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<document id="EF99CCD721BDBAFA9E4C65684997790E" ID-CLB-Dataset="9693" ID-DOI="10.5852/ejt.2020.721.1127" ID-GBIF-Dataset="0cd61b23-1089-4c1b-bfc0-487887abad84" ID-Zenodo-Dep="4122118" ID-ZooBank="89E62EF8-2E45-4C59-94B7-6A5603E8939B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1603474447689" checkinUser="valdenar" docAuthor="Telnov, Dmitry" docDate="2020" docId="1A3787F69D09924CFDF2FB21C9C72C7D" docLanguage="en" docName="ejt-721_telnov.pdf" docOrigin="European Journal of Taxonomy 721" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Maechidius aroae Heller 1914" docType="treatment" docVersion="10" lastPageNumber="24" masterDocId="E60EFF8E9D1C925BFF93FFBEC8562862" masterDocTitle="A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line" masterLastPageNumber="210" masterPageNumber="1" pageNumber="22" updateTime="1698846101371" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="40E26F07537F8029EFA3DD27F4D1AF33">A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line</mods:title>
</mods:titleInfo>
<mods:name id="A3D82759496222BDA8940627FBB169AB" type="personal">
<mods:role id="90DDD0D82BC9FC0B0D03FCB7ED6D0B3D">
<mods:roleTerm id="E68FF72D58FD95DBFDDFAD6B2E141AB4">Author</mods:roleTerm>
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<mods:namePart id="2AC35B537B33FF8E163953559679A9C4">Telnov, Dmitry</mods:namePart>
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<mods:title id="B3B238C5CB230117E52D414C108C5FA7">European Journal of Taxonomy</mods:title>
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<mods:date id="F8A44CA839CC6D7BC1F03BC6DAFCE128">2020</mods:date>
<mods:detail id="58515FEC3D8726ED01E168BE157204AE" type="pubDate">
<mods:number id="95EAA5CD62B60EA56815C9E75F84D34B">2020-10-19</mods:number>
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<mods:number id="91A60EFF60B86B54A24E2254FE8FEF4D">721</mods:number>
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<mods:classification id="C19099089E06F8849B4133D83E7DE5D9">journal article</mods:classification>
<mods:identifier id="0638E376C0B8D7AEB6E1B7C18EF0F5B5" type="CLB-Dataset">9693</mods:identifier>
<mods:identifier id="8E4EA9FB62B633B32F4F7CF738482AC5" type="DOI">10.5852/ejt.2020.721.1127</mods:identifier>
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<treatment id="1A3787F69D09924CFDF2FB21C9C72C7D" ID-DOI="http://doi.org/10.5281/zenodo.4344375" ID-GBIF-Taxon="168648183" ID-Zenodo-Dep="4344375" LSID="urn:lsid:plazi:treatment:1A3787F69D09924CFDF2FB21C9C72C7D" httpUri="http://treatment.plazi.org/id/1A3787F69D09924CFDF2FB21C9C72C7D" lastPageId="23" lastPageNumber="24" pageId="21" pageNumber="22">
<subSubSection id="DA84656B9D09924EFDF2FB21CB852CDB" box="[609,979,1183,1210]" pageId="21" pageNumber="22" type="nomenclature">
<paragraph id="922136E09D09924EFDF2FB21CB852CDB" blockId="21.[603,984,1183,1245]" box="[609,979,1183,1210]" pageId="21" pageNumber="22">
<heading id="C969818C9D09924EFDF2FB21CB852CDB" box="[609,979,1183,1210]" centered="true" fontSize="11" level="2" pageId="21" pageNumber="22" reason="2">
<taxonomicName id="559E4D639D09924EFDF2FB21CB852CDB" ID-CoL="3XFHC" authority="Heller, 1914" authorityName="Heller" authorityYear="1914" box="[609,979,1183,1210]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="aroae">
<emphasis id="A0EAEAF29D09924EFDF2FB21CB6C2CD8" bold="true" box="[609,826,1183,1210]" italics="true" pageId="21" pageNumber="22">Maechidius aroae</emphasis>
<bibRefCitation id="F60F4B119D09924EFCD2FB21CB852CDB" author="Heller K. M." box="[833,979,1183,1209]" pageId="21" pageNumber="22" pagination="615 - 666" refId="ref98213" refString="Heller K. M. 1914. Coleoptera. Nova Guinea. Resultats de l'Expedition scientifique neerlandaise a la Nouvelle-Guinee en 1907 et 1909 sous les auspices du Dr. H. A. Lorentz. Zoologie 9: 615 - 666." type="journal article" year="1914">Heller, 1914</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D09924EFDC8FB7CCB8E2CBF" box="[603,984,1218,1245]" pageId="21" pageNumber="22" type="description">
