828 lines
108 KiB
XML
828 lines
108 KiB
XML
<document id="E3EBBC2EFCDB7DD5163C1C81013A72BC" ID-CLB-Dataset="5750" ID-DOI="10.11646/zootaxa.4996.2.3" ID-GBIF-Dataset="f2e2b777-cc43-44fa-be59-3a443e93c204" ID-ISSN="1175-5326" ID-Zenodo-Dep="5069879" ID-ZooBank="C3724D94-327B-4701-9D97-30D98D8B4F18" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1625477987038" checkinUser="plazi" docAuthor="Kovačić, Marcelo, Bogorodsky, Sergey V., Zajonz, Uwe & Tornabene, Luke" docDate="2021" docId="8F7E87DCF631FFB6D5D865452293E7FD" docLanguage="en" docName="zootaxa.4996.2.3.pdf" docOrigin="Zootaxa 4996 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hetereleotris nasoramosa Kovačić & Bogorodsky & Zajonz & Tornabene 2021, sp. nov." docType="treatment" docVersion="5" lastPageNumber="294" masterDocId="7347FFA4F632FFBCD54F607E2272E434" masterDocTitle="A new species of Hetereleotris (Teleostei: Gobiidae) from the Socotra Archipelago (north-western Indian Ocean), a rare case of a hole-associated adaptation in gobiid fishes" masterLastPageNumber="300" masterPageNumber="284" pageNumber="287" updateTime="1699024520383" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="566A08323B8BB16C3A33FBC57EFF0FEB">
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<mods:title id="2D738156A134FA42C666469DB952E43A">A new species of Hetereleotris (Teleostei: Gobiidae) from the Socotra Archipelago (north-western Indian Ocean), a rare case of a hole-associated adaptation in gobiid fishes</mods:title>
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<mods:namePart id="FC61180D8169F2CDFAEC5652FC622C48">Kovačić, Marcelo</mods:namePart>
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<mods:nameIdentifier id="4EBF1E9D98E6A40E7BF8CAC464B0412E" type="ORCID">0000-0002-4049-9366</mods:nameIdentifier>
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<mods:affiliation id="CD6E1FB9BAE9302A547181758B36F3B3">Natural History Museum Rijeka, Lorenzov prolaz 1, HR- 51000 Rijeka, Croatia.</mods:affiliation>
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<mods:nameIdentifier id="ED854D26EA66AC9EB2E1AC2CAB7C721D" type="email">marcelo@prirodoslovni.com</mods:nameIdentifier>
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<mods:roleTerm id="F5D34B599040F57BF3EFC6752125CAD8">Author</mods:roleTerm>
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<mods:namePart id="EB1C43971D48310C9841E81C5ACEAC6F">Bogorodsky, Sergey V.</mods:namePart>
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<mods:affiliation id="598D2D6449658E8BB38D6AD680021BFD">Senckenberg Research Institute and Museum of Nature (SMF), Section Ichthyology & Senckenberg Biodiversity and Climate Research Centre (SBiK-F), Biogeography - Senckenberganlage 25, 60325 Frankfurt am Main, Germany. & Station of Naturalists, Omsk, Russia. ic 187196 @ yandex. ru</mods:affiliation>
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<mods:namePart id="390453D581161E8F12031431716A0B1A">Zajonz, Uwe</mods:namePart>
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<mods:affiliation id="3C8AC845429C88E305D107AA653A620F">Senckenberg Research Institute and Museum of Nature (SMF), Section Ichthyology & Senckenberg Biodiversity and Climate Research Centre (SBiK-F), Biogeography - Senckenberganlage 25, 60325 Frankfurt am Main, Germany. & Institute for Ecology, Diversity and Evolution, Faculty of Biological Sciences, Goethe University Frankfurt am Main, Max-von-Laue- Strasse 13, 60438 Frankfurt am Main, Germany;</mods:affiliation>
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<mods:namePart id="1C26672343D30C00DFE7579F42038C1D">Tornabene, Luke</mods:namePart>
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<mods:affiliation id="028ADD4A8BA824B59E23C284C5D95BE0">School of Aquatic and Fishery Sciences and the Burke Museum of Natural History and Culture, University of Washington, 1122 NE Boat Street, Seattle, WA 98105, USA.</mods:affiliation>
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<mods:title id="412EDDC9484414A689EA2726E1B2957C">Zootaxa</mods:title>
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<mods:date id="F58AC4A7AB7A327BFB696592A4EB03F2">2021</mods:date>
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<mods:number id="E394C8A9C729E13FD4FC6C74235C0680">2021-07-05</mods:number>
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<treatment id="8F7E87DCF631FFB6D5D865452293E7FD" ID-DOI="http://doi.org/10.5281/zenodo.5069881" ID-GBIF-Taxon="183318211" ID-Zenodo-Dep="5069881" LSID="urn:lsid:plazi:treatment:8F7E87DCF631FFB6D5D865452293E7FD" httpUri="http://treatment.plazi.org/id/8F7E87DCF631FFB6D5D865452293E7FD" lastPageId="10" lastPageNumber="294" pageId="3" pageNumber="287">
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<subSubSection id="4FCD6541F631FFBFD5D8654523C6E14D" pageId="3" pageNumber="287" type="nomenclature">
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<paragraph id="076836CAF631FFBFD5D865452041E161" blockId="3.[151,563,1339,1438]" box="[151,563,1339,1366]" pageId="3" pageNumber="287">
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<heading id="5C2081A6F631FFBFD5D865452041E161" bold="true" box="[151,563,1339,1366]" fontSize="11" level="1" pageId="3" pageNumber="287" reason="1">
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<emphasis id="35A3EAD8F631FFBFD5D865452041E161" bold="true" box="[151,563,1339,1366]" pageId="3" pageNumber="287">
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<taxonomicName id="C0D74D49F631FFBFD5D8654523BCE161" authority="Kovačić & Bogorodsky & Zajonz & Tornabene, 2021" authorityName="Kovačić & Bogorodsky & Zajonz & Tornabene" authorityYear="2021" box="[151,462,1339,1366]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="287" phylum="Chordata" rank="species" species="nasoramosa" status="sp. nov.">
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<emphasis id="35A3EAD8F631FFBFD5D8654523BCE161" bold="true" box="[151,462,1339,1366]" italics="true" pageId="3" pageNumber="287">Hetereleotris nasoramosa</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="2E9057A3F631FFBFD49A65452041E161" box="[469,563,1339,1365]" pageId="3" pageNumber="287" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="076836CAF631FFBFD5D8652123C6E14D" blockId="3.[151,563,1339,1438]" box="[151,436,1375,1401]" pageId="3" pageNumber="287">
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<collectingCountry id="7FC0765AF631FFBFD5D86521229FE14D" box="[151,237,1375,1401]" name="Yemen" pageId="3" pageNumber="287">Socotra</collectingCountry>
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hole-hiding goby
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</paragraph>
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</subSubSection>
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<subSubSection id="4FCD6541F631FFBFD5D865FD2312E1A9" box="[151,352,1411,1438]" pageId="3" pageNumber="287" type="description">
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<paragraph id="076836CAF631FFBFD5D865FD2312E1A9" blockId="3.[151,563,1339,1438]" box="[151,352,1411,1438]" pageId="3" pageNumber="287">
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<figureCitation id="9FEC2A4FF631FFBFD5D865FD2370E1AA" box="[151,258,1411,1438]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStartId-0="5.[151,250,1151,1175]" captionStartId-1="6.[151,250,1084,1108]" captionStartId-2="7.[151,250,1862,1886]" captionStartId-3="8.[151,250,1943,1967]" captionTargetBox-0="[151,1436,181,1127]" captionTargetBox-1="[159,1415,366,1049]" captionTargetBox-2="[151,1437,885,1839]" captionTargetBox-3="[192,1395,181,1920]" captionTargetId-0="figure-26@5.[151,1436,181,1127]" captionTargetId-1="figure-107@6.[159,1427,345,1061]" captionTargetId-2="figure-417@7.[151,1437,885,1839]" captionTargetId-3="figure-20@8.[192,1395,181,1920]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionTargetPageId-3="8" captionText-0="FIGURE 2. Hetereleotris nasoramosa sp. nov., holotype, SMF 39500 [sample tissue SOC18-191], female, 20.9 mm SL, Di Hamri, Socotra Island: A: head with AN - anterior nostril as a short tube with tentacle from the rim, PN - posterior nostril a long tube with very long anterior tentacle and shorter posterior tentacle extending from the rim, PAJ - posterior angle of jaws extends behind vertical through posterior edge of eye; B: pelvic fins completely separated and without frenum; C: caudal peduncle stained, with visible scales; D: nape stained, erected pores of anterior oculoscapular canal, terminology in text. Photos by S.V. Bogorodsky (A) and M. Kovačić (B-D)." captionText-1="FIGURE 3. Hetereleotris nasoramosa sp. nov. Cephalic sensory papillae, paratype, PMR VP4890, male, 22.3 mm SL, Di Hamri, Socotra Island. The right side illustrated, as it is in better condition, terminology in text. Drawing by M. Kovačić." captionText-2="FIGURE 4. Hetereleotris nasoramosa sp. nov., holotype, SMF 39500, female, 20.9 mm SL, Di Hamri, Socotra Island: A: freshly collected specimen; B: preserved specimen. Photos by S.V. Bogorodsky (A) and M. Kovačić (B)." captionText-3="FIGURE 5. Hetereleotris nasoramosa sp. nov.; A: SMF 39501, paratype, female, 24.9 mm SL, alive specimen, Di Hamri, Socotra Island; B: the same specimen, freshly collected; C: paratype, SMF 39503, male, 21.1 mm SL, alive specimen, Di Timri, Socotra Island; D: presumably male, alive specimen, not collected, Di Timri, Socotra Island. Photos by S.V. Bogorodsky." figureDoi-0="http://doi.org/10.5281/zenodo.5069885" figureDoi-1="http://doi.org/10.5281/zenodo.5069887" figureDoi-2="http://doi.org/10.5281/zenodo.5069889" figureDoi-3="http://doi.org/10.5281/zenodo.5069891" httpUri-0="https://zenodo.org/record/5069885/files/figure.png" httpUri-1="https://zenodo.org/record/5069887/files/figure.png" httpUri-2="https://zenodo.org/record/5069889/files/figure.png" httpUri-3="https://zenodo.org/record/5069891/files/figure.png" pageId="3" pageNumber="287">Figs. 2–5</figureCitation>
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,
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<tableCitation id="4A550371F631FFBFD44265FD2312E1A9" box="[269,352,1411,1437]" captionStart="TABLE 1" captionStartId="2.[151,239,151,175]" captionText="TABLE 1. Morphometric characters (% SL) of Hetereleotris nasoramosa sp. nov. Characters are sorted in alphabetic order. Non-overlapping ranges between male and females marked with *." pageId="3" pageNumber="287">Table 1</tableCitation>
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</paragraph>
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</subSubSection>
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<paragraph id="076836CAF631FFBFD5D865B5228DE21A" blockId="3.[151,1437,1482,2014]" pageId="3" pageNumber="287">
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<materialsCitation id="B7BF3C97F631FFBFD5D865B5228DE21A" ID-GBIF-Occurrence="3313884301" accessionNumber="SMF 39500" collectingDate="2018-05-05" collectionCode="SMF, V" collectorName="Bogorodsky & F. N. Saeed" country="Socotra Island" elevation="12" latitude="12.671866" location="Delisha" longLatPrecision="1" longitude="54.153534" pageId="3" pageNumber="287" specimenCount="1" typeStatus="holotype">
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<typeStatus id="D86C8868F631FFBFD5D865B52376E1D1" box="[151,260,1483,1509]" pageId="3" pageNumber="287" type="holotype">Holotype</typeStatus>
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</emphasis>
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.
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<collectionCode id="61C6AE0FF631FFBFD45C65B52339E1D1" box="[275,331,1483,1509]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="3" pageNumber="287">SMF</collectionCode>
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39500 [sample tissue SOC18-191], female, 20.9+
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<quantity id="C02F9B2FF631FFBFD6CC65B521AEE1D2" box="[899,988,1483,1510]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="3" pageNumber="287" unit="mm" value="4.8">4.8 mm</quantity>
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,
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<collectingCountry id="7FC0765AF631FFBFD6A765B526FFE1D1" box="[1000,1165,1483,1509]" name="Socotra Island" pageId="3" pageNumber="287">Socotra Island</collectingCountry>
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,
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<location id="02086011F631FFBFD1D665B52683E1D1" LSID="urn:lsid:plazi:treatment:8F7E87DCF631FFB6D5D865452293E7FD:02086011F631FFBFD1D665B52683E1D1" box="[1177,1265,1483,1509]" country="Socotra Island" latitude="12.671866" longLatPrecision="1" longitude="54.153534" name="Delisha" pageId="3" pageNumber="287">Delisha</location>
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,
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,
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, a small stone on sand flat, depth
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</quantity>
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, coll. S.
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<collectionCode id="61C6AE0FF631FFBFD622659121F6E23D" box="[877,900,1519,1545]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="287" type="Museum">V</collectionCode>
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.
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<collectorName id="AA22531CF631FFBFD6DF6591266BE23D" box="[912,1049,1519,1545]" pageId="3" pageNumber="287">Bogorodsky</collectorName>
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&
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<collectorName id="AA22531CF631FFBFD171658E26CEE23D" box="[1086,1212,1519,1546]" pageId="3" pageNumber="287">F.N. Saeed</collectorName>
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,
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<date id="7369100AF631FFBFD1876591272DE23D" box="[1224,1375,1519,1546]" pageId="3" pageNumber="287" value="2018-05-05">
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<collectingDate id="632DE9E2F631FFBFD1876591272DE23D" box="[1224,1375,1519,1546]" pageId="3" pageNumber="287" value="2018-05-05">05 May 2018</collectingDate>
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</date>
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(
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<figureCitation id="9FEC2A4FF631FFBFD02065912283E21A" captionStart="FIGURE 4" captionStartId="7.[151,250,1862,1886]" captionTargetBox="[151,1437,885,1839]" captionTargetId="figure-417@7.[151,1437,885,1839]" captionTargetPageId="7" captionText="FIGURE 4. Hetereleotris nasoramosa sp. nov., holotype, SMF 39500, female, 20.9 mm SL, Di Hamri, Socotra Island: A: freshly collected specimen; B: preserved specimen. Photos by S.V. Bogorodsky (A) and M. Kovačić (B)." figureDoi="http://doi.org/10.5281/zenodo.5069889" httpUri="https://zenodo.org/record/5069889/files/figure.png" pageId="3" pageNumber="287">Fig. 4A & B</figureCitation>
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).
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</materialsCitation>
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</paragraph>
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<paragraph id="076836CAF631FFBFD5886649239BE346" blockId="3.[151,1437,1482,2014]" pageId="3" pageNumber="287">
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<emphasis id="35A3EAD8F631FFBFD5886649234DE265" bold="true" box="[199,319,1591,1617]" pageId="3" pageNumber="287">
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<typeStatus id="D86C8868F631FFBFD5886649234DE265" box="[199,319,1591,1617]" pageId="3" pageNumber="287" type="paratype">Paratypes</typeStatus>
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</emphasis>
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(
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<specimenCount id="11D1FD43F631FFBFD41F664923ACE265" box="[336,478,1591,1618]" pageId="3" pageNumber="287" type="generic">5 specimens</specimenCount>
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,
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<quantity id="C02F9B2FF631FFBFD4A6664920FFE266" box="[489,653,1591,1618]" metricMagnitude="-2" metricUnit="m" metricValue="2.175" metricValueMax="2.4899999999999998" metricValueMin="1.86" pageId="3" pageNumber="287" unit="mm" value="21.75" valueMax="24.9" valueMin="18.6">18.6–24.9 mm</quantity>
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SL).
