400 lines
29 KiB
XML
400 lines
29 KiB
XML
<document ID-DOI="http://doi.org/10.5281/zenodo.5630219" ID-GBIF-Dataset="1db94db9-0aa3-41e5-a48a-b48152648f35" ID-GBIF-Taxon="120674243" ID-Zenodo-Dep="5630219" LSID="urn:lsid:zoobank.org:act:211BFA99-3B5E-4B7A-875F-3713EE0E804B" checkinTime="1462802976960" checkinUser="plazi" docAuthor="T. F. Flannery & C. P. Groves" docDate="1998" docId="03D60959FFF8FFFEFDC03A76FA06AD9E" docLanguage="en" docName="mamm.1998.62.3.367_text_over_image.pdf" docOrigin="Mammalia 62 (3)" docStyle="DocumentStyle{}" docTitle="Zaglossus attenboroughi Flannery & Groves, 1998, n. sp." docType="treatment" docUuid="211BFA99-3B5E-4B7A-875F-3713EE0E804B" docUuidSource="ZooBank" docVersion="11" lastPageId="23" lastPageNumber="389" masterDocId="FFEF7121FFECFFE9FFFE383EFFF0AC5C" masterDocTitle="A revision of the genus Zaglossus (Monotremata, Tachyglossidae), with description of new species and subspecies" masterLastPageNumber="396" masterPageNumber="367" pageId="20" pageNumber="387" updateTime="1635648814651" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A revision of the genus Zaglossus (Monotremata, Tachyglossidae), with description of new species and subspecies</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>T. F. Flannery</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>C. P. Groves</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Mammalia</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>1998</mods:date>
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<mods:detail type="volume">
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<mods:number>62</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>3</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>367</mods:start>
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<mods:end>396</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.1515/mamm.1998.62.3.367</mods:identifier>
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<mods:identifier type="GBIF-Dataset">1db94db9-0aa3-41e5-a48a-b48152648f35</mods:identifier>
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<mods:identifier type="Zenodo-Dep">270023</mods:identifier>
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<mods:identifier type="ZooBank">2285F206-66A5-4DA9-9ADE-EF15961FB919</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.5630219" ID-GBIF-Taxon="120674243" ID-Zenodo-Dep="5630219" LSID="urn:lsid:zoobank.org:act:211BFA99-3B5E-4B7A-875F-3713EE0E804B" httpUri="http://treatment.plazi.org/id/03D60959FFF8FFFEFDC03A76FA06AD9E" lastPageId="23" lastPageNumber="389" pageId="20" pageNumber="387">
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<subSubSection box="[574,1087,584,618]" pageId="20" pageNumber="387" type="nomenclature">
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<paragraph blockId="20.[573,1087,580,675]" box="[574,1087,584,618]" pageId="20" pageNumber="387">
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<heading box="[574,1087,584,618]" centered="true" fontSize="10" level="8" pageId="20" pageNumber="387" reason="4">
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<taxonomicName box="[574,984,584,618]" class="Mammalia" family="Tachyglossidae" genus="Zaglossus" kingdom="Animalia" order="Monotremata" pageId="20" pageNumber="387" phylum="Chordata" rank="species" species="attenboroughi" status="n. sp.">
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<emphasis box="[574,984,584,618]" italics="true" pageId="20" pageNumber="387">Zaglossus attenboroughi</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="20" pageNumber="387">
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<taxonomicNameLabel box="[1000,1087,584,618]" pageId="20" pageNumber="387" rank="species">n. sp.</taxonomicNameLabel>
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<subSubSection box="[620,1042,641,675]" pageId="20" pageNumber="387" type="description">
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<paragraph blockId="20.[573,1087,580,675]" box="[620,1042,641,675]" pageId="20" pageNumber="387">
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<heading box="[620,1042,641,675]" centered="true" fontSize="10" level="8" pageId="20" pageNumber="387" reason="4">
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(Appendix 1,
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<figureCitation box="[852,981,641,675]" captionStart="Fig. 10" captionStartId="20.[339,388,2447,2477]" captionTargetBox="[114,1526,1574,2394]" captionTargetPageId="20" captionText="Fig. 10. - X-ray of skull and pes of Zaglossus attenboroughi n. sp." pageId="20" pageNumber="387">Figs 10</figureCitation>
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<emphasis bold="true" box="[981,992,641,675]" pageId="20" pageNumber="387">-</emphasis>
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11
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<emphasis bold="true" box="[1031,1042,641,675]" pageId="20" pageNumber="387">)</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="20" pageNumber="387" type="materials_examined">
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<paragraph blockId="20.[95,1573,726,1491]" pageId="20" pageNumber="387">
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<emphasis bold="true" box="[179,359,726,760]" pageId="20" pageNumber="387">Holotype:</emphasis>
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RMNH 17301. Skin,
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<emphasis bold="true" box="[756,877,726,760]" pageId="20" pageNumber="387">dentary</emphasis>
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and
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<emphasis bold="true" box="[975,1173,726,760]" pageId="20" pageNumber="387">fragmentary</emphasis>
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rostral
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<emphasis bold="true" box="[1316,1419,726,760]" pageId="20" pageNumber="387">region</emphasis>
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of
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<emphasis bold="true" box="[1490,1567,726,760]" pageId="20" pageNumber="387">skull</emphasis>
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of
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<emphasis bold="true" box="[146,181,770,804]" pageId="20" pageNumber="387">an</emphasis>
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adult, sex not recorded.
