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<document id="056A7780C4A20F19590A1899BCA0CF11" ID-CLB-Dataset="9693" ID-DOI="10.5852/ejt.2020.721.1127" ID-GBIF-Dataset="0cd61b23-1089-4c1b-bfc0-487887abad84" ID-Zenodo-Dep="4122118" ID-ZooBank="89E62EF8-2E45-4C59-94B7-6A5603E8939B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1603474447689" checkinUser="valdenar" docAuthor="Telnov, Dmitry" docDate="2020" docId="1A3787F69D609226FDE7FA9FCA4F2F8A" docLanguage="en" docName="ejt-721_telnov.pdf" docOrigin="European Journal of Taxonomy 721" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Maechidius weigeli Telnov 2020, sp. nov." docType="treatment" docUuid="9C17E972-B606-419C-AC72-6713B01D5546" docUuidSource="ZooBank" docVersion="10" lastPageNumber="126" masterDocId="E60EFF8E9D1C925BFF93FFBEC8562862" masterDocTitle="A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallaces Line" masterLastPageNumber="210" masterPageNumber="1" pageNumber="125" updateTime="1698846101371" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="B7D994D1479236B73F0C70382314081E">A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallaces Line</mods:title>
</mods:titleInfo>
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<mods:namePart id="9611F7BAC6A90870670F7B0781A31D28">Telnov, Dmitry</mods:namePart>
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<mods:title id="5CAA799F09B3CA3B073A28EC8ED22A76">European Journal of Taxonomy</mods:title>
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<mods:part id="0523CF030ACA17DA96193988031A19C8">
<mods:date id="5C56666E25EF56430C03F0EB9E8F62CF">2020</mods:date>
<mods:detail id="DD65188CA98360A0310705B1942D6634" type="pubDate">
<mods:number id="471F65D8076F2FCD1FD359F5E07F269A">2020-10-19</mods:number>
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<mods:classification id="61AD6912DC9F5AF7CC2F768F80038E52">journal article</mods:classification>
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<mods:identifier id="3C07A928F60B878803A9891DF7F2E05C" type="DOI">10.5852/ejt.2020.721.1127</mods:identifier>
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<treatment id="1A3787F69D609226FDE7FA9FCA4F2F8A" ID-DOI="http://doi.org/10.5281/zenodo.4344339" ID-GBIF-Taxon="168648174" ID-Zenodo-Dep="4344339" LSID="urn:lsid:zoobank.org:act:9C17E972-B606-419C-AC72-6713B01D5546" httpUri="http://treatment.plazi.org/id/1A3787F69D609226FDE7FA9FCA4F2F8A" lastPageId="125" lastPageNumber="126" pageId="124" pageNumber="125">
<subSubSection id="DA84656B9D609227FDE7FA9FCB962D5E" box="[628,960,1313,1340]" pageId="124" pageNumber="125" type="nomenclature">
<paragraph id="922136E09D609227FDE7FA9FCB962D5E" blockId="124.[372,1215,1313,1411]" box="[628,960,1313,1340]" pageId="124" pageNumber="125">
<heading id="C969818C9D609227FDE7FA9FCB962D5E" box="[628,960,1313,1340]" centered="true" fontSize="11" level="2" pageId="124" pageNumber="125" reason="2">
<taxonomicName id="559E4D639D609227FDE7FA9FCB082D5E" ID-CoL="7X3XY" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[628,862,1313,1340]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="124" pageNumber="125" phylum="Arthropoda" rank="species" species="weigeli" status="sp. nov.">
<emphasis id="A0EAEAF29D609227FDE7FA9FCB082D5E" bold="true" box="[628,862,1313,1340]" italics="true" pageId="124" pageNumber="125">Maechidius weigeli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D609227FCF6FA9CCB962D5E" box="[869,960,1314,1340]" pageId="124" pageNumber="125" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D609227FEE7FAFBCC0A2DE0" pageId="124" pageNumber="125" type="description">
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<uri id="E60F3AE29D609227FEE7FAFBCCE92D02" box="[372,1215,1349,1376]" pageId="124" pageNumber="125">
urn:lsid:zoobank.org:act:
<uuid id="E6380C359D609227FD07FAFBCCE92D02" box="[660,1215,1349,1376]" pageId="124" pageNumber="125">9C17E972-B606-419C-AC72-6713B01D5546</uuid>
</uri>
</paragraph>
<paragraph id="922136E09D609227FE44FAD6CC0A2DE0" blockId="124.[372,1215,1313,1411]" box="[471,1116,1384,1411]" pageId="124" pageNumber="125">
<figureCitation id="0AA52A659D609227FE44FAD6CA742DE0" box="[471,546,1384,1410]" captionStart="Figs 27" captionStartId="7.[189,241,1893,1919]" captionTargetBox="[295,1291,270,1861]" captionTargetId="figure-76@7.[295,1294,265,1861]" captionTargetPageId="7" captionText="Figs 27. Maechidiini Burmeister, 1855 features. 2. Maechidius esau Heller, 1914, head, ventro-lateral view. 3. Same, head and pronotum, ventral view. 4. Maechidius sp., same. 5. M. bilobiceps (Fairmaire, 1877) comb. nov., prosternum and pronotal hypomera, ventral view (BMNH). 6. M. weigeli sp. nov., paratype, ♂, head, lateral view (NME). 7. M. spurius Macleay, 1819, mesoventrite. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122124" httpUri="https://zenodo.org/record/4122124/files/figure.png" pageId="124" pageNumber="125">Figs 6</figureCitation>