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<materialsCitation id="B7BF3C97F631FFBFD782664626F7E241" ID-GBIF-Occurrence="3313884304" collectionCode="PMR" country="Socotra Island" elevation="9" latitude="12.667434" location="Di Hamri" longLatPrecision="1" longitude="54.1793" pageId="3" pageNumber="287" specimenCode="VP4890" specimenCount="1" typeStatus="paratype">
|
||
<collectionCode id="61C6AE0FF631FFBFD7826646217AE266" box="[717,776,1592,1618]" country="0" httpUri="http://grbio.org/cool/xq4z-q5gq" name="Prirodoslovni muzej Rijeka" pageId="3" pageNumber="287">PMR</collectionCode>
|
||
<specimenCode id="57719EB1F631FFBFD65F66492119E265" box="[784,875,1591,1617]" collectionCode="PMR" country="0" httpUri="http://grbio.org/cool/xq4z-q5gq" name="Prirodoslovni muzej Rijeka" pageId="3" pageNumber="287">VP4890</specimenCode>
|
||
[sample tissue SOC19-355], male, 22.3+
|
||
<quantity id="C02F9B2FF631FFBFD00F664927EAE266" box="[1344,1432,1591,1618]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="3" pageNumber="287" unit="mm" value="5.1">5.1 mm</quantity>
|
||
,
|
||
<collectingCountry id="7FC0765AF631FFBFD5D866252348E241" box="[151,314,1627,1653]" name="Socotra Island" pageId="3" pageNumber="287">Socotra Island</collectingCountry>
|
||
,
|
||
<location id="02086011F631FFBFD40A662523C0E241" LSID="urn:lsid:plazi:treatment:8F7E87DCF631FFB6D5D865452293E7FD:02086011F631FFBFD40A662523C0E241" box="[325,434,1627,1653]" country="Socotra Island" latitude="12.667434" longLatPrecision="1" longitude="54.1793" name="Di Hamri" pageId="3" pageNumber="287">Di Hamri</location>
|
||
,
|
||
<geoCoordinate id="62E3500DF631FFBFD4F266252024E242" box="[445,598,1627,1654]" degrees="12" direction="north" minutes="40.046" orientation="latitude" pageId="3" pageNumber="287" precision="1" value="12.667434">12°40.046’ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="62E3500DF631FFBFD72E66252084E242" box="[609,758,1627,1654]" degrees="54" direction="east" minutes="10.758" orientation="longitude" pageId="3" pageNumber="287" precision="1" value="54.1793">54°10.758’ E</geoCoordinate>
|
||
, a small rock, depth
|
||
<quantity id="C02F9B2FF631FFBFD69266252641E242" box="[989,1075,1627,1654]" metricMagnitude="0" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="3" pageNumber="287" unit="m" value="9.0" valueMax="10.0" valueMin="8.0">
|
||
<elevation id="8CFAD1F9F631FFBFD69266252641E242" box="[989,1075,1627,1654]" metricMagnitude="0" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="3" pageNumber="287" unit="m" value="9.0" valueMax="10.0" valueMin="8.0">8–10 m</elevation>
|
||
</quantity>
|
||
, coll. S
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="B7BF3C97F631FFBFD1C5662522A0E2AD" ID-GBIF-Occurrence="3313884306" collectingDate="2019-04-02" collectionCode="V" country="Socotra Island" location="Bogorodsky" pageId="3" pageNumber="287" specimenCount="1" typeStatus="paratype">
|
||
<collectionCode id="61C6AE0FF631FFBFD1C5662526D3E241" box="[1162,1185,1627,1653]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="287" type="Museum">V</collectionCode>
|
||
.
|
||
<location id="02086011F631FFBFD1E466252747E241" LSID="urn:lsid:plazi:treatment:8F7E87DCF631FFB6D5D865452293E7FD:02086011F631FFBFD1E466252747E241" box="[1195,1333,1627,1653]" country="Socotra Island" name="Bogorodsky" pageId="3" pageNumber="287">Bogorodsky</location>
|
||
,
|
||
<date id="7369100AF631FFBFD00F662522A0E2AD" pageId="3" pageNumber="287" value="2019-04-02">
|
||
<collectingDate id="632DE9E2F631FFBFD00F662522A0E2AD" pageId="3" pageNumber="287" value="2019-04-02">02 April 2019</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="B7BF3C97F631FFBFD59066FE20A1E289" ID-GBIF-Occurrence="3313884303" collectionCode="PMR" country="Socotra Island" elevation="9" latitude="12.667434" location="Di Hamri" longLatPrecision="1" longitude="54.1793" pageId="3" pageNumber="287" specimenCode="VP4891" specimenCount="1" typeStatus="paratype">
|
||
<collectionCode id="61C6AE0FF631FFBFD59066FE2368E2AE" box="[223,282,1664,1690]" country="0" httpUri="http://grbio.org/cool/xq4z-q5gq" name="Prirodoslovni muzej Rijeka" pageId="3" pageNumber="287">PMR</collectionCode>
|
||
<specimenCode id="57719EB1F631FFBFD46D6601230CE2AD" box="[290,382,1663,1689]" collectionCode="PMR" country="0" httpUri="http://grbio.org/cool/xq4z-q5gq" name="Prirodoslovni muzej Rijeka" pageId="3" pageNumber="287">VP4891</specimenCode>
|
||
[sample tissue SOC19-358], female, 18.6+
|
||
<quantity id="C02F9B2FF631FFBFD625660121B6E2AE" box="[874,964,1663,1690]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="3" pageNumber="287" unit="mm" value="4.3">4.3 mm</quantity>
|
||
,
|
||
<collectingCountry id="7FC0765AF631FFBFD69F66012607E2AD" box="[976,1141,1663,1689]" name="Socotra Island" pageId="3" pageNumber="287">Socotra Island</collectingCountry>
|
||
,
|
||
<location id="02086011F631FFBFD1CD66012682E2AD" LSID="urn:lsid:plazi:treatment:8F7E87DCF631FFB6D5D865452293E7FD:02086011F631FFBFD1CD66012682E2AD" box="[1154,1264,1663,1689]" country="Socotra Island" latitude="12.667434" longLatPrecision="1" longitude="54.1793" name="Di Hamri" pageId="3" pageNumber="287">Di Hamri</location>
|
||
,
|
||
<geoCoordinate id="62E3500DF631FFBFD1B2660127EAE2AE" box="[1277,1432,1663,1690]" degrees="12" direction="north" minutes="40.046" orientation="latitude" pageId="3" pageNumber="287" precision="1" value="12.667434">12°40.046’ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="62E3500DF631FFBFD5D866DD235DE28A" box="[151,303,1699,1726]" degrees="54" direction="east" minutes="10.758" orientation="longitude" pageId="3" pageNumber="287" precision="1" value="54.1793">54°10.758’ E</geoCoordinate>
|
||
, a small rock, depth
|
||
<quantity id="C02F9B2FF631FFBFD76C66DD200EE28A" box="[547,636,1699,1726]" metricMagnitude="0" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="3" pageNumber="287" unit="m" value="9.0" valueMax="10.0" valueMin="8.0">
|
||
<elevation id="8CFAD1F9F631FFBFD76C66DD200EE28A" box="[547,636,1699,1726]" metricMagnitude="0" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="3" pageNumber="287" unit="m" value="9.0" valueMax="10.0" valueMin="8.0">8–10 m</elevation>
|
||
</quantity>
|
||
, coll. S
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="B7BF3C97F631FFBFD79866DD2644E289" ID-GBIF-Occurrence="3313884312" box="[727,1078,1699,1725]" collectingDate="2019-04-02" collectionCode="V" country="Socotra Island" location="Bogorodsky" pageId="3" pageNumber="287" specimenCount="1" typeStatus="paratype">
|
||
<collectionCode id="61C6AE0FF631FFBFD79866DD209CE289" box="[727,750,1699,1725]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="287" type="Museum">V</collectionCode>
|
||
.
|
||
<location id="02086011F631FFBFD7B366DD21F4E289" LSID="urn:lsid:plazi:treatment:8F7E87DCF631FFB6D5D865452293E7FD:02086011F631FFBFD7B366DD21F4E289" box="[764,902,1699,1725]" country="Socotra Island" name="Bogorodsky" pageId="3" pageNumber="287">Bogorodsky</location>
|
||
,
|
||
<date id="7369100AF631FFBFD6DC66DD2644E289" box="[915,1078,1699,1725]" pageId="3" pageNumber="287" value="2019-04-02">
|
||
<collectingDate id="632DE9E2F631FFBFD6DC66DD2644E289" box="[915,1078,1699,1725]" pageId="3" pageNumber="287" value="2019-04-02">02 April 2019</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="B7BF3C97F631FFBFD10B66DD26B7E2D5" ID-GBIF-Occurrence="3313884310" accessionNumber="SMF 39501" collectionCode="SMF" country="Socotra Island" elevation="9" latitude="12.667434" location="Di Hamri" longLatPrecision="1" longitude="54.1793" pageId="3" pageNumber="287" specimenCount="1" typeStatus="paratype">
|
||
<collectionCode id="61C6AE0FF631FFBFD10B66DD260EE289" box="[1092,1148,1699,1725]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="3" pageNumber="287">SMF</collectionCode>
|
||
39501, female, 24.9+
|
||
<quantity id="C02F9B2FF631FFBFD03566DD22B7E2D6" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="3" pageNumber="287" unit="mm" value="5.3">5.3 mm</quantity>
|
||
,
|
||
<collectingCountry id="7FC0765AF631FFBFD59E66B92306E2D5" box="[209,372,1735,1761]" name="Socotra Island" pageId="3" pageNumber="287">Socotra Island</collectingCountry>
|
||
,
|
||
<location id="02086011F631FFBFD4CF66B9239FE2D5" LSID="urn:lsid:plazi:treatment:8F7E87DCF631FFB6D5D865452293E7FD:02086011F631FFBFD4CF66B9239FE2D5" box="[384,493,1735,1761]" country="Socotra Island" latitude="12.667434" longLatPrecision="1" longitude="54.1793" name="Di Hamri" pageId="3" pageNumber="287">Di Hamri</location>
|
||
,
|
||
<geoCoordinate id="62E3500DF631FFBFD4B766B920E0E2D6" box="[504,658,1735,1762]" degrees="12" direction="north" minutes="40.046" orientation="latitude" pageId="3" pageNumber="287" precision="1" value="12.667434">12°40.046’ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="62E3500DF631FFBFD7D266B92141E2D6" box="[669,819,1735,1762]" degrees="54" direction="east" minutes="10.758" orientation="longitude" pageId="3" pageNumber="287" precision="1" value="54.1793">54°10.758’ E</geoCoordinate>
|
||
, a small rock, depth
|
||
<quantity id="C02F9B2FF631FFBFD15466B92600E2D6" box="[1051,1138,1735,1762]" metricMagnitude="0" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="3" pageNumber="287" unit="m" value="9.0" valueMax="10.0" valueMin="8.0">
|
||
<elevation id="8CFAD1F9F631FFBFD15466B92600E2D6" box="[1051,1138,1735,1762]" metricMagnitude="0" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="3" pageNumber="287" unit="m" value="9.0" valueMax="10.0" valueMin="8.0">8–10 m</elevation>
|
||
</quantity>
|
||
, coll. S
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="B7BF3C97F631FFBFD18666B923CEE332" ID-GBIF-Occurrence="3313884305" collectingDate="2019-04-02" collectionCode="V" country="Socotra Island" location="Bogorodsky" pageId="3" pageNumber="287" specimenCount="1" typeStatus="paratype">
|
||
<collectionCode id="61C6AE0FF631FFBFD18666B92692E2D5" box="[1225,1248,1735,1761]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="287" type="Museum">V</collectionCode>
|
||
.
|
||
<location id="02086011F631FFBFD1A466B92707E2D5" LSID="urn:lsid:plazi:treatment:8F7E87DCF631FFB6D5D865452293E7FD:02086011F631FFBFD1A466B92707E2D5" box="[1259,1397,1735,1761]" country="Socotra Island" name="Bogorodsky" pageId="3" pageNumber="287">Bogorodsky</location>
|
||
,
|
||
<date id="7369100AF631FFBFD0CF66B92360E331" pageId="3" pageNumber="287" value="2019-04-02">
|
||
<collectingDate id="632DE9E2F631FFBFD0CF66B92360E331" pageId="3" pageNumber="287" value="2019-04-02">02 April 2019</collectingDate>
|
||
</date>
|
||
(
|
||
<figureCitation id="9FEC2A4FF631FFBFD46D669523C0E332" box="[290,434,1771,1798]" captionStart="FIGURE 5" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[192,1395,181,1920]" captionTargetId="figure-20@8.[192,1395,181,1920]" captionTargetPageId="8" captionText="FIGURE 5. Hetereleotris nasoramosa sp. nov.; A: SMF 39501, paratype, female, 24.9 mm SL, alive specimen, Di Hamri, Socotra Island; B: the same specimen, freshly collected; C: paratype, SMF 39503, male, 21.1 mm SL, alive specimen, Di Timri, Socotra Island; D: presumably male, alive specimen, not collected, Di Timri, Socotra Island. Photos by S.V. Bogorodsky." figureDoi="http://doi.org/10.5281/zenodo.5069891" httpUri="https://zenodo.org/record/5069891/files/figure.png" pageId="3" pageNumber="287">Fig. 5A & B</figureCitation>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="B7BF3C97F631FFBFD486669520B2E31D" ID-GBIF-Occurrence="3313884308" accessionNumber="SMF 39502" collectionCode="SMF" country="Socotra Island" elevation="9" latitude="12.667434" location="Di Hamri" longLatPrecision="1" longitude="54.1793" pageId="3" pageNumber="287" specimenCount="1" typeStatus="paratype">
|
||
<collectionCode id="61C6AE0FF631FFBFD48666952073E331" box="[457,513,1771,1797]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="3" pageNumber="287">SMF</collectionCode>
|
||
39502, presumably male, 20.7+
|
||
<quantity id="C02F9B2FF631FFBFD622669521B5E332" box="[877,967,1771,1798]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="3" pageNumber="287" unit="mm" value="4.6">4.6 mm</quantity>
|
||
,
|
||
<collectingCountry id="7FC0765AF631FFBFD69C66952605E331" box="[979,1143,1771,1797]" name="Socotra Island" pageId="3" pageNumber="287">Socotra Island</collectingCountry>
|
||
,
|
||
<location id="02086011F631FFBFD1CB66952683E331" LSID="urn:lsid:plazi:treatment:8F7E87DCF631FFB6D5D865452293E7FD:02086011F631FFBFD1CB66952683E331" box="[1156,1265,1771,1797]" country="Socotra Island" latitude="12.667434" longLatPrecision="1" longitude="54.1793" name="Di Hamri" pageId="3" pageNumber="287">Di Hamri</location>
|
||
,
|
||
<geoCoordinate id="62E3500DF631FFBFD1B1669527EAE332" box="[1278,1432,1771,1798]" degrees="12" direction="north" minutes="40.046" orientation="latitude" pageId="3" pageNumber="287" precision="1" value="12.667434">12°40.046’ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="62E3500DF631FFBFD5D86771235FE31E" box="[151,301,1807,1834]" degrees="54" direction="east" minutes="10.758" orientation="longitude" pageId="3" pageNumber="287" precision="1" value="54.1793">54°10.758’ E</geoCoordinate>
|
||
, a small rock, depth
|
||
<quantity id="C02F9B2FF631FFBFD7596771201FE31E" box="[534,621,1807,1834]" metricMagnitude="0" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="3" pageNumber="287" unit="m" value="9.0" valueMax="10.0" valueMin="8.0">
|
||
<elevation id="8CFAD1F9F631FFBFD7596771201FE31E" box="[534,621,1807,1834]" metricMagnitude="0" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="3" pageNumber="287" unit="m" value="9.0" valueMax="10.0" valueMin="8.0">8–10 m</elevation>
|
||
</quantity>
|
||
, coll. S
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="B7BF3C97F631FFBFD78B67712669E31D" ID-GBIF-Occurrence="3313884311" box="[708,1051,1807,1833]" collectingDate="2019-04-02" collectionCode="V" country="Socotra Island" location="Bogorodsky" pageId="3" pageNumber="287" specimenCount="1" typeStatus="paratype">
|
||
<collectionCode id="61C6AE0FF631FFBFD78B677120A9E31D" box="[708,731,1807,1833]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="287" type="Museum">V</collectionCode>
|
||
.