<paragraph id="922136E09D09924EFDC8FB7CCB8E2CBF" blockId="21.[603,984,1183,1245]" box="[603,984,1218,1245]" pageId="21" pageNumber="22">
<figureCitation id="0AA52A659D09924EFDC8FB7CCAE32CBF" box="[603,693,1218,1245]" captionStart="Figs 22–24" captionStartId="22.[189,241,1928,1954]" captionTargetBox="[354,1240,272,1893]" captionTargetId="figure-55@22.[344,1244,264,1901]" captionTargetPageId="22" captionText="Figs 22–24. Papuan and Wallacean Maechidius Macleay, 1819, habitus, dorsal view. 22. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 23. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 24. M. awu sp. nov., holotype, ♂ (IECA). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122132" httpUri="https://zenodo.org/record/4122132/files/figure.png" pageId="21" pageNumber="22">Figs 23</figureCitation>
,
<figureCitation id="0AA52A659D09924EFD52FB7DCAB92CBF" box="[705,751,1219,1245]" captionStart="Figs 97–108" captionStartId="137.[189,241,1786,1812]" captionTargetBox="[351,1228,270,1756]" captionTargetId="figure-127@137.[351,1237,265,1772]" captionTargetPageId="137" captionText="Figs 97–108. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal view. 97. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 98. M. aiyura sp. nov., holotype, ♂ (BMNH). 99. M. alesbezdeki sp. nov., holotype, ♂ (NME). 100. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 101. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 102. M. bintang sp. nov., holotype, ♂ (NME). 103. M. awu sp. nov., holotype, ♂ (IECA). 104. Same, paratype, ♀ (IECA). 105. M. boessnecki sp. nov., holotype, ♂ (SNSD). 106. M. babyrousa sp. nov., holotype, ♂ (BMNH). 107. Same, paratype, ♀ (BMNH). 108. M. caperatus sp. nov., holotype, ♂ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122173" httpUri="https://zenodo.org/record/4122173/files/figure.png" pageId="21" pageNumber="22">101</figureCitation>
,
<figureCitation id="0AA52A659D09924EFD6FFB7DCB7D2CBF" box="[764,811,1219,1245]" captionStart="Figs 193–207" captionStartId="145.[189,241,1716,1742]" captionTargetBox="[385,1203,267,1681]" captionTargetId="figure-159@145.[385,1203,265,1694]" captionTargetPageId="145" captionText="Figs 193–207. Papuan and Wallacean Maechidius Macleay, 1819, head (193–196) and pronotum (197– 207), dorsal view. 193. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 194. Same, ♂ (SNSD). 195. M. yamdena sp. nov., paratype, ♂ (BMNH). 196. Same, paratype, ♀ (BMNH). 197. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 198. M. aiyura sp. nov., holotype, ♂ (BMNH). 199. M. alesbezdeki sp. nov., holotype, ♂ (NME). 200. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 201. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 202. M. awu sp. nov., holotype, ♂ (IECA). 203. M. babyrousa sp. nov., holotype, ♂ (BMNH). 204. M. bintang sp. nov., holotype, ♂ (NME). 205. M. boessnecki sp. nov., holotype, ♂ (SNSD). 206. M. brocki sp. nov., holotype, ♂ (BMNH). 207. M. crypticus sp. nov., paratype, ♀ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122189" httpUri="https://zenodo.org/record/4122189/files/figure.png" pageId="21" pageNumber="22">201</figureCitation>
,
<figureCitation id="0AA52A659D09924EFCA4FB7DCB302CBF" box="[823,870,1219,1245]" captionStart="Figs 286–301" captionStartId="152.[189,241,1716,1742]" captionTargetBox="[364,1224,266,1689]" captionTargetId="figure-172@152.[364,1224,266,1695]" captionTargetPageId="152" captionText="Figs 286–301. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 286. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 287. M. aiyura sp. nov., holotype, ♂ (BMNH). 288. M. alesbezdeki sp. nov., holotype, ♂ (NME). 289. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 290. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 291. M. paupianus Heller, 1910, ♂ (BMNH). 292. M. awu sp. nov., holotype, ♂ (IECA). 293. M. babyrousa sp. nov., holotype, ♂ (BMNH). 294. M. bintang sp. nov., holotype, ♂ (NME). 295. M. boessnecki sp. nov., holotype, ♂ (SNSD). 296. M. brocki sp. nov., holotype, ♂ (BMNH). 297. M. caperatus sp. nov., holotype, ♂ (NME). 298. M. ciliatus sp. nov., holotype, ♂ (IECA). 299. M. crypticus sp. nov., paratype, ♀ (BMNH). 300. M. dani sp. nov., holotype, ♂ (IECA). 301. M. deltouri sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122206" httpUri="https://zenodo.org/record/4122206/files/figure.png" pageId="21" pageNumber="22">290</figureCitation>
, 363, 456
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D09924EFF2EFABACA502E5D" pageId="21" pageNumber="22" type="materials_examined">
<paragraph id="922136E09D09924EFF2EFABAC93A2D7C" blockId="21.[189,364,1284,1310]" box="[189,364,1284,1310]" pageId="21" pageNumber="22">
<heading id="C969818C9D09924EFF2EFABAC93A2D7C" bold="true" box="[189,364,1284,1310]" fontSize="11" level="3" pageId="21" pageNumber="22" reason="3">
<emphasis id="A0EAEAF29D09924EFF2EFABAC93A2D7C" bold="true" box="[189,364,1284,1310]" pageId="21" pageNumber="22">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D09924EFF79FA8DCA3A2D2F" blockId="21.[189,1398,1330,1428]" box="[234,620,1330,1357]" pageId="21" pageNumber="22">
<emphasis id="A0EAEAF29D09924EFF79FA8DC9302D2F" bold="true" box="[234,358,1331,1357]" pageId="21" pageNumber="22">
<typeStatus id="4D2588429D09924EFF79FA8DC9302D2F" box="[234,358,1331,1357]" pageId="21" pageNumber="22" type="lectotype">Lectotype</typeStatus>
</emphasis>
[herewith designated]
</paragraph>
<paragraph id="922136E09D09924EFF2EFAE8CA602DF6" blockId="21.[189,1398,1330,1428]" pageId="21" pageNumber="22">
<materialsCitation id="22F63CBD9D09924EFF2EFAE8CA602DF6" ID-GBIF-Occurrence="2898878368" collectionCode="SNSD" country="Papua New Guinea" location="PAPUA NEW GUINEA" pageId="21" pageNumber="22" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="typus">
<collectingCountry id="EA8976709D09924EFF2EFAE8C98A2D12" box="[189,476,1366,1393]" name="Papua New Guinea" pageId="21" pageNumber="22">PAPUA NEW GUINEA</collectingCountry>
•
<specimenCount id="8498FD699D09924EFE6BFAE9CA592D13" box="[504,527,1367,1393]" pageId="21" pageNumber="22" type="female">♀</specimenCount>
; “14716 [h] //
<typeStatus id="4D2588429D09924EFD2BFAE8CB502D12" box="[696,774,1366,1392]" pageId="21" pageNumber="22">Typus</typeStatus>
<specimenCount id="8498FD699D09924EFC95FAE8CB5C2D12" box="[774,778,1366,1392]" pageId="21" pageNumber="22" type="male">♂</specimenCount>
[p]
<taxonomicName id="559E4D639D09924EFCADFAE9CB802D12" authority="Heller" authorityName="Heller" authorityYear="1914" box="[830,982,1366,1393]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="185" phylum="Arthropoda" rank="species" species="aroae">
<emphasis id="A0EAEAF29D09924EFCADFAE9CBD32D13" box="[830,901,1367,1393]" italics="true" pageId="21" pageNumber="22">Aroae</emphasis>
Heller
</taxonomicName>
[h, red label]// Staatl. Museum für Tierkunde.Dresden [p]”;
<collectionCode id="F48FAE259D09924EFE77FAC4CA652DF6" box="[484,563,1402,1428]" collectionName="SNSD" pageId="21" pageNumber="22">SNSD</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="922136E09D09924EFF2EFA05CA502E5D" blockId="21.[189,1399,1467,1600]" pageId="21" pageNumber="22">
<bibRefCitation id="F60F4B119D09924EFF2EFA05C9CC2DB4" author="Heller K. M." box="[189,410,1467,1494]" pageId="21" pageNumber="22" pagination="615 - 666" refId="ref98213" refString="Heller K. M. 1914. Coleoptera. Nova Guinea. Resultats de l'Expedition scientifique neerlandaise a la Nouvelle-Guinee en 1907 et 1909 sous les auspices du Dr. H. A. Lorentz. Zoologie 9: 615 - 666." type="journal article" year="1914">Heller (1914: 627)</bibRefCitation>
based his description of
