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<document id="F521610D82C1F92604FB877305804FCA" ID-CLB-Dataset="9693" ID-DOI="10.5852/ejt.2020.721.1127" ID-GBIF-Dataset="0cd61b23-1089-4c1b-bfc0-487887abad84" ID-Zenodo-Dep="4122118" ID-ZooBank="89E62EF8-2E45-4C59-94B7-6A5603E8939B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1603474447689" checkinUser="valdenar" docAuthor="Telnov, Dmitry" docDate="2020" docId="1A3787F69D749232FDB9FD4BC9FD2B34" docLanguage="en" docName="ejt-721_telnov.pdf" docOrigin="European Journal of Taxonomy 721" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Maechidius seriepunctatus Moser 1920" docType="treatment" docVersion="10" lastPageNumber="106" masterDocId="E60EFF8E9D1C925BFF93FFBEC8562862" masterDocTitle="A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line" masterLastPageNumber="210" masterPageNumber="1" pageNumber="105" updateTime="1698846101371" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="E9AAA4A85AB8AF7A8D9DCDB84F40F0DB">A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line</mods:title>
</mods:titleInfo>
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<mods:namePart id="A8FEF298A9D2F316F2AE748AEDB8B0D0">Telnov, Dmitry</mods:namePart>
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<mods:title id="9631A4485BD4C9559B901F5D12EF2978">European Journal of Taxonomy</mods:title>
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<mods:date id="7E678E02955F609B9F41BF1CB9F39F36">2020</mods:date>
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<mods:number id="FC19C71BE07709E99A526527229E7D8F">2020-10-19</mods:number>
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<mods:number id="60F27A515B447686CF8B47071B9F3E69">721</mods:number>
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<mods:classification id="42A9DDC3578EE336E909C79016247551">journal article</mods:classification>
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<mods:identifier id="D8E93244A88340F8B616A42BA82D1760" type="DOI">10.5852/ejt.2020.721.1127</mods:identifier>
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<treatment id="1A3787F69D749232FDB9FD4BC9FD2B34" ID-DOI="http://doi.org/10.5281/zenodo.4344461" ID-GBIF-Taxon="168648126" ID-Zenodo-Dep="4344461" LSID="urn:lsid:plazi:treatment:1A3787F69D749232FDB9FD4BC9FD2B34" httpUri="http://treatment.plazi.org/id/1A3787F69D749232FDB9FD4BC9FD2B34" lastPageId="105" lastPageNumber="106" pageId="104" pageNumber="105">
<subSubSection id="DA84656B9D749233FDB9FD4BCC5C2B72" box="[554,1034,757,784]" pageId="104" pageNumber="105" type="nomenclature">
<paragraph id="922136E09D749233FDB9FD4BCC5C2B72" blockId="104.[554,1034,757,821]" box="[554,1034,757,784]" pageId="104" pageNumber="105">
<heading id="C969818C9D749233FDB9FD4BCC5C2B72" box="[554,1034,757,784]" centered="true" fontSize="11" level="2" pageId="104" pageNumber="105" reason="2">
<taxonomicName id="559E4D639D749233FDB9FD4BCC5C2B72" ID-CoL="3XFL9" authority="Moser, 1920" authorityName="Moser" authorityYear="1920" box="[554,1034,757,784]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="104" pageNumber="105" phylum="Arthropoda" rank="species" species="seriepunctatus">
<emphasis id="A0EAEAF29D749233FDB9FD4BCB392B6D" bold="true" box="[554,879,757,784]" italics="true" pageId="104" pageNumber="105">Maechidius seriepunctatus</emphasis>
<bibRefCitation id="F60F4B119D749233FCE5FD48CC5C2B72" author="Moser J." box="[886,1034,758,784]" pageId="104" pageNumber="105" pagination="3 - 28" refId="ref98398" refString="Moser J. 1920. Beitrag zur Kenntnis der Melolonthiden XI. Stettiner Entomologische Zeitung 81: 3 - 28." type="journal article" year="1920">Moser, 1920</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D749233FDADFCA4CBA32B57" box="[574,1013,794,821]" pageId="104" pageNumber="105" type="description">