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<emphasis bold="true" box="[615,770,770,804]" pageId="20" pageNumber="387">Collected</emphasis>
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“
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<emphasis bold="true" box="[817,969,770,804]" pageId="20" pageNumber="387">Oost-top,</emphasis>
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Berg
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<emphasis bold="true" box="[1083,1168,770,804]" pageId="20" pageNumber="387">Rara,</emphasis>
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op 1,600 m ”, Cyclops Mountains. Dutch New
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<emphasis bold="true" box="[500,626,813,847]" pageId="20" pageNumber="387">Guinea,</emphasis>
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on July 4th, 1961 by P.
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<emphasis bold="true" box="[1045,1102,813,847]" pageId="20" pageNumber="387">van</emphasis>
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Royen.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="20" pageNumber="387" type="etymology">
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<paragraph blockId="20.[95,1573,726,1491]" pageId="20" pageNumber="387">
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<emphasis bold="true" box="[179,389,876,910]" pageId="20" pageNumber="387">Etymology:</emphasis>
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For
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<emphasis bold="true" box="[494,541,876,910]" pageId="20" pageNumber="387">Sir</emphasis>
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David
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<emphasis bold="true" box="[685,922,876,910]" pageId="20" pageNumber="387">Attenborough,</emphasis>
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who has contributed greatly to the public appreciation of the New
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<emphasis bold="true" box="[627,871,919,953]" pageId="20" pageNumber="387">Guinean fauna</emphasis>
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and flora.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="20" pageNumber="387" type="diagnosis">
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<paragraph blockId="20.[95,1573,726,1491]" pageId="20" pageNumber="387">
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<emphasis bold="true" box="[180,370,982,1016]" pageId="20" pageNumber="387">Diagnosis:</emphasis>
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The
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<emphasis bold="true" box="[474,606,982,1016]" pageId="20" pageNumber="387">smallest</emphasis>
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member of the genus, with a beak length
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<emphasis bold="true" box="[1345,1514,982,1016]" pageId="20" pageNumber="387">(measured</emphasis>
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on the skin, see below
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<emphasis bold="true" box="[422,433,1026,1060]" pageId="20" pageNumber="387">)</emphasis>
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of 70
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<emphasis bold="true" box="[548,635,1026,1060]" pageId="20" pageNumber="387">mm (</emphasis>
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as
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<emphasis bold="true" box="[685,821,1026,1060]" pageId="20" pageNumber="387">opposed</emphasis>
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to x
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<emphasis bold="true" box="[901,925,1026,1060]" pageId="20" pageNumber="387">=</emphasis>
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99.1
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<emphasis box="[1020,1041,1026,1060]" italics="true" pageId="20" pageNumber="387">±</emphasis>
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13.6, range 8Î.0
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<emphasis bold="true" box="[1322,1333,1026,1060]" pageId="20" pageNumber="387">-</emphasis>
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127.5, N
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<emphasis bold="true" box="[1498,1519,1026,1060]" pageId="20" pageNumber="387">=</emphasis>
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23 for all other
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<emphasis box="[325,511,1070,1104]" italics="true" pageId="20" pageNumber="387">
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<taxonomicName box="[325,489,1070,1104]" class="Mammalia" family="Tachyglossidae" genus="Zaglossus" kingdom="Animalia" order="Monotremata" pageId="20" pageNumber="387" phylum="Chordata" rank="genus">Zaglossus</taxonomicName>
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<emphasis bold="true" box="[489,587,1070,1104]" pageId="20" pageNumber="387">), and</emphasis>
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</emphasis>
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a dentary
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<emphasis bold="true" box="[785,885,1070,1104]" pageId="20" pageNumber="387">length</emphasis>
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of 107.5 mm
