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<document id="18E43B29017DF2FBF8C4AE69C1ED5A71" ID-CLB-Dataset="9693" ID-DOI="10.5852/ejt.2020.721.1127" ID-GBIF-Dataset="0cd61b23-1089-4c1b-bfc0-487887abad84" ID-Zenodo-Dep="4122118" ID-ZooBank="89E62EF8-2E45-4C59-94B7-6A5603E8939B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1603474447689" checkinUser="valdenar" docAuthor="Telnov, Dmitry" docDate="2020" docId="1A3787F69D08924EFDA8F971C9FD2C03" docLanguage="en" docName="ejt-721_telnov.pdf" docOrigin="European Journal of Taxonomy 721" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Maechidius angusticeps Arrow 1941" docType="treatment" docVersion="10" lastPageNumber="22" masterDocId="E60EFF8E9D1C925BFF93FFBEC8562862" masterDocTitle="A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line" masterLastPageNumber="210" masterPageNumber="1" pageNumber="21" updateTime="1698846101371" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="E89A057AE1EC8A5D64DD89EEB319E5B7">A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line</mods:title>
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<mods:namePart id="037DD29CE89593116D74C9812A58387A">Telnov, Dmitry</mods:namePart>
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<mods:title id="662DC8DE9C39048376929AA78DC8B8C0">European Journal of Taxonomy</mods:title>
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<mods:date id="60B61668CA9944B08E481BF61ABEC785">2020</mods:date>
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<mods:number id="13DCC093A96F75E2FB8ED8AF43AC1BE7">2020-10-19</mods:number>
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<treatment id="1A3787F69D08924EFDA8F971C9FD2C03" ID-DOI="http://doi.org/10.5281/zenodo.4344431" ID-GBIF-Taxon="168648184" ID-Zenodo-Dep="4344431" LSID="urn:lsid:plazi:treatment:1A3787F69D08924EFDA8F971C9FD2C03" httpUri="http://treatment.plazi.org/id/1A3787F69D08924EFDA8F971C9FD2C03" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21">
<subSubSection id="DA84656B9D08924FFDA8F971CBAF2E88" box="[571,1017,1743,1770]" pageId="20" pageNumber="21" type="nomenclature">
<paragraph id="922136E09D08924FFDA8F971CBAF2E88" blockId="20.[545,1042,1743,1806]" box="[571,1017,1743,1770]" pageId="20" pageNumber="21">
<heading id="C969818C9D08924FFDA8F971CBAF2E88" box="[571,1017,1743,1770]" centered="true" fontSize="11" level="2" pageId="20" pageNumber="21" reason="2">
<taxonomicName id="559E4D639D08924FFDA8F971CBAF2E88" ID-CoL="72TG2" authority="Arrow, 1941" authorityName="Arrow" authorityYear="1941" box="[571,1017,1743,1770]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="angusticeps">
<emphasis id="A0EAEAF29D08924FFDA8F971CB082E88" bold="true" box="[571,862,1743,1770]" italics="true" pageId="20" pageNumber="21">Maechidius angusticeps</emphasis>
<bibRefCitation id="F60F4B119D08924FFCF6F96ECBAF2E88" author="Arrow G. E." box="[869,1017,1744,1770]" pageId="20" pageNumber="21" pagination="448 - 464" refId="ref97558" refString="Arrow G. E. 1941. XXIX. - Some undescribed species of melolonthid Coleoptera from Western New Guinea and the adjacent islands of Waigeu and Japen. Annals and Magazine of Natural History, Series 11 7 (41): 448 - 464. https: // doi. org / 10.1080 / 03745481.1941.9727945" type="journal article" year="1941">Arrow, 1941</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D08924FFDB2F94DCC442F6C" box="[545,1042,1779,1806]" pageId="20" pageNumber="21" type="description">
<paragraph id="922136E09D08924FFDB2F94DCC442F6C" blockId="20.[545,1042,1743,1806]" box="[545,1042,1779,1806]" pageId="20" pageNumber="21">
<figureCitation id="0AA52A659D08924FFDB2F94DCA2D2F6F" box="[545,635,1779,1805]" captionStart="Figs 22–24" captionStartId="22.[189,241,1928,1954]" captionTargetBox="[354,1240,272,1893]" captionTargetId="figure-55@22.[344,1244,264,1901]" captionTargetPageId="22" captionText="Figs 22–24. Papuan and Wallacean Maechidius Macleay, 1819, habitus, dorsal view. 22. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 23. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 24. M. awu sp. nov., holotype, ♂ (IECA). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122132" httpUri="https://zenodo.org/record/4122132/files/figure.png" pageId="20" pageNumber="21">Figs 22</figureCitation>
