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<document id="16CA1C054C8815AA5B2BD0FADCD1A896" ID-CLB-Dataset="9693" ID-DOI="10.5852/ejt.2020.721.1127" ID-GBIF-Dataset="0cd61b23-1089-4c1b-bfc0-487887abad84" ID-Zenodo-Dep="4122118" ID-ZooBank="89E62EF8-2E45-4C59-94B7-6A5603E8939B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1603474447689" checkinUser="valdenar" docAuthor="Telnov, Dmitry" docDate="2020" docId="1A3787F69D349272FDFCFC75CAE12F8A" docLanguage="en" docName="ejt-721_telnov.pdf" docOrigin="European Journal of Taxonomy 721" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Maechidius deltouri Telnov 2020, sp. nov." docType="treatment" docUuid="9F79D2BF-8ADF-4856-9400-2AFCA33DB0D1" docUuidSource="ZooBank" docVersion="10" lastPageNumber="42" masterDocId="E60EFF8E9D1C925BFF93FFBEC8562862" masterDocTitle="A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line" masterLastPageNumber="210" masterPageNumber="1" pageNumber="41" updateTime="1698846101371" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="8BEAA582FA082497CB5B190C1696E8C1" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="433215C3ECB60DE752C148F6250695B3">
<mods:title id="48C242D666E6FA81DC63AB3696C01080">A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line</mods:title>
</mods:titleInfo>
<mods:name id="D356784CC278882CBF149C24277D6051" type="personal">
<mods:role id="E90CEB2EFA0DDB008819952416D90176">
<mods:roleTerm id="47F7CEB89940DECC50E6C947CD498D2D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="19FEBBB40F88350ED62B1F35AA77ADDC">Telnov, Dmitry</mods:namePart>
</mods:name>
<mods:typeOfResource id="A2CAD2657A918C4D618F2E8E457FD7AC">text</mods:typeOfResource>
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<mods:titleInfo id="3A1A8141A90F1F9B8031824EF066705D">
<mods:title id="5F2055C1B784AE7B19329A8127649CED">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="F0989637BCE6D8F6F261AC1DAD352C86">
<mods:date id="02223BBDE011662713837D056A412265">2020</mods:date>
<mods:detail id="466434BDDE78A8ACED3F5F555765EFB4" type="pubDate">
<mods:number id="62BB5E30FBB883A3AE5C9F82F1A71E45">2020-10-19</mods:number>
</mods:detail>
<mods:detail id="F845D88A5C7188918C0A97FA809ED84F" type="volume">
<mods:number id="FA6A70878C86554BCBD224AA97459341">721</mods:number>
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<mods:classification id="D3789A3C8886A8E09D040FDD12E111A5">journal article</mods:classification>
<mods:identifier id="7574A62871966313B282622B47C0961B" type="CLB-Dataset">9693</mods:identifier>
<mods:identifier id="F1CD71143F25416E30328D08C7FAA7BB" type="DOI">10.5852/ejt.2020.721.1127</mods:identifier>
<mods:identifier id="F8E15AA9F323CF35B69DA2FB4818929A" type="GBIF-Dataset">0cd61b23-1089-4c1b-bfc0-487887abad84</mods:identifier>
<mods:identifier id="D27A5F8B2BBAC6C1D2DF9446C931A7D1" type="Zenodo-Dep">4122118</mods:identifier>
<mods:identifier id="5349F0349AF5D6F4DBD64FCF2D7118A1" type="ZooBank">89E62EF8-2E45-4C59-94B7-6A5603E8939B</mods:identifier>
</mods:mods>
<treatment id="1A3787F69D349272FDFCFC75CAE12F8A" ID-DOI="http://doi.org/10.5281/zenodo.4344359" ID-GBIF-Taxon="168648145" ID-Zenodo-Dep="4344359" LSID="urn:lsid:zoobank.org:act:9F79D2BF-8ADF-4856-9400-2AFCA33DB0D1" httpUri="http://treatment.plazi.org/id/1A3787F69D349272FDFCFC75CAE12F8A" lastPageId="41" lastPageNumber="42" pageId="40" pageNumber="41">
<subSubSection id="DA84656B9D349273FDFCFC75CB932B84" box="[623,965,971,998]" pageId="40" pageNumber="41" type="nomenclature">
<paragraph id="922136E09D349273FDFCFC75CB932B84" blockId="40.[359,1228,971,1073]" box="[623,965,971,998]" pageId="40" pageNumber="41">
<heading id="C969818C9D349273FDFCFC75CB932B84" box="[623,965,971,998]" centered="true" fontSize="11" level="2" pageId="40" pageNumber="41" reason="2">
<taxonomicName id="559E4D639D349273FDFCFC75CB352B84" ID-CoL="7X3WM" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[623,867,971,998]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="deltouri" status="sp. nov.">