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<emphasis bold="true" box="[1151,1160,1070,1104]" pageId="20" pageNumber="387">(</emphasis>
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x
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<emphasis bold="true" box="[1210,1231,1070,1104]" pageId="20" pageNumber="387">=</emphasis>
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138.1, range 118.0
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<emphasis bold="true" box="[1560,1570,1070,1104]" pageId="20" pageNumber="387">-</emphasis>
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157.0, N
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<emphasis bold="true" box="[256,277,1113,1147]" pageId="20" pageNumber="387">=</emphasis>
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22 for all other
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<taxonomicName box="[560,723,1113,1147]" class="Mammalia" family="Tachyglossidae" genus="Zaglossus" kingdom="Animalia" order="Monotremata" pageId="20" pageNumber="387" phylum="Chordata" rank="genus">
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<emphasis box="[560,723,1113,1147]" italics="true" pageId="20" pageNumber="387">Zaglossus</emphasis>
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</taxonomicName>
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<emphasis bold="true" box="[739,1003,1113,1147]" pageId="20" pageNumber="387">combined). The</emphasis>
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short beak is
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<emphasis bold="true" box="[1250,1486,1113,1147]" pageId="20" pageNumber="387">also straighter</emphasis>
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than in other
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<emphasis box="[241,413,1158,1192]" italics="true" pageId="20" pageNumber="387">
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<taxonomicName box="[241,407,1158,1192]" class="Mammalia" family="Tachyglossidae" genus="Zaglossus" kingdom="Animalia" order="Monotremata" pageId="20" pageNumber="387" phylum="Chordata" rank="genus">Zaglossus</taxonomicName>
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.
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</emphasis>
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There are 5 claws on each foot.
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<emphasis bold="true" box="[980,1043,1158,1192]" pageId="20" pageNumber="387">The</emphasis>
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fur is shorter,
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<emphasis bold="true" box="[1300,1573,1158,1192]" pageId="20" pageNumber="387">denser and finer</emphasis>
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than in other
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<taxonomicName box="[329,492,1202,1236]" class="Mammalia" family="Tachyglossidae" genus="Zaglossus" kingdom="Animalia" order="Monotremata" pageId="20" pageNumber="387" phylum="Chordata" rank="genus">
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<emphasis box="[329,492,1202,1236]" italics="true" pageId="20" pageNumber="387">Zaglossus</emphasis>
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</taxonomicName>
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and differs from
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<emphasis bold="true" box="[798,858,1202,1236]" pageId="20" pageNumber="387">that</emphasis>
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of other species
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<emphasis box="[1158,1185,1202,1236]" italics="true" pageId="20" pageNumber="387">in</emphasis>
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<emphasis bold="true" box="[1202,1292,1202,1236]" pageId="20" pageNumber="387">being</emphasis>
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raw umber
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<emphasis bold="true" box="[1504,1570,1202,1236]" pageId="20" pageNumber="387">dorsally</emphasis>
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and fawn ventrally.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="20" pageNumber="387" type="distribution">
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<paragraph blockId="20.[95,1573,726,1491]" pageId="20" pageNumber="387">
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<emphasis bold="true" box="[180,416,1308,1342]" pageId="20" pageNumber="387">Distribution:</emphasis>
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The higher
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<emphasis bold="true" box="[670,762,1308,1342]" pageId="20" pageNumber="387">peaks</emphasis>
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of the Cyclops Mountains near Jayapura, extreme northeastern Irian Jaya
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<emphasis bold="true" box="[637,648,1352,1386]" pageId="20" pageNumber="387">(</emphasis>
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<figureCitation box="[647,750,1352,1386]" captionStart="Fig. 1" captionStartId="1.[154,203,2404,2434]" captionTargetBox="[155,1587,1325,2361]" captionTargetPageId="1" captionText="Fig. 1. - Map of New Guinea showing the approximate distribution of the species and subspecies of Zaglossus." pageId="20" pageNumber="387">
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Fig.