<paragraph id="922136E09D749233FDADFCA4CBA32B57" blockId="104.[554,1034,757,821]" box="[574,1013,794,821]" pageId="104" pageNumber="105">
<figureCitation id="0AA52A659D749233FDADFCA4CAC12B56" box="[574,663,794,820]" captionStart="Figs 78–81" captionStartId="100.[189,241,1928,1954]" captionTargetBox="[341,1237,264,1897]" captionTargetId="figure-63@100.[336,1259,251,1916]" captionTargetPageId="100" captionText="Figs 78–81. Papuan Maechidius Macleay, 1819, habitus, dorsal view. 78. M. riedeli sp. nov., holotype, ♂ (SMNS). 79. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 80. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 81. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122163" httpUri="https://zenodo.org/record/4122163/files/figure.png" pageId="104" pageNumber="105">Figs 81</figureCitation>
,
<figureCitation id="0AA52A659D749233FD37FCA4CA852B56" box="[676,723,794,820]" captionStart="Figs 169–180" captionStartId="143.[189,241,1787,1813]" captionTargetBox="[359,1217,271,1750]" captionTargetId="figure-123@143.[359,1228,265,1755]" captionTargetPageId="143" captionText="Figs 169–180. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal view. 169. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 170. Same, paratype, ♂ (BMNH). 171. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 172. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 173. Same, ♂ (ZMHB). 174. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). 175. M. similis sp. nov., paratype, ♂ (BMNH). 176. Same, paratype, ♀ (BMNH). 177. M. skalei sp. nov., paratype, ♂ (NME). 178. M. sougb sp. nov., holotype, ♂ (NME). 179. M. speciosus (Frey, 1969) comb. nov., ♂ (NHMB). 180. Same, ♀ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122185" httpUri="https://zenodo.org/record/4122185/files/figure.png" pageId="104" pageNumber="105">174</figureCitation>
,
<figureCitation id="0AA52A659D749233FD4CFCA4CB582B57" box="[735,782,794,821]" captionStart="Figs 242–258" captionStartId="148.[189,241,1646,1672]" captionTargetBox="[394,1193,265,1617]" captionTargetId="figure-193@148.[394,1193,265,1626]" captionTargetPageId="148" captionText="Figs 242–258. Papuan and Wallacean Maechidius Macleay, 1819, pronotum, dorsal view. 242. M. paupianus Heller, 1910, lectotype, ♀ (SNSD). 243. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 244. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 245. M. peregrinus Lansberge, 1886, ♂ (DTC). 246. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 247. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 248. M. riedeli sp. nov., holotype, ♂ (SMNS). 249. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 250. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 251. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). 252. M. similis sp. nov., paratype, ♂ (BMNH). 253. M. skalei sp. nov., paratype, ♂ (NME). 254. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 255. M. sougb sp. nov., holotype, ♂ (NME). 256. M. speciosus (Frey, 1969) comb. nov., ♀ (BMNH). 257. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). 258. M. suwawa sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122196" httpUri="https://zenodo.org/record/4122196/files/figure.png" pageId="104" pageNumber="105">251</figureCitation>
,
<figureCitation id="0AA52A659D749233FC8AFCA4CB1E2B56" box="[793,840,794,820]" captionStart="Figs 278–285" captionStartId="151.[189,241,1697,1723]" captionTargetBox="[211,1375,265,1597]" captionTargetId="figure-104@151.[211,1376,265,1614]" captionTargetPageId="151" captionText="Figs 278–285. Papuan and Wallacean Maechidius Macleay, 1819, pronotal hypomeron, ventro-lateral view. 278. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 279. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 280. M. seriepunctatus Moser, 1920, lectotype, ♂ (SNSD). 281. M. speciosus (Frey, 1969) comb. nov., ♀ (BMNH). 282. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). 283. M. trivialis sp. nov., holotype, ♂ (BMNH). 284. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 285. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122202" httpUri="https://zenodo.org/record/4122202/files/figure.png" pageId="104" pageNumber="105">280</figureCitation>