<emphasis id="A0EAEAF29D349273FDFCFC75CB352B84" bold="true" box="[623,867,971,998]" italics="true" pageId="40" pageNumber="41">Maechidius deltouri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D349273FCF9FC72CB932B84" box="[874,965,972,998]" pageId="40" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D349273FEF4FC4FCC792C53" pageId="40" pageNumber="41" type="description">
<paragraph id="922136E09D349273FEF4FC4FCC9A2C6E" blockId="40.[359,1228,971,1073]" box="[359,1228,1009,1036]" pageId="40" pageNumber="41">
<uri id="E60F3AE29D349273FEF4FC4FCC9A2C6E" box="[359,1228,1009,1036]" pageId="40" pageNumber="41">
urn:lsid:zoobank.org:act:
<uuid id="E6380C359D349273FD15FC4FCC9A2C6E" box="[646,1228,1009,1036]" pageId="40" pageNumber="41">9F79D2BF-8ADF-4856-9400-2AFCA33DB0D1</uuid>
</uri>
</paragraph>
<paragraph id="922136E09D349273FD96FBA9CC792C53" blockId="40.[359,1228,971,1073]" box="[517,1071,1046,1073]" pageId="40" pageNumber="41">
<figureCitation id="0AA52A659D349273FD96FBA9CA082C53" box="[517,606,1047,1073]" captionStart="Figs 31–34" captionStartId="36.[189,241,1928,1954]" captionTargetBox="[337,1255,261,1900]" captionTargetId="figure-64@36.[325,1262,257,1921]" captionTargetPageId="36" captionText="Figs 31–34. Papuan and Wallacean Maechidius Macleay, 1819, habitus, dorsal view. 31. M. ciliatus sp. nov., holotype, ♂ (IECA). 32. M. crypticus sp. nov., paratype, ♂ (BMNH). 33. M. dani sp. nov., holotype, ♂ (IECA). 34. M. deltouri sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122138" httpUri="https://zenodo.org/record/4122138/files/figure.png" pageId="40" pageNumber="41">Figs 34</figureCitation>
,
<figureCitation id="0AA52A659D349273FDF9FBA8CAC12C53" box="[618,663,1046,1073]" captionStart="Figs 109–120" captionStartId="138.[189,241,1750,1776]" captionTargetBox="[365,1214,275,1722]" captionTargetId="figure-15@138.[365,1222,265,1730]" captionTargetPageId="138" captionText="Figs 109–120. Papuan andWallacean MaechidiusMacleay, 1819,head, dorsal (109–119) and frontal (120) views. 109. M. brocki sp. nov., holotype, ♂ (BMNH). 110. Same, paratype, ♀ (BMNH). 111. M. ciliatus sp. nov., holotype, ♂ (IECA). 112. M. crypticus sp. nov., paratype, ♂ (BMNH). 113. Same, paratype, ♀ (BMNH). 114. M. dani sp. nov., holotype, ♂ (IECA). 115. M. deltouri sp. nov., holotype, ♂ (BMNH). 116. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 117. M. fraterculus Moser, 1920, lectotype, ♀ (ZMHB). 118. M. esau Heller, 1914, lectotype, ♂ (SNSD). 119. Same, ♂ (lectotype of M. setosus Moser, 1920) (ZMHB). 120. M. gressitti Frey, 1969, holotype, ♂ (photograph courtesy BPBM) (BPBM). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122175" httpUri="https://zenodo.org/record/4122175/files/figure.png" pageId="40" pageNumber="41">115</figureCitation>
,
<figureCitation id="0AA52A659D349273FD37FBA9CA842C53" box="[676,722,1047,1073]" captionStart="Figs 208–225" captionStartId="146.[189,241,1644,1670]" captionTargetBox="[414,1187,265,1610]" captionTargetId="figure-199@146.[414,1187,265,1618]" captionTargetPageId="146" captionText="Figs 208–225. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, pronotum, dorsal view. 208. M.caperatus sp. nov., holotype, ♂ (NME). 209. M.ciliatus sp. nov., holotype, ♂ (IECA). 210. M.dani sp. nov., holotype, ♂ (IECA). 211. M. deltouri sp. nov., holotype, ♂ (BMNH). 212. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 213. M. esau Heller, 1914 (lectotype, ♂ of M. setosus Moser, 1920) (ZMHB). 214. M. fraterculus Moser, 1920, lectotype, ♀ (ZMHB). 215. M. hamatus sp. nov., holotype, ♂ (IECA). 216. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 217. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 218. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 219. M. interruptocarinulatus Heller, 1914, holotype, ♀ (SNSD). 220. M. jobiensis Moser, 1920, lectotype, ♂ (ZMHB). 221. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 222. M. konjo sp. nov., holotype, ♂ (NMPC). 223. M. lapsus sp. nov., holotype, ♂ (BMNH). 224. M. legalovi sp. nov., holotype, ♂ (BMNH). 225. M. leucopsar sp. nov., holotype, ♂ (IECA). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122191" httpUri="https://zenodo.org/record/4122191/files/figure.png" pageId="40" pageNumber="41">211</figureCitation>
