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<mods:title id="443308100D45BC053B510B1ADCF869C4">A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico</mods:title>
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<mods:namePart id="EF22258B9A34BC96D4BF68BB7971008A">Luna-Cozar, Jesús</mods:namePart>
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<mods:namePart id="5FA925E05195BA7E5F0EA5C5049B927D">Jones, Robert W.</mods:namePart>
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<taxonomicName id="8DD74D6AFF9E6109FF14DE0E25E3FA90" ID-CoL="8HSKH" authority="Champion 1905" authorityName="Champion" authorityYear="1905" box="[151,476,1340,1366]" class="Insecta" family="Curculionidae" genus="Tylodinus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="78A3EAFBFF9E6109FF14DE0E25E3FA90" bold="true" box="[151,476,1340,1366]" pageId="5" pageNumber="6">
<emphasis id="78A3EAFBFF9E6109FF14DE0E2531FA90" bold="true" box="[151,270,1340,1366]" italics="true" pageId="5" pageNumber="6">Tylodinus</emphasis>
<bibRefCitation id="2E464B18FF9E6109FE96DE0E25E3FA90" author="Champion, C. G." box="[277,476,1340,1366]" pageId="5" pageNumber="6" pagination="441 - 600" refId="ref56589" refString="Champion, C. G. (1905) Biologia Centrali-Americana. Insecta. Coleoptera. Rhynchophora. Curculionidae. Curculioninae. Vol. 4. Part 4. pp. 441 - 600." type="book chapter" year="1905">Champion 1905</bibRefCitation>
</emphasis>
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</paragraph>
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<treatmentCitation id="CB7610F8FF9E6109FF14DEB025F9FA51" author="Champion, C. G." box="[151,454,1410,1431]" page="464" pageId="5" pageNumber="6" year="1905">
<taxonomicName id="8DD74D6AFF9E6109FF14DEB025F9FA51" ID-CoL="8HSKH" authority="Champion, 1905: 464" authorityName="Champion" authorityPageNumber="464" authorityYear="1905" box="[151,454,1410,1431]" class="Insecta" family="Curculionidae" genus="Tylodinus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="78A3EAFBFF9E6109FF14DEB024C9FA51" box="[151,246,1410,1431]" italics="true" pageId="5" pageNumber="6">Tylodinus</emphasis>
<bibRefCitation id="2E464B18FF9E6109FF7ADEB025F9FA51" author="Champion, C. G." box="[249,454,1410,1431]" pageId="5" pageNumber="6" pagination="441 - 600" refId="ref56589" refString="Champion, C. G. (1905) Biologia Centrali-Americana. Insecta. Coleoptera. Rhynchophora. Curculionidae. Curculioninae. Vol. 4. Part 4. pp. 441 - 600." type="book chapter" year="1905">Champion, 1905:464</bibRefCitation>
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; Fiedler 1940: 299–300; 
<treatmentCitation id="CB7610F8FF9E6109FD35DEB027AFFA51" author="Blackwelder, R. E." box="[694,912,1410,1431]" page="860" pageId="5" pageNumber="6" year="1947">
<bibRefCitation id="2E464B18FF9E6109FD35DEB027AFFA51" author="Blackwelder, R. E." box="[694,912,1410,1431]" pageId="5" pageNumber="6" pagination="765 - 925" refId="ref56413" refString="Blackwelder, R. E. (1947) Checklist of the coleopterous insects of Mexico, Central America, the West Indies and South America. Part. 5. Smithsonian Institution United States National Museum Bulletin, (185), I - IV, 765 - 925." type="journal article" year="1947">Blackwelder 1947:860</bibRefCitation>
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(checklist); 
<treatmentCitation id="CB7610F8FF9E6109FB84DEB02083FA51" author="Kuschel, G." box="[1031,1212,1410,1431]" page="323" pageId="5" pageNumber="6" year="1956">
<bibRefCitation id="2E464B18FF9E6109FB84DEB02083FA51" author="Kuschel, G." box="[1031,1212,1410,1431]" pageId="5" pageNumber="6" pagination="319 - 339" refId="ref56862" refString="Kuschel, G. (1956) Attelabidae und Curculionidae aus El Salvador. (Ins. Col. Curculionoidea, 21. Beitrag). Senckenbergiana Biologica, 37 (3 - 4), 319 - 339." type="journal article" year="1956">Kuschel 1956: 323</bibRefCitation>
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(new species); 
<treatmentCitation id="CB7610F8FF9E6109FAD2DEB0259DFA7C" author="O'Brien, C. W. &amp; Wibmer, G. J." page="139" pageId="5" pageNumber="6" year="1982">
<bibRefCitation id="2E464B18FF9E6109FAD2DEB0259DFA7C" author="O'Brien, C. W. &amp; Wibmer, G. J." pageId="5" pageNumber="6" pagination="1 - 382" refId="ref56933" refString="O'Brien, C. W. &amp; Wibmer, G. J. (1982) Annotated checklist of the weevils (Curculionidae sensu lato) of North America, Central America and the West Indies (Coleoptera: Curculionidae). Memoirs of the American Entomological Institute, 34, i - ix, 1 - 382." type="journal article" year="1982">O'Brien and Wibmer 1982:139</bibRefCitation>
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(checklist); Alonso-Zarazaga &amp; Lyal 1999:134 (catalog).
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</paragraph>
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<emphasis id="78A3EAFBFF9E6109FF14DED8250DF9C5" bold="true" box="[151,306,1514,1539]" pageId="5" pageNumber="6">
<typeStatus id="956C884BFF9E6109FF14DED824EDF9C5" box="[151,210,1514,1539]" pageId="5" pageNumber="6">Type</typeStatus>
species.
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<taxonomicName id="8DD74D6AFF9E6109FEBADED826D6F9C5" ID-CoL="8R2M" authority="Boheman 1837: 330" authorityName="Boheman" authorityPageNumber="330" authorityYear="1837" box="[313,745,1514,1539]" class="Insecta" family="Curculionidae" genus="Acalles" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="nodulosus">
<emphasis id="78A3EAFBFF9E6109FEBADED8263BF9C5" box="[313,516,1514,1539]" italics="true" pageId="5" pageNumber="6">Acalles nodulosus</emphasis>
<bibRefCitation id="2E464B18FF9E6109FD88DED826D6F9C5" author="Boheman, C. H. &amp; Schoenherr, C. J. &amp; Gyllenhaal, L." box="[523,745,1514,1539]" pageId="5" pageNumber="6" refId="ref56464" refString="Boheman, C. H., Schoenherr, C. J. &amp; Gyllenhaal, L. (1837) Genera et species curculionidum, cum synonymia hujus familiae. Species novae aut hactenus minus cognitae; descriptionibus a Dom L. Gyllenhal, C. H. Boheman, O. J. Fahraeus et entomologis aliis illustratae. Vol. 4. Part 1. Roret, 600 pp." type="book" year="1837">Boheman 1837:330</bibRefCitation>
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, by original designation. Gender masculine.
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<emphasis id="78A3EAFBFF9E6109FF44DD3C257EF9E1" bold="true" box="[199,321,1550,1575]" pageId="5" pageNumber="6">Diagnosis.</emphasis>
Distinguishable among 
<taxonomicName id="8DD74D6AFF9E6109FDD8DD3C2681F9E1" box="[603,702,1550,1575]" class="Gastropoda" family="Tylodinidae" kingdom="Animalia" order="Umbraculida" pageId="5" pageNumber="8" phylum="Mollusca" rank="subTribe" subTribe="Tylodina">Tylodina</taxonomicName>
by the metepisternum distinctly visible; the elytra clothed with elongate scales, moderately dense to sparse, more dense on the middle of disc and on the base, becoming more scattered toward the flanks; intervals weakly convex, less convex to flat toward the flanks; striae shallow, tubercles round or laminate; internal sac of male genitalia with diminutive round and acute sclerites.
</paragraph>
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<subSubSection id="02CD6562FF9E610BFF44DDAC25E1FD46" lastPageId="7" lastPageNumber="8" pageId="5" pageNumber="6" type="description">
<paragraph id="4A6836E9FF9E610BFF44DDAC25E1FD46" blockId="5.[151,1437,1514,2029]" lastBlockId="7.[151,1437,151,1685]" lastPageId="7" lastPageNumber="8" pageId="5" pageNumber="6">
<emphasis id="78A3EAFBFF9E6109FF44DDAC2567F971" bold="true" box="[199,344,1694,1719]" pageId="5" pageNumber="6">Description.</emphasis>
Length 2.0– 
<quantity id="8D2F9B0CFF9E6109FE64DDAC2674F970" box="[487,587,1694,1719]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" pageId="5" pageNumber="6" unit="mm" value="11.5">11.5 mm</quantity>
. Width 0.9–7.0 mm. Body oblong-ovate, 1.8–2.7x longer than wide, clothed with elongate scales, short or long. 
<emphasis id="78A3EAFBFF9E6109FDC0DDF726BCF918" bold="true" box="[579,643,1733,1758]" pageId="5" pageNumber="6">Head</emphasis>
with red, dark red or black integument; scales on vertex dense to contiguous or overlapping, same color as head scales, yellow, light brown or whitish; frons concave to moderately concave; rostrum moderately robust or narrow, surface moderately carinate to rugose, punctures small or large, deep or shallow, longer or as long as head punctures, with an apical area glabrous (male with apical 1/2, female with apical 2/3 (
<figureCitation id="D2EC2A6CFF9E6109FEDEDC6D25EDF8BE" box="[349,466,1887,1912]" captionStart="FIGURES 12–19. 12" captionStartId="9.[151,264,1919,1941]" captionTargetBox="[219,1377,209,1898]" captionTargetId="figure-86@9.[219,1377,197,1898]" captionTargetPageId="9" captionText="FIGURES 12–19. 12. Tylodinus pusillus head a) male, b) female. 13–14. Dorsal habitus of pronotum showing the lateral side of pronotum 13. Tylodinus andersoni, 14. Tylodinus porvenirensis. 15–17. Dorsal habitus: 15. Tylodinus rugosus, 16. Tylodinus exiguus, 17. Tylodinus jonesi. 18. Tylodinus coapillensis note tubercle on elytra declivity base, 19. Tylodinus elongatus showing metasternal tooth." figureDoi="http://doi.org/10.5281/zenodo.4913765" httpUri="https://zenodo.org/record/4913765/files/figure.png" pageId="5" pageNumber="6">Figure 12</figureCitation>
)), puncticulate (very fine and widely spaced punctures), punctate (fine impressed punctures) or smooth; antennal scape in resting position not reaching anterior margin of eye, funicle with articles 1 and 2 subequal in length, articles 3–4 slightly longer than wide, articles 5–7 submoniliform; eyes separated by about the width of an eye, moderately convex or flush with outline of head. 
<emphasis id="78A3EAFBFF9E6109FBC7DCE62082F82B" bold="true" box="[1092,1213,2004,2029]" pageId="5" pageNumber="6">Pronotum</emphasis>
with red or black integument; in lateral view with one of three patterns: 1) outline with anterior constriction and without posterior constriction (
<figureCitation id="D2EC2A6CFF9D610AFEAFDB8C259FFF11" box="[300,416,190,215]" captionStart="FIGURES 1–11. 1–5" captionStartId="8.[151,264,1743,1765]" captionTargetBox="[293,1295,205,1711]" captionTargetId="figure-208@8.[280,1300,195,1713]" captionTargetPageId="8" captionText="FIGURES 1–11. 1–5. Dorsal outline: 1. Generalized Tylodinus, indicating terminology used in this study. Key: letters indicate areas of pronotum. A, anterior area; B, median area; C, posterior area; a, anterior of lateral area; b, median section of lateral area; c, median section of dorsal area; d, anterior section of dorsal area; e, posterior section of lateral area; f, posterior section of dorsal area. 2. Tylodinus variabilis, note elytra basal margin straight. 3. Tylodinus ixchel, note elytra basal margin slightly sinuate. 4. Tylodinus immundus, note elytral basal margin sinuate. 5. Tylodinus elongatus, note elytra basal margin very sinuate. 6–11. Lateral outline: 6. Tylodinus complicatus, male, note ventrites 3–5 in same plane as ventrites 1–2. 7. Tylodinus variabilis, male, note ventrites 3–5 not in same plane as ventrites 1–2. 8. Tylodinus coapillensis, female, note ventrites 1–2 strongly convex, ventrites 3–5 not in same plane as ventrites 1–2. 9. Tylodinus zilchi (schematic). 10. Tylodinus sepulturaensis (schematic) 11. Tylodinus rugosus (schematic)." figureDoi="http://doi.org/10.5281/zenodo.4913763" httpUri="https://zenodo.org/record/4913763/files/figure.png" pageId="6" pageNumber="7">Figure 10</figureCitation>
), 2) outline with anterior and posterior constrictions (
<figureCitation id="D2EC2A6CFF9D610AFB8DDB8C20BFFF11" box="[1038,1152,190,215]" captionStart="FIGURES 1–11. 1–5" captionStartId="8.[151,264,1743,1765]" captionTargetBox="[293,1295,205,1711]" captionTargetId="figure-208@8.[280,1300,195,1713]" captionTargetPageId="8" captionText="FIGURES 1–11. 1–5. Dorsal outline: 1. Generalized Tylodinus, indicating terminology used in this study. Key: letters indicate areas of pronotum. A, anterior area; B, median area; C, posterior area; a, anterior of lateral area; b, median section of lateral area; c, median section of dorsal area; d, anterior section of dorsal area; e, posterior section of lateral area; f, posterior section of dorsal area. 2. Tylodinus variabilis, note elytra basal margin straight. 3. Tylodinus ixchel, note elytra basal margin slightly sinuate. 4. Tylodinus immundus, note elytral basal margin sinuate. 5. Tylodinus elongatus, note elytra basal margin very sinuate. 6–11. Lateral outline: 6. Tylodinus complicatus, male, note ventrites 3–5 in same plane as ventrites 1–2. 7. Tylodinus variabilis, male, note ventrites 3–5 not in same plane as ventrites 1–2. 8. Tylodinus coapillensis, female, note ventrites 1–2 strongly convex, ventrites 3–5 not in same plane as ventrites 1–2. 9. Tylodinus zilchi (schematic). 10. Tylodinus sepulturaensis (schematic) 11. Tylodinus rugosus (schematic)." figureDoi="http://doi.org/10.5281/zenodo.4913763" httpUri="https://zenodo.org/record/4913763/files/figure.png" pageId="6" pageNumber="7">Figure 11</figureCitation>
), and 3) outline slanting downward from highest point to the apex and the base (
<figureCitation id="D2EC2A6CFF9D610AFC8ADBD72754FF38" box="[777,875,229,254]" captionStart="FIGURES 1–11. 1–5" captionStartId="8.[151,264,1743,1765]" captionTargetBox="[293,1295,205,1711]" captionTargetId="figure-208@8.[280,1300,195,1713]" captionTargetPageId="8" captionText="FIGURES 1–11. 1–5. Dorsal outline: 1. Generalized Tylodinus, indicating terminology used in this study. Key: letters indicate areas of pronotum. A, anterior area; B, median area; C, posterior area; a, anterior of lateral area; b, median section of lateral area; c, median section of dorsal area; d, anterior section of dorsal area; e, posterior section of lateral area; f, posterior section of dorsal area. 2. Tylodinus variabilis, note elytra basal margin straight. 3. Tylodinus ixchel, note elytra basal margin slightly sinuate. 4. Tylodinus immundus, note elytral basal margin sinuate. 5. Tylodinus elongatus, note elytra basal margin very sinuate. 6–11. Lateral outline: 6. Tylodinus complicatus, male, note ventrites 3–5 in same plane as ventrites 1–2. 7. Tylodinus variabilis, male, note ventrites 3–5 not in same plane as ventrites 1–2. 8. Tylodinus coapillensis, female, note ventrites 1–2 strongly convex, ventrites 3–5 not in same plane as ventrites 1–2. 9. Tylodinus zilchi (schematic). 10. Tylodinus sepulturaensis (schematic) 11. Tylodinus rugosus (schematic)." figureDoi="http://doi.org/10.5281/zenodo.4913763" httpUri="https://zenodo.org/record/4913763/files/figure.png" pageId="6" pageNumber="7">Figure 9</figureCitation>
), feebly, moderately or strongly convex; in dorsal view widest at about midlength; with one of two patterns: 1) sides slightly divergent from base to midlength, then convergent to apex, constricted beyond the middle on the sides (
<figureCitation id="D2EC2A6CFF9D610AFCE1DA0027EEFE8D" box="[866,977,306,331]" captionStart="FIGURES 12–19. 12" captionStartId="9.[151,264,1919,1941]" captionTargetBox="[219,1377,209,1898]" captionTargetId="figure-86@9.[219,1377,197,1898]" captionTargetPageId="9" captionText="FIGURES 12–19. 12. Tylodinus pusillus head a) male, b) female. 13–14. Dorsal habitus of pronotum showing the lateral side of pronotum 13. Tylodinus andersoni, 14. Tylodinus porvenirensis. 15–17. Dorsal habitus: 15. Tylodinus rugosus, 16. Tylodinus exiguus, 17. Tylodinus jonesi. 18. Tylodinus coapillensis note tubercle on elytra declivity base, 19. Tylodinus elongatus showing metasternal tooth." figureDoi="http://doi.org/10.5281/zenodo.4913765" httpUri="https://zenodo.org/record/4913765/files/figure.png" pageId="6" pageNumber="7">Figure 14</figureCitation>
), or 2) similar to previous pattern but not constricted beyond the middle (