,
<figureCitation id="0AA52A659D609227FDBCFAD6CA192DE0" box="[559,591,1384,1410]" captionStart="Figs 9496" captionStartId="126.[189,241,1928,1954]" captionTargetBox="[346,1191,267,1904]" captionTargetId="figure-55@126.[339,1249,265,1909]" captionTargetPageId="126" captionText="Figs 9496. Papuan and Wallacean Maechidius Macleay, 1819, habitus, dorsal view. 94. M. weigeli sp. nov., holotype, ♂ (NME). 95. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 96. M. yamdena sp. nov., paratype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122171" httpUri="https://zenodo.org/record/4122171/files/figure.png" pageId="124" pageNumber="125">94</figureCitation>
,
<figureCitation id="0AA52A659D609227FDC8FAD6CA922DE0" box="[603,708,1384,1410]" captionStart="Figs 181192" captionStartId="144.[189,241,1787,1813]" captionTargetBox="[338,1249,274,1742]" captionTargetId="figure-122@144.[338,1249,265,1749]" captionTargetPageId="144" captionText="Figs 181192. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal view. 181. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 182. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). 183. Same, ♀ (SNSD). 184. M. suwawa sp. nov., holotype, ♂ (BMNH). 185. Same, paratype, ♀ (BMNH). 186. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 187. M. trivialis sp. nov., holotype, ♂ (BMNH). 188. M. ursus sp. nov., holotype, ♀ (NME). 189. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 190. Same, ♂ (SNSD). 191. M. weigeli sp. nov., holotype, ♂ (NME). 192. Same, paratype, ♀ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122187" httpUri="https://zenodo.org/record/4122187/files/figure.png" pageId="124" pageNumber="125">191192</figureCitation>
,
<figureCitation id="0AA52A659D609227FD43FAD6CAA92DE0" box="[720,767,1384,1410]" captionStart="Figs 259265" captionStartId="149.[189,241,1372,1398]" captionTargetBox="[189,1379,265,1308]" captionTargetId="figure-88@149.[189,1398,265,1311]" captionTargetPageId="149" captionText="Figs 259265. Papuan and Wallacean Maechidius Macleay, 1819, pronotum, dorsal view. 259. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 260. M. trivialis sp. nov., holotype, ♂ (BMNH). 261. M. ursus sp. nov., holotype, ♀ (NME). 262. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 263. M. weigeli sp. nov., holotype, ♂ (NME). 264. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 265. M. yamdena sp. nov., paratype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122198" httpUri="https://zenodo.org/record/4122198/files/figure.png" pageId="124" pageNumber="125">263</figureCitation>
,
<figureCitation id="0AA52A659D609227FC98FAD6CB6C2DE1" box="[779,826,1384,1411]" captionStart="Figs 349358" captionStartId="156.[189,241,1786,1812]" captionTargetBox="[287,1299,265,1743]" captionTargetId="figure-123@156.[287,1301,265,1760]" captionTargetPageId="156" captionText="Figs 349358. Papuan and Wallacean Maechidius Macleay, 1819, elytron (349356) and pronotal setation (357358), dorsal view. 349. M. suwawa sp. nov., paratype, ♀ (BMNH). 350. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 351. M. trivialis sp. nov., holotype, ♂ (BMNH). 352. M. ursus sp. nov., holotype, ♀ (NME). 353. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 354. M. weigeli sp. nov., holotype, ♂ (NME). 355. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 356. M. yamdena sp. nov., paratype, ♂ (BMNH). 357. M. leucopsar sp. nov., holotype, ♂ (IECA). 358. M. ursus sp. nov., holotype, ♀ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122214" httpUri="https://zenodo.org/record/4122214/files/figure.png" pageId="124" pageNumber="125">354</figureCitation>
,
