treatments-xml/data/03/F6/87/03F687FE435EBC750CD7FF69FAA174E1.xml

<document id="F2D2FC0CD6AAF2B9D0EFF939186A870C" ID-CLB-Dataset="2609" ID-DOI="10.11646/zootaxa.5091.2.3" ID-GBIF-Dataset="36ace73e-babc-4c20-a78f-fa772ce5f539" ID-ISSN="1175-5326" ID-Zenodo-Dep="5843837" ID-ZooBank="C8081B60-C872-4A19-9291-22A42DC8B763" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1642061981252" checkinUser="plazi" docAuthor="Mai, Zuqi, Hu, Jian, Minoshima, Yûsuke N., Jia, Fenglong &amp; Fikáček, Martin" docDate="2022" docId="03F687FE435EBC750CD7FF69FAA174E1" docLanguage="en" docName="zootaxa.5091.2.3.pdf" docOrigin="Zootaxa 5091 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Dactylosternum latum Fikacek et al. 2015" docType="treatment" docVersion="7" lastPageNumber="289" masterDocId="FFCFFF86434EBC610C40FFFEFF88770C" masterDocTitle="Review of Dactylosternum Wollaston, 1854 from China and Japan (Coleoptera, Hydrophilidae, Sphaeridiinae)" masterLastPageNumber="300" masterPageNumber="269" pageNumber="285" updateTime="1699302224171" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="4F9F620E4381DBAAD4B4DA9FAA8E3888">Review of Dactylosternum Wollaston, 1854 from China and Japan (Coleoptera, Hydrophilidae, Sphaeridiinae)</mods:title>
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<mods:namePart id="78774DC2C9FB1ABADB9F16B1C592CEC0">Mai, Zuqi</mods:namePart>
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<mods:affiliation id="055E22331691D9FAEF5AAD735054E4EC">School of Agriculture, Sun Yat-sen University, Guangzhou, 511436, Guangdong, China. &amp; maizq 5 @ mail 2. sysu. edu. cn; http: // orcid. org / 0000 - 0003 - 3124 - 2021</mods:affiliation>
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<mods:namePart id="5FD15986AE0BFC91F793FDC7DD9A8538">Hu, Jian</mods:namePart>
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<mods:affiliation id="33A1BAB4C2558C4C0A91F4DB8E65C2D8">School of Agriculture, Sun Yat-sen University, Guangzhou, 511436, Guangdong, China. &amp; lsshj @ mail. sysu. edu. cn; http: // orcid. org / 0000 - 0002 - 4122 - 8533</mods:affiliation>
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<mods:namePart id="1001F067F3DF86FA15AFBA144D93719F">Minoshima, Yûsuke N.</mods:namePart>
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<mods:affiliation id="2FA2F8961A9707A54A2C275577E345D2">Natural History Division, Kitakyushu Museum of Natural History and Human History, 2 - 4 - 1 Higashida, Yahatahigashi-ku, Kitakyushushi, Fukuoka, 805 - 0071 Japan. minoshima @ kmnh. jp; http: // orcid. org / 0000 - 0002 - 2575 - 4082</mods:affiliation>
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<mods:namePart id="8604758AED3E2E863E5EFCB191E1F8B8">Jia, Fenglong</mods:namePart>
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<mods:affiliation id="224B4474FF9C4CECDE9ACBAEE22A78E5">School of Agriculture, Sun Yat-sen University, Guangzhou, 511436, Guangdong, China. &amp; Institute of Entomology, Life Sciences School, Sun Yat-sen University, Guangzhou, 510275, Guangdong, China. lssjfl @ mail. sysu. edu. cn; fenglongjia @ alilyun. com; http: // orcid. org / 0000 - 0003 - 2391 - 5038</mods:affiliation>
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<mods:namePart id="CA39502D028175504B6CFD1308E516A2">Fikáček, Martin</mods:namePart>
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<mods:affiliation id="42855B36F32D5CAE4E04CA3DF661E6F7">Department of Biological Sciences, National Sun Yat-sen University, No. 70 Lien-hai Rd., Kaohsiung City 80424. mfikacek @ gmail. com; http: // orcid. org / 0000 - 0002 - 2078 - 6798 &amp; National Museum, Department of Entomology, Cirkusová 1740, CZ- 19300 Praha 9, Czech Republic &amp; Department of Zoology, Faculty of Science, Charles University in Prague, Viničná 7, CZ- 12843 Praha 2, Czech Republic Corresponding author</mods:affiliation>
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<taxonomicName id="4C5F4D6B435EBC710CD7FF69FDDD77BD" ID-CoL="33WWS" authority="(Sharp, 1873)" authorityName="Fikacek et al." authorityYear="2015" baseAuthorityName="Sharp" baseAuthorityYear="1873" box="[151,597,151,178]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="285" phylum="Arthropoda" rank="species" species="latum">
<emphasis id="B92BEAFA435EBC710CD7FF69FDDD77BD" bold="true" box="[151,597,151,178]" italics="true" pageId="16" pageNumber="285">
Dactylosternum latum (
<bibRefCitation id="EFCE4B19435EBC710DF4FF69FDC577BD" author="Sharp, D." box="[436,589,151,177]" pageId="16" pageNumber="285" pagination="64 - 65" refId="ref23391" refString="Sharp, D. (1873) The water beetles of Japan. Transactions of the Entomological Society of London, 1874, 64 - 65." type="journal article" year="1873">Sharp, 1873</bibRefCitation>
)
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<paragraph id="8BE036E8435EBC710CD7FF45FE4077DA" blockId="16.[151,597,151,214]" box="[151,456,187,214]" pageId="16" pageNumber="285">
(
<figureCitation id="13642A6D435EBC710CDFFF45FF6777D9" box="[159,239,187,213]" captionStart="FIGURES 5–10" captionStartId="8.[151,264,1933,1957]" captionTargetBox="[260,1303,188,1900]" captionTargetId="figure-30@8.[214,1373,181,1908]" captionTargetPageId="8" captionText="FIGURES 5–10. Habitus of Dactylosternum spp. from China and Japan. 5: D. songxiaobini sp. nov., 6: D. hydrophiloides (MacLeay, 1825), 7: D. pseudolatum sp. nov., 8: D. salvazai Orchymont, 1925, 9: D. latum (Sharp, 1873), 10: D. frater sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5843841" httpUri="https://zenodo.org/record/5843841/files/figure.png" pageId="16" pageNumber="285">Figs. 9</figureCitation>
, 
<figureCitation id="13642A6D435EBC710CBBFF45FECB77D9" box="[251,323,187,213]" captionStart-0="FIGURES 30–32" captionStart-1="FIGURES 33–38" captionStartId-0="17.[151,264,1701,1725]" captionStartId-1="18.[151,264,1947,1971]" captionTargetBox-0="[223,1377,191,1614]" captionTargetBox-1="[225,1404,347,1884]" captionTargetId-0="figure-19@17.[151,1436,181,1677]" captionTargetId-1="figure-85@18.[151,1436,309,1924]" captionTargetPageId-0="17" captionTargetPageId-1="18" captionText-0="FIGURES 30–32. Aedeagus of D. latum. 30: from Yunnan, 31: from Hunan, 32: syntype (30a, 31a, 32a: dorsal view, 30b, 31b: dorsal view, 30c, 31c: ventral plate of the median lobe, 32b: labels of syntype)" captionText-1="FIGURES 33–38. Aedeagus of D. latum (dorsal view). 33: from Hunan, Shimen County, 34: syntype, 35: from Guangxi, 36: holotype of D. vitalisi, 37: from Hunan, Shimen County, 38: from Yunnan" figureDoi-0="http://doi.org/10.5281/zenodo.5843869" figureDoi-1="http://doi.org/10.5281/zenodo.5843873" httpUri-0="https://zenodo.org/record/5843869/files/figure.png" httpUri-1="https://zenodo.org/record/5843873/files/figure.png" pageId="16" pageNumber="285">30–38</figureCitation>
, 
<figureCitation id="13642A6D435EBC710D0FFF45FE4877DA" box="[335,448,187,214]" captionStart="FIGURES 48–63" captionStartId="21.[151,264,1495,1519]" captionTargetBox="[153,1430,183,1463]" captionTargetId="figure-19@21.[151,1436,181,1471]" captionTargetPageId="21" captionText="FIGURES 48–63. Morphology characters of Dactylosternum spp. SEM micrographs. 48–50: mediobasal portion of elytron (48: D. songxiaobini sp. nov. (red arrows show primary serial punctures, green arrows show secondary serial punctures), 49: D. nanlingensis sp. nov., 50: D. corbetti), 51–55: punctures on posterior half of elytron (51: D. hydrophiloides (the red arrow indicates the ground puncture, the blue arrow indicates the micropuncture), 52: D. pseudolatum sp. nov., 53: D. salvazai, 54: D. latum, 55: D. frater sp. nov.), 56–59: anteromedial portion of metaventrite (56: D. pseudolatum sp. nov., 57: D. salvazai, 58: D. latum, 59: D. frater sp. nov.), 60–63: posteromedial portion of ventrite 5 (60: D. pseudolatum sp. nov., 61: D. salvazai, 62: D. latum, 63: D. frater sp. nov.)" figureDoi="http://doi.org/10.5281/zenodo.5843883" httpUri="https://zenodo.org/record/5843883/files/figure.png" pageId="16" pageNumber="285">54, 58, 62</figureCitation>
)
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<treatmentCitationGroup id="AB4F11C6435EBC710CD7FEFEFC177614" box="[151,927,255,280]" pageId="16" pageNumber="285">
<treatmentCitation id="0AFE10F9435EBC710CD7FEFEFE74761B" author="Sharp, D." box="[151,508,255,280]" page="64" pageId="16" pageNumber="285" year="1873">
<taxonomicName id="4C5F4D6B435EBC710CD7FEFEFE74761B" ID-CoL="32V82" authority="Sharp, 1873: 64" authorityName="Sharp" authorityPageNumber="64" authorityYear="1873" box="[151,508,255,280]" class="Insecta" family="Hydrophilidae" genus="Cyclonotum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="285" phylum="Arthropoda" rank="species" species="latum">
<emphasis id="B92BEAFA435EBC710CD7FEFEFEDA7614" box="[151,338,256,280]" italics="true" pageId="16" pageNumber="285">Cyclonotum latum</emphasis>
<materialsCitation id="3B373CB5435EBC710D18FEFEFE74761B" ID-GBIF-Occurrence="3436469395" box="[344,508,255,280]" country="Japan" location="Kyushu" pageId="16" pageNumber="285" specimenCount="1" stateProvince="Nagasaki">
<bibRefCitation id="EFCE4B19435EBC710D18FEFEFE74761B" author="Sharp, D." box="[344,508,255,280]" pageId="16" pageNumber="285" pagination="64 - 65" refId="ref23391" refString="Sharp, D. (1873) The water beetles of Japan. Transactions of the Entomological Society of London, 1874, 64 - 65." type="journal article" year="1873">Sharp, 1873: 64</bibRefCitation>
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</treatmentCitation>
. 
<typeStatus id="54E4884A435EBC710E45FEFEFDBF7614" box="[517,567,256,280]" pageId="16" pageNumber="285">Type</typeStatus>
locality: 
<collectingCountry id="F3487678435EBC710ED7FEFEFD5B761B" box="[663,723,256,279]" name="Japan" pageId="16" pageNumber="285">Japan</collectingCountry>
, 
<location id="8E806033435EBC710E9EFEFEFCA77614" LSID="urn:lsid:plazi:treatment:03F687FE435EBC750CD7FF69FAA174E1:8E806033435EBC710E9EFEFEFCA77614" box="[734,815,256,280]" country="Japan" name="Kyushu" pageId="16" pageNumber="285" stateProvince="Nagasaki">Kyushu</location>
, 
<collectingRegion id="499BF80A435EBC710F79FEFEFC137614" box="[825,923,256,280]" country="Japan" name="Nagasaki" pageId="16" pageNumber="285">Nagasaki</collectingRegion>
.
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<treatmentCitationGroup id="AB4F11C6435EBC710CD7FEE0FD08763A" box="[151,640,286,310]" pageId="16" pageNumber="285">
<taxonomicName id="4C5F4D6B435EBC710CD7FEE0FE2E763A" authority="(Sharp)" baseAuthorityName="Sharp" box="[151,422,286,310]" class="Insecta" family="Hydrophilidae" genus="Coelostoma" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="285" phylum="Arthropoda" rank="species" species="latum">
<emphasis id="B92BEAFA435EBC710CD7FEE0FED9763A" box="[151,337,286,310]" italics="true" pageId="16" pageNumber="285">Coelostoma latum</emphasis>
(Sharp)
</taxonomicName>
: 
<treatmentCitation id="0AFE10F9435EBC710DF0FEE0FDF5763A" author="Zaitzev, F. A." box="[432,637,286,310]" page="403" pageId="16" pageNumber="285" year="1908">
<bibRefCitation id="EFCE4B19435EBC710DF0FEE0FDF5763A" author="Zaitzev, F. A." box="[432,637,286,310]" pageId="16" pageNumber="285" pagination="283 - 420" refId="ref23642" refString="Zaitzev, F. A. (1908) Catalogue des Coleopteres aquatiques des familles Dryopidae, Georyssidae, Cyathoceridae, Heteroceridae et Hydrophilidae. Horae Societatis Entomologicae Sossicae, 38, 283 - 420." type="journal article" year="1908">Zaitzev (1908: 403)</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
<paragraph id="8BE036E8435EBC710CD7FEC3FD587659" blockId="16.[151,1097,255,372]" box="[151,720,317,341]" pageId="16" pageNumber="285">
<taxonomicName id="4C5F4D6B435EBC710CD7FEC3FE477659" ID-CoL="33WWS" authority="(Sharp)" authorityName="Fikacek et al." authorityYear="2015" baseAuthorityName="Sharp" baseAuthorityYear="1873" box="[151,463,317,341]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="285" phylum="Arthropoda" rank="species" species="latum">
<emphasis id="B92BEAFA435EBC710CD7FEC3FEF17659" box="[151,377,317,341]" italics="true" pageId="16" pageNumber="285">Dactylosternum latum</emphasis>
(Sharp)
</taxonomicName>
: Orchymont (1919: 108).