<figureCitation id="D2EC2A6CFF9D610AFE7FDA6B2652FEB4" box="[508,621,345,370]" captionStart="FIGURES 12–19. 12" captionStartId="9.[151,264,1919,1941]" captionTargetBox="[219,1377,209,1898]" captionTargetId="figure-86@9.[219,1377,197,1898]" captionTargetPageId="9" captionText="FIGURES 12–19. 12. Tylodinus pusillus head a) male, b) female. 13–14. Dorsal habitus of pronotum showing the lateral side of pronotum 13. Tylodinus andersoni, 14. Tylodinus porvenirensis. 15–17. Dorsal habitus: 15. Tylodinus rugosus, 16. Tylodinus exiguus, 17. Tylodinus jonesi. 18. Tylodinus coapillensis note tubercle on elytra declivity base, 19. Tylodinus elongatus showing metasternal tooth." figureDoi="http://doi.org/10.5281/zenodo.4913765" httpUri="https://zenodo.org/record/4913765/files/figure.png" pageId="6" pageNumber="7">Figure 13</figureCitation>
); apical margin in lateral view extended over hind portion of head; disc with one of four patterns: 1) deeply excavate (
<figureCitation id="D2EC2A6CFF9D610AFD24DA4D2743FE5E" box="[679,892,383,408]" captionStart="FIGURES 28–35" captionStartId="16.[151,264,1975,1997]" captionTargetBox="[197,1353,202,1921]" captionTargetId="figure-52@16.[196,1390,193,1947]" captionTargetPageId="16" captionText="FIGURES 28–35. Dorsal and lateral habitus: 28–29. Tylodinus nodulosus, 30–31. Tylodinus zilchi, 32–33. Tylodinus andersoni, 34–35. Tylodinus cavicrus." figureDoi="http://doi.org/10.5281/zenodo.4913769" httpUri="https://zenodo.org/record/4913769/files/figure.png" pageId="6" pageNumber="7">Figures 28, 30, 32</figureCitation>
), 2) distinctly hollowed (
<figureCitation id="D2EC2A6CFF9D610AFB18DA4D2128FE5E" box="[1179,1303,383,408]" captionStart="FIGURES 28–35" captionStartId="16.[151,264,1975,1997]" captionTargetBox="[197,1353,202,1921]" captionTargetId="figure-52@16.[196,1390,193,1947]" captionTargetPageId="16" captionText="FIGURES 28–35. Dorsal and lateral habitus: 28–29. Tylodinus nodulosus, 30–31. Tylodinus zilchi, 32–33. Tylodinus andersoni, 34–35. Tylodinus cavicrus." figureDoi="http://doi.org/10.5281/zenodo.4913769" httpUri="https://zenodo.org/record/4913769/files/figure.png" pageId="6" pageNumber="7">Figures 34</figureCitation>
, 
<figureCitation id="D2EC2A6CFF9D610AFAA7DA4D217DFE5E" box="[1316,1346,383,408]" captionStart="FIGURES 36–43" captionStartId="20.[151,264,1974,1996]" captionTargetBox="[197,1389,213,1947]" captionTargetId="figure-55@20.[196,1390,195,1949]" captionTargetPageId="20" captionText="FIGURES 36–43. Dorsal and lateral habitus: 36–37. Tylodinus pseudocavicrus, 38–39. Tylodinus rugosus. 40–41. Tylodinus buchanani. 42–43. Tylodinus exiguus." figureDoi="http://doi.org/10.5281/zenodo.4913771" httpUri="https://zenodo.org/record/4913771/files/figure.png" pageId="6" pageNumber="7">38</figureCitation>
, 
<figureCitation id="D2EC2A6CFF9D610AFAD3DA4D2150FE5E" box="[1360,1391,383,408]" captionStart="FIGURES 76–83" captionStartId="45.[151,264,1974,1996]" captionTargetBox="[205,1321,214,1948]" captionTargetId="figure-50@45.[196,1390,195,1949]" captionTargetPageId="45" captionText="FIGURES 76–83. Dorsal and lateral habitus: 76–77. Tylodinus rufus, 78–79. Tylodinus dominicus, 80–81. Tylodinus noctis, 82–83. Tylodinus kuscheli." figureDoi="http://doi.org/10.5281/zenodo.4913785" httpUri="https://zenodo.org/record/4913785/files/figure.png" pageId="6" pageNumber="7">76</figureCitation>
), 3) somewhat hollowed (
<figureCitation id="D2EC2A6CFF9D610AFE0DDA942604FE79" box="[398,571,422,447]" captionStart="FIGURES 60–67" captionStartId="36.[151,264,1974,1996]" captionTargetBox="[199,1383,203,1937]" captionTargetId="figure-53@36.[196,1390,195,1949]" captionTargetPageId="36" captionText="FIGURES 60–67. Dorsal and lateral habitus: 60–61. Tylodinus lum, 62–63. Tylodinus spiniventris, 64–65. Tylodinus mutabilis, 66–67. Tylodinus parvus." figureDoi="http://doi.org/10.5281/zenodo.4913781" httpUri="https://zenodo.org/record/4913781/files/figure.png" pageId="6" pageNumber="7">Figures 64, 66</figureCitation>
, 
<figureCitation id="D2EC2A6CFF9D610AFDC9DA942656FE79" box="[586,617,422,447]" captionStart="FIGURES 76–83" captionStartId="45.[151,264,1974,1996]" captionTargetBox="[205,1321,214,1948]" captionTargetId="figure-50@45.[196,1390,195,1949]" captionTargetPageId="45" captionText="FIGURES 76–83. Dorsal and lateral habitus: 76–77. Tylodinus rufus, 78–79. Tylodinus dominicus, 80–81. Tylodinus noctis, 82–83. Tylodinus kuscheli." figureDoi="http://doi.org/10.5281/zenodo.4913785" httpUri="https://zenodo.org/record/4913785/files/figure.png" pageId="6" pageNumber="7">78</figureCitation>
), or 4) lacking impressions (
<figureCitation id="D2EC2A6CFF9D610AFC46DA94204CFE79" box="[965,1139,422,447]" captionStart="FIGURES 44–51" captionStartId="24.[151,264,1974,1996]" captionTargetBox="[209,1345,216,1924]" captionTargetId="figure-55@24.[196,1390,195,1949]" captionTargetPageId="24" captionText="FIGURES 44–51. Dorsal and lateral habitus: 44–45. Tylodinus ixchel, 46–47. Tylodinus jonesi, 48–49. Tylodinus wibmeri, 50–51. Tylodinus variabilis." figureDoi="http://doi.org/10.5281/zenodo.4913773" httpUri="https://zenodo.org/record/4913773/files/figure.png" pageId="6" pageNumber="7">Figures 44, 48</figureCitation>
); surface smooth, weakly carinate to rugose, without granules or granulate, with large (
<figureCitation id="D2EC2A6CFF9D610AFCD0DAFF27FBFE20" box="[851,964,461,486]" captionStart="FIGURES 12–19. 12" captionStartId="9.[151,264,1919,1941]" captionTargetBox="[219,1377,209,1898]" captionTargetId="figure-86@9.[219,1377,197,1898]" captionTargetPageId="9" captionText="FIGURES 12–19. 12. Tylodinus pusillus head a) male, b) female. 13–14. Dorsal habitus of pronotum showing the lateral side of pronotum 13. Tylodinus andersoni, 14. Tylodinus porvenirensis. 15–17. Dorsal habitus: 15. Tylodinus rugosus, 16. Tylodinus exiguus, 17. Tylodinus jonesi. 18. Tylodinus coapillensis note tubercle on elytra declivity base, 19. Tylodinus elongatus showing metasternal tooth." figureDoi="http://doi.org/10.5281/zenodo.4913765" httpUri="https://zenodo.org/record/4913765/files/figure.png" pageId="6" pageNumber="7">Figure 16</figureCitation>
), medium (
<figureCitation id="D2EC2A6CFF9D610AFBC4DAFF2087FE20" box="[1095,1208,461,486]" captionStart="FIGURES 12–19. 12" captionStartId="9.[151,264,1919,1941]" captionTargetBox="[219,1377,209,1898]" captionTargetId="figure-86@9.[219,1377,197,1898]" captionTargetPageId="9" captionText="FIGURES 12–19. 12. Tylodinus pusillus head a) male, b) female. 13–14. Dorsal habitus of pronotum showing the lateral side of pronotum 13. Tylodinus andersoni, 14. Tylodinus porvenirensis. 15–17. Dorsal habitus: 15. Tylodinus rugosus, 16. Tylodinus exiguus, 17. Tylodinus jonesi. 18. Tylodinus coapillensis note tubercle on elytra declivity base, 19. Tylodinus elongatus showing metasternal tooth." figureDoi="http://doi.org/10.5281/zenodo.4913765" httpUri="https://zenodo.org/record/4913765/files/figure.png" pageId="6" pageNumber="7">Figure 14</figureCitation>
) or small punctures (
<figureCitation id="D2EC2A6CFF9D610AFF1DDAC6252FFDCB" box="[158,272,500,525]" captionStart="FIGURES 12–19. 12" captionStartId="9.[151,264,1919,1941]" captionTargetBox="[219,1377,209,1898]" captionTargetId="figure-86@9.[219,1377,197,1898]" captionTargetPageId="9" captionText="FIGURES 12–19. 12. Tylodinus pusillus head a) male, b) female. 13–14. Dorsal habitus of pronotum showing the lateral side of pronotum 13. Tylodinus andersoni, 14. Tylodinus porvenirensis. 15–17. Dorsal habitus: 15. Tylodinus rugosus, 16. Tylodinus exiguus, 17. Tylodinus jonesi. 18. Tylodinus coapillensis note tubercle on elytra declivity base, 19. Tylodinus elongatus showing metasternal tooth." figureDoi="http://doi.org/10.5281/zenodo.4913765" httpUri="https://zenodo.org/record/4913765/files/figure.png" pageId="6" pageNumber="7">Figure 17</figureCitation>
), deep or shallow, with one of two patterns: 1) contiguous at the base and progressively more widely spaced, smaller and shallower to the apex (
<figureCitation id="D2EC2A6CFF9D610AFD11D92827A7FDF5" box="[658,920,538,563]" captionStart="FIGURES 12–19. 12" captionStartId="9.[151,264,1919,1941]" captionTargetBox="[219,1377,209,1898]" captionTargetId="figure-86@9.[219,1377,197,1898]" captionTargetPageId="9" captionText="FIGURES 12–19. 12. Tylodinus pusillus head a) male, b) female. 13–14. Dorsal habitus of pronotum showing the lateral side of pronotum 13. Tylodinus andersoni, 14. Tylodinus porvenirensis. 15–17. Dorsal habitus: 15. Tylodinus rugosus, 16. Tylodinus exiguus, 17. Tylodinus jonesi. 18. Tylodinus coapillensis note tubercle on elytra declivity base, 19. Tylodinus elongatus showing metasternal tooth." figureDoi="http://doi.org/10.5281/zenodo.4913765" httpUri="https://zenodo.org/record/4913765/files/figure.png" pageId="6" pageNumber="7">Figures 14, 16 and 17</figureCitation>
), or 2) dense to contiguous on lateral sides, progressively more widely spaced and less conspicuous in the median, anterior and posterior sections (
<figureCitation id="D2EC2A6CFF9D610AFA9ED97321B1FD9C" box="[1309,1422,577,602]" captionStart="FIGURES 12–19. 12" captionStartId="9.[151,264,1919,1941]" captionTargetBox="[219,1377,209,1898]" captionTargetId="figure-86@9.[219,1377,197,1898]" captionTargetPageId="9" captionText="FIGURES 12–19. 12. Tylodinus pusillus head a) male, b) female. 13–14. Dorsal habitus of pronotum showing the lateral side of pronotum 13. Tylodinus andersoni, 14. Tylodinus porvenirensis. 15–17. Dorsal habitus: 15. Tylodinus rugosus, 16. Tylodinus exiguus, 17. Tylodinus jonesi. 18. Tylodinus coapillensis note tubercle on elytra declivity base, 19. Tylodinus elongatus showing metasternal tooth." figureDoi="http://doi.org/10.5281/zenodo.4913765" httpUri="https://zenodo.org/record/4913765/files/figure.png" pageId="6" pageNumber="7">Figure 15</figureCitation>
); scales sparse, moderately dense to dense, distributed in one of three patterns: 1) light scales (generally yellow, light brown or white) very dense on lateral sides and flanks, anterior area and forming a thick longitudinal fringe along the middle and a narrow strip that diverges from the middle base to the flanks, and with dark scales on the middle and posterior areas of the disc (dark brown or black) (
<figureCitation id="D2EC2A6CFF9D610AFD7AD9EE279FFD33" box="[761,928,732,757]" captionStart="FIGURES 12–19. 12" captionStartId="9.[151,264,1919,1941]" captionTargetBox="[219,1377,209,1898]" captionTargetId="figure-86@9.[219,1377,197,1898]" captionTargetPageId="9" captionText="FIGURES 12–19. 12. Tylodinus pusillus head a) male, b) female. 13–14. Dorsal habitus of pronotum showing the lateral side of pronotum 13. Tylodinus andersoni, 14. Tylodinus porvenirensis. 15–17. Dorsal habitus: 15. Tylodinus rugosus, 16. Tylodinus exiguus, 17. Tylodinus jonesi. 18. Tylodinus coapillensis note tubercle on elytra declivity base, 19. Tylodinus elongatus showing metasternal tooth." figureDoi="http://doi.org/10.5281/zenodo.4913765" httpUri="https://zenodo.org/record/4913765/files/figure.png" pageId="6" pageNumber="7">Figures 14, 17</figureCitation>
), occasionally the pattern is not well defined (
<figureCitation id="D2EC2A6CFF9D610AFF1DD8302531FCDD" box="[158,270,770,795]" captionStart="FIGURES 44–51" captionStartId="24.[151,264,1974,1996]" captionTargetBox="[209,1345,216,1924]" captionTargetId="figure-55@24.[196,1390,195,1949]" captionTargetPageId="24" captionText="FIGURES 44–51. Dorsal and lateral habitus: 44–45. Tylodinus ixchel, 46–47. Tylodinus jonesi, 48–49. Tylodinus wibmeri, 50–51. Tylodinus variabilis." figureDoi="http://doi.org/10.5281/zenodo.4913773" httpUri="https://zenodo.org/record/4913773/files/figure.png" pageId="6" pageNumber="7">Figure 48</figureCitation>
, 
<figureCitation id="D2EC2A6CFF9D610AFE99D8302506FCDD" box="[282,313,770,795]" captionStart="FIGURES 52–59" captionStartId="28.[151,264,1974,1996]" captionTargetBox="[201,1375,209,1931]" captionTargetId="figure-55@28.[196,1390,195,1949]" captionTargetPageId="28" captionText="FIGURES 52–59. Dorsal and lateral habitus: 52–53. Tylodinus coapillensis, 54–55. Tylodinus leoncortesi, 56–57. Tylodinus triumforium, 58–59. Tylodinus sepulturaensis." figureDoi="http://doi.org/10.5281/zenodo.4913775" httpUri="https://zenodo.org/record/4913775/files/figure.png" pageId="6" pageNumber="7">56</figureCitation>
, 
<figureCitation id="D2EC2A6CFF9D610AFEC7D830255CFCDD" box="[324,355,770,795]" captionStart="FIGURES 76–83" captionStartId="45.[151,264,1974,1996]" captionTargetBox="[205,1321,214,1948]" captionTargetId="figure-50@45.[196,1390,195,1949]" captionTargetPageId="45" captionText="FIGURES 76–83. Dorsal and lateral habitus: 76–77. Tylodinus rufus, 78–79. Tylodinus dominicus, 80–81. Tylodinus noctis, 82–83. Tylodinus kuscheli." figureDoi="http://doi.org/10.5281/zenodo.4913785" httpUri="https://zenodo.org/record/4913785/files/figure.png" pageId="6" pageNumber="7">78</figureCitation>
), 2) similar to last pattern but without dark brown scales (
<figureCitation id="D2EC2A6CFF9D610AFC77D830205CFCDD" box="[1012,1123,770,795]" captionStart="FIGURES 76–83" captionStartId="45.[151,264,1974,1996]" captionTargetBox="[205,1321,214,1948]" captionTargetId="figure-50@45.[196,1390,195,1949]" captionTargetPageId="45" captionText="FIGURES 76–83. Dorsal and lateral habitus: 76–77. Tylodinus rufus, 78–79. Tylodinus dominicus, 80–81. Tylodinus noctis, 82–83. Tylodinus kuscheli." figureDoi="http://doi.org/10.5281/zenodo.4913785" httpUri="https://zenodo.org/record/4913785/files/figure.png" pageId="6" pageNumber="7">Figure 76</figureCitation>
), and 3) when excavation is present, scales dense on edge of pronotal excavation, more widely separated towards lateral and median areas of disc (
<figureCitation id="D2EC2A6CFF9D610AFF57D87D2597FCAE" box="[212,424,847,872]" captionStart="FIGURES 28–35" captionStartId="16.[151,264,1975,1997]" captionTargetBox="[197,1353,202,1921]" captionTargetId="figure-52@16.[196,1390,193,1947]" captionTargetPageId="16" captionText="FIGURES 28–35. Dorsal and lateral habitus: 28–29. Tylodinus nodulosus, 30–31. Tylodinus zilchi, 32–33. Tylodinus andersoni, 34–35. Tylodinus cavicrus." figureDoi="http://doi.org/10.5281/zenodo.4913769" httpUri="https://zenodo.org/record/4913769/files/figure.png" pageId="6" pageNumber="7">Figures 28, 30, 32</figureCitation>
); scales dense (
<figureCitation id="D2EC2A6CFF9D610AFDDBD87D26F4FCAE" box="[600,715,847,872]" captionStart="FIGURES 76–83" captionStartId="45.[151,264,1974,1996]" captionTargetBox="[205,1321,214,1948]" captionTargetId="figure-50@45.[196,1390,195,1949]" captionTargetPageId="45" captionText="FIGURES 76–83. Dorsal and lateral habitus: 76–77. Tylodinus rufus, 78–79. Tylodinus dominicus, 80–81. Tylodinus noctis, 82–83. Tylodinus kuscheli." figureDoi="http://doi.org/10.5281/zenodo.4913785" httpUri="https://zenodo.org/record/4913785/files/figure.png" pageId="6" pageNumber="7">Figure 78</figureCitation>
), moderately dense (
<figureCitation id="D2EC2A6CFF9D610AFC35D87D2019FCAE" box="[950,1062,847,872]" captionStart="FIGURES 60–67" captionStartId="36.[151,264,1974,1996]" captionTargetBox="[199,1383,203,1937]" captionTargetId="figure-53@36.[196,1390,195,1949]" captionTargetPageId="36" captionText="FIGURES 60–67. Dorsal and lateral habitus: 60–61. Tylodinus lum, 62–63. Tylodinus spiniventris, 64–65. Tylodinus mutabilis, 66–67. Tylodinus parvus." figureDoi="http://doi.org/10.5281/zenodo.4913781" httpUri="https://zenodo.org/record/4913781/files/figure.png" pageId="6" pageNumber="7">Figure 64</figureCitation>
) to scattered (
<figureCitation id="D2EC2A6CFF9D610AFB4BD87D2104FCAE" box="[1224,1339,847,872]" captionStart="FIGURES 76–83" captionStartId="45.[151,264,1974,1996]" captionTargetBox="[205,1321,214,1948]" captionTargetId="figure-50@45.[196,1390,195,1949]" captionTargetPageId="45" captionText="FIGURES 76–83. Dorsal and lateral habitus: 76–77. Tylodinus rufus, 78–79. Tylodinus dominicus, 80–81. Tylodinus noctis, 82–83. Tylodinus kuscheli." figureDoi="http://doi.org/10.5281/zenodo.4913785" httpUri="https://zenodo.org/record/4913785/files/figure.png" pageId="6" pageNumber="7">Figure 76</figureCitation>
). 