<taxonomicName id="559E4D639D09924EFD56FA02CB662DB7" authorityName="Heller" authorityYear="1914" box="[709,816,1468,1493]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="aroae">
<emphasis id="A0EAEAF29D09924EFD56FA02CB662DB7" box="[709,816,1468,1493]" italics="true" pageId="21" pageNumber="22">M. aroae</emphasis>
</taxonomicName>
on an unstated number of specimens. The single
<collectionCode id="F48FAE259D09924EFF2EFA61C95E2D9B" box="[189,264,1503,1529]" pageId="21" pageNumber="22">SNSD</collectionCode>
specimen labelled “Typus♂” is herewith designated as the
<typeStatus id="4D2588429D09924EFC4CFA60CC1D2D9A" box="[991,1099,1502,1528]" pageId="21" pageNumber="22" type="lectotype">lectotype</typeStatus>
in order to enhance the stability of nomenclature and fix the specimen that I based my redescription on, in case other
<typeStatus id="4D2588429D09924EFA83F9BCCD212E7E" box="[1296,1399,1538,1564]" pageId="21" pageNumber="22" type="syntype">syntypes</typeStatus>
are discovered in the future.
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D09924CFF2EF9D8C9C72C7D" lastPageId="23" lastPageNumber="24" pageId="21" pageNumber="22" type="description">
<paragraph id="922136E09D09924EFF2EF9D8C9182EE2" blockId="21.[189,334,1638,1664]" box="[189,334,1638,1664]" pageId="21" pageNumber="22">
<heading id="C969818C9D09924EFF2EF9D8C9182EE2" bold="true" box="[189,334,1638,1664]" fontSize="11" level="3" pageId="21" pageNumber="22" reason="3">
<emphasis id="A0EAEAF29D09924EFF2EF9D8C9182EE2" bold="true" box="[189,334,1638,1664]" pageId="21" pageNumber="22">Description</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D09924EFF2EF928CB112ED2" blockId="21.[189,839,1685,1712]" box="[189,839,1685,1712]" pageId="21" pageNumber="22">
MEASUREMENTS. Total body length
<quantity id="55669B059D09924EFDC2F92BCAEF2ED2" box="[593,697,1685,1712]" metricMagnitude="-3" metricUnit="m" metricValue="9.6" pageId="21" pageNumber="22" unit="mm" value="9.6">9.60 mm</quantity>
(
<typeStatus id="4D2588429D09924EFD5BF92BCB6D2ED2" box="[712,827,1685,1712]" pageId="21" pageNumber="22" type="lectotype">lectotype</typeStatus>
).
</paragraph>
<paragraph id="922136E09D09924CFF2EF969CCC22BC9" blockId="21.[189,1399,1751,2025]" lastBlockId="23.[189,1399,265,939]" lastPageId="23" lastPageNumber="24" pageId="21" pageNumber="22">
Dorsum and venter uniformly black-brown with exception of paler brown appendages and labroclypeus. Head flattened dorsally, subopaque dorsally and ventrally. Female labroclypeus very broadly emarginate anteriorly (
<figureCitation id="0AA52A659D09924EFED3F8A0C9F12F5A" box="[320,423,1822,1848]" captionStart="Figs 97–108" captionStartId="137.[189,241,1786,1812]" captionTargetBox="[351,1228,270,1756]" captionTargetId="figure-127@137.[351,1237,265,1772]" captionTargetPageId="137" captionText="Figs 97–108. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal view. 97. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 98. M. aiyura sp. nov., holotype, ♂ (BMNH). 99. M. alesbezdeki sp. nov., holotype, ♂ (NME). 100. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 101. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 102. M. bintang sp. nov., holotype, ♂ (NME). 103. M. awu sp. nov., holotype, ♂ (IECA). 104. Same, paratype, ♀ (IECA). 105. M. boessnecki sp. nov., holotype, ♂ (SNSD). 106. M. babyrousa sp. nov., holotype, ♂ (BMNH). 107. Same, paratype, ♀ (BMNH). 108. M. caperatus sp. nov., holotype, ♂ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122173" httpUri="https://zenodo.org/record/4122173/files/figure.png" pageId="21" pageNumber="22">Fig. 101</figureCitation>
), its lateral margins nearly straight in dorsal, sinuous in lateral view. Anterolateral angles of labroclypeus very broadly rounded, not protruding, raised up at angle of nearly 90° to frons in lateral view. Anterior and lateral margins of labroclypeus smooth. Underside of labroclypeus with sparse long setae along anterior and lateral margins. Canthus very broadly rounded in dorsal view. Punctures of frons circular, moderately large and shallow. Intervening spaces microreticulate, generally much larger than punctures. Pubescence delicate, variably long, appressed. Some setae very short, much shorter than corresponding punctures. Each seta rises from anterior margin of corresponding puncture. Female antenna 8-segmented, club 3-lamellate (antenna 7-segmented according to
<bibRefCitation id="F60F4B119D0B924CFC46FE8ECC392928" author="Frey G." box="[981,1135,303,330]" pageId="23" pageNumber="24" pagination="494 - 509" refId="ref98001" refString="Frey G. 1969. Revision der Maechidius - Arten Neu-Guineas (Col., Melolonth.). Entomologische Arbeiten aus dem Museum G. Frey 20: 494 - 509." type="journal article" year="1969">Frey (1969))</bibRefCitation>
. Pronotum transverse, flattened dorsally, subopaque dorsally and laterally. Anterior margin emarginate with anterolateral angles protruding anteriad. Basal margin subtruncate. Lateral margin of pronotum broadly rounded, somewhat more strongly constricted in basal fourth, crenulate in anterior, denticulate in posterior half (
<figureCitation id="0AA52A659D0B924CFF55FE79C97D2983" box="[198,299,455,481]" captionStart="Figs 193–207" captionStartId="145.[189,241,1716,1742]" captionTargetBox="[385,1203,267,1681]" captionTargetId="figure-159@145.[385,1203,265,1694]" captionTargetPageId="145" captionText="Figs 193–207. Papuan and Wallacean Maechidius Macleay, 1819, head (193–196) and pronotum (197– 207), dorsal view. 193. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 194. Same, ♂ (SNSD). 195. M. yamdena sp. nov., paratype, ♂ (BMNH). 196. Same, paratype, ♀ (BMNH). 197. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 198. M. aiyura sp. nov., holotype, ♂ (BMNH). 199. M. alesbezdeki sp. nov., holotype, ♂ (NME). 200. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 201. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 202. M. awu sp. nov., holotype, ♂ (IECA). 203. M. babyrousa sp. nov., holotype, ♂ (BMNH). 204. M. bintang sp. nov., holotype, ♂ (NME). 205. M. boessnecki sp. nov., holotype, ♂ (SNSD). 206. M. brocki sp. nov., holotype, ♂ (BMNH). 207. M. crypticus sp. nov., paratype, ♀ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122189" httpUri="https://zenodo.org/record/4122189/files/figure.png" pageId="23" pageNumber="24">Fig. 201</figureCitation>
). Long distally curved erect seta present between every two crenulae / denticles. Lateral margin of pronotum slightly sinuous in lateral view. Punctures circular to ovoid, shallow. Intervening spaces as large as to larger than punctures. Hypomeron separated from prosternum by low straight carina, with extraordinarily long setae on anterolateral margin opposite to compound eye. Antennal pocket shallow. Scutellar shield subtriangular, narrowly rounded apically. Elytron glossy, with vague tracks of two vague glabrous longitudinal carinae. Sutural carinae not present. Punctures of disc sparse, narrow and rather deep, linear incision-shaped (
<figureCitation id="0AA52A659D0B924CFD1CFD15CAA02AA4" box="[655,758,683,710]" captionStart="Figs 286–301" captionStartId="152.[189,241,1716,1742]" captionTargetBox="[364,1224,266,1689]" captionTargetId="figure-172@152.[364,1224,266,1695]" captionTargetPageId="152" captionText="Figs 286–301. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 286. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 287. M. aiyura sp. nov., holotype, ♂ (BMNH). 288. M. alesbezdeki sp. nov., holotype, ♂ (NME). 289. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 290. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 291. M. paupianus Heller, 1910, ♂ (BMNH). 292. M. awu sp. nov., holotype, ♂ (IECA). 293. M. babyrousa sp. nov., holotype, ♂ (BMNH). 294. M. bintang sp. nov., holotype, ♂ (NME). 295. M. boessnecki sp. nov., holotype, ♂ (SNSD). 296. M. brocki sp. nov., holotype, ♂ (BMNH). 297. M. caperatus sp. nov., holotype, ♂ (NME). 298. M. ciliatus sp. nov., holotype, ♂ (IECA). 299. M. crypticus sp. nov., paratype, ♀ (BMNH). 300. M. dani sp. nov., holotype, ♂ (IECA). 301. M. deltouri sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122206" httpUri="https://zenodo.org/record/4122206/files/figure.png" pageId="23" pageNumber="24">Fig. 290</figureCitation>
). Inconspicuous suberect seta rises from under anterior margin of each puncture; setae generally longer than corresponding punctures. Intervening spaces in part microreticulate, micropunctate or wrinkled. Setae inconspicuous, short, suberect. Female pygidium flattened dorsally, with large annular punctures (
<figureCitation id="0AA52A659D0B924CFC81FCA0CB292B5A" box="[786,895,797,824]" captionStart="Figs 451–468" captionStartId="163.[189,241,1681,1707]" captionTargetBox="[456,1130,262,1650]" captionTargetId="figure-178@163.[456,1131,262,1657]" captionTargetPageId="163" captionText="Figs 451–468. Papuan and Wallacean Maechidius Macleay, 1819, pygidium, dorsal view. 451. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 452. M. aiyura sp. nov., holotype, ♂ (BMNH). 453. Same, different aspect (BMNH). 454. M. alesbezdeki sp. nov., holotype, ♂ (NME). 455. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 456. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 457. M. awu sp. nov., holotype, ♂ (IECA). 458. M. babyrousa sp. nov., holotype, ♂ (BMNH). 459. Same, paratype, ♀ (BMNH). 460. M. bintang sp. nov., holotype, ♂ (NME). 461. M. boessnecki sp. nov., holotype, ♂ (SNSD). 462. M. brocki sp. nov., holotype, ♂ (BMNH). 463. M. caperatus sp. nov., holotype, ♂ (NME). 464. M. ciliatus sp. nov., holotype, ♂ (IECA). 465. M. crypticus sp. nov., paratype, ♂ (BMNH). 466. Same, paratype, ♀ (BMNH). 467. M. dani sp. nov., holotype, ♂ (IECA). 468. M. deltouri sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122222" httpUri="https://zenodo.org/record/4122222/files/figure.png" pageId="23" pageNumber="24">Fig. 456</figureCitation>
). Intervening spaces glossy, smaller than punctures except along midline. Female protibia widened distally, with three large external teeth: two larger acute distal and one obtuse basal (Fig. 363). Protibial terminal spur not present. Female metatibial terminal spurs short, apically rounded. Tarsal claws with pulvilli (in females only?).
</paragraph>
<caption id="C6E166689D0A924DFF2EF836CAE52F8A" ID-DOI="http://doi.org/10.5281/zenodo.4122132" ID-Zenodo-Dep="4122132" httpUri="https://zenodo.org/record/4122132/files/figure.png" pageId="22" pageNumber="23" startId="22.[189,241,1928,1954]" targetBox="[354,1240,272,1893]" targetPageId="22">
<paragraph id="922136E09D0A924DFF2EF836CAE52F8A" blockId="22.[189,1399,1928,2025]" pageId="22" pageNumber="23">
<emphasis id="A0EAEAF29D0A924DFF2EF836C91E2FC0" bold="true" box="[189,328,1928,1954]" pageId="22" pageNumber="23">Figs 22–24.</emphasis>
Papuan and Wallacean
<taxonomicName id="559E4D639D0A924DFDCFF836CBCF2FC0" authority="Macleay, 1819" authorityName="Macleay" authorityYear="1819" box="[604,921,1928,1954]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="A0EAEAF29D0A924DFDCFF836CAB32FC0" box="[604,741,1928,1954]" italics="true" pageId="22" pageNumber="23">Maechidius</emphasis>
<bibRefCitation id="F60F4B119D0A924DFD78F836CBCF2FC0" author="Macleay W. S." box="[747,921,1928,1954]" pageId="22" pageNumber="23" refId="ref98319" refString="Macleay W. S. 1819. Horae Entomologicae: or Essays on the Annulose Animals. Volume I, Part I. Containing General Observations on the Geography, Manners, and the Affinities of the Insects which compose the Genus Scarabaeus of Linnaeus; to which are added a few incidental Remarks on the Genera Lucanus and Hister of the same Author. With Appendix and Plates. R. and A. Taylor, London." type="book" year="1819">Macleay, 1819</bibRefCitation>
</taxonomicName>
, habitus, dorsal view.