,
<figureCitation id="0AA52A659D749233FCC7FCA4CBD52B56" box="[852,899,794,820]" captionStart="Figs 333–348" captionStartId="155.[189,241,1682,1708]" captionTargetBox="[394,1193,265,1641]" captionTargetId="figure-177@155.[394,1193,265,1655]" captionTargetPageId="155" captionText="Figs 333–348. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 333. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 334. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 335. M. peregrinus Lansberge, 1886, ♂ (DTC). 336. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 337. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 338. Same, enlarged (SMNS). 339. M. riedeli sp. nov., holotype, ♂ (SMNS). 340. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 341. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 342. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). 343. M. similis sp. nov., paratype, ♂ (BMNH). 344. M. skalei sp. nov., paratype, ♂ (NME). 345. M. sougb sp. nov., holotype, ♂ (NME). 346. M. speciosus (Frey, 1969) comb. nov., ♀(BMNH). 347. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 348. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122212" httpUri="https://zenodo.org/record/4122212/files/figure.png" pageId="104" pageNumber="105">342</figureCitation>
, 414, 522
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D749233FF2EFCE2C92F2C71" pageId="104" pageNumber="105" type="materials_examined">
<paragraph id="922136E09D749233FF2EFCE2C93A2B14" blockId="104.[189,364,860,886]" box="[189,364,860,886]" pageId="104" pageNumber="105">
<heading id="C969818C9D749233FF2EFCE2C93A2B14" bold="true" box="[189,364,860,886]" fontSize="11" level="3" pageId="104" pageNumber="105" reason="3">
<emphasis id="A0EAEAF29D749233FF2EFCE2C93A2B14" bold="true" box="[189,364,860,886]" pageId="104" pageNumber="105">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D749233FF79FC32CA3A2BC4" blockId="104.[189,1398,907,1043]" box="[234,620,907,934]" pageId="104" pageNumber="105">
<emphasis id="A0EAEAF29D749233FF79FC32C9302BC4" bold="true" box="[234,358,908,934]" pageId="104" pageNumber="105">
<typeStatus id="4D2588429D749233FF79FC32C9302BC4" box="[234,358,908,934]" pageId="104" pageNumber="105" type="lectotype">Lectotype</typeStatus>
</emphasis>
[herewith designated]
</paragraph>
<paragraph id="922136E09D749233FF2EFC0EC92F2C71" blockId="104.[189,1398,907,1043]" pageId="104" pageNumber="105">
<materialsCitation id="22F63CBD9D749233FF2EFC0EC92F2C71" ID-GBIF-Occurrence="2898878482" collectionCode="ZMHB" collectorName="N. Lauenburg Mioko C. Ribbe" country="Papua New Guinea" location="PAPUA NEW GUINEA" pageId="104" pageNumber="105" specimenCode="MFNB 2019" specimenCount="1" typeStatus="syntype">
<collectingCountry id="EA8976709D749233FF2EFC0EC9822BA8" box="[189,468,944,970]" name="Papua New Guinea" pageId="104" pageNumber="105">PAPUA NEW GUINEA</collectingCountry>
•
<specimenCount id="8498FD699D749233FE74FC0EC9A92BA8" box="[487,511,944,970]" pageId="104" pageNumber="105">♂</specimenCount>
; “
<collectorName id="3F6B53369D749233FD87FC0ECB3C2BA8" box="[532,874,944,970]" pageId="104" pageNumber="105">N.Lauenburg Mioko C.Ribbe</collectorName>
[p, black frame] //
<taxonomicName id="559E4D639D749233FBADFC0EC8AF2B8D" authority="Mos." authorityName="Moser" authorityYear="1920" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="104" pageNumber="105" phylum="Arthropoda" rank="species" species="seriepunctatus">
<emphasis id="A0EAEAF29D749233FBADFC0ECD202BA8" box="[1086,1398,944,970]" italics="true" pageId="104" pageNumber="105">Maechidius seriepunctatus</emphasis>
Mos.