,
<figureCitation id="0AA52A659D349273FD4DFBA9CB5B2C53" box="[734,781,1047,1073]" captionStart="Figs 286–301" captionStartId="152.[189,241,1716,1742]" captionTargetBox="[364,1224,266,1689]" captionTargetId="figure-172@152.[364,1224,266,1695]" captionTargetPageId="152" captionText="Figs 286–301. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 286. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 287. M. aiyura sp. nov., holotype, ♂ (BMNH). 288. M. alesbezdeki sp. nov., holotype, ♂ (NME). 289. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 290. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 291. M. paupianus Heller, 1910, ♂ (BMNH). 292. M. awu sp. nov., holotype, ♂ (IECA). 293. M. babyrousa sp. nov., holotype, ♂ (BMNH). 294. M. bintang sp. nov., holotype, ♂ (NME). 295. M. boessnecki sp. nov., holotype, ♂ (SNSD). 296. M. brocki sp. nov., holotype, ♂ (BMNH). 297. M. caperatus sp. nov., holotype, ♂ (NME). 298. M. ciliatus sp. nov., holotype, ♂ (IECA). 299. M. crypticus sp. nov., paratype, ♀ (BMNH). 300. M. dani sp. nov., holotype, ♂ (IECA). 301. M. deltouri sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122206" httpUri="https://zenodo.org/record/4122206/files/figure.png" pageId="40" pageNumber="41">301</figureCitation>
, 374, 468, 542, 607–610
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D349273FF2EFBE4C9892D7A" pageId="40" pageNumber="41" type="diagnosis">
<paragraph id="922136E09D349273FF2EFBE4C99F2C16" blockId="40.[189,457,1114,1140]" box="[189,457,1114,1140]" pageId="40" pageNumber="41">
<heading id="C969818C9D349273FF2EFBE4C99F2C16" bold="true" box="[189,457,1114,1140]" fontSize="11" level="3" pageId="40" pageNumber="41" reason="3">
<emphasis id="A0EAEAF29D349273FF2EFBE4C99F2C16" bold="true" box="[189,457,1114,1140]" pageId="40" pageNumber="41">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D349273FF2EFB32C9892D7A" blockId="40.[189,1399,1164,1304]" pageId="40" pageNumber="41">
This species is most similar to
<taxonomicName id="559E4D639D349273FDB2FB33CB1F2CC4" authority="Lansberge" authorityName="Lansberge" authorityYear="1886" box="[545,841,1164,1190]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="peregrinus">
<emphasis id="A0EAEAF29D349273FDB2FB33CA932CC4" box="[545,709,1165,1190]" italics="true" pageId="40" pageNumber="41">M. peregrinus</emphasis>
Lansberge
</taxonomicName>
,
<taxonomicName id="559E4D639D349273FCC0FB33CBA22CC4" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[851,1012,1164,1190]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="babyrousa" status="sp. nov.">
<emphasis id="A0EAEAF29D349273FCC0FB33CBA22CC4" box="[851,1012,1164,1190]" italics="true" pageId="40" pageNumber="41">M. babyrousa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D349273FC69FB32CC012CC4" box="[1018,1111,1164,1190]" pageId="40" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="559E4D639D349273FBF2FB33CCB12CC4" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[1121,1255,1164,1190]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="legalovi" status="sp. nov.">
<emphasis id="A0EAEAF29D349273FBF2FB33CCB12CC4" box="[1121,1255,1164,1190]" italics="true" pageId="40" pageNumber="41">M. legalovi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D349273FB7FFB32CD102CC4" box="[1260,1350,1164,1190]" pageId="40" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="559E4D639D349273FF2EFB0CC9132CA9" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[189,325,1202,1227]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="suwawa" status="sp. nov.">
<emphasis id="A0EAEAF29D349273FF2EFB0CC9132CA9" box="[189,325,1202,1227]" italics="true" pageId="40" pageNumber="41">M. suwawa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D349273FEDDFB0CC9FC2CAE" box="[334,426,1202,1228]" pageId="40" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