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<emphasis bold="true" box="[733,769,1352,1386]" pageId="20" pageNumber="387">1).</emphasis>
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</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="22" lastPageNumber="389" pageId="20" pageNumber="387" type="description">
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<paragraph blockId="20.[95,1573,726,1491]" lastBlockId="21.[81,1595,1562,2479]" lastPageId="21" lastPageNumber="388" pageId="20" pageNumber="387">
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<emphasis bold="true" box="[180,405,1413,1447]" pageId="20" pageNumber="387">Description:</emphasis>
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<emphasis box="[429,491,1413,1447]" italics="true" pageId="20" pageNumber="387">The</emphasis>
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type and only specimen consists of a complete skin
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<emphasis bold="true" box="[1445,1503,1413,1447]" pageId="20" pageNumber="387">and</emphasis>
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the rostral portions of a badly crushed cranium
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<emphasis bold="true" box="[839,850,1457,1491]" pageId="20" pageNumber="387">(</emphasis>
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<figureCitation box="[850,973,1457,1491]" captionStart="Fig. 10" captionStartId="20.[339,388,2447,2477]" captionTargetBox="[114,1526,1574,2394]" captionTargetPageId="20" captionText="Fig. 10. - X-ray of skull and pes of Zaglossus attenboroughi n. sp." pageId="20" pageNumber="387">Fig. 10</figureCitation>
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<emphasis bold="true" box="[972,990,1457,1491]" pageId="20" pageNumber="387">).</emphasis>
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The fleshy part of the beak
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<emphasis bold="true" box="[1478,1488,1457,1491]" pageId="20" pageNumber="387">(</emphasis>
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measured along ils dorsal edge, from the tip to the margin of the fur
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<emphasis bold="true" box="[1221,1231,1567,1601]" pageId="21" pageNumber="388">)</emphasis>
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is 70 mm long. The beak
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<emphasis box="[179,204,1611,1645]" italics="true" pageId="21" pageNumber="388">is</emphasis>
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straight, but has been broken and possibly somewhat distorted
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<emphasis bold="true" box="[1319,1330,1611,1645]" pageId="21" pageNumber="388">(</emphasis>
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<figureCitation box="[1330,1453,1611,1645]" captionStart="Fig. 10" captionStartId="20.[339,388,2447,2477]" captionTargetBox="[114,1526,1574,2394]" captionTargetPageId="20" captionText="Fig. 10. - X-ray of skull and pes of Zaglossus attenboroughi n. sp." pageId="21" pageNumber="388">Fig. 10</figureCitation>
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<emphasis bold="true" box="[1453,1471,1611,1645]" pageId="21" pageNumber="388">).</emphasis>
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Nonetheless, it it likely that it was straighter
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<emphasis bold="true" box="[823,833,1655,1689]" pageId="21" pageNumber="388">(</emphasis>
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and certainly shorter
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<emphasis bold="true" box="[1204,1214,1655,1689]" pageId="21" pageNumber="388">)</emphasis>
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in life than in other
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<taxonomicName box="[81,252,1700,1734]" class="Mammalia" family="Tachyglossidae" genus="Zaglossus" kingdom="Animalia" order="Monotremata" pageId="21" pageNumber="388" phylum="Chordata" rank="genus">
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<emphasis box="[81,252,1700,1734]" italics="true" pageId="21" pageNumber="388">Zaglossus</emphasis>
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</taxonomicName>
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.
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</paragraph>
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<caption box="[339,1342,2446,2478]" pageId="20" pageNumber="387" targetBox="[114,1526,1574,2394]" targetPageId="20">
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<paragraph blockId="20.[338,1342,2442,2479]" box="[339,1342,2446,2478]" pageId="20" pageNumber="387">
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Fig. 10.
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<emphasis bold="true" box="[464,475,2447,2477]" pageId="20" pageNumber="387">-</emphasis>
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X-ray of skull and pes of
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<taxonomicName box="[884,1254,2446,2478]" class="Mammalia" family="Tachyglossidae" genus="Zaglossus" kingdom="Animalia" order="Monotremata" pageId="20" pageNumber="387" phylum="Chordata" rank="species" species="attenboroughi" status="n. sp.">
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<emphasis box="[884,1254,2446,2478]" italics="true" pageId="20" pageNumber="387">Zaglossus attenboroughi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[1266,1342,2447,2477]" pageId="20" pageNumber="387" rank="species">n. sp.</taxonomicNameLabel>
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</paragraph>
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</caption>
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<caption box="[428,1251,1452,1484]" httpUri="https://zenodo.org/record/3099694/files/figure.png" pageId="21" pageNumber="388" targetBox="[541,1128,291,1319]" targetPageId="21">
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<paragraph blockId="21.[428,1251,1449,1484]" box="[428,1251,1452,1484]" pageId="21" pageNumber="388">
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Fig.