,
<figureCitation id="0AA52A659D08924FFD14F94DCAE02F6F" box="[647,694,1779,1805]" captionStart="Figs 97–108" captionStartId="137.[189,241,1786,1812]" captionTargetBox="[351,1228,270,1756]" captionTargetId="figure-127@137.[351,1237,265,1772]" captionTargetPageId="137" captionText="Figs 97–108. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal view. 97. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 98. M. aiyura sp. nov., holotype, ♂ (BMNH). 99. M. alesbezdeki sp. nov., holotype, ♂ (NME). 100. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 101. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 102. M. bintang sp. nov., holotype, ♂ (NME). 103. M. awu sp. nov., holotype, ♂ (IECA). 104. Same, paratype, ♀ (IECA). 105. M. boessnecki sp. nov., holotype, ♂ (SNSD). 106. M. babyrousa sp. nov., holotype, ♂ (BMNH). 107. Same, paratype, ♀ (BMNH). 108. M. caperatus sp. nov., holotype, ♂ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122173" httpUri="https://zenodo.org/record/4122173/files/figure.png" pageId="20" pageNumber="21">100</figureCitation>
,
<figureCitation id="0AA52A659D08924FFD52F94DCAA62F6F" box="[705,752,1779,1805]" captionStart="Figs 193–207" captionStartId="145.[189,241,1716,1742]" captionTargetBox="[385,1203,267,1681]" captionTargetId="figure-159@145.[385,1203,265,1694]" captionTargetPageId="145" captionText="Figs 193–207. Papuan and Wallacean Maechidius Macleay, 1819, head (193–196) and pronotum (197– 207), dorsal view. 193. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 194. Same, ♂ (SNSD). 195. M. yamdena sp. nov., paratype, ♂ (BMNH). 196. Same, paratype, ♀ (BMNH). 197. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 198. M. aiyura sp. nov., holotype, ♂ (BMNH). 199. M. alesbezdeki sp. nov., holotype, ♂ (NME). 200. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 201. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 202. M. awu sp. nov., holotype, ♂ (IECA). 203. M. babyrousa sp. nov., holotype, ♂ (BMNH). 204. M. bintang sp. nov., holotype, ♂ (NME). 205. M. boessnecki sp. nov., holotype, ♂ (SNSD). 206. M. brocki sp. nov., holotype, ♂ (BMNH). 207. M. crypticus sp. nov., paratype, ♀ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122189" httpUri="https://zenodo.org/record/4122189/files/figure.png" pageId="20" pageNumber="21">200</figureCitation>
,
<figureCitation id="0AA52A659D08924FFD6FF94DCB7D2F6F" box="[764,811,1779,1805]" captionStart="Figs 286–301" captionStartId="152.[189,241,1716,1742]" captionTargetBox="[364,1224,266,1689]" captionTargetId="figure-172@152.[364,1224,266,1695]" captionTargetPageId="152" captionText="Figs 286–301. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 286. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 287. M. aiyura sp. nov., holotype, ♂ (BMNH). 288. M. alesbezdeki sp. nov., holotype, ♂ (NME). 289. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 290. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 291. M. paupianus Heller, 1910, ♂ (BMNH). 292. M. awu sp. nov., holotype, ♂ (IECA). 293. M. babyrousa sp. nov., holotype, ♂ (BMNH). 294. M. bintang sp. nov., holotype, ♂ (NME). 295. M. boessnecki sp. nov., holotype, ♂ (SNSD). 296. M. brocki sp. nov., holotype, ♂ (BMNH). 297. M. caperatus sp. nov., holotype, ♂ (NME). 298. M. ciliatus sp. nov., holotype, ♂ (IECA). 299. M. crypticus sp. nov., paratype, ♀ (BMNH). 300. M. dani sp. nov., holotype, ♂ (IECA). 301. M. deltouri sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122206" httpUri="https://zenodo.org/record/4122206/files/figure.png" pageId="20" pageNumber="21">289</figureCitation>
, 362, 455, 576–578
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D08924FFF2EF88BC9C32F8B" pageId="20" pageNumber="21" type="materials_examined">
<paragraph id="922136E09D08924FFF2EF88BC93A2F2C" blockId="20.[189,364,1844,1871]" box="[189,364,1844,1871]" pageId="20" pageNumber="21">