(all from
<collectingRegion id="505AF8029D349273FDB5FB0FCAC32CAE" box="[550,661,1201,1228]" country="Indonesia" name="Sulawesi Utara" pageId="40" pageNumber="41">Sulawesi</collectingRegion>
). It readily differs from them and other congeners only in the shape of the male genital organs. Two vague circular impressions on the pronotal disc are also present in
<taxonomicName id="559E4D639D349273FF48FB40C92B2D75" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[219,381,1277,1303]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="babyrousa" status="sp. nov.">
<emphasis id="A0EAEAF29D349273FF48FB40C92B2D75" box="[219,381,1277,1303]" italics="true" pageId="40" pageNumber="41">M. babyrousa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D349273FE17FB40C9892D7A" box="[388,479,1278,1304]" pageId="40" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D349273FF2EFAFFCBA52DD0" pageId="40" pageNumber="41" type="etymology">
<paragraph id="922136E09D349273FF2EFAFFC9122D39" blockId="40.[189,324,1345,1371]" box="[189,324,1345,1371]" pageId="40" pageNumber="41">
<heading id="C969818C9D349273FF2EFAFFC9122D39" bold="true" box="[189,324,1345,1371]" fontSize="11" level="3" pageId="40" pageNumber="41" reason="3">
<emphasis id="A0EAEAF29D349273FF2EFAFFC9122D39" bold="true" box="[189,324,1345,1371]" pageId="40" pageNumber="41">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D349273FF2EFACCCBA52DD0" blockId="40.[189,1399,1394,1458]" pageId="40" pageNumber="41">
Patronymic. This species is named after Gaёtan Deltour (Naturevolution, La Rochelle,
<collectingCountry id="EA8976709D349273FB6AFACDCD1D2DEF" box="[1273,1355,1395,1421]" name="France" pageId="40" pageNumber="41">France</collectingCountry>
) to commemorate his efforts in the conservation of Sulawesi biodiversity.
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D349273FF2EFA62CCA72E11" pageId="40" pageNumber="41" type="materials_examined">
<paragraph id="922136E09D349273FF2EFA62C93A2D94" blockId="40.[189,364,1500,1526]" box="[189,364,1500,1526]" pageId="40" pageNumber="41">
<heading id="C969818C9D349273FF2EFA62C93A2D94" bold="true" box="[189,364,1500,1526]" fontSize="11" level="3" pageId="40" pageNumber="41" reason="3">
<emphasis id="A0EAEAF29D349273FF2EFA62C93A2D94" bold="true" box="[189,364,1500,1526]" pageId="40" pageNumber="41">Type material</emphasis>
</heading>
</paragraph>
<materialsCitation id="22F63CBD9D349273FF79F9B3CCA72E11" ID-GBIF-Occurrence="2898878520" collectingDate="1985-05-18" collectingDateMax="1985-05-24" collectingDateMin="1985-05-18" collectingMethod="Yellow pan trap" collectionCode="BMNH" country="Indonesia" location="Kotamobagu" pageId="40" pageNumber="41" specimenCount="1" specimenCount-male="1" stateProvince="SULAWESI UTARA" typeStatus="holotype">
<paragraph id="922136E09D349273FF79F9B3C90A2E45" blockId="40.[189,1399,1549,1651]" box="[234,348,1549,1575]" pageId="40" pageNumber="41">
<emphasis id="A0EAEAF29D349273FF79F9B3C90A2E45" bold="true" box="[234,348,1549,1575]" pageId="40" pageNumber="41">
<typeStatus id="4D2588429D349273FF79F9B3C90A2E45" box="[234,348,1549,1575]" pageId="40" pageNumber="41" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="922136E09D349273FF2EF98DCCA72E11" blockId="40.[189,1399,1549,1651]" pageId="40" pageNumber="41">
<collectingCountry id="EA8976709D349273FF2EF98DC90D2E2F" box="[189,347,1587,1613]" name="Indonesia" pageId="40" pageNumber="41">INDONESIA</collectingCountry>
•
<specimenCount id="8498FD699D349273FEE7F98DC9DA2E2F" box="[372,396,1587,1613]" pageId="40" pageNumber="41" type="male">♂</specimenCount>
; “
<collectingCountry id="EA8976709D349273FE30F98DCA122E2F" box="[419,580,1587,1613]" name="Indonesia" pageId="40" pageNumber="41">INDONESIA</collectingCountry>
<collectingRegion id="505AF8029D349273FDDFF98DCB102E2F" box="[588,838,1587,1613]" country="Indonesia" name="Sulawesi Utara" pageId="40" pageNumber="41">SULAWESI UTARA</collectingRegion>
Gng. Ambang F.R. nr.