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<emphasis bold="true" box="[505,576,1453,1483]" pageId="21" pageNumber="388">1 1.-</emphasis>
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Skin of holotype of
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<taxonomicName box="[899,1157,1452,1484]" class="Mammalia" family="Tachyglossidae" genus="Zaglossus" kingdom="Animalia" order="Monotremata" pageId="21" pageNumber="388" phylum="Chordata" rank="species" species="attenboroughi">
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<emphasis box="[899,922,1453,1483]" italics="true" pageId="21" pageNumber="388">Z</emphasis>
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|
.
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<emphasis box="[942,1157,1452,1484]" italics="true" pageId="21" pageNumber="388">attenboroughi</emphasis>
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</taxonomicName>
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». sp.
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</paragraph>
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</caption>
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<paragraph blockId="21.[81,1595,1562,2479]" pageId="21" pageNumber="388">
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In overall size, the specimen conforms closely with
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<emphasis box="[1074,1477,1743,1777]" italics="true" pageId="21" pageNumber="388">
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Tac
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<emphasis bold="true" box="[1134,1151,1743,1777]" italics="true" pageId="21" pageNumber="388">/</emphasis>
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iyglossus
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<taxonomicName box="[1315,1477,1743,1777]" class="Mammalia" family="Tachyglossidae" genus="Tachyglossus" kingdom="Animalia" order="Monotremata" pageId="21" pageNumber="388" phylum="Chordata" rank="species" species="aculeatus">aculeatus</taxonomicName>
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</emphasis>
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. being much smaller than other
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<emphasis box="[546,723,1786,1820]" italics="true" pageId="21" pageNumber="388">
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<taxonomicName box="[546,717,1786,1820]" class="Mammalia" family="Tachyglossidae" genus="Zaglossus" kingdom="Animalia" order="Monotremata" pageId="21" pageNumber="388" phylum="Chordata" rank="genus">Zaglossus</taxonomicName>
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.
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</emphasis>
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On the basis of comparison with similar-sized
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<emphasis box="[84,318,1829,1863]" italics="true" pageId="21" pageNumber="388">ï'achyglossus.</emphasis>
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its weight
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<emphasis box="[536,560,1829,1863]" italics="true" pageId="21" pageNumber="388">is</emphasis>
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estimated to have been approximately two to three kilograms.
|
|
</paragraph>
|
|
<paragraph blockId="21.[81,1595,1562,2479]" pageId="21" pageNumber="388">
|
|
There are five claws on each foot. There is a small spur
|
|
<emphasis bold="true" box="[1168,1179,1917,1951]" pageId="21" pageNumber="388">(</emphasis>
|
|
4.4 mm long x 1.7 mm wide at its base
|
|
<emphasis bold="true" box="[355,366,1960,1994]" pageId="21" pageNumber="388">)</emphasis>
|
|
present on the left ankle. The base is shielded by a fleshy tubercle. A remnant of a similar spur is present on the right foot. These spurs suggest that the specimen is male.
|
|
</paragraph>
|
|
<paragraph blockId="21.[81,1595,1562,2479]" pageId="21" pageNumber="388">
|
|
The fur is dense, very fine and short, in all of these characteristics differing from other
|
|
<taxonomicName box="[188,356,2136,2170]" class="Mammalia" family="Tachyglossidae" genus="Zaglossus" kingdom="Animalia" order="Monotremata" pageId="21" pageNumber="388" phylum="Chordata" rank="genus">
|
|
<emphasis box="[188,356,2136,2170]" italics="true" pageId="21" pageNumber="388">Zaglossus</emphasis>
|
|
</taxonomicName>
|
|
<emphasis bold="true" box="[379,390,2136,2170]" pageId="21" pageNumber="388">(</emphasis>
|
|
where the fur is longer, coarser and less dense
|
|
<emphasis bold="true" box="[1247,1265,2136,2170]" pageId="21" pageNumber="388">).</emphasis>
|
|
It is a distinctive brown colour
|
|
<emphasis bold="true" box="[325,336,2180,2214]" pageId="21" pageNumber="388">(</emphasis>
|
|
close to raw umber
|
|
<emphasis bold="true" box="[671,681,2180,2214]" pageId="21" pageNumber="388">)</emphasis>
|
|
unmatched in any other
|
|
<taxonomicName box="[1114,1286,2180,2214]" class="Mammalia" family="Tachyglossidae" genus="Zaglossus" kingdom="Animalia" order="Monotremata" pageId="21" pageNumber="388" phylum="Chordata" rank="genus">
|
|
<emphasis box="[1114,1286,2180,2214]" italics="true" pageId="21" pageNumber="388">Zaglossus</emphasis>
|
|
</taxonomicName>
|
|
, which vary from reddish brown to black, with an occasional very pale variant. The venter is fawn, again differing from any other member of the genus. The fur of the feet, tail and forehead are yellowish in colour.