<emphasis id="78A3EAFBFF9D610AFAD3D87D21A3FCAE" bold="true" box="[1360,1436,847,872]" pageId="6" pageNumber="7">Elytra</emphasis>
with the basal margin straight to very sinuate (
<figureCitation id="D2EC2A6CFF9D610AFD26D844270FFC49" box="[677,816,886,911]" captionStart="FIGURES 1–11. 1–5" captionStartId="8.[151,264,1743,1765]" captionTargetBox="[293,1295,205,1711]" captionTargetId="figure-208@8.[280,1300,195,1713]" captionTargetPageId="8" captionText="FIGURES 1–11. 1–5. Dorsal outline: 1. Generalized Tylodinus, indicating terminology used in this study. Key: letters indicate areas of pronotum. A, anterior area; B, median area; C, posterior area; a, anterior of lateral area; b, median section of lateral area; c, median section of dorsal area; d, anterior section of dorsal area; e, posterior section of lateral area; f, posterior section of dorsal area. 2. Tylodinus variabilis, note elytra basal margin straight. 3. Tylodinus ixchel, note elytra basal margin slightly sinuate. 4. Tylodinus immundus, note elytral basal margin sinuate. 5. Tylodinus elongatus, note elytra basal margin very sinuate. 6–11. Lateral outline: 6. Tylodinus complicatus, male, note ventrites 3–5 in same plane as ventrites 1–2. 7. Tylodinus variabilis, male, note ventrites 3–5 not in same plane as ventrites 1–2. 8. Tylodinus coapillensis, female, note ventrites 1–2 strongly convex, ventrites 3–5 not in same plane as ventrites 1–2. 9. Tylodinus zilchi (schematic). 10. Tylodinus sepulturaensis (schematic) 11. Tylodinus rugosus (schematic)." figureDoi="http://doi.org/10.5281/zenodo.4913763" httpUri="https://zenodo.org/record/4913763/files/figure.png" pageId="6" pageNumber="7">Figures 2–5</figureCitation>
); widest just posterior to basal margin then convergent to apex; apex narrowly truncate, clothed with sparse to moderately dense scales, more dense on middle of disc and on base, becoming more scattered toward the flanks; elytral declivity with scales denser on interval 1–2 (
<figureCitation id="D2EC2A6CFF9D610AFAC4D8F62509FBC5" captionStart="FIGURES 52–59" captionStartId="28.[151,264,1974,1996]" captionTargetBox="[201,1375,209,1931]" captionTargetId="figure-55@28.[196,1390,195,1949]" captionTargetPageId="28" captionText="FIGURES 52–59. Dorsal and lateral habitus: 52–53. Tylodinus coapillensis, 54–55. Tylodinus leoncortesi, 56–57. Tylodinus triumforium, 58–59. Tylodinus sepulturaensis." figureDoi="http://doi.org/10.5281/zenodo.4913775" httpUri="https://zenodo.org/record/4913775/files/figure.png" pageId="6" pageNumber="7">Figures 56–57, 58–59</figureCitation>
, 
<figureCitation id="D2EC2A6CFF9D610AFEC0D8D825B3FBC5" box="[323,396,1002,1027]" captionStart="FIGURES 68–75" captionStartId="39.[151,264,1969,1991]" captionTargetBox="[201,1389,216,1946]" captionTargetId="figure-52@39.[196,1390,193,1947]" captionTargetPageId="39" captionText="FIGURES 68–75. Dorsal and lateral habitus: 68–69. Tylodinus pusillus, 70–71. Tylodinus porvenirensis, 72–73. Tylodinus elongatus, 74–75. Tylodinus complicatus." figureDoi="http://doi.org/10.5281/zenodo.4913783" httpUri="https://zenodo.org/record/4913783/files/figure.png" pageId="6" pageNumber="7">72–73</figureCitation>
, 
<figureCitation id="D2EC2A6CFF9D610AFE1AD8D825DEFBC5" box="[409,481,1002,1027]" captionStart="FIGURES 76–83" captionStartId="45.[151,264,1974,1996]" captionTargetBox="[205,1321,214,1948]" captionTargetId="figure-50@45.[196,1390,195,1949]" captionTargetPageId="45" captionText="FIGURES 76–83. Dorsal and lateral habitus: 76–77. Tylodinus rufus, 78–79. Tylodinus dominicus, 80–81. Tylodinus noctis, 82–83. Tylodinus kuscheli." figureDoi="http://doi.org/10.5281/zenodo.4913785" httpUri="https://zenodo.org/record/4913785/files/figure.png" pageId="6" pageNumber="7">76–77</figureCitation>
) or with dense patch on intervals 1–3 (
<figureCitation id="D2EC2A6CFF9D610AFC26D8D82072FBC5" box="[933,1101,1002,1027]" captionStart="FIGURES 44–51" captionStartId="24.[151,264,1974,1996]" captionTargetBox="[209,1345,216,1924]" captionTargetId="figure-55@24.[196,1390,195,1949]" captionTargetPageId="24" captionText="FIGURES 44–51. Dorsal and lateral habitus: 44–45. Tylodinus ixchel, 46–47. Tylodinus jonesi, 48–49. Tylodinus wibmeri, 50–51. Tylodinus variabilis." figureDoi="http://doi.org/10.5281/zenodo.4913773" httpUri="https://zenodo.org/record/4913773/files/figure.png" pageId="6" pageNumber="7">Figures 46–47</figureCitation>
, 
<figureCitation id="D2EC2A6CFF9D610AFBDAD8D8209EFBC5" box="[1113,1185,1002,1027]" captionStart="FIGURES 68–75" captionStartId="39.[151,264,1969,1991]" captionTargetBox="[201,1389,216,1946]" captionTargetId="figure-52@39.[196,1390,193,1947]" captionTargetPageId="39" captionText="FIGURES 68–75. Dorsal and lateral habitus: 68–69. Tylodinus pusillus, 70–71. Tylodinus porvenirensis, 72–73. Tylodinus elongatus, 74–75. Tylodinus complicatus." figureDoi="http://doi.org/10.5281/zenodo.4913783" httpUri="https://zenodo.org/record/4913783/files/figure.png" pageId="6" pageNumber="7">74–75</figureCitation>
) and on intervals 1–5 (
<figureCitation id="D2EC2A6CFF9D610AFF1DDF232578FBEC" box="[158,327,1041,1066]" captionStart="FIGURES 44–51" captionStartId="24.[151,264,1974,1996]" captionTargetBox="[209,1345,216,1924]" captionTargetId="figure-55@24.[196,1390,195,1949]" captionTargetPageId="24" captionText="FIGURES 44–51. Dorsal and lateral habitus: 44–45. Tylodinus ixchel, 46–47. Tylodinus jonesi, 48–49. Tylodinus wibmeri, 50–51. Tylodinus variabilis." figureDoi="http://doi.org/10.5281/zenodo.4913773" httpUri="https://zenodo.org/record/4913773/files/figure.png" pageId="6" pageNumber="7">Figures 48–49</figureCitation>
, 
<figureCitation id="D2EC2A6CFF9D610AFED6DF2325A3FBEC" box="[341,412,1041,1066]" captionStart="FIGURES 76–83" captionStartId="45.[151,264,1974,1996]" captionTargetBox="[205,1321,214,1948]" captionTargetId="figure-50@45.[196,1390,195,1949]" captionTargetPageId="45" captionText="FIGURES 76–83. Dorsal and lateral habitus: 76–77. Tylodinus rufus, 78–79. Tylodinus dominicus, 80–81. Tylodinus noctis, 82–83. Tylodinus kuscheli." figureDoi="http://doi.org/10.5281/zenodo.4913785" httpUri="https://zenodo.org/record/4913785/files/figure.png" pageId="6" pageNumber="7">78–79</figureCitation>
) at base of declivity, base with scales similar those of disc or with light brown or white scales on tubercles or only on first tubercle; integument color with one of three patterns: 1) black (
<figureCitation id="D2EC2A6CFF9D610AFB6ADF0521B0FB96" box="[1257,1423,1079,1104]" captionStart-0="FIGURES 52–59" captionStart-1="FIGURES 60–67" captionStartId-0="28.[151,264,1974,1996]" captionStartId-1="36.[151,264,1974,1996]" captionTargetBox-0="[201,1375,209,1931]" captionTargetBox-1="[199,1383,203,1937]" captionTargetId-0="figure-55@28.[196,1390,195,1949]" captionTargetId-1="figure-53@36.[196,1390,195,1949]" captionTargetPageId-0="28" captionTargetPageId-1="36" captionText-0="FIGURES 52–59. Dorsal and lateral habitus: 52–53. Tylodinus coapillensis, 54–55. Tylodinus leoncortesi, 56–57. Tylodinus triumforium, 58–59. Tylodinus sepulturaensis." captionText-1="FIGURES 60–67. Dorsal and lateral habitus: 60–61. Tylodinus lum, 62–63. Tylodinus spiniventris, 64–65. Tylodinus mutabilis, 66–67. Tylodinus parvus." figureDoi-0="http://doi.org/10.5281/zenodo.4913775" figureDoi-1="http://doi.org/10.5281/zenodo.4913781" httpUri-0="https://zenodo.org/record/4913775/files/figure.png" httpUri-1="https://zenodo.org/record/4913781/files/figure.png" pageId="6" pageNumber="7">Figures 52–63</figureCitation>
), 2) black with red coloration at the base and along intervals 1–2 and on declivity of intervals 1–5 (
<figureCitation id="D2EC2A6CFF9D610AFB6BDF6C21AFFBB1" box="[1256,1424,1118,1143]" captionStart-0="FIGURES 68–75" captionStart-1="FIGURES 76–83" captionStartId-0="39.[151,264,1969,1991]" captionStartId-1="45.[151,264,1974,1996]" captionTargetBox-0="[201,1389,216,1946]" captionTargetBox-1="[205,1321,214,1948]" captionTargetId-0="figure-52@39.[196,1390,193,1947]" captionTargetId-1="figure-50@45.[196,1390,195,1949]" captionTargetPageId-0="39" captionTargetPageId-1="45" captionText-0="FIGURES 68–75. Dorsal and lateral habitus: 68–69. Tylodinus pusillus, 70–71. Tylodinus porvenirensis, 72–73. Tylodinus elongatus, 74–75. Tylodinus complicatus." captionText-1="FIGURES 76–83. Dorsal and lateral habitus: 76–77. Tylodinus rufus, 78–79. Tylodinus dominicus, 80–81. Tylodinus noctis, 82–83. Tylodinus kuscheli." figureDoi-0="http://doi.org/10.5281/zenodo.4913783" figureDoi-1="http://doi.org/10.5281/zenodo.4913785" httpUri-0="https://zenodo.org/record/4913783/files/figure.png" httpUri-1="https://zenodo.org/record/4913785/files/figure.png" pageId="6" pageNumber="7">Figures 74–77</figureCitation>
), and 3) black with red coloration at the base and along middle line, elytral declivity and flanks (
<figureCitation id="D2EC2A6CFF9D610AFB6CDFB721A7FB58" box="[1263,1432,1157,1182]" captionStart="FIGURES 44–51" captionStartId="24.[151,264,1974,1996]" captionTargetBox="[209,1345,216,1924]" captionTargetId="figure-55@24.[196,1390,195,1949]" captionTargetPageId="24" captionText="FIGURES 44–51. Dorsal and lateral habitus: 44–45. Tylodinus ixchel, 46–47. Tylodinus jonesi, 48–49. Tylodinus wibmeri, 50–51. Tylodinus variabilis." figureDoi="http://doi.org/10.5281/zenodo.4913773" httpUri="https://zenodo.org/record/4913773/files/figure.png" pageId="6" pageNumber="7">Figures 44–51</figureCitation>
, 
<figureCitation id="D2EC2A6CFF9D610AFF14DF9E24DDFB03" box="[151,226,1196,1221]" captionStart="FIGURES 60–67" captionStartId="36.[151,264,1974,1996]" captionTargetBox="[199,1383,203,1937]" captionTargetId="figure-53@36.[196,1390,195,1949]" captionTargetPageId="36" captionText="FIGURES 60–67. Dorsal and lateral habitus: 60–61. Tylodinus lum, 62–63. Tylodinus spiniventris, 64–65. Tylodinus mutabilis, 66–67. Tylodinus parvus." figureDoi="http://doi.org/10.5281/zenodo.4913781" httpUri="https://zenodo.org/record/4913781/files/figure.png" pageId="6" pageNumber="7">64–65</figureCitation>
). Intervals weakly convex, less convex to flat toward flanks, intervals 2, 3, 7, 8 and 9 convex on declivity near apex; striae shallow; stria 10 complete, incomplete or absent; tubercles on elytra disc variously developed, laminate (
<figureCitation id="D2EC2A6CFF9D610AFE84DFCB25E5FAD4" box="[263,474,1273,1298]" captionStart="FIGURES 28–35" captionStartId="16.[151,264,1975,1997]" captionTargetBox="[197,1353,202,1921]" captionTargetId="figure-52@16.[196,1390,193,1947]" captionTargetPageId="16" captionText="FIGURES 28–35. Dorsal and lateral habitus: 28–29. Tylodinus nodulosus, 30–31. Tylodinus zilchi, 32–33. Tylodinus andersoni, 34–35. Tylodinus cavicrus." figureDoi="http://doi.org/10.5281/zenodo.4913769" httpUri="https://zenodo.org/record/4913769/files/figure.png" pageId="6" pageNumber="7">Figures 29, 31, 33</figureCitation>
) or round; declivity with tubercle on interval 4 laminate (
<figureCitation id="D2EC2A6CFF9D610AFBEDDFCB2109FAD4" box="[1134,1334,1273,1298]" captionStart="FIGURES 28–35" captionStartId="16.[151,264,1975,1997]" captionTargetBox="[197,1353,202,1921]" captionTargetId="figure-52@16.[196,1390,193,1947]" captionTargetPageId="16" captionText="FIGURES 28–35. Dorsal and lateral habitus: 28–29. Tylodinus nodulosus, 30–31. Tylodinus zilchi, 32–33. Tylodinus andersoni, 34–35. Tylodinus cavicrus." figureDoi="http://doi.org/10.5281/zenodo.4913769" httpUri="https://zenodo.org/record/4913769/files/figure.png" pageId="6" pageNumber="7">Figures 29,31, 33</figureCitation>
and 
<figureCitation id="D2EC2A6CFF9D610AFAF3DFCB21B0FAD4" box="[1392,1423,1273,1298]" captionStart="FIGURES 92–99. 92–99" captionStartId="54.[151,264,1902,1924]" captionTargetBox="[175,1413,195,1889]" captionTargetId="figure-87@54.[170,1422,195,1889]" captionTargetPageId="54" captionText="FIGURES 92–99. 92–99. Dorsal and lateral habitus: 92–93. Tylodinus gibbosus, 94–95. Tylodinus kissingeri, 96–97. Tylodinus pinguis. 98. Median lobe apex of: a) Tylodinus elongatus, b) Tylodinus exiguus, c) Tylodinus pseudocavicrus, d) Tylodinus rufus, e) Tylodinus rufus (squashed), 99. Tylodinus zilchi, ventral and lateral habitus of median lobe." figureDoi="http://doi.org/10.5281/zenodo.4913789" httpUri="https://zenodo.org/record/4913789/files/figure.png" pageId="6" pageNumber="7">97</figureCitation>
), acute (
<figureCitation id="D2EC2A6CFF9D610AFF63DE2D2572FAFE" box="[224,333,1311,1336]" captionStart="FIGURES 12–19. 12" captionStartId="9.[151,264,1919,1941]" captionTargetBox="[219,1377,209,1898]" captionTargetId="figure-86@9.[219,1377,197,1898]" captionTargetPageId="9" captionText="FIGURES 12–19. 12. Tylodinus pusillus head a) male, b) female. 13–14. Dorsal habitus of pronotum showing the lateral side of pronotum 13. Tylodinus andersoni, 14. Tylodinus porvenirensis. 15–17. Dorsal habitus: 15. Tylodinus rugosus, 16. Tylodinus exiguus, 17. Tylodinus jonesi. 18. Tylodinus coapillensis note tubercle on elytra declivity base, 19. Tylodinus elongatus showing metasternal tooth." figureDoi="http://doi.org/10.5281/zenodo.4913765" httpUri="https://zenodo.org/record/4913765/files/figure.png" pageId="6" pageNumber="7">Figure 18</figureCitation>
) or round; tubercle on interval 2 absent, longer, smaller or same size as tubercle in same position at interval 4; surface with or without granules. Scutellar shield absent. Wings absent. 
