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<document id="96721780E0EA73FA1B4634FE1994616D" ID-DOI="10.11646/zootaxa.5028.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5453967" ID-ZooBank="C678F0FE-1B62-4F34-8A66-449CF9806B50" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630655426570" checkinUser="plazi" docAuthor="Santos, Bruna S., Marques, Mariana P., Bauer, Aaron M. &amp; Ceríaco, Luis M. P." docDate="2021" docId="B007F528FFC9FFC416D7FC56FE435F2A" docLanguage="en" docName="zootaxa.5028.1.1.pdf" docOrigin="Zootaxa 5028 (1)" docSource="http://zoobank.org/c678f0fe-1b62-4f34-8a66-449cf9806b50" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Tomopterna ahli" docType="treatment" docVersion="3" lastPageNumber="18" masterDocId="4C3E8D50FFD8FFD51640FFA1FFD2597B" masterDocTitle="Herpetological results of Francisco Newton’s Zoological Expedition to Angola (1903 – 1906): a taxonomic revision and new records of a forgotten collection" masterLastPageNumber="80" masterPageNumber="1" pageNumber="18" updateTime="1699281765633" updateUser="plazi">
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<mods:title id="0E0061043FEA067AAB6994350FB3331D">Herpetological results of Francisco Newton’s Zoological Expedition to Angola (1903 – 1906): a taxonomic revision and new records of a forgotten collection</mods:title>
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<mods:namePart id="8A60DAF3AEEFDB79255E54CB17AA5FA5">Santos, Bruna S.</mods:namePart>
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<mods:namePart id="468C8AA4422485A44472D7CA352A9F0E">Marques, Mariana P.</mods:namePart>
<mods:nameIdentifier id="50E86347E63692638E7F47EB2E01C8B6" type="ORCID">0000-0002-1712-2632</mods:nameIdentifier>
<mods:nameIdentifier id="FB4F7C78E809EB4A2FDC8E8AB36677F4" type="email">mptlmarques@gmail.com</mods:nameIdentifier>
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<mods:namePart id="89042C14E979F60660911CE862DB5780">Bauer, Aaron M.</mods:namePart>
<mods:nameIdentifier id="C44C1FCCBF037CA6BE6A4F37EABBF895" type="ORCID">0000-0001-6839-8025</mods:nameIdentifier>
<mods:nameIdentifier id="67D6CFB4E6B692C152C283298859127F" type="email">aaron.bauer@villanova.edu</mods:nameIdentifier>
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<mods:roleTerm id="73340EAE1C22EC5CF4967235CAC6A2E6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0FD0C3A8A10D2E1A69DD2B4AD346C0D2">Ceríaco, Luis M. P.</mods:namePart>
<mods:nameIdentifier id="84FCA87921567BFE2D25DEAA82E9260C" type="ORCID">0000-0002-3798-2664</mods:nameIdentifier>
<mods:nameIdentifier id="E7C1852AC9D57CCC76E6E8C56DBC200D" type="email">brunascesantos@gmail.com</mods:nameIdentifier>
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<mods:title id="1DF78DBD1B271706DA2E409A670A3718">Zootaxa</mods:title>
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<mods:date id="734A16BAB9CD846837D374B8DD1E258E">2021</mods:date>
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<mods:number id="5CC927A2B7D052A31D0EBDAEED664889">2021-09-03</mods:number>
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<mods:number id="5D343CABE399AA114E9750741A2CB34C">5028</mods:number>
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<mods:identifier id="FCDBDF05929431AABEC1A69A0323B0F8" type="DOI">10.11646/zootaxa.5028.1.1</mods:identifier>
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<treatment id="B007F528FFC9FFC416D7FC56FE435F2A" LSID="urn:lsid:plazi:treatment:B007F528FFC9FFC416D7FC56FE435F2A" httpUri="http://treatment.plazi.org/id/B007F528FFC9FFC416D7FC56FE435F2A" lastPageNumber="18" pageId="17" pageNumber="18">
<subSubSection id="70B417B5FFC9FFC416D7FC56FDF55D6A" box="[151,551,1015,1042]" pageId="17" pageNumber="18" type="nomenclature">
<paragraph id="3811443EFFC9FFC416D7FC56FDF55D6A" blockId="17.[151,551,1015,1042]" box="[151,551,1015,1042]" pageId="17" pageNumber="18">
<heading id="6359F352FFC9FFC416D7FC56FDF55D6A" bold="true" box="[151,551,1015,1042]" fontSize="11" level="2" pageId="17" pageNumber="18" reason="4">
<taxonomicName id="FFAE3FBDFFC9FFC416D7FC56FDF55D6A" ID-CoL="GYPB" authority="(Deckert, 1938)" baseAuthorityName="Deckert" baseAuthorityYear="1938" box="[151,551,1015,1042]" class="Amphibia" family="Pyxicephalidae" genus="Tomopterna" kingdom="Animalia" order="Anura" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="ahli">
<emphasis id="0ADA982CFFC9FFC416D7FC56FDF55D6A" bold="true" box="[151,551,1015,1042]" pageId="17" pageNumber="18">
<emphasis id="0ADA982CFFC9FFC416D7FC56FE8D5D69" bold="true" box="[151,351,1015,1042]" italics="true" pageId="17" pageNumber="18">Tomopterna ahli</emphasis>
(Deckert, 1938)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="70B417B5FFC9FFC416D7FB9EFA9F5D05" pageId="17" pageNumber="18" type="materials_examined">
<paragraph id="3811443EFFC9FFC416D7FB9EFA9F5D05" blockId="17.[151,1437,1087,1437]" pageId="17" pageNumber="18">
<emphasis id="0ADA982CFFC9FFC416D7FB9EFE115D22" bold="true" box="[151,451,1087,1113]" pageId="17" pageNumber="18">
Material (
<specimenCount id="2EA88FB7FFC9FFC4174FFB9EFE645D22" box="[271,438,1087,1113]" pageId="17" pageNumber="18" type="generic">one specimen</specimenCount>
):
</emphasis>
<materialsCitation id="88C64E63FFC9FFC4178BFB9EFA9A5D05" collectionCode="ANF" elevation="24" location="Namibe Province" pageId="17" pageNumber="18" specimenCode="ANF 225" specimenCount="1" stateProvince="Namibe">
One adult specimen (MHNCUP/
<specimenCode id="6808EC45FFC9FFC4157CFB9EFC7B5D21" box="[828,937,1087,1114]" collectionCode="ANF" pageId="17" pageNumber="18">ANF 225</specimenCode>
, formerly UP-MHNFCP-017341;
<figureCitation id="A09558BBFFC9FFC4136CFB9EFA5D5D21" box="[1324,1423,1087,1114]" captionStart="FIGURE S" captionStartId="54.[151,249,2015,2039]" captionTargetBox="[249,1337,1105,1989]" captionTargetId="figure-51@54.[249,1337,1105,1989]" captionTargetPageId="54" captionText="FIGURE S26. Specimen of Tomopterna ahli (MHNC-UP/ANF225), from Moçâmedes. Photo by Bruna Santos." figureDoi="http://doi.org/10.5281/zenodo.5454035" httpUri="https://zenodo.org/record/5454035/files/figure.png" pageId="17" pageNumber="18">Fig. S26</figureCitation>
), collected from Mossamedes [=Moçâmedes] [-15.20000º, 12.15000º,
<elevation id="B383A30DFFC9FFC415DBFBC2FBC25D06" box="[923,1040,1122,1150]" metricMagnitude="1" metricUnit="m" metricValue="2.4" pageId="17" pageNumber="18" unit="m" value="24.0">
<quantity id="FF56E9DBFFC9FFC415DBFBC2FC065D05" box="[923,980,1123,1150]" metricMagnitude="1" metricUnit="m" metricValue="2.4" pageId="17" pageNumber="18" unit="m" value="24.0">24 m</quantity>
a.s.l.
</elevation>
]
<collectingRegion id="FA6A8ADCFFC9FFC4125FFBC2FB375D06" box="[1055,1253,1123,1149]" country="Angola" name="Namibe" pageId="17" pageNumber="18">Namibe Province</collectingRegion>
, in 1905
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="70B417B5FFC9FFC41687FB26FE435F2A" pageId="17" pageNumber="18" type="discussion">
<paragraph id="3811443EFFC9FFC41687FB26FB695CE6" blockId="17.[151,1437,1087,1437]" pageId="17" pageNumber="18">
<emphasis id="0ADA982CFFC9FFC41687FB26FE805DDA" bold="true" box="[199,338,1159,1185]" pageId="17" pageNumber="18">Comments:</emphasis>
<taxonomicName id="FFAE3FBDFFC9FFC4171BFB29FDC55DDA" baseAuthorityName="Deckert" baseAuthorityYear="1938" box="[347,535,1159,1185]" class="Amphibia" family="Pyxicephalidae" genus="Tomopterna" kingdom="Animalia" order="Anura" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="ahli">
<emphasis id="0ADA982CFFC9FFC4171BFB29FDC55DDA" box="[347,535,1159,1185]" italics="true" pageId="17" pageNumber="18">Tomopterna ahli</emphasis>
</taxonomicName>
is a species found in southwestern
<collectingCountry id="40B904AEFFC9FFC415F1FB26FBD45DDA" box="[945,1030,1159,1185]" name="Angola" pageId="17" pageNumber="18">Angola</collectingCountry>
, extending to northern and western
<collectingCountry id="40B904AEFFC9FFC416D7FB0AFF255DBE" box="[151,247,1195,1221]" name="Namibia" pageId="17" pageNumber="18">Namibia</collectingCountry>
(
<bibRefCitation id="5C3F39CFFFC9FFC41747FB0AFE3D5DBE" author="Marques, M. P. &amp; Ceriaco, L. M. P. &amp; Blackburn, D. C. &amp; Bauer, A. M." box="[263,495,1195,1221]" pageId="17" pageNumber="18" pagination="1 - 501" refId="ref38562" refString="Marques, M. P., Ceriaco, L. M. P., Blackburn, D. C. &amp; Bauer, A. M. (2018) Diversity and distribution of the amphibians and terrestrial reptiles of Angola atlas of historical and bibliographic records (1840 - 2017). Proceedings of the California Academy of Sciences, Series 4, 65 (Supplement 2), 1 - 501." type="journal article" year="2018">
Marques
<emphasis id="0ADA982CFFC9FFC41733FB0DFE7E5DBE" box="[371,428,1195,1221]" italics="true" pageId="17" pageNumber="18">et al.</emphasis>
2018
</bibRefCitation>
;
<bibRefCitation id="5C3F39CFFFC9FFC417BCFB0DFDA85DBE" author="Frost, D. R." box="[508,634,1195,1222]" pageId="17" pageNumber="18" refId="ref37539" refString="Frost, D. R. (2020) Amphibian Species of the World: an online reference. Version 6. 0. American Museum of Natural History, New York. Available from: http: // research. amnh. org / herpetology / amphibia / index. html (accessed 19 September 2020)" type="url" year="2020">Frost 2020</bibRefCitation>
). This specimen is part of the shipment of specimens made by Newton from
<collectingRegion id="FA6A8ADCFFC9FFC41695FB6EFE485D92" box="[213,410,1231,1257]" country="Angola" name="Namibe" pageId="17" pageNumber="18">Namibe Province</collectingRegion>
and therefore was never published by Ferreira or any subsequent authors.
<taxonomicName id="FFAE3FBDFFC9FFC412A6FB71FAFC5D92" baseAuthorityName="Deckert" baseAuthorityYear="1938" box="[1254,1326,1231,1257]" class="Amphibia" family="Pyxicephalidae" genus="Tomopterna" kingdom="Animalia" order="Anura" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="ahli">
<emphasis id="0ADA982CFFC9FFC412A6FB71FAFC5D92" box="[1254,1326,1231,1257]" italics="true" pageId="17" pageNumber="18">T. ahli</emphasis>
</taxonomicName>
was until very recently known as
<taxonomicName id="FFAE3FBDFFC9FFC417E1FB55FC3F5C76" authority="Dawood &amp; Channing, 2002" authorityName="Dawood &amp; Channing" authorityYear="2002" box="[417,1005,1267,1293]" class="Amphibia" family="Pyxicephalidae" genus="Tomopterna" kingdom="Animalia" order="Anura" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="damarensis">
<emphasis id="0ADA982CFFC9FFC417E1FB55FD7C5C76" box="[417,686,1267,1293]" italics="true" pageId="17" pageNumber="18">Tomopterna damarensis</emphasis>
Dawood &amp; Channing, 2002
</taxonomicName>
. The previous name is now considered a junior synonym (
<bibRefCitation id="5C3F39CFFFC9FFC41733FAB6FD705C4A" author="Channing, A. &amp; Becker, F." box="[371,674,1303,1329]" pageId="17" pageNumber="18" pagination="549 - 560" refId="ref36220" refString="Channing, A. &amp; Becker, F. (2019) Correction to the type locality of Tomopterna ahli (Deckert, 1938) (Anura: Pyxicephalidae), with the designation of a neotype. Zootaxa, 4688 (4), 549 - 560. https: // doi. org / 10.11646 / zootaxa. 4688.4.6" type="journal article" year="2019">Channing &amp; Becker 2019</bibRefCitation>
). The first account for the species in
<collectingCountry id="40B904AEFFC9FFC41214FAB6FB7B5C4A" box="[1108,1193,1303,1329]" name="Angola" pageId="17" pageNumber="18">Angola</collectingCountry>
, based on specimens collected in Iona National Park, southern
<collectingRegion id="FA6A8ADCFFC9FFC4142CFA9AFCE25C2E" box="[620,816,1339,1365]" country="Angola" name="Namibe" pageId="17" pageNumber="18">Namibe Province</collectingRegion>
was published in 2016 (
<bibRefCitation id="5C3F39CFFFC9FFC41203FA9AFAF95C2E" author="Ceriaco, L. M. P. &amp; De Sa, S. A. C. &amp; Bandeira, S. &amp; Valerio, H. &amp; Stanley, E. L. &amp; Kuhn, A. L. &amp; Marques, M. P. &amp; Vindum, J. V. &amp; Blackburn, D. C. &amp; Bauer, A. M." box="[1091,1323,1339,1365]" pageId="17" pageNumber="18" pagination="15 - 61" refId="ref35490" refString="Ceriaco, L. M. P., De Sa, S. A. C., Bandeira, S., Valerio, H., Stanley, E. L., Kuhn, A. L., Marques, M. P., Vindum, J. V., Blackburn, D. C. &amp; Bauer, A. M. (2016 a) Herpetological survey of the Iona National Park and Namibe National Park, Angola, with a synoptic list of the amphibians and reptiles of Namibe Province, southwestern Angola. Proceedings of the California Academy of Sciences, Series 4, 63 (2), 15 - 61." type="journal article" year="2016">
Ceríaco
<emphasis id="0ADA982CFFC9FFC412E3FA9DFB0E5C2E" box="[1187,1244,1339,1365]" italics="true" pageId="17" pageNumber="18">et al.</emphasis>
2016a
</bibRefCitation>
;
<bibRefCitation id="5C3F39CFFFC9FFC41377FA9AFEC05C02" author="Heinicke, M. P. &amp; Ceriaco, L. M. P. &amp; Moore, I. M. &amp; Bauer, A. M. &amp; Blackburn, D. C." pageId="17" pageNumber="18" pagination="461 - 465" refId="ref37959" refString="Heinicke, M. P., Ceriaco, L. M. P., Moore, I. M., Bauer, A. M. &amp; Blackburn, D. C. (2017) Tomopterna damarensis (Anura: Pyxicephalidae) is broadly distributed in Namibia and Angola. Salamandra, 53 (3), 461 - 465." type="journal article" year="2017">
Heinicke
<emphasis id="0ADA982CFFC9FFC416D7FAC1FF1D5C02" box="[151,207,1375,1401]" italics="true" pageId="17" pageNumber="18">et al.</emphasis>
2017
</bibRefCitation>
).
<bibRefCitation id="5C3F39CFFFC9FFC41764FAFEFE2B5C02" author="Butler, B. O. &amp; Ceriaco, L. M. P. &amp; Marques, M. P. &amp; Bandeira, S. &amp; Julio, T. &amp; Heinicke, M. P. &amp; Bauer, A. M." box="[292,505,1375,1401]" pageId="17" pageNumber="18" pagination="225 - 240" refId="ref35071" refString="Butler, B. O., Ceriaco, L. M. P., Marques, M. P., Bandeira, S., Julio, T., Heinicke, M. P. &amp; Bauer, A. M. (2019) Herpetological survey of Huila Province, southwest Angola, including first records from Bicuar National Park. Herpetological Review, 50 (2), 225 - 240." type="journal article" year="2019">
Butler
<emphasis id="0ADA982CFFC9FFC41731FAC1FE7A5C02" box="[369,424,1375,1401]" italics="true" pageId="17" pageNumber="18">et al.</emphasis>
(2019)
</bibRefCitation>
provided additional record from Bicuar National Park,
<collectingRegion id="FA6A8ADCFFC9FFC41224FAFEFAC25C02" box="[1124,1296,1375,1401]" country="Angola" name="Huila" pageId="17" pageNumber="18">Huíla Province</collectingRegion>
. The present specimen extends the known distribution of the species northwest towards Moçâmedes region.
</paragraph>
<paragraph id="3811443EFFC9FFC416D7FA4EFF285F72" blockId="17.[151,250,1519,1545]" box="[151,250,1519,1545]" pageId="17" pageNumber="18">
<taxonomicName id="FFAE3FBDFFC9FFC416D7FA4EFF285F72" box="[151,250,1519,1545]" class="Reptilia" kingdom="Animalia" pageId="17" pageNumber="18" phylum="Chordata" rank="class">
<emphasis id="0ADA982CFFC9FFC416D7FA4EFF285F72" bold="true" box="[151,250,1519,1545]" pageId="17" pageNumber="18">Reptilia</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="3811443EFFC9FFC416D7F996FE435F2A" blockId="17.[151,401,1591,1617]" box="[151,401,1591,1617]" pageId="17" pageNumber="18">
<taxonomicName id="FFAE3FBDFFC9FFC416D7F996FE435F2A" authority="(Lizards)" baseAuthorityName="Lizards" box="[151,401,1591,1617]" class="Reptilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="18" phylum="Chordata" rank="order">
<emphasis id="0ADA982CFFC9FFC416D7F996FE435F2A" bold="true" box="[151,401,1591,1617]" pageId="17" pageNumber="18">Squamata (Lizards)</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>
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