</paragraph>
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<taxonomicName id="4C5F4D6B435EBC710CD7FEA2FDFE7678" ID-CoL="33WXN" authority="Orchymont, 1925: 271" authorityName="d' Orchymont" authorityPageNumber="271" authorityYear="1925" box="[151,630,347,372]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="285" phylum="Arthropoda" rank="species" species="vitalisi">
<emphasis id="B92BEAFA435EBC710CD7FEA2FE0E7678" box="[151,390,348,372]" italics="true" pageId="16" pageNumber="285">Dactylosternum Vitalisi</emphasis>
Orchymont, 1925: 271
</taxonomicName>
. 
<typeStatus id="54E4884A435EBC710E3FFEA2FD397678" box="[639,689,348,372]" pageId="16" pageNumber="285">Type</typeStatus>
locality: 
<collectingCountry id="F3487678435EBC710F51FEA2FCCD767F" box="[785,837,348,371]" name="Laos" pageId="16" pageNumber="285">Laos</collectingCountry>
, Ban Sai. 
<emphasis id="B92BEAFA435EBC710FEBFEA2FBC17678" bold="true" box="[939,1097,348,372]" pageId="16" pageNumber="285">New synonym.</emphasis>
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<emphasis id="B92BEAFA435EBC710CD7FE5CFE4F76B7" bold="true" box="[151,455,417,444]" pageId="16" pageNumber="285">Type material examined:</emphasis>
<taxonomicName id="4C5F4D6B435EBC710D90FE5CFD4576B0" authorityName="Fikacek et al." authorityYear="2015" baseAuthorityName="Sharp" baseAuthorityYear="1873" box="[464,717,418,444]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="285" phylum="Arthropoda" rank="species" species="latum">
<emphasis id="B92BEAFA435EBC710D90FE5CFD4576B0" box="[464,717,418,444]" italics="true" pageId="16" pageNumber="285">Dactylosternum latum</emphasis>
</taxonomicName>
: 
<materialsCitation id="3B373CB5435EBC710E9EFE5FFDA37508" ID-GBIF-Occurrence="3436469305" collectionCode="BMNH" country="Japan" determinerName="YN Minoshima" location="Japan" pageId="16" pageNumber="285" specimenCount="1" specimenCount-male="1" typeStatus="syntype">
<emphasis id="B92BEAFA435EBC710E9EFE5FFCF176B7" bold="true" box="[734,889,417,443]" pageId="16" pageNumber="285">
<typeStatus id="54E4884A435EBC710E9EFE5FFCFD76B7" box="[734,885,417,443]" pageId="16" pageNumber="285" type="syntype">SYNTYPES</typeStatus>
:
</emphasis>
<specimenCount id="9D59FD61435EBC710FC3FE5CFC5976B0" box="[899,977,418,444]" pageId="16" pageNumber="285" type="male">1 male</specimenCount>
(
<collectionCode id="ED4EAE2D435EBC710FA3FE5CFBB676B0" box="[995,1086,418,444]" pageId="16" pageNumber="285">BMNH</collectionCode>
): “ 
<collectingCountry id="F3487678435EBC71081CFE5CFB1776B0" box="[1116,1183,418,444]" name="Japan" pageId="16" pageNumber="285">Japan</collectingCountry>
/ Lewis // Sharp Coll / 1905–313 // Type // 
<taxonomicName id="4C5F4D6B435EBC710DD4FE38FDE076EC" authorityName="Sharp" authorityYear="1873" box="[404,616,454,480]" class="Insecta" family="Hydrophilidae" genus="Cyclonotum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="285" phylum="Arthropoda" rank="species" species="latum">Cyclonotum latum</taxonomicName>
/ type D.S. // 
<typeStatus id="54E4884A435EBC710F4FFE38FC0576EC" box="[783,909,454,480]" pageId="16" pageNumber="285" type="syntype">SYNTYPE</typeStatus>
/ 
<taxonomicName id="4C5F4D6B435EBC710FE8FE38FBE676EC" box="[936,1134,454,480]" class="Insecta" family="Hydrophilidae" genus="Cyclonotum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="285" phylum="Arthropoda" rank="genus">CYCLONOTUM</taxonomicName>
/ 
<taxonomicName id="4C5F4D6B435EBC7108C8FE38FAD276EC" authority="Sharp, 1873" authorityName="Sharp" authorityYear="1873" box="[1160,1370,454,480]" class="Insecta" family="Hydrophilidae" genus="Cyclonotum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="285" phylum="Arthropoda" rank="species" species="latum">
latum 
<bibRefCitation id="EFCE4B19435EBC710891FE38FAD276EC" author="Sharp, D." box="[1233,1370,454,480]" pageId="16" pageNumber="285" pagination="64 - 65" refId="ref23391" refString="Sharp, D. (1873) The water beetles of Japan. Transactions of the Entomological Society of London, 1874, 64 - 65." type="journal article" year="1873">Sharp, 1873</bibRefCitation>
</taxonomicName>
/ 
<determinerName id="7E3CDA2C435EBC710934FE38FE9F7508" pageId="16" pageNumber="285">YN Minoshima</determinerName>
det. 2021” (
<figureCitation id="13642A6D435EBC710DE0FE14FE7F7508" box="[416,503,490,516]" captionStart="FIGURES 30–32" captionStartId="17.[151,264,1701,1725]" captionTargetBox="[223,1377,191,1614]" captionTargetId="figure-19@17.[151,1436,181,1677]" captionTargetPageId="17" captionText="FIGURES 30–32. Aedeagus of D. latum. 30: from Yunnan, 31: from Hunan, 32: syntype (30a, 31a, 32a: dorsal view, 30b, 31b: dorsal view, 30c, 31c: ventral plate of the median lobe, 32b: labels of syntype)" figureDoi="http://doi.org/10.5281/zenodo.5843869" httpUri="https://zenodo.org/record/5843869/files/figure.png" pageId="16" pageNumber="285">Figs 32</figureCitation>
, 
<figureCitation id="13642A6D435EBC710E42FE14FDAA7508" box="[514,546,490,516]" captionStart="FIGURES 33–38" captionStartId="18.[151,264,1947,1971]" captionTargetBox="[225,1404,347,1884]" captionTargetId="figure-85@18.[151,1436,309,1924]" captionTargetPageId="18" captionText="FIGURES 33–38. Aedeagus of D. latum (dorsal view). 33: from Hunan, Shimen County, 34: syntype, 35: from Guangxi, 36: holotype of D. vitalisi, 37: from Hunan, Shimen County, 38: from Yunnan" figureDoi="http://doi.org/10.5281/zenodo.5843873" httpUri="https://zenodo.org/record/5843873/files/figure.png" pageId="16" pageNumber="285">34</figureCitation>
)
</materialsCitation>
; 
<materialsCitation id="3B373CB5435EBC710E76FE14FBEF7524" ID-GBIF-Occurrence="3436469389" collectionCode="BMNH" collectorName="G. Lewis" country="Japan" determinerName="YN Minoshima" location="Japan" pageId="16" pageNumber="285" specimenCount="1" specimenCount-male="1" typeStatus="syntype">
<specimenCount id="9D59FD61435EBC710E76FE14FD097508" box="[566,641,490,516]" pageId="16" pageNumber="285" type="male">1 male</specimenCount>
(
<collectionCode id="ED4EAE2D435EBC710ED0FE14FD627508" box="[656,746,490,516]" pageId="16" pageNumber="285">BMNH</collectionCode>
), “ 
<collectingCountry id="F3487678435EBC710F46FE14FCC17508" box="[774,841,490,516]" name="Japan" pageId="16" pageNumber="285">Japan</collectingCountry>
/ 
<collectorName id="26AA533E435EBC710F1EFE14FC4C7508" box="[862,964,490,516]" pageId="16" pageNumber="285">G. Lewis</collectorName>
/ 1910—320 // 
<taxonomicName id="4C5F4D6B435EBC710822FE14FB647508" box="[1122,1260,490,516]" class="Insecta" family="Hydrophilidae" genus="Cyclonotum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="285" phylum="Arthropoda" rank="genus">Cyclonotum</taxonomicName>
/ latum / mihi D.S. // 
<typeStatus id="54E4884A435EBC710CA7FDF0FEED7524" box="[231,357,526,552]" pageId="16" pageNumber="285" type="syntype">SYNTYPE</typeStatus>
/ 
<taxonomicName id="4C5F4D6B435EBC710D3AFDF0FDC87524" box="[378,576,526,552]" class="Insecta" family="Hydrophilidae" genus="Cyclonotum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="285" phylum="Arthropoda" rank="genus">CYCLONOTUM</taxonomicName>
/ 
<taxonomicName id="4C5F4D6B435EBC710E16FDF0FCAC7524" authority="Sharp, 1873" authorityName="Sharp" authorityYear="1873" box="[598,804,526,552]" class="Insecta" family="Hydrophilidae" genus="Cyclonotum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="285" phylum="Arthropoda" rank="species" species="latum">
latum 
<bibRefCitation id="EFCE4B19435EBC710EDCFDF0FCAC7524" author="Sharp, D." box="[668,804,526,552]" pageId="16" pageNumber="285" pagination="64 - 65" refId="ref23391" refString="Sharp, D. (1873) The water beetles of Japan. Transactions of the Entomological Society of London, 1874, 64 - 65." type="journal article" year="1873">Sharp, 1873</bibRefCitation>
</taxonomicName>
/ 
<determinerName id="7E3CDA2C435EBC710F78FDF0FC607524" box="[824,1000,526,552]" pageId="16" pageNumber="285">YN Minoshima</determinerName>
det. 2021”
</materialsCitation>
.
</paragraph>
<paragraph id="8BE036E8435EBC710C87FDCCFE2175B4" blockId="16.[151,1437,417,1957]" pageId="16" pageNumber="285">
<taxonomicName id="4C5F4D6B435EBC710C87FDCCFE477540" authorityName="d' Orchymont" authorityYear="1925" box="[199,463,562,588]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="285" phylum="Arthropoda" rank="species" species="vitalisi">
<emphasis id="B92BEAFA435EBC710C87FDCCFE477540" box="[199,463,562,588]" italics="true" pageId="16" pageNumber="285">Dactylosternum vitalisi</emphasis>
</taxonomicName>
: 
<materialsCitation id="3B373CB5435EBC710DA0FDCFFF487599" ID-GBIF-Occurrence="3436469356" collectingDate="1918-08-12" collectionCode="IRSNB" country="Laos" determinerName="A. d'Orchymont" location="Ban Sai" pageId="16" pageNumber="285" specimenCount="1" typeStatus="holotype">
<emphasis id="B92BEAFA435EBC710DA0FDCFFD007547" bold="true" box="[480,648,561,587]" pageId="16" pageNumber="285">
<typeStatus id="54E4884A435EBC710DA0FDCFFD0B7547" box="[480,643,561,587]" pageId="16" pageNumber="285" type="holotype">HOLOTYPE</typeStatus>
:
</emphasis>
male (
<collectionCode id="ED4EAE2D435EBC710E98FDCCFCB97540" box="[728,817,562,588]" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="16" pageNumber="285">IRSNB</collectionCode>
): “ 
<collectingCountry id="F3487678435EBC710F11FDCCFC067540" box="[849,910,562,588]" name="Laos" pageId="16" pageNumber="285">Laos</collectingCountry>
: / 
<location id="8E806033435EBC710FEAFDCCFB8C7540" LSID="urn:lsid:plazi:treatment:03F687FE435EBC750CD7FF69FAA174E1:8E806033435EBC710FEAFDCCFB8C7540" box="[938,1028,562,588]" country="Laos" name="Ban Sai" pageId="16" pageNumber="285">Ban Sai</location>
/ 
<date id="FFE11028435EBC71085EFDCCFB067540" box="[1054,1166,562,588]" pageId="16" pageNumber="285" value="1918-08-12">
<collectingDate id="EFA5E9C0435EBC71085EFDCCFB067540" box="[1054,1166,562,588]" pageId="16" pageNumber="285" value="1918-08-12">8.12.1918</collectingDate>
</date>
/ Vitalis // Ban Sai / 8- XII-18 / No 2199 // TYPE // TYPE // 
<determinerName id="7E3CDA2C435EBC710E0DFDA8FC81757C" box="[589,777,598,624]" pageId="16" pageNumber="285">A. d´Orchymont</determinerName>
det. / 
<taxonomicName id="4C5F4D6B435EBC710F14FDA8FBC0757C" authorityName="Wollaston" authorityYear="1854" box="[852,1096,598,624]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="285" phylum="Arthropoda" rank="genus">Dactylosternum 1922</taxonomicName>
/ 
<taxonomicName id="4C5F4D6B435EBC710821FDA8FADC757C" authority="d'Orchymont" authorityName="d' Orchymont" authorityYear="1925" box="[1121,1364,598,624]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="285" phylum="Arthropoda" rank="species" species="vitalisi">Vitalisi d´Orchymont</taxonomicName>
” (
<figureCitation id="13642A6D435EBC71092FFDA8FF3F7599" captionStart="FIGURES 33–38" captionStartId="18.[151,264,1947,1971]" captionTargetBox="[225,1404,347,1884]" captionTargetId="figure-85@18.[151,1436,309,1924]" captionTargetPageId="18" captionText="FIGURES 33–38. Aedeagus of D. latum (dorsal view). 33: from Hunan, Shimen County, 34: syntype, 35: from Guangxi, 36: holotype of D. vitalisi, 37: from Hunan, Shimen County, 38: from Yunnan" figureDoi="http://doi.org/10.5281/zenodo.5843873" httpUri="https://zenodo.org/record/5843873/files/figure.png" pageId="16" pageNumber="285">Fig. 36</figureCitation>
)
</materialsCitation>
; 
<materialsCitation id="3B373CB5435EBC710C8BFD87FE2175B4" ID-GBIF-Occurrence="3436469301" collectionCode="BMNH" country="Laos" determinerName="A. d' Orchymont" pageId="16" pageNumber="285" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<emphasis id="B92BEAFA435EBC710C8BFD87FED7759F" bold="true" box="[203,351,633,659]" pageId="16" pageNumber="285">
<typeStatus id="54E4884A435EBC710C8BFD87FED7759F" box="[203,351,633,659]" pageId="16" pageNumber="285" type="paratype">PARATYPE</typeStatus>
</emphasis>
: 
<specimenCount id="9D59FD61435EBC710D2EFD84FE317598" box="[366,441,634,660]" pageId="16" pageNumber="285" type="male">1 male</specimenCount>
(
<collectionCode id="ED4EAE2D435EBC710D89FD84FDAC7598" box="[457,548,634,660]" pageId="16" pageNumber="285">BMNH</collectionCode>
): “Fokien // 
<determinerName id="7E3CDA2C435EBC710EEDFD84FCEC7598" box="[685,868,634,660]" pageId="16" pageNumber="285">A.d’ Orchymont</determinerName>
det / 
<taxonomicName id="4C5F4D6B435EBC710FE3FD84FB1D7598" authorityName="Wollaston" authorityYear="1854" box="[931,1173,634,660]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="285" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEAFA435EBC710FE3FD84FBDE7598" box="[931,1110,634,660]" italics="true" pageId="16" pageNumber="285">Dactylosternum</emphasis>
1922
</taxonomicName>
/ 
<taxonomicName id="4C5F4D6B435EBC7108EBFD84FA157598" authority="d' Orchymont" authorityName="d' Orchymont" authorityYear="1925" box="[1195,1437,634,660]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="285" phylum="Arthropoda" rank="species" species="vitalisi">
<emphasis id="B92BEAFA435EBC7108EBFD84FB727598" box="[1195,1274,634,660]" italics="true" pageId="16" pageNumber="285">Vitalisi</emphasis>
d’ Orchymont
</taxonomicName>
// Co-type // co- TYPE.”
</materialsCitation>
</paragraph>
<paragraph id="8BE036E8435EBC710C87FD3CFE897460" blockId="16.[151,1437,417,1957]" pageId="16" pageNumber="285">
<emphasis id="B92BEAFA435EBC710C87FD3CFE8E75D0" bold="true" box="[199,262,706,732]" pageId="16" pageNumber="285">Note.</emphasis>
<materialsCitation id="3B373CB5435EBC710D4DFD3CFE927428" ID-GBIF-Occurrence="3436469329" county="Sharp" location="On" municipality="Japanese" pageId="16" pageNumber="285" specimenCount="1" typeStatus="syntype">
<location id="8E806033435EBC710D4DFD3CFEA775D0" LSID="urn:lsid:plazi:treatment:03F687FE435EBC750CD7FF69FAA174E1:8E806033435EBC710D4DFD3CFEA775D0" box="[269,303,706,732]" county="Sharp" municipality="Japanese" name="On" pageId="16" pageNumber="285">On</location>
the request of loan of the type specimens of 
<taxonomicName id="4C5F4D6B435EBC710F67FD3DFC0175D0" authorityName="Sharp" authorityYear="1873" box="[807,905,706,732]" class="Insecta" family="Hydrophilidae" genus="Cyclonotum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="285" phylum="Arthropoda" rank="species" species="latum">
<emphasis id="B92BEAFA435EBC710F67FD3DFC0175D0" box="[807,905,706,732]" italics="true" pageId="16" pageNumber="285">D. latum</emphasis>
</taxonomicName>
, we received two 
<collectingMunicipality id="6B84AC92435EBC710816FD3CFB3275D0" box="[1110,1210,706,732]" pageId="16" pageNumber="285">Japanese</collectingMunicipality>
specimens of 
<collectingCounty id="62814E64435EBC71091BFD3CFA1475D0" box="[1371,1436,706,732]" pageId="16" pageNumber="285">Sharp</collectingCounty>
collection from BMNH. 
<bibRefCitation id="EFCE4B19435EBC710DE9FD18FDB3740C" author="Sharp, D." box="[425,571,742,768]" pageId="16" pageNumber="285" pagination="64 - 65" refId="ref23391" refString="Sharp, D. (1873) The water beetles of Japan. Transactions of the Entomological Society of London, 1874, 64 - 65." type="journal article" year="1873">Sharp (1873)</bibRefCitation>
mentioned that he examined 
<specimenCount id="9D59FD61435EBC710FC2FD18FBA2740C" box="[898,1066,742,768]" pageId="16" pageNumber="285" type="generic">two specimens</specimenCount>
, therefore they can be considered as 
<typeStatus id="54E4884A435EBC710CF4FCF4FE927428" box="[180,282,778,804]" pageId="16" pageNumber="285" type="syntype">syntypes</typeStatus>
</materialsCitation>
. 
<materialsCitation id="3B373CB5435EBC710D63FCF4FEAA7444" ID-GBIF-Occurrence="3436469309" country="Japan" location="Both" municipality="The" pageId="16" pageNumber="285" specimenCount="1" typeStatus="syntype">
<location id="8E806033435EBC710D63FCF4FED17428" LSID="urn:lsid:plazi:treatment:03F687FE435EBC750CD7FF69FAA174E1:8E806033435EBC710D63FCF4FED17428" box="[291,345,778,804]" country="Japan" municipality="The" name="Both" pageId="16" pageNumber="285">Both</location>
specimens are male, morphologically identical and with labels indicating that they are from 
<collectingCountry id="F3487678435EBC710915FCF4FA107428" box="[1365,1432,778,804]" name="Japan" pageId="16" pageNumber="285">Japan</collectingCountry>
. 
<collectingMunicipality id="6B84AC92435EBC710CD7FCD0FF4B7444" box="[151,195,814,840]" pageId="16" pageNumber="285">The</collectingMunicipality>
<typeStatus id="54E4884A435EBC710C8BFCD0FEAA7444" box="[203,290,814,840]" pageId="16" pageNumber="285" type="syntype">syntype</typeStatus>
</materialsCitation>
status of both specimens does not cause any taxonomic confusion and we hence retain their status of 
<typeStatus id="54E4884A435EBC710CD7FCACFF757460" box="[151,253,850,876]" pageId="16" pageNumber="285" type="syntype">syntypes</typeStatus>
.
</paragraph>
<paragraph id="8BE036E8435EBC710C87FC8BFC9573BC" blockId="16.[151,1437,417,1957]" pageId="16" pageNumber="285">
<materialsCitation id="3B373CB5435EBC710C87FC8BFD9074B8" ID-GBIF-Occurrence="3436469391" collectingDate="2013-07-13" collectionCode="SYSU" collectorName="Qiu &amp; Xu" country="China" county="Tangkou County" location="Hougu" municipality="Huangshan Prefecture" pageId="16" pageNumber="285" specimenCount="1" specimenCount-female="1" stateProvince="Anhui">
<emphasis id="B92BEAFA435EBC710C87FC8BFD747483" bold="true" box="[199,764,885,911]" pageId="16" pageNumber="285">
Additional material examined. 
<collectingCountry id="F3487678435EBC710E7CFC8BFD167483" box="[572,670,885,911]" name="China" pageId="16" pageNumber="285">CHINA</collectingCountry>
: 
<collectingRegion id="499BF80A435EBC710EE9FC8BFD707483" box="[681,760,885,911]" country="China" name="Anhui" pageId="16" pageNumber="285">Anhui</collectingRegion>
:
</emphasis>
<specimenCount id="9D59FD61435EBC710F44FC88FCED749C" box="[772,869,886,912]" pageId="16" pageNumber="285" type="female">1 female</specimenCount>
(
<collectionCode id="ED4EAE2D435EBC710F35FC88FC4A749C" box="[885,962,886,912]" pageId="16" pageNumber="285">SYSU</collectionCode>
), 
<collectingMunicipality id="6B84AC92435EBC710F95FC88FB59749C" box="[981,1233,886,912]" pageId="16" pageNumber="285">Huangshan Prefecture</collectingMunicipality>
, 
<collectingCounty id="62814E64435EBC71089CFC88FA11749C" box="[1244,1433,886,912]" pageId="16" pageNumber="285">Tangkou County</collectingCounty>
, 
<location id="8E806033435EBC710CD7FC64FF6E74B8" LSID="urn:lsid:plazi:treatment:03F687FE435EBC750CD7FF69FAA174E1:8E806033435EBC710CD7FC64FF6E74B8" box="[151,230,922,948]" country="China" county="Tangkou County" municipality="Huangshan Prefecture" name="Hougu" pageId="16" pageNumber="285" stateProvince="Anhui">Hougu</location>
, 
<date id="FFE11028435EBC710CADFC64FEF174B8" box="[237,377,922,948]" pageId="16" pageNumber="285" value="2013-07-13">
<collectingDate id="EFA5E9C0435EBC710CADFC64FEF174B8" box="[237,377,922,948]" pageId="16" pageNumber="285" value="2013-07-13">13.VII.2013</collectingDate>
</date>
, 
<collectorName id="26AA533E435EBC710DC0FC64FE2274B8" box="[384,426,922,948]" pageId="16" pageNumber="285">Qiu</collectorName>
&amp; 
<collectorName id="26AA533E435EBC710D86FC64FE6074B8" box="[454,488,922,948]" pageId="16" pageNumber="285">Xu</collectorName>
leg.
</materialsCitation>
; 
<materialsCitation id="3B373CB5435EBC710E60FC67FEE474D4" ID-GBIF-Occurrence="3436469404" collectingDate="2002-05-27" collectionCode="HBUM" collectorName="Xiujuan Yang" country="China" county="Tianlin County" elevation="1250" location="Jiudongpin" pageId="16" pageNumber="285" specimenCount="1" specimenCount-male="1" stateProvince="Guangxi">
<emphasis id="B92BEAFA435EBC710E60FC67FD1874BF" bold="true" box="[544,656,921,947]" pageId="16" pageNumber="285">
<collectingRegion id="499BF80A435EBC710E60FC67FD0374BF" box="[544,651,921,947]" country="China" name="Guangxi" pageId="16" pageNumber="285">Guangxi</collectingRegion>
:
</emphasis>
<specimenCount id="9D59FD61435EBC710ED3FC64FD5274B8" box="[659,730,922,948]" pageId="16" pageNumber="285" type="male">1male</specimenCount>
(
<collectionCode id="ED4EAE2D435EBC710EA5FC64FCB774B8" box="[741,831,922,948]" country="China" name="College of Life Sciences Hebei Univesity, Baoding" pageId="16" pageNumber="285">HBUM</collectionCode>
), 
<collectingCounty id="62814E64435EBC710F0DFC64FC7C74B8" box="[845,1012,922,948]" pageId="16" pageNumber="285">Tianlin County</collectingCounty>
, 
<location id="8E806033435EBC710FBBFC64FBF274B8" LSID="urn:lsid:plazi:treatment:03F687FE435EBC750CD7FF69FAA174E1:8E806033435EBC710FBBFC64FBF274B8" box="[1019,1146,922,948]" country="China" county="Tianlin County" name="Jiudongpin" pageId="16" pageNumber="285" stateProvince="Guangxi">Jiudongpin</location>
, 
<quantity id="4CA79B0D435EBC7108C2FC64FA9374B8" box="[1154,1307,922,948]" metricMagnitude="3" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="16" pageNumber="285" unit="m" value="1250.0" valueMax="1300.0" valueMin="1200.0">
<elevation id="0072D1DB435EBC7108C2FC64FA9374B8" box="[1154,1307,922,948]" metricMagnitude="3" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="16" pageNumber="285" unit="m" value="1250.0" valueMax="1300.0" valueMin="1200.0">1200–1300m</elevation>
</quantity>
, 
<date id="FFE11028435EBC710963FC64FA1074B8" box="[1315,1432,922,948]" pageId="16" pageNumber="285" value="2002-05-27">
<collectingDate id="EFA5E9C0435EBC710963FC64FA1074B8" box="[1315,1432,922,948]" pageId="16" pageNumber="285" value="2002-05-27">27.V.2002</collectingDate>
</date>
, 
<collectorName id="26AA533E435EBC710CD7FC40FEBD74D4" box="[151,309,958,984]" pageId="16" pageNumber="285">Xiujuan Yang</collectorName>
leg.
</materialsCitation>
; 
<materialsCitation id="3B373CB5435EBC710D3CFC40FDAE74F0" ID-GBIF-Occurrence="3436469387" collectingDate="2004-08-17" collectingDateMax="2004-08-20" collectingDateMin="2004-08-17" collectionCode="HBUM" collectorName="Jianfeng Wang &amp; Jiliang Wang" country="China" county="Shimen County" location="Huping Mountain" pageId="16" pageNumber="285" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Hunan">
<emphasis id="B92BEAFA435EBC710D3CFC40FE5074D4" bold="true" box="[380,472,958,984]" pageId="16" pageNumber="285">
<collectingRegion id="499BF80A435EBC710D3CFC40FE5B74D4" box="[380,467,958,984]" country="China" name="Hunan" pageId="16" pageNumber="285">Hunan</collectingRegion>
:
</emphasis>
<specimenCount id="9D59FD61435EBC710DA3FC40FDC874D4" box="[483,576,958,984]" pageId="16" pageNumber="285" type="male">2 males</specimenCount>
, 
<specimenCount id="9D59FD61435EBC710E0FFC40FD3C74D4" box="[591,692,958,984]" pageId="16" pageNumber="285" type="female">1 female</specimenCount>
(
<collectionCode id="ED4EAE2D435EBC710E87FC40FCA974D4" box="[711,801,958,984]" country="China" name="College of Life Sciences Hebei Univesity, Baoding" pageId="16" pageNumber="285">HBUM</collectionCode>
), 
<collectingCounty id="62814E64435EBC710F78FC40FC6574D4" box="[824,1005,958,984]" pageId="16" pageNumber="285">Shimen County</collectingCounty>
, 
<location id="8E806033435EBC710FBBFC40FB4274D4" LSID="urn:lsid:plazi:treatment:03F687FE435EBC750CD7FF69FAA174E1:8E806033435EBC710FBBFC40FB4274D4" box="[1019,1226,958,984]" country="China" county="Shimen County" name="Huping Mountain" pageId="16" pageNumber="285" stateProvince="Hunan">Huping Mountain</location>
, 
<date id="FFE11028435EBC710899FC40FA1074D4" box="[1241,1432,958,984]" pageId="16" pageNumber="285" value="2004-08-17" valueMax="2004-08-20" valueMin="2004-08-17">
<collectingDate id="EFA5E9C0435EBC710899FC40FA1074D4" box="[1241,1432,958,984]" pageId="16" pageNumber="285" value="2004-08-17" valueMax="2004-08-20" valueMin="2004-08-17">17–20.VIII.2004</collectingDate>
</date>
, 
<collectorName id="26AA533E435EBC710CD7FC1CFEB674F0" box="[151,318,994,1020]" pageId="16" pageNumber="285">Jianfeng Wang</collectorName>
&amp; 
<collectorName id="26AA533E435EBC710D21FC1CFE7B74F0" box="[353,499,994,1020]" pageId="16" pageNumber="285">Jiliang Wang</collectorName>
leg.
</materialsCitation>
; 
<materialsCitation id="3B373CB5435EBC710E72FC1CFECA732C" ID-GBIF-Occurrence="3436469349" collectingDate="2017-06-06" collectionCode="SYSU" collectorName="Qiu &amp; Xu" country="China" county="Shimen County" location="Maozhuhe" municipality="Huping Mountain" pageId="16" pageNumber="285" specimenCount="2" specimenCount-female="2" stateProvince="Hunan">
<specimenCount id="9D59FD61435EBC710E72FC1CFD1674F0" box="[562,670,994,1020]" pageId="16" pageNumber="285" type="female">2 females</specimenCount>
(
<collectionCode id="ED4EAE2D435EBC710EEEFC1CFD7374F0" box="[686,763,994,1020]" pageId="16" pageNumber="285">SYSU</collectionCode>
), 
<collectingCounty id="62814E64435EBC710F4EFC1CFC3774F0" box="[782,959,994,1020]" pageId="16" pageNumber="285">Shimen County</collectingCounty>
, 
<collectingMunicipality id="6B84AC92435EBC710F89FC1CFB1C74F0" box="[969,1172,994,1020]" pageId="16" pageNumber="285">Huping Mountain</collectingMunicipality>
, 
<location id="8E806033435EBC7108DFFC1CFA9074F0" LSID="urn:lsid:plazi:treatment:03F687FE435EBC750CD7FF69FAA174E1:8E806033435EBC7108DFFC1CFA9074F0" box="[1183,1304,994,1020]" country="China" county="Shimen County" municipality="Huping Mountain" name="Maozhuhe" pageId="16" pageNumber="285" stateProvince="Hunan">Maozhuhe</location>
, 
<date id="FFE11028435EBC710964FC1CFA1F74F0" box="[1316,1431,994,1020]" pageId="16" pageNumber="285" value="2017-06-06">
<collectingDate id="EFA5E9C0435EBC710964FC1CFA1F74F0" box="[1316,1431,994,1020]" pageId="16" pageNumber="285" value="2017-06-06">6.VI.2017</collectingDate>
</date>
, 
<collectorName id="26AA533E435EBC710CD7FBF8FF49732C" box="[151,193,1030,1056]" pageId="16" pageNumber="285">Qiu</collectorName>
&amp; 
<collectorName id="26AA533E435EBC710CAAFBF8FE84732C" box="[234,268,1030,1056]" pageId="16" pageNumber="285">Xu</collectorName>
leg.
</materialsCitation>
; 
<materialsCitation id="3B373CB5435EBC710D10FBF8FF4B7348" ID-GBIF-Occurrence="3436469370" collectingDate="2004-07-30" collectionCode="HBUM" collectorName="Jishan Xu &amp; Lingxiao Chang" country="China" county="Fenghuang County" location="Liangtouyang" pageId="16" pageNumber="285" specimenCount="1" specimenCount-male="1" stateProvince="Hunan">
<specimenCount id="9D59FD61435EBC710D10FBF8FE15732C" box="[336,413,1030,1056]" pageId="16" pageNumber="285" type="male">1 male</specimenCount>
(
<collectionCode id="ED4EAE2D435EBC710DEFFBF8FD81732C" box="[431,521,1030,1056]" country="China" name="College of Life Sciences Hebei Univesity, Baoding" pageId="16" pageNumber="285">HBUM</collectionCode>
), 
<collectingCounty id="62814E64435EBC710E5EFBF8FD70732C" box="[542,760,1030,1056]" pageId="16" pageNumber="285">Fenghuang County</collectingCounty>
, 
<location id="8E806033435EBC710F45FBF8FC2B732C" LSID="urn:lsid:plazi:treatment:03F687FE435EBC750CD7FF69FAA174E1:8E806033435EBC710F45FBF8FC2B732C" box="[773,931,1030,1056]" country="China" county="Fenghuang County" name="Liangtouyang" pageId="16" pageNumber="285" stateProvince="Hunan">Liangtouyang</location>
, 
<date id="FFE11028435EBC710FF1FBF8FBB4732C" box="[945,1084,1030,1056]" pageId="16" pageNumber="285" value="2004-07-30">
<collectingDate id="EFA5E9C0435EBC710FF1FBF8FBB4732C" box="[945,1084,1030,1056]" pageId="16" pageNumber="285" value="2004-07-30">30.VII.2004</collectingDate>
</date>
, 
<collectorName id="26AA533E435EBC71080AFBF8FB34732C" box="[1098,1212,1030,1056]" pageId="16" pageNumber="285">Jishan Xu</collectorName>
&amp; 
<collectorName id="26AA533E435EBC7108A5FBF8FA14732C" box="[1253,1436,1030,1056]" pageId="16" pageNumber="285">Lingxiao Chang</collectorName>
leg.
</materialsCitation>
; 
<materialsCitation id="3B373CB5435EBC710C90FBD4FE847364" ID-GBIF-Occurrence="3436469317" collectingDate="2016-05-22" collectionCode="SYSU" country="China" elevation="1751" latitude="24.91" location="Gaoligong Natural Park" longLatPrecision="749" longitude="98.81" municipality="Baoshan Prefecture" pageId="16" pageNumber="285" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Yunnan">
<emphasis id="B92BEAFA435EBC710C90FBD4FEBF7348" bold="true" box="[208,311,1066,1092]" pageId="16" pageNumber="285">
<collectingRegion id="499BF80A435EBC710C90FBD4FEBA7348" box="[208,306,1066,1092]" country="China" name="Yunnan" pageId="16" pageNumber="285">Yunnan</collectingRegion>
:
</emphasis>
<specimenCount id="9D59FD61435EBC710D7FFBD4FE057348" box="[319,397,1066,1092]" pageId="16" pageNumber="285" type="male">1 male</specimenCount>
, 
<specimenCount id="9D59FD61435EBC710DD9FBD4FE737348" box="[409,507,1066,1092]" pageId="16" pageNumber="285" type="female">1 female</specimenCount>
(
<collectionCode id="ED4EAE2D435EBC710E4AFBD4FDDF7348" box="[522,599,1066,1092]" pageId="16" pageNumber="285">SYSU</collectionCode>
), 
<collectingMunicipality id="6B84AC92435EBC710E2BFBD4FCC17348" box="[619,841,1066,1092]" pageId="16" pageNumber="285">Baoshan Prefecture</collectingMunicipality>
, 
<location id="8E806033435EBC710F15FBD4FBEC7348" LSID="urn:lsid:plazi:treatment:03F687FE435EBC750CD7FF69FAA174E1:8E806033435EBC710F15FBD4FBEC7348" box="[853,1124,1066,1092]" country="China" latitude="24.91" longLatPrecision="749" longitude="98.81" municipality="Baoshan Prefecture" name="Gaoligong Natural Park" pageId="16" pageNumber="285" stateProvince="Yunnan">Gaoligong Natural Park</location>
, 
<geoCoordinate id="EE6B502F435EBC710830FBD4FB467348" box="[1136,1230,1066,1092]" degrees="24.91" direction="north" orientation="latitude" pageId="16" pageNumber="285" precision="555" value="24.91">24.91°N</geoCoordinate>
<geoCoordinate id="EE6B502F435EBC710896FBD4FABB7348" box="[1238,1331,1066,1092]" degrees="98.81" direction="east" orientation="longitude" pageId="16" pageNumber="285" precision="555" value="98.81">98.81°E</geoCoordinate>
, 
<quantity id="4CA79B0D435EBC710900FBD4FA1F7348" box="[1344,1431,1066,1093]" metricMagnitude="3" metricUnit="m" metricValue="1.751" pageId="16" pageNumber="285" unit="m" value="1751.0">
<elevation id="0072D1DB435EBC710900FBD4FA1F7348" box="[1344,1431,1066,1093]" metricMagnitude="3" metricUnit="m" metricValue="1.751" pageId="16" pageNumber="285" unit="m" value="1751.0">1751 m</elevation>
</quantity>
, 
<date id="FFE11028435EBC710CD7FBB0FE847364" box="[151,268,1102,1128]" pageId="16" pageNumber="285" value="2016-05-22">
<collectingDate id="EFA5E9C0435EBC710CD7FBB0FE847364" box="[151,268,1102,1128]" pageId="16" pageNumber="285" value="2016-05-22">22.V.2016</collectingDate>
</date>
</materialsCitation>
, Yudan Tang &amp; Ruijuan Zhang. 
<emphasis id="B92BEAFA435EBC710E3DFBB3FCEB736B" bold="true" box="[637,867,1101,1127]" pageId="16" pageNumber="285">Locality uncertain:</emphasis>
<materialsCitation id="3B373CB5435EBC710F2BFBB0FC9573BC" ID-GBIF-Occurrence="3436469318" collectionCode="BMNH" country="China" location="Precise" municipality="Note" pageId="16" pageNumber="285" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan">
<specimenCount id="9D59FD61435EBC710F2BFBB0FC467364" box="[875,974,1102,1128]" pageId="16" pageNumber="285" type="female">1 female</specimenCount>
(
<collectionCode id="ED4EAE2D435EBC710F9EFBB0FBB07364" box="[990,1080,1102,1128]" pageId="16" pageNumber="285">BMNH</collectionCode>
), “China. // Sharp Coll. / 1905- 313.” [
<collectingMunicipality id="6B84AC92435EBC710CA5FB8CFEAA7380" box="[229,290,1138,1164]" pageId="16" pageNumber="285">Note</collectingMunicipality>
: 
<location id="8E806033435EBC710D6FFB8CFE087380" LSID="urn:lsid:plazi:treatment:03F687FE435EBC750CD7FF69FAA174E1:8E806033435EBC710D6FFB8CFE087380" box="[303,384,1138,1164]" country="China" municipality="Note" name="Precise" pageId="16" pageNumber="285" stateProvince="Yunnan">Precise</location>
identification of the species without examination of aedeagus is difficult, but the specimen agrees with the 
<typeStatus id="54E4884A435EBC710D08FB68FEF073BC" box="[328,376,1174,1200]" pageId="16" pageNumber="285">type</typeStatus>
specimens in all external characters.]