<emphasis id="78A3EAFBFF9D610AFB09DE742114FA99" bold="true" box="[1162,1323,1350,1375]" pageId="6" pageNumber="7">Metasternum</emphasis>
concave, moderately concave or flat, glabrous on median section or squamous; metepimeron visible; metasternal tooth present (
<figureCitation id="D2EC2A6CFF9D610AFF74DEA62559FA6B" box="[247,358,1428,1453]" captionStart="FIGURES 12–19. 12" captionStartId="9.[151,264,1919,1941]" captionTargetBox="[219,1377,209,1898]" captionTargetId="figure-86@9.[219,1377,197,1898]" captionTargetPageId="9" captionText="FIGURES 12–19. 12. Tylodinus pusillus head a) male, b) female. 13–14. Dorsal habitus of pronotum showing the lateral side of pronotum 13. Tylodinus andersoni, 14. Tylodinus porvenirensis. 15–17. Dorsal habitus: 15. Tylodinus rugosus, 16. Tylodinus exiguus, 17. Tylodinus jonesi. 18. Tylodinus coapillensis note tubercle on elytra declivity base, 19. Tylodinus elongatus showing metasternal tooth." figureDoi="http://doi.org/10.5281/zenodo.4913765" httpUri="https://zenodo.org/record/4913765/files/figure.png" pageId="6" pageNumber="7">Figure 19</figureCitation>
) or absent. 
<emphasis id="78A3EAFBFF9D610AFE68DEA62660FA6B" bold="true" box="[491,607,1428,1453]" pageId="6" pageNumber="7">Abdomen</emphasis>
with scales narrow and elongate (
<figureCitation id="D2EC2A6CFF9D610AFC5CDEA62047FA6B" box="[991,1144,1428,1453]" captionStart="FIGURES 20–27. 20–21" captionStartId="10.[151,264,1914,1936]" captionTargetBox="[217,1351,197,1862]" captionTargetId="figure-81@10.[216,1351,197,1863]" captionTargetPageId="10" captionText="FIGURES 20–27. 20–21. Tylodinus spiniventris: 20. meso- and metathorax (male) 21. ventrite 5 (female), 22. Tylodinus elongatus ventrite 5 (female), 23. Tylodinus triumforium, ventrite 2 (male). 24. Tylodinus complicatus, ventrite 2 (female). 25. Tylodinus complicatus, ventrite 2 (male), 26. Hind tibia of Tylodinus triumforium, 27. Front tibia of Tylodinus rugosus." figureDoi="http://doi.org/10.5281/zenodo.4913767" httpUri="https://zenodo.org/record/4913767/files/figure.png" pageId="6" pageNumber="7">Figure 20–25</figureCitation>
), scales wider, longer and dense at sides; male with ventrites 
<quantity id="8D2F9B0CFF9D610AFDA5DE882651FA15" box="[550,622,1466,1491]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" metricValueMax="1.27" metricValueMin="0.762" pageId="6" pageNumber="7" unit="in" value="4.0" valueMax="5.0" valueMin="3.0">3–5 in</quantity>
same plane as ventrites 1–2 or not (
<figureCitation id="D2EC2A6CFF9D610AFB88DE8820A8FA15" box="[1035,1175,1466,1491]" captionStart="FIGURES 1–11. 1–5" captionStartId="8.[151,264,1743,1765]" captionTargetBox="[293,1295,205,1711]" captionTargetId="figure-208@8.[280,1300,195,1713]" captionTargetPageId="8" captionText="FIGURES 1–11. 1–5. Dorsal outline: 1. Generalized Tylodinus, indicating terminology used in this study. Key: letters indicate areas of pronotum. A, anterior area; B, median area; C, posterior area; a, anterior of lateral area; b, median section of lateral area; c, median section of dorsal area; d, anterior section of dorsal area; e, posterior section of lateral area; f, posterior section of dorsal area. 2. Tylodinus variabilis, note elytra basal margin straight. 3. Tylodinus ixchel, note elytra basal margin slightly sinuate. 4. Tylodinus immundus, note elytral basal margin sinuate. 5. Tylodinus elongatus, note elytra basal margin very sinuate. 6–11. Lateral outline: 6. Tylodinus complicatus, male, note ventrites 3–5 in same plane as ventrites 1–2. 7. Tylodinus variabilis, male, note ventrites 3–5 not in same plane as ventrites 1–2. 8. Tylodinus coapillensis, female, note ventrites 1–2 strongly convex, ventrites 3–5 not in same plane as ventrites 1–2. 9. Tylodinus zilchi (schematic). 10. Tylodinus sepulturaensis (schematic) 11. Tylodinus rugosus (schematic)." figureDoi="http://doi.org/10.5281/zenodo.4913763" httpUri="https://zenodo.org/record/4913763/files/figure.png" pageId="6" pageNumber="7">Figures 6–7</figureCitation>
); female with ventrites 3–5 not in same plane as 1–2 (
<figureCitation id="D2EC2A6CFF9D610AFD8FDED3264EFA3C" box="[524,625,1505,1530]" captionStart="FIGURES 1–11. 1–5" captionStartId="8.[151,264,1743,1765]" captionTargetBox="[293,1295,205,1711]" captionTargetId="figure-208@8.[280,1300,195,1713]" captionTargetPageId="8" captionText="FIGURES 1–11. 1–5. Dorsal outline: 1. Generalized Tylodinus, indicating terminology used in this study. Key: letters indicate areas of pronotum. A, anterior area; B, median area; C, posterior area; a, anterior of lateral area; b, median section of lateral area; c, median section of dorsal area; d, anterior section of dorsal area; e, posterior section of lateral area; f, posterior section of dorsal area. 2. Tylodinus variabilis, note elytra basal margin straight. 3. Tylodinus ixchel, note elytra basal margin slightly sinuate. 4. Tylodinus immundus, note elytral basal margin sinuate. 5. Tylodinus elongatus, note elytra basal margin very sinuate. 6–11. Lateral outline: 6. Tylodinus complicatus, male, note ventrites 3–5 in same plane as ventrites 1–2. 7. Tylodinus variabilis, male, note ventrites 3–5 not in same plane as ventrites 1–2. 8. Tylodinus coapillensis, female, note ventrites 1–2 strongly convex, ventrites 3–5 not in same plane as ventrites 1–2. 9. Tylodinus zilchi (schematic). 10. Tylodinus sepulturaensis (schematic) 11. Tylodinus rugosus (schematic)." figureDoi="http://doi.org/10.5281/zenodo.4913763" httpUri="https://zenodo.org/record/4913763/files/figure.png" pageId="6" pageNumber="7">Figure 8</figureCitation>
); ventrite 1 concave, moderately concave, feebly concave or weakly convex, glabrous on median section, squamous or with scattered scales or with an anterior fringe of scales; ventrite 2 feebly concave, concave or weakly convex, longer than or as long as ventrites 3–4 combined, glabrous (
<figureCitation id="D2EC2A6CFF9D610AFAD2DD1C2489F9A8" captionStart="FIGURES 20–27. 20–21" captionStartId="10.[151,264,1914,1936]" captionTargetBox="[217,1351,197,1862]" captionTargetId="figure-81@10.[216,1351,197,1863]" captionTargetPageId="10" captionText="FIGURES 20–27. 20–21. Tylodinus spiniventris: 20. meso- and metathorax (male) 21. ventrite 5 (female), 22. Tylodinus elongatus ventrite 5 (female), 23. Tylodinus triumforium, ventrite 2 (male). 24. Tylodinus complicatus, ventrite 2 (female). 25. Tylodinus complicatus, ventrite 2 (male), 26. Hind tibia of Tylodinus triumforium, 27. Front tibia of Tylodinus rugosus." figureDoi="http://doi.org/10.5281/zenodo.4913767" httpUri="https://zenodo.org/record/4913767/files/figure.png" pageId="6" pageNumber="7">Figure 25</figureCitation>
), or with two transverse lines of scales interrupted medially (
<figureCitation id="D2EC2A6CFF9D610AFC05DD6727C7F9A8" box="[902,1016,1621,1646]" captionStart="FIGURES 20–27. 20–21" captionStartId="10.[151,264,1914,1936]" captionTargetBox="[217,1351,197,1862]" captionTargetId="figure-81@10.[216,1351,197,1863]" captionTargetPageId="10" captionText="FIGURES 20–27. 20–21. Tylodinus spiniventris: 20. meso- and metathorax (male) 21. ventrite 5 (female), 22. Tylodinus elongatus ventrite 5 (female), 23. Tylodinus triumforium, ventrite 2 (male). 24. Tylodinus complicatus, ventrite 2 (female). 25. Tylodinus complicatus, ventrite 2 (male), 26. Hind tibia of Tylodinus triumforium, 27. Front tibia of Tylodinus rugosus." figureDoi="http://doi.org/10.5281/zenodo.4913767" httpUri="https://zenodo.org/record/4913767/files/figure.png" pageId="6" pageNumber="7">Figure 23</figureCitation>
) or not interrupted, female of some species with transverse line of scales interrupted medially (
<figureCitation id="D2EC2A6CFF9D610AFCB2DD4E279BF953" box="[817,932,1660,1685]" captionStart="FIGURES 20–27. 20–21" captionStartId="10.[151,264,1914,1936]" captionTargetBox="[217,1351,197,1862]" captionTargetId="figure-81@10.[216,1351,197,1863]" captionTargetPageId="10" captionText="FIGURES 20–27. 20–21. Tylodinus spiniventris: 20. meso- and metathorax (male) 21. ventrite 5 (female), 22. Tylodinus elongatus ventrite 5 (female), 23. Tylodinus triumforium, ventrite 2 (male). 24. Tylodinus complicatus, ventrite 2 (female). 25. Tylodinus complicatus, ventrite 2 (male), 26. Hind tibia of Tylodinus triumforium, 27. Front tibia of Tylodinus rugosus." figureDoi="http://doi.org/10.5281/zenodo.4913767" httpUri="https://zenodo.org/record/4913767/files/figure.png" pageId="6" pageNumber="7">Figure 24</figureCitation>
); female with ventrites 1–2 strongly convex; ventrites 3–4 squamous; ventrite 5 convex along median line or convex at anterior 1/2 and flat on posterior 1/2 or flat, as long or longer than ventrite 1, with one of three patterns: 1) with an apical impression, 2) with apical and lateral impression or 3) without impression, in some species, female with acute projections (
<figureCitation id="D2EC2A6CFF9D610AFB29DDDD216FF8CE" box="[1194,1360,1775,1800]" captionStart="FIGURES 20–27. 20–21" captionStartId="10.[151,264,1914,1936]" captionTargetBox="[217,1351,197,1862]" captionTargetId="figure-81@10.[216,1351,197,1863]" captionTargetPageId="10" captionText="FIGURES 20–27. 20–21. Tylodinus spiniventris: 20. meso- and metathorax (male) 21. ventrite 5 (female), 22. Tylodinus elongatus ventrite 5 (female), 23. Tylodinus triumforium, ventrite 2 (male). 24. Tylodinus complicatus, ventrite 2 (female). 25. Tylodinus complicatus, ventrite 2 (male), 26. Hind tibia of Tylodinus triumforium, 27. Front tibia of Tylodinus rugosus." figureDoi="http://doi.org/10.5281/zenodo.4913767" httpUri="https://zenodo.org/record/4913767/files/figure.png" pageId="6" pageNumber="7">Figures 21–22</figureCitation>
). 