</taxonomicName>
<typeStatus id="4D2588429D749233FE91FC6BC96A2B8D" box="[258,316,981,1007]" pageId="104" pageNumber="105">Type</typeStatus>
[h]//
<typeStatus id="4D2588429D749233FE13FC6ACA522B8C" box="[384,516,980,1006]" pageId="104" pageNumber="105" type="syntype">SYNTYPE</typeStatus>
<taxonomicName id="559E4D639D749233FD9DFC6ACBBB2B8C" authority="Moser, 1920" authorityName="Moser" authorityYear="1920" box="[526,1005,980,1007]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="104" pageNumber="105" phylum="Arthropoda" rank="species" species="seriepunctatus">
<emphasis id="A0EAEAF29D749233FD9DFC6ACB1A2B8C" box="[526,844,980,1006]" italics="true" pageId="104" pageNumber="105">Maechidius seriepunctatus</emphasis>
<bibRefCitation id="F60F4B119D749233FCC4FC6BCBBB2B8C" author="Moser J." box="[855,1005,980,1007]" pageId="104" pageNumber="105" pagination="3 - 28" refId="ref98398" refString="Moser J. 1920. Beitrag zur Kenntnis der Melolonthiden XI. Stettiner Entomologische Zeitung 81: 3 - 28." type="journal article" year="1920">Moser, 1920</bibRefCitation>
</taxonomicName>
labelled by
<specimenCode id="C2389E9B9D749233FB16FC6BCD4B2B8C" box="[1157,1309,980,1007]" collectionCode="MFNB" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="104" pageNumber="105">
<collectionCode id="F48FAE259D749233FB16FC6BCC8E2B8D" box="[1157,1240,981,1007]" collectionName="Italy, Udine, Museo Friulano di Storia Naturale" pageId="104" pageNumber="105">MFNB</collectionCode>
2019
</specimenCode>
[p, red label]”;
<collectionCode id="F48FAE259D749233FE8FFC47C9232C71" box="[284,373,1017,1043]" collectionName="Germany, Berlin, Museum fuer Naturkunde der Humboldt-Universitaet" pageId="104" pageNumber="105">ZMHB</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D749233FF2EFB85C9022C93" pageId="104" pageNumber="105" type="discussion">
<paragraph id="922136E09D749233FF2EFB85C9792C37" blockId="104.[189,303,1083,1109]" box="[189,303,1083,1109]" pageId="104" pageNumber="105">
<heading id="C969818C9D749233FF2EFB85C9792C37" bold="true" box="[189,303,1083,1109]" fontSize="11" level="3" pageId="104" pageNumber="105" reason="3">
<emphasis id="A0EAEAF29D749233FF2EFB85C9792C37" bold="true" box="[189,303,1083,1109]" pageId="104" pageNumber="105">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D749233FF2EFBD5C9022C93" blockId="104.[189,1399,1130,1265]" pageId="104" pageNumber="105">
<bibRefCitation id="F60F4B119D749233FF2EFBD5C9C52CE7" author="Moser J." box="[189,403,1130,1157]" pageId="104" pageNumber="105" pagination="3 - 28" refId="ref98398" refString="Moser J. 1920. Beitrag zur Kenntnis der Melolonthiden XI. Stettiner Entomologische Zeitung 81: 3 - 28." type="journal article" year="1920">Moser (1920: 16)</bibRefCitation>
based his description of
<taxonomicName id="559E4D639D749233FD5FFBD5CBF52CE6" authorityName="Moser" authorityYear="1920" box="[716,931,1131,1156]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="104" pageNumber="105" phylum="Arthropoda" rank="species" species="seriepunctatus">
<emphasis id="A0EAEAF29D749233FD5FFBD5CBF52CE6" box="[716,931,1131,1156]" italics="true" pageId="104" pageNumber="105">M. seriepunctatus</emphasis>
</taxonomicName>
on an unstated number of specimens. The single ZMHB
<typeStatus id="4D2588429D749233FE3AFB31CA522CCB" box="[425,516,1167,1193]" pageId="104" pageNumber="105" type="syntype">syntype</typeStatus>
is therefore designated as
<typeStatus id="4D2588429D749233FCC1FB30CBE82CCA" box="[850,958,1166,1192]" pageId="104" pageNumber="105" type="lectotype">lectotype</typeStatus>
in order to enhance the stability of nomenclature and fix the specimen I based my redescription on, in case other
<typeStatus id="4D2588429D749233FBCFFB0DCC952CAF" box="[1116,1219,1203,1229]" pageId="104" pageNumber="105" type="syntype">syntypes</typeStatus>
are discovered in the future.