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3456563435EBC710C87FB47FD3572F8" pageId="16" pageNumber="285" type="diagnosis">
<paragraph id="8BE036E8435EBC710C87FB47FD3572F8" blockId="16.[151,1437,417,1957]" pageId="16" pageNumber="285">
<emphasis id="B92BEAFA435EBC710C87FB47FECA73DF" bold="true" box="[199,322,1209,1235]" pageId="16" pageNumber="285">Diagnosis.</emphasis>
Length 6.9–8.0 mm. Body oval, anterior half of elytra nearly parallel-sided (
<figureCitation id="13642A6D435EBC7108FFFB44FA8E73D8" box="[1215,1286,1210,1236]" captionStart="FIGURES 5–10" captionStartId="8.[151,264,1933,1957]" captionTargetBox="[260,1303,188,1900]" captionTargetId="figure-30@8.[214,1373,181,1908]" captionTargetPageId="8" captionText="FIGURES 5–10. Habitus of Dactylosternum spp. from China and Japan. 5: D. songxiaobini sp. nov., 6: D. hydrophiloides (MacLeay, 1825), 7: D. pseudolatum sp. nov., 8: D. salvazai Orchymont, 1925, 9: D. latum (Sharp, 1873), 10: D. frater sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5843841" httpUri="https://zenodo.org/record/5843841/files/figure.png" pageId="16" pageNumber="285">Fig. 9</figureCitation>
), moderately convex. Dorsal black, surface with fine punctures, comparatively shiny.Anterior margin of clypeus without marginal rim. Antenna yellow-brown, club compact. Elytra with sutural stria and ten distinct rows of serial punctures, all nearly reaching elytral base. Anteromedial portion of metaventral process with a strongly depressed pit (
<figureCitation id="13642A6D435EBC71097AFAD8FA18724D" box="[1338,1424,1318,1345]" captionStart="FIGURES 48–63" captionStartId="21.[151,264,1495,1519]" captionTargetBox="[153,1430,183,1463]" captionTargetId="figure-19@21.[151,1436,181,1471]" captionTargetPageId="21" captionText="FIGURES 48–63. Morphology characters of Dactylosternum spp. SEM micrographs. 48–50: mediobasal portion of elytron (48: D. songxiaobini sp. nov. (red arrows show primary serial punctures, green arrows show secondary serial punctures), 49: D. nanlingensis sp. nov., 50: D. corbetti), 51–55: punctures on posterior half of elytron (51: D. hydrophiloides (the red arrow indicates the ground puncture, the blue arrow indicates the micropuncture), 52: D. pseudolatum sp. nov., 53: D. salvazai, 54: D. latum, 55: D. frater sp. nov.), 56–59: anteromedial portion of metaventrite (56: D. pseudolatum sp. nov., 57: D. salvazai, 58: D. latum, 59: D. frater sp. nov.), 60–63: posteromedial portion of ventrite 5 (60: D. pseudolatum sp. nov., 61: D. salvazai, 62: D. latum, 63: D. frater sp. nov.)" figureDoi="http://doi.org/10.5281/zenodo.5843883" httpUri="https://zenodo.org/record/5843883/files/figure.png" pageId="16" pageNumber="285">Fig. 58</figureCitation>
). Posterior margin of abdominal ventrite 5 with fine marginal bead, without stria behind the marginal bead (
<figureCitation id="13642A6D435EBC71097CFAB4FA187269" box="[1340,1424,1354,1381]" captionStart="FIGURES 48–63" captionStartId="21.[151,264,1495,1519]" captionTargetBox="[153,1430,183,1463]" captionTargetId="figure-19@21.[151,1436,181,1471]" captionTargetPageId="21" captionText="FIGURES 48–63. Morphology characters of Dactylosternum spp. SEM micrographs. 48–50: mediobasal portion of elytron (48: D. songxiaobini sp. nov. (red arrows show primary serial punctures, green arrows show secondary serial punctures), 49: D. nanlingensis sp. nov., 50: D. corbetti), 51–55: punctures on posterior half of elytron (51: D. hydrophiloides (the red arrow indicates the ground puncture, the blue arrow indicates the micropuncture), 52: D. pseudolatum sp. nov., 53: D. salvazai, 54: D. latum, 55: D. frater sp. nov.), 56–59: anteromedial portion of metaventrite (56: D. pseudolatum sp. nov., 57: D. salvazai, 58: D. latum, 59: D. frater sp. nov.), 60–63: posteromedial portion of ventrite 5 (60: D. pseudolatum sp. nov., 61: D. salvazai, 62: D. latum, 63: D. frater sp. nov.)" figureDoi="http://doi.org/10.5281/zenodo.5843883" httpUri="https://zenodo.org/record/5843883/files/figure.png" pageId="16" pageNumber="285">Fig. 62</figureCitation>
). 
<emphasis id="B92BEAFA435EBC710CD7FA93FE8F7284" bold="true" box="[151,263,1389,1416]" italics="true" pageId="16" pageNumber="285">Aedeagus</emphasis>
(
<figureCitation id="13642A6D435EBC710D58FA90FE2B7284" box="[280,419,1390,1416]" captionStart-0="FIGURES 30–32" captionStart-1="FIGURES 33–38" captionStartId-0="17.[151,264,1701,1725]" captionStartId-1="18.[151,264,1947,1971]" captionTargetBox-0="[223,1377,191,1614]" captionTargetBox-1="[225,1404,347,1884]" captionTargetId-0="figure-19@17.[151,1436,181,1677]" captionTargetId-1="figure-85@18.[151,1436,309,1924]" captionTargetPageId-0="17" captionTargetPageId-1="18" captionText-0="FIGURES 30–32. Aedeagus of D. latum. 30: from Yunnan, 31: from Hunan, 32: syntype (30a, 31a, 32a: dorsal view, 30b, 31b: dorsal view, 30c, 31c: ventral plate of the median lobe, 32b: labels of syntype)" captionText-1="FIGURES 33–38. Aedeagus of D. latum (dorsal view). 33: from Hunan, Shimen County, 34: syntype, 35: from Guangxi, 36: holotype of D. vitalisi, 37: from Hunan, Shimen County, 38: from Yunnan" figureDoi-0="http://doi.org/10.5281/zenodo.5843869" figureDoi-1="http://doi.org/10.5281/zenodo.5843873" httpUri-0="https://zenodo.org/record/5843869/files/figure.png" httpUri-1="https://zenodo.org/record/5843873/files/figure.png" pageId="16" pageNumber="285">Figs. 30–38</figureCitation>
). Morphologically variable, phallobase shorter than parameres. Parameres slightly sinuate on outer margin. Median lobe widest in the basal one-fourths or at midlength, a little shorter than parameres; ventral plate of the median lobe almost parallel-sided in apical half, apex rounded, gonopore situated at apex of ventral plate (
<figureCitation id="13642A6D435EBC710CDFFA24FEC872F8" box="[159,320,1498,1524]" captionStart="FIGURES 30–32" captionStartId="17.[151,264,1701,1725]" captionTargetBox="[223,1377,191,1614]" captionTargetId="figure-19@17.[151,1436,181,1677]" captionTargetPageId="17" captionText="FIGURES 30–32. Aedeagus of D. latum. 30: from Yunnan, 31: from Hunan, 32: syntype (30a, 31a, 32a: dorsal view, 30b, 31b: dorsal view, 30c, 31c: ventral plate of the median lobe, 32b: labels of syntype)" figureDoi="http://doi.org/10.5281/zenodo.5843869" httpUri="https://zenodo.org/record/5843869/files/figure.png" pageId="16" pageNumber="285">Figs. 30c, 31c</figureCitation>
), reaching ca. 0.8 of median lobe.
</paragraph>
</subSubSection>
<subSubSection id="C3456563435EBC750C87FA03FEA07611" lastPageId="20" lastPageNumber="289" pageId="16" pageNumber="285" type="description">
<paragraph id="8BE036E8435EBC710C87FA03FAC57114" blockId="16.[151,1437,417,1957]" box="[199,1357,1533,1561]" pageId="16" pageNumber="285">
<emphasis id="B92BEAFA435EBC710C87FA03FEFD711B" bold="true" box="[199,373,1533,1559]" pageId="16" pageNumber="285">Redescription.</emphasis>
Total length 6.9–8.0 mm; maximum width 
<quantity id="4CA79B0D435EBC710F1DFA00FC6F7114" box="[861,999,1534,1561]" metricMagnitude="-3" metricUnit="m" metricValue="4.65" metricValueMax="4.8" metricValueMin="4.5" pageId="16" pageNumber="285" unit="mm" value="4.65" valueMax="4.8" valueMin="4.5">4.5–4.8 mm</quantity>
; body oval, moderately convex.
</paragraph>
<paragraph id="8BE036E8435EBC710C87F9DFFC2C716C" blockId="16.[151,1437,417,1957]" pageId="16" pageNumber="285">
<emphasis id="B92BEAFA435EBC710C87F9DFFEC37130" box="[199,331,1569,1596]" italics="true" pageId="16" pageNumber="285">
<emphasis id="B92BEAFA435EBC710C87F9DFFECB7130" bold="true" box="[199,323,1569,1596]" italics="true" pageId="16" pageNumber="285">Coloration</emphasis>
.
</emphasis>
Dorsal black, with lateral margin of elytron reddish brown (
<figureCitation id="13642A6D435EBC710FBDF9DCFBCB7130" box="[1021,1091,1570,1596]" captionStart="FIGURES 5–10" captionStartId="8.[151,264,1933,1957]" captionTargetBox="[260,1303,188,1900]" captionTargetId="figure-30@8.[214,1373,181,1908]" captionTargetPageId="8" captionText="FIGURES 5–10. Habitus of Dactylosternum spp. from China and Japan. 5: D. songxiaobini sp. nov., 6: D. hydrophiloides (MacLeay, 1825), 7: D. pseudolatum sp. nov., 8: D. salvazai Orchymont, 1925, 9: D. latum (Sharp, 1873), 10: D. frater sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5843841" httpUri="https://zenodo.org/record/5843841/files/figure.png" pageId="16" pageNumber="285">Fig. 9</figureCitation>
). Maxillary palps and antenna yellowish brown. Ventral black to reddish brown, legs reddish brown.
</paragraph>
<paragraph id="8BE036E8435EBC710C87F997FAB370A8" blockId="16.[151,1437,417,1957]" pageId="16" pageNumber="285">
<emphasis id="B92BEAFA435EBC710C87F997FE847188" bold="true" box="[199,268,1641,1668]" italics="true" pageId="16" pageNumber="285">Head.</emphasis>
Clypeus weakly convex, surface with dense and fine punctures, anterior portion with denser punctures. Interstices between punctures smooth. Anterior margin of clypeus without marginal rim. Frons with punctures sparser than those on clypeus. Eyes of moderate size, separated by ca. 5× as wide as one eye, distinctly emarginate anteriorly in lateral view, smooth medially. Maxilla with glabrous basistipes. Mentum transverse, glabrous, strongly impressed anteromedially, lateral sides with series of fine setae. Submentum reddish brown, pubescent. Labial palps rather stout, palpomere 1 minute, palpomere 2 strongly widened distally, with dense brush of long yellow setae on its inner face and distal margin, palpomere 3 longest. Antennal scapus longest, rather thick, ca. 7.5× as long as pedicellus, antennomere 3 thin and a little shorter than pedicellus, antennomeres 4 and 5 almost equal in length, slightly widening distally, club densely pubescent, with few long setae, compact, ca. 2.5× as long as wide.
</paragraph>
<caption id="DF206660435FBC700CD7F95BFC3971ED" ID-DOI="http://doi.org/10.5281/zenodo.5843869" ID-Zenodo-Dep="5843869" httpUri="https://zenodo.org/record/5843869/files/figure.png" pageId="17" pageNumber="286" startId="17.[151,264,1701,1725]" targetBox="[223,1377,191,1614]" targetPageId="17">
<paragraph id="8BE036E8435FBC700CD7F95BFC3971ED" blockId="17.[151,1437,1701,1761]" pageId="17" pageNumber="286">
<emphasis id="B92BEAFA435FBC700CD7F95BFEDC71B1" bold="true" box="[151,340,1701,1725]" pageId="17" pageNumber="286">FIGURES 30–32.</emphasis>
Aedeagus of 
<taxonomicName id="4C5F4D6B435FBC700D9DF958FDBE71B1" authorityName="Sharp" authorityYear="1873" box="[477,566,1701,1725]" class="Insecta" family="Hydrophilidae" genus="Cyclonotum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="286" phylum="Arthropoda" rank="species" species="latum">
<emphasis id="B92BEAFA435FBC700D9DF958FDBE71B1" box="[477,566,1701,1725]" italics="true" pageId="17" pageNumber="286">D. latum</emphasis>
</taxonomicName>
. 30: from Yunnan, 31: from Hunan, 32: syntype (30a, 31a, 32a: dorsal view, 30b, 31b: dorsal view, 30c, 31c: ventral plate of the median lobe, 32b: labels of syntype)
</paragraph>
</caption>
<paragraph id="8BE036E8435FBC700C87F8F1FCFC707D" blockId="17.[151,1436,1807,2014]" pageId="17" pageNumber="286">
<emphasis id="B92BEAFA435FBC700C87F8F1FECC7026" bold="true" box="[199,324,1807,1834]" italics="true" pageId="17" pageNumber="286">Prothorax.</emphasis>
Pronotum moderately convex, widest basally, arcuately narrowed in front, with punctures sparser than those on vertex, interstices between punctures smooth; posterior margin slightly arcuate. Prosternum strongly tectiform, with median carina and a distinct tooth anteromedially.
</paragraph>
<paragraph id="8BE036E8435FBC730C87F882FE757611" blockId="17.[151,1436,1807,2014]" lastBlockId="18.[151,1437,150,285]" lastPageId="18" lastPageNumber="287" pageId="17" pageNumber="286">
<emphasis id="B92BEAFA435FBC700C87F882FE58709A" bold="true" box="[199,464,1915,1942]" italics="true" pageId="17" pageNumber="286">Meso- and metathorax.</emphasis>
Scutellum distinctly longer than wide, apex rounded, in shape of isosceles triangle, with punctation similar to those on pronotum. Elytron moderately convex, sutural stria deeply impressed in posterior half; each elytron with 10 longitudinal serial punctures, deeply impressed in posterior half, intervals with fine ground punctures, each puncture attached with two short transverse microsculpture especially in posterior half (
<figureCitation id="13642A6D435CBC73097AFF69FA0777BD" box="[1338,1423,150,177]" captionStart="FIGURES 48–63" captionStartId="21.[151,264,1495,1519]" captionTargetBox="[153,1430,183,1463]" captionTargetId="figure-19@21.[151,1436,181,1471]" captionTargetPageId="21" captionText="FIGURES 48–63. Morphology characters of Dactylosternum spp. SEM micrographs. 48–50: mediobasal portion of elytron (48: D. songxiaobini sp. nov. (red arrows show primary serial punctures, green arrows show secondary serial punctures), 49: D. nanlingensis sp. nov., 50: D. corbetti), 51–55: punctures on posterior half of elytron (51: D. hydrophiloides (the red arrow indicates the ground puncture, the blue arrow indicates the micropuncture), 52: D. pseudolatum sp. nov., 53: D. salvazai, 54: D. latum, 55: D. frater sp. nov.), 56–59: anteromedial portion of metaventrite (56: D. pseudolatum sp. nov., 57: D. salvazai, 58: D. latum, 59: D. frater sp. nov.), 60–63: posteromedial portion of ventrite 5 (60: D. pseudolatum sp. nov., 61: D. salvazai, 62: D. latum, 63: D. frater sp. nov.)" figureDoi="http://doi.org/10.5281/zenodo.5843883" httpUri="https://zenodo.org/record/5843883/files/figure.png" pageId="18" pageNumber="287">Fig. 54</figureCitation>
), series 3+4 and 6+7 linked together subposteriorly, others serial punctures reaching end of elytron, the outer 4 serial punctures larger and deeper than the inner 6 serial punctures. Anteromedial portion of metaventral process with a strongly depressed pit (
<figureCitation id="13642A6D435CBC730DDCFEFDFE797611" box="[412,497,258,285]" captionStart="FIGURES 48–63" captionStartId="21.[151,264,1495,1519]" captionTargetBox="[153,1430,183,1463]" captionTargetId="figure-19@21.[151,1436,181,1471]" captionTargetPageId="21" captionText="FIGURES 48–63. Morphology characters of Dactylosternum spp. SEM micrographs. 48–50: mediobasal portion of elytron (48: D. songxiaobini sp. nov. (red arrows show primary serial punctures, green arrows show secondary serial punctures), 49: D. nanlingensis sp. nov., 50: D. corbetti), 51–55: punctures on posterior half of elytron (51: D. hydrophiloides (the red arrow indicates the ground puncture, the blue arrow indicates the micropuncture), 52: D. pseudolatum sp. nov., 53: D. salvazai, 54: D. latum, 55: D. frater sp. nov.), 56–59: anteromedial portion of metaventrite (56: D. pseudolatum sp. nov., 57: D. salvazai, 58: D. latum, 59: D. frater sp. nov.), 60–63: posteromedial portion of ventrite 5 (60: D. pseudolatum sp. nov., 61: D. salvazai, 62: D. latum, 63: D. frater sp. nov.)" figureDoi="http://doi.org/10.5281/zenodo.5843883" httpUri="https://zenodo.org/record/5843883/files/figure.png" pageId="18" pageNumber="287">Fig. 58</figureCitation>
).
</paragraph>
<caption id="DF206660435CBC730CD7F865FC0C70DB" ID-DOI="http://doi.org/10.5281/zenodo.5843873" ID-Zenodo-Dep="5843873" httpUri="https://zenodo.org/record/5843873/files/figure.png" pageId="18" pageNumber="287" startId="18.[151,264,1947,1971]" targetBox="[225,1404,347,1884]" targetPageId="18">
<paragraph id="8BE036E8435CBC730CD7F865FC0C70DB" blockId="18.[151,1437,1947,2007]" pageId="18" pageNumber="287">
<emphasis id="B92BEAFA435CBC730CD7F865FEDD70BF" bold="true" box="[151,341,1947,1971]" pageId="18" pageNumber="287">FIGURES 33–38.</emphasis>
Aedeagus of 
<taxonomicName id="4C5F4D6B435CBC730DA2F862FDB470BF" authorityName="Sharp" authorityYear="1873" box="[482,572,1947,1971]" class="Insecta" family="Hydrophilidae" genus="Cyclonotum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="287" phylum="Arthropoda" rank="species" species="latum">
<emphasis id="B92BEAFA435CBC730DA2F862FDB470BF" box="[482,572,1947,1971]" italics="true" pageId="18" pageNumber="287">D. latum</emphasis>
</taxonomicName>
(dorsal view). 33: from Hunan, Shimen County, 34: syntype, 35: from Guangxi, 36: holotype of 
<taxonomicName id="4C5F4D6B435CBC730D52F83EFEFE70DB" authorityName="d' Orchymont" authorityYear="1925" box="[274,374,1983,2007]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="287" phylum="Arthropoda" rank="species" species="vitalisi">
<emphasis id="B92BEAFA435CBC730D52F83EFEFE70DB" box="[274,374,1983,2007]" italics="true" pageId="18" pageNumber="287">D. vitalisi</emphasis>
</taxonomicName>
, 37: from Hunan, Shimen County, 38: from Yunnan
</paragraph>
</caption>
<caption id="DF206660435DBC720CD7FA44FA1F716E" ID-DOI="http://doi.org/10.5281/zenodo.5843879" ID-Zenodo-Dep="5843879" httpUri="https://zenodo.org/record/5843879/files/figure.png" pageId="19" pageNumber="288" startId="19.[151,264,1466,1490]" targetBox="[151,1431,183,1437]" targetPageId="19">
<paragraph id="8BE036E8435DBC720CD7FA44FA1F716E" blockId="19.[151,1437,1466,1634]" pageId="19" pageNumber="288">
<emphasis id="B92BEAFA435DBC720CD7FA44FEDB72DE" bold="true" box="[151,339,1466,1490]" pageId="19" pageNumber="288">FIGURES 39–47.</emphasis>
Morphology characters of 
<taxonomicName id="4C5F4D6B435DBC720E21FA44FCBA72DD" box="[609,818,1466,1490]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="288" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B92BEAFA435DBC720E21FA44FC8B72DE" box="[609,771,1466,1490]" italics="true" pageId="19" pageNumber="288">Dactylosternum</emphasis>
spp.
</taxonomicName>
SEM micrographs. 39–40: antennal club (39: 
<taxonomicName id="4C5F4D6B435DBC7208BEFA45FA1472DE" authority="Mai &amp; Hu &amp; Minoshima &amp; Jia &amp; Fikáček, 2022" authorityName="Mai &amp; Hu &amp; Minoshima &amp; Jia &amp; Fikáček" authorityYear="2022" box="[1278,1436,1466,1490]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="288" phylum="Arthropoda" rank="species" species="nanlingensis" status="sp. nov.">
<emphasis id="B92BEAFA435DBC7208BEFA45FA1472DE" box="[1278,1436,1466,1490]" italics="true" pageId="19" pageNumber="288">D. nanlingensis</emphasis>
</taxonomicName>
<emphasis id="B92BEAFA435DBC720CD7FA20FF6F72FA" bold="true" box="[151,231,1502,1526]" pageId="19" pageNumber="288">
<taxonomicNameLabel id="A2185781435DBC720CD7FA20FF6F72FA" box="[151,231,1502,1526]" pageId="19" pageNumber="288" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, 40: 
<taxonomicName id="4C5F4D6B435DBC720D59FA21FE2772FA" baseAuthorityName="Fabricius" baseAuthorityYear="1792" box="[281,431,1502,1526]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="288" phylum="Arthropoda" rank="species" species="abdominale">
<emphasis id="B92BEAFA435DBC720D59FA21FE2772FA" box="[281,431,1502,1526]" italics="true" pageId="19" pageNumber="288">D. abdominale</emphasis>
</taxonomicName>
), 41–42: ventral surface of mesofemora (41: 
<taxonomicName id="4C5F4D6B435DBC720F35FA21FC3B72FA" authorityName="Jia. This" authorityYear="2002" box="[885,947,1503,1526]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="288" phylum="Arthropoda" rank="species" species="pui">
<emphasis id="B92BEAFA435DBC720F35FA21FC3B72FA" box="[885,947,1503,1526]" italics="true" pageId="19" pageNumber="288">D. pui</emphasis>
</taxonomicName>
, 42: 
<taxonomicName id="4C5F4D6B435DBC720FA6FA21FB0C72FA" authority="Mai &amp; Hu &amp; Minoshima &amp; Jia &amp; Fikáček, 2022" authorityName="Mai &amp; Hu &amp; Minoshima &amp; Jia &amp; Fikáček" authorityYear="2022" box="[998,1156,1502,1526]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="288" phylum="Arthropoda" rank="species" species="nanlingensis" status="sp. nov.">
<emphasis id="B92BEAFA435DBC720FA6FA21FB0C72FA" box="[998,1156,1502,1526]" italics="true" pageId="19" pageNumber="288">D. nanlingensis</emphasis>
</taxonomicName>
<emphasis id="B92BEAFA435DBC7208C9FA20FB5172FA" bold="true" box="[1161,1241,1502,1526]" pageId="19" pageNumber="288">
<taxonomicNameLabel id="A2185781435DBC7208C9FA20FB5172FA" box="[1161,1241,1502,1526]" pageId="19" pageNumber="288" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
), 43–47: punctures on pronotum (43: 
<taxonomicName id="4C5F4D6B435DBC720D0EF9FDFE057116" authorityName="Jia. This" authorityYear="2002" box="[334,397,1539,1562]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="288" phylum="Arthropoda" rank="species" species="pui">
<emphasis id="B92BEAFA435DBC720D0EF9FDFE057116" box="[334,397,1539,1562]" italics="true" pageId="19" pageNumber="288">D. pui</emphasis>
</taxonomicName>
, 44: 
<taxonomicName id="4C5F4D6B435DBC720D80F9FDFDEA7116" authority="Mai &amp; Hu &amp; Minoshima &amp; Jia &amp; Fikáček, 2022" authorityName="Mai &amp; Hu &amp; Minoshima &amp; Jia &amp; Fikáček" authorityYear="2022" box="[448,610,1538,1562]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="288" phylum="Arthropoda" rank="species" species="songxiaobini" status="sp. nov.">
<emphasis id="B92BEAFA435DBC720D80F9FDFDEA7116" box="[448,610,1538,1562]" italics="true" pageId="19" pageNumber="288">D. songxiaobini</emphasis>
</taxonomicName>
<emphasis id="B92BEAFA435DBC720E27F9FCFD307116" bold="true" box="[615,696,1538,1562]" pageId="19" pageNumber="288">
<taxonomicNameLabel id="A2185781435DBC720E27F9FCFD307116" box="[615,696,1538,1562]" pageId="19" pageNumber="288" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(surface with two sizes of punctures, the blue arrow indicates the coarser puncture, the yellow arrow indicates the finer puncture), 45: 
<taxonomicName id="4C5F4D6B435DBC720EBAF9D9FC117132" authority="Mai &amp; Hu &amp; Minoshima &amp; Jia &amp; Fikáček, 2022" authorityName="Mai &amp; Hu &amp; Minoshima &amp; Jia &amp; Fikáček" authorityYear="2022" box="[762,921,1574,1598]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="288" phylum="Arthropoda" rank="species" species="nanlingensis" status="sp. nov.">
<emphasis id="B92BEAFA435DBC720EBAF9D9FC117132" box="[762,921,1574,1598]" italics="true" pageId="19" pageNumber="288">D. nanlingensis</emphasis>
</taxonomicName>
<emphasis id="B92BEAFA435DBC720FDFF9D8FC677132" bold="true" box="[927,1007,1574,1598]" pageId="19" pageNumber="288">
<taxonomicNameLabel id="A2185781435DBC720FDFF9D8FC677132" box="[927,1007,1574,1598]" pageId="19" pageNumber="288" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, 46: 
<taxonomicName id="4C5F4D6B435DBC720861F9D9FB077132" authorityName="Balfour-Browne" authorityYear="1942" box="[1057,1167,1574,1598]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="288" phylum="Arthropoda" rank="species" species="corbetti">
<emphasis id="B92BEAFA435DBC720861F9D9FB077132" box="[1057,1167,1574,1598]" italics="true" pageId="19" pageNumber="288">D. corbetti</emphasis>
</taxonomicName>
(arrow indicates two short divergent lines connected with a puncture), 47: 
<taxonomicName id="4C5F4D6B435DBC720E39F9B5FC98716E" baseAuthorityName="Fabricius" baseAuthorityYear="1792" box="[633,784,1610,1634]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="288" phylum="Arthropoda" rank="species" species="abdominale">
<emphasis id="B92BEAFA435DBC720E39F9B5FC98716E" box="[633,784,1610,1634]" italics="true" pageId="19" pageNumber="288">D. abdominale</emphasis>
</taxonomicName>
(arrow indicates mesh-like microsculpture between punctures))
</paragraph>
</caption>
<paragraph id="8BE036E8435DBC720C87F96FFF6D701A" blockId="19.[151,1437,1680,2031]" pageId="19" pageNumber="288">
<emphasis id="B92BEAFA435DBC720C87F96FFE8D71A7" bold="true" box="[199,261,1681,1707]" italics="true" pageId="19" pageNumber="288">Legs.</emphasis>
Reddish brown, rather short. Procoxae globular, pubescent; profemora without pubescence ventrally; protarsomeres unpubescent ventrally. Ventral surface of meso- and metafemora without pubescence, with sparse punctures and microsculptures; meso- and metatarsomeres with sparse pubescence ventrally, almost equal in length.
</paragraph>
<paragraph id="8BE036E8435DBC720C87F8DEFE5F708F" blockId="19.[151,1437,1680,2031]" pageId="19" pageNumber="288">
<emphasis id="B92BEAFA435DBC720C87F8DEFEB47037" bold="true" box="[199,316,1824,1851]" italics="true" pageId="19" pageNumber="288">Abdomen.</emphasis>
Pubescent, ventrite 1 with complete median carina, other ventrites uniformly convex. Posteromedial portion of ventrite 5 unpubescent, with fine punctures, posterior margin of ventrite 5 rounded, with fine marginal bead, without stria (
<figureCitation id="13642A6D435DBC720D36F896FE43708F" box="[374,459,1896,1923]" captionStart="FIGURES 48–63" captionStartId="21.[151,264,1495,1519]" captionTargetBox="[153,1430,183,1463]" captionTargetId="figure-19@21.[151,1436,181,1471]" captionTargetPageId="21" captionText="FIGURES 48–63. Morphology characters of Dactylosternum spp. SEM micrographs. 48–50: mediobasal portion of elytron (48: D. songxiaobini sp. nov. (red arrows show primary serial punctures, green arrows show secondary serial punctures), 49: D. nanlingensis sp. nov., 50: D. corbetti), 51–55: punctures on posterior half of elytron (51: D. hydrophiloides (the red arrow indicates the ground puncture, the blue arrow indicates the micropuncture), 52: D. pseudolatum sp. nov., 53: D. salvazai, 54: D. latum, 55: D. frater sp. nov.), 56–59: anteromedial portion of metaventrite (56: D. pseudolatum sp. nov., 57: D. salvazai, 58: D. latum, 59: D. frater sp. nov.), 60–63: posteromedial portion of ventrite 5 (60: D. pseudolatum sp. nov., 61: D. salvazai, 62: D. latum, 63: D. frater sp. nov.)" figureDoi="http://doi.org/10.5281/zenodo.5843883" httpUri="https://zenodo.org/record/5843883/files/figure.png" pageId="19" pageNumber="288">Fig. 62</figureCitation>
).
</paragraph>
<paragraph id="8BE036E8435DBC750C87F872FEA07611" blockId="19.[151,1437,1680,2031]" lastBlockId="20.[151,1437,150,1006]" lastPageId="20" lastPageNumber="289" pageId="19" pageNumber="288">
<emphasis id="B92BEAFA435DBC720C87F872FEE670AB" bold="true" box="[199,366,1932,1959]" italics="true" pageId="19" pageNumber="288">Male genitalia</emphasis>
(
<figureCitation id="13642A6D435DBC720D3DF872FD8F70AA" box="[381,519,1932,1958]" captionStart-0="FIGURES 30–32" captionStart-1="FIGURES 33–38" captionStartId-0="17.[151,264,1701,1725]" captionStartId-1="18.[151,264,1947,1971]" captionTargetBox-0="[223,1377,191,1614]" captionTargetBox-1="[225,1404,347,1884]" captionTargetId-0="figure-19@17.[151,1436,181,1677]" captionTargetId-1="figure-85@18.[151,1436,309,1924]" captionTargetPageId-0="17" captionTargetPageId-1="18" captionText-0="FIGURES 30–32. Aedeagus of D. latum. 30: from Yunnan, 31: from Hunan, 32: syntype (30a, 31a, 32a: dorsal view, 30b, 31b: dorsal view, 30c, 31c: ventral plate of the median lobe, 32b: labels of syntype)" captionText-1="FIGURES 33–38. Aedeagus of D. latum (dorsal view). 33: from Hunan, Shimen County, 34: syntype, 35: from Guangxi, 36: holotype of D. vitalisi, 37: from Hunan, Shimen County, 38: from Yunnan" figureDoi-0="http://doi.org/10.5281/zenodo.5843869" figureDoi-1="http://doi.org/10.5281/zenodo.5843873" httpUri-0="https://zenodo.org/record/5843869/files/figure.png" httpUri-1="https://zenodo.org/record/5843873/files/figure.png" pageId="19" pageNumber="288">Figs. 30–38</figureCitation>
). Aedeagus around 
<quantity id="4CA79B0D435DBC720EA2F872FCB070AB" box="[738,824,1932,1959]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="19" pageNumber="288" unit="mm" value="1.2">1.2 mm</quantity>
long, ca. 2.7× as long as wide. Phallobase shorter than parameres, with short manubrium and rounded posteriorly. Parameres widest basally, gradually narrowing toward apex, outer margin straight, slightly sinuate, slightly or strongly curved inwards apically. Median lobe ca. 0.8–0.9× as long as parameres, widest in the basal 0.25 or at midlength, outer margin nearly parallel-side or slightly narrowing towards apex, rounded anteriorly, dully pointed apically; ventral plate of median lobe widest basally, almost parallelsided in apical half, apex rounded, gonopore situated at apex of ventral plate (
<figureCitation id="13642A6D435ABC750853FF21FB3977F5" box="[1043,1201,223,249]" captionStart="FIGURES 30–32" captionStartId="17.[151,264,1701,1725]" captionTargetBox="[223,1377,191,1614]" captionTargetId="figure-19@17.[151,1436,181,1677]" captionTargetPageId="17" captionText="FIGURES 30–32. Aedeagus of D. latum. 30: from Yunnan, 31: from Hunan, 32: syntype (30a, 31a, 32a: dorsal view, 30b, 31b: dorsal view, 30c, 31c: ventral plate of the median lobe, 32b: labels of syntype)" figureDoi="http://doi.org/10.5281/zenodo.5843869" httpUri="https://zenodo.org/record/5843869/files/figure.png" pageId="20" pageNumber="289">Figs 30c, 31c</figureCitation>
), reaching ca. 0.8 of median lobe.
</paragraph>
</subSubSection>
<subSubSection id="C3456563435ABC750C87FED8FB517531" pageId="20" pageNumber="289" type="biology_ecology">
<paragraph id="8BE036E8435ABC750C87FED8FB517531" blockId="20.[151,1437,150,1006]" pageId="20" pageNumber="289">
<emphasis id="B92BEAFA435ABC750C87FED8FEAF764C" bold="true" box="[199,295,294,320]" pageId="20" pageNumber="289">Biology.</emphasis>
Unknown. The species was considered as aquatic in the original description (
<bibRefCitation id="EFCE4B19435ABC7508DCFED9FAAA764D" author="Sharp, D." box="[1180,1314,295,321]" pageId="20" pageNumber="289" pagination="64 - 65" refId="ref23391" refString="Sharp, D. (1873) The water beetles of Japan. Transactions of the Entomological Society of London, 1874, 64 - 65." type="journal article" year="1873">Sharp 1873</bibRefCitation>
), based on the information from the collector (G. Lewis) that it is ‘common in ponds’. 
<bibRefCitation id="EFCE4B19435ABC750FB7FEB5FBF4766A" author="Sato, M." box="[1015,1148,331,358]" pageId="20" pageNumber="289" pagination="21 - 22" refId="ref23240" refString="Sato, M. (1960) Notes on the tribe Sphaeridiini of Japan (Coleoptera: Hydrophilidae). Ageha, 8, 21 - 22. [in Japanese]" type="journal article" year="1960">Satô (1960)</bibRefCitation>
questioned the habitat of 
<taxonomicName id="4C5F4D6B435ABC750CD7FE91FF727685" authorityName="Sharp" authorityYear="1873" box="[151,250,367,393]" class="Insecta" family="Hydrophilidae" genus="Cyclonotum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="289" phylum="Arthropoda" rank="species" species="latum">
<emphasis id="B92BEAFA435ABC750CD7FE91FF727685" box="[151,250,367,393]" italics="true" pageId="20" pageNumber="289">D. latum</emphasis>
</taxonomicName>
and suspected the species to be a synonym of 
<taxonomicName id="4C5F4D6B435ABC750F4AFE91FC5C7685" baseAuthorityName="MacLeay" baseAuthorityYear="1825" box="[778,980,367,393]" class="Insecta" family="Hydrophilidae" genus="Sphaeridium" kingdom="Fungi" order="Coleoptera" pageId="20" pageNumber="289" phylum="Ascomycota" rank="species" species="hydrophiloides">
<emphasis id="B92BEAFA435ABC750F4AFE91FC5C7685" box="[778,980,367,393]" italics="true" pageId="20" pageNumber="289">D. hydrophiloides</emphasis>
</taxonomicName>
. We suspect that the habitat information in the original description was likely based on the confusion with some 
<taxonomicName id="4C5F4D6B435ABC750F98FE6DFB3076A1" box="[984,1208,403,429]" class="Insecta" family="Hydrophilidae" genus="Coelostoma" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="289" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B92BEAFA435ABC750F98FE6DFBD676A1" box="[984,1118,403,429]" italics="true" pageId="20" pageNumber="289">Coelostoma</emphasis>
species
</taxonomicName>
which are common in 
<collectingRegion id="499BF80A435ABC750CF8FE49FEAB76DD" box="[184,291,439,465]" country="Japan" name="Nagasaki" pageId="20" pageNumber="289">Nagasaki</collectingRegion>
, of the similar morphology, coloration and body size as 
<taxonomicName id="4C5F4D6B435ABC750F80FE49FBAE76DD" authorityName="Sharp" authorityYear="1873" box="[960,1062,439,465]" class="Insecta" family="Hydrophilidae" genus="Cyclonotum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="289" phylum="Arthropoda" rank="species" species="latum">
<emphasis id="B92BEAFA435ABC750F80FE49FBAE76DD" box="[960,1062,439,465]" italics="true" pageId="20" pageNumber="289">D. latum</emphasis>
</taxonomicName>
, and are aquatic. All species of 
<taxonomicName id="4C5F4D6B435ABC750CD7FE25FEC276F9" authorityName="Wollaston" authorityYear="1854" box="[151,330,475,501]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="289" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEAFA435ABC750CD7FE25FEC276F9" box="[151,330,475,501]" italics="true" pageId="20" pageNumber="289">Dactylosternum</emphasis>
</taxonomicName>
known so far are terrestrial species (
<bibRefCitation id="EFCE4B19435ABC750F42FE25FC1676F9" author="Hansen, M." box="[770,926,475,501]" pageId="20" pageNumber="289" pagination="1 - 368" refId="ref22450" refString="Hansen, M. (1991) The hydrophiloid beetles. Phylogeny, classification and a revision of the genera (Coleoptera, Hydrophiloidea). Biologiske Skrifter, 40, 1 - 368." type="journal article" year="1991">Hansen 1991</bibRefCitation>
; 
<bibRefCitation id="EFCE4B19435ABC750FEDFE25FBC576F9" author="Fikacek, M." box="[941,1101,475,501]" pageId="20" pageNumber="289" refId="ref22128" refString="Fikacek, M. (2010) Hydrophilidae: Sphaeridiinae (Coleoptera). In: Jach, M. A. &amp; Balke, M. (Eds.), Water Beetles of New Caledonia. Volume 1. Zoologisch-Botanische Gesellschaft in Osterreich and Wiener Coleopterologenverein, Wien, pp. 323 - 364." type="book" year="2010">Fikáček 2010</bibRefCitation>
), including the other species treated here (
<emphasis id="B92BEAFA435ABC750D6BFE01FDC27515" box="[299,586,511,537]" italics="true" pageId="20" pageNumber="289">
<taxonomicName id="4C5F4D6B435ABC750D6BFE01FE067515" authorityName="Mai &amp; Hu &amp; Minoshima &amp; Jia &amp; Fikáček" authorityYear="2022" box="[299,398,511,537]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="289" phylum="Arthropoda" rank="species" species="frater">D. frater</taxonomicName>
, 
<taxonomicName id="4C5F4D6B435ABC750DD9FE01FDC27515" authorityName="Mai &amp; Hu &amp; Minoshima &amp; Jia &amp; Fikáček" authorityYear="2022" box="[409,586,511,537]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="289" phylum="Arthropoda" rank="species" species="pseudolatum">D. pseudolatum</taxonomicName>
</emphasis>
and 
<taxonomicName id="4C5F4D6B435ABC750EC1FE01FC827515" box="[641,778,511,537]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="289" phylum="Arthropoda" rank="species" species="salvarzai">
<emphasis id="B92BEAFA435ABC750EC1FE01FC827515" box="[641,778,511,537]" italics="true" pageId="20" pageNumber="289">D. salvarzai</emphasis>
</taxonomicName>
) which are very similar and likely rather closely related to 
<emphasis id="B92BEAFA435ABC750CD7FDDDFE897531" box="[151,257,547,573]" italics="true" pageId="20" pageNumber="289">
<taxonomicName id="4C5F4D6B435ABC750CD7FDDDFF747531" authorityName="Sharp" authorityYear="1873" box="[151,252,547,573]" class="Insecta" family="Hydrophilidae" genus="Cyclonotum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="289" phylum="Arthropoda" rank="species" species="latum">D. latum</taxonomicName>
.
</emphasis>
We hence assume that 
<taxonomicName id="4C5F4D6B435ABC750E45FDDDFDE07531" authorityName="Sharp" authorityYear="1873" box="[517,616,547,573]" class="Insecta" family="Hydrophilidae" genus="Cyclonotum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="289" phylum="Arthropoda" rank="species" species="latum">
<emphasis id="B92BEAFA435ABC750E45FDDDFDE07531" box="[517,616,547,573]" italics="true" pageId="20" pageNumber="289">D. latum</emphasis>
</taxonomicName>
is a terrestrial species living in decaying plant material.
</paragraph>
</subSubSection>
<subSubSection id="C3456563435ABC750C87FDB9FF6D74A9" pageId="20" pageNumber="289" type="discussion">
<paragraph id="8BE036E8435ABC750C87FDB9FC397451" blockId="20.[151,1437,150,1006]" pageId="20" pageNumber="289">
<emphasis id="B92BEAFA435ABC750C87FDB9FEB3756D" bold="true" box="[199,315,583,609]" pageId="20" pageNumber="289">Remarks.</emphasis>
This species was described based on 
<specimenCount id="9D59FD61435ABC750EB6FDB9FC17756D" box="[758,927,583,609]" pageId="20" pageNumber="289" type="generic">two specimens</specimenCount>
collected by George Lewis in 
<collectingCountry id="F3487678435ABC750948FDB9FAC3756D" box="[1288,1355,583,609]" name="Japan" pageId="20" pageNumber="289">Japan</collectingCountry>
. It had not been reported since the original description (
<bibRefCitation id="EFCE4B19435ABC750E89FD95FCDA7589" author="Sharp, D." box="[713,850,619,645]" pageId="20" pageNumber="289" pagination="64 - 65" refId="ref23391" refString="Sharp, D. (1873) The water beetles of Japan. Transactions of the Entomological Society of London, 1874, 64 - 65." type="journal article" year="1873">Sharp 1873</bibRefCitation>
). Orchymont (1925) described 
<taxonomicName id="4C5F4D6B435ABC7508FBFD95FAA47589" authorityName="d' Orchymont" authorityYear="1925" box="[1211,1324,619,645]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="289" phylum="Arthropoda" rank="species" species="vitalisi">
<emphasis id="B92BEAFA435ABC7508FBFD95FAA47589" box="[1211,1324,619,645]" italics="true" pageId="20" pageNumber="289">D. vitalisi</emphasis>
</taxonomicName>
as a new species from 
<collectingCountry id="F3487678435ABC750D70FD71FEEE75A5" box="[304,358,655,681]" name="Laos" pageId="20" pageNumber="289">Laos</collectingCountry>
(type locality) and Fujian without examining the type specimens of 
<taxonomicName id="4C5F4D6B435ABC750836FD71FB5275A5" authorityName="Sharp" authorityYear="1873" box="[1142,1242,655,681]" class="Insecta" family="Hydrophilidae" genus="Cyclonotum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="289" phylum="Arthropoda" rank="species" species="latum">
<emphasis id="B92BEAFA435ABC750836FD71FB5275A5" box="[1142,1242,655,681]" italics="true" pageId="20" pageNumber="289">D. latum</emphasis>
</taxonomicName>
. 
<taxonomicName id="4C5F4D6B435ABC7508A9FD71FF6B75C1" authorityName="d' Orchymont" authorityYear="1925" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="289" phylum="Arthropoda" rank="species" species="vitalisi">
<emphasis id="B92BEAFA435ABC7508A9FD71FF6B75C1" italics="true" pageId="20" pageNumber="289">Dactylosternum vitalisi</emphasis>
</taxonomicName>
is also a large-sized species (more than 
<quantity id="4CA79B0D435ABC750EF4FD4DFD7E75C1" box="[692,758,691,717]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="20" pageNumber="289" unit="mm" value="6.0">6 mm</quantity>
long) and is similar with 
<taxonomicName id="4C5F4D6B435ABC750860FD4DFB0C75C1" authorityName="Sharp" authorityYear="1873" box="[1056,1156,691,717]" class="Insecta" family="Hydrophilidae" genus="Cyclonotum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="289" phylum="Arthropoda" rank="species" species="latum">
<emphasis id="B92BEAFA435ABC750860FD4DFB0C75C1" box="[1056,1156,691,717]" italics="true" pageId="20" pageNumber="289">D. latum</emphasis>
</taxonomicName>
. The examination of the 
<typeStatus id="54E4884A435ABC750CD7FD29FF7175FD" box="[151,249,727,753]" pageId="20" pageNumber="289" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="4C5F4D6B435ABC750D61FD29FE1975FD" authorityName="d' Orchymont" authorityYear="1925" box="[289,401,727,753]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="289" phylum="Arthropoda" rank="species" species="vitalisi">
<emphasis id="B92BEAFA435ABC750D61FD29FE1975FD" box="[289,401,727,753]" italics="true" pageId="20" pageNumber="289">D. vitalisi</emphasis>
</taxonomicName>
from 
<collectingCountry id="F3487678435ABC750D99FD29FD8775FD" box="[473,527,727,753]" name="Laos" pageId="20" pageNumber="289">Laos</collectingCountry>
confirmed it represents the same species as the examined type specimens of 
<taxonomicName id="4C5F4D6B435ABC7509C1FD29FF5F7419" authorityName="Sharp" authorityYear="1873" class="Insecta" family="Hydrophilidae" genus="Cyclonotum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="289" phylum="Arthropoda" rank="species" species="latum">
<emphasis id="B92BEAFA435ABC7509C1FD29FF5F7419" italics="true" pageId="20" pageNumber="289">D. latum</emphasis>
</taxonomicName>
. Surprisingly, the examined 
<typeStatus id="54E4884A435ABC750E5EFD05FDF67419" box="[542,638,763,789]" pageId="20" pageNumber="289" type="paratype">paratype</typeStatus>
of 
<taxonomicName id="4C5F4D6B435ABC750EE6FD05FC9D7419" authorityName="d' Orchymont" authorityYear="1925" box="[678,789,763,789]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="289" phylum="Arthropoda" rank="species" species="vitalisi">
<emphasis id="B92BEAFA435ABC750EE6FD05FC9D7419" box="[678,789,763,789]" italics="true" pageId="20" pageNumber="289">D. vitalisi</emphasis>
</taxonomicName>
is not identical to the 
<typeStatus id="54E4884A435ABC750855FD05FBFF7419" box="[1045,1143,763,789]" pageId="20" pageNumber="289" type="holotype">holotype</typeStatus>
and is more similar to 
<taxonomicName id="4C5F4D6B435ABC7509C1FD05FEAE7435" authority="Mai &amp; Hu &amp; Minoshima &amp; Jia &amp; Fikáček, 2022" authorityName="Mai &amp; Hu &amp; Minoshima &amp; Jia &amp; Fikáček" authorityYear="2022" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="289" phylum="Arthropoda" rank="species" species="pseudolatum" status="sp. nov.">
<emphasis id="B92BEAFA435ABC7509C1FD05FEAE7435" italics="true" pageId="20" pageNumber="289">D. pseudolatum</emphasis>
</taxonomicName>
<emphasis id="B92BEAFA435ABC750D6EFCE1FE0F7435" bold="true" box="[302,391,799,825]" pageId="20" pageNumber="289">
<taxonomicNameLabel id="A2185781435ABC750D6EFCE1FE0F7435" box="[302,391,799,825]" pageId="20" pageNumber="289" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, especially in the shape of aedeagus. This indicates the type series of 
<taxonomicName id="4C5F4D6B435ABC7508D8FCE1FA8E7435" authorityName="d' Orchymont" authorityYear="1925" box="[1176,1286,799,825]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="289" phylum="Arthropoda" rank="species" species="vitalisi">
<emphasis id="B92BEAFA435ABC7508D8FCE1FA8E7435" box="[1176,1286,799,825]" italics="true" pageId="20" pageNumber="289">D. vitalisi</emphasis>
</taxonomicName>
contains two species. We hence consider 
<taxonomicName id="4C5F4D6B435ABC750D8FFCBDFDB57451" authorityName="d' Orchymont" authorityYear="1925" box="[463,573,835,861]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="289" phylum="Arthropoda" rank="species" species="vitalisi">
<emphasis id="B92BEAFA435ABC750D8FFCBDFDB57451" box="[463,573,835,861]" italics="true" pageId="20" pageNumber="289">D. vitalisi</emphasis>
</taxonomicName>
is a junior synonym of 
<emphasis id="B92BEAFA435ABC750F08FCBDFC397451" box="[840,945,835,861]" italics="true" pageId="20" pageNumber="289">
<taxonomicName id="4C5F4D6B435ABC750F08FCBDFC257451" authorityName="Sharp" authorityYear="1873" box="[840,941,835,861]" class="Insecta" family="Hydrophilidae" genus="Cyclonotum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="289" phylum="Arthropoda" rank="species" species="latum">D. latum</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="8BE036E8435ABC750C87FC98FF6D74A9" blockId="20.[151,1437,150,1006]" pageId="20" pageNumber="289">
<emphasis id="B92BEAFA435ABC750C87FC98FEC5748C" bold="true" box="[199,333,870,896]" pageId="20" pageNumber="289">Variability.</emphasis>
This species is morphologically variable, especially in the shape of aedeagus, as illustrated in 
<figureCitation id="13642A6D435ABC750925FC99FF6974A9" captionStart="FIGURES 33–38" captionStartId="18.[151,264,1947,1971]" captionTargetBox="[225,1404,347,1884]" captionTargetId="figure-85@18.[151,1436,309,1924]" captionTargetPageId="18" captionText="FIGURES 33–38. Aedeagus of D. latum (dorsal view). 33: from Hunan, Shimen County, 34: syntype, 35: from Guangxi, 36: holotype of D. vitalisi, 37: from Hunan, Shimen County, 38: from Yunnan" figureDoi="http://doi.org/10.5281/zenodo.5843873" httpUri="https://zenodo.org/record/5843873/files/figure.png" pageId="20" pageNumber="289">Figs. 33–38</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3456563435ABC750C87FC50FAA174E1" pageId="20" pageNumber="289" type="distribution">
<paragraph id="8BE036E8435ABC750C87FC50FAA174E1" blockId="20.[151,1437,150,1006]" pageId="20" pageNumber="289">
<emphasis id="B92BEAFA435ABC750C87FC50FEE974C4" bold="true" box="[199,353,942,968]" pageId="20" pageNumber="289">Distribution.</emphasis>
China (
<collectingRegion id="499BF80A435ABC750DF7FC51FD8974C5" box="[439,513,943,969]" country="China" name="Anhui" pageId="20" pageNumber="289">Anhui</collectingRegion>
, 
<collectingRegion id="499BF80A435ABC750E4BFC51FDDE74C5" box="[523,598,943,969]" country="China" name="Fujian" pageId="20" pageNumber="289">Fujian</collectingRegion>
, 
<collectingRegion id="499BF80A435ABC750E1FFC51FD2474C5" box="[607,684,943,969]" country="China" name="Hunan" pageId="20" pageNumber="289">Hunan</collectingRegion>
, 
<collectingRegion id="499BF80A435ABC750EF6FC51FC9174C5" box="[694,793,943,969]" country="China" name="Guangxi" pageId="20" pageNumber="289">Guangxi</collectingRegion>
, 
<collectingRegion id="499BF80A435ABC750F63FC51FCF674C5" box="[803,894,943,969]" country="China" name="Yunnan" pageId="20" pageNumber="289">Yunnan</collectingRegion>
); Japan (Nagasaki, based on historical specimens only). 
<emphasis id="B92BEAFA435ABC750CA0FC2DFE1F74E0" bold="true" box="[224,407,978,1005]" pageId="20" pageNumber="289">
New for 
<collectingCountry id="F3487678435ABC750D06FC2CFE1974E0" box="[326,401,978,1004]" name="China" pageId="20" pageNumber="289">China</collectingCountry>
.
</emphasis>
Outside of China and Japan, the species if known from: Laos (Orchymont 1925).
</paragraph>
</subSubSection>
</treatment>
</document>