, 2020" authorityName="Purkayastha &amp; Das &amp; Bohra &amp; Bauer &amp; Agarwal" authorityYear="2020" box="[723,849,1392,1419]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="387" phylum="Chordata" rank="species" species="urbanus" status="sp. nov.">
<emphasis id="A0E6F12C2469FFC8FD0C8E5472547AF3" box="[723,849,1392,1419]" italics="true" pageId="12" pageNumber="387">C. urbanus</emphasis>
</taxonomicName>
<emphasis id="A0E6F12C2469FFC8FC878E5472B77AF2" bold="true" box="[856,946,1393,1419]" pageId="12" pageNumber="387">
<taxonomicNameLabel id="BBD54C572469FFC8FC878E5472B77AF2" box="[856,946,1393,1419]" pageId="12" pageNumber="387" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is bilobed, elongated and muscular, but the lobes are not clearly distinguishable or demarcated. The apex seems bulbous and almost round in shape with no ornamentation on the sulcal surface. The length of the lobe of the hemipenis is almost 1/4
<superScript id="65E780762469FFC8FB818E92756E7ABC" attach="left" box="[1118,1131,1463,1477]" fontSize="6" pageId="12" pageNumber="387">th</superScript>
of its total length (hemepenis length=
<quantity id="556A80DB2469FFC8FEC38EF970707A8E" box="[284,373,1500,1527]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="12" pageNumber="387" unit="mm" value="5.2">5.2 mm</quantity>
, lobe length=
<quantity id="556A80DB2469FFC8FDCF8EF9736C7A8E" box="[528,617,1500,1527]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="12" pageNumber="387" unit="mm" value="2.1">2.1 mm</quantity>
). The sulcus spermaticus is clearly bifurcated, goes around the lobe head before entering it. The sulcus spermaticus is deeply grooved. The asulcal surface has mildly rough, granules present on the lobe head. The pedicel is unadorned on both sulcal and asulcal surfaces.
</paragraph>
</subSubSection>
<subSubSection id="DA887EB52469FFC8FF188D6D738678FB" pageId="12" pageNumber="387" type="distribution">
<paragraph id="922D2D3E2469FFC8FF188D6D738678FB" blockId="12.[151,1436,1392,1922]" pageId="12" pageNumber="387">
<emphasis id="A0E6F12C2469FFC8FF188D6D735F791A" bold="true" box="[199,602,1608,1635]" pageId="12" pageNumber="387">Distribution and Natural History.</emphasis>
This species is only known from degraded moist deciduous forests in and around Guwahati,
<collectingRegion id="5056E3DC2469FFC8FEBA8D4870B079FE" box="[357,437,1645,1671]" country="India" name="Assam" pageId="12" pageNumber="387">Assam</collectingRegion>
; based on voucher specimens from the
<typeStatus id="4D29939C2469FFC8FCB58D48729F79FE" box="[874,922,1645,1671]" pageId="12" pageNumber="387">type</typeStatus>
locality, Basistha, and photographic vouchers from Deeporbeel, Garbhanga Reserve Forest, Gotanagar and Rani Reserve Forest. The
<typeStatus id="4D29939C2469FFC8FB5F8DB475B579D2" box="[1152,1200,1681,1707]" pageId="12" pageNumber="387">type</typeStatus>
series was collected from around the Basistha Temple in Guwahati (
<figureCitation id="0AA931BB2469FFC8FD678D9173F879B7" box="[696,765,1716,1742]" captionStart="FIGURE 9" captionStartId="15.[151,250,1536,1561]" captionTargetBox="[151,1436,553,1517]" captionTargetId="figure@15.[151,1436,553,1517]" captionTargetPageId="15" captionText="FIGURE 9. Habitat of Cyrtodactylus urbanus sp. nov., Basistha, Guwahati, Assam" figureDoi="http://doi.org/10.5281/zenodo.3667227" httpUri="https://zenodo.org/record/3667227/files/figure.png" pageId="12" pageNumber="387">Fig. 9</figureCitation>
) on
<date id="E62C0BFE2469FFC8FCEC8D9172C979B7" box="[819,972,1716,1743]" pageId="12" pageNumber="387" value="2018-06-13">13 June 2018</date>
between 1900h and 2100h. The animals were seen to be active between the months of March and November. Being a nocturnal species, they are most active two to five hours after dark and are mostly seen inside rock crevices and small rocky caves which are associated with running hill streams. Gravid females were observed from mid-April to early August. The diet of this species primarily consists of spiders, crickets and other small arthropods. In one instance we also found semi-digested ecdysed skin of
<taxonomicName id="559256BD2469FFC8FEEB8C4D70D478FB" authorityName="Gray" authorityYear="1827" box="[308,465,1896,1922]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="387" phylum="Chordata" rank="genus">
<emphasis id="A0E6F12C2469FFC8FEEB8C4D70D478FB" box="[308,465,1896,1922]" italics="true" pageId="12" pageNumber="387">Cyrtodactylus</emphasis>
</taxonomicName>
in the stomach.
</paragraph>
</subSubSection>
</treatment>
</document>