<figureCitation id="0AA52A659D609227FCD6FAD6CB222DE0" box="[837,884,1384,1410]" captionStart="Figs 424435" captionStartId="161.[189,241,1394,1420]" captionTargetBox="[238,1349,265,1318]" captionTargetId="figure-136@161.[217,1354,265,1322]" captionTargetPageId="161" captionText="Figs 424435. Papuan and Wallacean Maechidius Macleay, 1819, protibia, dorsal view (424428), and abdominal ventrites (429435), ventral and lateral views. 424. M. ursus sp. nov., holotype, ♀ (NME). 425. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 426. M. weigeli sp. nov., holotype, ♂ (NME). 427. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 428. M. yamdena sp. nov., paratype, ♂ (BMNH). 429. M.dani sp. nov., holotype, ♂ (IECA). 430. Same, different aspect (IECA). 431. M. hamatus sp. nov., holotype, ♂ (IECA). 432. Same, different aspect (IECA). 433. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 434. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). 435. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122216" httpUri="https://zenodo.org/record/4122216/files/figure.png" pageId="124" pageNumber="125">426</figureCitation>
, 531532, 739741
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D609227FF2EFA17CD212D90" pageId="124" pageNumber="125" type="diagnosis">
<paragraph id="922136E09D609227FF2EFA17C99F2DA1" blockId="124.[189,457,1449,1475]" box="[189,457,1449,1475]" pageId="124" pageNumber="125">
<heading id="C969818C9D609227FF2EFA17C99F2DA1" bold="true" box="[189,457,1449,1475]" fontSize="11" level="3" pageId="124" pageNumber="125" reason="3">
<emphasis id="A0EAEAF29D609227FF2EFA17C99F2DA1" bold="true" box="[189,457,1449,1475]" pageId="124" pageNumber="125">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D609227FF2EFA66CD212D90" blockId="124.[189,1399,1496,1523]" box="[189,1399,1496,1523]" pageId="124" pageNumber="125">Differs from all congeners in the shape of the aedeagus and in the setation and structure of the pygidium.</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D609227FF2EF9A4CAFE2EE4" pageId="124" pageNumber="125" type="etymology">
<paragraph id="922136E09D609227FF2EF9A4C9122E56" blockId="124.[189,324,1562,1588]" box="[189,324,1562,1588]" pageId="124" pageNumber="125">
<heading id="C969818C9D609227FF2EF9A4C9122E56" bold="true" box="[189,324,1562,1588]" fontSize="11" level="3" pageId="124" pageNumber="125" reason="3">
<emphasis id="A0EAEAF29D609227FF2EF9A4C9122E56" bold="true" box="[189,324,1562,1588]" pageId="124" pageNumber="125">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D609227FF2EF9F6CAFE2EE4" blockId="124.[189,1398,1608,1670]" pageId="124" pageNumber="125">
Patronymic. This species is named after Andreas Weigel (Wernburg,
<collectingCountry id="EA8976709D609227FC4DF9F7CC062E01" box="[990,1104,1609,1635]" name="Germany" pageId="124" pageNumber="125">Germany</collectingCountry>
), a famous expert on oldworld
<taxonomicName id="559E4D639D609227FE9AF9D2C9E42EE4" authorityName="Latreille" authorityYear="1802" box="[265,434,1644,1670]" class="Insecta" family="Cerambycidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="124" pageNumber="125" phylum="Arthropoda" rank="family">Cerambycidae</taxonomicName>
and my good friend.
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D609227FF2EF910CAD52F8A" pageId="124" pageNumber="125" type="materials_examined">
<paragraph id="922136E09D609227FF2EF910C93A2EA5" blockId="124.[189,364,1709,1736]" box="[189,364,1709,1736]" pageId="124" pageNumber="125">
<heading id="C969818C9D609227FF2EF910C93A2EA5" bold="true" box="[189,364,1709,1736]" fontSize="11" level="3" pageId="124" pageNumber="125" reason="3">
<emphasis id="A0EAEAF29D609227FF2EF910C93A2EA5" bold="true" box="[189,364,1709,1736]" pageId="124" pageNumber="125">Type material</emphasis>
</heading>
</paragraph>
<materialsCitation id="22F63CBD9D609227FF79F962C9232F5F" ID-GBIF-Occurrence="2898878365" collectingDate="1996-01-06" collectionCode="NME" collectorName="A. Weigel" country="Indonesia" elevation="1150" location="Nabire Epomani" pageId="124" pageNumber="125" specimenCount="1" stateProvince="Papua Barat" typeStatus="holotype">
<paragraph id="922136E09D609227FF79F962C90A2E94" blockId="124.[189,1398,1756,1853]" box="[234,348,1756,1782]" pageId="124" pageNumber="125">
<emphasis id="A0EAEAF29D609227FF79F962C90A2E94" bold="true" box="[234,348,1756,1782]" pageId="124" pageNumber="125">
<typeStatus id="4D2588429D609227FF79F962C90A2E94" box="[234,348,1756,1782]" pageId="124" pageNumber="125" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="922136E09D609227FF2EF8BEC9232F5F" blockId="124.[189,1398,1756,1853]" pageId="124" pageNumber="125">
<collectingCountry id="EA8976709D609227FF2EF8BEC90D2F78" box="[189,347,1792,1818]" name="Indonesia" pageId="124" pageNumber="125">INDONESIA</collectingCountry>
<specimenCount id="8498FD699D609227FEE8F8BEC9C52F78" box="[379,403,1792,1818]" pageId="124" pageNumber="125"></specimenCount>
; “
<collectingCountry id="EA8976709D609227FE3EF8BECA182F78" box="[429,590,1792,1818]" name="Indonesia" pageId="124" pageNumber="125">INDONESIA</collectingCountry>
or.
<collectingRegion id="505AF8029D609227FD11F941CAAE2F78" box="[642,760,1791,1818]" country="Indonesia" name="Papua Barat" pageId="124" pageNumber="125">Irian Jaya</collectingRegion>
<quantity id="55669B059D609227FC91F941CB022F7B" box="[770,852,1791,1817]" metricMagnitude="5" metricUnit="m" metricValue="1.7" pageId="124" pageNumber="125" unit="km" value="170.0">170km</quantity>
S
<location id="9741603B9D609227FCEAF941CC682F7B" LSID="urn:lsid:plazi:treatment:1A3787F69D609226FDE7FA9FCA4F2F8A:9741603B9D609227FCEAF941CC682F7B" box="[889,1086,1791,1817]" country="Indonesia" name="Nabire Epomani" pageId="124" pageNumber="125" stateProvince="Papua Barat">Nabire Epomani</location>
<quantity id="55669B059D609227FBDBF941CCCA2F78" box="[1096,1180,1791,1818]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="124" pageNumber="125" unit="m" value="1150.0">
<elevation id="19B3D1D39D609227FBDBF941CCCA2F78" box="[1096,1180,1791,1818]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="124" pageNumber="125" unit="m" value="1150.0">1150m</elevation>
</quantity>
,
<date id="E62010209D609227FB38F941CD4D2F7B" box="[1195,1307,1791,1817]" pageId="124" pageNumber="125" value="1996-01-06">
<collectingDate id="F664E9C89D609227FB38F941CD4D2F7B" box="[1195,1307,1791,1817]" pageId="124" pageNumber="125" value="1996-01-06">06.I.1996</collectingDate>
</date>
leg.
<collectorName id="3F6B53369D609227FAC9F8BEC9402F5F" pageId="124" pageNumber="125">A. Weigel</collectorName>
”;
<collectionCode id="F48FAE259D609227FEBFF89DC9272F5F" box="[300,369,1827,1853]" country="Germany" httpUri="http://grbio.org/cool/87iw-xebj" name="Sammlung des Naturkundemseum Erfurt" pageId="124" pageNumber="125">NME</collectionCode>
.
</paragraph>
</materialsCitation>
<paragraph id="922136E09D609227FF79F8DACA452F1D" blockId="124.[189,1398,1892,2024]" box="[234,531,1892,1919]" pageId="124" pageNumber="125">
<emphasis id="A0EAEAF29D609227FF79F8DAC9312F1C" bold="true" box="[234,359,1892,1918]" pageId="124" pageNumber="125">
<typeStatus id="4D2588429D609227FF79F8DAC9312F1C" box="[234,359,1892,1918]" pageId="124" pageNumber="125" type="paratype">Paratypes</typeStatus>
</emphasis>
(
<specimenCount id="8498FD699D609227FEEAF8DACA5D2F1D" box="[377,523,1892,1919]" pageId="124" pageNumber="125" type="generic">5 specimens</specimenCount>
)
</paragraph>
<paragraph id="922136E09D609227FF2EF836CAD52F8A" blockId="124.[189,1398,1892,2024]" pageId="124" pageNumber="125">
<materialsCitation id="22F63CBD9D609227FF2EF836CBD72FC0" ID-GBIF-Occurrence="2898878549" box="[189,897,1928,1954]" collectionCode="NME" country="Indonesia" location="INDONESIA" pageId="124" pageNumber="125" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<collectingCountry id="EA8976709D609227FF2EF836C90D2FC0" box="[189,347,1928,1954]" name="Indonesia" pageId="124" pageNumber="125">INDONESIA</collectingCountry>
<specimenCount id="8498FD699D609227FEE4F836C9F02FC0" box="[375,422,1928,1954]" pageId="124" pageNumber="125" type="male">1 ♂</specimenCount>
,
<specimenCount id="8498FD699D609227FE26F836C9B22FC0" box="[437,484,1928,1954]" pageId="124" pageNumber="125" type="female">1 ♀</specimenCount>
; same label as for
<typeStatus id="4D2588429D609227FD5BF836CB642FC0" box="[712,818,1928,1954]" pageId="124" pageNumber="125" type="holotype">holotype</typeStatus>
;
<collectionCode id="F48FAE259D609227FCD3F836CBD72FC0" box="[832,897,1928,1954]" country="Germany" httpUri="http://grbio.org/cool/87iw-xebj" name="Sammlung des Naturkundemseum Erfurt" pageId="124" pageNumber="125">NME</collectionCode>
</materialsCitation>
<materialsCitation id="22F63CBD9D609227FC0DF836CD352FC0" ID-GBIF-Occurrence="2898878599" box="[926,1379,1928,1954]" collectionCode="DTC" country="Indonesia" pageId="124" pageNumber="125" specimenCount="1" typeStatus="holotype">
<specimenCount id="8498FD699D609227FC0DF836CB9B2FC0" box="[926,973,1928,1954]" pageId="124" pageNumber="125" type="male">1 ♂</specimenCount>
; same label as for
<typeStatus id="4D2588429D609227FB22F836CD4D2FC0" box="[1201,1307,1928,1954]" pageId="124" pageNumber="125" type="holotype">holotype</typeStatus>
;
<collectionCode id="F48FAE259D609227FABBF836CD352FC0" box="[1320,1379,1928,1954]" collectionName="Dmitrijs Telnovs" pageId="124" pageNumber="125">DTC</collectionCode>
</materialsCitation>
<materialsCitation id="22F63CBD9D609227FF2EF815CAD62F8A" ID-GBIF-Occurrence="2898878402" collectingDate="1999-03-01" collectingDateMax="1999-03-02" collectingDateMin="1999-03-01" collectionCode="SMNS" collectorName="Waropen Pr. Wapoga Riv." country="Indonesia" elevation="100" location="IRIAN JAYA" pageId="124" pageNumber="125" specimenCount="1" stateProvince="Papua Barat" typeStatus="holotype">
<specimenCount id="8498FD699D609227FF2EF815C9562FA4" box="[189,256,1963,1990]" pageId="124" pageNumber="125" type="male">2 ♂♂</specimenCount>
; “
<collectingRegion id="505AF8029D609227FE84F815C9E82FA7" box="[279,446,1963,1989]" country="Indonesia" name="Papua Barat" pageId="124" pageNumber="125">IRIAN JAYA</collectingRegion>
:J.-
<collectorName id="3F6B53369D609227FE44F815CB5C2FA7" box="[471,778,1963,1990]" pageId="124" pageNumber="125">Waropen Pr. Wapoga Riv.</collectorName>
,
<quantity id="55669B059D609227FC87F815CB0C2FA7" box="[788,858,1963,1989]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="124" pageNumber="125" unit="m" value="100.0">
<elevation id="19B3D1D39D609227FC87F815CB0C2FA7" box="[788,858,1963,1989]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="124" pageNumber="125" unit="m" value="100.0">100m</elevation>
</quantity>
, Kwadewa, loading [sic
<specimenCount id="8498FD699D609227FBE4F815CC2D2FA7" box="[1143,1147,1963,1989]" pageId="124" pageNumber="125"></specimenCount>
logging] road, km
<date id="E62010209D609227FACAF815C95C2F8A" pageId="124" pageNumber="125" value="1980-03-01" valueMax="1980-03-02" valueMin="1980-03-01">
80
<collectingDate id="F664E9C89D609227FF2EF870C95C2F8A" box="[189,266,1998,2024]" pageId="124" pageNumber="125" value="1999-03-01" valueMax="1999-03-02" valueMin="1999-03-01">1-2.III.</collectingDate>
</date>
1999, leg. A.RIEDEL”;
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</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D619226FF2EFEB6CAEE2EBD" pageId="125" pageNumber="126" type="description">
<paragraph id="922136E09D619226FF2EFEB6C9182940" blockId="125.[189,334,264,290]" box="[189,334,264,290]" pageId="125" pageNumber="126">
<heading id="C969818C9D619226FF2EFEB6C9182940" bold="true" box="[189,334,264,290]" fontSize="11" level="3" pageId="125" pageNumber="126" reason="3">
<emphasis id="A0EAEAF29D619226FF2EFEB6C9182940" bold="true" box="[189,334,264,290]" pageId="125" pageNumber="126">Description</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D619226FF2EFE87CB1829F9" blockId="125.[189,1399,312,412]" pageId="125" pageNumber="126">
MEASUREMENTS. Male
<typeStatus id="4D2588429D619226FE52FE86CA7D2930" box="[449,555,312,338]" pageId="125" pageNumber="126" type="holotype">holotype</typeStatus>
, total body length
<quantity id="55669B059D619226FC9AFE86CB202931" box="[777,886,312,339]" metricMagnitude="-3" metricUnit="m" metricValue="6.6" pageId="125" pageNumber="126" unit="mm" value="6.6">6.60 mm</quantity>
. Head
<quantity id="55669B059D619226FC5AFE86CC652931" box="[969,1075,312,339]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="125" pageNumber="126" unit="mm" value="1.4">1.40 mm</quantity>
long, across eyes
<quantity id="55669B059D619226FA9EFE86CD212931" box="[1293,1399,312,339]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="125" pageNumber="126" unit="mm" value="1.6">1.60 mm</quantity>
wide. Pronotum
<quantity id="55669B059D619226FEEDFEE3C9B02915" box="[382,486,349,375]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="125" pageNumber="126" unit="mm" value="1.4">1.40 mm</quantity>
long, maximum width
<quantity id="55669B059D619226FD66FEE3CB092915" box="[757,863,349,375]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="125" pageNumber="126" unit="mm" value="2.1">2.10 mm</quantity>
. Elytral length
<quantity id="55669B059D619226FB83FEE3CC2D2915" box="[1040,1147,349,375]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="125" pageNumber="126" unit="mm" value="3.8">3.80 mm</quantity>
, maximum combined width
<quantity id="55669B059D619226FE94FE3FC92529FE" box="[263,371,385,412]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="125" pageNumber="126" unit="mm" value="2.9">2.90 mm</quantity>
. Selected
<typeStatus id="4D2588429D619226FE7AFE3CCA0F29FE" box="[489,601,386,412]" pageId="125" pageNumber="126" type="paratype">paratypes</typeStatus>
<quantity id="55669B059D619226FDF2FE3FCB5D29FE" box="[609,779,385,412]" metricMagnitude="-3" metricUnit="m" metricValue="6.1" metricValueMax="6.4" metricValueMin="5.8" pageId="125" pageNumber="126" unit="mm" value="6.1" valueMax="6.4" valueMin="5.8">5.806.40 mm</quantity>
long.