<heading id="C969818C9D08924FFF2EF88BC93A2F2C" bold="true" box="[189,364,1844,1871]" fontSize="11" level="3" pageId="20" pageNumber="21" reason="3">
<emphasis id="A0EAEAF29D08924FFF2EF88BC93A2F2C" bold="true" box="[189,364,1844,1871]" pageId="20" pageNumber="21">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D08924FFF79F8DAC90A2F1C" blockId="20.[189,1399,1892,2025]" box="[234,348,1892,1918]" pageId="20" pageNumber="21">
<emphasis id="A0EAEAF29D08924FFF79F8DAC90A2F1C" bold="true" box="[234,348,1892,1918]" pageId="20" pageNumber="21">
<typeStatus id="4D2588429D08924FFF79F8DAC90A2F1C" box="[234,348,1892,1918]" pageId="20" pageNumber="21" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="922136E09D08924FFF2EF839C9C32F8B" blockId="20.[189,1399,1892,2025]" pageId="20" pageNumber="21">
<materialsCitation id="22F63CBD9D08924FFF2EF839C9C32F8B" ID-GBIF-Occurrence="2898878436" collectingDate="1937-07" collectionCode="BMNH" collectorName="W. Suber." country="Indonesia" location="Humboldt Bay Dist. Bewani Mts." municipality="Humboldt Bay Dist. Bewani Mts." pageId="20" pageNumber="21" specimenCount="1" specimenCount-male="1" typeStatus="type">
<collectingCountry id="EA8976709D08924FFF2EF839C90D2FC3" box="[189,347,1927,1953]" name="Indonesia" pageId="20" pageNumber="21">INDONESIA</collectingCountry>
•
<specimenCount id="8498FD699D08924FFEE5F839C9D82FC3" box="[374,398,1927,1953]" pageId="20" pageNumber="21" type="male">♂</specimenCount>
; “
<typeStatus id="4D2588429D08924FFE35F839C9B52FC3" box="[422,483,1927,1953]" pageId="20" pageNumber="21">Type</typeStatus>
[p, label circular, red frame] //
<collectingCountry id="EA8976709D08924FFCDCF839CC2A2FC3" box="[847,1148,1927,1953]" pageId="20" pageNumber="21">DUTCH NEW GUINEA</collectingCountry>
:
<collectingMunicipality id="7245AC9A9D08924FFB1BF839C9032FA6" pageId="20" pageNumber="21">Humboldt Bay Dist. Bewani Mts.</collectingMunicipality>
400 meters.
<date id="E62010209D08924FFE61F814CA0E2FA6" box="[498,600,1962,1988]" pageId="20" pageNumber="21" value="1937-07">
<collectingDate id="F664E9C89D08924FFE61F814CA0E2FA6" box="[498,600,1962,1988]" pageId="20" pageNumber="21" value="1937-07">Vii.1937</collectingDate>
</date>
[p] //
<collectorName id="3F6B53369D08924FFD09F814CB532FA6" box="[666,773,1962,1988]" pageId="20" pageNumber="21">W.Süber.</collectorName>
B.M.1938-177. [p]//
<taxonomicName id="559E4D639D08924FFB96F814CD212FA6" authority="Arrow" authorityName="Arrow" authorityYear="1941" box="[1029,1399,1962,1988]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="angusticeps">
<emphasis id="A0EAEAF29D08924FFB96F814CD752FA6" box="[1029,1315,1962,1988]" italics="true" pageId="20" pageNumber="21">Maechidius angusticeps</emphasis>
Arrow
</taxonomicName>
<typeStatus id="4D2588429D08924FFF2EF871C8A62F8B" box="[189,240,1999,2025]" pageId="20" pageNumber="21">type</typeStatus>
[h]”;
<collectionCode id="F48FAE259D08924FFEA5F871C9C42F8B" box="[310,402,1999,2025]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="20" pageNumber="21">BMNH</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D09924EFF2EFEB6C9FD2C03" pageId="21" pageNumber="22" type="description">
<paragraph id="922136E09D09924EFF2EFEB6C9182940" blockId="21.[189,334,264,290]" box="[189,334,264,290]" pageId="21" pageNumber="22">
<heading id="C969818C9D09924EFF2EFEB6C9182940" bold="true" box="[189,334,264,290]" fontSize="11" level="3" pageId="21" pageNumber="22" reason="3">
<emphasis id="A0EAEAF29D09924EFF2EFEB6C9182940" bold="true" box="[189,334,264,290]" pageId="21" pageNumber="22">Description</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D09924EFF50FE86CD212B93" blockId="21.[189,1399,311,1009]" pageId="21" pageNumber="22">
Dorsum and venter uniformly black-brown, labroclypeus castaneous brown. Male labroclypeus broadly emarginate, anterolateral angles obtuse angulate, protruding anteriad (
<figureCitation id="0AA52A659D09924EFB86FEE4CCD62917" box="[1045,1152,346,373]" captionStart="Figs 97–108" captionStartId="137.[189,241,1786,1812]" captionTargetBox="[351,1228,270,1756]" captionTargetId="figure-127@137.[351,1237,265,1772]" captionTargetPageId="137" captionText="Figs 97–108. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal view. 97. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 98. M. aiyura sp. nov., holotype, ♂ (BMNH). 99. M. alesbezdeki sp. nov., holotype, ♂ (NME). 100. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 101. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 102. M. bintang sp. nov., holotype, ♂ (NME). 103. M. awu sp. nov., holotype, ♂ (IECA). 104. Same, paratype, ♀ (IECA). 105. M. boessnecki sp. nov., holotype, ♂ (SNSD). 106. M. babyrousa sp. nov., holotype, ♂ (BMNH). 107. Same, paratype, ♀ (BMNH). 108. M. caperatus sp. nov., holotype, ♂ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122173" httpUri="https://zenodo.org/record/4122173/files/figure.png" pageId="21" pageNumber="22">Fig. 100</figureCitation>
), its lateral margins strongly sinuous in lateral view. Punctures of frons ovoid, deep. Setae inconspicuous, scale-like, narrow and short (not or hardly surpassing length of corresponding punctures), each seta rises from anterior margin of corresponding puncture. Lateral margin of pronotum broadly rounded, shallowly emarginate prebasally, delicately crenulate all along (
<figureCitation id="0AA52A659D09924EFD38FE56CB422A60" box="[683,788,488,514]" captionStart="Figs 193–207" captionStartId="145.[189,241,1716,1742]" captionTargetBox="[385,1203,267,1681]" captionTargetId="figure-159@145.[385,1203,265,1694]" captionTargetPageId="145" captionText="Figs 193–207. Papuan and Wallacean Maechidius Macleay, 1819, head (193–196) and pronotum (197– 207), dorsal view. 193. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 194. Same, ♂ (SNSD). 195. M. yamdena sp. nov., paratype, ♂ (BMNH). 196. Same, paratype, ♀ (BMNH). 197. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 198. M. aiyura sp. nov., holotype, ♂ (BMNH). 199. M. alesbezdeki sp. nov., holotype, ♂ (NME). 200. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 201. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 202. M. awu sp. nov., holotype, ♂ (IECA). 203. M. babyrousa sp. nov., holotype, ♂ (BMNH). 204. M. bintang sp. nov., holotype, ♂ (NME). 205. M. boessnecki sp. nov., holotype, ♂ (SNSD). 206. M. brocki sp. nov., holotype, ♂ (BMNH). 207. M. crypticus sp. nov., paratype, ♀ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122189" httpUri="https://zenodo.org/record/4122189/files/figure.png" pageId="21" pageNumber="22">Fig. 200</figureCitation>
). Appressed scale-like clavate seta present between every two crenulae. Lateral margin of pronotum sinuous in lateral view. Hypomeron sinuous, long setose opposite to compound eye, flange-like. Antennal pocket deep. Punctures larger and deeper than those on head. Intervening spaces glossy, narrow, wrinkled. All setae much shorter than corresponding punctures, each seta rises from anterior margin of corresponding puncture. Basal margin laterally with fringe of longer clavate scale-like setae. Lateral margins, antero- and posterolateral angles completely covered with microscopical velvety pubescence. Punctures of elytra ovoid, irregular in size, less deep than those on forebody (
<figureCitation id="0AA52A659D09924EFE7AFD61CA022A9B" box="[489,596,735,761]" captionStart="Figs 286–301" captionStartId="152.[189,241,1716,1742]" captionTargetBox="[364,1224,266,1689]" captionTargetId="figure-172@152.[364,1224,266,1695]" captionTargetPageId="152" captionText="Figs 286–301. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 286. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 287. M. aiyura sp. nov., holotype, ♂ (BMNH). 288. M. alesbezdeki sp. nov., holotype, ♂ (NME). 289. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 290. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 291. M. paupianus Heller, 1910, ♂ (BMNH). 292. M. awu sp. nov., holotype, ♂ (IECA). 293. M. babyrousa sp. nov., holotype, ♂ (BMNH). 294. M. bintang sp. nov., holotype, ♂ (NME). 295. M. boessnecki sp. nov., holotype, ♂ (SNSD). 296. M. brocki sp. nov., holotype, ♂ (BMNH). 297. M. caperatus sp. nov., holotype, ♂ (NME). 298. M. ciliatus sp. nov., holotype, ♂ (IECA). 299. M. crypticus sp. nov., paratype, ♀ (BMNH). 300. M. dani sp. nov., holotype, ♂ (IECA). 301. M. deltouri sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122206" httpUri="https://zenodo.org/record/4122206/files/figure.png" pageId="21" pageNumber="22">Fig. 289</figureCitation>
). Elytral setae generally very short (only few setae are as long as corresponding punctures), each seta rises from anterior margin of corresponding puncture. Most of punctures partly or completely encircled by delicate and extremely short velvety pubescence, composing circular to subcircular patterns around perimeter of punctures. Male pygidium slightly convex dorsally, with dense ovoid deep punctures and suberect scale-like clavate setae not or hardly surpassing length of corresponding punctures (
<figureCitation id="0AA52A659D09924EFD8AFC2ECAD32BC9" box="[537,645,912,939]" captionStart="Figs 451–468" captionStartId="163.[189,241,1681,1707]" captionTargetBox="[456,1130,262,1650]" captionTargetId="figure-178@163.[456,1131,262,1657]" captionTargetPageId="163" captionText="Figs 451–468. Papuan and Wallacean Maechidius Macleay, 1819, pygidium, dorsal view. 451. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 452. M. aiyura sp. nov., holotype, ♂ (BMNH). 453. Same, different aspect (BMNH). 454. M. alesbezdeki sp. nov., holotype, ♂ (NME). 455. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 456. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 457. M. awu sp. nov., holotype, ♂ (IECA). 458. M. babyrousa sp. nov., holotype, ♂ (BMNH). 459. Same, paratype, ♀ (BMNH). 460. M. bintang sp. nov., holotype, ♂ (NME). 461. M. boessnecki sp. nov., holotype, ♂ (SNSD). 462. M. brocki sp. nov., holotype, ♂ (BMNH). 463. M. caperatus sp. nov., holotype, ♂ (NME). 464. M. ciliatus sp. nov., holotype, ♂ (IECA). 465. M. crypticus sp. nov., paratype, ♂ (BMNH). 466. Same, paratype, ♀ (BMNH). 467. M. dani sp. nov., holotype, ♂ (IECA). 468. M. deltouri sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122222" httpUri="https://zenodo.org/record/4122222/files/figure.png" pageId="21" pageNumber="22">Fig. 455</figureCitation>
). Male protibia with flat dorsal carina and two distal teeth on external margin (Fig. 362). Male upper metatibial terminal spur longer than half length of male basal metatarsomere, surpassing distal margin of metatibia in dorsal view. Male aedeagus as in
<figureCitation id="0AA52A659D09924EFB5CFC69CD252B93" box="[1231,1395,982,1009]" captionStart="Figs 570–585" captionStartId="170.[189,241,1892,1918]" captionTargetBox="[425,1155,265,1858]" captionTargetId="figure-74@170.[416,1172,265,1864]" captionTargetPageId="170" captionText="Figs 570–585. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 570– 572. M. aiyura sp. nov., holotype, ♂ (BMNH). 573–575. M. alesbezdeki sp. nov., holotype, ♂ (NME). 576–578. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 579–581. M. awu sp. nov., holotype, ♂ (IECA). 582–585. M. babyrousa sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122232" httpUri="https://zenodo.org/record/4122232/files/figure.png" pageId="21" pageNumber="22">Figs 576–578</figureCitation>
.
</paragraph>
<paragraph id="922136E09D09924EFF2EFBA6C9E62C50" blockId="21.[189,432,1048,1074]" box="[189,432,1048,1074]" pageId="21" pageNumber="22">
<heading id="C969818C9D09924EFF2EFBA6C9E62C50" bold="true" box="[189,432,1048,1074]" fontSize="11" level="3" pageId="21" pageNumber="22" reason="3">
<emphasis id="A0EAEAF29D09924EFF2EFBA6C9E62C50" bold="true" box="[189,432,1048,1074]" pageId="21" pageNumber="22">Sexual dimorphism</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D09924EFF2EFBF9C9FD2C03" blockId="21.[189,427,1095,1121]" box="[189,427,1095,1121]" pageId="21" pageNumber="22">Female is unknown.</paragraph>
</subSubSection>
</treatment>
</document>
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