<emphasis id="78A3EAFBFF9D610AFAE7DDDD21A3F8CE" bold="true" box="[1380,1436,1775,1800]" pageId="6" pageNumber="7">Legs</emphasis>
narrowly elongate (3.6–5 x longer than wide); with feeble punctures or densely, deeply punctuate throughout, punctures elongate-oval and linearly arranged giving legs (especially femora) a striated appearance, with fine granules between punctures; hind femur with length variable, from not reaching elytral apex to reaching beyond the elytra apex; femora armed with a small, distally inclined tooth ventrally or unarmed; middle and hind tibiae with outer margin subparallel or an external tooth (
<figureCitation id="D2EC2A6CFF9D610AFD28DC83271FF80C" box="[683,800,1969,1994]" captionStart="FIGURES 20–27. 20–21" captionStartId="10.[151,264,1914,1936]" captionTargetBox="[217,1351,197,1862]" captionTargetId="figure-81@10.[216,1351,197,1863]" captionTargetPageId="10" captionText="FIGURES 20–27. 20–21. Tylodinus spiniventris: 20. meso- and metathorax (male) 21. ventrite 5 (female), 22. Tylodinus elongatus ventrite 5 (female), 23. Tylodinus triumforium, ventrite 2 (male). 24. Tylodinus complicatus, ventrite 2 (female). 25. Tylodinus complicatus, ventrite 2 (male), 26. Hind tibia of Tylodinus triumforium, 27. Front tibia of Tylodinus rugosus." figureDoi="http://doi.org/10.5281/zenodo.4913767" httpUri="https://zenodo.org/record/4913767/files/figure.png" pageId="6" pageNumber="7">Figure 26</figureCitation>
); front tibia straight to sinuate or very deeply excavate apically (
<figureCitation id="D2EC2A6CFF9D610AFF7CDCEA254CF837" box="[255,371,2008,2033]" captionStart="FIGURES 20–27. 20–21" captionStartId="10.[151,264,1914,1936]" captionTargetBox="[217,1351,197,1862]" captionTargetId="figure-81@10.[216,1351,197,1863]" captionTargetPageId="10" captionText="FIGURES 20–27. 20–21. Tylodinus spiniventris: 20. meso- and metathorax (male) 21. ventrite 5 (female), 22. Tylodinus elongatus ventrite 5 (female), 23. Tylodinus triumforium, ventrite 2 (male). 24. Tylodinus complicatus, ventrite 2 (female). 25. Tylodinus complicatus, ventrite 2 (male), 26. Hind tibia of Tylodinus triumforium, 27. Front tibia of Tylodinus rugosus." figureDoi="http://doi.org/10.5281/zenodo.4913767" httpUri="https://zenodo.org/record/4913767/files/figure.png" pageId="6" pageNumber="7">Figure 27</figureCitation>
); premucro present or absent. 
<emphasis id="78A3EAFBFF9D610AFD48DCEA2706F837" bold="true" box="[715,825,2008,2033]" pageId="6" pageNumber="7">Genitalia</emphasis>
greatly varied, male with median lobe in lateral view with curvature and width varied, from weakly to strongly curved, very slender to stout, apex straight to abruptly curved downward; width of apical process very slender to robust; apical process form almost linear, slightly curved, sinuate or curved upward; occasionally with a longitudinal notch on lateral base. Median lobe length in ventral view from longer than to ½ as long as the length of the apodemes, sides straight or convex near the base (less than of 0.25 of length from the base) to near the middle (&gt;0.27 to 0.5 of length from the base), convergent (base of median lobe wider than subbasal portion of apex), divergent (subbasal portion of apex wider than base of median lobe) or subparallel (base of median lobe as wide as subbasal portion of apex), apex of median lobe acute (
<figureCitation id="D2EC2A6CFF9C610BFF1DDA942505FE79" box="[158,314,422,447]" captionStart="FIGURES 92–99. 92–99" captionStartId="54.[151,264,1902,1924]" captionTargetBox="[175,1413,195,1889]" captionTargetId="figure-87@54.[170,1422,195,1889]" captionTargetPageId="54" captionText="FIGURES 92–99. 92–99. Dorsal and lateral habitus: 92–93. Tylodinus gibbosus, 94–95. Tylodinus kissingeri, 96–97. Tylodinus pinguis. 98. Median lobe apex of: a) Tylodinus elongatus, b) Tylodinus exiguus, c) Tylodinus pseudocavicrus, d) Tylodinus rufus, e) Tylodinus rufus (squashed), 99. Tylodinus zilchi, ventral and lateral habitus of median lobe." figureDoi="http://doi.org/10.5281/zenodo.4913789" httpUri="https://zenodo.org/record/4913789/files/figure.png" pageId="7" pageNumber="8">Figure 98d–e</figureCitation>
), truncate (
<figureCitation id="D2EC2A6CFF9C610BFE3ADA942609FE79" box="[441,566,422,447]" captionStart="FIGURES 92–99. 92–99" captionStartId="54.[151,264,1902,1924]" captionTargetBox="[175,1413,195,1889]" captionTargetId="figure-87@54.[170,1422,195,1889]" captionTargetPageId="54" captionText="FIGURES 92–99. 92–99. Dorsal and lateral habitus: 92–93. Tylodinus gibbosus, 94–95. Tylodinus kissingeri, 96–97. Tylodinus pinguis. 98. Median lobe apex of: a) Tylodinus elongatus, b) Tylodinus exiguus, c) Tylodinus pseudocavicrus, d) Tylodinus rufus, e) Tylodinus rufus (squashed), 99. Tylodinus zilchi, ventral and lateral habitus of median lobe." figureDoi="http://doi.org/10.5281/zenodo.4913789" httpUri="https://zenodo.org/record/4913789/files/figure.png" pageId="7" pageNumber="8">Figure 98c</figureCitation>
) or blunt (
<figureCitation id="D2EC2A6CFF9C610BFD32DA94270EFE79" box="[689,817,422,447]" captionStart="FIGURES 92–99. 92–99" captionStartId="54.[151,264,1902,1924]" captionTargetBox="[175,1413,195,1889]" captionTargetId="figure-87@54.[170,1422,195,1889]" captionTargetPageId="54" captionText="FIGURES 92–99. 92–99. Dorsal and lateral habitus: 92–93. Tylodinus gibbosus, 94–95. Tylodinus kissingeri, 96–97. Tylodinus pinguis. 98. Median lobe apex of: a) Tylodinus elongatus, b) Tylodinus exiguus, c) Tylodinus pseudocavicrus, d) Tylodinus rufus, e) Tylodinus rufus (squashed), 99. Tylodinus zilchi, ventral and lateral habitus of median lobe." figureDoi="http://doi.org/10.5281/zenodo.4913789" httpUri="https://zenodo.org/record/4913789/files/figure.png" pageId="7" pageNumber="8">Figure 98a</figureCitation>
); apical process constricted beyond the median orifice (
<figureCitation id="D2EC2A6CFF9C610BFF1DDAFF2563FE20" box="[158,348,461,486]" captionStart="FIGURES 92–99. 92–99" captionStartId="54.[151,264,1902,1924]" captionTargetBox="[175,1413,195,1889]" captionTargetId="figure-87@54.[170,1422,195,1889]" captionTargetPageId="54" captionText="FIGURES 92–99. 92–99. Dorsal and lateral habitus: 92–93. Tylodinus gibbosus, 94–95. Tylodinus kissingeri, 96–97. Tylodinus pinguis. 98. Median lobe apex of: a) Tylodinus elongatus, b) Tylodinus exiguus, c) Tylodinus pseudocavicrus, d) Tylodinus rufus, e) Tylodinus rufus (squashed), 99. Tylodinus zilchi, ventral and lateral habitus of median lobe." figureDoi="http://doi.org/10.5281/zenodo.4913789" httpUri="https://zenodo.org/record/4913789/files/figure.png" pageId="7" pageNumber="8">Figure 98a,c,d–e</figureCitation>
) or not, sides sub-parallel, convergent and sometimes rhomboid-shaped subapically (
<figureCitation id="D2EC2A6CFF9C610BFA91DAFF21B0FE20" box="[1298,1423,461,486]" captionStart="FIGURES 92–99. 92–99" captionStartId="54.[151,264,1902,1924]" captionTargetBox="[175,1413,195,1889]" captionTargetId="figure-87@54.[170,1422,195,1889]" captionTargetPageId="54" captionText="FIGURES 92–99. 92–99. Dorsal and lateral habitus: 92–93. Tylodinus gibbosus, 94–95. Tylodinus kissingeri, 96–97. Tylodinus pinguis. 98. Median lobe apex of: a) Tylodinus elongatus, b) Tylodinus exiguus, c) Tylodinus pseudocavicrus, d) Tylodinus rufus, e) Tylodinus rufus (squashed), 99. Tylodinus zilchi, ventral and lateral habitus of median lobe." figureDoi="http://doi.org/10.5281/zenodo.4913789" httpUri="https://zenodo.org/record/4913789/files/figure.png" pageId="7" pageNumber="8">Figure 98e</figureCitation>
). Female with tergite eight trapezoidal or constricted, apex invaginate, obtuse or slightly curved; sternite eight oblong, doliform, or trapezoidal, with the length of median membranous area from 1/3 as long as basal plate to subequal in length, spermatheca very broad to very shallowly curved, corpus convex to slightly convex, ramus and nodulus distinct or indistinct.
</paragraph>
</subSubSection>
<subSubSection id="02CD6562FF9C610BFF44D9BC24C9FA40" pageId="7" pageNumber="8" type="discussion">
<paragraph id="4A6836E9FF9C610BFF44D9BC200AFC70" blockId="7.[151,1437,151,1685]" pageId="7" pageNumber="8">
<emphasis id="78A3EAFBFF9C610BFF44D9BC2570FD61" bold="true" box="[199,335,654,679]" pageId="7" pageNumber="8">Comments.</emphasis>
Members of this genus can be confused with other genera of the subtribe 
<taxonomicName id="8DD74D6AFF9C610BFB15D9BC20C6FD61" box="[1174,1273,654,679]" class="Gastropoda" family="Tylodinidae" kingdom="Animalia" order="Umbraculida" pageId="7" pageNumber="8" phylum="Mollusca" rank="subTribe" subTribe="Tylodina">Tylodina</taxonomicName>
that also have tuberculate elytra, such as 
<taxonomicName id="8DD74D6AFF9C610BFE57D9872616FD08" box="[468,553,693,718]" class="Insecta" family="Curculionidae" genus="Tylodes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="78A3EAFBFF9C610BFE57D9872616FD08" box="[468,553,693,718]" italics="true" pageId="7" pageNumber="8">Tylodes</emphasis>
</taxonomicName>
, 
<taxonomicName id="8DD74D6AFF9C610BFDBFD9842684FD0B" authorityName="Faust" authorityYear="1896" box="[572,699,694,717]" class="Insecta" family="Curculionidae" genus="Xenosomus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="78A3EAFBFF9C610BFDBFD9842684FD0B" box="[572,699,694,717]" italics="true" pageId="7" pageNumber="8">Xenosomus</emphasis>
</taxonomicName>
, or 
<emphasis id="78A3EAFBFF9C610BFD71D9872794FD08" box="[754,939,693,718]" italics="true" pageId="7" pageNumber="8">
<taxonomicName id="8DD74D6AFF9C610BFD71D9872798FD08" authorityName="Faust" authorityYear="1896" box="[754,935,693,718]" class="Insecta" family="Curculionidae" genus="Phymatophosus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">Phymatophosus</taxonomicName>
.
</emphasis>
Species of 
<taxonomicName id="8DD74D6AFF9C610BFBB8D9872097FD08" authorityName="Champion" authorityYear="1905" box="[1083,1192,693,718]" class="Insecta" family="Curculionidae" genus="Tylodinus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="78A3EAFBFF9C610BFBB8D9872097FD08" box="[1083,1192,693,718]" italics="true" pageId="7" pageNumber="8">Tylodinus</emphasis>
</taxonomicName>
are most commonly confused with 
<taxonomicName id="8DD74D6AFF9C610BFEB8D9EE25B2FD33" authorityName="Schoenherr" authorityYear="1825" box="[315,397,732,757]" class="Insecta" family="Curculionidae" genus="Acalles" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="78A3EAFBFF9C610BFEB8D9EE25B2FD33" box="[315,397,732,757]" italics="true" pageId="7" pageNumber="8">Acalles</emphasis>
</taxonomicName>
, because so many species were erroneously originally placed in this genus (Champion 1904, 
<bibRefCitation id="2E464B18FF9C610BFF14D83025E5FCDD" author="O'Brien, C. W. &amp; Wibmer, G. J." box="[151,474,770,795]" pageId="7" pageNumber="8" pagination="1 - 382" refId="ref56933" refString="O'Brien, C. W. &amp; Wibmer, G. J. (1982) Annotated checklist of the weevils (Curculionidae sensu lato) of North America, Central America and the West Indies (Coleoptera: Curculionidae). Memoirs of the American Entomological Institute, 34, i - ix, 1 - 382." type="journal article" year="1982">O´Brien and Wibmer 1982</bibRefCitation>
); however, the metepisternum of these genera is not externally visible, and this character separates these genera from 
<taxonomicName id="8DD74D6AFF9C610BFDC0D81B2690FC84" authorityName="Champion" authorityYear="1905" box="[579,687,809,834]" class="Insecta" family="Curculionidae" genus="Tylodinus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="78A3EAFBFF9C610BFDC0D81B2690FC84" box="[579,687,809,834]" italics="true" pageId="7" pageNumber="8">Tylodinus</emphasis>
</taxonomicName>
. Fiedler (1940) placed 
<taxonomicName id="8DD74D6AFF9C610BFC34D81B201CFC84" authorityName="Champion" authorityYear="1905" box="[951,1059,809,834]" class="Insecta" family="Curculionidae" genus="Tylodinus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="78A3EAFBFF9C610BFC34D81B201CFC84" box="[951,1059,809,834]" italics="true" pageId="7" pageNumber="8">Tylodinus</emphasis>
</taxonomicName>
near 
<taxonomicName id="8DD74D6AFF9C610BFBE0D81820FCFC87" authorityName="Casey" authorityYear="1892" box="[1123,1219,810,833]" class="Insecta" family="Curculionidae" genus="Canistes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="78A3EAFBFF9C610BFBE0D81820FCFC87" box="[1123,1219,810,833]" italics="true" pageId="7" pageNumber="8">Canistes</emphasis>
</taxonomicName>
and 
<taxonomicName id="8DD74D6AFF9C610BFB79D81B21A3FC84" authorityName="Blatchley" authorityYear="1916" box="[1274,1436,809,834]" class="Insecta" family="Curculionidae" genus="Pseudoacalles" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="78A3EAFBFF9C610BFB79D81B21A3FC84" box="[1274,1436,809,834]" italics="true" pageId="7" pageNumber="8">Pseudoacalles</emphasis>
</taxonomicName>
because these genera have the metepisternum distinctly visible, but other characters given in the diagnosis will separate 
<taxonomicName id="8DD74D6AFF9C610BFF79D8442559FC49" authorityName="Champion" authorityYear="1905" box="[250,358,886,911]" class="Insecta" family="Curculionidae" genus="Tylodinus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="78A3EAFBFF9C610BFF79D8442559FC49" box="[250,358,886,911]" italics="true" pageId="7" pageNumber="8">Tylodinus</emphasis>
</taxonomicName>
from 
<taxonomicName id="8DD74D6AFF9C610BFE2FD8452633FC48" authorityName="Casey" authorityYear="1892" box="[428,524,887,910]" class="Insecta" family="Curculionidae" genus="Canistes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="78A3EAFBFF9C610BFE2FD8452633FC48" box="[428,524,887,910]" italics="true" pageId="7" pageNumber="8">Canistes</emphasis>
</taxonomicName>
and 
<taxonomicName id="8DD74D6AFF9C610BFDC7D84426D9FC49" authorityName="Blatchley" authorityYear="1916" box="[580,742,886,911]" class="Insecta" family="Curculionidae" genus="Pseudoacalles" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="78A3EAFBFF9C610BFDC7D84426D9FC49" box="[580,742,886,911]" italics="true" pageId="7" pageNumber="8">Pseudoacalles</emphasis>
</taxonomicName>
. In a forthcoming paper on the phylogenetic relationships of 
<taxonomicName id="8DD74D6AFF9C610BFF14D8AF253CFC70" authorityName="Champion" authorityYear="1905" box="[151,259,925,950]" class="Insecta" family="Curculionidae" genus="Tylodinus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="78A3EAFBFF9C610BFF14D8AF253CFC70" box="[151,259,925,950]" italics="true" pageId="7" pageNumber="8">Tylodinus</emphasis>
</taxonomicName>
, a number of new genera, closely related to 
<taxonomicName id="8DD74D6AFF9C610BFD77D8AF275FFC70" authorityName="Champion" authorityYear="1905" box="[756,864,925,950]" class="Insecta" family="Curculionidae" genus="Tylodinus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="78A3EAFBFF9C610BFD77D8AF275FFC70" box="[756,864,925,950]" italics="true" pageId="7" pageNumber="8">Tylodinus</emphasis>
</taxonomicName>
, will be described.