<location id="9741603B9D349273FBC5F98DCCB82E2F" LSID="urn:lsid:plazi:treatment:1A3787F69D349272FDFCFC75CAE12F8A:9741603B9D349273FBC5F98DCCB82E2F" box="[1110,1262,1587,1613]" country="Indonesia" name="Kotamobagu" pageId="40" pageNumber="41" stateProvince="SULAWESI UTARA">Kotamobagu</location>
<date id="E62010209D349273FB66F98DC8AD2E11" pageId="40" pageNumber="41" value="1985-05-18" valueMax="1985-05-24" valueMin="1985-05-18">
<collectingDate id="F664E9C89D349273FB66F98DC8AD2E11" pageId="40" pageNumber="41" value="1985-05-18" valueMax="1985-05-24" valueMin="1985-05-18">18-24 May 1985</collectingDate>
</date>
. //
<collectingMethod id="4BDF4EF79D349273FE85F9E6C9862E11" box="[278,464,1624,1651]" pageId="40" pageNumber="41">Yellow pan trap</collectingMethod>
// R.Ent.Soc.Lond. PROJECT WALLACE B.M. 1985-10.”;
<collectionCode id="F48FAE259D349273FB02F9E7CCBB2E11" box="[1169,1261,1625,1651]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="40" pageNumber="41">BMNH</collectionCode>
.
</paragraph>
</materialsCitation>
</subSubSection>
<subSubSection id="DA84656B9D349272FF2EF922C9FD2E98" lastPageId="41" lastPageNumber="42" pageId="40" pageNumber="41" type="description">
<paragraph id="922136E09D349273FF2EF922C9182ED4" blockId="40.[189,334,1692,1718]" box="[189,334,1692,1718]" pageId="40" pageNumber="41">
<heading id="C969818C9D349273FF2EF922C9182ED4" bold="true" box="[189,334,1692,1718]" fontSize="11" level="3" pageId="40" pageNumber="41" reason="3">
<emphasis id="A0EAEAF29D349273FF2EF922C9182ED4" bold="true" box="[189,334,1692,1718]" pageId="40" pageNumber="41">Description</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D349273FF2EF970CBD92F56" blockId="40.[189,1399,1741,1844]" pageId="40" pageNumber="41">
MEASUREMENTS (exposed abdominal ventrites not included). Male
<typeStatus id="4D2588429D349273FC50F973CC7B2E85" box="[963,1069,1741,1767]" pageId="40" pageNumber="41" type="holotype">holotype</typeStatus>
, total body length
<quantity id="55669B059D349273FA94F973CD242E8A" box="[1287,1394,1741,1768]" metricMagnitude="-3" metricUnit="m" metricValue="8.45" pageId="40" pageNumber="41" unit="mm" value="8.45">8.45 mm</quantity>
. Head
<quantity id="55669B059D349273FE97F94DC9382F6C" box="[260,366,1779,1806]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="40" pageNumber="41" unit="mm" value="1.6">1.60 mm</quantity>
long, across eyes
<quantity id="55669B059D349273FDD9F94DCAE22F6C" box="[586,692,1779,1806]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="40" pageNumber="41" unit="mm" value="1.8">1.80 mm</quantity>
wide. Pronotum
<quantity id="55669B059D349273FC10F94DCBBB2F6C" box="[899,1005,1779,1806]" metricMagnitude="-3" metricUnit="m" metricValue="1.65" pageId="40" pageNumber="41" unit="mm" value="1.65">1.65 mm</quantity>
long, maximum width
<quantity id="55669B059D349273FA96F94DCD252F6C" box="[1285,1395,1779,1806]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="40" pageNumber="41" unit="mm" value="2.8">2.80 mm</quantity>
. Elytral length
<quantity id="55669B059D349273FEF6F8A7C9862F56" box="[357,464,1817,1844]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="40" pageNumber="41" unit="mm" value="5.2">5.20 mm</quantity>
, maximum combined width
<quantity id="55669B059D349273FC8CF8A7CBDD2F56" box="[799,907,1817,1844]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="40" pageNumber="41" unit="mm" value="3.8">3.80 mm</quantity>
.