|
|
</paragraph>
|
|
<paragraph blockId="21.[81,1595,1562,2479]" lastBlockId="22.[96,1602,288,1488]" lastPageId="22" lastPageNumber="389" pageId="21" pageNumber="388">
|
|
There are two hair whorls on the ventral surface on either side of the neck. The short fur covers the spines in the middle of the back, which are extremely sparse, and large portions of the midback entirely lack spines. The spines are narrow
|
|
<emphasis bold="true" box="[1415,1425,2444,2478]" pageId="21" pageNumber="388">(</emphasis>
|
|
averaging 1.6 mm in diameter at their thickest point
|
|
<emphasis bold="true" box="[823,833,293,327]" pageId="22" pageNumber="389">)</emphasis>
|
|
and are all whitish in colour. The thickest
|
|
<emphasis box="[96,196,337,371]" italics="true" pageId="22" pageNumber="389">spines</emphasis>
|
|
occur near the rump. Spines are absent from the ventral surface.
|
|
</paragraph>
|
|
<paragraph blockId="22.[96,1602,288,1488]" pageId="22" pageNumber="389">
|
|
Only small fragments of the skull remain, the most important being the dentaries, a crushed rostrum and portions of the orbital and frontal regions. The region of the naso frontal suture is well preserved, and although portions of the suture are visible through the slightly translucent bone, the suture is entirely closed, indicating that the specimen
|
|
<emphasis box="[270,293,556,590]" italics="true" pageId="22" pageNumber="389">is</emphasis>
|
|
adult
|
|
<emphasis bold="true" box="[409,420,556,590]" pageId="22" pageNumber="389">(</emphasis>
|
|
<figureCitation box="[420,545,556,590]" captionStart="Fig. 12" captionStartId="22.[194,243,2449,2479]" captionTargetBox="[566,1122,1542,2398]" captionTargetPageId="22" captionText="Fig. 12. - Left orbital region (distal end downwards) of Zaglossus attenboroughi n. sp." pageId="22" pageNumber="389">Fig. 12</figureCitation>
|
|
<emphasis bold="true" box="[544,563,556,590]" pageId="22" pageNumber="389">).</emphasis>
|
|
The sutures present on other fragments are all closed.
|
|
<emphasis box="[1531,1594,556,590]" italics="true" pageId="22" pageNumber="389">The</emphasis>
|
|
dentary differs from juveniles in the form of its articulation
|
|
<emphasis bold="true" box="[1137,1148,600,634]" pageId="22" pageNumber="389">(</emphasis>
|
|
which is well-defined
|
|
<emphasis bold="true" box="[1538,1596,600,634]" pageId="22" pageNumber="389">and</emphasis>
|
|
arched sharply above the horizontal ramus
|
|
<emphasis bold="true" box="[824,917,644,678]" pageId="22" pageNumber="389">), and</emphasis>
|
|
in
|
|
<emphasis bold="true" box="[980,1015,644,678]" pageId="22" pageNumber="389">its</emphasis>
|
|
more gradual taper. The articular facet is more rounded than in other
|
|
<taxonomicName box="[706,872,689,723]" class="Mammalia" family="Tachyglossidae" genus="Zaglossus" kingdom="Animalia" order="Monotremata" pageId="22" pageNumber="389" phylum="Chordata" rank="genus">
|
|
<emphasis box="[706,872,689,723]" italics="true" pageId="22" pageNumber="389">Zaglossus</emphasis>
|
|
</taxonomicName>
|
|
<emphasis bold="true" box="[888,1059,689,723]" pageId="22" pageNumber="389">examined.</emphasis>
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection lastPageId="23" lastPageNumber="390" pageId="22" pageNumber="389" type="discussion">
|
|
<paragraph blockId="22.[96,1602,288,1488]" pageId="22" pageNumber="389">
|
|
<emphasis bold="true" box="[183,387,750,784]" pageId="22" pageNumber="389">Discussion:</emphasis>
|
|
This species is sample 6
|
|
<emphasis box="[855,882,750,784]" italics="true" pageId="22" pageNumber="389">in</emphasis>
|
|
our analysis. The small size and narrow
|
|
<emphasis box="[101,200,794,828]" italics="true" pageId="22" pageNumber="389">spines</emphasis>
|
|
of the holotype made us suspect initially that
|
|
<emphasis bold="true" box="[1017,1036,794,828]" pageId="22" pageNumber="389">it</emphasis>
|
|
might be juvenile. Examination of the cranial fragments and X-ray of the feet soon convinced us, however, that it is adult. Even older subadult
|
|
<taxonomicName box="[570,736,882,916]" class="Mammalia" family="Tachyglossidae" genus="Zaglossus" kingdom="Animalia" order="Monotremata" pageId="22" pageNumber="389" phylum="Chordata" rank="genus">
|
|
<emphasis box="[570,736,882,916]" italics="true" pageId="22" pageNumber="389">Zaglossus</emphasis>
|
|
</taxonomicName>
|
|
<emphasis bold="true" box="[756,766,882,916]" pageId="22" pageNumber="389">(</emphasis>
|
|
such as AM M5993
|
|
<emphasis bold="true" box="[1140,1151,882,916]" pageId="22" pageNumber="389">(</emphasis>
|
|
<taxonomicName box="[1150,1457,882,916]" class="Mammalia" family="Tachyglossidae" genus="Zaglossus" kingdom="Animalia" order="Monotremata" pageId="22" pageNumber="389" phylum="Chordata" rank="subSpecies" species="hartoni" subSpecies="bartoni">
|
|
<emphasis box="[1150,1175,882,916]" italics="true" pageId="22" pageNumber="389">Z</emphasis>
|
|
.
|
|
<emphasis box="[1196,1457,882,916]" italics="true" pageId="22" pageNumber="389">hartoni bartoni</emphasis>
|
|
</taxonomicName>
|
|
<emphasis bold="true" box="[1458,1478,882,916]" pageId="22" pageNumber="389">),</emphasis>
|
|
which has a dentary length of 120, retains open sutures and a juvenile-like dentary structure. Furthermore, juveniles typically have very long fur. This is in contrast to the short, dense fur of the holotype.
|
|
</paragraph>
|
|
<paragraph blockId="22.[96,1602,288,1488]" pageId="22" pageNumber="389">
|
|
The discovery of
|
|
<taxonomicName box="[510,927,1057,1091]" class="Mammalia" family="Tachyglossidae" genus="Zaglossus" kingdom="Animalia" order="Monotremata" pageId="22" pageNumber="389" phylum="Chordata" rank="species" species="attenboroughi">
|
|
<emphasis box="[510,927,1057,1091]" italics="true" pageId="22" pageNumber="389">Zaglossus attenboroughi</emphasis>
|
|
</taxonomicName>
|
|
in the higher peaks of the Cyclops Mountains was unexpected. The nearest populations of
|
|
<taxonomicName box="[1062,1228,1101,1135]" class="Mammalia" family="Tachyglossidae" genus="Zaglossus" kingdom="Animalia" order="Monotremata" pageId="22" pageNumber="389" phylum="Chordata" rank="genus">
|
|
<emphasis box="[1062,1228,1101,1135]" italics="true" pageId="22" pageNumber="389">Zaglossus</emphasis>
|
|
</taxonomicName>
|
|
<emphasis bold="true" box="[1248,1259,1101,1135]" pageId="22" pageNumber="389">(</emphasis>
|
|
<emphasis box="[1259,1284,1101,1135]" italics="true" pageId="22" pageNumber="389">Z</emphasis>
|
|
.
|
|
<emphasis box="[1305,1506,1101,1135]" italics="true" pageId="22" pageNumber="389">b. clianiomli</emphasis>
|
|
<emphasis bold="true" box="[1522,1532,1101,1135]" pageId="22" pageNumber="389">-</emphasis>
|
|
the largest of all
|
|
<emphasis box="[329,507,1145,1179]" italics="true" pageId="22" pageNumber="389">
|
|
<taxonomicName box="[329,495,1145,1179]" class="Mammalia" family="Tachyglossidae" genus="Zaglossus" kingdom="Animalia" order="Monotremata" pageId="22" pageNumber="389" phylum="Chordata" rank="genus">Zaglossus</taxonomicName>
|
|
<emphasis bold="true" box="[494,507,1145,1179]" italics="true" pageId="22" pageNumber="389">)</emphasis>
|
|
</emphasis>
|
|
occupy the mid-elevation forests above 1,300 metres in elevation on the northern slopes of the central cordillera, which lies some 200 km to the south.
|
|
<taxonomicName box="[226,391,1233,1267]" class="Mammalia" family="Tachyglossidae" genus="Zaglossus" kingdom="Animalia" order="Monotremata" pageId="22" pageNumber="389" phylum="Chordata" rank="genus">
|
|
<emphasis box="[226,391,1233,1267]" italics="true" pageId="22" pageNumber="389">Zaglossus</emphasis>
|
|
</taxonomicName>
|
|
is certainly absent from the North Coast Ranges, including the Bewani Mountains
|
|
<emphasis bold="true" box="[438,449,1277,1311]" pageId="22" pageNumber="389">(</emphasis>
|
|
which are no more than 50 km east of the Cyclops Mountains
|
|
<emphasis bold="true" box="[1540,1550,1277,1311]" pageId="22" pageNumber="389">)</emphasis>
|
|
as extensive survey work
|
|
<emphasis bold="true" box="[495,506,1321,1355]" pageId="22" pageNumber="389">(</emphasis>
|
|
by T.F.F. and others
|
|
<emphasis bold="true" box="[857,867,1321,1355]" pageId="22" pageNumber="389">)</emphasis>
|
|
has failed to reveal its presence there, and echidna-like animals are unknown to local hunters.
|
|
</paragraph>
|
|
<paragraph blockId="22.[96,1602,288,1488]" lastBlockId="23.[122,1600,280,451]" lastPageId="23" lastPageNumber="390" pageId="22" pageNumber="389">
|
|
The occurrence of
|
|
<taxonomicName box="[505,923,1409,1443]" class="Mammalia" family="Tachyglossidae" genus="Zaglossus" kingdom="Animalia" order="Monotremata" pageId="22" pageNumber="389" phylum="Chordata" rank="species" species="attenboroughi">
|
|
<emphasis box="[505,923,1409,1443]" italics="true" pageId="22" pageNumber="389">Zaglossus attenboroughi</emphasis>
|
|
</taxonomicName>
|
|
in the Cyclops Mountains is congruent with a number of endemics which are restricted to one or more peaks in the North Coast Ranges. These include
|
|
<emphasis box="[641,1270,285,319]" italics="true" pageId="23" pageNumber="390">
|
|
<taxonomicName box="[641,790,285,319]" class="Mammalia" family="Petauridae" genus="Petaurus" kingdom="Animalia" order="Diprotodontia" pageId="23" pageNumber="390" phylum="Chordata" rank="genus">Petaurus</taxonomicName>
|
|
abid'u
|
|
<taxonomicName box="[921,1270,285,319]" class="Mammalia" family="Macropodidae" genus="Dendrolagus" kingdom="Animalia" order="Diprotodontia" pageId="23" pageNumber="390" phylum="Chordata" rank="species" species="scottae">Dendrolagus scottae</taxonomicName>
|
|
</emphasis>
|
|
and
|
|
<emphasis italics="true" pageId="23" pageNumber="390">
|
|
<taxonomicName class="Mammalia" family="Muridae" genus="Paraleptomys" kingdom="Animalia" order="Rodentia" pageId="23" pageNumber="390" phylum="Chordata" rank="species" species="rufilatus">Paraleptomys rufilatus</taxonomicName>
|
|
.
|
|
</emphasis>
|
|
There are no other
|
|
<emphasis box="[624,740,328,362]" italics="true" pageId="23" pageNumber="390">species</emphasis>
|
|
of mammals, however, which are endemic solely to the Cyclops Mountains. It seems possible that
|
|
<taxonomicName box="[988,1265,372,406]" class="Mammalia" family="Tachyglossidae" genus="Zaglossus" kingdom="Animalia" order="Monotremata" pageId="23" pageNumber="390" phylum="Chordata" rank="species" species="attenboroughi">
|
|
<emphasis box="[988,1012,372,406]" italics="true" pageId="23" pageNumber="390">Z</emphasis>
|
|
.
|
|
<emphasis box="[1032,1265,372,406]" italics="true" pageId="23" pageNumber="390">attenboroughi</emphasis>
|
|
</taxonomicName>
|
|
was possibly once widespread in the North Coast Ranges, and has found its last refuge in the Cyclops.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |