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<document id="FC538ABA78EC693EA210AC34ACB39429" ID-CLB-Dataset="9693" ID-DOI="10.5852/ejt.2020.721.1127" ID-GBIF-Dataset="0cd61b23-1089-4c1b-bfc0-487887abad84" ID-Zenodo-Dep="4122118" ID-ZooBank="89E62EF8-2E45-4C59-94B7-6A5603E8939B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1603474447689" checkinUser="valdenar" docAuthor="Telnov, Dmitry" docDate="2020" docId="1A3787F69D459202FDA3FEB6C9FD2F47" docLanguage="en" docName="ejt-721_telnov.pdf" docOrigin="European Journal of Taxonomy 721" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Maechidius parallelicollis Moser 1920" docType="treatment" docVersion="10" lastPageNumber="90" masterDocId="E60EFF8E9D1C925BFF93FFBEC8562862" masterDocTitle="A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line" masterLastPageNumber="210" masterPageNumber="1" pageNumber="90" updateTime="1698846101371" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo id="FB33DB0BF98112395B5247DDEB5212B7">
<mods:title id="1F26DED4A7B342C5BDBB0989AA58B3B2">A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line</mods:title>
</mods:titleInfo>
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<mods:namePart id="E0764528B7D0C298DAFD628296EA66DE">Telnov, Dmitry</mods:namePart>
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<mods:title id="D649DC2B156EC71DFF2FF15E9BDDAF98">European Journal of Taxonomy</mods:title>
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<mods:date id="A92083103B114E12E6857566806CA646">2020</mods:date>
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<mods:number id="AA18879D0F47759769639F0C4894D4E4">2020-10-19</mods:number>
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<mods:number id="6C475EA97526C214727FB146149230EF">721</mods:number>
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<mods:classification id="DBDC387AA94F36F9F08B20D231139627">journal article</mods:classification>
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<mods:identifier id="F4E9F4C22C838E0CCCD38005FFB82FD6" type="DOI">10.5852/ejt.2020.721.1127</mods:identifier>
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<treatment id="1A3787F69D459202FDA3FEB6C9FD2F47" ID-DOI="http://doi.org/10.5281/zenodo.4344453" ID-GBIF-Taxon="168648153" ID-Zenodo-Dep="4344453" LSID="urn:lsid:plazi:treatment:1A3787F69D459202FDA3FEB6C9FD2F47" httpUri="http://treatment.plazi.org/id/1A3787F69D459202FDA3FEB6C9FD2F47" lastPageNumber="90" pageId="89" pageNumber="90">
<subSubSection id="DA84656B9D459202FDA3FEB6CC522941" box="[560,1028,264,291]" pageId="89" pageNumber="90" type="nomenclature">
<paragraph id="922136E09D459202FDA3FEB6CC522941" blockId="89.[560,1028,264,329]" box="[560,1028,264,291]" pageId="89" pageNumber="90">
<heading id="C969818C9D459202FDA3FEB6CC522941" box="[560,1028,264,291]" centered="true" fontSize="11" level="2" pageId="89" pageNumber="90" reason="2">
<taxonomicName id="559E4D639D459202FDA3FEB6CC522941" ID-CoL="3XFKL" authority="Moser, 1920" authorityName="Moser" authorityYear="1920" box="[560,1028,264,291]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="parallelicollis">
<emphasis id="A0EAEAF29D459202FDA3FEB6CB3C2941" bold="true" box="[560,874,264,291]" italics="true" pageId="89" pageNumber="90">Maechidius parallelicollis</emphasis>
<bibRefCitation id="F60F4B119D459202FCE2FEB7CC522941" author="Moser J." box="[881,1028,265,291]" pageId="89" pageNumber="90" pagination="3 - 28" refId="ref98398" refString="Moser J. 1920. Beitrag zur Kenntnis der Melolonthiden XI. Stettiner Entomologische Zeitung 81: 3 - 28." type="journal article" year="1920">Moser, 1920</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D459202FDEAFE90CBED292B" box="[633,955,302,329]" pageId="89" pageNumber="90" type="description">
<paragraph id="922136E09D459202FDEAFE90CBED292B" blockId="89.[560,1028,264,329]" box="[633,955,302,329]" pageId="89" pageNumber="90">
<figureCitation id="0AA52A659D459202FDEAFE90CA85292A" box="[633,723,302,328]" captionStart="Figs 66–69" captionStartId="86.[189,241,1892,1918]" captionTargetBox="[360,1220,270,1856]" captionTargetId="figure-64@86.[356,1231,265,1862]" captionTargetPageId="86" captionText="Figs 66–69. Papuan Maechidius Macleay, 1819, habitus, dorsal view. 66. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 67. M. paupianus Heller, 1910, lectotype, ♀ (SNSD). 68. Same, paratype, ♂ (M. arrowi Frey, 1969) (BMNH). 69. M. parallelicollis Moser, 1920, lectotype, ♂ (ZMHB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122156" httpUri="https://zenodo.org/record/4122156/files/figure.png" pageId="89" pageNumber="90">Figs 69</figureCitation>
,
<figureCitation id="0AA52A659D459202FD4CFE90CB5B292B" box="[735,781,302,329]" captionStart="Figs 157–168" captionStartId="142.[189,241,1752,1778]" captionTargetBox="[338,1248,270,1715]" captionTargetId="figure-140@142.[338,1249,265,1724]" captionTargetPageId="142" captionText="Figs 157–168. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal (157–159, 161–167) and frontal (160, 168) views. 157. M. parallelicollis Moser, 1920, lectotype, ♂ (ZMHB). 158. M. paupianus Heller, 1910, lectotype, ♀ (SNSD). 159. Same, ♂ (paratype of M. arrowi Frey, 1969) (BMNH). 160. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 161. Same, ♀ (SNSD). 162. M. peregrinus Lansberge, 1886, ♂ (DTC). 163. M. pedarioides Arrow, 1941, syntype, ♀ (BMNH). 164. Same, syntype, ♂ (BMNH). 165. M. riedeli sp. nov., holotype, ♂ (SMNS). 166. M. perlatus (Frey, 1969) comb. nov., holotype, ♂ (NHMB). 167. Same, ♀ (NHMB). 168. M. simplex Frey, 1969, holotype, ♂ (photograph courtesy BPBM) (BPBM). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122183" httpUri="https://zenodo.org/record/4122183/files/figure.png" pageId="89" pageNumber="90">157</figureCitation>
,
<figureCitation id="0AA52A659D459202FC8AFE90CB1E292A" box="[793,840,302,328]" captionStart="Figs 226–241" captionStartId="147.[189,241,1681,1707]" captionTargetBox="[391,1192,268,1647]" captionTargetId="figure-170@147.[391,1196,265,1656]" captionTargetPageId="147" captionText="Figs 226–241. Papuan and Wallacean Maechidius Macleay, 1819, pronotum, dorsal view. 226. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 227. M. lobaticeps Frey, 1969, ♂ (DTC). 228. M. longipes sp. nov., holotype, ♂ (BMNH). 229. M. luniceps Fairmaire, 1883, ♂ (BMNH). 230. M. mailu sp. nov., holotype, ♂ (NHMB). 231. M. maleo sp. nov., holotype, ♂ (IRSN). 232. M. merdeka sp. nov., holotype, ♂ (BMNH). 233. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 234. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 235. M. muticus Arrow, 1941, syntype, ♂ (BMNH). 236. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 237. M. nepenthephilus sp. nov., holotype, ♂ (NME). 238. M. opatroides Arrow, 1941 comb. rest., lectotype, ♀ (BMNH). 239. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 240. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 241. M. parallelicollis Moser, 1920, lectotype, ♂ (ZMHB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122193" httpUri="https://zenodo.org/record/4122193/files/figure.png" pageId="89" pageNumber="90">241</figureCitation>
,
<figureCitation id="0AA52A659D459202FCC7FE90CBD5292A" box="[852,899,302,328]" captionStart="Figs 317–332" captionStartId="154.[189,241,1681,1707]" captionTargetBox="[401,1187,265,1655]" captionTargetId="figure-172@154.[401,1187,265,1669]" captionTargetPageId="154" captionText="Figs 317–332. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 317. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 318. M. lobaticeps Frey, 1969, ♂ (DTC). 319. M. longipes sp. nov., holotype, ♂ (BMNH). 320. M. luniceps Fairmaire, 1883, ♂ (BMNH). 321. M. mailu sp. nov., holotype, ♂ (NHMB). 322. M. maleo sp. nov., holotype, ♂ (IRSN). 323. M. merdeka sp. nov., holotype, ♂ (BMNH). 324. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 325. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 326. M. muticusArrow, 1941, syntype, ♂ (BMNH). 327. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 328. M. nepenthephilus sp. nov., holotype, ♂ (NME). 329. M. opatroides Arrow, 1941 comb. rest., lectotype, ♀ (BMNH). 330. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 331. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 332. M. parallelicollis Moser, 1920, lectotype, ♂ (ZMHB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122210" httpUri="https://zenodo.org/record/4122210/files/figure.png" pageId="89" pageNumber="90">332</figureCitation>
,
<figureCitation id="0AA52A659D459202FC1CFE90CBED292B" box="[911,955,302,329]" captionStart="Figs487–503" captionStartId="165.[189,241,1681,1707]" captionTargetBox="[422,1160,265,1655]" captionTargetId="figure-171@165.[421,1167,265,1658]" captionTargetPageId="165" captionText="Figs487–503.Papuan andWallaceanMaechidius Macleay, 1819,pygidium, dorsalview.487.M.lobaticeps Frey, 1969, ♂ (DTC). 488. M. longipes sp. nov., holotype, ♂ (BMNH). 489. M. luniceps Fairmaire, 1883, ♂ (BMNH). 490. M. mailu sp. nov., holotype, ♂ (NHMB). 491. M. maleo sp. nov., holotype, ♂ (IRSN). 492. M. merdeka sp. nov., holotype, ♂ (BMNH). 493. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 494. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 495. M. muticus Arrow, 1941, syntype, ♂ (BMNH). 496. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 497. M. nepenthephilus sp. nov., holotype, ♂ (NME). 498. M. opatroidesArrow, 1941 comb. rest., lectotype, ♀ (BMNH). 499. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 500. Same, paratype, ♀ (BMNH). 501. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 502. Same, different aspect. 503. M. parallelicollis Moser, 1920, lectotype, ♂ (ZMHB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122226" httpUri="https://zenodo.org/record/4122226/files/figure.png" pageId="89" pageNumber="90">503</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D459202FF2EFECCC97C2A4F" pageId="89" pageNumber="90" type="materials_examined">
<paragraph id="922136E09D459202FF2EFECCC93A29E9" blockId="89.[189,364,369,396]" box="[189,364,369,396]" pageId="89" pageNumber="90">
<heading id="C969818C9D459202FF2EFECCC93A29E9" bold="true" box="[189,364,369,396]" fontSize="11" level="3" pageId="89" pageNumber="90" reason="3">
<emphasis id="A0EAEAF29D459202FF2EFECCC93A29E9" bold="true" box="[189,364,369,396]" pageId="89" pageNumber="90">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D459202FF79FE1CCA3A29DF" blockId="89.[189,1399,418,557]" box="[234,620,418,445]" pageId="89" pageNumber="90">
<emphasis id="A0EAEAF29D459202FF79FE1CC93029DE" bold="true" box="[234,358,418,444]" pageId="89" pageNumber="90">
<typeStatus id="4D2588429D459202FF79FE1CC93029DE" box="[234,358,418,444]" pageId="89" pageNumber="90" type="lectotype">Lectotype</typeStatus>
</emphasis>
[herewith designated]
</paragraph>
<paragraph id="922136E09D459202FF2EFE76C97C2A4F" blockId="89.[189,1399,418,557]" pageId="89" pageNumber="90">
<materialsCitation id="22F63CBD9D459202FF2EFE76C97C2A4F" ID-GBIF-Occurrence="2898878616" collectionCode="ZMHB" country="Indonesia" location="INDONESIA" pageId="89" pageNumber="90" specimenCode="MFNB 2019" specimenCount="1" typeStatus="syntype">
<collectingCountry id="EA8976709D459202FF2EFE76C90D2980" box="[189,347,456,482]" name="Indonesia" pageId="89" pageNumber="90">INDONESIA</collectingCountry>
•
<specimenCount id="8498FD699D459202FE11FE76C9CC2980" box="[386,410,456,482]" pageId="89" pageNumber="90" type="male">♂</specimenCount>
; “Roon [p] //
<taxonomicName id="559E4D639D459202FDD7FE76CB9D2980" authority="Mos." authorityName="Moser" authorityYear="1920" box="[580,971,456,482]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="parallelicollis">
<emphasis id="A0EAEAF29D459202FDD7FE76CBD72980" box="[580,897,456,482]" italics="true" pageId="89" pageNumber="90">Maechidius parallelicollis</emphasis>
Mos.
</taxonomicName>
<typeStatus id="4D2588429D459202FC4BFE76CC442980" box="[984,1042,456,482]" pageId="89" pageNumber="90">Type</typeStatus>
[h] //
<typeStatus id="4D2588429D459202FBC8FE76CC892980" box="[1115,1247,456,482]" pageId="89" pageNumber="90" type="syntype">SYNTYPE</typeStatus>
<taxonomicName id="559E4D639D459202FB7DFE76CA572A65" authority="Moser, 1920" authorityName="Moser" authorityYear="1920" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="parallelicollis">
<emphasis id="A0EAEAF29D459202FB7DFE76C9342A65" italics="true" pageId="89" pageNumber="90">Maechidius parallelicollis</emphasis>
<bibRefCitation id="F60F4B119D459202FEF8FE53CA572A65" author="Moser J." box="[363,513,493,519]" pageId="89" pageNumber="90" pagination="3 - 28" refId="ref98398" refString="Moser J. 1920. Beitrag zur Kenntnis der Melolonthiden XI. Stettiner Entomologische Zeitung 81: 3 - 28." type="journal article" year="1920">Moser, 1920</bibRefCitation>
</taxonomicName>
labelled by
<specimenCode id="C2389E9B9D459202FD0AFE53CB672A65" box="[665,817,493,519]" collectionCode="MFNB" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="89" pageNumber="90">MFNB 2019</specimenCode>
[p, red label]”;
<collectionCode id="F48FAE259D459202FC65FE53CC1D2A65" box="[1014,1099,493,519]" collectionName="Germany, Berlin, Museum fuer Naturkunde der Humboldt-Universitaet" pageId="89" pageNumber="90">ZMHB</collectionCode>
[forelegs are completely missing].
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D459202FF2EFDE8C9022B72" pageId="89" pageNumber="90" type="discussion">
<paragraph id="922136E09D459202FF2EFDE8C9792A12" blockId="89.[189,303,598,624]" box="[189,303,598,624]" pageId="89" pageNumber="90">
<heading id="C969818C9D459202FF2EFDE8C9792A12" bold="true" box="[189,303,598,624]" fontSize="11" level="3" pageId="89" pageNumber="90" reason="3">
<emphasis id="A0EAEAF29D459202FF2EFDE8C9792A12" bold="true" box="[189,303,598,624]" pageId="89" pageNumber="90">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D459202FF2EFD39C9022B72" blockId="89.[189,1399,646,784]" pageId="89" pageNumber="90">
<bibRefCitation id="F60F4B119D459202FF2EFD39C9DA2AC3" author="Moser J." box="[189,396,646,673]" pageId="89" pageNumber="90" pagination="3 - 28" refId="ref98398" refString="Moser J. 1920. Beitrag zur Kenntnis der Melolonthiden XI. Stettiner Entomologische Zeitung 81: 3 - 28." type="journal article" year="1920">Moser (1920: 15)</bibRefCitation>
based his description of
<taxonomicName id="559E4D639D459202FD26FD39CBD42AC2" authorityName="Moser" authorityYear="1920" box="[693,898,646,672]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="parallelicollis">
<emphasis id="A0EAEAF29D459202FD26FD39CBD42AC2" box="[693,898,646,672]" italics="true" pageId="89" pageNumber="90">M. parallelicollis</emphasis>
</taxonomicName>
on an unstated number of specimens. The
<typeStatus id="4D2588429D459202FF2EFD15C97F2AA7" box="[189,297,683,709]" pageId="89" pageNumber="90" type="lectotype">lectotype</typeStatus>
is therefore designated for the single
<typeStatus id="4D2588429D459202FD60FD12CB182AA4" box="[755,846,684,710]" pageId="89" pageNumber="90" type="syntype">syntype</typeStatus>
in ZMHB in order to enhance the stability of nomenclature and fix the specimen I based my redescription on, in case other
<typeStatus id="4D2588429D459202FBCFFD6FCC952A89" box="[1116,1219,721,747]" pageId="89" pageNumber="90" type="syntype">syntypes</typeStatus>
are discovered in the future.
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D459202FF2EFC87C9FD2F47" pageId="89" pageNumber="90" type="description">
<paragraph id="922136E09D459202FF2EFC87C9182B31" blockId="89.[189,334,825,851]" box="[189,334,825,851]" pageId="89" pageNumber="90">
<heading id="C969818C9D459202FF2EFC87C9182B31" bold="true" box="[189,334,825,851]" fontSize="11" level="3" pageId="89" pageNumber="90" reason="3">
<emphasis id="A0EAEAF29D459202FF2EFC87C9182B31" bold="true" box="[189,334,825,851]" pageId="89" pageNumber="90">Description</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D459202FF2EFCD5CB112BE7" blockId="89.[189,839,874,901]" box="[189,839,874,901]" pageId="89" pageNumber="90">
MEASUREMENTS. Total body length
<quantity id="55669B059D459202FDC2FCD4CAEF2BE7" box="[593,697,874,901]" metricMagnitude="-3" metricUnit="m" metricValue="7.45" pageId="89" pageNumber="90" unit="mm" value="7.45">7.45 mm</quantity>
(
<typeStatus id="4D2588429D459202FD5BFCD4CB6D2BE7" box="[712,827,874,901]" pageId="89" pageNumber="90" type="lectotype">lectotype</typeStatus>
).
</paragraph>
<paragraph id="922136E09D459202FF2EFC10CB5C2ED0" blockId="89.[189,1399,942,1714]" pageId="89" pageNumber="90">
Dorsum and venter uniformly brown except for paler appendages and antennae. Head flattened dorsally on frons between eyes, glossy dorsally and ventrally. Male labroclypeus (
<figureCitation id="0AA52A659D459202FBB2FC6DCCD32B8C" box="[1057,1157,979,1006]" captionStart="Figs 157–168" captionStartId="142.[189,241,1752,1778]" captionTargetBox="[338,1248,270,1715]" captionTargetId="figure-140@142.[338,1249,265,1724]" captionTargetPageId="142" captionText="Figs 157–168. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal (157–159, 161–167) and frontal (160, 168) views. 157. M. parallelicollis Moser, 1920, lectotype, ♂ (ZMHB). 158. M. paupianus Heller, 1910, lectotype, ♀ (SNSD). 159. Same, ♂ (paratype of M. arrowi Frey, 1969) (BMNH). 160. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 161. Same, ♀ (SNSD). 162. M. peregrinus Lansberge, 1886, ♂ (DTC). 163. M. pedarioides Arrow, 1941, syntype, ♀ (BMNH). 164. Same, syntype, ♂ (BMNH). 165. M. riedeli sp. nov., holotype, ♂ (SMNS). 166. M. perlatus (Frey, 1969) comb. nov., holotype, ♂ (NHMB). 167. Same, ♀ (NHMB). 168. M. simplex Frey, 1969, holotype, ♂ (photograph courtesy BPBM) (BPBM). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122183" httpUri="https://zenodo.org/record/4122183/files/figure.png" pageId="89" pageNumber="90">Fig. 157</figureCitation>
) shallowly and very broadly emarginate anteriorly, its lateral margins sinuous in both dorsal and lateral views. Anterolateral angles almost right-angled, hardly protruding, raised up at angle of nearly 90–95° to frons in lateral view. Anterior and lateral margins of labroclypeus smooth. Upper- and underside of labroclypeus with sparse moderately long setae along anterior and lateral margins. Canthus slightly sinuous in dorsal view. Punctures of frons large, of irregular shape, annular, shallow and dense. Intervening spaces glossy, distinctly smaller than punctures. Each seta rises from anterior margin of corresponding puncture. Male antenna 9-segmented, club 3-lamellate. Pronotum transverse, glossy dorsally and laterally. Anterior margin of pronotum broadly emarginate with protruding anterolateral angles, basal margin slightly sinuous. Lateral margin of pronotum nearly straight, delicately crenulate all along (about 20 flat crenulae on either side) (
<figureCitation id="0AA52A659D459202FEE6FAF6C98A2D00" box="[373,476,1352,1378]" captionStart="Figs 226–241" captionStartId="147.[189,241,1681,1707]" captionTargetBox="[391,1192,268,1647]" captionTargetId="figure-170@147.[391,1196,265,1656]" captionTargetPageId="147" captionText="Figs 226–241. Papuan and Wallacean Maechidius Macleay, 1819, pronotum, dorsal view. 226. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 227. M. lobaticeps Frey, 1969, ♂ (DTC). 228. M. longipes sp. nov., holotype, ♂ (BMNH). 229. M. luniceps Fairmaire, 1883, ♂ (BMNH). 230. M. mailu sp. nov., holotype, ♂ (NHMB). 231. M. maleo sp. nov., holotype, ♂ (IRSN). 232. M. merdeka sp. nov., holotype, ♂ (BMNH). 233. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 234. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 235. M. muticus Arrow, 1941, syntype, ♂ (BMNH). 236. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 237. M. nepenthephilus sp. nov., holotype, ♂ (NME). 238. M. opatroides Arrow, 1941 comb. rest., lectotype, ♀ (BMNH). 239. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 240. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 241. M. parallelicollis Moser, 1920, lectotype, ♂ (ZMHB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122193" httpUri="https://zenodo.org/record/4122193/files/figure.png" pageId="89" pageNumber="90">Fig. 241</figureCitation>
). Inconspicuous erect seta present between every two crenulae. Lateral margin of pronotum hardly sinuous in lateral view. Hypomeron slightly arched, separated from prosternum by moderately high partly acute carina. Antennal pocket moderately deep. Pronotal punctures larger than those on head, annular, stronger ovoid at lateral sides and base. Setae inconspicuous, erect, hardly surpassing length of corresponding punctures. Scutellar shield narrowly rounded apically. Elytron subopaque, without tracks of longitudinal carinae. Elytral punctures very elongate and narrow, dense, incision-shaped (
<figureCitation id="0AA52A659D459202FE19F999C9A12E20" box="[394,503,1575,1602]" captionStart="Figs 317–332" captionStartId="154.[189,241,1681,1707]" captionTargetBox="[401,1187,265,1655]" captionTargetId="figure-172@154.[401,1187,265,1669]" captionTargetPageId="154" captionText="Figs 317–332. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 317. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 318. M. lobaticeps Frey, 1969, ♂ (DTC). 319. M. longipes sp. nov., holotype, ♂ (BMNH). 320. M. luniceps Fairmaire, 1883, ♂ (BMNH). 321. M. mailu sp. nov., holotype, ♂ (NHMB). 322. M. maleo sp. nov., holotype, ♂ (IRSN). 323. M. merdeka sp. nov., holotype, ♂ (BMNH). 324. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 325. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 326. M. muticusArrow, 1941, syntype, ♂ (BMNH). 327. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 328. M. nepenthephilus sp. nov., holotype, ♂ (NME). 329. M. opatroides Arrow, 1941 comb. rest., lectotype, ♀ (BMNH). 330. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 331. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 332. M. parallelicollis Moser, 1920, lectotype, ♂ (ZMHB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122210" httpUri="https://zenodo.org/record/4122210/files/figure.png" pageId="89" pageNumber="90">Fig. 332</figureCitation>
). Setae inconspicuous, not surpassing length of corresponding punctures, appressed. Male pygidium slightly convex dorsally, densely annularly punctate and with short erect setae (
<figureCitation id="0AA52A659D459202FE9AF9CCC9242EEF" box="[265,370,1650,1677]" captionStart="Figs487–503" captionStartId="165.[189,241,1681,1707]" captionTargetBox="[422,1160,265,1655]" captionTargetId="figure-171@165.[421,1167,265,1658]" captionTargetPageId="165" captionText="Figs487–503.Papuan andWallaceanMaechidius Macleay, 1819,pygidium, dorsalview.487.M.lobaticeps Frey, 1969, ♂ (DTC). 488. M. longipes sp. nov., holotype, ♂ (BMNH). 489. M. luniceps Fairmaire, 1883, ♂ (BMNH). 490. M. mailu sp. nov., holotype, ♂ (NHMB). 491. M. maleo sp. nov., holotype, ♂ (IRSN). 492. M. merdeka sp. nov., holotype, ♂ (BMNH). 493. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 494. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 495. M. muticus Arrow, 1941, syntype, ♂ (BMNH). 496. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 497. M. nepenthephilus sp. nov., holotype, ♂ (NME). 498. M. opatroidesArrow, 1941 comb. rest., lectotype, ♀ (BMNH). 499. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 500. Same, paratype, ♀ (BMNH). 501. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 502. Same, different aspect. 503. M. parallelicollis Moser, 1920, lectotype, ♂ (ZMHB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122226" httpUri="https://zenodo.org/record/4122226/files/figure.png" pageId="89" pageNumber="90">Fig. 503</figureCitation>
). Male metatibial terminal spurs long, subequal in length. Tarsal claws with pulvilli at least on middle and posterior legs (in male only?).
</paragraph>
<paragraph id="922136E09D459202FF2EF964C9E62E96" blockId="89.[189,432,1754,1780]" box="[189,432,1754,1780]" pageId="89" pageNumber="90">
<heading id="C969818C9D459202FF2EF964C9E62E96" bold="true" box="[189,432,1754,1780]" fontSize="11" level="3" pageId="89" pageNumber="90" reason="3">
<emphasis id="A0EAEAF29D459202FF2EF964C9E62E96" bold="true" box="[189,432,1754,1780]" pageId="89" pageNumber="90">Sexual dimorphism</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D459202FF2EF8B5C9FD2F47" blockId="89.[189,427,1803,1829]" box="[189,427,1803,1829]" pageId="89" pageNumber="90">Female is unknown.</paragraph>
</subSubSection>
</treatment>
</document>
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