</paragraph>
<paragraph id="922136E09D349272FF2EF8E0CC102EE0" blockId="40.[189,1399,1885,2024]" lastBlockId="41.[189,1399,265,1667]" lastPageId="41" lastPageNumber="42" pageId="40" pageNumber="41">
Dorsum uniformly brown with castaneous labroclypeus, mouthparts, legs and venter. Head transverse, trapezoid, flattened on frons, convex on vertex, glossy dorsally and ventrally, with large not prominent compound eyes occupying nearly half side of head. Male labroclypeus (
<figureCitation id="0AA52A659D349273FB88F816CCD62FA1" box="[1051,1152,1960,1987]" captionStart="Figs 109–120" captionStartId="138.[189,241,1750,1776]" captionTargetBox="[365,1214,275,1722]" captionTargetId="figure-15@138.[365,1222,265,1730]" captionTargetPageId="138" captionText="Figs 109–120. Papuan andWallacean MaechidiusMacleay, 1819,head, dorsal (109–119) and frontal (120) views. 109. M. brocki sp. nov., holotype, ♂ (BMNH). 110. Same, paratype, ♀ (BMNH). 111. M. ciliatus sp. nov., holotype, ♂ (IECA). 112. M. crypticus sp. nov., paratype, ♂ (BMNH). 113. Same, paratype, ♀ (BMNH). 114. M. dani sp. nov., holotype, ♂ (IECA). 115. M. deltouri sp. nov., holotype, ♂ (BMNH). 116. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 117. M. fraterculus Moser, 1920, lectotype, ♀ (ZMHB). 118. M. esau Heller, 1914, lectotype, ♂ (SNSD). 119. Same, ♂ (lectotype of M. setosus Moser, 1920) (ZMHB). 120. M. gressitti Frey, 1969, holotype, ♂ (photograph courtesy BPBM) (BPBM). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122175" httpUri="https://zenodo.org/record/4122175/files/figure.png" pageId="40" pageNumber="41">Fig. 115</figureCitation>
) broadly emarginate anteriorly, its lateral margins smooth, slightly sinuous in both dorsal and lateral views. Anterolateral angles strongly protruding anteriad, almost right-angled, bent up perpendicularly to axis of frons in lateral view. Head dorsal punctures irregularly circular to hexagonal, larger in median part. Intervening spaces glossy, much smaller than punctures. Pubescence brownish, rather long and erect but sparse; each seta rising from different part of puncture. Labroclypeus laterally and frontally with delicate yellowish setae. Labroclypeus laterally and frontally with delicate yellowish setae, longest of which present near compound eyes. Male antenna 9-segmented. Scape large, widened on upper side in distal half, provided with few long erect setae on posterior and distal margin. Antennomere 2 short and transverse. Club 3-lamellate. Pronotum transverse, glossy.Anterior margin of pronotum slightly sinuous with anterolateral angles (stronger) and mesal portion (slighter) protruding anteriad. Basal margin of pronotum broadly rounded. Lateral margin of pronotum evenly broadly rounded, largely crenulate all along except for delicately crenulate prebasal area (
<figureCitation id="0AA52A659D359272FDCBFD32CAE92AC4" box="[600,703,652,678]" captionStart="Figs 208–225" captionStartId="146.[189,241,1644,1670]" captionTargetBox="[414,1187,265,1610]" captionTargetId="figure-199@146.[414,1187,265,1618]" captionTargetPageId="146" captionText="Figs 208–225. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, pronotum, dorsal view. 208. M.caperatus sp. nov., holotype, ♂ (NME). 209. M.ciliatus sp. nov., holotype, ♂ (IECA). 210. M.dani sp. nov., holotype, ♂ (IECA). 211. M. deltouri sp. nov., holotype, ♂ (BMNH). 212. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 213. M. esau Heller, 1914 (lectotype, ♂ of M. setosus Moser, 1920) (ZMHB). 214. M. fraterculus Moser, 1920, lectotype, ♀ (ZMHB). 215. M. hamatus sp. nov., holotype, ♂ (IECA). 216. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 217. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 218. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 219. M. interruptocarinulatus Heller, 1914, holotype, ♀ (SNSD). 220. M. jobiensis Moser, 1920, lectotype, ♂ (ZMHB). 221. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 222. M. konjo sp. nov., holotype, ♂ (NMPC). 223. M. lapsus sp. nov., holotype, ♂ (BMNH). 224. M. legalovi sp. nov., holotype, ♂ (BMNH). 225. M. leucopsar sp. nov., holotype, ♂ (IECA). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122191" httpUri="https://zenodo.org/record/4122191/files/figure.png" pageId="41" pageNumber="42">Fig. 211</figureCitation>
). Disc with two vague circular postmedian impressions on either side. Lateral margin of pronotum nearly straight in lateral view. Punctures irregularly circular to ovoid, deep and coarse; inner margin with delicate membrane in part covered by microscopical velvety pubescence and partly encircling puncture. Intervening spaces much smaller than punctures, glossy, partly wrinkled. Setation as on head, stronger curved; long seta between each two lateral crenulae and all along basal margin. Hypomeron smooth, sparsely punctured with circular punctures, emarginate opposite to eye (to receive canthus separating compound eye) and covered with long setae, with anterior margin which is flange-like produced. Antennal pocket deep. Scutellar shield triangular, pointed apically. Elytra cylindrical, maximum width in postbasal ¾, glossy and slightly convex dorsally, with rounded humeri and elevated omoplates. Indistinct track of one flat longitudinal carina on each elytron. Sutural carinae not indicated. Rather large obtuse hump on each elytron near apex. Elytral disc and lateral sides of elytra irregularly densely punctured, punctures ovoid; inner margin with delicate membrane in part or completely covered by microscopical velvety pubescence and partly or completely encircles puncture (
<figureCitation id="0AA52A659D359272FEA6FBC4C9CB2CF6" box="[309,413,1146,1172]" captionStart="Figs 286–301" captionStartId="152.[189,241,1716,1742]" captionTargetBox="[364,1224,266,1689]" captionTargetId="figure-172@152.[364,1224,266,1695]" captionTargetPageId="152" captionText="Figs 286–301. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 286. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 287. M. aiyura sp. nov., holotype, ♂ (BMNH). 288. M. alesbezdeki sp. nov., holotype, ♂ (NME). 289. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 290. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 291. M. paupianus Heller, 1910, ♂ (BMNH). 292. M. awu sp. nov., holotype, ♂ (IECA). 293. M. babyrousa sp. nov., holotype, ♂ (BMNH). 294. M. bintang sp. nov., holotype, ♂ (NME). 295. M. boessnecki sp. nov., holotype, ♂ (SNSD). 296. M. brocki sp. nov., holotype, ♂ (BMNH). 297. M. caperatus sp. nov., holotype, ♂ (NME). 298. M. ciliatus sp. nov., holotype, ♂ (IECA). 299. M. crypticus sp. nov., paratype, ♀ (BMNH). 300. M. dani sp. nov., holotype, ♂ (IECA). 301. M. deltouri sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122206" httpUri="https://zenodo.org/record/4122206/files/figure.png" pageId="41" pageNumber="42">Fig. 301</figureCitation>
). Setae inconspicuous and short, appressed (except suberect longer slightly clavate scale-like setae on disc and elytral apices), generally longer than on forebody (seta as long as length of corresponding puncture, not or hardly surpassing length of corresponding puncture). Intervening spaces glossy, vary from smaller than up to 2 × as large as punctures. An inconspicuous suberect seta rises from anterior margin of each puncture. Epipleuron with row or two (in broad anterior part) of suberect setae. Male pygidium with rather flat mesal hump at apical margin, with shallow annular punctures and partly microscopically velvety pubescent intervening spaces (
<figureCitation id="0AA52A659D359272FC5CFAE0CC6E2D1B" box="[975,1080,1374,1401]" captionStart="Figs 451–468" captionStartId="163.[189,241,1681,1707]" captionTargetBox="[456,1130,262,1650]" captionTargetId="figure-178@163.[456,1131,262,1657]" captionTargetPageId="163" captionText="Figs 451–468. Papuan and Wallacean Maechidius Macleay, 1819, pygidium, dorsal view. 451. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 452. M. aiyura sp. nov., holotype, ♂ (BMNH). 453. Same, different aspect (BMNH). 454. M. alesbezdeki sp. nov., holotype, ♂ (NME). 455. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 456. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 457. M. awu sp. nov., holotype, ♂ (IECA). 458. M. babyrousa sp. nov., holotype, ♂ (BMNH). 459. Same, paratype, ♀ (BMNH). 460. M. bintang sp. nov., holotype, ♂ (NME). 461. M. boessnecki sp. nov., holotype, ♂ (SNSD). 462. M. brocki sp. nov., holotype, ♂ (BMNH). 463. M. caperatus sp. nov., holotype, ♂ (NME). 464. M. ciliatus sp. nov., holotype, ♂ (IECA). 465. M. crypticus sp. nov., paratype, ♂ (BMNH). 466. Same, paratype, ♀ (BMNH). 467. M. dani sp. nov., holotype, ♂ (IECA). 468. M. deltouri sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122222" httpUri="https://zenodo.org/record/4122222/files/figure.png" pageId="41" pageNumber="42">Fig. 468</figureCitation>
