treatments-xml/data/E8/10/B9/E810B924E00E5676A21F5D5ED71C4EEC.xml

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<mods:title id="FE5AD08FA0891E8B1775061087102BAF">Molecular phylogeny of the riffle beetle genus Hexanchorus revealed a presence of a new genus (Coleoptera: Elmidae)</mods:title>
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<mods:namePart id="841BD1A019E65F60697EBADAF830FE86">Linsky, Marek</mods:namePart>
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<mods:namePart id="9592CAE6A57A6BE101F2E9708B70EF98">Ciamporova-Zatovicova, Zuzana</mods:namePart>
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<mods:namePart id="F89D962E3AE3AFAD0F070B0FD6E81E46">Lassova, Kristina</mods:namePart>
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<paragraph id="E3E3EDE80FF2A371525321B84F1EB301" pageId="0" pageNumber="575">
3.2.6. 
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<taxonomicNameLabel id="E2B6268E7AB434331EA3050E849E4AD6" pageId="0" pageNumber="575">sp. nov.</taxonomicNameLabel>
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<paragraph id="89956F60050C4D00C8A929C595C34215" pageId="0" pageNumber="575">
<figureCitation id="E6AFADDD014142528BDE8AB965361791" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal habiti of Rumilara gen. nov. species: a R. obscura sp. nov. male holotype; b R. suppressa sp. nov. male holotype; c R. riberai sp. nov. male holotype; d R. paterna sp. nov. male holotype; e R. leleupi comb. nov. male holotype. Scale: 0.5 mm." figureDoi="10.3897/asp.80.e84013.figure2" httpUri="https://binary.pensoft.net/fig/770425" pageId="0" pageNumber="575">Figs 2d</figureCitation>
<figureCitation id="4CF55757946BA1D28E0253F8E6CDFB93" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Lateral habiti of Rumilara gen. nov. species. a R. obscura sp. nov. male holotype; b R. suppressa sp. nov. male holotype; c R. riberai sp. nov. male holotype; d R. paterna sp. nov. male holotype; e R. leleupi comb. nov. male holotype. Scale: 0.5 mm." figureDoi="10.3897/asp.80.e84013.figure3" httpUri="https://binary.pensoft.net/fig/770426" pageId="0" pageNumber="575">, 3d</figureCitation>
<figureCitation id="FFC404D3269AAE95FBB004CF73C3D934" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Antennae of Rumilara gen. nov. and Hexanchorus: a R. obscura sp. nov.; b R. suppressa sp. nov.; c R. riberai sp. nov.; d R. paterna sp. nov.; e R. leleupi comb. nov.; f H. mcdiarmidi; g H. virilis; h H. shannoni; i H. angeli; j H. rostratus; k H. cordillierae; l H. onorei; m H. tarsalis; n H. usitatus; o H. gracilipes; p H. caraibus; q H. crinitus; r H. dimorphus; s H. emarginatus; t H. browni. (a, k based on dissected antennae; b-j, l-r based on photographs, not to scale, setae omitted)." figureDoi="10.3897/asp.80.e84013.figure5" httpUri="https://binary.pensoft.net/fig/770428" pageId="0" pageNumber="575">, 5d</figureCitation>
<figureCitation id="58DD20EDF7FDAE204AE23859A79A50E7" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Pronota of Rumilara gen. nov. species: a R. obscura sp. nov. male holotype; b R. suppressa sp. nov. male holotype; c R. riberai sp. nov. male holotype; d R. paterna sp. nov. male holotype; e R. leleupi comb. nov. male holotype. Scale: 0.2 mm." figureDoi="10.3897/asp.80.e84013.figure6" httpUri="https://binary.pensoft.net/fig/770429" pageId="0" pageNumber="575">, 6d</figureCitation>
<figureCitation id="41111A9E6836891D3E017A1311EDD10A" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Ventral habiti of Rumilara gen. nov. species: a R. obscura sp. nov. male paratype; b R. suppressa sp. nov. male paratype; c R. riberai sp. nov. female paratype; d R. paterna sp. nov. paratype; e R. leleupi comb. nov. male holotype. Scale: 0.5 mm." figureDoi="10.3897/asp.80.e84013.figure7" httpUri="https://binary.pensoft.net/fig/770430" pageId="0" pageNumber="575">, 7d</figureCitation>
<figureCitation id="07D88E00381E1776A0EA1D306C795F08" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Abdomens of Rumilara gen. nov. species: a R. obscura sp. nov. male holotype; b R. suppressa sp. nov. male holotype; c R. riberai sp. nov. male holotype; d R. paterna sp. nov. male holotype; e R. leleupi comb. nov. male holotype. 0.2 mm." figureDoi="10.3897/asp.80.e84013.figure8" httpUri="https://binary.pensoft.net/fig/770431" pageId="0" pageNumber="575">, 8d</figureCitation>
<figureCitation id="20F0F015879FB49BB0E390454756C1A2" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Larvae of Rumilara gen. nov. species: a R. obscura sp. nov., dorsal; b R. suppressa sp. nov., dorsal; c R. riberai sp. nov., dorsal; d R. paterna sp. nov., dorsal; e R. obscura sp. nov., ventral; f R. suppressa sp. nov., ventral. Scale: 1 mm." figureDoi="10.3897/asp.80.e84013.figure9" httpUri="https://binary.pensoft.net/fig/770432" pageId="0" pageNumber="575">, 9d</figureCitation>
<figureCitation id="B1745BC4703D4BEB055E4B875992096A" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Rumilara paterna sp. nov.: a aedeagus, left lateral view; b aedeagus, ventral view; c aedeagus, right lateral view; d male sternite VIII. Scale: 0.1 mm." figureDoi="10.3897/asp.80.e84013.figure14" httpUri="https://binary.pensoft.net/fig/770437" pageId="0" pageNumber="575">, 14</figureCitation>
</paragraph>
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<paragraph id="CF2E9C7A2556EF7EA0FEEDAAB0F4C4E7" pageId="0" pageNumber="575">Material examined.</paragraph>
<paragraph id="3F7C61934B70AD7621DF4FC56C99BA09" pageId="0" pageNumber="575">
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<emphasis id="3DAF5DFC4C3438F0B6418D8BE7978780" italics="true" pageId="0" pageNumber="575">Type material</emphasis>
: 
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<typeStatus id="54D64D75F8B25B74E65B8DBF5BA2C32F">Holotype</typeStatus>
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(PUCE) &quot; 
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, 
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, 
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env.,, 
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, 
<geoCoordinate id="788B43D3A5350866E1761387ACEEB49B" degrees="78" direction="west" minutes="32" orientation="longitude" precision="1" seconds="10.5" value="-78.53625">78°32′10.5″ W</geoCoordinate>
, 
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<quantity id="B4CAC9E78476C0EAFF3CBBFAF052BF85" metricMagnitude="3" metricUnit="m" metricValue="1.299" unit="m" value="1299.0">1299m</quantity>
a.s.l.
</elevation>
, 
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, stream ca 
<quantity id="9CD710910E5341751F2DC419E767AF09" metricMagnitude="0" metricUnit="m" metricValue="5.0" unit="m" value="5.0">
<elevation id="18EFC457C07A9D7856CA5FD96AD4A27B" metricMagnitude="0" metricUnit="m" metricValue="5.0" unit="m" value="5.0">5m</elevation>
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wide, with gravel, boulders, leaf debris, submerged wood, 
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Jr &amp; 
<normalizedToken id="B814F68CD8CB4831FF5DEA67597205D9" originalValue="Čiamporová-Zaťovičová">Ciamporova-Zatovicova</normalizedToken>
lgt.&quot;
</materialsCitation>
</emphasis>
; 
<materialsCitation id="657EA62ED92E2B806EF06F5814B6CB5D" ID-GBIF-Occurrence="3968369309" collectingDate="2013-08-17" collectingDateMax="2013-08-25" collectingDateMin="2013-08-17" collectorName="Ciampor Jr, Ciamporova-Zatovicova" country="Ecuador" elevation="698" latitude="-3.0874166" location="Cascada la Encanada" longLatPrecision="1" longitude="-78.41" municipality="Santa Rosa de Mamanguy" specimenCount="2" specimenCount-larva="1" stateProvince="Morona-Santiago prov." typeStatus="Paratype">
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<typeStatus id="6E0661FF8BD8A15F7EF1C3A2B8F7D17E">Paratype</typeStatus>
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(CCB) &quot; 
<collectingCountry id="0237219803D77F235E0457451247DE00" name="Ecuador">Ecuador</collectingCountry>
, 
<collectingRegion id="2499BE26552794F1D244CF028638607F" country="Ecuador" name="Morona-Santiago">Morona-Santiago prov.</collectingRegion>
, 
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env., 
<location id="DAC632C7B24553185168171B44B2C698" LSID="urn:lsid:plazi:treatment:E810B924E00E5676A21F5D5ED71C4EEC:DAC632C7B24553185168171B44B2C698" country="Ecuador" latitude="-3.0874166" longLatPrecision="1" longitude="-78.41" municipality="Santa Rosa de Mamanguy" name="Cascada la Encanada" stateProvince="Morona-Santiago prov.">
Cascada la 
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, 
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, 
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, 
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<quantity id="53365CE208FE45828C1125D98094DEA3" metricMagnitude="2" metricUnit="m" metricValue="6.9799999999999995" unit="m" value="698.0">698m</quantity>
a.s.l.
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, 
<collectingDate id="0756B776279A62D37CB489D072BEB7BE" value="2013-08-25">25.8.2013</collectingDate>
, stream ca 
<quantity id="29A0C093D4A2C4E0B5339308DDD8FD22" metricMagnitude="0" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" unit="m" value="2.5" valueMax="3.0" valueMin="2.0">
<elevation id="FBF8E2BB479F3B58CB6CBD370DF73137" metricMagnitude="0" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" unit="m" value="2.5" valueMax="3.0" valueMin="2.0">2-3m</elevation>
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wide under waterfall, with clay, gravel, stones, submerged wood, 
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Jr
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&amp; 
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lgt.&quot; 
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<typeStatus id="2E5561885A2A20AA430912546E37DAC1" isOtherTypeStatus="true">Other material</typeStatus>
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: 
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&quot; 
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, 
<collectingRegion id="F95A5F315D01301FC5C5EDF449D7F66B" country="Ecuador" name="Napo">Napo prov.</collectingRegion>
, road to Coca, 
<geoCoordinate id="C98ED1A4865AF3BCBF9F1E4014FD5302" degrees="00" direction="south" minutes="43" orientation="latitude" precision="1" seconds="39.6" value="-0.7276666">00°43′39.6″ S</geoCoordinate>
, 
<geoCoordinate id="C2DBD4FF619250C0B819F2462E820491" degrees="77" direction="west" minutes="45" orientation="longitude" precision="1" seconds="56.1" value="-77.76559">77°45′56.1″ W</geoCoordinate>
, 
<elevation id="85C02937C0740B0BBDFA1BAAEA7F6C84" metricMagnitude="3" metricUnit="m" metricValue="1.129" unit="m" value="1129.0">
<quantity id="CED33CEF4FE8707C32427E0791164D72" metricMagnitude="3" metricUnit="m" metricValue="1.129" unit="m" value="1129.0">1129m</quantity>
a.s.l.
</elevation>
, 
<collectingDate id="EC1950202E16146993F49F4D04FB460F" value="2013-08-17">17.8.2013</collectingDate>
, small waterfall, 
<normalizedToken id="D62D0F8B705D5A55F5F3F05877DF9E95" originalValue="Čiampor">Ciampor</normalizedToken>
Jr &amp; 
<normalizedToken id="C1EA7EC2D35EDC005D5F0A7E62219EE4" originalValue="Čiamporová-Zaťovičová">Ciamporova-Zatovicova</normalizedToken>
lgt.&quot;
</materialsCitation>
; 
<materialsCitation id="B4874D3D7515BEE89828CD8DCDE7D5B7" ID-GBIF-Occurrence="3968369319" collectingDate="2013-08-24" country="Ecuador" elevation="979" latitude="-3.0491667" location="Rio Crusado" longLatPrecision="1" longitude="-78.50089" specimenCount="2" specimenCount-larva="2" stateProvince="Morona-Santiago prov." typeStatus="paratype">
<specimenCount id="172D6465E1ECC12719A150EF94767564" type="larva">2 larvae</specimenCount>
&quot; 
<collectingCountry id="E88E649AB34A4713E4E0C90563471FF2" name="Ecuador">Ecuador</collectingCountry>
, 
<collectingRegion id="405133BCDEA5BC53D559F4D8CD5ADC93" country="Ecuador" name="Morona-Santiago">Morona-Santiago prov.</collectingRegion>
, tributary of 
<location id="95B3477B895E4CE199A4B4EDFDA86623" LSID="urn:lsid:plazi:treatment:E810B924E00E5676A21F5D5ED71C4EEC:95B3477B895E4CE199A4B4EDFDA86623" country="Ecuador" latitude="-3.0491667" longLatPrecision="1" longitude="-78.50089" name="Rio Crusado" stateProvince="Morona-Santiago prov.">
<normalizedToken id="6124B09F9BCBBB414342E6D6DC8BC16C" originalValue="Río">Rio</normalizedToken>
Crusado
</location>
, 
<geoCoordinate id="095EA6B0072694C696099B98272AD849" degrees="03" direction="south" minutes="02" orientation="latitude" precision="1" seconds="57.0" value="-3.0491667">03°02′57.0″ S</geoCoordinate>
, 
<geoCoordinate id="98A602A393CF7972BFFE4601A671648E" degrees="78" direction="west" minutes="30" orientation="longitude" precision="1" seconds="03.2" value="-78.50089">78°30′03.2″ W</geoCoordinate>
, 
<elevation id="0F3C53EAED2CE95B4510D45E9854C44C" metricMagnitude="2" metricUnit="m" metricValue="9.79" unit="m" value="979.0">
<quantity id="E88F5A6C51F710B0E98BB6AFDAC7EA01" metricMagnitude="2" metricUnit="m" metricValue="9.79" unit="m" value="979.0">979m</quantity>
a.s.l.
</elevation>
, 
<collectingDate id="35185143FFE6B3735D80FC8F29D20F9C" value="2013-08-24">24.8.2013</collectingDate>
, stream ca 
<quantity id="F51036E752A0525A7E7A1AAD4E014828" metricMagnitude="0" metricUnit="m" metricValue="2.0" unit="m" value="2.0">
<elevation id="767348CE032043F3348C56310BFA5590" metricMagnitude="0" metricUnit="m" metricValue="2.0" unit="m" value="2.0">2m</elevation>
</quantity>
wide, in forest, with gravel, stones, leaf debris, submerged wood with moss, 
<normalizedToken id="B4D5264DB8DC086E7912255C5DF20160" originalValue="Čiampor">Ciampor</normalizedToken>
Jr &amp; 
<normalizedToken id="AECA75E07EF5EEC008505287C408AF06" originalValue="Čiamporová-Zaťovičová">Ciamporova-Zatovicova</normalizedToken>
lgt.&quot;
</materialsCitation>
; 
<materialsCitation id="543BD7A5E2B42AC9EEFECE601A106282" ID-GBIF-Occurrence="3968369305" collectingDate="2013-08-25" country="Ecuador" elevation="698" latitude="-3.0874166" location="Cascada la Encanada" longLatPrecision="1" longitude="-78.41" municipality="Santa Rosa de Mamanguy" specimenCount="1" specimenCount-larva="1" stateProvince="Morona-Santiago prov." typeStatus="paratype">
<specimenCount id="91B6F2E2276E432805CED54184046065" type="larva">1 larva</specimenCount>
&quot; 
<collectingCountry id="F58D182A89855B2F00BA6E5C21E51D2C" name="Ecuador">Ecuador</collectingCountry>
, 
<collectingRegion id="93044082D2D66A76CEDC1C99E6CA98BB" country="Ecuador" name="Morona-Santiago">Morona-Santiago prov.</collectingRegion>
, 
<collectingMunicipality id="9F48A1074998C6D313F8F07894276C8B">Santa Rosa de Mamanguy</collectingMunicipality>
env., 
<location id="FA47464094779A3B31C10291A9829D60" LSID="urn:lsid:plazi:treatment:E810B924E00E5676A21F5D5ED71C4EEC:FA47464094779A3B31C10291A9829D60" country="Ecuador" latitude="-3.0874166" longLatPrecision="1" longitude="-78.41" municipality="Santa Rosa de Mamanguy" name="Cascada la Encanada" stateProvince="Morona-Santiago prov.">
Cascada la 
<normalizedToken id="61D35A7CD3A8AE09B40A7BFCBD7D4F06" originalValue="Encañada">Encanada</normalizedToken>
</location>
, 
<geoCoordinate id="E978F4CF00ED0010F62EBEAFF3C7FE1D" degrees="03" direction="south" minutes="05" orientation="latitude" precision="1" seconds="14.7" value="-3.0874166">03°05′14.7″ S</geoCoordinate>
, 
<geoCoordinate id="558DB56E90976B7EC787B9C45A65DFA4" degrees="78" direction="west" minutes="24" orientation="longitude" precision="1" seconds="36.0" value="-78.41">78°24′36.0″ W</geoCoordinate>
, 
<elevation id="28894FA35E4D3671F3D4797B215EE177" metricMagnitude="2" metricUnit="m" metricValue="6.9799999999999995" unit="m" value="698.0">
<quantity id="0E653877E559FBDE46BB79BA2D524AE9" metricMagnitude="2" metricUnit="m" metricValue="6.9799999999999995" unit="m" value="698.0">698m</quantity>
a.s.l.
</elevation>
, 
<collectingDate id="0152A1870EC3D8CAC689E2A2EBAAABB3" value="2013-08-25">25.8.2013</collectingDate>
, stream ca 
<quantity id="AFDAA158D31F1E3371EBFAA0F0689C22" metricMagnitude="0" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" unit="m" value="2.5" valueMax="3.0" valueMin="2.0">
<elevation id="FEF4B05134CC63780747393D0029CD79" metricMagnitude="0" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" unit="m" value="2.5" valueMax="3.0" valueMin="2.0">2-3m</elevation>
</quantity>
wide under waterfall, with clay, gravel, stones, submerged wood, 
<normalizedToken id="6936FA66F20B3A5EA86160DF7D093548" originalValue="Čiampor">Ciampor</normalizedToken>
Jr &amp; 
<normalizedToken id="90CF2685DC06EBEEC3C0D8A09072C567" originalValue="Čiamporová-Zaťovičová">Ciamporova-Zatovicova</normalizedToken>
lgt.&quot;
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="BF10FE9382EF11BCB9561165D5C4CD96" pageId="0" pageNumber="575" type="adult diagnosis">
<paragraph id="E862153193D705A2CDEF24EC506E6C26" pageId="0" pageNumber="575">Adult diagnosis.</paragraph>
<paragraph id="842070DE4C7DA6249A9851A29587418C" pageId="0" pageNumber="575">The species can be distinguished by a combination of following characters: 1) head and clypeus fully clothed by pubescence; 2) femora fully pubescent; 3) elytra with dense pubescence, not arranged in stripes; 4) aedeagus with rod-like sclerite; 5) parameres moderately asymmetrical.</paragraph>
</subSubSection>
<subSubSection id="AE8212279178B457BD1F3981082A2247" pageId="0" pageNumber="575" type="description">
<paragraph id="A230C589B1C2ED643DAA90EF56D9F5BE" pageId="0" pageNumber="575">Adult diagnostic description.</paragraph>
<paragraph id="F21F3CAD8A2A0395161B7BCFEC33C460" pageId="0" pageNumber="575">
Male. 
<emphasis id="49A0CC000D756AE895A212D04486CDE0" bold="true" italics="true" pageId="0" pageNumber="575">Colour</emphasis>
: Body (Figs 
<figureCitation id="9B2817F411DC97ABFD32240FBF41AD89" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal habiti of Rumilara gen. nov. species: a R. obscura sp. nov. male holotype; b R. suppressa sp. nov. male holotype; c R. riberai sp. nov. male holotype; d R. paterna sp. nov. male holotype; e R. leleupi comb. nov. male holotype. Scale: 0.5 mm." figureDoi="10.3897/asp.80.e84013.figure2" httpUri="https://binary.pensoft.net/fig/770425" pageId="0" pageNumber="575">2d</figureCitation>
, 
<figureCitation id="85B6EDB5CBC39E3638DC6511C0B85125" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Lateral habiti of Rumilara gen. nov. species. a R. obscura sp. nov. male holotype; b R. suppressa sp. nov. male holotype; c R. riberai sp. nov. male holotype; d R. paterna sp. nov. male holotype; e R. leleupi comb. nov. male holotype. Scale: 0.5 mm." figureDoi="10.3897/asp.80.e84013.figure3" httpUri="https://binary.pensoft.net/fig/770426" pageId="0" pageNumber="575">3d</figureCitation>
) dark brown; coxae, trochanters, femora, tibiae, and tarsi brown; tarsal claws pale brown with reddish tincture. 
<emphasis id="6CCE9D1AE81378AFD93A53C7D7991854" bold="true" italics="true" pageId="0" pageNumber="575">Pubescence</emphasis>
: Head and clypeus fully clothed with short, prone pubescence, most densely set behind eyes, and with numerous slightly longer, curved, semi-erect, hair-like setae. Pronotum (Fig. 
<figureCitation id="6942065DF0E4F973490F3A0B6EC4523D" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Pronota of Rumilara gen. nov. species: a R. obscura sp. nov. male holotype; b R. suppressa sp. nov. male holotype; c R. riberai sp. nov. male holotype; d R. paterna sp. nov. male holotype; e R. leleupi comb. nov. male holotype. Scale: 0.2 mm." figureDoi="10.3897/asp.80.e84013.figure6" httpUri="https://binary.pensoft.net/fig/770429" pageId="0" pageNumber="575">6d</figureCitation>
) covered by a short, moderately dense, prone pubescence, and with moderately long, curved, semi-erect, hair-like setae. Elytra with densely arranged, short, recumbent pubescence, and numerous moderately long, curved, semi-erect, hair-like setae. Femora, pro- and metatibiae densely covered with moderately long, recumbent, hair-like setae; mesotibiae with pubescence only at extreme base. Metaventrite densely clothed with short, prone pubescence. 
<emphasis id="3A61B9B16E7B9EAC95F84EB576448D12" bold="true" italics="true" pageId="0" pageNumber="575">Surface</emphasis>
: Head and clypeus sparsely microreticulate, concealed by pubescence. Pronotum densely punctate, partly concealed by pubescence. Elytra with rows of punctures shallowly impressed; intervals nearly flat. Prosternal process (Fig. 
<figureCitation id="5EDE2900FFB03F8237DBCAFE92840C53" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Ventral habiti of Rumilara gen. nov. species: a R. obscura sp. nov. male paratype; b R. suppressa sp. nov. male paratype; c R. riberai sp. nov. female paratype; d R. paterna sp. nov. paratype; e R. leleupi comb. nov. male holotype. Scale: 0.5 mm." figureDoi="10.3897/asp.80.e84013.figure7" httpUri="https://binary.pensoft.net/fig/770430" pageId="0" pageNumber="575">7d</figureCitation>
) subparallel between procoxae, then subtriangular with rounded apex. 
<emphasis id="523D3780CC2F0B9C6F6206DB2E1E515D" bold="true" italics="true" pageId="0" pageNumber="575">Aedeagus</emphasis>
(Figs 
<figureCitation id="EC5D8D11D8884EDE5F3E995C76983B93" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Rumilara paterna sp. nov.: a aedeagus, left lateral view; b aedeagus, ventral view; c aedeagus, right lateral view; d male sternite VIII. Scale: 0.1 mm." figureDoi="10.3897/asp.80.e84013.figure14" httpUri="https://binary.pensoft.net/fig/770437" pageId="0" pageNumber="575">14a-c</figureCitation>
) with long, dorsal, rod-like sclerite on penis; parameres asymmetrical, left one slightly longer. - Female unknown.
</paragraph>
</subSubSection>
<subSubSection id="6DDEC71D5608217230629AAED97B6270" pageId="0" pageNumber="575" type="description">
<paragraph id="9474402DC7FFB6ACE61D37289FD61181" pageId="0" pageNumber="575">Measurements.</paragraph>
<paragraph id="4BA5225CB452D1AE18C81DA0019CB348" pageId="0" pageNumber="575">♂ - CL: 2.67 mm; PL: 0.61 mm; PW: 0.75 mm; EL: 2.07 mm; EW: 0.83 mm.</paragraph>
</subSubSection>
<subSubSection id="AF434FC603F13E85397E0C8FC7091E01" pageId="0" pageNumber="575" type="variation">
<paragraph id="4C1C8587034A0DC6BA97665CC2B7A2A8" pageId="0" pageNumber="575">Variation.</paragraph>
<paragraph id="360F942A4BBA7389A2C8CD50261CAC26" pageId="0" pageNumber="575">No significant differences were observed between the two known specimens.</paragraph>
</subSubSection>
<subSubSection id="23BAEB7EFB347F5553A3C12EEF528D68" pageId="0" pageNumber="575" type="etymology">
<paragraph id="DB33D1E64C98270ADB86B6B7F04F657B" pageId="0" pageNumber="575">Etymology.</paragraph>
<paragraph id="7E6BC18BB873AAE76250279218CE1100" pageId="0" pageNumber="575">
From Latin 
<emphasis id="B123901343272D261F4BA769BD9DF880" bold="true" pageId="0" pageNumber="575">paternus</emphasis>
(of or connected with 
<normalizedToken id="3464C1874C6992BB8063E7B373897867" originalValue="one’s">one's</normalizedToken>
forefathers, ancestral) due to its similarity with the genus 
<taxonomicName id="D8DF53003377305EC84EFDD4E1FBA071" authorityName="Sharp" authorityYear="1882" class="Insecta" family="Elmidae" genus="Hexanchorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hexanchorus" order="Coleoptera" pageId="0" pageNumber="575" phylum="Arthropoda" rank="genus">
<emphasis id="25B037BC82AF73EBCE33212E911A2B86" italics="true" pageId="0" pageNumber="575">Hexanchorus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="FCE51C4284A6E3398BBC2E3F4D64674D" pageId="0" pageNumber="575" type="larva">
<paragraph id="6B47CFEFA088220ECF184C714034A247" pageId="0" pageNumber="575">Larva</paragraph>
<paragraph id="9D88512E0D2118E6AABB4180D1C53B22" pageId="0" pageNumber="575">
(Fig. 
<figureCitation id="40E5ED89D13BB793A5C167179B91E4A1" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Larvae of Rumilara gen. nov. species: a R. obscura sp. nov., dorsal; b R. suppressa sp. nov., dorsal; c R. riberai sp. nov., dorsal; d R. paterna sp. nov., dorsal; e R. obscura sp. nov., ventral; f R. suppressa sp. nov., ventral. Scale: 1 mm." figureDoi="10.3897/asp.80.e84013.figure9" httpUri="https://binary.pensoft.net/fig/770432" pageId="0" pageNumber="575">9d</figureCitation>
) with length 3.48-3.55 mm, greatest width 0.99-1.10 mm. Pleurites on abdominal segments 1-5. Abdominal segments 1-7 explanate. Surface around the frontal pair of pronotal gibbosities paler than the rest of pronotum. Tergum of abdominal segments 1-7 with a pair of distinct, longitudinally arranged small gibbosities.
</paragraph>
<caption id="F897E66BBFEE9EE5079EC3EA7792B388" doi="10.3897/asp.80.e84013.figure16" httpUri="https://binary.pensoft.net/fig/770439" pageId="0" pageNumber="575" start="Figure 16" startId="F16">
<paragraph id="73CD68ABDC6A8DE7AD24AD59EBAF2FA2" pageId="0" pageNumber="575">
<emphasis id="6FAB01764F7969B77AEE37716AE884F3" bold="true" pageId="0" pageNumber="575">Figure 16.</emphasis>
Sampling sites of 
<taxonomicName id="16C741C03CB2C27EE94ED3FC97CA4CC6" authorityName="Linský &amp; Čiamporová-Zaťovičová &amp; Laššová &amp; Čiampor Jr" authorityYear="2022" class="Insecta" family="Elmidae" genus="Rumilara" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rumilara" order="Coleoptera" pageId="0" pageNumber="575" phylum="Arthropoda" rank="genus">
<emphasis id="7EDB93F40096B1A7F8D5BD201AEBA529" italics="true" pageId="0" pageNumber="575">Rumilara</emphasis>
</taxonomicName>
<emphasis id="48A9A130446AABC2108CB0FAB27AC6BC" bold="true" pageId="0" pageNumber="575">gen. nov.</emphasis>
in Ecuador: 
<emphasis id="894AE321EA44237B6E3C96A502AEB674" bold="true" pageId="0" pageNumber="575">a</emphasis>
Cotopaxi prov., Otonga env., 
<normalizedToken id="6C1ADAAF97EA489CD701731AEC4ECB8C" originalValue="Río">Rio</normalizedToken>
Esmeralda; 
<emphasis id="AD577E5393069920FF8CDF29E8EBCCFD" bold="true" pageId="0" pageNumber="575">b</emphasis>
Cotopaxi prov., Artos env., stream above confluence with 
<normalizedToken id="2A5BEC26F32A39A47D38226785AF635C" originalValue="Río">Rio</normalizedToken>
Esmeralda; 
<emphasis id="81F67ACAD1099C93296EDA8ECB076486" bold="true" pageId="0" pageNumber="575">c</emphasis>
Santo Domingo de los Tsachilas prov., Otongachi env., tributary of 
<normalizedToken id="8DEBBDE3B60385BB57918BD45AC15DAD" originalValue="Río">Rio</normalizedToken>
Toachi; 
<emphasis id="53732062A1A61E07A209498C20C5EA22" bold="true" pageId="0" pageNumber="575">d</emphasis>
Santo Domingo de los Tsachilas prov., thin water layer on rock, Otongachi env.; 
<emphasis id="DFA66F01F9E106DBF3B66D65DD8F7428" bold="true" pageId="0" pageNumber="575">e</emphasis>
Morona-Santiago prov., small stream near Indanza; 
<emphasis id="42C30679C26F5378EE85ECA2320AF15B" bold="true" pageId="0" pageNumber="575">f</emphasis>
Morona-Santiago prov., Santa Rosa de Mamanguy env., Cascada la 
<normalizedToken id="22530AC0650DB857C8EC90625CFDC602" originalValue="Encañada">Encanada</normalizedToken>
; 
<emphasis id="BCAAD1F80676E9DC5F1BFCA2226DF5B3" bold="true" pageId="0" pageNumber="575">g</emphasis>
fast-flowing stream in Napo prov.; 
<emphasis id="0833A527855D6CC9B0F07E3B6FA531DF" bold="true" pageId="0" pageNumber="575">h</emphasis>
Morona-Santiago prov., tributary of 
<normalizedToken id="5915870C5EBC8C986E05759B26F3B5C8" originalValue="Río">Rio</normalizedToken>
Tinajillas; 
<emphasis id="0111DB57AD8097C7193782043AAF72C3" bold="true" pageId="0" pageNumber="575">i</emphasis>
Morona-Santiago prov., tributary of 
<normalizedToken id="9F8BEC506C4C98B10462FFC146DE46C6" originalValue="Río">Rio</normalizedToken>
Tinajillas; (localities of 
<taxonomicName id="AA36154798D615A0DC461C4217EC856B" lsidName="R. obscura" pageId="0" pageNumber="575" rank="species" species="obscura">
<emphasis id="8044F36A42B2446D86FDF6EE9ABB9E98" italics="true" pageId="0" pageNumber="575">R. obscura</emphasis>
</taxonomicName>
- 
<emphasis id="64284867E48878D22B9FFA55AD099EE2" bold="true" pageId="0" pageNumber="575">a</emphasis>
, 
<emphasis id="4EA11D7ADC8F3BD7A1403DF0497EC6AD" bold="true" pageId="0" pageNumber="575">b</emphasis>
; 
<taxonomicName id="346ED46B142DE9058F58AB971EFF1B40" lsidName="R. riberai" pageId="0" pageNumber="575" rank="species" species="riberai">
<emphasis id="C35A697E12CAC54549658470A779BFBF" italics="true" pageId="0" pageNumber="575">R. riberai</emphasis>
</taxonomicName>
- 
<emphasis id="CB2D80176BE1C88F921F49D7731AB1BF" bold="true" pageId="0" pageNumber="575">c</emphasis>
, 
<emphasis id="F736D0854F724168A8B8963BFD05ED70" bold="true" pageId="0" pageNumber="575">d</emphasis>
; 
<taxonomicName id="7BCF0D1EFB779A715C97F6252E5E6919" lsidName="R. paterna" pageId="0" pageNumber="575" rank="species" species="paterna">
<emphasis id="97C74DE628F9BC05B753E37B69167E81" italics="true" pageId="0" pageNumber="575">R. paterna</emphasis>
</taxonomicName>
- 
<emphasis id="141474F815DC0259715A55B575E62BDD" bold="true" pageId="0" pageNumber="575">e</emphasis>
, 
<emphasis id="BE747C0508B9CB8BD45DFB201393C4BE" bold="true" pageId="0" pageNumber="575">f</emphasis>
; 
<taxonomicName id="09A94F7FF7225AB3D5537B62FDC22AFD" lsidName="R. suppressa" pageId="0" pageNumber="575" rank="species" species="suppressa">
<emphasis id="787A3BADC9A74BA149F62B1F8339E0E8" italics="true" pageId="0" pageNumber="575">R. suppressa</emphasis>
</taxonomicName>
- 
<emphasis id="12A61C7AF94BF3C9EF9F945E683F6E30" bold="true" pageId="0" pageNumber="575">g</emphasis>
- 
<emphasis id="7BDD35C998DD42F8BF0E058598217015" bold="true" pageId="0" pageNumber="575">i</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="1173FA79D0516390CB7076A49001B786" pageId="0" pageNumber="575" type="distribution">
<paragraph id="587D4393650E5385FB612D01F8B339E1" pageId="0" pageNumber="575">Distribution.</paragraph>
<paragraph id="EFFA15D2B78E0FDBC67C8B929E131EB5" pageId="0" pageNumber="575">
Known from one locality in Napo Province and three localities in Morona-Santiago Province (Figs 
<figureCitation id="F898ED37F5E60E27E9549950B3485EE1" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Sampling sites of Rumilara gen. nov. in Ecuador: a Cotopaxi prov., Otonga env., Rio Esmeralda; b Cotopaxi prov., Artos env., stream above confluence with Rio Esmeralda; c Santo Domingo de los Tsachilas prov., Otongachi env., tributary of Rio Toachi; d Santo Domingo de los Tsachilas prov., thin water layer on rock, Otongachi env.; e Morona-Santiago prov., small stream near Indanza; f Morona-Santiago prov., Santa Rosa de Mamanguy env., Cascada la Encanada; g fast-flowing stream in Napo prov.; h Morona-Santiago prov., tributary of Rio Tinajillas; i Morona-Santiago prov., tributary of Rio Tinajillas; (localities of R. obscura - a, b; R. riberai - c, d; R. paterna - e, f; R. suppressa - g - i)." figureDoi="10.3897/asp.80.e84013.figure16" httpUri="https://binary.pensoft.net/fig/770439" pageId="0" pageNumber="575">16e-f</figureCitation>
).
</paragraph>
<caption id="6E71482D9FD8852AC26207DF56807C29" doi="10.3897/asp.80.e84013.figure17" httpUri="https://binary.pensoft.net/fig/770440" pageId="0" pageNumber="575" start="Figure 17" startId="F17">
<paragraph id="0814D073D107E94C6D2FF8A3762392BD" pageId="0" pageNumber="575">
<emphasis id="6E51303248B9F130FB452121C7B4E18C" bold="true" pageId="0" pageNumber="575">Figure 17.</emphasis>
Altitudinal distribution of 
<taxonomicName id="4BAEEA0420410E8B9B6251E116B06BEE" authorityName="Sharp" authorityYear="1882" class="Insecta" family="Elmidae" genus="Hexanchorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hexanchorus" order="Coleoptera" pageId="0" pageNumber="575" phylum="Arthropoda" rank="genus">
<emphasis id="0D3E3AF41F1F9DFC2D1AB600CB701F1A" italics="true" pageId="0" pageNumber="575">Hexanchorus</emphasis>
</taxonomicName>
and 
<taxonomicName id="DE1E73AF82A56A6A965F2A8E4EAF6372" authorityName="Linský &amp; Čiamporová-Zaťovičová &amp; Laššová &amp; Čiampor Jr" authorityYear="2022" class="Insecta" family="Elmidae" genus="Rumilara" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rumilara" order="Coleoptera" pageId="0" pageNumber="575" phylum="Arthropoda" rank="genus">
<emphasis id="4ABAED9A7D1F7423FD630FA45BDBF6C4" italics="true" pageId="0" pageNumber="575">Rumilara</emphasis>
</taxonomicName>
<emphasis id="9B5FB6D65AB710D3D5683635240499FA" bold="true" pageId="0" pageNumber="575">gen. nov.</emphasis>
in Ecuador (with one record from Colombia); circles - 
<taxonomicName id="2F53833EB7D3F70DD1BBB571EE898426" authorityName="Sharp" authorityYear="1882" class="Insecta" family="Elmidae" genus="Hexanchorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hexanchorus" order="Coleoptera" pageId="0" pageNumber="575" phylum="Arthropoda" rank="genus">
<emphasis id="5F22C6F11FD441472D3D6D6C0554D1BD" italics="true" pageId="0" pageNumber="575">Hexanchorus</emphasis>
</taxonomicName>
, triangles - 
<taxonomicName id="7CFB026A4A6A8F96778046FE385B8C75" authorityName="Linský &amp; Čiamporová-Zaťovičová &amp; Laššová &amp; Čiampor Jr" authorityYear="2022" class="Insecta" family="Elmidae" genus="Rumilara" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rumilara" order="Coleoptera" pageId="0" pageNumber="575" phylum="Arthropoda" rank="genus">
<emphasis id="220D96BAC3BCE5AE4F3E8ECD866DE6E1" italics="true" pageId="0" pageNumber="575">Rumilara</emphasis>
</taxonomicName>
<emphasis id="E0017A2A30B0DC8026985E563B60EF0E" bold="true" pageId="0" pageNumber="575">gen. nov.</emphasis>
Hollow symbols represent material not examined in this study (Modified from Google Earth).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="ECFC789947B884AB1544F596AA5E0818" pageId="0" pageNumber="575" type="remarks">
<paragraph id="E4EC62DF1DA8E7A45D6AA29C7CF16BDB" pageId="0" pageNumber="575">Remarks.</paragraph>
<paragraph id="62B5C6CAAD7B416E80423314D7FCEDB5" pageId="0" pageNumber="575">Only two specimens are known, one of them of unidentified sex, with abdomen lacking last two segments.</paragraph>
</subSubSection>
</treatment>
</document>