|
||
<location id="02086011F631FFBFD7A967712102E31D" LSID="urn:lsid:plazi:treatment:8F7E87DCF631FFB6D5D865452293E7FD:02086011F631FFBFD7A967712102E31D" box="[742,880,1807,1833]" country="Socotra Island" name="Bogorodsky" pageId="3" pageNumber="287">Bogorodsky</location>
|
||
,
|
||
<date id="7369100AF631FFBFD63367712669E31D" box="[892,1051,1807,1833]" pageId="3" pageNumber="287" value="2019-04-02">
|
||
<collectingDate id="632DE9E2F631FFBFD63367712669E31D" box="[892,1051,1807,1833]" pageId="3" pageNumber="287" value="2019-04-02">02 April 2019</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="B7BF3C97F631FFBFD1686771276BE379" ID-GBIF-Occurrence="3313884302" accessionNumber="SMF 39503" collectionCode="SMF" country="Socotra Island" elevation="10" latitude="12.62025" location="Di Timri" longLatPrecision="1" longitude="54.2867" pageId="3" pageNumber="287" specimenCount="1" typeStatus="paratype">
|
||
<collectionCode id="61C6AE0FF631FFBFD1686771262DE31D" box="[1063,1119,1807,1833]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="3" pageNumber="287">SMF</collectionCode>
|
||
39503, male, 21.1+
|
||
<quantity id="C02F9B2FF631FFBFD070677127EAE31E" box="[1343,1432,1807,1834]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="3" pageNumber="287" unit="mm" value="4.5">4.5 mm</quantity>
|
||
,
|
||
<collectingCountry id="7FC0765AF631FFBFD5D8674D2348E379" box="[151,314,1843,1869]" name="Socotra Island" pageId="3" pageNumber="287">Socotra Island</collectingCountry>
|
||
,
|
||
<location id="02086011F631FFBFD40A674D23DBE379" LSID="urn:lsid:plazi:treatment:8F7E87DCF631FFB6D5D865452293E7FD:02086011F631FFBFD40A674D23DBE379" box="[325,425,1843,1869]" country="Socotra Island" latitude="12.62025" longLatPrecision="1" longitude="54.2867" name="Di Timri" pageId="3" pageNumber="287">Di Timri</location>
|
||
,
|
||
<geoCoordinate id="62E3500DF631FFBFD4FB674D203FE37A" box="[436,589,1843,1870]" degrees="12" direction="north" minutes="37.215" orientation="latitude" pageId="3" pageNumber="287" precision="1" value="12.62025">12°37.215’ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="62E3500DF631FFBFD718674D209EE37A" box="[599,748,1843,1870]" degrees="54" direction="east" minutes="17.202" orientation="longitude" pageId="3" pageNumber="287" precision="1" value="54.2867">54°17.202’ E</geoCoordinate>
|
||
, rocky ridge with large stones, depth
|
||
<quantity id="C02F9B2FF631FFBFD1C2674D26B5E37A" box="[1165,1223,1843,1870]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="287" unit="m" value="10.0">
|
||
<elevation id="8CFAD1F9F631FFBFD1C2674D26B5E37A" box="[1165,1223,1843,1870]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="287" unit="m" value="10.0">10 m</elevation>
|
||
</quantity>
|
||
, coll. S
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="B7BF3C97F631FFBFD052674D2306E345" ID-GBIF-Occurrence="3313884307" collectingDate="2019-04-08" collectionCode="V" country="Socotra Island" location="Bogorodsky" pageId="3" pageNumber="287" specimenCount="1" typeStatus="paratype">
|
||
<collectionCode id="61C6AE0FF631FFBFD052674D2746E379" box="[1309,1332,1843,1869]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="287" type="Museum">V</collectionCode>
|
||
.
|
||
<location id="02086011F631FFBFD070674A22BFE345" LSID="urn:lsid:plazi:treatment:8F7E87DCF631FFB6D5D865452293E7FD:02086011F631FFBFD070674A22BFE345" country="Socotra Island" name="Bogorodsky" pageId="3" pageNumber="287">Bogorodsky</location>
|
||
,
|
||
<date id="7369100AF631FFBFD59767292306E345" box="[216,372,1879,1905]" pageId="3" pageNumber="287" value="2019-04-08">
|
||
<collectingDate id="632DE9E2F631FFBFD59767292306E345" box="[216,372,1879,1905]" pageId="3" pageNumber="287" value="2019-04-08">08 April 2019</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
(
|
||
<figureCitation id="9FEC2A4FF631FFBFD4CC672923AEE346" box="[387,476,1879,1906]" captionStart="FIGURE 5" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[192,1395,181,1920]" captionTargetId="figure-20@8.[192,1395,181,1920]" captionTargetPageId="8" captionText="FIGURE 5. Hetereleotris nasoramosa sp. nov.; A: SMF 39501, paratype, female, 24.9 mm SL, alive specimen, Di Hamri, Socotra Island; B: the same specimen, freshly collected; C: paratype, SMF 39503, male, 21.1 mm SL, alive specimen, Di Timri, Socotra Island; D: presumably male, alive specimen, not collected, Di Timri, Socotra Island. Photos by S.V. Bogorodsky." figureDoi="http://doi.org/10.5281/zenodo.5069891" httpUri="https://zenodo.org/record/5069891/files/figure.png" pageId="3" pageNumber="287">Fig. 5C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4FCD6541F631FFB8D58867052676E5BD" lastPageId="4" lastPageNumber="288" pageId="3" pageNumber="287" type="diagnosis">
|
||
<paragraph id="076836CAF631FFB8D58867052676E5BD" blockId="3.[151,1437,1482,2014]" lastBlockId="4.[151,1436,151,393]" lastPageId="4" lastPageNumber="288" pageId="3" pageNumber="287">
|
||
<emphasis id="35A3EAD8F631FFBFD58867052348E3A1" bold="true" box="[199,314,1915,1941]" pageId="3" pageNumber="287">Diagnosis</emphasis>
|
||
. Dorsal-fin rays VI+I,12; anal-fin rays I,11; pectoral-fin rays 15–16, all rays branched; pelvic-fin rays I,5, fins separated and without frenum, fifth ray unbranched; anterior nostril a moderately long tube with the process extending from median side of rim (as tentacle longer than tube) or as a slender flap slightly shorter than tube; posterior nostril a long tube with two tentacles, anterior long and posterior shorter; no tentacle above eye; mouth moderately large, posterior angle of jaws behind vertical through posterior edge of eye; no opercular spine; small mental frenum present; pelvic fins ending distantly well in front of anus; head and body mainly naked except for several ctenoid scales on caudal peduncle at caudal-fin base; head with anterior oculoscapular canal and preopercular canal, posterior oculoscapular canal absent; anterior oculoscapular canal pores with erected rim; suborbital rows of papillae in five transverse rows; nearly rounded dark blue or blackish spot as large as pupil above opercle; a large irregular white blotch on postorbital head, cheek and lower preopercle.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="53A86642F636FFB8D5D867F923B0E3D3" ID-DOI="http://doi.org/10.5281/zenodo.5069883" ID-Zenodo-Dep="5069883" httpUri="https://zenodo.org/record/5069883/files/figure.png" pageId="4" pageNumber="288" startId="4.[151,250,1927,1951]" targetBox="[194,1388,423,1894]" targetPageId="4">
|
||
<paragraph id="076836CAF636FFB8D5D867F923B0E3D3" blockId="4.[151,1436,1927,2023]" pageId="4" pageNumber="288">
|
||
<emphasis id="35A3EAD8F636FFB8D5D867F92361E3AB" bold="true" box="[151,275,1927,1951]" pageId="4" pageNumber="288">FIGURE 1.</emphasis>
|
||
Bayesian molecular phylogenetic analysis based on combined mtDNA and nuclear genes. Colored circles at nodes depict posterior probability support values. Nodes with support less than 0.50 are collapsed into polytomies. Names of lineages follow
|
||
<bibRefCitation id="63464B3BF636FFB8D59167B123CCE3D3" author="Agorreta, A. & San Mauro, D. & Schliewen, U. & Van Tassell, J. L. & Kovacic, M. & Zardoya, R. & Ruber, L." box="[222,446,1999,2023]" pageId="4" pageNumber="288" pagination="619 - 633" refId="ref12280" refString="Agorreta, A., San Mauro, D., Schliewen, U., Van Tassell, J. L., Kovacic, M., Zardoya, R. & Ruber, L. (2013) Molecular phy- logenetics of Gobioidei and phylogenetic placement of European gobies. Molecular Phylogenetics and Evolution, 69, 619 - 633. https: // doi. org / 10.1016 / j. ympev. 2013.07.017" type="journal article" year="2013">
|
||
Agorreta
|
||
<emphasis id="35A3EAD8F636FFB8D47067AE2319E3D3" box="[319,363,1999,2023]" italics="true" pageId="4" pageNumber="288">et al</emphasis>
|
||
. (2013)
|
||
</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="53A86642F637FFB9D5D86401205CE17F" ID-DOI="http://doi.org/10.5281/zenodo.5069885" ID-Zenodo-Dep="5069885" httpUri="https://zenodo.org/record/5069885/files/figure.png" pageId="5" pageNumber="289" startId="5.[151,250,1151,1175]" targetBox="[151,1436,181,1127]" targetPageId="5">
|
||
<paragraph id="076836CAF637FFB9D5D86401205CE17F" blockId="5.[151,1437,1151,1355]" pageId="5" pageNumber="289">
|
||
<emphasis id="35A3EAD8F637FFB9D5D864012367E0A3" bold="true" box="[151,277,1151,1175]" pageId="5" pageNumber="289">FIGURE 2.</emphasis>
|
||
<taxonomicName id="C0D74D49F637FFB9D45264012057E0A3" authority="Kovačić & Bogorodsky & Zajonz & Tornabene, 2021" authorityName="Kovačić & Bogorodsky & Zajonz & Tornabene" authorityYear="2021" box="[285,549,1151,1175]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="289" phylum="Chordata" rank="species" species="nasoramosa" status="sp. nov.">
|
||
<emphasis id="35A3EAD8F637FFB9D45264012057E0A3" box="[285,549,1151,1175]" italics="true" pageId="5" pageNumber="289">Hetereleotris nasoramosa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="35A3EAD8F637FFB9D7626401200DE0A3" bold="true" box="[557,639,1151,1175]" pageId="5" pageNumber="289">
|
||
<taxonomicNameLabel id="2E9057A3F637FFB9D7626401200DE0A3" box="[557,639,1151,1175]" pageId="5" pageNumber="289" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype, SMF 39500 [sample tissue SOC18-191], female, 20.9 mm SL, Di Hamri, Socotra Island:
|
||
<emphasis id="35A3EAD8F637FFB9D4CE64DD23E1E08F" bold="true" box="[385,403,1187,1211]" pageId="5" pageNumber="289">A</emphasis>
|
||
: head with AN - anterior nostril as a short tube with tentacle from the rim, PN - posterior nostril a long tube with very long anterior tentacle and shorter posterior tentacle extending from the rim, PAJ - posterior angle of jaws extends behind vertical through posterior edge of eye;
|
||
<emphasis id="35A3EAD8F637FFB9D731649520FDE137" bold="true" box="[638,655,1259,1283]" pageId="5" pageNumber="289">B</emphasis>
|
||
: pelvic fins completely separated and without frenum;
|
||
<emphasis id="35A3EAD8F637FFB9D19B64952694E137" bold="true" box="[1236,1254,1259,1283]" pageId="5" pageNumber="289">C</emphasis>
|
||
: caudal peduncle stained, with visible scales;
|
||
<emphasis id="35A3EAD8F637FFB9D4FA657123B5E113" bold="true" box="[437,455,1295,1319]" pageId="5" pageNumber="289">D</emphasis>
|
||
: nape stained, erected pores of anterior oculoscapular canal, terminology in text. Photos by S.V. Bogorodsky (
|
||
<emphasis id="35A3EAD8F637FFB9D46D654D2346E17F" bold="true" box="[290,308,1331,1355]" pageId="5" pageNumber="289">A</emphasis>
|
||
) and M. Kovačić (
|
||
<emphasis id="35A3EAD8F637FFB9D4BB654D2052E17F" bold="true" box="[500,544,1331,1355]" pageId="5" pageNumber="289">B-D</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="4FCD6541F637FFB5D5886507218BE219" lastPageId="9" lastPageNumber="293" pageId="5" pageNumber="289" type="description">
|
||
<paragraph id="076836CAF637FFBAD58865072775E575" blockId="5.[151,1437,1401,2040]" lastBlockId="6.[151,1437,150,321]" lastPageId="6" lastPageNumber="290" pageId="5" pageNumber="289">
|
||
<emphasis id="35A3EAD8F637FFB9D58865072323E1A7" bold="true" box="[199,337,1401,1427]" pageId="5" pageNumber="289">Description</emphasis>
|
||
(The morphometric values in the text are presented first for the
|
||
<typeStatus id="D86C8868F637FFB9D17F650726E0E1A7" box="[1072,1170,1401,1427]" pageId="5" pageNumber="289" type="holotype">holotype</typeStatus>
|
||
followed by ranges for
|
||
<typeStatus id="D86C8868F637FFB9D5D865E02374E18C" box="[151,262,1438,1464]" pageId="5" pageNumber="289" type="paratype">paratypes</typeStatus>
|
||
; meristic values, if variable, the same). Body elongate, depth at anal-fin origin 5.6 (6.0–6.3) in SL, laterally compressed posteriorly, caudal peduncle deep and short, caudal peduncle depth 1.4 (1.3–1.4 female
|
||
<typeStatus id="D86C8868F637FFB9D06565BC27EAE1E8" box="[1322,1432,1474,1500]" pageId="5" pageNumber="289" type="paratype">paratypes</typeStatus>
|
||
, 1.5–1.6 male
|
||
<typeStatus id="D86C8868F637FFB9D460659823EEE234" box="[303,412,1510,1536]" pageId="5" pageNumber="289" type="paratype">paratypes</typeStatus>
|
||
) of body depth at anal-fin origin, caudal peduncle depth 0.9 (0.9 female
|
||
<typeStatus id="D86C8868F637FFB9D18C65982743E234" box="[1219,1329,1510,1536]" pageId="5" pageNumber="289" type="paratype">paratypes</typeStatus>
|
||
, 1.1 male
|
||
<typeStatus id="D86C8868F637FFB9D5D866742376E210" box="[151,260,1546,1572]" pageId="5" pageNumber="289" type="paratype">paratypes</typeStatus>
|
||
) of caudal peduncle length (
|
||