<emphasis id="A0EAEAF29D0A924DFB09F836CCE12FC0" bold="true" box="[1178,1207,1928,1954]" pageId="22" pageNumber="23">22</emphasis>
.
<taxonomicName id="559E4D639D0A924DFB55F837C90E2FA7" authority="Arrow, 1941" authorityName="Arrow" authorityYear="1941" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="angusticeps">
<emphasis id="A0EAEAF29D0A924DFB55F837CD212FC0" box="[1222,1399,1929,1954]" italics="true" pageId="22" pageNumber="23">M. angusticeps</emphasis>
<bibRefCitation id="F60F4B119D0A924DFF2EF815C90E2FA7" author="Arrow G. E." box="[189,344,1963,1989]" pageId="22" pageNumber="23" pagination="448 - 464" refId="ref97558" refString="Arrow G. E. 1941. XXIX. - Some undescribed species of melolonthid Coleoptera from Western New Guinea and the adjacent islands of Waigeu and Japen. Annals and Magazine of Natural History, Series 11 7 (41): 448 - 464. https: // doi. org / 10.1080 / 03745481.1941.9727945" type="journal article" year="1941">Arrow, 1941</bibRefCitation>
</taxonomicName>
, holotype, ♂ (BMNH).
<emphasis id="A0EAEAF29D0A924DFD10F815CAF62FA7" bold="true" box="[643,672,1963,1989]" pageId="22" pageNumber="23">23</emphasis>
.
<taxonomicName id="559E4D639D0A924DFD20F812CB902FA7" authority="Heller, 1914" authorityName="Heller" authorityYear="1914" box="[691,966,1963,1989]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="aroae">
<emphasis id="A0EAEAF29D0A924DFD20F812CB742FA7" box="[691,802,1964,1989]" italics="true" pageId="22" pageNumber="23">M. aroae</emphasis>
<bibRefCitation id="F60F4B119D0A924DFCBDF815CB902FA7" author="Heller K. M." box="[814,966,1963,1989]" pageId="22" pageNumber="23" pagination="615 - 666" refId="ref98213" refString="Heller K. M. 1914. Coleoptera. Nova Guinea. Resultats de l'Expedition scientifique neerlandaise a la Nouvelle-Guinee en 1907 et 1909 sous les auspices du Dr. H. A. Lorentz. Zoologie 9: 615 - 666." type="journal article" year="1914">Heller, 1914</bibRefCitation>
</taxonomicName>
, lectotype, ♀ (SNSD).
<emphasis id="A0EAEAF29D0A924DFB7AF815CD502FA7" bold="true" box="[1257,1286,1963,1989]" pageId="22" pageNumber="23">24</emphasis>
.
<taxonomicName id="559E4D639D0A924DFA89F812CD212FA7" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[1306,1399,1964,1989]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="awu" status="sp. nov.">
<emphasis id="A0EAEAF29D0A924DFA89F812CD212FA7" box="[1306,1399,1964,1989]" italics="true" pageId="22" pageNumber="23">M. awu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D0A924DFF2EF871C94C2F8B" box="[189,282,1999,2025]" pageId="22" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂ (IECA). Not to scale.
</paragraph>
</caption>
<paragraph id="922136E09D0B924CFF2EFC6AC9E62B8C" blockId="23.[189,432,980,1006]" box="[189,432,980,1006]" pageId="23" pageNumber="24">
<heading id="C969818C9D0B924CFF2EFC6AC9E62B8C" bold="true" box="[189,432,980,1006]" fontSize="11" level="3" pageId="23" pageNumber="24" reason="3">
<emphasis id="A0EAEAF29D0B924CFF2EFC6AC9E62B8C" bold="true" box="[189,432,980,1006]" pageId="23" pageNumber="24">Sexual dimorphism</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D0B924CFF2EFBBBC9C72C7D" blockId="23.[189,401,1029,1055]" box="[189,401,1029,1055]" pageId="23" pageNumber="24">Male is unknown.</paragraph>
</subSubSection>
</treatment>
</document>
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