). Setae of pygidium rather long, suberect. Male abdominal ventrites glossy, covered with large annular punctures, each provided with short seta. Legs long and slender, femora and tibiae covered with delicate whitish setae. Pro- and metatibiae rather strongly thickened distally. Outer margins of meso- and metatibiae densely denticulate. Two strong teeth on external margin of male protibia (Fig. 374). Male lower meso- and metatibial terminal spur strongly curved. Distal margin of metatibia in both sexes with projection over basal metatarsomere. Male metatibial spurs shorter than basal metatarsomere. Male tarsal claws with pulvilli. Spiculum gastrale as in Fig. 542. Male aedeagus as in
<figureCitation id="0AA52A659D359272FC0DF9D6CC142EE0" box="[926,1090,1640,1666]" captionStart="Figs 601–616" captionStartId="172.[189,241,1892,1918]" captionTargetBox="[458,1119,274,1854]" captionTargetId="figure-73@172.[450,1138,265,1868]" captionTargetPageId="172" captionText="Figs 601–616. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 601– 603. M. brocki sp. nov., holotype, ♂ (BMNH). 604–606. M. dani sp. nov., holotype, ♂ (IECA). 607–610. M. deltouri sp. nov., holotype, ♂ (BMNH). 611–613. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 614–616. M. esau Heller, 1914, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122238" httpUri="https://zenodo.org/record/4122238/files/figure.png" pageId="41" pageNumber="42">Figs 607–610</figureCitation>
.
</paragraph>
<paragraph id="922136E09D359272FF2EF913C9E62EA5" blockId="41.[189,432,1709,1735]" box="[189,432,1709,1735]" pageId="41" pageNumber="42">
<heading id="C969818C9D359272FF2EF913C9E62EA5" bold="true" box="[189,432,1709,1735]" fontSize="11" level="3" pageId="41" pageNumber="42" reason="3">
<emphasis id="A0EAEAF29D359272FF2EF913C9E62EA5" bold="true" box="[189,432,1709,1735]" pageId="41" pageNumber="42">Sexual dimorphism</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D359272FF2EF95EC9FD2E98" blockId="41.[189,427,1760,1786]" box="[189,427,1760,1786]" pageId="41" pageNumber="42">Female is unknown.</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D359272FF2EF89BCD402F13" box="[189,1302,1829,1906]" pageId="41" pageNumber="42" type="biology_ecology">
<paragraph id="922136E09D359272FF2EF89BC9762F5D" blockId="41.[189,288,1829,1855]" box="[189,288,1829,1855]" pageId="41" pageNumber="42">
<heading id="C969818C9D359272FF2EF89BC9762F5D" bold="true" box="[189,288,1829,1855]" fontSize="11" level="3" pageId="41" pageNumber="42" reason="3">
<emphasis id="A0EAEAF29D359272FF2EF89BC9762F5D" bold="true" box="[189,288,1829,1855]" pageId="41" pageNumber="42">Ecology</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D359272FF2EF8E9CD402F13" blockId="41.[189,1302,1879,1906]" box="[189,1302,1879,1906]" pageId="41" pageNumber="42">Inhabits lowland rainforests. Attracted to yellow pan traps (anthophilous?). Possibly nocturnal.</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D359272FF2EF822CAE12F8A" box="[189,695,1948,2024]" pageId="41" pageNumber="42" type="distribution">
<paragraph id="922136E09D359272FF2EF822C9002FD4" blockId="41.[189,342,1948,1974]" box="[189,342,1948,1974]" pageId="41" pageNumber="42">
<heading id="C969818C9D359272FF2EF822C9002FD4" bold="true" box="[189,342,1948,1974]" fontSize="11" level="3" pageId="41" pageNumber="42" reason="3">
<emphasis id="A0EAEAF29D359272FF2EF822C9002FD4" bold="true" box="[189,342,1948,1974]" pageId="41" pageNumber="42">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D359272FF2EF870CAE12F8A" blockId="41.[189,695,1998,2024]" box="[189,695,1998,2024]" pageId="41" pageNumber="42">
Hitherto only known from
<collectingRegion id="505AF8029D359272FE6AF870CAE52F8A" box="[505,691,1998,2024]" country="Indonesia" name="Sulawesi Utara" pageId="41" pageNumber="42">North Sulawesi</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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