</paragraph>
<paragraph id="922136E09D619226FF2EFE7ACAB92E2A" blockId="125.[189,1399,452,1608]" pageId="125" pageNumber="126">
With general features of the
<taxonomicName id="559E4D639D619226FDB6FE7ACAC729BC" authorityName="Heller" authorityYear="1910" box="[549,657,452,478]" class="Insecta" family="Scarabaeidae" genus="Paramaechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="125" pageNumber="126" phylum="Arthropoda" rank="species" species="pauxillus">
<emphasis id="A0EAEAF29D619226FDB6FE7ACAC729BC" box="[549,657,452,478]" italics="true" pageId="125" pageNumber="126">pauxillus</emphasis>
</taxonomicName>
group. Dorsal surface opaque except on anterior margin of labroclypeus. Frons with paired hump. Male labroclypeus (
<figureCitation id="0AA52A659D619226FCE9FE56CB882A60" box="[890,990,488,514]" captionStart="Figs 181192" captionStartId="144.[189,241,1787,1813]" captionTargetBox="[338,1249,274,1742]" captionTargetId="figure-122@144.[338,1249,265,1749]" captionTargetPageId="144" captionText="Figs 181192. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal view. 181. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 182. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). 183. Same, ♀ (SNSD). 184. M. suwawa sp. nov., holotype, ♂ (BMNH). 185. Same, paratype, ♀ (BMNH). 186. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 187. M. trivialis sp. nov., holotype, ♂ (BMNH). 188. M. ursus sp. nov., holotype, ♀ (NME). 189. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 190. Same, ♂ (SNSD). 191. M. weigeli sp. nov., holotype, ♂ (NME). 192. Same, paratype, ♀ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122187" httpUri="https://zenodo.org/record/4122187/files/figure.png" pageId="125" pageNumber="126">Fig. 191</figureCitation>
) broadly rather deeply emarginate on anterior margin. Anterolateral angles of labroclypeus acute in male, strongly protruding, raised up at angle of nearly 90° to frons in lateral view (
<figureCitation id="0AA52A659D619226FD53FD8FCB5E2A2E" box="[704,776,561,588]" captionStart="Figs 27" captionStartId="7.[189,241,1893,1919]" captionTargetBox="[295,1291,270,1861]" captionTargetId="figure-76@7.[295,1294,265,1861]" captionTargetPageId="7" captionText="Figs 27. Maechidiini Burmeister, 1855 features. 2. Maechidius esau Heller, 1914, head, ventro-lateral view. 3. Same, head and pronotum, ventral view. 4. Maechidius sp., same. 5. M. bilobiceps (Fairmaire, 1877) comb. nov., prosternum and pronotal hypomera, ventral view (BMNH). 6. M. weigeli sp. nov., paratype, ♂, head, lateral view (NME). 7. M. spurius Macleay, 1819, mesoventrite. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122124" httpUri="https://zenodo.org/record/4122124/files/figure.png" pageId="125" pageNumber="126">Fig. 6</figureCitation>
). Female labroclypeus with comparatively less acute anterolateral angles (
<figureCitation id="0AA52A659D619226FE27FDE8CA4D2A12" box="[436,539,598,624]" captionStart="Figs 181192" captionStartId="144.[189,241,1787,1813]" captionTargetBox="[338,1249,274,1742]" captionTargetId="figure-122@144.[338,1249,265,1749]" captionTargetPageId="144" captionText="Figs 181192. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal view. 181. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 182. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). 183. Same, ♀ (SNSD). 184. M. suwawa sp. nov., holotype, ♂ (BMNH). 185. Same, paratype, ♀ (BMNH). 186. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 187. M. trivialis sp. nov., holotype, ♂ (BMNH). 188. M. ursus sp. nov., holotype, ♀ (NME). 189. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 190. Same, ♂ (SNSD). 191. M. weigeli sp. nov., holotype, ♂ (NME). 192. Same, paratype, ♀ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122187" httpUri="https://zenodo.org/record/4122187/files/figure.png" pageId="125" pageNumber="126">Fig. 192</figureCitation>
). Lateral margins of labroclypeus slightly sinuous in dorsal, but strongly in lateral view. Head dorsum except on anterior margin of labroclypeus covered with microscopic velvety pubescence. Head punctures elongate, very deep, moderately dense. Head with moderately long, appressed scale-like setae. Setae becoming distinctly longer, suberect to erect and stronger clavate on humps of frons (
<figureCitation id="0AA52A659D619226FE2FFD59CA3F2B60" box="[444,617,743,770]" captionStart="Figs 181192" captionStartId="144.[189,241,1787,1813]" captionTargetBox="[338,1249,274,1742]" captionTargetId="figure-122@144.[338,1249,265,1749]" captionTargetPageId="144" captionText="Figs 181192. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal view. 181. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 182. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). 183. Same, ♀ (SNSD). 184. M. suwawa sp. nov., holotype, ♂ (BMNH). 185. Same, paratype, ♀ (BMNH). 186. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 187. M. trivialis sp. nov., holotype, ♂ (BMNH). 188. M. ursus sp. nov., holotype, ♀ (NME). 189. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 190. Same, ♂ (SNSD). 191. M. weigeli sp. nov., holotype, ♂ (NME). 192. Same, paratype, ♀ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122187" httpUri="https://zenodo.org/record/4122187/files/figure.png" pageId="125" pageNumber="126">Figs 191192</figureCitation>
). Antenna 9-segmented, club 3-lamellate. Pronotum transverse, slightly sinuous on anterior margin with protruding anterolateral angles. Basal margin broadly rounded, posterolateral angles right-angled. Lateral margin of pronotum in dorsal view almost straight to gradually widened towards middle, moderately strongly emarginate postmedially, crenulate all along (
<figureCitation id="0AA52A659D619226FA92FCEBCD3F2B12" box="[1281,1385,853,880]" captionStart="Figs 259265" captionStartId="149.[189,241,1372,1398]" captionTargetBox="[189,1379,265,1308]" captionTargetId="figure-88@149.[189,1398,265,1311]" captionTargetPageId="149" captionText="Figs 259265. Papuan and Wallacean Maechidius Macleay, 1819, pronotum, dorsal view. 259. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 260. M. trivialis sp. nov., holotype, ♂ (BMNH). 261. M. ursus sp. nov., holotype, ♀ (NME). 262. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 263. M. weigeli sp. nov., holotype, ♂ (NME). 264. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 265. M. yamdena sp. nov., paratype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122198" httpUri="https://zenodo.org/record/4122198/files/figure.png" pageId="125" pageNumber="126">Fig. 263</figureCitation>
). Lateral margin of pronotum sinuous in lateral view. Two flat humps on anterior margin opposite to those on frons (
<figureCitation id="0AA52A659D619226FEA0FC20C9CA2BDB" box="[307,412,926,953]" captionStart="Figs 259265" captionStartId="149.[189,241,1372,1398]" captionTargetBox="[189,1379,265,1308]" captionTargetId="figure-88@149.[189,1398,265,1311]" captionTargetPageId="149" captionText="Figs 259265. Papuan and Wallacean Maechidius Macleay, 1819, pronotum, dorsal view. 259. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 260. M. trivialis sp. nov., holotype, ♂ (BMNH). 261. M. ursus sp. nov., holotype, ♀ (NME). 262. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 263. M. weigeli sp. nov., holotype, ♂ (NME). 264. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 265. M. yamdena sp. nov., paratype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122198" httpUri="https://zenodo.org/record/4122198/files/figure.png" pageId="125" pageNumber="126">Fig. 263</figureCitation>
). Pronotal punctures ovoid, variably large, very deep, dense. Pronotal dorsum and hypomeron covered with microscopic velvety pubescence. Pronotal setae minute, rise from anterior margin of each puncture, generally not surpassing midlength of corresponding puncture. Few slightly longer scale-like setae scattered over pronotal disc, not surpassing length of corresponding punctures. Posterolateral angles of pronotum with moderately long scale-like clavate setae. Setae on pronotal humps also long, erect, scale-like and clavate. Small group of large appressed setae on either side of pronotum along postmedian emargination. Hypomeron flange-like produced, slightly emarginate and very long setose opposite to compound eye. Antennal pocket deep. Scutellar shield rounded apically. Elytron with glabrous elevated broadly interrupted track of sutural carina. Elytral punctures irregularly shaped, deep and dense (
<figureCitation id="0AA52A659D619226FED2FB58C9F02D63" box="[321,422,1254,1281]" captionStart="Figs 349358" captionStartId="156.[189,241,1786,1812]" captionTargetBox="[287,1299,265,1743]" captionTargetId="figure-123@156.[287,1301,265,1760]" captionTargetPageId="156" captionText="Figs 349358. Papuan and Wallacean Maechidius Macleay, 1819, elytron (349356) and pronotal setation (357358), dorsal view. 349. M. suwawa sp. nov., paratype, ♀ (BMNH). 350. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 351. M. trivialis sp. nov., holotype, ♂ (BMNH). 352. M. ursus sp. nov., holotype, ♀ (NME). 353. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 354. M. weigeli sp. nov., holotype, ♂ (NME). 355. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 356. M. yamdena sp. nov., paratype, ♂ (BMNH). 357. M. leucopsar sp. nov., holotype, ♂ (IECA). 358. M. ursus sp. nov., holotype, ♀ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122214" httpUri="https://zenodo.org/record/4122214/files/figure.png" pageId="125" pageNumber="126">Fig. 354</figureCitation>
) and generally smaller than those on pronotum. Setae minute, arising from anterior margin of each puncture, not surpassing midlength of corresponding punctures. Much longer suberect scale-like clavate setae arranged in irregular longitudinal rows across elytra. Male pygidium flattened dorsally, slightly impressed in anterior third, dense and deep moderately large punctate (Fig. 531). Intervening spaces opaque, microreticulate, covered with microscopical velvety pubescence. Setae of pygidium minute, with paired group of scale-like clavate suberect pubescence in median part. Female pygidium with setae more regular in size and shape (Fig. 532). Protibia with inconspicuous longitudinal dorsal carina, with two inconspicuous distal teeth (
<figureCitation id="0AA52A659D619226FCBCFA5BCBCA2E62" box="[815,924,1509,1536]" captionStart="Figs 424435" captionStartId="161.[189,241,1394,1420]" captionTargetBox="[238,1349,265,1318]" captionTargetId="figure-136@161.[217,1354,265,1322]" captionTargetPageId="161" captionText="Figs 424435. Papuan and Wallacean Maechidius Macleay, 1819, protibia, dorsal view (424428), and abdominal ventrites (429435), ventral and lateral views. 424. M. ursus sp. nov., holotype, ♀ (NME). 425. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 426. M. weigeli sp. nov., holotype, ♂ (NME). 427. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 428. M. yamdena sp. nov., paratype, ♂ (BMNH). 429. M.dani sp. nov., holotype, ♂ (IECA). 430. Same, different aspect (IECA). 431. M. hamatus sp. nov., holotype, ♂ (IECA). 432. Same, different aspect (IECA). 433. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 434. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). 435. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122216" httpUri="https://zenodo.org/record/4122216/files/figure.png" pageId="125" pageNumber="126">Fig. 426</figureCitation>
). Male protibial terminal spur strongly curved, female one shorter and nearly straight. Male lower meso- and metatibial terminal spurs curved, rather short. Male aedeagus as in
<figureCitation id="0AA52A659D619226FDD5F990CABD2E2A" box="[582,747,1582,1608]" captionStart="Figs 739750" captionStartId="181.[189,241,1892,1918]" captionTargetBox="[360,1207,271,1867]" captionTargetId="figure-66@181.[347,1240,265,1871]" captionTargetPageId="181" captionText="Figs 739750. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 739 741. M. weigeli sp. nov., paratype, ♂ (NME). 742744. M. woodlarkianus Heller, 1914, ♂ (SNSD). 745747. M. yamdena sp. nov., paratype, ♂ (BMNH). 748750. M. paupianus Heller, 1910 (paratype ♂ of M. arrowi Frey, 1969) (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122256" httpUri="https://zenodo.org/record/4122256/files/figure.png" pageId="125" pageNumber="126">Figs 739741</figureCitation>
.