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D749232FF2EFAA7C9FD2B34" lastPageId="105" lastPageNumber="106" pageId="104" pageNumber="105" type="description">
<paragraph id="922136E09D749233FF2EFAA7C9182D51" blockId="104.[189,334,1305,1331]" box="[189,334,1305,1331]" pageId="104" pageNumber="105">
<heading id="C969818C9D749233FF2EFAA7C9182D51" bold="true" box="[189,334,1305,1331]" fontSize="11" level="3" pageId="104" pageNumber="105" reason="3">
<emphasis id="A0EAEAF29D749233FF2EFAA7C9182D51" bold="true" box="[189,334,1305,1331]" pageId="104" pageNumber="105">Description</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D749233FF2EFAF7CB112D06" blockId="104.[189,839,1353,1380]" box="[189,839,1353,1380]" pageId="104" pageNumber="105">
MEASUREMENTS. Total body length
<quantity id="55669B059D749233FDC2FAF7CAEF2D01" box="[593,697,1353,1379]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="104" pageNumber="105" unit="mm" value="6.8">6.80 mm</quantity>
(
<typeStatus id="4D2588429D749233FD5BFAF7CB6D2D06" box="[712,827,1353,1380]" pageId="104" pageNumber="105" type="lectotype">lectotype</typeStatus>
).
</paragraph>
<paragraph id="922136E09D749232FF2EFA32CA0F2ABF" blockId="104.[189,1399,1419,2025]" lastBlockId="105.[189,1399,265,733]" lastPageId="105" lastPageNumber="106" pageId="104" pageNumber="105">
Dorsum and venter uniformly castaneous brown. Frons convex dorsally between compound eyes, subopaque dorsally and ventrally. Male labroclypeus (
<figureCitation id="0AA52A659D749233FCBCFA11CBC72DA8" box="[815,913,1455,1482]" captionStart="Figs 169–180" captionStartId="143.[189,241,1787,1813]" captionTargetBox="[359,1217,271,1750]" captionTargetId="figure-123@143.[359,1228,265,1755]" captionTargetPageId="143" captionText="Figs 169–180. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal view. 169. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 170. Same, paratype, ♂ (BMNH). 171. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 172. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 173. Same, ♂ (ZMHB). 174. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). 175. M. similis sp. nov., paratype, ♂ (BMNH). 176. Same, paratype, ♀ (BMNH). 177. M. skalei sp. nov., paratype, ♂ (NME). 178. M. sougb sp. nov., holotype, ♂ (NME). 179. M. speciosus (Frey, 1969) comb. nov., ♂ (NHMB). 180. Same, ♀ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122185" httpUri="https://zenodo.org/record/4122185/files/figure.png" pageId="104" pageNumber="105">Fig. 174</figureCitation>
) broadly shallowly emarginate anteriorly, its lateral margins sinuous in both dorsal and lateral views. Anterolateral angles rounded, slightly protruding, raised up at angle of nearly 95° to frons in lateral view. Anterior and lateral margins of labroclypeus smooth. Upper- and underside of labroclypeus with sparse moderately long setae along anterior and lateral margins. Canthus nearly straight in dorsal view. Punctures of frons ovoid, moderately large, rather flat. Intervening spaces microreticulate, variably large. Head setae scale-like, appressed to suberect, those along internal margin of compound eye surpassing length of corresponding punctures, those on labroclypeus, most of frons and vertex minute, rising from anterior margin of corresponding punctures, not surpassing their length. Antenna 9-segmented, club 3-lamellate. Pronotum transverse, flattened dorsally, subopaque dorsally and laterally. Anterior margin of pronotum broadly emarginate with protruding anterolateral angles, basal margin broadly rounded. Lateral margin of pronotum broadly rounded, delicately crenulate all along (
<figureCitation id="0AA52A659D749233FDEFF883CA892F3A" box="[636,735,1853,1880]" captionStart="Figs 242–258" captionStartId="148.[189,241,1646,1672]" captionTargetBox="[394,1193,265,1617]" captionTargetId="figure-193@148.[394,1193,265,1626]" captionTargetPageId="148" captionText="Figs 242–258. Papuan and Wallacean Maechidius Macleay, 1819, pronotum, dorsal view. 242. M. paupianus Heller, 1910, lectotype, ♀ (SNSD). 243. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 244. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 245. M. peregrinus Lansberge, 1886, ♂ (DTC). 246. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 247. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 248. M. riedeli sp. nov., holotype, ♂ (SMNS). 249. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 250. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 251. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). 252. M. similis sp. nov., paratype, ♂ (BMNH). 253. M. skalei sp. nov., paratype, ♂ (NME). 254. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 255. M. sougb sp. nov., holotype, ♂ (NME). 256. M. speciosus (Frey, 1969) comb. nov., ♀ (BMNH). 257. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). 258. M. suwawa sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122196" httpUri="https://zenodo.org/record/4122196/files/figure.png" pageId="104" pageNumber="105">Fig.251</figureCitation>