<figureCitation id="9FEC2A4FF637FFB9D70E667720F4E210" box="[577,646,1545,1572]" captionStart="FIGURE 4" captionStartId="7.[151,250,1862,1886]" captionTargetBox="[151,1437,885,1839]" captionTargetId="figure-417@7.[151,1437,885,1839]" captionTargetPageId="7" captionText="FIGURE 4. Hetereleotris nasoramosa sp. nov., holotype, SMF 39500, female, 20.9 mm SL, Di Hamri, Socotra Island: A: freshly collected specimen; B: preserved specimen. Photos by S.V. Bogorodsky (A) and M. Kovačić (B)." figureDoi="http://doi.org/10.5281/zenodo.5069889" httpUri="https://zenodo.org/record/5069889/files/figure.png" pageId="5" pageNumber="289">Fig. 4</figureCitation>
|
||
). Head of moderate size, its length 3.4 (3.2–3.4) in SL, width 4.7 (4.8– 5.0) in SL, depth 5.1 (5.1–5.2) in SL, not depressed, its depth 1.1 (1.0–1.1) of width. Snout very steep in profile from lateral view, and short, length 0.5 (0.5) of eye diameter, 7.7 (7.7–8.1) in head length, no unpaired horn-like tentacle at the level of nostrils on snout midline. Anterior nostril a moderately long tube set over tip of upper lip with process extending from median side of rim (as tentacle longer than tube) or as slender flap slightly shorter than tube. Posterior nostril long tube, with tentacle longer than tube extending from anterior side of rim and tentacle shorter than tube extending from posterior side of rim (
|
||
<figureCitation id="9FEC2A4FF637FFB9D73E669F20B9E2C8" box="[625,715,1761,1788]" captionStart="FIGURE 2" captionStartId="5.[151,250,1151,1175]" captionTargetBox="[151,1436,181,1127]" captionTargetId="figure-26@5.[151,1436,181,1127]" captionTargetPageId="5" captionText="FIGURE 2. Hetereleotris nasoramosa sp. nov., holotype, SMF 39500 [sample tissue SOC18-191], female, 20.9 mm SL, Di Hamri, Socotra Island: A: head with AN - anterior nostril as a short tube with tentacle from the rim, PN - posterior nostril a long tube with very long anterior tentacle and shorter posterior tentacle extending from the rim, PAJ - posterior angle of jaws extends behind vertical through posterior edge of eye; B: pelvic fins completely separated and without frenum; C: caudal peduncle stained, with visible scales; D: nape stained, erected pores of anterior oculoscapular canal, terminology in text. Photos by S.V. Bogorodsky (A) and M. Kovačić (B-D)." figureDoi="http://doi.org/10.5281/zenodo.5069885" httpUri="https://zenodo.org/record/5069885/files/figure.png" pageId="5" pageNumber="289">Fig. 2A</figureCitation>
|
||
). Eyes dorsolateral, moderately large, eye diameter 4.0 (3.7–4.0) in head length, orbit projecting forward and slightly elevated above dorsal profile of head. Interorbital narrow, 4.4 (4.1–5.3) of eye diameter. No tentacle above eye. Nape straight and flat. Mouth slightly subterminal, oblique, jaws ending anteriorly unequally, lower jaw slightly retracted anteriorly. Mouth moderately large, posterior angle of jaws behind vertical through posterior edge of eye. Upper lip wider than cheek, slightly inflated, in preserved specimens flattened and extending upwards on cheek. Cheek very narrow, suborbital depth less than upper lip width. Each side at front of upper jaw with outer row of 4–5 large recurved caniniform teeth, followed by irregularly scattered band of small conical inner teeth. Side of jaw with a single lateral row of about 10 medium-sized caniniform teeth. Sides of front of lower jaw with outer row of 4–5 large caniniform teeth, followed by irregularly scattered band of small conical median teeth and inner row of 5–6 large widely-spaced caniniform teeth. Single row of about ten mediumsized caniniform teeth continuous laterally on side of jaw. Small mental frenum midventrally behind lower lip. Branchiostegal membranes fused from isthmus to below pectoral-fin base, gill openings restricted to pectoral-fin base. Most of lower limb of first gill arch joined to gill cover by membrane. No spines on preopercle.
|
||
</paragraph>
|
||
<caption id="53A86642F634FFBAD5D864422713E04D" ID-DOI="http://doi.org/10.5281/zenodo.5069887" ID-Zenodo-Dep="5069887" httpUri="https://zenodo.org/record/5069887/files/figure.png" pageId="6" pageNumber="290" startId="6.[151,250,1084,1108]" targetBox="[159,1415,366,1049]" targetPageId="6">
|
||
<paragraph id="076836CAF634FFBAD5D864422713E04D" blockId="6.[151,1436,1084,1145]" pageId="6" pageNumber="290">
|
||
<emphasis id="35A3EAD8F634FFBAD5D864422364E060" bold="true" box="[151,278,1084,1108]" pageId="6" pageNumber="290">FIGURE 3.</emphasis>
|
||
<taxonomicName id="C0D74D49F634FFBAD4506443205BE060" authority="Kovačić & Bogorodsky & Zajonz & Tornabene, 2021" authorityName="Kovačić & Bogorodsky & Zajonz & Tornabene" authorityYear="2021" box="[287,553,1085,1109]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="290" phylum="Chordata" rank="species" species="nasoramosa" status="sp. nov.">
|
||
<emphasis id="35A3EAD8F634FFBAD4506443205BE060" box="[287,553,1085,1109]" italics="true" pageId="6" pageNumber="290">Hetereleotris nasoramosa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="35A3EAD8F634FFBAD77D644320F7E061" bold="true" box="[562,645,1085,1109]" pageId="6" pageNumber="290">
|
||
<taxonomicNameLabel id="2E9057A3F634FFBAD77D644320F7E061" box="[562,645,1085,1109]" pageId="6" pageNumber="290" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Cephalic sensory papillae, paratype, PMR VP4890, male, 22.3 mm SL, Di Hamri, Socotra Island. The right side illustrated, as it is in better condition, terminology in text. Drawing by M. Kovačić.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="076836CAF634FFBAD58864D626F4E279" blockId="6.[151,1437,1191,2010]" pageId="6" pageNumber="290">
|
||
<emphasis id="35A3EAD8F634FFBAD58864D6228BE0F5" box="[199,249,1192,1217]" italics="true" pageId="6" pageNumber="290">Fins</emphasis>
|
||
. First dorsal fin with VI spines, second dorsal fin I,12; anal fin I,11; branched caudal-fin rays 15, segmented 17 (16:1, 17:3). Pectoral-fin rays left 16 and right cut (15:4, 16:2, right side cut in
|
||
<specimenCount id="11D1FD43F634FFBAD16A64B526B7E0D1" box="[1061,1221,1227,1253]" pageId="6" pageNumber="290" type="generic" typeStatus="paratypes">two paratypes</specimenCount>
|
||
), all rays branched, not free at tips. Pectoral girdle without flaps on anterior edge. Pelvic fins I,5+5,I, left and right fin completely separated, distant and without frenum (
|
||
<figureCitation id="9FEC2A4FF634FFBAD76F656D200BE119" box="[544,633,1299,1325]" captionStart="FIGURE 2" captionStartId="5.[151,250,1151,1175]" captionTargetBox="[151,1436,181,1127]" captionTargetId="figure-26@5.[151,1436,181,1127]" captionTargetPageId="5" captionText="FIGURE 2. Hetereleotris nasoramosa sp. nov., holotype, SMF 39500 [sample tissue SOC18-191], female, 20.9 mm SL, Di Hamri, Socotra Island: A: head with AN - anterior nostril as a short tube with tentacle from the rim, PN - posterior nostril a long tube with very long anterior tentacle and shorter posterior tentacle extending from the rim, PAJ - posterior angle of jaws extends behind vertical through posterior edge of eye; B: pelvic fins completely separated and without frenum; C: caudal peduncle stained, with visible scales; D: nape stained, erected pores of anterior oculoscapular canal, terminology in text. Photos by S.V. Bogorodsky (A) and M. Kovačić (B-D)." figureDoi="http://doi.org/10.5281/zenodo.5069885" httpUri="https://zenodo.org/record/5069885/files/figure.png" pageId="6" pageNumber="290">Fig. 2B</figureCitation>
|
||
). Pelvic-fin rays branched, except fifth ray, fourth ray the longest, rays progressively shorter towards the first ray and pelvic spine. Spines of first dorsal fin not elongate or filamentous, first to fifth spine of first dorsal fin about equal in length, third always slightly longer than others, sixth spine short; fourth or fifth to sixth spine of first dorsal fin barely reaching origin of second dorsal fin when folded down. Origin of first dorsal fin behind vertical at pectoral-fin base. Origin of anal fin posterior to vertical at origin of second dorsal fin, i.e. opposite about first segmented ray of second dorsal fin. Pectoral fins not reaching posteriorly to below origin of second dorsal fin or just reaching to vertical at the second dorsal-fin spine. Pelvic fins short, ending well in front of anus in both sexes, 0.6 (0.6–0.7) of distance between origin of fins and anus, shorter than pectoral fins, 0.9 (0.9) of pectoral-fin length. Caudal fin rounded, shorter than head, 1.3 (1.3–1.4) in head length.
|
||
</paragraph>
|
||
<paragraph id="076836CAF634FFBAD588662923E1E28D" blockId="6.[151,1437,1191,2010]" pageId="6" pageNumber="290">
|
||
<emphasis id="35A3EAD8F634FFBAD5886629233FE245" box="[199,333,1623,1649]" italics="true" pageId="6" pageNumber="290">Squamation</emphasis>
|
||
. Head and body naked except for several ctenoid scales on caudal peduncle at caudal-fin base: four to eight scales ventrally on lateral side and one scale on the dorsal lateral side, most specimens with one more scale midlaterally (
|
||
<figureCitation id="9FEC2A4FF634FFBAD46166E123F5E28D" box="[302,391,1695,1721]" captionStart="FIGURE 2" captionStartId="5.[151,250,1151,1175]" captionTargetBox="[151,1436,181,1127]" captionTargetId="figure-26@5.[151,1436,181,1127]" captionTargetPageId="5" captionText="FIGURE 2. Hetereleotris nasoramosa sp. nov., holotype, SMF 39500 [sample tissue SOC18-191], female, 20.9 mm SL, Di Hamri, Socotra Island: A: head with AN - anterior nostril as a short tube with tentacle from the rim, PN - posterior nostril a long tube with very long anterior tentacle and shorter posterior tentacle extending from the rim, PAJ - posterior angle of jaws extends behind vertical through posterior edge of eye; B: pelvic fins completely separated and without frenum; C: caudal peduncle stained, with visible scales; D: nape stained, erected pores of anterior oculoscapular canal, terminology in text. Photos by S.V. Bogorodsky (A) and M. Kovačić (B-D)." figureDoi="http://doi.org/10.5281/zenodo.5069885" httpUri="https://zenodo.org/record/5069885/files/figure.png" pageId="6" pageNumber="290">Fig. 2C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="076836CAF634FFBBD58866BD202CE721" blockId="6.[151,1437,1191,2010]" lastBlockId="7.[151,1437,150,861]" lastPageId="7" lastPageNumber="291" pageId="6" pageNumber="290">
|
||
<emphasis id="35A3EAD8F634FFBAD58866BD2395E2E9" box="[199,487,1731,1757]" italics="true" pageId="6" pageNumber="290">Cephalic sensory systems</emphasis>
|
||
(
|
||
<figureCitation id="9FEC2A4FF634FFBAD4B966BD202BE2E9" box="[502,601,1731,1757]" captionStart="FIGURE 2" captionStartId="5.[151,250,1151,1175]" captionTargetBox="[151,1436,181,1127]" captionTargetId="figure-26@5.[151,1436,181,1127]" captionTargetPageId="5" captionText="FIGURE 2. Hetereleotris nasoramosa sp. nov., holotype, SMF 39500 [sample tissue SOC18-191], female, 20.9 mm SL, Di Hamri, Socotra Island: A: head with AN - anterior nostril as a short tube with tentacle from the rim, PN - posterior nostril a long tube with very long anterior tentacle and shorter posterior tentacle extending from the rim, PAJ - posterior angle of jaws extends behind vertical through posterior edge of eye; B: pelvic fins completely separated and without frenum; C: caudal peduncle stained, with visible scales; D: nape stained, erected pores of anterior oculoscapular canal, terminology in text. Photos by S.V. Bogorodsky (A) and M. Kovačić (B-D)." figureDoi="http://doi.org/10.5281/zenodo.5069885" httpUri="https://zenodo.org/record/5069885/files/figure.png" pageId="6" pageNumber="290">Figs. 2D</figureCitation>
|
||
&
|
||
<figureCitation id="9FEC2A4FF634FFBAD73266BD20FFE2E9" box="[637,653,1731,1757]" captionStart="FIGURE 3" captionStartId="6.[151,250,1084,1108]" captionTargetBox="[159,1415,366,1049]" captionTargetId="figure-107@6.[159,1427,345,1061]" captionTargetPageId="6" captionText="FIGURE 3. Hetereleotris nasoramosa sp. nov. Cephalic sensory papillae, paratype, PMR VP4890, male, 22.3 mm SL, Di Hamri, Socotra Island. The right side illustrated, as it is in better condition, terminology in text. Drawing by M. Kovačić." figureDoi="http://doi.org/10.5281/zenodo.5069887" httpUri="https://zenodo.org/record/5069887/files/figure.png" pageId="6" pageNumber="290">3</figureCitation>
|
||
). Head with anterior oculoscapular canal and preopercular canal with pores
|
||
<emphasis id="35A3EAD8F634FFBAD591669923E2E335" bold="true" box="[222,400,1766,1793]" italics="true" pageId="6" pageNumber="290">σ, λ, κ, ω, α, ρ,</emphasis>
|
||
and
|
||
<emphasis id="35A3EAD8F634FFBAD48366992386E335" bold="true" box="[460,500,1767,1793]" italics="true" pageId="6" pageNumber="290">δ, ε</emphasis>
|
||
respectively. Pores in anterior oculoscapular canal with erected rim, nearly short tubes in some specimens (e.g.