</paragraph>
<paragraph id="4A6836E9FF9C610BFF44D8F620D8FB89" blockId="7.[151,1437,151,1685]" pageId="7" pageNumber="8">
<emphasis id="78A3EAFBFF9C610BFF44D8F62609FC1B" bold="true" box="[199,566,964,989]" pageId="7" pageNumber="8">Derivation of generic name. C</emphasis>
hampion (1905) did not indicate the derivation of the name 
<taxonomicName id="8DD74D6AFF9C610BFB69D8F62168FC1B" authorityName="Champion" authorityYear="1905" box="[1258,1367,964,989]" class="Insecta" family="Curculionidae" genus="Tylodinus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="78A3EAFBFF9C610BFB69D8F62168FC1B" box="[1258,1367,964,989]" italics="true" pageId="7" pageNumber="8">Tylodinus</emphasis>
</taxonomicName>
but it appears to be derived from the combination of two words: 
<taxonomicName id="8DD74D6AFF9C610BFCCBD8D827A0FBC5" box="[840,927,1002,1027]" class="Insecta" family="Curculionidae" genus="Tylodes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="78A3EAFBFF9C610BFCCBD8D827A0FBC5" box="[840,927,1002,1027]" italics="true" pageId="7" pageNumber="8">Tylodes</emphasis>
</taxonomicName>
= derived from Greek “ 
<emphasis id="78A3EAFBFF9C610BFB38D8D82135FBC5" box="[1211,1290,1002,1027]" italics="true" pageId="7" pageNumber="8">tylodes</emphasis>
” = tubercle, probably referring to the tubercles on the body of the genus 
<taxonomicName id="8DD74D6AFF9C610BFCB1DF2327B8FBEC" box="[818,903,1041,1066]" class="Insecta" family="Curculionidae" genus="Tylodes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="78A3EAFBFF9C610BFCB1DF2327B8FBEC" box="[818,903,1041,1066]" italics="true" pageId="7" pageNumber="8">Tylodes</emphasis>
</taxonomicName>
and from the Latin suffix 
<emphasis id="78A3EAFBFF9C610BFB2FDF2020E5FBEF" box="[1196,1242,1042,1065]" italics="true" pageId="7" pageNumber="8">inus</emphasis>
= like; 
<taxonomicName id="8DD74D6AFF9C610BFAACDF2321A3FBEC" authorityName="Champion" authorityYear="1905" box="[1327,1436,1041,1066]" class="Insecta" family="Curculionidae" genus="Tylodinus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="78A3EAFBFF9C610BFAACDF2321A3FBEC" box="[1327,1436,1041,1066]" italics="true" pageId="7" pageNumber="8">Tylodinus</emphasis>
</taxonomicName>
= like a 
<taxonomicName id="8DD74D6AFF9C610BFF71DF052578FB96" box="[242,327,1079,1104]" class="Insecta" family="Curculionidae" genus="Tylodes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="78A3EAFBFF9C610BFF71DF052578FB96" box="[242,327,1079,1104]" italics="true" pageId="7" pageNumber="8">Tylodes</emphasis>
</taxonomicName>
. The name refers apparently to possible confusion between 
<taxonomicName id="8DD74D6AFF9C610BFC6BDF05206BFB96" authorityName="Champion" authorityYear="1905" box="[1000,1108,1079,1104]" class="Insecta" family="Curculionidae" genus="Tylodinus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="78A3EAFBFF9C610BFC6BDF05206BFB96" box="[1000,1108,1079,1104]" italics="true" pageId="7" pageNumber="8">Tylodinus</emphasis>
</taxonomicName>
and 
<taxonomicName id="8DD74D6AFF9C610BFB08DF0520DFFB96" box="[1163,1248,1079,1104]" class="Insecta" family="Curculionidae" genus="Tylodes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="78A3EAFBFF9C610BFB08DF0520DFFB96" box="[1163,1248,1079,1104]" italics="true" pageId="7" pageNumber="8">Tylodes</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="4A6836E9FF9C610BFF44DF6C24C9FA40" blockId="7.[151,1437,151,1685]" pageId="7" pageNumber="8">
<emphasis id="78A3EAFBFF9C610BFF44DF6C25BAFBB1" bold="true" box="[199,389,1118,1143]" pageId="7" pageNumber="8">Natural history.</emphasis>
Very few details are available on the natural history of any species of 
<taxonomicName id="8DD74D6AFF9C610BFB18DF6C2138FBB1" authorityName="Champion" authorityYear="1905" box="[1179,1287,1118,1143]" class="Insecta" family="Curculionidae" genus="Tylodinus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="78A3EAFBFF9C610BFB18DF6C2138FBB1" box="[1179,1287,1118,1143]" italics="true" pageId="7" pageNumber="8">Tylodinus</emphasis>
</taxonomicName>
. Adults have been collected almost exclusively in Berlese or Winkler extractions of sifted leaf litter and other ground substrate debris. Some specimens have been collected beating dead vegetation and numbers of larger-sized species have been hand collected on dead wood, apparently in association with 
<taxonomicName id="8DD74D6AFF9C610BFC22DFE02018FB2D" box="[929,1063,1234,1259]" class="Sordariomycetes" family="Xylariaceae" kingdom="Fungi" order="Xylariales" pageId="7" pageNumber="8" phylum="Ascomycota" rank="family">Xylariaceae</taxonomicName>
fungi, especially those with the crustose fruiting bodies, which the black, tuberculate body form of the weevil greatly resembles. Immature stages are unknown. Given what is known of the habits of related taxa, it is likely that the larvae mine in dead, decaying wood or other plant debris on the forest floor. Nearly all species are associated with a variety of montane wet forest habitats.
</paragraph>
</subSubSection>
<subSubSection id="02CD6562FF9C6107FF44DEA625C3FB1C" lastPageId="11" lastPageNumber="12" pageId="7" pageNumber="8" type="distribution">
<paragraph id="4A6836E9FF9C610BFF44DEA62603F953" blockId="7.[151,1437,151,1685]" pageId="7" pageNumber="8">
<emphasis id="78A3EAFBFF9C610BFF44DEA6255FFA6B" bold="true" box="[199,352,1428,1453]" pageId="7" pageNumber="8">Distribution.</emphasis>
<taxonomicName id="8DD74D6AFF9C610BFEEADEA625EAFA6B" authorityName="Champion" authorityYear="1905" box="[361,469,1428,1453]" class="Insecta" family="Curculionidae" genus="Tylodinus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="78A3EAFBFF9C610BFEEADEA625EAFA6B" box="[361,469,1428,1453]" italics="true" pageId="7" pageNumber="8">Tylodinus</emphasis>
</taxonomicName>
occur throughout the Sierra Madre Oriental from Tamaulipas south to 
<collectingCountry id="32C07679FF9C610BFA8BDEA6215EFA6D" box="[1288,1377,1428,1451]" name="Panama" pageId="7" pageNumber="8">Panama</collectingCountry>
with many species undescribed. At present, 20 species have been described with an additional 32 species described herein, and numerous other new species from other geographic areas are known. Thus, at present there 52 species known to date, 40 are known to occur in 
<collectingCountry id="32C07679FF9C610BFDDFDD35268EF9E6" box="[604,689,1543,1568]" name="Mexico" pageId="7" pageNumber="8">México</collectingCountry>
and of these, 38 species have only been recorded from 
<collectingCountry id="32C07679FF9C610BFA9BDD352152F9E6" box="[1304,1389,1543,1568]" name="Mexico" pageId="7" pageNumber="8">México</collectingCountry>
and two extend from 
<collectingCountry id="32C07679FF9C610BFEDBDD1C2592F981" box="[344,429,1582,1607]" name="Mexico" pageId="7" pageNumber="8">México</collectingCountry>
south into Central America. As discussed, 
<taxonomicName id="8DD74D6AFF9C610BFC10DD1C2067F981" box="[915,1112,1582,1607]" class="Insecta" family="Curculionidae" genus="Tylodinus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="78A3EAFBFF9C610BFC10DD1C27C0F981" box="[915,1023,1582,1607]" italics="true" pageId="7" pageNumber="8">Tylodinus</emphasis>
species
</taxonomicName>
in 
<collectingRegion id="8813F80BFF9C610BFBFFDD1C20E9F981" box="[1148,1238,1582,1607]" country="Mexico" name="Chiapas" pageId="7" pageNumber="8">Chiapas</collectingRegion>
are well sampled and were collected at elevations from 
<quantity id="8D2F9B0CFF9C610BFDE8DD67270EF9AB" box="[619,817,1621,1646]" metricMagnitude="3" metricUnit="m" metricValue="2.025" metricValueMax="2.9699999999999998" metricValueMin="1.08" pageId="7" pageNumber="8" unit="m" value="2025.0" valueMax="2970.0" valueMin="1080.0">1080 to 2970 m</quantity>
in 20 municipalities; the species numbers in these municipalities are detailed in 
<tableCitation id="07550352FF9C610BFE61DD4E2609F953" box="[482,566,1660,1685]" captionStart="TABLE 1" captionStartId="11.[151,244,151,176]" captionTargetPageId="11" captionText="TABLE 1. Number of Tylodinus species by municipality of Chiapas, Mexico." httpUri="http://table.plazi.org/id/1EA86661FF906107FF14DBA527C1FF76" pageId="7" pageNumber="8" tableUuid="1EA86661FF906107FF14DBA527C1FF76">Table 1</tableCitation>
.
</paragraph>
<caption id="1EA86661FF936104FF14DDFD25C4F836" ID-DOI="http://doi.org/10.5281/zenodo.4913763" ID-Zenodo-Dep="4913763" httpUri="https://zenodo.org/record/4913763/files/figure.png" pageId="8" pageNumber="9" startId="8.[151,264,1743,1765]" targetBox="[293,1295,205,1711]" targetPageId="8">
<paragraph id="4A6836E9FF936104FF14DDFD25C4F836" blockId="8.[151,1437,1743,2032]" pageId="8" pageNumber="9">
<emphasis id="78A3EAFBFF936104FF14DDFD2546F923" bold="true" box="[151,377,1743,1765]" pageId="8" pageNumber="9">FIGURES 1–11. 1–5.</emphasis>
Dorsal outline: 
<emphasis id="78A3EAFBFF936104FD9FDDFD2610F923" bold="true" box="[540,559,1743,1765]" pageId="8" pageNumber="9">1.</emphasis>
Generalized 
<taxonomicName id="8DD74D6AFF936104FD35DDFD2725F923" authorityName="Champion" authorityYear="1905" box="[694,794,1743,1765]" class="Insecta" family="Curculionidae" genus="Tylodinus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="78A3EAFBFF936104FD35DDFD2725F923" box="[694,794,1743,1765]" italics="true" pageId="8" pageNumber="9">Tylodinus</emphasis>
</taxonomicName>
, indicating terminology used in this study. Key: letters indicate areas of pronotum. A, anterior area; B, median area; C, posterior area; a, anterior of lateral area; b, median section of lateral area; c, median section of dorsal area; d, anterior section of dorsal area; e, posterior section of lateral area; f, posterior section of dorsal area. 
<emphasis id="78A3EAFBFF936104FE92DC01251BF88F" bold="true" box="[273,292,1843,1865]" pageId="8" pageNumber="9">2.</emphasis>
<taxonomicName id="8DD74D6AFF936104FEA9DC0125CAF88F" authorityName="Luna-Cozar &amp; Anderson &amp; Jones &amp; León-Cortés" authorityYear="2014" box="[298,501,1843,1865]" class="Insecta" family="Curculionidae" genus="Tylodinus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="variabilis">
<emphasis id="78A3EAFBFF936104FEA9DC0125CAF88F" box="[298,501,1843,1865]" italics="true" pageId="8" pageNumber="9">Tylodinus variabilis</emphasis>
</taxonomicName>
, note elytra basal margin straight. 
<emphasis id="78A3EAFBFF936104FCD0DC012759F88F" bold="true" box="[851,870,1843,1865]" pageId="8" pageNumber="9">3.</emphasis>
<taxonomicName id="8DD74D6AFF936104FCEFDC01202EF88F" authorityName="Luna-Cozar &amp; Anderson &amp; Jones &amp; León-Cortés" authorityYear="2014" box="[876,1041,1843,1865]" class="Insecta" family="Curculionidae" genus="Tylodinus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="ixchel">
<emphasis id="78A3EAFBFF936104FCEFDC01202EF88F" box="[876,1041,1843,1865]" italics="true" pageId="8" pageNumber="9">Tylodinus ixchel</emphasis>
</taxonomicName>
, note elytra basal margin slightly sinuate. 
<emphasis id="78A3EAFBFF936104FF40DC6624E9F8AC" bold="true" box="[195,214,1876,1898]" pageId="8" pageNumber="9">4.</emphasis>
<taxonomicName id="8DD74D6AFF936104FF5DDC66258DF8AC" authorityName="Luna-Cozar &amp; Anderson &amp; Jones &amp; León-Cortés" authorityYear="2014" box="[222,434,1876,1898]" class="Insecta" family="Curculionidae" genus="Tylodinus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="immundus">
<emphasis id="78A3EAFBFF936104FF5DDC66258DF8AC" box="[222,434,1876,1898]" italics="true" pageId="8" pageNumber="9">Tylodinus immundus</emphasis>
</taxonomicName>
, note elytral basal margin sinuate. 
<emphasis id="78A3EAFBFF936104FC9CDC66270DF8AC" bold="true" box="[799,818,1876,1898]" pageId="8" pageNumber="9">5.</emphasis>
<taxonomicName id="8DD74D6AFF936104FCB9DC662037F8AC" authorityName="Luna-Cozar &amp; Anderson &amp; Jones &amp; León-Cortés" authorityYear="2014" box="[826,1032,1876,1898]" class="Insecta" family="Curculionidae" genus="Tylodinus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="elongatus">
<emphasis id="78A3EAFBFF936104FCB9DC662037F8AC" box="[826,1032,1876,1898]" italics="true" pageId="8" pageNumber="9">Tylodinus elongatus</emphasis>
</taxonomicName>
, note elytra basal margin very sinuate. 
<emphasis id="78A3EAFBFF936104FF14DC4424EFF84A" bold="true" box="[151,208,1910,1932]" pageId="8" pageNumber="9">6–11.</emphasis>
Lateral outline: 
<emphasis id="78A3EAFBFF936104FEF5DC4425B6F84A" bold="true" box="[374,393,1910,1932]" pageId="8" pageNumber="9">6.</emphasis>
<taxonomicName id="8DD74D6AFF936104FE0CDC44264CF84A" authorityName="Luna-Cozar &amp; Anderson &amp; Jones &amp; León-Cortés" authorityYear="2014" box="[399,627,1910,1932]" class="Insecta" family="Curculionidae" genus="Tylodinus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="complicatus">
<emphasis id="78A3EAFBFF936104FE0CDC44264CF84A" box="[399,627,1910,1932]" italics="true" pageId="8" pageNumber="9">Tylodinus complicatus</emphasis>
</taxonomicName>
, male, note ventrites 3 
<emphasis id="78A3EAFBFF936104FCDBDC44275BF84A" bold="true" box="[856,868,1910,1932]" pageId="8" pageNumber="9">–</emphasis>
5 in same plane as ventrites 1 
<emphasis id="78A3EAFBFF936104FB0FDC4420A6F84A" bold="true" box="[1164,1177,1910,1932]" pageId="8" pageNumber="9">–</emphasis>
2. 
<emphasis id="78A3EAFBFF936104FB31DC4420FAF84A" bold="true" box="[1202,1221,1910,1932]" pageId="8" pageNumber="9">7.</emphasis>
<taxonomicName id="8DD74D6AFF936104FB48DC4421AAF84A" authorityName="Luna-Cozar &amp; Anderson &amp; Jones &amp; León-Cortés" authorityYear="2014" box="[1227,1429,1910,1932]" class="Insecta" family="Curculionidae" genus="Tylodinus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="variabilis">
<emphasis id="78A3EAFBFF936104FB48DC4421AAF84A" box="[1227,1429,1910,1932]" italics="true" pageId="8" pageNumber="9">Tylodinus variabilis</emphasis>
</taxonomicName>
, male, note ventrites 3 
<emphasis id="78A3EAFBFF936104FEF0DCA52540F86B" bold="true" box="[371,383,1943,1965]" pageId="8" pageNumber="9">–</emphasis>
5 not in same plane as ventrites 1 
<emphasis id="78A3EAFBFF936104FD57DCA526DEF86B" bold="true" box="[724,737,1943,1965]" pageId="8" pageNumber="9">–</emphasis>
2. 
<emphasis id="78A3EAFBFF936104FD78DCA52732F86B" bold="true" box="[763,781,1943,1965]" pageId="8" pageNumber="9">8.</emphasis>
<taxonomicName id="8DD74D6AFF936104FC97DCA527C7F86B" authorityName="Luna-Cozar &amp; Anderson &amp; Jones &amp; León-Cortés" authorityYear="2014" box="[788,1016,1943,1965]" class="Insecta" family="Curculionidae" genus="Tylodinus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="coapillensis">
<emphasis id="78A3EAFBFF936104FC97DCA527C7F86B" box="[788,1016,1943,1965]" italics="true" pageId="8" pageNumber="9">Tylodinus coapillensis</emphasis>
</taxonomicName>
, female, note ventrites 1 
<emphasis id="78A3EAFBFF936104FB76DCA5213EF86B" bold="true" box="[1269,1281,1943,1965]" pageId="8" pageNumber="9">–</emphasis>
2 strongly convex, ventrites 3 
<emphasis id="78A3EAFBFF936104FEB0DC8B257FF809" bold="true" box="[307,320,1977,1999]" pageId="8" pageNumber="9">–</emphasis>
5 not in same plane as ventrites 1 
<emphasis id="78A3EAFBFF936104FD11DC8B26A0F809" bold="true" box="[658,671,1977,1999]" pageId="8" pageNumber="9">–</emphasis>
2. 
<emphasis id="78A3EAFBFF936104FD3BDC8B26F4F809" bold="true" box="[696,715,1977,1999]" pageId="8" pageNumber="9">9.</emphasis>
<taxonomicName id="8DD74D6AFF936104FD51DC8B274CF809" authorityName="Kuschel" authorityYear="1956" box="[722,883,1977,1999]" class="Insecta" family="Curculionidae" genus="Tylodinus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="zilchi">
<emphasis id="78A3EAFBFF936104FD51DC8B274CF809" box="[722,883,1977,1999]" italics="true" pageId="8" pageNumber="9">Tylodinus zilchi</emphasis>
</taxonomicName>
(schematic). 
<emphasis id="78A3EAFBFF936104FC7FDC8B2023F809" bold="true" box="[1020,1052,1977,1999]" pageId="8" pageNumber="9">10.</emphasis>
<taxonomicName id="8DD74D6AFF936104FBA0DC8B2120F809" authorityName="Luna-Cozar &amp; Anderson &amp; Jones &amp; León-Cortés" authorityYear="2014" box="[1059,1311,1977,1999]" class="Insecta" family="Curculionidae" genus="Tylodinus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="sepulturaensis">
<emphasis id="78A3EAFBFF936104FBA0DC8B2120F809" box="[1059,1311,1977,1999]" italics="true" pageId="8" pageNumber="9">Tylodinus sepulturaensis</emphasis>
</taxonomicName>
(schematic) 
<emphasis id="78A3EAFBFF936104FF14DCE82489F836" bold="true" box="[151,182,2010,2032]" pageId="8" pageNumber="9">11.</emphasis>
<taxonomicName id="8DD74D6AFF936104FF3EDCE82548F836" authorityName="Luna-Cozar &amp; Anderson &amp; Jones &amp; León-Cortés" authorityYear="2014" box="[189,375,2010,2032]" class="Insecta" family="Curculionidae" genus="Tylodinus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="rugosus">
<emphasis id="78A3EAFBFF936104FF3EDCE82548F836" box="[189,375,2010,2032]" italics="true" pageId="8" pageNumber="9">Tylodinus rugosus</emphasis>
</taxonomicName>
(schematic).
</paragraph>
</caption>
<caption id="1EA86661FF926105FF14DC4D2596F837" ID-DOI="http://doi.org/10.5281/zenodo.4913765" ID-Zenodo-Dep="4913765" httpUri="https://zenodo.org/record/4913765/files/figure.png" pageId="9" pageNumber="10" startId="9.[151,264,1919,1941]" targetBox="[219,1377,209,1898]" targetPageId="9">
<paragraph id="4A6836E9FF926105FF14DC4D2596F837" blockId="9.[151,1436,1919,2033]" pageId="9" pageNumber="10">
<emphasis id="78A3EAFBFF926105FF14DC4D2545F853" bold="true" box="[151,378,1919,1941]" pageId="9" pageNumber="10">FIGURES 12–19. 12.</emphasis>
<taxonomicName id="8DD74D6AFF926105FE02DC4D2604F853" authorityName="Luna-Cozar &amp; Anderson &amp; Jones &amp; León-Cortés" authorityYear="2014" box="[385,571,1919,1941]" class="Insecta" family="Curculionidae" genus="Tylodinus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="pusillus">
<emphasis id="78A3EAFBFF926105FE02DC4D2604F853" box="[385,571,1919,1941]" italics="true" pageId="9" pageNumber="10">Tylodinus pusillus</emphasis>
</taxonomicName>
head a) male, b) female. 
<emphasis id="78A3EAFBFF926105FCBCDC4D27BAF853" bold="true" box="[831,901,1919,1941]" pageId="9" pageNumber="10">13–14.</emphasis>
Dorsal habitus of pronotum showing the lateral side of pronotum 
<emphasis id="78A3EAFBFF926105FE9EDCAC2502F872" bold="true" box="[285,317,1950,1972]" pageId="9" pageNumber="10">13.</emphasis>
<taxonomicName id="8DD74D6AFF926105FEC6DCAC262BF872" authorityName="Luna-Cozar &amp; Anderson &amp; Jones &amp; León-Cortés" authorityYear="2014" box="[325,532,1950,1972]" class="Insecta" family="Curculionidae" genus="Tylodinus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="andersoni">
<emphasis id="78A3EAFBFF926105FEC6DCAC262BF872" box="[325,532,1950,1972]" italics="true" pageId="9" pageNumber="10">Tylodinus andersoni</emphasis>
</taxonomicName>
, 
<emphasis id="78A3EAFBFF926105FDA1DCAC267DF872" bold="true" box="[546,578,1950,1972]" pageId="9" pageNumber="10">14.</emphasis>
<taxonomicName id="8DD74D6AFF926105FDCADCAC2701F872" authorityName="Luna-Cozar &amp; Anderson &amp; Jones &amp; León-Cortés" authorityYear="2014" box="[585,830,1950,1972]" class="Insecta" family="Curculionidae" genus="Tylodinus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="porvenirensis">
<emphasis id="78A3EAFBFF926105FDCADCAC2701F872" box="[585,830,1950,1972]" italics="true" pageId="9" pageNumber="10">Tylodinus porvenirensis</emphasis>
</taxonomicName>
. 
<emphasis id="78A3EAFBFF926105FCCFDCAC27ADF872" bold="true" box="[844,914,1950,1972]" pageId="9" pageNumber="10">15–17.</emphasis>
Dorsal habitus: 
<emphasis id="78A3EAFBFF926105FBBFDCAC2064F872" bold="true" box="[1084,1115,1950,1972]" pageId="9" pageNumber="10">15.</emphasis>
<emphasis id="78A3EAFBFF926105FBE0DCAC211AF872" box="[1123,1317,1950,1972]" italics="true" pageId="9" pageNumber="10">
<taxonomicName id="8DD74D6AFF926105FBE0DCAC211EF872" authorityName="Luna-Cozar &amp; Anderson &amp; Jones &amp; León-Cortés" authorityYear="2014" box="[1123,1313,1950,1972]" class="Insecta" family="Curculionidae" genus="Tylodinus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="rugosus">Tylodinus rugosus</taxonomicName>
,
</emphasis>
<emphasis id="78A3EAFBFF926105FAAFDCAC2174F872" bold="true" box="[1324,1355,1950,1972]" pageId="9" pageNumber="10">16.</emphasis>
<taxonomicName id="8DD74D6AFF926105FAD0DCAC252FF814" authorityName="Luna-Cozar &amp; Anderson &amp; Jones &amp; León-Cortés" authorityYear="2014" class="Insecta" family="Curculionidae" genus="Tylodinus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="exiguus">
<emphasis id="78A3EAFBFF926105FAD0DCAC252FF814" italics="true" pageId="9" pageNumber="10">Tylodinus exiguus</emphasis>
</taxonomicName>
, 
<emphasis id="78A3EAFBFF926105FE9CDC8E2501F814" bold="true" box="[287,318,1980,2002]" pageId="9" pageNumber="10">17.</emphasis>
<taxonomicName id="8DD74D6AFF926105FEC4DC8E25CFF814" authorityName="Luna-Cozar &amp; Anderson &amp; Jones &amp; León-Cortés" authorityYear="2014" box="[327,496,1980,2002]" class="Insecta" family="Curculionidae" genus="Tylodinus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="jonesi">
<emphasis id="78A3EAFBFF926105FEC4DC8E25CFF814" box="[327,496,1980,2002]" italics="true" pageId="9" pageNumber="10">Tylodinus jonesi</emphasis>
</taxonomicName>
. 
<emphasis id="78A3EAFBFF926105FE7CDC8E2621F814" bold="true" box="[511,542,1980,2002]" pageId="9" pageNumber="10">18.</emphasis>
<taxonomicName id="8DD74D6AFF926105FDA4DC8E2732F814" authorityName="Luna-Cozar &amp; Anderson &amp; Jones &amp; León-Cortés" authorityYear="2014" box="[551,781,1980,2002]" class="Insecta" family="Curculionidae" genus="Tylodinus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="coapillensis">
<emphasis id="78A3EAFBFF926105FDA4DC8E2732F814" box="[551,781,1980,2002]" italics="true" pageId="9" pageNumber="10">Tylodinus coapillensis</emphasis>
</taxonomicName>
note tubercle on elytra declivity base, 
<emphasis id="78A3EAFBFF926105FB26DC8E20FAF814" bold="true" box="[1189,1221,1980,2002]" pageId="9" pageNumber="10">19.</emphasis>
<taxonomicName id="8DD74D6AFF926105FB4EDC8E21A3F814" authorityName="Luna-Cozar &amp; Anderson &amp; Jones &amp; León-Cortés" authorityYear="2014" box="[1229,1436,1980,2002]" class="Insecta" family="Curculionidae" genus="Tylodinus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="elongatus">
<emphasis id="78A3EAFBFF926105FB4EDC8E21A3F814" box="[1229,1436,1980,2002]" italics="true" pageId="9" pageNumber="10">Tylodinus elongatus</emphasis>
</taxonomicName>
showing metasternal tooth.
</paragraph>
</caption>
<caption id="1EA86661FF916106FF14DC482171F814" ID-DOI="http://doi.org/10.5281/zenodo.4913767" ID-Zenodo-Dep="4913767" httpUri="https://zenodo.org/record/4913767/files/figure.png" pageId="10" pageNumber="11" startId="10.[151,264,1914,1936]" targetBox="[217,1351,197,1862]" targetPageId="10">
<paragraph id="4A6836E9FF916106FF14DC482171F814" blockId="10.[151,1436,1914,2002]" pageId="10" pageNumber="11">
<emphasis id="78A3EAFBFF916106FF14DC48259FF856" bold="true" box="[151,416,1914,1936]" pageId="10" pageNumber="11">FIGURES 20–27. 20–21.</emphasis>
<taxonomicName id="8DD74D6AFF916106FE26DC4826B9F856" authorityName="Luna-Cozar &amp; Anderson &amp; Jones &amp; León-Cortés" authorityYear="2014" box="[421,646,1914,1936]" class="Insecta" family="Curculionidae" genus="Tylodinus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="spiniventris">
<emphasis id="78A3EAFBFF916106FE26DC4826B9F856" box="[421,646,1914,1936]" italics="true" pageId="10" pageNumber="11">Tylodinus spiniventris</emphasis>
</taxonomicName>
: 
<emphasis id="78A3EAFBFF916106FD10DC48268CF856" bold="true" box="[659,691,1914,1936]" pageId="10" pageNumber="11">20.</emphasis>
meso- and metathorax (male) 
<emphasis id="78A3EAFBFF916106FC6BDC482037F856" bold="true" box="[1000,1032,1914,1936]" pageId="10" pageNumber="11">21.</emphasis>
ventrite 5 (female), 
<emphasis id="78A3EAFBFF916106FB5BDC4820C7F856" bold="true" box="[1240,1272,1914,1936]" pageId="10" pageNumber="11">22.</emphasis>
<taxonomicName id="8DD74D6AFF916106FB7DDC4824F1F877" authorityName="Luna-Cozar &amp; Anderson &amp; Jones &amp; León-Cortés" authorityYear="2014" class="Insecta" family="Curculionidae" genus="Tylodinus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="elongatus">
<emphasis id="78A3EAFBFF916106FB7DDC4824F1F877" italics="true" pageId="10" pageNumber="11">Tylodinus elongatus</emphasis>
</taxonomicName>
ventrite 5 (female), 
<emphasis id="78A3EAFBFF916106FE29DCA925F5F877" bold="true" box="[426,458,1947,1969]" pageId="10" pageNumber="11">23.</emphasis>
<taxonomicName id="8DD74D6AFF916106FE50DCA92684F877" authorityName="Luna-Cozar &amp; Anderson &amp; Jones &amp; León-Cortés" authorityYear="2014" box="[467,699,1947,1969]" class="Insecta" family="Curculionidae" genus="Tylodinus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="triumforium">
<emphasis id="78A3EAFBFF916106FE50DCA92684F877" box="[467,699,1947,1969]" italics="true" pageId="10" pageNumber="11">Tylodinus triumforium</emphasis>
</taxonomicName>
, ventrite 2 (male). 
<emphasis id="78A3EAFBFF916106FC0ADCA92796F877" bold="true" box="[905,937,1947,1969]" pageId="10" pageNumber="11">24.</emphasis>
<taxonomicName id="8DD74D6AFF916106FC31DCA920A5F877" authorityName="Luna-Cozar &amp; Anderson &amp; Jones &amp; León-Cortés" authorityYear="2014" box="[946,1178,1947,1969]" class="Insecta" family="Curculionidae" genus="Tylodinus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="complicatus">
<emphasis id="78A3EAFBFF916106FC31DCA920A5F877" box="[946,1178,1947,1969]" italics="true" pageId="10" pageNumber="11">Tylodinus complicatus</emphasis>
</taxonomicName>
, ventrite 2 (female). 
<emphasis id="78A3EAFBFF916106FAFFDCA921A3F877" bold="true" box="[1404,1436,1947,1969]" pageId="10" pageNumber="11">25.</emphasis>
<taxonomicName id="8DD74D6AFF916106FF14DC8E2544F814" authorityName="Luna-Cozar &amp; Anderson &amp; Jones &amp; León-Cortés" authorityYear="2014" box="[151,379,1980,2002]" class="Insecta" family="Curculionidae" genus="Tylodinus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="complicatus">
<emphasis id="78A3EAFBFF916106FF14DC8E2544F814" box="[151,379,1980,2002]" italics="true" pageId="10" pageNumber="11">Tylodinus complicatus</emphasis>
</taxonomicName>
, ventrite 2 (male), 
<emphasis id="78A3EAFBFF916106FDBEDC8E2662F814" bold="true" box="[573,605,1980,2002]" pageId="10" pageNumber="11">26.</emphasis>
Hind tibia of 
<taxonomicName id="8DD74D6AFF916106FD68DC8E27F0F814" authorityName="Luna-Cozar &amp; Anderson &amp; Jones &amp; León-Cortés" authorityYear="2014" box="[747,975,1980,2002]" class="Insecta" family="Curculionidae" genus="Tylodinus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="triumforium">
<emphasis id="78A3EAFBFF916106FD68DC8E27F0F814" box="[747,975,1980,2002]" italics="true" pageId="10" pageNumber="11">Tylodinus triumforium</emphasis>
</taxonomicName>
, 
<emphasis id="78A3EAFBFF916106FC5FDC8E27C3F814" bold="true" box="[988,1020,1980,2002]" pageId="10" pageNumber="11">27.</emphasis>
Front tibia of 
<taxonomicName id="8DD74D6AFF916106FB0DDC8E2177F814" authorityName="Luna-Cozar &amp; Anderson &amp; Jones &amp; León-Cortés" authorityYear="2014" box="[1166,1352,1980,2002]" class="Insecta" family="Curculionidae" genus="Tylodinus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="rugosus">
<emphasis id="78A3EAFBFF916106FB0DDC8E2177F814" box="[1166,1352,1980,2002]" italics="true" pageId="10" pageNumber="11">Tylodinus rugosus</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<caption id="1EA86661FF906107FF14DBA527C1FF76" ID-Table-UUID="1EA86661FF906107FF14DBA527C1FF76" box="[151,1022,151,176]" httpUri="http://table.plazi.org/id/1EA86661FF906107FF14DBA527C1FF76" pageId="11" pageNumber="12" startId="11.[151,244,151,176]" targetBox="[157,1250,245,1148]" targetIsTable="true" targetPageId="11">
<paragraph id="4A6836E9FF906107FF14DBA527C1FF76" blockId="11.[151,1022,151,176]" box="[151,1022,151,176]" pageId="11" pageNumber="12">
<emphasis id="78A3EAFBFF906107FF14DBA5252FFF76" bold="true" box="[151,272,151,176]" pageId="11" pageNumber="12">TABLE 1.</emphasis>
Number of 
<taxonomicName id="8DD74D6AFF906107FE1BDBA52663FF76" box="[408,604,151,176]" class="Insecta" family="Curculionidae" genus="Tylodinus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="78A3EAFBFF906107FE1BDBA5263AFF76" box="[408,517,151,176]" italics="true" pageId="11" pageNumber="12">Tylodinus</emphasis>
species
</taxonomicName>
by municipality of Chiapas, Mexico.
</paragraph>
</caption>
<paragraph id="4A6836E9FF906107FF1EDBC727EAFBBA" pageId="11" pageNumber="12">
<table id="38D7C449FF909EF3FF1EDBC720DDFBBA" box="[157,1250,245,1148]" gridcols="3" gridrows="21" pageId="11" pageNumber="12">
<tr id="F4E734ABFF909EF3FF1EDBC720DDFECD" box="[157,1250,245,267]" gridrow="0" pageId="11" pageNumber="12">
<th id="B7365DD7FF909EF3FF1EDBC725FEFECD" box="[157,449,245,267]" gridcol="0" gridrow="0" pageId="11" pageNumber="12">Municipality</th>
<th id="B7365DD7FF909EF3FDF1DBC7270FFECD" box="[626,816,245,267]" gridcol="1" gridrow="0" pageId="11" pageNumber="12">Number of species</th>
<th id="B7365DD7FF909EF3FC4BDBC720DDFECD" box="[968,1250,245,267]" gridcol="2" gridrow="0" pageId="11" pageNumber="12">Number of endemic species</th>
</tr>
<tr id="F4E734ABFF909EF3FF1EDA1320DDFEF1" box="[157,1250,289,311]" gridrow="1" pageId="11" pageNumber="12">
<th id="B7365DD7FF909EF3FF1EDA1325FEFEF1" box="[157,449,289,311]" gridcol="0" gridrow="1" pageId="11" pageNumber="12">Ángel Albino Corzo</th>
<td id="B7365DD7FF909EF3FDF1DA13270FFEF1" box="[626,816,289,311]" gridcol="1" gridrow="1" pageId="11" pageNumber="12">4</td>
<td id="B7365DD7FF909EF3FC4BDA1320DDFEF1" box="[968,1250,289,311]" gridcol="2" gridrow="1" pageId="11" pageNumber="12">2</td>
</tr>
<tr id="F4E734ABFF909EF3FF1EDA7F20DDFEA5" box="[157,1250,333,355]" gridrow="2" pageId="11" pageNumber="12">
<th id="B7365DD7FF909EF3FF1EDA7F25FEFEA5" box="[157,449,333,355]" gridcol="0" gridrow="2" pageId="11" pageNumber="12">Bochil</th>
<td id="B7365DD7FF909EF3FDF1DA7F270FFEA5" box="[626,816,333,355]" gridcol="1" gridrow="2" pageId="11" pageNumber="12">5</td>
<td id="B7365DD7FF909EF3FC4BDA7F20DDFEA5" box="[968,1250,333,355]" gridcol="2" gridrow="2" pageId="11" pageNumber="12">1</td>
</tr>
<tr id="F4E734ABFF909EF3FF1EDA4B20DDFE49" box="[157,1250,377,399]" gridrow="3" pageId="11" pageNumber="12">
<th id="B7365DD7FF909EF3FF1EDA4B25FEFE49" box="[157,449,377,399]" gridcol="0" gridrow="3" pageId="11" pageNumber="12">Chalchihuitán</th>
<td id="B7365DD7FF909EF3FDF1DA4B270FFE49" box="[626,816,377,399]" gridcol="1" gridrow="3" pageId="11" pageNumber="12">3</td>
<td id="B7365DD7FF909EF3FC4BDA4B20DDFE49" box="[968,1250,377,399]" gridcol="2" gridrow="3" pageId="11" pageNumber="12">0</td>
</tr>
<tr id="F4E734ABFF909EF3FF1EDA9720DDFE7D" box="[157,1250,421,443]" gridrow="4" pageId="11" pageNumber="12">
<th id="B7365DD7FF909EF3FF1EDA9725FEFE7D" box="[157,449,421,443]" gridcol="0" gridrow="4" pageId="11" pageNumber="12">Coapilla</th>
<td id="B7365DD7FF909EF3FDF1DA97270FFE7D" box="[626,816,421,443]" gridcol="1" gridrow="4" pageId="11" pageNumber="12">7</td>
<td id="B7365DD7FF909EF3FC4BDA9720DDFE7D" box="[968,1250,421,443]" gridcol="2" gridrow="4" pageId="11" pageNumber="12">1</td>
</tr>
<tr id="F4E734ABFF909EF3FF1EDAE320DDFE21" box="[157,1250,465,487]" gridrow="5" pageId="11" pageNumber="12">
<th id="B7365DD7FF909EF3FF1EDAE325FEFE21" box="[157,449,465,487]" gridcol="0" gridrow="5" pageId="11" pageNumber="12">El Bosque</th>
<td id="B7365DD7FF909EF3FDF1DAE3270FFE21" box="[626,816,465,487]" gridcol="1" gridrow="5" pageId="11" pageNumber="12">2</td>
<td id="B7365DD7FF909EF3FC4BDAE320DDFE21" box="[968,1250,465,487]" gridcol="2" gridrow="5" pageId="11" pageNumber="12">0</td>
</tr>
<tr id="F4E734ABFF909EF3FF1EDACF20DDFDD5" box="[157,1250,509,531]" gridrow="6" pageId="11" pageNumber="12">
<th id="B7365DD7FF909EF3FF1EDACF25FEFDD5" box="[157,449,509,531]" gridcol="0" gridrow="6" pageId="11" pageNumber="12">El Porvenir</th>
<td id="B7365DD7FF909EF3FDF1DACF270FFDD5" box="[626,816,509,531]" gridcol="1" gridrow="6" pageId="11" pageNumber="12">1</td>
<td id="B7365DD7FF909EF3FC4BDACF20DDFDD5" box="[968,1250,509,531]" gridcol="2" gridrow="6" pageId="11" pageNumber="12">1</td>
</tr>
<tr id="F4E734ABFF909EF3FF1ED91B20DDFDF9" box="[157,1250,553,575]" gridrow="7" pageId="11" pageNumber="12">
<th id="B7365DD7FF909EF3FF1ED91B25FEFDF9" box="[157,449,553,575]" gridcol="0" gridrow="7" pageId="11" pageNumber="12">Huixtán</th>
<td id="B7365DD7FF909EF3FDF1D91B270FFDF9" box="[626,816,553,575]" gridcol="1" gridrow="7" pageId="11" pageNumber="12">5</td>
<td id="B7365DD7FF909EF3FC4BD91B20DDFDF9" box="[968,1250,553,575]" gridcol="2" gridrow="7" pageId="11" pageNumber="12">0</td>
</tr>
<tr id="F4E734ABFF909EF3FF1ED96720DDFDAD" box="[157,1250,597,619]" gridrow="8" pageId="11" pageNumber="12">
<th id="B7365DD7FF909EF3FF1ED96725FEFDAD" box="[157,449,597,619]" gridcol="0" gridrow="8" pageId="11" pageNumber="12">La Concordia</th>
<td id="B7365DD7FF909EF3FDF1D967270FFDAD" box="[626,816,597,619]" gridcol="1" gridrow="8" pageId="11" pageNumber="12">7</td>
<td id="B7365DD7FF909EF3FC4BD96720DDFDAD" box="[968,1250,597,619]" gridcol="2" gridrow="8" pageId="11" pageNumber="12">4</td>
</tr>
<tr id="F4E734ABFF909EF3FF1ED9B320DDFD51" box="[157,1250,641,663]" gridrow="9" pageId="11" pageNumber="12">
<th id="B7365DD7FF909EF3FF1ED9B325FEFD51" box="[157,449,641,663]" gridcol="0" gridrow="9" pageId="11" pageNumber="12">Motozintla</th>
<td id="B7365DD7FF909EF3FDF1D9B3270FFD51" box="[626,816,641,663]" gridcol="1" gridrow="9" pageId="11" pageNumber="12">2</td>
<td id="B7365DD7FF909EF3FC4BD9B320DDFD51" box="[968,1250,641,663]" gridcol="2" gridrow="9" pageId="11" pageNumber="12">2</td>
</tr>
<tr id="F4E734ABFF909EF3FF1ED99F20DDFD05" box="[157,1250,685,707]" gridrow="10" pageId="11" pageNumber="12">
<th id="B7365DD7FF909EF3FF1ED99F25FEFD05" box="[157,449,685,707]" gridcol="0" gridrow="10" pageId="11" pageNumber="12">Ocozocuautla</th>
<td id="B7365DD7FF909EF3FDF1D99F270FFD05" box="[626,816,685,707]" gridcol="1" gridrow="10" pageId="11" pageNumber="12">1</td>
<td id="B7365DD7FF909EF3FC4BD99F20DDFD05" box="[968,1250,685,707]" gridcol="2" gridrow="10" pageId="11" pageNumber="12">0</td>
</tr>
<tr id="F4E734ABFF909EF3FF1ED9E820DDFD36" box="[157,1250,730,752]" gridrow="11" pageId="11" pageNumber="12">
<th id="B7365DD7FF909EF3FF1ED9E825FEFD36" box="[157,449,730,752]" gridcol="0" gridrow="11" pageId="11" pageNumber="12">Pueblo Nuevo Solistahuacan</th>
<td id="B7365DD7FF909EF3FDF1D9E8270FFD36" box="[626,816,730,752]" gridcol="1" gridrow="11" pageId="11" pageNumber="12">2</td>
<td id="B7365DD7FF909EF3FC4BD9E820DDFD36" box="[968,1250,730,752]" gridcol="2" gridrow="11" pageId="11" pageNumber="12">0</td>
</tr>
<tr id="F4E734ABFF909EF3FF1ED83720DDFCDD" box="[157,1250,773,795]" gridrow="12" pageId="11" pageNumber="12">
<th id="B7365DD7FF909EF3FF1ED83725FEFCDD" box="[157,449,773,795]" gridcol="0" gridrow="12" pageId="11" pageNumber="12">Rayón</th>
<td id="B7365DD7FF909EF3FDF1D837270FFCDD" box="[626,816,773,795]" gridcol="1" gridrow="12" pageId="11" pageNumber="12">3</td>
<td id="B7365DD7FF909EF3FC4BD83720DDFCDD" box="[968,1250,773,795]" gridcol="2" gridrow="12" pageId="11" pageNumber="12">0</td>
</tr>
<tr id="F4E734ABFF909EF3FF1ED80020DDFC8E" box="[157,1250,818,840]" gridrow="13" pageId="11" pageNumber="12">
<th id="B7365DD7FF909EF3FF1ED80025FEFC8E" box="[157,449,818,840]" gridcol="0" gridrow="13" pageId="11" pageNumber="12">San Cristóbal de Las Casas</th>
<td id="B7365DD7FF909EF3FDF1D800270FFC8E" box="[626,816,818,840]" gridcol="1" gridrow="13" pageId="11" pageNumber="12">7</td>
<td id="B7365DD7FF909EF3FC4BD80020DDFC8E" box="[968,1250,818,840]" gridcol="2" gridrow="13" pageId="11" pageNumber="12">1</td>
</tr>
<tr id="F4E734ABFF909EF3FF1ED86F20DDFCB5" box="[157,1250,861,883]" gridrow="14" pageId="11" pageNumber="12">
<th id="B7365DD7FF909EF3FF1ED86F25FEFCB5" box="[157,449,861,883]" gridcol="0" gridrow="14" pageId="11" pageNumber="12">San Juan Chamula</th>
<td id="B7365DD7FF909EF3FDF1D86F270FFCB5" box="[626,816,861,883]" gridcol="1" gridrow="14" pageId="11" pageNumber="12">3</td>
<td id="B7365DD7FF909EF3FC4BD86F20DDFCB5" box="[968,1250,861,883]" gridcol="2" gridrow="14" pageId="11" pageNumber="12">0</td>
</tr>
<tr id="F4E734ABFF909EF3FF1ED8B820DDFC66" box="[157,1250,906,928]" gridrow="15" pageId="11" pageNumber="12">
<th id="B7365DD7FF909EF3FF1ED8B825FEFC66" box="[157,449,906,928]" gridcol="0" gridrow="15" pageId="11" pageNumber="12">Tapalapa</th>
<td id="B7365DD7FF909EF3FDF1D8B8270FFC66" box="[626,816,906,928]" gridcol="1" gridrow="15" pageId="11" pageNumber="12">3</td>
<td id="B7365DD7FF909EF3FC4BD8B820DDFC66" box="[968,1250,906,928]" gridcol="2" gridrow="15" pageId="11" pageNumber="12">0</td>
</tr>
<tr id="F4E734ABFF909EF3FF1ED88720DDFC0D" box="[157,1250,949,971]" gridrow="16" pageId="11" pageNumber="12">
<th id="B7365DD7FF909EF3FF1ED88725FEFC0D" box="[157,449,949,971]" gridcol="0" gridrow="16" pageId="11" pageNumber="12">Tenejapa</th>
<td id="B7365DD7FF909EF3FDF1D887270FFC0D" box="[626,816,949,971]" gridcol="1" gridrow="16" pageId="11" pageNumber="12">3</td>
<td id="B7365DD7FF909EF3FC4BD88720DDFC0D" box="[968,1250,949,971]" gridcol="2" gridrow="16" pageId="11" pageNumber="12">0</td>
</tr>
<tr id="F4E734ABFF909EF3FF1ED8D020DDFC3E" box="[157,1250,994,1016]" gridrow="17" pageId="11" pageNumber="12">
<th id="B7365DD7FF909EF3FF1ED8D025FEFC3E" box="[157,449,994,1016]" gridcol="0" gridrow="17" pageId="11" pageNumber="12">Trinitaria</th>
<td id="B7365DD7FF909EF3FDF1D8D0270FFC3E" box="[626,816,994,1016]" gridcol="1" gridrow="17" pageId="11" pageNumber="12">1</td>
<td id="B7365DD7FF909EF3FC4BD8D020DDFC3E" box="[968,1250,994,1016]" gridcol="2" gridrow="17" pageId="11" pageNumber="12">1</td>
</tr>
<tr id="F4E734ABFF909EF3FF1EDF3C20DDFBE2" box="[157,1250,1038,1060]" gridrow="18" pageId="11" pageNumber="12">
<th id="B7365DD7FF909EF3FF1EDF3C25FEFBE2" box="[157,449,1038,1060]" gridcol="0" gridrow="18" pageId="11" pageNumber="12">Tuxtla/SCLC</th>
<td id="B7365DD7FF909EF3FDF1DF3C270FFBE2" box="[626,816,1038,1060]" gridcol="1" gridrow="18" pageId="11" pageNumber="12">1</td>
<td id="B7365DD7FF909EF3FC4BDF3C20DDFBE2" box="[968,1250,1038,1060]" gridcol="2" gridrow="18" pageId="11" pageNumber="12">1</td>
</tr>
<tr id="F4E734ABFF909EF3FF1EDF0820DDFB96" box="[157,1250,1082,1104]" gridrow="19" pageId="11" pageNumber="12">
<th id="B7365DD7FF909EF3FF1EDF0825FEFB96" box="[157,449,1082,1104]" gridcol="0" gridrow="19" pageId="11" pageNumber="12">Unión Juárez</th>
<td id="B7365DD7FF909EF3FDF1DF08270FFB96" box="[626,816,1082,1104]" gridcol="1" gridrow="19" pageId="11" pageNumber="12">6</td>
<td id="B7365DD7FF909EF3FC4BDF0820DDFB96" box="[968,1250,1082,1104]" gridcol="2" gridrow="19" pageId="11" pageNumber="12">4</td>
</tr>
<tr id="F4E734ABFF909EF3FF1EDF5420DDFBBA" box="[157,1250,1126,1148]" gridrow="20" pageId="11" pageNumber="12">
<th id="B7365DD7FF909EF3FF1EDF5425FEFBBA" box="[157,449,1126,1148]" gridcol="0" gridrow="20" pageId="11" pageNumber="12">Villacorzo</th>
<td id="B7365DD7FF909EF3FDF1DF54270FFBBA" box="[626,816,1126,1148]" gridcol="1" gridrow="20" pageId="11" pageNumber="12">5</td>
<td id="B7365DD7FF909EF3FC4BDF5420DDFBBA" box="[968,1250,1126,1148]" gridcol="2" gridrow="20" pageId="11" pageNumber="12">2</td>
</tr>
</table>
</paragraph>
<paragraph id="4A6836E9FF906107FF14DFF225C3FB1C" blockId="11.[151,508,1216,1242]" box="[151,508,1216,1242]" pageId="11" pageNumber="12">
<emphasis id="78A3EAFBFF906107FF14DFF225C3FB1C" bold="true" box="[151,508,1216,1242]" pageId="11" pageNumber="12">Checklist of included species</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>