).Lateral margin of pronotum sinuous in lateral view.Short thickened suberect seta present between every two crenulae. Lateral margins, antero- and posterolateral angles completely covered with microscopical velvety pubescence. Hypomeron emarginate and long setose on anterior margin which is flange-like (
<figureCitation id="0AA52A659D749233FD6EF814CB3E2FA6" box="[765,872,1962,1988]" captionStart="Figs 278–285" captionStartId="151.[189,241,1697,1723]" captionTargetBox="[211,1375,265,1597]" captionTargetId="figure-104@151.[211,1376,265,1614]" captionTargetPageId="151" captionText="Figs 278–285. Papuan and Wallacean Maechidius Macleay, 1819, pronotal hypomeron, ventro-lateral view. 278. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 279. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 280. M. seriepunctatus Moser, 1920, lectotype, ♂ (SNSD). 281. M. speciosus (Frey, 1969) comb. nov., ♀ (BMNH). 282. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). 283. M. trivialis sp. nov., holotype, ♂ (BMNH). 284. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 285. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122202" httpUri="https://zenodo.org/record/4122202/files/figure.png" pageId="104" pageNumber="105">Fig. 280</figureCitation>
). Antennal pocket deep. Pronotal punctures ovoid, deep and dense, intervening spaces glossy, larger than punctures on anterior half, smaller than those on posterior half of disc. Setae minute, arise from anterior margin of corresponding punctures, not surpassing length of punctures. Scutellar shield rounded apically. Elytron subopaque, without tracks of longitudinal carinae. Elytral punctures elongate, dense and deep, arranged in irregular longitudinal rows (
<figureCitation id="0AA52A659D759232FE9BFE3FC92729F9" box="[264,369,385,411]" captionStart="Figs 333–348" captionStartId="155.[189,241,1682,1708]" captionTargetBox="[394,1193,265,1641]" captionTargetId="figure-177@155.[394,1193,265,1655]" captionTargetPageId="155" captionText="Figs 333–348. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 333. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 334. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 335. M. peregrinus Lansberge, 1886, ♂ (DTC). 336. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 337. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 338. Same, enlarged (SMNS). 339. M. riedeli sp. nov., holotype, ♂ (SMNS). 340. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 341. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 342. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). 343. M. similis sp. nov., paratype, ♂ (BMNH). 344. M. skalei sp. nov., paratype, ♂ (NME). 345. M. sougb sp. nov., holotype, ♂ (NME). 346. M. speciosus (Frey, 1969) comb. nov., ♀(BMNH). 347. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 348. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122212" httpUri="https://zenodo.org/record/4122212/files/figure.png" pageId="105" pageNumber="106">Fig. 342</figureCitation>
). Intervening spaces glossy, generally smaller than punctures, in part wrinkled. Elytral setae inconspicuous, scale-like, appressed; seta rises from anterior margin of each puncture, generally not surpassing length of corresponding puncture. Some longer scale-like clavate scattered over disc and especially abundant along lateral sides. Male pygidium flattened dorsally, very vaguely medially longitudinally impressed, with dense rather deep ovoid punctures (Fig. 522). Intervening spaces glossy, variably large. Setae of pygidium scale-like, in part clavate, sparse, suberect, those on distal half longer than corresponding punctures. Male protibia widened distally, with two obtuse external teeth (Fig. 414). Male protibial terminal spur straight. Male metatibial terminal spurs long, subequal, pointed. Tarsal claws with pulvilli (in male only?).
</paragraph>
<paragraph id="922136E09D759232FF2EFCB6C9E62B40" blockId="105.[189,432,776,802]" box="[189,432,776,802]" pageId="105" pageNumber="106">
<heading id="C969818C9D759232FF2EFCB6C9E62B40" bold="true" box="[189,432,776,802]" fontSize="11" level="3" pageId="105" pageNumber="106" reason="3">
<emphasis id="A0EAEAF29D759232FF2EFCB6C9E62B40" bold="true" box="[189,432,776,802]" pageId="105" pageNumber="106">Sexual dimorphism</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D759232FF2EFC82C9FD2B34" blockId="105.[189,427,828,854]" box="[189,427,828,854]" pageId="105" pageNumber="106">Female is unknown.</paragraph>
</subSubSection>
</treatment>
</document>
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