|
||
<typeStatus id="D86C8868F634FFBAD4BF67752026E312" box="[496,596,1803,1830]" pageId="6" pageNumber="290" type="holotype">holotype</typeStatus>
|
||
) (
|
||
<figureCitation id="9FEC2A4FF634FFBAD724677520B4E311" box="[619,710,1803,1829]" captionStart="FIGURE 2" captionStartId="5.[151,250,1151,1175]" captionTargetBox="[151,1436,181,1127]" captionTargetId="figure-26@5.[151,1436,181,1127]" captionTargetPageId="5" captionText="FIGURE 2. Hetereleotris nasoramosa sp. nov., holotype, SMF 39500 [sample tissue SOC18-191], female, 20.9 mm SL, Di Hamri, Socotra Island: A: head with AN - anterior nostril as a short tube with tentacle from the rim, PN - posterior nostril a long tube with very long anterior tentacle and shorter posterior tentacle extending from the rim, PAJ - posterior angle of jaws extends behind vertical through posterior edge of eye; B: pelvic fins completely separated and without frenum; C: caudal peduncle stained, with visible scales; D: nape stained, erected pores of anterior oculoscapular canal, terminology in text. Photos by S.V. Bogorodsky (A) and M. Kovačić (B-D)." figureDoi="http://doi.org/10.5281/zenodo.5069885" httpUri="https://zenodo.org/record/5069885/files/figure.png" pageId="6" pageNumber="290">Fig. 2D</figureCitation>
|
||
). Pore
|
||
<emphasis id="35A3EAD8F634FFBAD65A6775215BE311" bold="true" box="[789,809,1803,1829]" italics="true" pageId="6" pageNumber="290">ω</emphasis>
|
||
distant from eye, pore
|
||
<emphasis id="35A3EAD8F634FFBAD17F6775264FE311" bold="true" box="[1072,1085,1803,1829]" italics="true" pageId="6" pageNumber="290">ρ</emphasis>
|
||
positioned quite dorsally close to the level of pore
|
||
<emphasis id="35A3EAD8F634FFBAD43B675123FAE37D" bold="true" box="[372,392,1839,1865]" italics="true" pageId="6" pageNumber="290">ω</emphasis>
|
||
, pore
|
||
<emphasis id="35A3EAD8F634FFBAD49F675023ACE37D" bold="true" box="[464,478,1838,1865]" italics="true" pageId="6" pageNumber="290">β</emphasis>
|
||
absent. Posterior oculoscapular canal absent. Rows of head sensory papillae counted on all types, at least one side was in sufficient condition to count (
|
||
<typeStatus id="D86C8868F634FFBAD6C9672D2198E359" box="[902,1002,1875,1901]" pageId="6" pageNumber="290" type="holotype">holotype</typeStatus>
|
||
with better right side used for
|
||
<figureCitation id="9FEC2A4FF634FFBAD005672D27FCE359" box="[1354,1422,1875,1901]" captionStart="FIGURE 3" captionStartId="6.[151,250,1084,1108]" captionTargetBox="[159,1415,366,1049]" captionTargetId="figure-107@6.[159,1427,345,1061]" captionTargetPageId="6" captionText="FIGURE 3. Hetereleotris nasoramosa sp. nov. Cephalic sensory papillae, paratype, PMR VP4890, male, 22.3 mm SL, Di Hamri, Socotra Island. The right side illustrated, as it is in better condition, terminology in text. Drawing by M. Kovačić." figureDoi="http://doi.org/10.5281/zenodo.5069887" httpUri="https://zenodo.org/record/5069887/files/figure.png" pageId="6" pageNumber="290">Fig. 3</figureCitation>
|
||
).
|
||
<emphasis id="35A3EAD8F634FFBAD5D867092335E3A5" box="[151,327,1911,1937]" italics="true" pageId="6" pageNumber="290">Preorbital rows</emphasis>
|
||
: snout with four median preorbital series, upper row
|
||
<emphasis id="35A3EAD8F634FFBAD6D0670621D8E3A5" box="[927,938,1912,1937]" italics="true" pageId="6" pageNumber="290">r</emphasis>
|
||
(1–2) middorsally between eyes above pore
|
||
<emphasis id="35A3EAD8F634FFBAD5D867E522D7E381" bold="true" box="[151,165,1947,1973]" italics="true" pageId="6" pageNumber="290">σ</emphasis>
|
||
and posterior nostrils, row
|
||
<emphasis id="35A3EAD8F634FFBAD49667E2239EE39D" box="[473,492,1947,1973]" italics="true" pageId="6" pageNumber="290">
|
||
s
|
||
<superScript id="F0A29B82F634FFBAD4AB67E5239EE39D" attach="left" box="[484,492,1947,1961]" fontSize="6" pageId="6" pageNumber="290">1</superScript>
|
||
</emphasis>
|
||
(1) at posterior nostril, row
|
||
<emphasis id="35A3EAD8F634FFBAD66667E2214EE39D" box="[809,828,1947,1973]" italics="true" pageId="6" pageNumber="290">
|
||
s
|
||
<superScript id="F0A29B82F634FFBAD67B67E5214EE39D" attach="left" box="[820,828,1947,1961]" fontSize="6" pageId="6" pageNumber="290">2</superScript>
|
||
</emphasis>
|
||
(1) below posterior nostril and row
|
||
<emphasis id="35A3EAD8F634FFBAD19D67E22697E39D" box="[1234,1253,1947,1973]" italics="true" pageId="6" pageNumber="290">
|
||
s
|
||
<superScript id="F0A29B82F634FFBAD19267E52697E39D" attach="left" box="[1245,1253,1947,1961]" fontSize="6" pageId="6" pageNumber="290">3</superScript>
|
||
</emphasis>
|
||
(2–3) above upper lip (rows
|
||
<emphasis id="35A3EAD8F634FFBAD47E67BE2336E3F9" box="[305,324,1983,2009]" italics="true" pageId="6" pageNumber="290">
|
||
s
|
||
<superScript id="F0A29B82F634FFBAD47367C12336E3F9" attach="left" box="[316,324,1983,1997]" fontSize="6" pageId="6" pageNumber="290">1</superScript>
|
||
</emphasis>
|
||
and
|
||
<emphasis id="35A3EAD8F634FFBAD43167BE23E2E3F9" box="[382,400,1983,2009]" italics="true" pageId="6" pageNumber="290">
|
||
s
|
||
<superScript id="F0A29B82F634FFBAD4C767C123E2E3F9" attach="left" box="[392,400,1983,1997]" fontSize="6" pageId="6" pageNumber="290">2</superScript>
|
||
</emphasis>
|
||
on
|
||
<figureCitation id="9FEC2A4FF634FFBAD4F267C12072E3ED" box="[445,512,1983,2009]" captionStart="FIGURE 3" captionStartId="6.[151,250,1084,1108]" captionTargetBox="[159,1415,366,1049]" captionTargetId="figure-107@6.[159,1427,345,1061]" captionTargetPageId="6" captionText="FIGURE 3. Hetereleotris nasoramosa sp. nov. Cephalic sensory papillae, paratype, PMR VP4890, male, 22.3 mm SL, Di Hamri, Socotra Island. The right side illustrated, as it is in better condition, terminology in text. Drawing by M. Kovačić." figureDoi="http://doi.org/10.5281/zenodo.5069887" httpUri="https://zenodo.org/record/5069887/files/figure.png" pageId="6" pageNumber="290">Fig. 3</figureCitation>
|
||
hidden by nostrils and tentacles). Lateral series
|
||
<emphasis id="35A3EAD8F634FFBAD16867BE2641E3ED" box="[1063,1075,1984,2009]" italics="true" pageId="6" pageNumber="290">c</emphasis>
|
||
on snout in four parts, superior rows: upper row
|
||
<emphasis id="35A3EAD8F635FFBBD41860E92319E490" box="[343,363,150,176]" italics="true" pageId="7" pageNumber="291">
|
||
c
|
||
<superScript id="F0A29B82F635FFBBD42C60E82319E490" attach="left" box="[355,363,150,164]" fontSize="6" pageId="7" pageNumber="291">2</superScript>
|
||
</emphasis>
|
||
(2) between anterior and posterior nostrils, lower row
|
||
<emphasis id="35A3EAD8F635FFBBD69D60E92195E490" box="[978,999,150,176]" italics="true" pageId="7" pageNumber="291">
|
||
c
|
||
<superScript id="F0A29B82F635FFBBD69060E82195E490" attach="left" box="[991,999,150,164]" fontSize="6" pageId="7" pageNumber="291">1</superScript>
|
||
</emphasis>
|
||
(2–3) at anterior nostril; inferior rows: upper horizontal
|
||
<emphasis id="35A3EAD8F635FFBBD41960C52319E4EF" box="[342,363,187,219]" italics="true" pageId="7" pageNumber="291">
|
||
c
|
||
<subScript id="9B53348FF635FFBBD42C60B32319E4EF" attach="left" box="[355,363,205,219]" fontSize="6" pageId="7" pageNumber="291">2</subScript>
|
||
</emphasis>
|
||
(3) anteriorly, and lower horizontal
|
||
<emphasis id="35A3EAD8F635FFBBD7B060C52161E4EF" box="[767,787,187,219]" italics="true" pageId="7" pageNumber="291">
|
||
c
|
||
<subScript id="9B53348FF635FFBBD64460B32161E4EF" attach="left" box="[779,787,205,219]" fontSize="6" pageId="7" pageNumber="291">1</subScript>
|
||
</emphasis>
|
||
(1–2) posteriorly above upper lip.
|
||
<emphasis id="35A3EAD8F635FFBBD1D560C5273EE4E0" box="[1178,1356,187,213]" italics="true" pageId="7" pageNumber="291">Suborbital rows</emphasis>
|
||
: row
|
||
<emphasis id="35A3EAD8F635FFBBD0C160C527EEE4E0" box="[1422,1436,187,212]" italics="true" pageId="7" pageNumber="291">a</emphasis>
|
||
absent; row
|
||
<emphasis id="35A3EAD8F635FFBBD45060A1235FE4CD" box="[287,301,223,249]" italics="true" pageId="7" pageNumber="291">b</emphasis>
|
||
short, behind vertical of posterior edge of eye and anteriorly close to suborbital row
|
||
<emphasis id="35A3EAD8F635FFBBD19360A12698E4CD" box="[1244,1258,223,249]" italics="true" pageId="7" pageNumber="291">4</emphasis>
|
||
; five transverse suborbital rows of sensory papillae (
|
||
<emphasis id="35A3EAD8F635FFBBD77F617D2028E529" box="[560,602,259,285]" italics="true" pageId="7" pageNumber="291">1–5</emphasis>
|
||
) present, row
|
||
<emphasis id="35A3EAD8F635FFBBD7B4617D217BE529" box="[763,777,259,285]" italics="true" pageId="7" pageNumber="291">1</emphasis>
|
||
oblique, reaching upper lip, rows
|
||
<emphasis id="35A3EAD8F635FFBBD1C5617D26EAE529" box="[1162,1176,259,285]" italics="true" pageId="7" pageNumber="291">2</emphasis>
|
||
and
|
||
<emphasis id="35A3EAD8F635FFBBD181617D26AEE529" box="[1230,1244,259,285]" italics="true" pageId="7" pageNumber="291">3</emphasis>
|
||
short and distant from eye, close to row
|
||
<emphasis id="35A3EAD8F635FFBBD4DB615923D0E575" box="[404,418,295,321]" italics="true" pageId="7" pageNumber="291">d</emphasis>
|
||
, row
|
||
<emphasis id="35A3EAD8F635FFBBD4AF6159239CE575" box="[480,494,295,321]" italics="true" pageId="7" pageNumber="291">4</emphasis>
|
||
long, curved, from posterior edge of eye slanting to posterior end of row
|
||
<emphasis id="35A3EAD8F635FFBBD05861592757E575" box="[1303,1317,295,321]" italics="true" pageId="7" pageNumber="291">d</emphasis>
|
||
or slightly below it, row
|
||
<emphasis id="35A3EAD8F635FFBBD47B61352330E551" box="[308,322,331,357]" italics="true" pageId="7" pageNumber="291">5</emphasis>
|
||
close to pore
|
||
<emphasis id="35A3EAD8F635FFBBD4AD61352383E551" bold="true" box="[482,497,331,357]" italics="true" pageId="7" pageNumber="291">α</emphasis>
|
||
(
|
||
<emphasis id="35A3EAD8F635FFBBD74E6135207DE551" box="[513,527,331,357]" italics="true" pageId="7" pageNumber="291">1</emphasis>
|
||
: 6–9,
|
||
<emphasis id="35A3EAD8F635FFBBD71861352017E551" box="[599,613,331,357]" italics="true" pageId="7" pageNumber="291">2</emphasis>
|
||
: 4–5,
|
||
<emphasis id="35A3EAD8F635FFBBD7E2613520C9E551" box="[685,699,331,357]" italics="true" pageId="7" pageNumber="291">3</emphasis>
|
||
: 5–7,
|
||
<emphasis id="35A3EAD8F635FFBBD64C61352163E551" box="[771,785,331,357]" italics="true" pageId="7" pageNumber="291">4</emphasis>
|
||
: 13–19,
|
||
<emphasis id="35A3EAD8F635FFBBD63A613521FEE551" box="[885,908,331,357]" italics="true" pageId="7" pageNumber="291">5:</emphasis>
|
||
4); row
|
||
<emphasis id="35A3EAD8F635FFBBD6A261352189E551" box="[1005,1019,331,357]" italics="true" pageId="7" pageNumber="291">d</emphasis>
|
||
(6+10 – 7+13) long, above posterior part of upper lip, and continuing on lower cheek to meet suborbital row
|
||
<emphasis id="35A3EAD8F635FFBBD68B611121A0E5BD" box="[964,978,367,393]" italics="true" pageId="7" pageNumber="291">4</emphasis>
|
||
.
|
||
<emphasis id="35A3EAD8F635FFBBD6AF61112740E5BC" box="[992,1330,367,393]" italics="true" pageId="7" pageNumber="291">Preoperculo-mandibular rows</emphasis>
|
||
: external row
|
||
<emphasis id="35A3EAD8F635FFBBD58561ED22A4E598" box="[202,214,403,428]" italics="true" pageId="7" pageNumber="291">e</emphasis>
|
||
(14+10 – 18+18) longitudinal and uniserial, divided into anterior and posterior sections; internal row
|
||
<emphasis id="35A3EAD8F635FFBBD01761ED2712E598" box="[1368,1376,403,428]" italics="true" pageId="7" pageNumber="291">i</emphasis>
|
||
(7+8 – 10+8) longitudinal and uniserial, divided into anterior and posterior sections, mental row
|
||
<emphasis id="35A3EAD8F635FFBBD1FB61C926CEE5E5" box="[1204,1212,439,465]" italics="true" pageId="7" pageNumber="291">f</emphasis>
|
||
(4) behind frenum.