</paragraph>
<paragraph id="922136E09D619226FF2EF9CEC9E62EE8" blockId="125.[189,432,1648,1674]" box="[189,432,1648,1674]" pageId="125" pageNumber="126">
<heading id="C969818C9D619226FF2EF9CEC9E62EE8" bold="true" box="[189,432,1648,1674]" fontSize="11" level="3" pageId="125" pageNumber="126" reason="3">
<emphasis id="A0EAEAF29D619226FF2EF9CEC9E62EE8" bold="true" box="[189,432,1648,1674]" pageId="125" pageNumber="126">Sexual dimorphism</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D619226FF2EF91ECAEE2EBD" blockId="125.[189,1398,1696,1759]" pageId="125" pageNumber="126">
Female lamellae of antennal club shorter; anterior margin of labroclypeus nearly subtruncate, shallower (
<figureCitation id="0AA52A659D619226FF55F97BC97B2EBD" box="[198,301,1733,1759]" captionStart="Figs 181192" captionStartId="144.[189,241,1787,1813]" captionTargetBox="[338,1249,274,1742]" captionTargetId="figure-122@144.[338,1249,265,1749]" captionTargetPageId="144" captionText="Figs 181192. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal view. 181. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 182. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). 183. Same, ♀ (SNSD). 184. M. suwawa sp. nov., holotype, ♂ (BMNH). 185. Same, paratype, ♀ (BMNH). 186. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 187. M. trivialis sp. nov., holotype, ♂ (BMNH). 188. M. ursus sp. nov., holotype, ♀ (NME). 189. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 190. Same, ♂ (SNSD). 191. M. weigeli sp. nov., holotype, ♂ (NME). 192. Same, paratype, ♀ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122187" httpUri="https://zenodo.org/record/4122187/files/figure.png" pageId="125" pageNumber="126">Fig. 192</figureCitation>
); tibial terminal spurs not curved.
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D619226FF2EF8B6CBC52F33" box="[189,915,1800,1874]" pageId="125" pageNumber="126" type="biology_ecology">
<paragraph id="922136E09D619226FF2EF8B6C9762F40" blockId="125.[189,288,1800,1826]" box="[189,288,1800,1826]" pageId="125" pageNumber="126">
<heading id="C969818C9D619226FF2EF8B6C9762F40" bold="true" box="[189,288,1800,1826]" fontSize="11" level="3" pageId="125" pageNumber="126" reason="3">
<emphasis id="A0EAEAF29D619226FF2EF8B6C9762F40" bold="true" box="[189,288,1800,1826]" pageId="125" pageNumber="126">Ecology</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D619226FF2EF889CBC52F33" blockId="125.[189,915,1847,1874]" box="[189,915,1847,1874]" pageId="125" pageNumber="126">
Occurs in lower montane rainforests at about
<quantity id="55669B059D619226FD47F889CB7A2F30" box="[724,812,1847,1874]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="125" pageNumber="126" unit="m" value="1150.0">1150 m</quantity>
altitude.
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D619226FF2EF8C4CA4F2F8A" pageId="125" pageNumber="126" type="distribution">
<paragraph id="922136E09D619226FF2EF8C4C9002FF6" blockId="125.[189,342,1914,1940]" box="[189,342,1914,1940]" pageId="125" pageNumber="126">
<heading id="C969818C9D619226FF2EF8C4C9002FF6" bold="true" box="[189,342,1914,1940]" fontSize="11" level="3" pageId="125" pageNumber="126" reason="3">
<emphasis id="A0EAEAF29D619226FF2EF8C4C9002FF6" bold="true" box="[189,342,1914,1940]" pageId="125" pageNumber="126">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D619226FF2EF814CA4F2F8A" blockId="125.[189,1398,1962,2025]" pageId="125" pageNumber="126">
Hitherto known from the westernmost part of the Central Cordillera (Nabire surroundings; Wapoga River valley) of New
<collectingCountry id="EA8976709D619226FE2EF870CA402F8A" box="[445,534,1998,2024]" name="Guinea-Bissau" pageId="125" pageNumber="126">Guinea</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>