|
||
<emphasis id="35A3EAD8F635FFBBD5D861A5230EE5C0" box="[151,380,475,501]" italics="true" pageId="7" pageNumber="291">Oculoscapular rows</emphasis>
|
||
: three larger papillae longitudinally arranged on position of missing posterior oculoscapular canal above opercle, marked
|
||
<emphasis id="35A3EAD8F635FFBBD4AF6181207AE62C" box="[480,520,511,536]" italics="true" pageId="7" pageNumber="291">poc</emphasis>
|
||
. Anterior longitudinal row
|
||
<emphasis id="35A3EAD8F635FFBBD67261812120E638" box="[829,850,510,536]" italics="true" pageId="7" pageNumber="291">
|
||
x
|
||
<superScript id="F0A29B82F635FFBBD60561802120E638" attach="left" box="[842,850,510,524]" fontSize="6" pageId="7" pageNumber="291">1</superScript>
|
||
</emphasis>
|
||
(3+1 – 4+3) above level and behind pore
|
||
<emphasis id="35A3EAD8F635FFBBD0636181274BE62D" bold="true" box="[1324,1337,511,537]" italics="true" pageId="7" pageNumber="291">ρ</emphasis>
|
||
, divided in two parts, posterior longitudinal row
|
||
<emphasis id="35A3EAD8F635FFBBD711625D2000E604" box="[606,626,546,572]" italics="true" pageId="7" pageNumber="291">
|
||
x
|
||
<superScript id="F0A29B82F635FFBBD725625C2000E604" attach="left" box="[618,626,546,560]" fontSize="6" pageId="7" pageNumber="291">2</superScript>
|
||
</emphasis>
|
||
(3) above posterior opercular edge, transverse row
|
||
<emphasis id="35A3EAD8F635FFBBD18F625D26B9E608" box="[1216,1227,547,572]" italics="true" pageId="7" pageNumber="291">z</emphasis>
|
||
(7–10) below and anterior to pore
|
||
<emphasis id="35A3EAD8F635FFBBD40162392329E655" bold="true" box="[334,347,583,609]" italics="true" pageId="7" pageNumber="291">ρ</emphasis>
|
||
, row
|
||
<emphasis id="35A3EAD8F635FFBBD4D1623923D8E654" box="[414,426,583,608]" italics="true" pageId="7" pageNumber="291">y</emphasis>
|
||
(1) below and behind row
|
||
<emphasis id="35A3EAD8F635FFBBD79062392086E660" box="[735,756,582,608]" italics="true" pageId="7" pageNumber="291">
|
||
x
|
||
<superScript id="F0A29B82F635FFBBD7A362382086E660" attach="right" box="[748,756,582,596]" fontSize="6" pageId="7" pageNumber="291">2</superScript>
|
||
</emphasis>
|
||
. Transverse axillary rows
|
||
<emphasis id="35A3EAD8F635FFBBD16F62392633E660" box="[1056,1089,582,608]" italics="true" pageId="7" pageNumber="291">
|
||
as
|
||
<superScript id="F0A29B82F635FFBBD17662382633E660" attach="left" box="[1081,1089,582,596]" fontSize="6" pageId="7" pageNumber="291">1</superScript>
|
||
</emphasis>
|
||
(3) and
|
||
<emphasis id="35A3EAD8F635FFBBD1D0623926B2E660" box="[1183,1216,582,608]" italics="true" pageId="7" pageNumber="291">
|
||
as
|
||
<superScript id="F0A29B82F635FFBBD1F7623826B2E660" attach="left" box="[1208,1216,582,596]" fontSize="6" pageId="7" pageNumber="291">2</superScript>
|
||
</emphasis>
|
||
(2–3) and
|
||
<emphasis id="35A3EAD8F635FFBBD07562392729E660" box="[1338,1371,582,608]" italics="true" pageId="7" pageNumber="291">
|
||
as
|
||
<superScript id="F0A29B82F635FFBBD01C62382729E660" attach="left" box="[1363,1371,582,596]" fontSize="6" pageId="7" pageNumber="291">3</superScript>
|
||
</emphasis>
|
||
(1–2) present, row
|
||
<emphasis id="35A3EAD8F635FFBBD4676215233BE64C" box="[296,329,618,645]" italics="true" pageId="7" pageNumber="291">
|
||
la
|
||
<superScript id="F0A29B82F635FFBBD40E6214233BE64C" attach="left" box="[321,329,618,632]" fontSize="6" pageId="7" pageNumber="291">1</superScript>
|
||
</emphasis>
|
||
(2) above between rows
|
||
<emphasis id="35A3EAD8F635FFBBD72E621520F0E64C" box="[609,642,618,644]" italics="true" pageId="7" pageNumber="291">
|
||
as
|
||
<superScript id="F0A29B82F635FFBBD735621420F0E64C" attach="left" box="[634,642,618,632]" fontSize="6" pageId="7" pageNumber="291">1</superScript>
|
||
</emphasis>
|
||
and
|
||
<emphasis id="35A3EAD8F635FFBBD7F8621520AAE64C" box="[695,728,618,644]" italics="true" pageId="7" pageNumber="291">
|
||
as
|
||
<superScript id="F0A29B82F635FFBBD79F621420AAE64C" attach="left" box="[720,728,618,632]" fontSize="6" pageId="7" pageNumber="291">2</superScript>
|
||
</emphasis>
|
||
not visible in all specimens and
|
||
<emphasis id="35A3EAD8F635FFBBD10C62152617E64C" box="[1091,1125,618,645]" italics="true" pageId="7" pageNumber="291">
|
||
la
|
||
<superScript id="F0A29B82F635FFBBD11262142617E64C" attach="none" box="[1117,1125,618,632]" fontSize="6" pageId="7" pageNumber="291">2</superScript>
|
||
</emphasis>
|
||
not present (row
|
||
<emphasis id="35A3EAD8F635FFBBD06962152735E64C" box="[1318,1351,618,644]" italics="true" pageId="7" pageNumber="291">
|
||
as
|
||
<superScript id="F0A29B82F635FFBBD07062142735E64C" attach="left" box="[1343,1351,618,632]" fontSize="6" pageId="7" pageNumber="291">3</superScript>
|
||
</emphasis>
|
||
on
|
||
<figureCitation id="9FEC2A4FF635FFBBD03F621522D7E69D" captionStart="FIGURE 3" captionStartId="6.[151,250,1084,1108]" captionTargetBox="[159,1415,366,1049]" captionTargetId="figure-107@6.[159,1427,345,1061]" captionTargetPageId="6" captionText="FIGURE 3. Hetereleotris nasoramosa sp. nov. Cephalic sensory papillae, paratype, PMR VP4890, male, 22.3 mm SL, Di Hamri, Socotra Island. The right side illustrated, as it is in better condition, terminology in text. Drawing by M. Kovačić." figureDoi="http://doi.org/10.5281/zenodo.5069887" httpUri="https://zenodo.org/record/5069887/files/figure.png" pageId="7" pageNumber="291">Fig. 3</figureCitation>
|
||
hidden by pectoral fin).
|
||
<emphasis id="35A3EAD8F635FFBBD4F262F1201CE69C" box="[445,622,655,681]" italics="true" pageId="7" pageNumber="291">Opercular rows</emphasis>
|
||
: single papilla at location of missing pore
|
||
<emphasis id="35A3EAD8F635FFBBD11E62F12611E69D" bold="true" box="[1105,1123,655,681]" italics="true" pageId="7" pageNumber="291">γ,</emphasis>
|
||
marked
|
||
<emphasis id="35A3EAD8F635FFBBD18962F1269AE69C" box="[1222,1256,655,680]" italics="true" pageId="7" pageNumber="291">pc.</emphasis>
|
||
Transverse row
|
||
<emphasis id="35A3EAD8F635FFBBD5D862CD22DFE6F8" box="[151,173,691,716]" italics="true" pageId="7" pageNumber="291">ot</emphasis>
|
||
(11–13); superior longitudinal row
|
||
<emphasis id="35A3EAD8F635FFBBD70B62CD202FE6F8" box="[580,605,691,716]" italics="true" pageId="7" pageNumber="291">os</emphasis>
|
||
(4–5); and inferior longitudinal row
|
||
<emphasis id="35A3EAD8F635FFBBD14D62CD266AE6F8" box="[1026,1048,691,716]" italics="true" pageId="7" pageNumber="291">oi</emphasis>
|
||
(3) not visible in every specimen.
|
||
<emphasis id="35A3EAD8F635FFBBD5D862A923F9E6C4" box="[151,395,727,753]" italics="true" pageId="7" pageNumber="291">Anterior dorsal rows:</emphasis>
|
||
transverse row
|
||
<emphasis id="35A3EAD8F635FFBBD77262A92039E6C4" box="[573,587,727,752]" italics="true" pageId="7" pageNumber="291">n</emphasis>
|
||
(3–4) behind pore
|
||
<emphasis id="35A3EAD8F635FFBBD66C62A92145E6C5" bold="true" box="[803,823,727,753]" italics="true" pageId="7" pageNumber="291">ω</emphasis>
|
||
, rows
|
||
<emphasis id="35A3EAD8F635FFBBD6CC62A921E3E6C4" box="[899,913,727,752]" italics="true" pageId="7" pageNumber="291">o</emphasis>
|
||
and
|
||
<emphasis id="35A3EAD8F635FFBBD68862A921A9E6C4" box="[967,987,727,752]" italics="true" pageId="7" pageNumber="291">m</emphasis>
|
||
absent; row
|
||
<emphasis id="35A3EAD8F635FFBBD12362A92608E6C4" box="[1132,1146,727,752]" italics="true" pageId="7" pageNumber="291">g</emphasis>
|
||
longitudinal (6–8); row
|
||
<emphasis id="35A3EAD8F635FFBBD0C162A927EEE6C5" box="[1422,1436,727,753]" italics="true" pageId="7" pageNumber="291">h</emphasis>
|
||
(3–5) in front of the first dorsal-fin base.
|
||
</paragraph>
|
||
<paragraph id="076836CAF635FFBBD58863612289E769" blockId="7.[151,1437,150,861]" pageId="7" pageNumber="291">
|
||
<emphasis id="35A3EAD8F635FFBBD5886361234DE70D" box="[199,319,799,825]" italics="true" pageId="7" pageNumber="291">Osteology.</emphasis>
|
||
There are 10 precaudal vertebrae and 17 caudal vertebrae. The dorsal-fin pterygiophore pattern is 3-22110.
|
||
</paragraph>
|
||
<caption id="53A86642F635FFBBD5D8673826CFE3B7" ID-DOI="http://doi.org/10.5281/zenodo.5069889" ID-Zenodo-Dep="5069889" httpUri="https://zenodo.org/record/5069889/files/figure.png" pageId="7" pageNumber="291" startId="7.[151,250,1862,1886]" targetBox="[151,1437,885,1839]" targetPageId="7">
|
||
<paragraph id="076836CAF635FFBBD5D8673826CFE3B7" blockId="7.[151,1436,1862,1923]" pageId="7" pageNumber="291">
|
||
<emphasis id="35A3EAD8F635FFBBD5D867382367E36B" bold="true" box="[151,277,1862,1887]" pageId="7" pageNumber="291">FIGURE 4.</emphasis>
|
||
<taxonomicName id="C0D74D49F635FFBBD4506739205AE36A" authority="Kovačić & Bogorodsky & Zajonz & Tornabene, 2021" authorityName="Kovačić & Bogorodsky & Zajonz & Tornabene" authorityYear="2021" box="[287,552,1863,1887]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="291" phylum="Chordata" rank="species" species="nasoramosa" status="sp. nov.">
|
||
<emphasis id="35A3EAD8F635FFBBD4506739205AE36A" box="[287,552,1863,1887]" italics="true" pageId="7" pageNumber="291">Hetereleotris nasoramosa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="35A3EAD8F635FFBBD77E673920F6E36B" bold="true" box="[561,644,1863,1887]" pageId="7" pageNumber="291">
|
||
<taxonomicNameLabel id="2E9057A3F635FFBBD77E673920F6E36B" box="[561,644,1863,1887]" pageId="7" pageNumber="291" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype, SMF 39500, female, 20.9 mm SL, Di Hamri, Socotra Island:
|
||
<emphasis id="35A3EAD8F635FFBBD0CC673827E7E36A" bold="true" box="[1411,1429,1862,1886]" pageId="7" pageNumber="291">A</emphasis>
|
||
: freshly collected specimen;
|
||
<emphasis id="35A3EAD8F635FFBBD4FC671523B6E3B7" bold="true" box="[435,452,1899,1923]" pageId="7" pageNumber="291">B</emphasis>
|
||
: preserved specimen. Photos by S.V. Bogorodsky (
|
||
<emphasis id="35A3EAD8F635FFBBD683671421ACE3B6" bold="true" box="[972,990,1898,1922]" pageId="7" pageNumber="291">A</emphasis>
|
||
) and M. Kovačić (
|
||
<emphasis id="35A3EAD8F635FFBBD1D1671526DDE3B7" bold="true" box="[1182,1199,1899,1923]" pageId="7" pageNumber="291">B</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="53A86642F63AFFB4D5D867E92716E3CC" ID-DOI="http://doi.org/10.5281/zenodo.5069891" ID-Zenodo-Dep="5069891" httpUri="https://zenodo.org/record/5069891/files/figure.png" pageId="8" pageNumber="292" startId="8.[151,250,1943,1967]" targetBox="[192,1395,181,1920]" targetPageId="8">
|
||
<paragraph id="076836CAF63AFFB4D5D867E92716E3CC" blockId="8.[151,1437,1943,2040]" pageId="8" pageNumber="292">
|
||
<emphasis id="35A3EAD8F63AFFB4D5D867E92367E384" bold="true" box="[151,277,1943,1968]" pageId="8" pageNumber="292">FIGURE 5.</emphasis>
|
||
<taxonomicName id="C0D74D49F63AFFB4D45167E62055E39B" authority="Kovačić & Bogorodsky & Zajonz & Tornabene, 2021" authorityName="Kovačić & Bogorodsky & Zajonz & Tornabene" authorityYear="2021" box="[286,551,1944,1968]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="292" phylum="Chordata" rank="species" species="nasoramosa" status="sp. nov.">
|
||
<emphasis id="35A3EAD8F63AFFB4D45167E62055E39B" box="[286,551,1944,1968]" italics="true" pageId="8" pageNumber="292">Hetereleotris nasoramosa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="35A3EAD8F63AFFB4D77F67E620F0E384" bold="true" box="[560,642,1944,1968]" pageId="8" pageNumber="292">
|
||
<taxonomicNameLabel id="2E9057A3F63AFFB4D77F67E620F0E384" box="[560,642,1944,1968]" pageId="8" pageNumber="292" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
;
|
||
<emphasis id="35A3EAD8F63AFFB4D7DC67E920DFE39B" bold="true" box="[659,685,1943,1967]" pageId="8" pageNumber="292">A:</emphasis>
|
||
SMF 39501, paratype, female, 24.9 mm SL, alive specimen, Di Hamri, Socotra Island;
|
||
<emphasis id="35A3EAD8F63AFFB4D47A67C2233CE3E0" bold="true" box="[309,334,1980,2004]" pageId="8" pageNumber="292">B:</emphasis>
|
||
the same specimen, freshly collected;
|
||
<emphasis id="35A3EAD8F63AFFB4D79E67C52091E3E7" bold="true" box="[721,739,1979,2003]" pageId="8" pageNumber="292">C</emphasis>
|
||
: paratype, SMF 39503, male, 21.1 mm SL, alive specimen, Di Timri, Socotra Island;
|
||
<emphasis id="35A3EAD8F63AFFB4D479679E2323E3CC" bold="true" box="[310,337,2016,2040]" pageId="8" pageNumber="292">D:</emphasis>
|
||
presumably male, alive specimen, not collected, Di Timri, Socotra Island. Photos by S.V. Bogorodsky.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="076836CAF63BFFB5D58860E9264CE70D" blockId="9.[151,1437,150,2013]" pageId="9" pageNumber="293">
|
||
<emphasis id="35A3EAD8F63BFFB5D58860E9206BE485" box="[199,537,151,177]" italics="true" pageId="9" pageNumber="293">Color of fresh and live female</emphasis>
|
||
(
|
||
<figureCitation id="9FEC2A4FF63BFFB5D76660E920F1E485" box="[553,643,151,177]" captionStart="FIGURE 4" captionStartId="7.[151,250,1862,1886]" captionTargetBox="[151,1437,885,1839]" captionTargetId="figure-417@7.[151,1437,885,1839]" captionTargetPageId="7" captionText="FIGURE 4. Hetereleotris nasoramosa sp. nov., holotype, SMF 39500, female, 20.9 mm SL, Di Hamri, Socotra Island: A: freshly collected specimen; B: preserved specimen. Photos by S.V. Bogorodsky (A) and M. Kovačić (B)." figureDoi="http://doi.org/10.5281/zenodo.5069889" httpUri="https://zenodo.org/record/5069889/files/figure.png" pageId="9" pageNumber="293">Fig. 4A</figureCitation>
|
||
,
|
||
<typeStatus id="D86C8868F63BFFB5D7C060E92086E485" box="[655,756,151,177]" pageId="9" pageNumber="293" type="holotype">holotype</typeStatus>
|
||
, SMF 39500;
|
||
<figureCitation id="9FEC2A4FF63BFFB5D6DA60E92656E485" box="[917,1060,150,177]" captionStart="FIGURE 5" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[192,1395,181,1920]" captionTargetId="figure-20@8.[192,1395,181,1920]" captionTargetPageId="8" captionText="FIGURE 5. Hetereleotris nasoramosa sp. nov.; A: SMF 39501, paratype, female, 24.9 mm SL, alive specimen, Di Hamri, Socotra Island; B: the same specimen, freshly collected; C: paratype, SMF 39503, male, 21.1 mm SL, alive specimen, Di Timri, Socotra Island; D: presumably male, alive specimen, not collected, Di Timri, Socotra Island. Photos by S.V. Bogorodsky." figureDoi="http://doi.org/10.5281/zenodo.5069891" httpUri="https://zenodo.org/record/5069891/files/figure.png" pageId="9" pageNumber="293">Fig. 5A & B</figureCitation>
|
||
, SMF 39501). Body in life transparent, with numerous yellow-brown or brown dots, a series of six or seven short, double, saddle-like dark brown marks along back, four series of 2–5 irregular iridescent white marks alternating with dark brown dashes along vertebral column, and five series of reflected white spots ventrally on body visible in alive specimens, the first series as 4–5 spots on abdomen, the last, fifth series as 1–2 very small spots above posterior part of anal fin; two obscure dusky blotches on abdomen, first beneath the pectoral fin, the second just behind it. An obvious nearly rounded dark blue to blackish spot as large as pupil above opercle over posterior opercular edge at the level of the pupil is most striking coloration pattern. Head brown, darker anteriorly, with a large oblique, irregular, white blotch on postorbital head and most of cheek, with ventroposterior extension ending at lower edge of preopercle; behind large blotch several smaller whitish spots of various size and shape present over preopercle and opercle and on nape and predorsal area, and a small white spot at angle of jaws; head covered with dark brown to blackish dots, becoming black and more dense on cheek and preopercle. First dorsal fin with two brown oblique bands, one in outer part, another short in the middle of fin from first to fourth spine, with white rectangular mark at base of first two spines and oblique white band between two brown bands, tips from fourth to sixth spines white. The second dorsal fin with transparent membranes and greenish brown spine and rays, with white dots on rays forming three irregular oblique rows. Anal fin mostly transparent with widely scattered small melanophores, mostly at base. Caudal fin transparent, with greenish rays, clearly visible in freshly dead specimens, and melanophores mostly at the fin base. Pectoral-fin base within a large white blotch, extending on base of fin except between sixth to eighth rays, rest of fin transparent, yellow in freshly dead specimens. Pelvic fins dark brown, densely dotted with melanophores.
|
||
</paragraph>
|
||
<paragraph id="076836CAF63BFFB5D588633D2723E7FD" blockId="9.[151,1437,150,2013]" pageId="9" pageNumber="293">The overall live surface coloration pattern is retained in freshly dead specimens, with white marks on body less obvious and brown dotted coloration more obvious than in live specimens. Iris mottled brown with four faint radiating orange-red bars, pupil green-black. Body opaque white, with scattered melanophores concentrated in six double, short, brown bars along dorsal-fin bases, two below the first dorsal fin and four below the second dorsal fin.</paragraph>
|
||
<paragraph id="076836CAF63BFFB5D58863AD27EEE06D" blockId="9.[151,1437,150,2013]" pageId="9" pageNumber="293">
|
||
<emphasis id="35A3EAD8F63BFFB5D58863AD23DBE7D9" box="[199,425,979,1005]" italics="true" pageId="9" pageNumber="293">Color of live male (</emphasis>
|
||
<figureCitation id="9FEC2A4FF63BFFB5D4E663AD2073E7D9" box="[425,513,978,1005]" captionStart="FIGURE 5" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[192,1395,181,1920]" captionTargetId="figure-20@8.[192,1395,181,1920]" captionTargetPageId="8" captionText="FIGURE 5. Hetereleotris nasoramosa sp. nov.; A: SMF 39501, paratype, female, 24.9 mm SL, alive specimen, Di Hamri, Socotra Island; B: the same specimen, freshly collected; C: paratype, SMF 39503, male, 21.1 mm SL, alive specimen, Di Timri, Socotra Island; D: presumably male, alive specimen, not collected, Di Timri, Socotra Island. Photos by S.V. Bogorodsky." figureDoi="http://doi.org/10.5281/zenodo.5069891" httpUri="https://zenodo.org/record/5069891/files/figure.png" pageId="9" pageNumber="293">Fig. 5C</figureCitation>
|
||
,
|
||
<typeStatus id="D86C8868F63BFFB5D74063AD2000E7D9" box="[527,626,979,1005]" pageId="9" pageNumber="293" type="paratype">paratype</typeStatus>
|
||
, SMF 39503,
|
||
<figureCitation id="9FEC2A4FF63BFFB5D65963AD2102E7D9" box="[790,880,978,1005]" captionStart="FIGURE 5" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[192,1395,181,1920]" captionTargetId="figure-20@8.[192,1395,181,1920]" captionTargetPageId="8" captionText="FIGURE 5. Hetereleotris nasoramosa sp. nov.; A: SMF 39501, paratype, female, 24.9 mm SL, alive specimen, Di Hamri, Socotra Island; B: the same specimen, freshly collected; C: paratype, SMF 39503, male, 21.1 mm SL, alive specimen, Di Timri, Socotra Island; D: presumably male, alive specimen, not collected, Di Timri, Socotra Island. Photos by S.V. Bogorodsky." figureDoi="http://doi.org/10.5281/zenodo.5069891" httpUri="https://zenodo.org/record/5069891/files/figure.png" pageId="9" pageNumber="293">Fig. 5D</figureCitation>
|
||
). Material is limited, hence sexual dichromatism cannot be confirmed. Based on photograph of a male (
|
||
<figureCitation id="9FEC2A4FF63BFFB5D64F6389212AE025" box="[768,856,1014,1041]" captionStart="FIGURE 5" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[192,1395,181,1920]" captionTargetId="figure-20@8.[192,1395,181,1920]" captionTargetPageId="8" captionText="FIGURE 5. Hetereleotris nasoramosa sp. nov.; A: SMF 39501, paratype, female, 24.9 mm SL, alive specimen, Di Hamri, Socotra Island; B: the same specimen, freshly collected; C: paratype, SMF 39503, male, 21.1 mm SL, alive specimen, Di Timri, Socotra Island; D: presumably male, alive specimen, not collected, Di Timri, Socotra Island. Photos by S.V. Bogorodsky." figureDoi="http://doi.org/10.5281/zenodo.5069891" httpUri="https://zenodo.org/record/5069891/files/figure.png" pageId="9" pageNumber="293">Fig. 5C</figureCitation>
|
||
), head is not darker anteriorly than rest of the head as in the female, black spot above posterior edge of opercle is faint, a single white blotch on abdomen (vs. series of clusters of reflected white spots ventrally on body in females), and two dark bands on first dorsal fin are indistinct.
|
||
</paragraph>
|
||
<paragraph id="076836CAF63BFFB5D588641D218BE219" blockId="9.[151,1437,150,2013]" pageId="9" pageNumber="293">
|
||
<emphasis id="35A3EAD8F63BFFB5D588641D2063E048" box="[199,529,1123,1149]" italics="true" pageId="9" pageNumber="293">Color of preserved specimens</emphasis>
|
||
(
|
||
<figureCitation id="9FEC2A4FF63BFFB5D750641D2005E049" box="[543,631,1123,1149]" captionStart="FIGURE 4" captionStartId="7.[151,250,1862,1886]" captionTargetBox="[151,1437,885,1839]" captionTargetId="figure-417@7.[151,1437,885,1839]" captionTargetPageId="7" captionText="FIGURE 4. Hetereleotris nasoramosa sp. nov., holotype, SMF 39500, female, 20.9 mm SL, Di Hamri, Socotra Island: A: freshly collected specimen; B: preserved specimen. Photos by S.V. Bogorodsky (A) and M. Kovačić (B)." figureDoi="http://doi.org/10.5281/zenodo.5069889" httpUri="https://zenodo.org/record/5069889/files/figure.png" pageId="9" pageNumber="293">Fig. 4B</figureCitation>
|
||
): Coloration lost in preservation, body yellowish white, with dark brown dots from melanophores. No sexual dichromatism observed. Head dotted brown. Ventral side and mouth darker than rest of head. Large oblique irregular whitish marking behind and below eye. Smaller whitish spots of various size and shape on head opercle, preopercle, dorsal and ventral side, not prominent, formed by less dense pattern of melanophores compared to darker parts. Eye with dark iris and white pupil. Body with scattered melanophores more concentrated in dorsal part, six marks along both dorsal-fin bases and ill-defined marks along lateral midline recognizable in some specimens. Intensive dark brown rounded mark above posterior opercular edge and pectoral fin is most conspicuous coloration pattern. Breast brown with many melanophores, rest of ventral side of body with more broadly spaced melanophores. First dorsal fin with two dark oblique bands, one in upper part, one in the middle of fin, alternating with whitish mark at anterior corner of fin base and oblique white band between two brown bands. Second dorsal fin marbled with dark and white spots. Anal fin transparent with rarely scattered small melanophores mostly at base. Caudal fin transparent, with melanophores at the fin base. Pectoral fins with few scattered melanophores, the rest of fin transparent. Pelvic fins dark dotted with melanophores.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4FCD6541F63BFFB5D5886649201CE379" pageId="9" pageNumber="293" type="etymology">
|
||
<paragraph id="076836CAF63BFFB5D58866492066E2AD" blockId="9.[151,1437,150,2013]" pageId="9" pageNumber="293">
|
||
<emphasis id="35A3EAD8F63BFFB5D5886649233AE265" bold="true" box="[199,328,1591,1617]" pageId="9" pageNumber="293">Etymology</emphasis>
|
||
. The specific name
|
||
<taxonomicName id="C0D74D49F63BFFB5D760664920CAE264" authorityName="Kovačić & Bogorodsky & Zajonz & Tornabene" authorityYear="2021" box="[559,696,1591,1616]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="293" phylum="Chordata" rank="species" species="nasoramosa">
|
||
<emphasis id="35A3EAD8F63BFFB5D760664920CAE264" box="[559,696,1591,1616]" italics="true" pageId="9" pageNumber="293">nasoramosa</emphasis>
|
||
</taxonomicName>
|
||
is formed from the combination of two Latin words
|
||
<emphasis id="35A3EAD8F63BFFB5D05D66492735E264" box="[1298,1351,1591,1616]" italics="true" pageId="9" pageNumber="293">naso</emphasis>
|
||
(nose), and
|
||
<emphasis id="35A3EAD8F63BFFB5D58966252368E240" box="[198,282,1627,1652]" italics="true" pageId="9" pageNumber="293">ramosa</emphasis>
|
||
(branched) in reference to the well-developed, branched process extending from the rim of the anterior and posterior nostril, respectively.
|
||
</paragraph>
|
||
<paragraph id="076836CAF63BFFB5D58866DC201CE379" blockId="9.[151,1437,150,2013]" pageId="9" pageNumber="293">
|
||
<emphasis id="35A3EAD8F63BFFB5D58866DC2382E288" bold="true" box="[199,496,1698,1725]" pageId="9" pageNumber="293">Habitat and Distribution</emphasis>
|
||
.
|
||
<materialsCitation id="B7BF3C97F63BFFB5D4B266DD2018E379" ID-GBIF-Occurrence="3313884309" country="Socotra Island" elevation="10" location="Specimens" municipality="The" pageId="9" pageNumber="293" specimenCount="2" typeStatus="holotype">
|
||
<location id="02086011F63BFFB5D4B266DD2004E289" LSID="urn:lsid:plazi:treatment:8F7E87DCF631FFB6D5D865452293E7FD:02086011F63BFFB5D4B266DD2004E289" box="[509,630,1699,1725]" country="Socotra Island" municipality="The" name="Specimens" pageId="9" pageNumber="293">Specimens</location>
|
||
of
|
||
<taxonomicName id="C0D74D49F63BFFB5D7D366DD2134E288" authorityName="Kovačić & Bogorodsky & Zajonz & Tornabene" authorityYear="2021" box="[668,838,1699,1724]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="293" phylum="Chordata" rank="species" species="nasoramosa">
|
||
<emphasis id="35A3EAD8F63BFFB5D7D366DD2134E288" box="[668,838,1699,1724]" italics="true" pageId="9" pageNumber="293">H. nasoramosa</emphasis>
|
||
</taxonomicName>
|
||
were collected on rubble-sand patches in sandy areas and in areas with mixed corals and stones or rocks at several localities from the north-eastern part of the
|
||
<collectingCountry id="7FC0765AF63BFFB5D00866B922ADE331" name="Socotra Island" pageId="9" pageNumber="293">Socotra Island</collectingCountry>
|
||
.
|
||
<collectingMunicipality id="E70CACB0F63BFFB5D5A466952365E331" box="[235,279,1771,1797]" pageId="9" pageNumber="293">The</collectingMunicipality>
|
||
<typeStatus id="D86C8868F63BFFB5D450669523F3E331" box="[287,385,1771,1797]" pageId="9" pageNumber="293" type="holotype">holotype</typeStatus>
|
||
and
|
||
<typeStatus id="D86C8868F63BFFB5D4F466952054E331" box="[443,550,1771,1797]" pageId="9" pageNumber="293" type="paratype">paratypes</typeStatus>
|
||
were found living inside small holes in moderately large pieces of coral rock covered with short algae at depths of
|
||
<quantity id="C02F9B2FF63BFFB5D775677120E3E31D" box="[570,657,1807,1833]" metricMagnitude="0" metricUnit="m" metricValue="9.5" metricValueMax="11.0" metricValueMin="8.0" pageId="9" pageNumber="293" unit="m" value="9.5" valueMax="11.0" valueMin="8.0">
|
||
<elevation id="8CFAD1F9F63BFFB5D775677120E3E31D" box="[570,657,1807,1833]" metricMagnitude="0" metricUnit="m" metricValue="9.5" metricValueMax="11.0" metricValueMin="8.0" pageId="9" pageNumber="293" unit="m" value="9.5" valueMax="11.0" valueMin="8.0">8–11 m</elevation>
|
||
</quantity>
|
||
. The species has not yet been recognized from elsewhere and appears to be endemic to the Socotra Archipelago
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4FCD6541F63BFFB6D58867292293E7FD" lastPageId="10" lastPageNumber="294" pageId="9" pageNumber="293" type="discussion">
|
||
<paragraph id="076836CAF63BFFB6D588672923BFE721" blockId="9.[151,1437,150,2013]" lastBlockId="10.[151,1437,151,969]" lastPageId="10" lastPageNumber="294" pageId="9" pageNumber="293">
|
||
<emphasis id="35A3EAD8F63BFFB5D58867292346E345" bold="true" box="[199,308,1879,1905]" pageId="9" pageNumber="293">Remarks</emphasis>
|
||
. Based on the molecular phylogeny and several diagnostic morphological characters (see Generic identification above), the new species was assigned to
|
||
<emphasis id="35A3EAD8F63BFFB5D64E670521E9E3A1" box="[769,923,1915,1941]" italics="true" pageId="9" pageNumber="293">
|
||
<taxonomicName id="C0D74D49F63BFFB5D64E670521E5E3A1" authorityName="Bleeker" authorityYear="1874" box="[769,919,1915,1941]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="293" phylum="Chordata" rank="genus">Hetereleotris</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
It is unique though among
|
||
<taxonomicName id="C0D74D49F63BFFB5D19C670522B1E38D" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="293" phylum="Chordata" rank="species" species="undetermined">
|
||
<emphasis id="35A3EAD8F63BFFB5D19C67052714E3A1" box="[1235,1382,1915,1941]" italics="true" pageId="9" pageNumber="293">Hetereleotris</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
in having a simple tentacle or slender flap extending from the anterior nostril and anterior and posterior tentacles extending from the posterior nostril, and by having five suborbital transverse papillae rows. An obvious large (as large as pupil), dark spot above the opercle is shared only with
|
||
<taxonomicName id="C0D74D49F638FFB6D6C960E926A9E485" authority="Tortonese, 1976" authorityName="Tortonese" authorityYear="1976" box="[902,1243,151,177]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="294" phylum="Chordata" rank="species" species="bipunctata">
|
||
<emphasis id="35A3EAD8F638FFB6D6C960E92652E485" box="[902,1056,151,177]" italics="true" pageId="10" pageNumber="294">H. bipunctata</emphasis>
|
||
<bibRefCitation id="63464B3BF638FFB6D16860E926A9E485" author="Tortonese, E." box="[1063,1243,151,177]" pageId="10" pageNumber="294" pagination="187 - 193" refId="ref13783" refString="Tortonese, E. (1976) Gobioid fishes from the Gulf of Aden. Monitore Zoologico Italiano, New Series, Supplement 7 (4), 187 - 193. https: // doi. org / 10.1080 / 03749444.1976.10736826" type="journal article" year="1976">Tortonese, 1976</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="C0D74D49F638FFB6D05D60E9233DE4E1" authority="Hoese, 1986" authorityName="Hoese" authorityYear="1986" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="294" phylum="Chordata" rank="species" species="margaretae">
|
||
<emphasis id="35A3EAD8F638FFB6D05D60E922CBE4E0" italics="true" pageId="10" pageNumber="294">H. margaretae</emphasis>
|
||
<bibRefCitation id="63464B3BF638FFB6D58E60C5233DE4E1" author="Hoese, D. F." box="[193,335,187,213]" pageId="10" pageNumber="294" pagination="1 - 25" refId="ref12719" refString="Hoese, D. F. (1986) Descriptions of two new species of Hetereleotris (Pisces: Gobiidae) from the western Indian Ocean, with discussion of related species. J. L. B. Smith Institute of Ichthyology, Special Publication, 41, 1 - 25." type="journal article" year="1986">Hoese, 1986</bibRefCitation>
|
||
</taxonomicName>
|
||
. However, both latter species have coloration patterns distinguishing them from the new species. Among twenty
|
||
<taxonomicName id="C0D74D49F638FFB6D40860A12041E4CD" box="[327,563,223,249]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="294" phylum="Chordata" rank="species" species="undetermined">
|
||
<emphasis id="35A3EAD8F638FFB6D40860A123A8E4CD" box="[327,474,223,249]" italics="true" pageId="10" pageNumber="294">Hetereleotris</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
presently recognized, only the new species and
|
||
<taxonomicName id="C0D74D49F638FFB6D11860A127EEE4CD" authority="(Smith, 1958)" baseAuthorityName="Smith" baseAuthorityYear="1958" box="[1111,1436,222,249]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="294" phylum="Chordata" rank="species" species="tentaculata">
|
||
<emphasis id="35A3EAD8F638FFB6D11860A1268AE4CD" box="[1111,1272,223,249]" italics="true" pageId="10" pageNumber="294">H. tentaculata</emphasis>
|
||
(
|
||
<bibRefCitation id="63464B3BF638FFB6D04760A127E6E4CD" author="Smith, J. L. B." box="[1288,1428,222,249]" pageId="10" pageNumber="294" pagination="137 - 163" refId="ref13744" refString="Smith, J. L. B. (1958) The fishes of the family Eleotridae in the western Indian Ocean. Ichthyological Bulletin, Department of Ichthyology, Rhodes University, 11, 137 - 163." type="journal article" year="1958">Smith, 1958</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
share forward-set elevated eyes, a very short snout, and the posterior angle of the jaws behind vertical through the posterior edge of eye. The combination of head canals and pores seen in the new species is present otherwise only in
|
||
<taxonomicName id="C0D74D49F638FFB6D5FC61352392E551" authority="(Smith, 1958)" baseAuthorityName="Smith" baseAuthorityYear="1958" box="[179,480,330,357]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="294" phylum="Chordata" rank="species" species="caminata">
|
||
<emphasis id="35A3EAD8F638FFB6D5FC6135234FE550" box="[179,317,331,356]" italics="true" pageId="10" pageNumber="294">H. caminata</emphasis>
|
||
(
|
||
<bibRefCitation id="63464B3BF638FFB6D404613523AAE551" author="Smith, J. L. B." box="[331,472,330,357]" pageId="10" pageNumber="294" pagination="137 - 163" refId="ref13744" refString="Smith, J. L. B. (1958) The fishes of the family Eleotridae in the western Indian Ocean. Ichthyological Bulletin, Department of Ichthyology, Rhodes University, 11, 137 - 163." type="journal article" year="1958">Smith, 1958</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="C0D74D49F638FFB6D4A4613520F9E551" baseAuthorityName="Smith" baseAuthorityYear="1958" box="[491,651,331,357]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="294" phylum="Chordata" rank="species" species="tentaculata">
|
||
<emphasis id="35A3EAD8F638FFB6D4A4613520F9E551" box="[491,651,331,357]" italics="true" pageId="10" pageNumber="294">H. tentaculata</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="35A3EAD8F638FFB6D78F61352137E551" box="[704,837,331,357]" italics="true" pageId="10" pageNumber="294">
|
||
<taxonomicName id="C0D74D49F638FFB6D78F61352132E551" baseAuthorityName="Klunzinger" baseAuthorityYear="1871" box="[704,832,331,357]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="294" phylum="Chordata" rank="species" species="vulgaris">H. vulgaris</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
The ctenoid scales, restricted to caudal peduncle only as a ventral patch and one or two individual scales dorsally, was reported among
|
||
<taxonomicName id="C0D74D49F638FFB6D15361112774E5BD" box="[1052,1286,367,393]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="294" phylum="Chordata" rank="species" species="undetermined">
|
||
<emphasis id="35A3EAD8F638FFB6D153611126DDE5BD" box="[1052,1199,367,393]" italics="true" pageId="10" pageNumber="294">Hetereleotris</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
only in some specimens of
|
||
<taxonomicName id="C0D74D49F638FFB6D47F61ED23A2E599" baseAuthorityName="Smith" baseAuthorityYear="1958" box="[304,464,403,429]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="294" phylum="Chordata" rank="species" species="tentaculata">
|
||
<emphasis id="35A3EAD8F638FFB6D47F61ED23A2E599" box="[304,464,403,429]" italics="true" pageId="10" pageNumber="294">H. tentaculata</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="63464B3BF638FFB6D4B561ED20E1E59A" author="Hoese, D. F." box="[506,659,403,430]" pageId="10" pageNumber="294" pagination="1 - 25" refId="ref12719" refString="Hoese, D. F. (1986) Descriptions of two new species of Hetereleotris (Pisces: Gobiidae) from the western Indian Ocean, with discussion of related species. J. L. B. Smith Institute of Ichthyology, Special Publication, 41, 1 - 25." type="journal article" year="1986">Hoese (1986)</bibRefCitation>
|
||
, in which it is variably present. The extreme reduction to a few scales is also present in
|
||
<taxonomicName id="C0D74D49F638FFB6D41361C92127E5E5" authority="Kovacic & Bogorodsky, 2019" authorityName="Kovacic & Bogorodsky" authorityYear="2019" box="[348,853,439,465]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="294" phylum="Chordata" rank="species" species="aurantiaca">
|
||
<emphasis id="35A3EAD8F638FFB6D41361C92388E5E4" box="[348,506,439,464]" italics="true" pageId="10" pageNumber="294">H. aurantiaca</emphasis>
|
||
Kovačić & Bogorodsky, 2019
|
||
</taxonomicName>
|
||
, but with a different pattern. All other
|
||
<taxonomicName id="C0D74D49F638FFB6D04561C9229AE5C1" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="294" phylum="Chordata" rank="species" species="undetermined">
|
||
<emphasis id="35A3EAD8F638FFB6D04561C927EFE5E5" box="[1290,1437,439,465]" italics="true" pageId="10" pageNumber="294">Hetereleotris</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
are more extensively scaled, at least on the caudal peduncle, or the body is completely scaleless.
|
||
<taxonomicName id="C0D74D49F638FFB6D06961A52339E62C" authorityName="Kovačić & Bogorodsky & Zajonz & Tornabene" authorityYear="2021" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="294" phylum="Chordata" rank="species" species="nasoramosa">
|
||
<emphasis id="35A3EAD8F638FFB6D06961A52339E62C" italics="true" pageId="10" pageNumber="294">Hetereleotris nasoramosa</emphasis>
|
||
</taxonomicName>
|
||
is morphologically most similar to
|
||
<taxonomicName id="C0D74D49F638FFB6D79A61812106E62D" baseAuthorityName="Smith" baseAuthorityYear="1958" box="[725,884,511,537]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="294" phylum="Chordata" rank="species" species="tentaculata">
|
||
<emphasis id="35A3EAD8F638FFB6D79A61812106E62D" box="[725,884,511,537]" italics="true" pageId="10" pageNumber="294">H. tentaculata</emphasis>
|
||
</taxonomicName>
|
||
, sharing the following combination of characters: dorsal-fin rays VI+I,12, anal fin rays I+I,11, presence of some kind of tentacles on the head, eyes forward-set and elevated, a very short snout, posterior angle of jaws behind vertical through posterior edge of eye, scales restricted to caudal peduncle in a similar pattern (at least in some
|
||
<taxonomicName id="C0D74D49F638FFB6D655621521CEE6B1" baseAuthorityName="Smith" baseAuthorityYear="1958" box="[794,956,619,645]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="294" phylum="Chordata" rank="species" species="tentaculata">
|
||
<emphasis id="35A3EAD8F638FFB6D655621521CEE6B1" box="[794,956,619,645]" italics="true" pageId="10" pageNumber="294">H. tentaculata</emphasis>
|
||
</taxonomicName>
|
||
specimens), and the same pattern of head canals and pores. However,
|
||
<taxonomicName id="C0D74D49F638FFB6D49862F1200BE69D" baseAuthorityName="Smith" baseAuthorityYear="1958" box="[471,633,655,681]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="294" phylum="Chordata" rank="species" species="tentaculata">
|
||
<emphasis id="35A3EAD8F638FFB6D49862F1200BE69D" box="[471,633,655,681]" italics="true" pageId="10" pageNumber="294">H. tentaculata</emphasis>
|
||
</taxonomicName>
|
||
has a tentacle over the eye, no tentacles on the nostrils and a different coloration, e.g. presence of a dark bar below first dorsal fin and absence of a dark humeral spot. The putatively close relationship of these two species based on morphology should be confirmed once
|
||
<taxonomicName id="C0D74D49F638FFB6D17962A926A5E6C5" baseAuthorityName="Smith" baseAuthorityYear="1958" box="[1078,1239,727,753]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="294" phylum="Chordata" rank="species" species="tentaculata">
|
||
<emphasis id="35A3EAD8F638FFB6D17962A926A5E6C5" box="[1078,1239,727,753]" italics="true" pageId="10" pageNumber="294">H. tentaculata</emphasis>
|
||
</taxonomicName>
|
||
material is available for molecular analysis.
|
||
</paragraph>
|
||
<paragraph id="076836CAF638FFB6D58863612293E7FD" blockId="10.[151,1437,151,969]" pageId="10" pageNumber="294">
|
||
<taxonomicName id="C0D74D49F638FFB6D588636123BDE70D" authorityName="Kovacic, Bogorodsky, Troyer & Tornabene" authorityYear="2019" box="[199,463,799,825]" class="Actinopterygii" family="Gobiidae" genus="Cerogobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="294" phylum="Chordata" rank="species" species="petrophilus">
|
||
<emphasis id="35A3EAD8F638FFB6D588636123BDE70D" box="[199,463,799,825]" italics="true" pageId="10" pageNumber="294">Cerogobius petrophilus</emphasis>
|
||
</taxonomicName>
|
||
was described in its own genus based on the presumed distinctive presence of a single tentacle on the snout midline between nostrils, with no other tentacles on the head. Compared to the new species,
|
||
<taxonomicName id="C0D74D49F638FFB6D5D86319234AE7B5" authorityName="Kovacic, Bogorodsky, Troyer & Tornabene" authorityYear="2019" box="[151,312,871,897]" class="Actinopterygii" family="Gobiidae" genus="Cerogobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="294" phylum="Chordata" rank="species" species="petrophilus">
|
||
<emphasis id="35A3EAD8F638FFB6D5D86319234AE7B5" box="[151,312,871,897]" italics="true" pageId="10" pageNumber="294">C. petrophilus</emphasis>
|
||
</taxonomicName>
|
||
is scaleless, with head canals and pores absent, with four suborbital transverse papillae rows and a different coloration (see
|
||
<bibRefCitation id="63464B3BF638FFB6D4E863F520E1E792" author="Kovacic, M. & Bogorodsky, S. V. & Troyer, E. M. & Tornabene, L." box="[423,659,907,934]" pageId="10" pageNumber="294" pagination="171 - 189" refId="ref13127" refString="Kovacic, M., Bogorodsky, S. V., Troyer, E. M. & Tornabene, L. (2019 b) Cerogobius petrophilus (Perciformes: Gobiidae), a new gobiid genus and species from the Red Sea. Zootaxa, 4565 (2), 171 - 189. https: // doi. org / 10.11646 / zootaxa. 4565.2.2" type="journal article" year="2019">
|
||
Kovačić
|
||
<emphasis id="35A3EAD8F638FFB6D74463F52049E791" box="[523,571,907,933]" italics="true" pageId="10" pageNumber="294">et al</emphasis>
|
||
. 2019b
|
||
</bibRefCitation>
|
||
). Its generic assignment is pending further investigation, as elaborated below.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |