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<document id="DCB3D9231F3D222A9275290E86F8F233" ID-DOI="10.5852/ejt.2024.967.2699" ID-ISSN="2118-9773" ID-Zenodo-Dep="13990819" ID-ZooBank="C08B8027-50CC-417E-BCD4-5183B9FF6738" IM.bibliography_approvedBy="guilherme" IM.illustrations_approvedBy="guilherme" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="guilherme" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="guilherme" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="guilherme" checkinTime="1729849508920" checkinUser="plazi" docAuthor="Zonstein, Sergei L." docDate="2024" docId="03A8B7235504FF95FD81E544FD49C9B2" docLanguage="en" docName="EJT.2024.967.1-185.pdf" docOrigin="European Journal of Taxonomy 967" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2699/12459" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Raveniola sororcula Zonstein, 2024, sp. nov." docType="treatment" docUuid="8ECBE7DA-43F6-4D99-96B6-C87823C6E986" docUuidSource="ZooBank" docVersion="4" lastPageNumber="48" masterDocId="FF91CF5B552AFFBAFFEFE72AFFB2CE5B" masterDocTitle="A revision of the spider genus Raveniola (Araneae, Nemesiidae). II. Species from Central Asia" masterLastPageNumber="185" masterPageNumber="1" pageNumber="47" updateTime="1730386529905" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="8A258B02E0B325A4817660EDBE1E7611" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="EE474DFFF7CA37CE1B5C535927E17A3C">A revision of the spider genus Raveniola (Araneae, Nemesiidae). II. Species from Central Asia</mods:title>
</mods:titleInfo>
<mods:name id="F65B089AF6FFD24FA475103183D43E4B" type="personal">
<mods:role id="A3BCB05E8F8061B16B4D3AF6453C9911">
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<mods:namePart id="5368F26FD7E743C19607DAEAB5186B34">Zonstein, Sergei L.</mods:namePart>
<mods:nameIdentifier id="22BEF3DABB3D603AC7F68B2C8BBD2636" type="LSID">urn:lsid:zoobank.org:author:EADD3607-30FF-49AE-93F5-8410630469BE</mods:nameIdentifier>
<mods:affiliation id="6AC9A5274F9D75C1DBB78961EB33B28B">Steinhardt Museum of Natural History, Tel-Aviv University, 69978 Tel-Aviv, Israel</mods:affiliation>
<mods:nameIdentifier id="3D6357EAE0ABB259BC417D7F884F11D5" type="email">znn@tauex.tau.ac.il</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="3BDCB7ECED321340AC19D5DD954A2440">text</mods:typeOfResource>
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<mods:title id="5DD0E313A31874CE5A28167B6402CB39">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="E8ECC5AF54435EE904FA6724BDBD1B12">
<mods:date id="D615240652A2AC13B267B6DCC3376903">2024</mods:date>
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<mods:number id="6784044AB55E6CF3D2CD66617F87CCDD">2024-10-24</mods:number>
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<mods:number id="6EA03BCE4E5AA08AFBA4B929397AA5C6">967</mods:number>
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<mods:url id="8A72F67312960820C6B0F1FA152BAE3B">https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2699/12459</mods:url>
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<mods:classification id="B39EABB12C888549792F23817C490135">journal article</mods:classification>
<mods:identifier id="D75F273C6AC5EEE8719C2F0D00CB7A37" type="DOI">10.5852/ejt.2024.967.2699</mods:identifier>
<mods:identifier id="A91841B669DD14B1185C5F22A9404C2A" type="ISSN">2118-9773</mods:identifier>
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<treatment id="03A8B7235504FF95FD81E544FD49C9B2" LSID="urn:lsid:zoobank.org:act:8ECBE7DA-43F6-4D99-96B6-C87823C6E986" httpUri="http://treatment.plazi.org/id/03A8B7235504FF95FD81E544FD49C9B2" lastPageId="47" lastPageNumber="48" pageId="46" pageNumber="47">
<subSubSection id="C31B55BE5504FF94FD81E544FB74CCF6" pageId="46" pageNumber="47" type="nomenclature">
<paragraph id="8BBE06355504FF94FD81E544FC74CCD2" blockId="46.[365,1222,622,722]" box="[622,966,622,649]" pageId="46" pageNumber="47">
<heading id="D0F6B1595504FF94FD81E544FC74CCD2" box="[622,966,622,649]" centered="true" fontSize="11" level="2" pageId="46" pageNumber="47" reason="2">
<taxonomicName id="4C017DB65504FF94FD81E544FCD6CCD2" authorityName="Zonstein" authorityYear="2024" box="[622,868,622,649]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="sororcula" status="sp. nov.">
<emphasis id="B975DA275504FF94FD81E544FCD6CCD2" bold="true" box="[622,868,622,649]" italics="true" pageId="46" pageNumber="47">Raveniola sororcula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A246675C5504FF94FC84E545FC74CCD2" box="[875,966,623,649]" pageId="46" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
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urn:lsid:zoobank.org:act:
<uuid id="FFA73CE05504FF94FD62E5B9FB74CCF6" box="[653,1222,659,685]" pageId="46" pageNumber="47">8ECBE7DA-43F6-4D99-96B6-C87823C6E986</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C31B55BE5504FF94FE26E59DFBD9CC89" box="[457,1131,695,722]" pageId="46" pageNumber="47" type="description">
<paragraph id="8BBE06355504FF94FE26E59DFBD9CC89" blockId="46.[365,1222,622,722]" box="[457,1131,695,722]" pageId="46" pageNumber="47">
<figureCitation id="133A1AB05504FF94FE26E59DFD91CC8A" box="[457,547,695,721]" captionStart="Figs 45–53" captionStartId="112.[189,241,1842,1868]" captionTargetBox="[190,1398,265,1803]" captionTargetId="figure-85@112.[189,1398,265,1803]" captionTargetPageId="112" captionText="Figs 45–53. Raveniola spp., ♀♀, paratypes (except 49, 51), habitus, dorsal views. 45–46. R. ornata sp. nov., from Ganjina and Sanglok (both SMNH), respectively. 47. R. ornatula sp. nov. (SMNH). 48. R. pamira sp. nov. (SMNH). 49. R. sororcula sp. nov., holotype (SMNH). 50. R. diluta sp. nov. (SMNH). 51. R. fedotovi(Charitonov, 1946), from Hojakurgan (SMNH). 52. R. pallenssp. nov. (SMNH). 53. R. zyuzini sp. nov. (SMNH). Scale bars: 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990836" httpUri="https://zenodo.org/record/13990836/files/figure.png" pageId="46" pageNumber="47">Figs 49</figureCitation>
,
<figureCitation id="133A1AB05504FF94FDC0E59DFDEFCC8A" box="[559,605,695,721]" captionStart="Figs 118–126" captionStartId="121.[189,241,1855,1881]" captionTargetBox="[189,1398,265,1816]" captionTargetId="figure-90@121.[189,1398,265,1816]" captionTargetPageId="121" captionText="Figs 118–126. Raveniola spp., ♀♀, paratypes (except 119, 123, 125), cephalothorax, dorsal views. 118. R. ignobilis sp. nov. (SMNH). 119. R. karategensis sp. nov., holotype (SMNH). 120. R. ornata sp. nov., from Sanglok (SMNH). 121. R. ornatula sp. nov. (SMNH). 122. R. pamira sp. nov. (SMNH). 123. R. sororculasp. nov., holotype (SMNH). 124. R. dilutasp. nov.(SMNH). 125. R. fedotovi(Charitonov, 1946), ♀ from Hojakurgan (SMNH). 126. R. pallens sp. nov. (SMNH). Scale bars: 2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990860" httpUri="https://zenodo.org/record/13990860/files/figure.png" pageId="46" pageNumber="47">123</figureCitation>
,
<figureCitation id="133A1AB05504FF94FD86E59DFD25CC8A" box="[617,663,695,721]" captionStart="Figs 172–183" captionStartId="126.[189,241,1749,1775]" captionTargetBox="[189,1399,265,1701]" captionTargetId="figure-107@126.[189,1399,265,1701]" captionTargetPageId="126" captionText="Figs 172–183. Raveniola spp., ♀♀, paratypes (except 176, 181, 183), clypeus and eye tubercle, dorsal views. 172. R. dolosa sp. nov. (SMNH). 173–174. R. hirta sp. nov. (SMNH). 175. R. ignobilis sp. nov. (SMNH). 176. R. karategensis sp. nov., holotype (SMNH). 177. R. karategensis, paratype (SMNH). 178. R. ornata sp. nov., from Sanglok (SMNH). 179. R. ornatula sp. nov. (SMNH). 180. R. pamira sp. nov. (SMNH). 181. R. sororcula sp. nov., holotype (SMNH). 182. R. diluta sp. nov. (SMNH). 183. R. fedotovi (Charitonov, 1946), ♀ from Hojakurgan (SMNH). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990872" httpUri="https://zenodo.org/record/13990872/files/figure.png" pageId="46" pageNumber="47">181</figureCitation>
,
<figureCitation id="133A1AB05504FF94FD4BE59DFD61CC8A" box="[676,723,695,721]" captionStart="Figs 238–246" captionStartId="133.[189,241,1820,1846]" captionTargetBox="[208,1381,265,1770]" captionTargetId="figure-91@133.[208,1381,265,1770]" captionTargetPageId="133" captionText="Figs 238–246. Raveniola spp., ♀♀, paratypes (except 238, 242, 244), sternum, labium and maxillae, ventral views. 238. R. karategensis sp. nov., holotype (SMNH). 239. R. ornata sp. nov., from Sanglok (SMNH). 240. R. ornatula sp. nov. (SMNH). 241. R. pamira sp. nov. (SMNH). 242. R. sororcula sp. nov., holotype (SMNH). 243. R. diluta sp. nov. (SMNH). 244. R. fedotovi (Charitonov, 1946), ♀ from Hojakurgan (SMNH). 245. R. pallens sp. nov. (SMNH). 246. R. zyuzini sp. nov. (SMNH). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990887" httpUri="https://zenodo.org/record/13990887/files/figure.png" pageId="46" pageNumber="47">242</figureCitation>
,
<figureCitation id="133A1AB05504FF94FD30E59DFCFACC89" box="[735,840,695,722]" captionStart="Figs 522–536" captionStartId="158.[189,241,1728,1754]" captionTargetBox="[189,1398,265,1688]" captionTargetId="figure-119@158.[189,1398,265,1688]" captionTargetPageId="158" captionText="Figs 522–536. Raveniola spp., ♀♀, spermathecae, dorsal views. 522–523. R. pamira sp. nov., paratype and female from Shahob, respectively (both SMNH). 524. R. sororcula sp. nov., holotype (SMNH). 525. R. sororcula, paratype (SMNH). 526–527. R. diluta sp. nov., paratype from Kondara (SMNH). 528–529. R. fedotovi (Charitonov, 1946), from Hojakurgan (SMNH). 530–531. R. pallens sp. nov., paratype (SMNH). 532–533. R. zyuzini sp. nov., paratype (SMNH). 534–536. R. ferghanensis (Zonstein, 1984), from Andijan (SMNH), non-type specimen from environs of Jalal-Abad (SMNH), and paratype from Jalal-Abad (SMNH), respectively. Abbreviations: ld = lateral diverticulum; sb = spermathecal base; st = spermathecal trunk. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990945" httpUri="https://zenodo.org/record/13990945/files/figure.png" pageId="46" pageNumber="47">524–525</figureCitation>
,
<figureCitation id="133A1AB05504FF94FCBBE59DFC0CCC89" box="[852,958,695,722]" captionStart="Figs 584–592" captionStartId="163.[189,241,1821,1847]" captionTargetBox="[189,1398,265,1789]" captionTargetId="figure-92@163.[189,1398,265,1789]" captionText="Figs 584–592. Raveniola spp., ♂♂ (584, 589, 591) and ♀♀ (585–588, 590, 592), holotypes (except 585–586, 590, 592). Spinnerets, ventral (584–585, 587, 589, 591–592) and lateral (586, 588, 590) views. 584. R. pamira sp. nov. (SMNH). 585–586. R. pamira, paratype (SMNH). 587–588. R. sororcula sp. nov. (SMNH). 589. R. diluta sp. nov. (SMNH). 590. R. diluta, paratype (SMNH). 591. R. fedotovi (Charitonov, 1946), paralectotype (PSU). 592. R. fedotovi, ♀ from Hojakurgan (SMNH). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14018408" httpUri="https://zenodo.org/record/14018408/files/figure.png" pageId="46" pageNumber="47">587–588</figureCitation>
, 691, 698, 754
</paragraph>
</subSubSection>
<subSubSection id="C31B55BE5504FF94FF52E5D3FB5ECAF7" pageId="46" pageNumber="47" type="diagnosis">
<paragraph id="8BBE06355504FF94FF52E5D3FE84CD48" blockId="46.[189,310,761,787]" box="[189,310,761,787]" pageId="46" pageNumber="47">
<heading id="D0F6B1595504FF94FF52E5D3FE84CD48" bold="true" box="[189,310,761,787]" fontSize="11" level="3" pageId="46" pageNumber="47" reason="3">
<emphasis id="B975DA275504FF94FF52E5D3FE84CD48" bold="true" box="[189,310,761,787]" pageId="46" pageNumber="47">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BBE06355504FF94FF52E403FB5ECAF7" blockId="46.[188,1399,809,1196]" pageId="46" pageNumber="47">
The rich and dense dorsal ornament of the abdomen in the new species resembles that in
<taxonomicName id="4C017DB65504FF94FAEFE403FEB9CD3C" authorityName="Zonstein" authorityYear="2024" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="ornata" status="sp. nov.">
<emphasis id="B975DA275504FF94FAEFE403FEB9CD3C" italics="true" pageId="46" pageNumber="47">Raveniola ornata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A246675C5504FF94FEFBE467FEDDCD3C" box="[276,367,845,871]" pageId="46" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="4C017DB65504FF94FE44E464FD82CD3C" authorityName="Zonstein" authorityYear="2024" box="[427,560,845,871]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="ornatula" status="sp. nov.">
<emphasis id="B975DA275504FF94FE44E464FE0ECD3C" box="[427,444,846,871]" italics="true" pageId="46" pageNumber="47">R</emphasis>
.
<emphasis id="B975DA275504FF94FE24E467FD82CD3C" box="[459,560,845,871]" italics="true" pageId="46" pageNumber="47">ornatula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A246675C5504FF94FDD6E467FD26CD3C" box="[569,660,845,871]" pageId="46" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="133A1AB05504FF94FD4AE467FD48CD3C" box="[677,762,845,871]" captionStart="Figs 45–53" captionStartId="112.[189,241,1842,1868]" captionTargetBox="[190,1398,265,1803]" captionTargetId="figure-85@112.[189,1398,265,1803]" captionTargetPageId="112" captionText="Figs 45–53. Raveniola spp., ♀♀, paratypes (except 49, 51), habitus, dorsal views. 45–46. R. ornata sp. nov., from Ganjina and Sanglok (both SMNH), respectively. 47. R. ornatula sp. nov. (SMNH). 48. R. pamira sp. nov. (SMNH). 49. R. sororcula sp. nov., holotype (SMNH). 50. R. diluta sp. nov. (SMNH). 51. R. fedotovi(Charitonov, 1946), from Hojakurgan (SMNH). 52. R. pallenssp. nov. (SMNH). 53. R. zyuzini sp. nov. (SMNH). Scale bars: 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990836" httpUri="https://zenodo.org/record/13990836/files/figure.png" pageId="46" pageNumber="47">Fig. 49</figureCitation>
cf.
<figureCitation id="133A1AB05504FF94FCC6E467FC00CD33" box="[809,946,845,872]" captionStart="Figs 45–53" captionStartId="112.[189,241,1842,1868]" captionTargetBox="[190,1398,265,1803]" captionTargetId="figure-85@112.[189,1398,265,1803]" captionTargetPageId="112" captionText="Figs 45–53. Raveniola spp., ♀♀, paratypes (except 49, 51), habitus, dorsal views. 45–46. R. ornata sp. nov., from Ganjina and Sanglok (both SMNH), respectively. 47. R. ornatula sp. nov. (SMNH). 48. R. pamira sp. nov. (SMNH). 49. R. sororcula sp. nov., holotype (SMNH). 50. R. diluta sp. nov. (SMNH). 51. R. fedotovi(Charitonov, 1946), from Hojakurgan (SMNH). 52. R. pallenssp. nov. (SMNH). 53. R. zyuzini sp. nov. (SMNH). Scale bars: 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990836" httpUri="https://zenodo.org/record/13990836/files/figure.png" pageId="46" pageNumber="47">Figs 45–47</figureCitation>
). However, unlike these congeners,
<taxonomicName id="4C017DB65504FF94FAB1E464FE9FCDD0" authorityName="Zonstein" authorityYear="2024" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="sororcula" status="sp. nov.">
<emphasis id="B975DA275504FF94FAB1E464FADDCD3C" box="[1374,1391,846,871]" italics="true" pageId="46" pageNumber="47">R</emphasis>
.
<emphasis id="B975DA275504FF94FF52E45BFE9FCDD0" box="[189,301,881,907]" italics="true" pageId="46" pageNumber="47">sororcula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A246675C5504FF94FEDCE45BFE3ECDD0" box="[307,396,881,907]" pageId="46" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
possesses a broadly oval sternum, medium-sized PMS and a very short apical segment of the PMS (vs a subcircular sternum, small PMS and digitiform apical segment of PLS;
<figureCitation id="133A1AB05504FF94FAEBE4BFFAC3CDF4" box="[1284,1393,917,943]" captionStart="Figs 238–246" captionStartId="133.[189,241,1820,1846]" captionTargetBox="[208,1381,265,1770]" captionTargetId="figure-91@133.[208,1381,265,1770]" captionTargetPageId="133" captionText="Figs 238–246. Raveniola spp., ♀♀, paratypes (except 238, 242, 244), sternum, labium and maxillae, ventral views. 238. R. karategensis sp. nov., holotype (SMNH). 239. R. ornata sp. nov., from Sanglok (SMNH). 240. R. ornatula sp. nov. (SMNH). 241. R. pamira sp. nov. (SMNH). 242. R. sororcula sp. nov., holotype (SMNH). 243. R. diluta sp. nov. (SMNH). 244. R. fedotovi (Charitonov, 1946), ♀ from Hojakurgan (SMNH). 245. R. pallens sp. nov. (SMNH). 246. R. zyuzini sp. nov. (SMNH). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990887" httpUri="https://zenodo.org/record/13990887/files/figure.png" pageId="46" pageNumber="47">Figs 242</figureCitation>
,
<figureCitation id="133A1AB05504FF94FF52E493FE96CD8F" box="[189,292,953,980]" captionStart="Figs 584–592" captionStartId="163.[189,241,1821,1847]" captionTargetBox="[189,1398,265,1789]" captionTargetId="figure-92@163.[189,1398,265,1789]" captionText="Figs 584–592. Raveniola spp., ♂♂ (584, 589, 591) and ♀♀ (585–588, 590, 592), holotypes (except 585–586, 590, 592). Spinnerets, ventral (584–585, 587, 589, 591–592) and lateral (586, 588, 590) views. 584. R. pamira sp. nov. (SMNH). 585–586. R. pamira, paratype (SMNH). 587–588. R. sororcula sp. nov. (SMNH). 589. R. diluta sp. nov. (SMNH). 590. R. diluta, paratype (SMNH). 591. R. fedotovi (Charitonov, 1946), paralectotype (PSU). 592. R. fedotovi, ♀ from Hojakurgan (SMNH). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14018408" httpUri="https://zenodo.org/record/14018408/files/figure.png" pageId="46" pageNumber="47">587–588</figureCitation>
cf.
<figureCitation id="133A1AB05504FF94FEBCE493FE45CD88" box="[339,503,953,979]" captionStart="Figs 238–246" captionStartId="133.[189,241,1820,1846]" captionTargetBox="[208,1381,265,1770]" captionTargetId="figure-91@133.[208,1381,265,1770]" captionTargetPageId="133" captionText="Figs 238–246. Raveniola spp., ♀♀, paratypes (except 238, 242, 244), sternum, labium and maxillae, ventral views. 238. R. karategensis sp. nov., holotype (SMNH). 239. R. ornata sp. nov., from Sanglok (SMNH). 240. R. ornatula sp. nov. (SMNH). 241. R. pamira sp. nov. (SMNH). 242. R. sororcula sp. nov., holotype (SMNH). 243. R. diluta sp. nov. (SMNH). 244. R. fedotovi (Charitonov, 1946), ♀ from Hojakurgan (SMNH). 245. R. pallens sp. nov. (SMNH). 246. R. zyuzini sp. nov. (SMNH). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990887" httpUri="https://zenodo.org/record/13990887/files/figure.png" pageId="46" pageNumber="47">Figs 239–240</figureCitation>
,
<figureCitation id="133A1AB05504FF94FDEBE493FDDCCD8F" box="[516,622,953,980]" captionStart="Figs 575–583" captionStartId="162.[189,241,1857,1883]" captionTargetBox="[189,1398,265,1816]" captionTargetId="figure-74@162.[189,1398,265,1816]" captionText="Figs 575–583. Raveniola spp., ♂♂ (575, 577, 581) and ♀♀ (576, 578–580, 582–583); holotypes (575– 577, 581) and paratypes (578–580, 582–583). Spinnerets (575–579, 581–582) and entire abdomen (580, 583), ventral (575–576, 579–580, 582–583) and lateral (577–578, 581) views. 575. R. insolita sp. nov. (ZISP). 576. R. karategensis sp. nov. (SMNH). 577–580. R. ornata sp. nov. (SMNH). 581– 583. R. ornatula sp. nov. (SMNH). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14018406" httpUri="https://zenodo.org/record/14018406/files/figure.png" pageId="46" pageNumber="47">579, 582</figureCitation>
) and lacks any more-or-less developed ventral abdominal pattern (such as shown in
<figureCitation id="133A1AB05504FF94FE79E4F7FD88CDA3" box="[406,570,989,1016]" captionStart="Figs 575–583" captionStartId="162.[189,241,1857,1883]" captionTargetBox="[189,1398,265,1816]" captionTargetId="figure-74@162.[189,1398,265,1816]" captionText="Figs 575–583. Raveniola spp., ♂♂ (575, 577, 581) and ♀♀ (576, 578–580, 582–583); holotypes (575– 577, 581) and paratypes (578–580, 582–583). Spinnerets (575–579, 581–582) and entire abdomen (580, 583), ventral (575–576, 579–580, 582–583) and lateral (577–578, 581) views. 575. R. insolita sp. nov. (ZISP). 576. R. karategensis sp. nov. (SMNH). 577–580. R. ornata sp. nov. (SMNH). 581– 583. R. ornatula sp. nov. (SMNH). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14018406" httpUri="https://zenodo.org/record/14018406/files/figure.png" pageId="46" pageNumber="47">Figs 580, 583</figureCitation>
). In the structure of the spermathecae,
<taxonomicName id="4C017DB65504FF94FC11E4F4FB3CCDAC" authorityName="Zonstein" authorityYear="2024" box="[1022,1166,989,1016]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="sororcula">
<emphasis id="B975DA275504FF94FC11E4F4FBA2CDAC" box="[1022,1040,990,1015]" italics="true" pageId="46" pageNumber="47">R</emphasis>
.
<emphasis id="B975DA275504FF94FBF1E4F7FB3CCDAC" box="[1054,1166,989,1015]" italics="true" pageId="46" pageNumber="47">sororcula</emphasis>
</taxonomicName>
seems to be similar to
<taxonomicName id="4C017DB65504FF94FF31E328FEE2CA40" authorityName="Zonstein" authorityYear="2024" box="[222,336,1025,1052]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="dolosa" status="sp. nov.">
<emphasis id="B975DA275504FF94FF31E328FF42CA40" box="[222,240,1026,1051]" italics="true" pageId="46" pageNumber="47">R</emphasis>
.
<emphasis id="B975DA275504FF94FEEDE32BFEE2CA40" box="[258,336,1025,1051]" italics="true" pageId="46" pageNumber="47">dolosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A246675C5504FF94FEB5E328FE0ACA47" box="[346,440,1026,1052]" pageId="46" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="133A1AB05504FF94FE25E32BFDDDCA47" box="[458,623,1025,1052]" captionStart="Figs 522–536" captionStartId="158.[189,241,1728,1754]" captionTargetBox="[189,1398,265,1688]" captionTargetId="figure-119@158.[189,1398,265,1688]" captionTargetPageId="158" captionText="Figs 522–536. Raveniola spp., ♀♀, spermathecae, dorsal views. 522–523. R. pamira sp. nov., paratype and female from Shahob, respectively (both SMNH). 524. R. sororcula sp. nov., holotype (SMNH). 525. R. sororcula, paratype (SMNH). 526–527. R. diluta sp. nov., paratype from Kondara (SMNH). 528–529. R. fedotovi (Charitonov, 1946), from Hojakurgan (SMNH). 530–531. R. pallens sp. nov., paratype (SMNH). 532–533. R. zyuzini sp. nov., paratype (SMNH). 534–536. R. ferghanensis (Zonstein, 1984), from Andijan (SMNH), non-type specimen from environs of Jalal-Abad (SMNH), and paratype from Jalal-Abad (SMNH), respectively. Abbreviations: ld = lateral diverticulum; sb = spermathecal base; st = spermathecal trunk. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990945" httpUri="https://zenodo.org/record/13990945/files/figure.png" pageId="46" pageNumber="47">Figs 524–525</figureCitation>
cf.
<figureCitation id="133A1AB05504FF94FD4DE32BFCF9CA47" box="[674,843,1025,1052]" captionStart="Figs 487–503" captionStartId="156.[189,241,1695,1721]" captionTargetBox="[189,1398,265,1653]" captionTargetId="figure-102@156.[189,1398,265,1653]" captionTargetPageId="156" captionText="Figs 487–503. Raveniola spp., ♀♀, paratypes (except 487–489), spermathecae, dorsal views. 487–488. R. redikorzevi (Spassky, 1937), Kepele (SMNH). 489. R. afghana sp. nov. (NHMD). 490. Same, holotype. 491–493. R. alajensis sp. nov., Berksu. 494. R. concolor Zonstein, 2000 (MCSN AR 19). 495–496. R. cucullata sp. nov., Shikildara (SMNH). 497. R. cucullata, Shahob (SMNH). 498– 499. R. cucullata, Sangdara (SMNH). 500–503. R. dolosa sp. nov. (SMNH). Abbreviations: isb = inner spermathecal branch; osb = outer spermathecal branch; sb = spermathecal base. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990941" httpUri="https://zenodo.org/record/13990941/files/figure.png" pageId="46" pageNumber="47">Figs 500–503</figureCitation>
); however, it differs from the latter species in possessing clearly larger PMS, compared with small PMS in
<taxonomicName id="4C017DB65504FF94FC65E30CFC4BCA64" authorityName="Zonstein" authorityYear="2024" box="[906,1017,1061,1088]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="dolosa">
<emphasis id="B975DA275504FF94FC65E30CFC2ECA64" box="[906,924,1062,1087]" italics="true" pageId="46" pageNumber="47">R</emphasis>
.
<emphasis id="B975DA275504FF94FC44E30FFC4BCA64" box="[939,1017,1061,1087]" italics="true" pageId="46" pageNumber="47">dolosa</emphasis>
</taxonomicName>
(see
<figureCitation id="133A1AB05504FF94FBD8E30FFB2ACA1B" box="[1079,1176,1061,1088]" captionStart="Figs 584–592" captionStartId="163.[189,241,1821,1847]" captionTargetBox="[189,1398,265,1789]" captionTargetId="figure-92@163.[189,1398,265,1789]" captionText="Figs 584–592. Raveniola spp., ♂♂ (584, 589, 591) and ♀♀ (585–588, 590, 592), holotypes (except 585–586, 590, 592). Spinnerets, ventral (584–585, 587, 589, 591–592) and lateral (586, 588, 590) views. 584. R. pamira sp. nov. (SMNH). 585–586. R. pamira, paratype (SMNH). 587–588. R. sororcula sp. nov. (SMNH). 589. R. diluta sp. nov. (SMNH). 590. R. diluta, paratype (SMNH). 591. R. fedotovi (Charitonov, 1946), paralectotype (PSU). 592. R. fedotovi, ♀ from Hojakurgan (SMNH). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14018408" httpUri="https://zenodo.org/record/14018408/files/figure.png" pageId="46" pageNumber="47">Fig. 587</figureCitation>
cf.
<figureCitation id="133A1AB05504FF94FB2AE30FFAD8CA1B" box="[1221,1386,1061,1088]" captionStart="Figs 565–574" captionStartId="161.[189,241,1825,1851]" captionTargetBox="[202,1386,265,1765]" captionTargetId="figure-100@161.[202,1386,265,1784]" captionText="Figs 565–574. Raveniola spp., ♂♂ (565, 568, 571, 573) and ♀♀ (566–567, 569–570, 572), paratypes (except 568, 571, 573). Spinnerets, ventral (565–566, 571–574), lateral (567–568) and ventrolateral (569–570) views. 565–566. R. cucullata sp. nov., paratypes from Shahob (SMNH). 567. R. cucullata, from Sangdara (SMNH). 568. R. dolosa sp. nov. (SMNH). 569–570. R. dolosa, paratypes (SMNH). 571. R. hirta sp. nov., holotype (SMNH). 572. R. hirta, paratype (SMNH). 573. R. ignobilis sp. nov. (SMNH). 574. R. ignobilis, paratype (SMNH). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14018403" httpUri="https://zenodo.org/record/14018403/files/figure.png" pageId="46" pageNumber="47">Figs 569–570</figureCitation>
). Finally, this new species differs from the sympatric congener,
<taxonomicName id="4C017DB65504FF94FC5CE360FB9ECA38" authorityName="Zonstein" authorityYear="2024" box="[947,1068,1098,1124]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="pamira" status="sp. nov.">
<emphasis id="B975DA275504FF94FC5CE360FC77CA38" box="[947,965,1098,1123]" italics="true" pageId="46" pageNumber="47">R</emphasis>
.
<emphasis id="B975DA275504FF94FC38E360FB9ECA38" box="[983,1068,1098,1123]" italics="true" pageId="46" pageNumber="47">pamira</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A246675C5504FF94FBD9E360FB24CA3F" box="[1078,1174,1098,1124]" pageId="46" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
, in having a weak cheliceral rastellum and a shorter apical segment of the PLS (vs the absence of rastellar setae and a noticeably longer apical segment of the PLS in the latter species; see
<figureCitation id="133A1AB05504FF94FC05E3BBFBFECAF7" box="[1002,1100,1169,1196]" captionStart="Figs 584–592" captionStartId="163.[189,241,1821,1847]" captionTargetBox="[189,1398,265,1789]" captionTargetId="figure-92@163.[189,1398,265,1789]" captionText="Figs 584–592. Raveniola spp., ♂♂ (584, 589, 591) and ♀♀ (585–588, 590, 592), holotypes (except 585–586, 590, 592). Spinnerets, ventral (584–585, 587, 589, 591–592) and lateral (586, 588, 590) views. 584. R. pamira sp. nov. (SMNH). 585–586. R. pamira, paratype (SMNH). 587–588. R. sororcula sp. nov. (SMNH). 589. R. diluta sp. nov. (SMNH). 590. R. diluta, paratype (SMNH). 591. R. fedotovi (Charitonov, 1946), paralectotype (PSU). 592. R. fedotovi, ♀ from Hojakurgan (SMNH). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14018408" httpUri="https://zenodo.org/record/14018408/files/figure.png" pageId="46" pageNumber="47">Fig. 586</figureCitation>
cf.
<figureCitation id="133A1AB05504FF94FB96E3BBFB6DCAF7" box="[1145,1247,1169,1196]" captionStart="Figs 584–592" captionStartId="163.[189,241,1821,1847]" captionTargetBox="[189,1398,265,1789]" captionTargetId="figure-92@163.[189,1398,265,1789]" captionText="Figs 584–592. Raveniola spp., ♂♂ (584, 589, 591) and ♀♀ (585–588, 590, 592), holotypes (except 585–586, 590, 592). Spinnerets, ventral (584–585, 587, 589, 591–592) and lateral (586, 588, 590) views. 584. R. pamira sp. nov. (SMNH). 585–586. R. pamira, paratype (SMNH). 587–588. R. sororcula sp. nov. (SMNH). 589. R. diluta sp. nov. (SMNH). 590. R. diluta, paratype (SMNH). 591. R. fedotovi (Charitonov, 1946), paralectotype (PSU). 592. R. fedotovi, ♀ from Hojakurgan (SMNH). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14018408" httpUri="https://zenodo.org/record/14018408/files/figure.png" pageId="46" pageNumber="47">Fig. 588</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C31B55BE5504FF94FF52E3FEFC1FCB19" pageId="46" pageNumber="47" type="etymology">
<paragraph id="8BBE06355504FF94FF52E3FEFEF6CAB5" blockId="46.[189,324,1236,1262]" box="[189,324,1236,1262]" pageId="46" pageNumber="47">
<heading id="D0F6B1595504FF94FF52E3FEFEF6CAB5" bold="true" box="[189,324,1236,1262]" fontSize="11" level="3" pageId="46" pageNumber="47" reason="3">
<emphasis id="B975DA275504FF94FF52E3FEFEF6CAB5" bold="true" box="[189,324,1236,1262]" pageId="46" pageNumber="47">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BBE06355504FF94FF52E229FC1FCB19" blockId="46.[189,1399,1283,1346]" pageId="46" pageNumber="47">
The specific epithet is a Latin noun meaning ‘a little sister’; the name refers to a smaller species size, compared to that of a larger sibling neighbor,
<taxonomicName id="4C017DB65504FF94FD39E202FCF9CB1A" authorityName="Zonstein" authorityYear="2024" box="[726,843,1320,1346]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="pamira" status="sp. nov.">
<emphasis id="B975DA275504FF94FD39E202FD5ACB1A" box="[726,744,1320,1345]" italics="true" pageId="46" pageNumber="47">R</emphasis>
.
<emphasis id="B975DA275504FF94FD19E202FCF9CB1A" box="[758,843,1320,1345]" italics="true" pageId="46" pageNumber="47">pamira</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A246675C5504FF94FCBDE202FC1FCB19" box="[850,941,1320,1346]" pageId="46" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C31B55BE5504FF94FF52E243FE73C8F1" pageId="46" pageNumber="47" type="materials_examined">
<paragraph id="8BBE06355504FF94FF52E243FE18CBD8" blockId="46.[189,426,1385,1411]" box="[189,426,1385,1411]" pageId="46" pageNumber="47">
<heading id="D0F6B1595504FF94FF52E243FE18CBD8" bold="true" box="[189,426,1385,1411]" fontSize="11" level="3" pageId="46" pageNumber="47" reason="3">
<emphasis id="B975DA275504FF94FF52E243FE18CBD8" bold="true" box="[189,426,1385,1411]" pageId="46" pageNumber="47">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BBE06355504FF94FF05E2B3FEEECBE8" blockId="46.[189,1398,1433,1532]" box="[234,348,1433,1459]" pageId="46" pageNumber="47">
<emphasis id="B975DA275504FF94FF05E2B3FEEECBE8" bold="true" box="[234,348,1433,1459]" pageId="46" pageNumber="47">
<typeStatus id="54BAB8975504FF94FF05E2B3FEEECBE8" box="[234,348,1433,1459]" pageId="46" pageNumber="47" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BBE06355504FF94FF52E297FAC4CBA7" blockId="46.[189,1398,1433,1532]" pageId="46" pageNumber="47">
<materialsCitation id="3B690C685504FF94FF52E297FAC4CBA7" collectingDate="2019-07-15" collectionCode="SMNH" collectorName="S. Zonstein &amp; A. Hakimov" country="Tajikistan" county="Hurk Village" elevation="1650" latitude="38.516666" location="Western Pamir" longLatPrecision="1175" longitude="71.03333" municipality="Obiviskharvi Canyon" pageId="46" pageNumber="47" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<collectingCountry id="F31646A55504FF94FF52E297FED2CB8C" box="[189,352,1469,1495]" name="Tajikistan" pageId="46" pageNumber="47">TAJIKISTAN</collectingCountry>
•
<specimenCount id="9D07CDBC5504FF94FE99E294FE3FCB83" box="[374,397,1470,1496]" count="1" pageId="46" pageNumber="47" type="female">♀</specimenCount>
;
<location id="8EDE50EE5504FF94FE75E294FDF5CB8C" LSID="urn:lsid:plazi:treatment:03A8B7235504FF95FD81E544FD49C9B2:8EDE50EE5504FF94FE75E294FDF5CB8C" box="[410,583,1469,1496]" country="Tajikistan" county="Hurk Village" latitude="38.516666" longLatPrecision="1175" longitude="71.03333" municipality="Obiviskharvi Canyon" name="Western Pamir" pageId="46" pageNumber="47">Western Pamir</location>
,
<location id="8EDE50EE5504FF94FDBEE294FD68CB83" LSID="urn:lsid:plazi:treatment:03A8B7235504FF95FD81E544FD49C9B2:8EDE50EE5504FF94FDBEE294FD68CB83" box="[593,730,1470,1496]" country="Tajikistan" county="Hurk Village" latitude="38.516666" longLatPrecision="1175" longitude="71.03333" municipality="Obiviskharvi Canyon" name="Darvaz Mts" pageId="46" pageNumber="47">Darvaz Mts</location>
(southern slope),
<collectingMunicipality id="6BDA9C4F5504FF94FC46E297FB14CB8C" box="[937,1190,1469,1495]" pageId="46" pageNumber="47">Obiviskharvi Canyon</collectingMunicipality>
, environs of
<collectingCounty id="62DF7EB95504FF94FAD5E297FEA7CBA0" pageId="46" pageNumber="47">Hurk Village</collectingCounty>
;
<geoCoordinate id="EE3560F25504FF94FECFE2CAFE39CBA7" box="[288,395,1504,1532]" degrees="38" direction="north" minutes="31" orientation="latitude" pageId="46" pageNumber="47" precision="925" value="38.516666">38°31′ N</geoCoordinate>
,
<geoCoordinate id="EE3560F25504FF94FE79E2CAFE4CCBA7" box="[406,510,1504,1532]" degrees="71" direction="east" minutes="02" orientation="longitude" pageId="46" pageNumber="47" precision="925" value="71.03333">71°02′ E</geoCoordinate>
;
<elevation id="002CE1065504FF94FDE6E2CBFD5BCBA0" box="[521,745,1505,1532]" metricMagnitude="3" metricUnit="m" metricValue="1.65" metricValueMax="1.7" metricValueMin="1.6" pageId="46" pageNumber="47" unit="m" value="1650.0" valueMax="1700.0" valueMin="1600.0">
<quantity id="4CF9ABD05504FF94FDE6E2CBFD18CBA7" box="[521,682,1505,1532]" metricMagnitude="3" metricUnit="m" metricValue="1.65" metricValueMax="1.7" metricValueMin="1.6" pageId="46" pageNumber="47" unit="m" value="1650.0" valueMax="1700.0" valueMin="1600.0">1600–1700 m</quantity>
a.s.l.
</elevation>
;
<date id="FFBF20F55504FF94FD1AE2CBFC34CBA0" box="[757,902,1505,1532]" pageId="46" pageNumber="47" value="2019-07-15">
<collectingDate id="EFFBD91D5504FF94FD1AE2CBFC34CBA0" box="[757,902,1505,1532]" pageId="46" pageNumber="47" value="2019-07-15">15 Jul. 2019</collectingDate>
</date>
;
<collectorName id="26F463E35504FF94FC7EE2CBFBA4CBA0" box="[913,1046,1505,1531]" pageId="46" pageNumber="47">S. Zonstein</collectorName>
and
<collectorName id="26F463E35504FF94FBA4E2CBFB69CBA0" box="[1099,1243,1505,1531]" pageId="46" pageNumber="47">A. Hakimov</collectorName>
leg.;
<collectionCode id="ED109EF05504FF94FAF5E2CBFADDCBA0" box="[1306,1391,1505,1531]" country="Sweden" httpUri="http://grbio.org/cool/8z6v-f8is" name="Department of Paleozoology, Swedish Museum of Natural History" pageId="46" pageNumber="47">SMNH</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BBE06355504FF94FF05E10EFAC5C8DD" blockId="46.[189,1399,1572,1706]" pageId="46" pageNumber="47">
<materialsCitation id="3B690C685504FF94FF05E10EFED2C839" country="Tajikistan" location="TAJIKISTAN" pageId="46" pageNumber="47" specimenCount="3" specimenCount-female="3" typeStatus="paratype">
<emphasis id="B975DA275504FF94FF05E10EFED5C865" bold="true" box="[234,359,1572,1598]" pageId="46" pageNumber="47">
<typeStatus id="54BAB8975504FF94FF05E10EFED5C865" box="[234,359,1572,1598]" pageId="46" pageNumber="47" type="paratype">Paratypes</typeStatus>
</emphasis>
(
<specimenCount id="9D07CDBC5504FF94FE96E10EFE0BC865" box="[377,441,1572,1598]" count="3" pageId="46" pageNumber="47" type="female">3 ♀♀</specimenCount>
)
<collectingCountry id="F31646A55504FF94FF52E162FED2C839" box="[189,352,1608,1634]" name="Tajikistan" pageId="46" pageNumber="47">TAJIKISTAN</collectingCountry>
</materialsCitation>
•
<materialsCitation id="3B690C685504FF94FE94E162FC53C839" box="[379,993,1608,1634]" collectingDate="2019-07-15" collectionCode="SMNH" collectorName="S. Zonstein &amp; A. Hakimov" country="Tajikistan" county="Hurk Village" elevation="1650" latitude="38.516666" location="Western Pamir" longLatPrecision="1175" longitude="71.03333" municipality="Obiviskharvi Canyon" pageId="46" pageNumber="47" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<specimenCount id="9D07CDBC5504FF94FE94E162FE0DC839" box="[379,447,1608,1634]" count="2" pageId="46" pageNumber="47" type="female">2 ♀♀</specimenCount>
; same collection data as for holotype;
<collectionCode id="ED109EF05504FF94FC63E162FC53C839" box="[908,993,1608,1634]" country="Sweden" httpUri="http://grbio.org/cool/8z6v-f8is" name="Department of Paleozoology, Swedish Museum of Natural History" pageId="46" pageNumber="47">SMNH</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B690C685504FF94FC13E162FAC0C8DD" collectingDate="1988-07-15" collectionCode="SMNH" collectorName="S. Zonstein &amp; A. Hakimov" country="Tajikistan" county="Hurk Village" elevation="2025" latitude="38.533333" location="Ubagan Village" longLatPrecision="1175" longitude="71.05" municipality="Obiviskharvi Canyon" pageId="46" pageNumber="47" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="9D07CDBC5504FF94FC13E162FB99C839" box="[1020,1067,1608,1634]" count="1" pageId="46" pageNumber="47" type="female">1 ♀</specimenCount>
; same collection data as for preceding, surroundings of
<location id="8EDE50EE5504FF94FDEAE146FD73C8DD" LSID="urn:lsid:plazi:treatment:03A8B7235504FF95FD81E544FD49C9B2:8EDE50EE5504FF94FDEAE146FD73C8DD" box="[517,705,1644,1670]" country="Tajikistan" latitude="38.533333" longLatPrecision="1175" longitude="71.05" name="Ubagan Village" pageId="46" pageNumber="47">Ubagan Village</location>
;
<geoCoordinate id="EE3560F25504FF94FD20E140FC8FC8DD" box="[719,829,1642,1670]" degrees="38" direction="north" minutes="32" orientation="latitude" pageId="46" pageNumber="47" precision="925" value="38.533333">38°32′ N</geoCoordinate>
,
<geoCoordinate id="EE3560F25504FF94FCA4E140FC04C8DD" box="[843,950,1642,1670]" degrees="71" direction="east" minutes="03" orientation="longitude" pageId="46" pageNumber="47" precision="925" value="71.05">71°03′ E</geoCoordinate>
;
<elevation id="002CE1065504FF94FC2BE146FB18C8DD" box="[964,1194,1644,1671]" metricMagnitude="3" metricUnit="m" metricValue="2.025" metricValueMax="2.1" metricValueMin="1.95" pageId="46" pageNumber="47" unit="m" value="2025.0" valueMax="2100.0" valueMin="1950.0">
<quantity id="4CF9ABD05504FF94FC2BE146FBDAC8DD" box="[964,1128,1644,1671]" metricMagnitude="3" metricUnit="m" metricValue="2.025" metricValueMax="2.1" metricValueMin="1.95" pageId="46" pageNumber="47" unit="m" value="2025.0" valueMax="2100.0" valueMin="1950.0">1950–2100 m</quantity>
a.s.l.
</elevation>
;
<date id="FFBF20F55504FF94FB56E146FAE3C8DD" box="[1209,1361,1644,1671]" pageId="46" pageNumber="47" value="1988-07-15">
<collectingDate id="EFFBD91D5504FF94FB56E146FAE3C8DD" box="[1209,1361,1644,1671]" pageId="46" pageNumber="47" value="1988-07-15">15 Jul. 1988</collectingDate>
</date>
; S
</materialsCitation>
.
</paragraph>
<paragraph id="8BBE06355504FF94FF50E1BAFE73C8F1" blockId="46.[189,1399,1572,1706]" box="[191,449,1680,1706]" pageId="46" pageNumber="47">
Zonstein leg.;
<collectionCode id="ED109EF05504FF94FE87E1BAFE09C8F1" box="[360,443,1680,1706]" country="Sweden" httpUri="http://grbio.org/cool/8z6v-f8is" name="Department of Paleozoology, Swedish Museum of Natural History" pageId="46" pageNumber="47">SMNH</collectionCode>
.
</paragraph>
</subSubSection>
<subSubSection id="C31B55BE5504FF95FF52E1F8FDEDC8B8" lastPageId="47" lastPageNumber="48" pageId="46" pageNumber="47" type="description">
<paragraph id="8BBE06355504FF94FF52E1F8FEFCC8B7" blockId="46.[189,334,1746,1772]" box="[189,334,1746,1772]" pageId="46" pageNumber="47">
<heading id="D0F6B1595504FF94FF52E1F8FEFCC8B7" bold="true" box="[189,334,1746,1772]" fontSize="11" level="3" pageId="46" pageNumber="47" reason="3">
<emphasis id="B975DA275504FF94FF52E1F8FEFCC8B7" bold="true" box="[189,334,1746,1772]" pageId="46" pageNumber="47">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BBE06355504FF94FF05E028FE75C940" blockId="46.[189,455,1793,1856]" box="[234,455,1793,1820]" pageId="46" pageNumber="47">
<emphasis id="B975DA275504FF94FF05E028FEF7C947" bold="true" box="[234,325,1794,1820]" pageId="46" pageNumber="47">Female</emphasis>
(
<typeStatus id="54BAB8975504FF94FEBAE02BFE0DC940" box="[341,447,1793,1819]" pageId="46" pageNumber="47" type="holotype">holotype</typeStatus>
)
</paragraph>
<paragraph id="8BBE06355504FF94FF52E00CFE05C91B" blockId="46.[189,455,1793,1856]" box="[189,439,1830,1856]" pageId="46" pageNumber="47">
<smallCapsWord id="8D5890E95504FF94FF52E00CFEAEC965" baselines="1849,1849" box="[189,284,1830,1856]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Habitus" pageId="46" pageNumber="47">HABITUS</smallCapsWord>
. See
<figureCitation id="133A1AB05504FF94FEB2E00CFE05C91B" box="[349,439,1830,1856]" captionStart="Figs 45–53" captionStartId="112.[189,241,1842,1868]" captionTargetBox="[190,1398,265,1803]" captionTargetId="figure-85@112.[189,1398,265,1803]" captionTargetPageId="112" captionText="Figs 45–53. Raveniola spp., ♀♀, paratypes (except 49, 51), habitus, dorsal views. 45–46. R. ornata sp. nov., from Ganjina and Sanglok (both SMNH), respectively. 47. R. ornatula sp. nov. (SMNH). 48. R. pamira sp. nov. (SMNH). 49. R. sororcula sp. nov., holotype (SMNH). 50. R. diluta sp. nov. (SMNH). 51. R. fedotovi(Charitonov, 1946), from Hojakurgan (SMNH). 52. R. pallenssp. nov. (SMNH). 53. R. zyuzini sp. nov. (SMNH). Scale bars: 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990836" httpUri="https://zenodo.org/record/13990836/files/figure.png" pageId="46" pageNumber="47">Fig. 49.</figureCitation>
</paragraph>
<paragraph id="8BBE06355504FF94FF52E042FB15C9D9" blockId="46.[189,1191,1896,1923]" box="[189,1191,1896,1923]" pageId="46" pageNumber="47">
<smallCapsWord id="8D5890E95504FF94FF52E042FEDEC9DB" baselines="1915,1915" box="[189,364,1896,1922]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="46" pageNumber="47">MEASUREMENTS</smallCapsWord>
. TBL 18.35, CL 6.77, CW 6.10, LL 0.64, LW 1.21, SL 3.55, SW 3.17.
</paragraph>
<paragraph id="8BBE06355504FF95FF52E080FCB2CF30" blockId="46.[189,1399,1962,2025]" lastBlockId="47.[189,1399,265,363]" lastPageId="47" lastPageNumber="48" pageId="46" pageNumber="47">
<smallCapsWord id="8D5890E95504FF94FF52E080FEAAC998" baselines="1981,1982" box="[189,280,1962,1988]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Colour" pageId="46" pageNumber="47">COLOUR</smallCapsWord>
. Carapace dark reddish brown; eye tubercle dark brown with eyes encircled with wide blackish brown rings; chelicerae very dark cherry brown; sternum, labium, maxillae, palps and legs I–IV medium to dark brownish orange; abdomen medium fawn brown, dorsally with rich and dense dark reddish brown reticulate ornament (similar in its shade and intensity to carapace colouration); book lungs, epigastrum and spinnerets light yellowish brown.
</paragraph>
<paragraph id="8BBE06355505FF95FF52E6B9FC75CC3B" blockId="47.[188,1399,403,608]" pageId="47" pageNumber="48">
<smallCapsWord id="8D5890E95505FF95FF52E6B9FECACFF0" baselines="422,422" box="[189,376,403,429]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Cephalothorax" pageId="47" pageNumber="48">CEPHALOTHORAX</smallCapsWord>
. Carapace and chelicerae as shown in
<figureCitation id="133A1AB05505FF95FCADE6B9FC1ACFF6" box="[834,936,403,429]" captionStart="Figs 118–126" captionStartId="121.[189,241,1855,1881]" captionTargetBox="[189,1398,265,1816]" captionTargetId="figure-90@121.[189,1398,265,1816]" captionTargetPageId="121" captionText="Figs 118–126. Raveniola spp., ♀♀, paratypes (except 119, 123, 125), cephalothorax, dorsal views. 118. R. ignobilis sp. nov. (SMNH). 119. R. karategensis sp. nov., holotype (SMNH). 120. R. ornata sp. nov., from Sanglok (SMNH). 121. R. ornatula sp. nov. (SMNH). 122. R. pamira sp. nov. (SMNH). 123. R. sororculasp. nov., holotype (SMNH). 124. R. dilutasp. nov.(SMNH). 125. R. fedotovi(Charitonov, 1946), ♀ from Hojakurgan (SMNH). 126. R. pallens sp. nov. (SMNH). Scale bars: 2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990860" httpUri="https://zenodo.org/record/13990860/files/figure.png" pageId="47" pageNumber="48">Fig. 123</figureCitation>
. Clypeus and eye group as in
<figureCitation id="133A1AB05505FF95FAE3E6B9FAC0CFF6" box="[1292,1394,403,429]" captionStart="Figs 172–183" captionStartId="126.[189,241,1749,1775]" captionTargetBox="[189,1399,265,1701]" captionTargetId="figure-107@126.[189,1399,265,1701]" captionTargetPageId="126" captionText="Figs 172–183. Raveniola spp., ♀♀, paratypes (except 176, 181, 183), clypeus and eye tubercle, dorsal views. 172. R. dolosa sp. nov. (SMNH). 173–174. R. hirta sp. nov. (SMNH). 175. R. ignobilis sp. nov. (SMNH). 176. R. karategensis sp. nov., holotype (SMNH). 177. R. karategensis, paratype (SMNH). 178. R. ornata sp. nov., from Sanglok (SMNH). 179. R. ornatula sp. nov. (SMNH). 180. R. pamira sp. nov. (SMNH). 181. R. sororcula sp. nov., holotype (SMNH). 182. R. diluta sp. nov. (SMNH). 183. R. fedotovi (Charitonov, 1946), ♀ from Hojakurgan (SMNH). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990872" httpUri="https://zenodo.org/record/13990872/files/figure.png" pageId="47" pageNumber="48">Fig. 181</figureCitation>
. Eye diameters and interdistances: AME 0.15(0.21), ALE 0.29, PLE 0.22, PME 0.15, AME–AME 0.16(0.10), ALE–AME 0.14(0.11), ALE–PLE 0.16, PLE–PME 0.10, PME–PME 0.47. Dorsodistal edge of chelicerae with a transverse row of ca 20 thickened and partially broken setae in front of fang base. Each cheliceral furrow with 8 promarginal teeth and 7–8 mesobasal denticles. Sternum, labium and maxillae as shown in
<figureCitation id="133A1AB05505FF95FE54E56CFD93CC3B" box="[443,545,582,608]" captionStart="Figs 238–246" captionStartId="133.[189,241,1820,1846]" captionTargetBox="[208,1381,265,1770]" captionTargetId="figure-91@133.[208,1381,265,1770]" captionTargetPageId="133" captionText="Figs 238–246. Raveniola spp., ♀♀, paratypes (except 238, 242, 244), sternum, labium and maxillae, ventral views. 238. R. karategensis sp. nov., holotype (SMNH). 239. R. ornata sp. nov., from Sanglok (SMNH). 240. R. ornatula sp. nov. (SMNH). 241. R. pamira sp. nov. (SMNH). 242. R. sororcula sp. nov., holotype (SMNH). 243. R. diluta sp. nov. (SMNH). 244. R. fedotovi (Charitonov, 1946), ♀ from Hojakurgan (SMNH). 245. R. pallens sp. nov. (SMNH). 246. R. zyuzini sp. nov. (SMNH). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990887" httpUri="https://zenodo.org/record/13990887/files/figure.png" pageId="47" pageNumber="48">Fig. 242</figureCitation>
. Maxillae with ca 80 cuspules each.
</paragraph>
<paragraph id="8BBE06355505FF95FF52E5A3FEE2CD56" blockId="47.[188,1401,648,781]" pageId="47" pageNumber="48">
<smallCapsWord id="8D5890E95505FF95FF52E5A3FF44CCFA" baselines="668,668" box="[189,246,649,675]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Legs" pageId="47" pageNumber="48">LEGS</smallCapsWord>
. Scopula: entire and distal on metatarsi I–II; entire on palpal tarsus and tarsi I–II; widely divided by setae on tarsi III–IV. Trichobothria: 2 rows of 9–10 each on tibiae, 15–18 on metatarsi, 11–16 on tarsi. Palpal claw with 5 promarginal teeth. PTC I–II and III–IV with 4–7 and 7–8 teeth on each margin, respectively.
</paragraph>
<paragraph id="8BBE06355505FF95FF52E41CFE28CD15" blockId="47.[189,410,822,848]" box="[189,410,822,848]" pageId="47" pageNumber="48">
<tableNote id="76E707BB5505FF95FF52E41CFE28CD15" box="[189,410,822,848]" pageId="47" pageNumber="48" targetBox="[197,952,861,1065]" targetPageId="47">
<smallCapsWord id="8D5890E95505FF95FF52E41CFF58CD15" baselines="841,841" box="[189,234,822,848]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Leg" pageId="47" pageNumber="48">LEG</smallCapsWord>
<smallCapsWord id="8D5890E95505FF95FF1DE411FE28CD15" baselines="841" box="[242,410,827,846]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="measurements" pageId="47" pageNumber="48">MEASUREMENTS</smallCapsWord>
</tableNote>
</paragraph>
<paragraph id="8BBE06355505FF95FED7E477FC0ACA73" pageId="47" pageNumber="48">
<table id="F901F49555050045FF2AE477FC0ACA72" box="[197,952,861,1065]" gridcols="7" gridrows="6" pageId="47" pageNumber="48">
<tr id="3531047755050045FF2AE477FC0ACD2C" box="[197,952,861,887]" gridrow="0" pageId="47" pageNumber="48" rowspan-0="1">
<th id="76E06D0B55050045FED7E477FE3FCD2C" box="[312,397,861,887]" gridcol="1" gridrow="0" pageId="47" pageNumber="48">
<emphasis id="B975DA275505FF95FED7E477FE3FCD2C" bold="true" box="[312,397,861,887]" pageId="47" pageNumber="48">Femur</emphasis>
</th>
<th id="76E06D0B55050045FE4DE477FE4ACD2C" box="[418,504,861,887]" gridcol="2" gridrow="0" pageId="47" pageNumber="48">
<emphasis id="B975DA275505FF95FE4DE477FE4ACD2C" bold="true" box="[418,504,861,887]" pageId="47" pageNumber="48">Patella</emphasis>
</th>
<th id="76E06D0B55050045FDFFE477FDE0CD2C" box="[528,594,861,887]" gridcol="3" gridrow="0" pageId="47" pageNumber="48">
<emphasis id="B975DA275505FF95FDFFE477FDE0CD2C" bold="true" box="[528,594,861,887]" pageId="47" pageNumber="48">Tibia</emphasis>
</th>
<th id="76E06D0B55050045FD88E477FD47CD2C" box="[615,757,861,887]" gridcol="4" gridrow="0" pageId="47" pageNumber="48">
<emphasis id="B975DA275505FF95FD88E477FD47CD2C" bold="true" box="[615,757,861,887]" pageId="47" pageNumber="48">Metatarsus</emphasis>
</th>
<th id="76E06D0B55050045FCE7E477FCEECD2C" box="[776,860,861,887]" gridcol="5" gridrow="0" pageId="47" pageNumber="48">
<emphasis id="B975DA275505FF95FCE7E477FCEECD2C" bold="true" box="[776,860,861,887]" pageId="47" pageNumber="48">Tarsus</emphasis>
</th>
<th id="76E06D0B55050045FC99E477FC0ACD2C" box="[886,952,861,887]" gridcol="6" gridrow="0" pageId="47" pageNumber="48">
<emphasis id="B975DA275505FF95FC98E477FC05CD2C" bold="true" box="[887,951,861,887]" pageId="47" pageNumber="48">Total</emphasis>
</th>
</tr>
<tr id="3531047755050045FF2AE4AAFC0ACDC0" box="[197,952,896,923]" gridrow="1" pageId="47" pageNumber="48">
<th id="76E06D0B55050045FF2AE4AAFEAACDC0" box="[197,280,896,923]" gridcol="0" gridrow="1" pageId="47" pageNumber="48">Palp</th>
<td id="76E06D0B55050045FED7E4AAFE3FCDC0" box="[312,397,896,923]" gridcol="1" gridrow="1" pageId="47" pageNumber="48">3.94</td>
<td id="76E06D0B55050045FE4DE4AAFE4ACDC0" box="[418,504,896,923]" gridcol="2" gridrow="1" pageId="47" pageNumber="48">1.85</td>
<td id="76E06D0B55050045FDFFE4AAFDE0CDC0" box="[528,594,896,923]" gridcol="3" gridrow="1" pageId="47" pageNumber="48">2.63</td>
<td id="76E06D0B55050045FD88E4AAFD47CDC0" box="[615,757,896,923]" gridcol="4" gridrow="1" pageId="47" pageNumber="48">–</td>
<td id="76E06D0B55050045FCE7E4AAFCEECDC0" box="[776,860,896,923]" gridcol="5" gridrow="1" pageId="47" pageNumber="48">2.34</td>
<td id="76E06D0B55050045FC99E4AAFC0ACDC0" box="[886,952,896,923]" gridcol="6" gridrow="1" pageId="47" pageNumber="48">10.76</td>
</tr>
<tr id="3531047755050045FF2AE48EFC0ACDE4" box="[197,952,932,959]" gridrow="2" pageId="47" pageNumber="48">
<th id="76E06D0B55050045FF2AE48EFEAACDE4" box="[197,280,932,959]" gridcol="0" gridrow="2" pageId="47" pageNumber="48">Leg I</th>
<td id="76E06D0B55050045FED7E48EFE3FCDE4" box="[312,397,932,959]" gridcol="1" gridrow="2" pageId="47" pageNumber="48">5.15</td>
<td id="76E06D0B55050045FE4DE48EFE4ACDE4" box="[418,504,932,959]" gridcol="2" gridrow="2" pageId="47" pageNumber="48">3.13</td>
<td id="76E06D0B55050045FDFFE48EFDE0CDE4" box="[528,594,932,959]" gridcol="3" gridrow="2" pageId="47" pageNumber="48">3.84</td>
<td id="76E06D0B55050045FD88E48EFD47CDE4" box="[615,757,932,959]" gridcol="4" gridrow="2" pageId="47" pageNumber="48">2.94</td>
<td id="76E06D0B55050045FCE7E48EFCEECDE4" box="[776,860,932,959]" gridcol="5" gridrow="2" pageId="47" pageNumber="48">2.03</td>
<td id="76E06D0B55050045FC99E48EFC0ACDE4" box="[886,952,932,959]" gridcol="6" gridrow="2" pageId="47" pageNumber="48">17.09</td>
</tr>
<tr id="3531047755050045FF2AE4EDFC0ACDB9" box="[197,952,967,994]" gridrow="3" pageId="47" pageNumber="48">
<th id="76E06D0B55050045FF2AE4EDFEAACDB9" box="[197,280,967,994]" gridcol="0" gridrow="3" pageId="47" pageNumber="48">Leg II</th>
<td id="76E06D0B55050045FED7E4EDFE3FCDB9" box="[312,397,967,994]" gridcol="1" gridrow="3" pageId="47" pageNumber="48">4.82</td>
<td id="76E06D0B55050045FE4DE4EDFE4ACDB9" box="[418,504,967,994]" gridcol="2" gridrow="3" pageId="47" pageNumber="48">3.02</td>
<td id="76E06D0B55050045FDFFE4EDFDE0CDB9" box="[528,594,967,994]" gridcol="3" gridrow="3" pageId="47" pageNumber="48">3.20</td>
<td id="76E06D0B55050045FD88E4EDFD47CDB9" box="[615,757,967,994]" gridcol="4" gridrow="3" pageId="47" pageNumber="48">3.03</td>
<td id="76E06D0B55050045FCE7E4EDFCEECDB9" box="[776,860,967,994]" gridcol="5" gridrow="3" pageId="47" pageNumber="48">2.12</td>
<td id="76E06D0B55050045FC99E4EDFC0ACDB9" box="[886,952,967,994]" gridcol="6" gridrow="3" pageId="47" pageNumber="48">16.19</td>
</tr>
<tr id="3531047755050045FF2AE4C1FC0ACA5D" box="[197,952,1003,1030]" gridrow="4" pageId="47" pageNumber="48">
<th id="76E06D0B55050045FF2AE4C1FEAACA5D" box="[197,280,1003,1030]" gridcol="0" gridrow="4" pageId="47" pageNumber="48">Leg III</th>
<td id="76E06D0B55050045FED7E4C1FE3FCA5D" box="[312,397,1003,1030]" gridcol="1" gridrow="4" pageId="47" pageNumber="48">3.95</td>
<td id="76E06D0B55050045FE4DE4C1FE4ACA5D" box="[418,504,1003,1030]" gridcol="2" gridrow="4" pageId="47" pageNumber="48">2.48</td>
<td id="76E06D0B55050045FDFFE4C1FDE0CA5D" box="[528,594,1003,1030]" gridcol="3" gridrow="4" pageId="47" pageNumber="48">2.43</td>
<td id="76E06D0B55050045FD88E4C1FD47CA5D" box="[615,757,1003,1030]" gridcol="4" gridrow="4" pageId="47" pageNumber="48">3.15</td>
<td id="76E06D0B55050045FCE7E4C1FCEECA5D" box="[776,860,1003,1030]" gridcol="5" gridrow="4" pageId="47" pageNumber="48">1.84</td>
<td id="76E06D0B55050045FC99E4C1FC0ACA5D" box="[886,952,1003,1030]" gridcol="6" gridrow="4" pageId="47" pageNumber="48">13.85</td>
</tr>
<tr id="3531047755050045FF2AE324FC0ACA72" box="[197,952,1038,1065]" gridrow="5" pageId="47" pageNumber="48">
<th id="76E06D0B55050045FF2AE324FEAACA72" box="[197,280,1038,1065]" gridcol="0" gridrow="5" pageId="47" pageNumber="48">Leg IV</th>
<td id="76E06D0B55050045FED7E324FE3FCA72" box="[312,397,1038,1065]" gridcol="1" gridrow="5" pageId="47" pageNumber="48">5.32</td>
<td id="76E06D0B55050045FE4DE324FE4ACA72" box="[418,504,1038,1065]" gridcol="2" gridrow="5" pageId="47" pageNumber="48">2.87</td>
<td id="76E06D0B55050045FDFFE324FDE0CA72" box="[528,594,1038,1065]" gridcol="3" gridrow="5" pageId="47" pageNumber="48">3.60</td>
<td id="76E06D0B55050045FD88E324FD47CA72" box="[615,757,1038,1065]" gridcol="4" gridrow="5" pageId="47" pageNumber="48">4.81</td>
<td id="76E06D0B55050045FCE7E324FCEECA72" box="[776,860,1038,1065]" gridcol="5" gridrow="5" pageId="47" pageNumber="48">2.37</td>
<td id="76E06D0B55050045FC99E324FC0ACA72" box="[886,952,1038,1065]" gridcol="6" gridrow="5" pageId="47" pageNumber="48">18.97</td>
</tr>
</table>
</paragraph>
<paragraph id="8BBE06355505FF95FF52E364FDA4CAAC" blockId="47.[188,1399,1102,1272]" pageId="47" pageNumber="48">
<smallCapsWord id="8D5890E95505FF95FF52E364FE9ECA3D" baselines="1121,1121" box="[189,300,1102,1128]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Spination" pageId="47" pageNumber="48">SPINATION</smallCapsWord>
. All femora I–II with one basodorsal spine and 3–4 dorsal bristles. Palpal patella, patella I and tarsi I–IV aspinose. Palp: femur pd1; tibia p2, v8(7); tarsus v5(4). Leg I: femur pd1; tibia p1, v5(4); metatarsus v6. Leg II: femur pd3; patella p1; tibia p2, v5; metatarsus v6. Leg III: femur pd3, rd2; patella p2, r1; tibia d2, p2, r3, v7; metatarsus d3, p4, r3, v7. Leg IV: femur pd1, rd1; patella r1; tibia p2, r3, v7; metatarsus d1, p3, r3, v10(8).
</paragraph>
<paragraph id="8BBE06355505FF95FF52E235FDCFCB05" blockId="47.[189,1398,1311,1374]" pageId="47" pageNumber="48">
<smallCapsWord id="8D5890E95505FF95FF52E235FED0CB63" baselines="1330,1331" box="[189,354,1311,1337]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Spermathecae" pageId="47" pageNumber="48">SPERMATHECAE</smallCapsWord>
. Each of paired spermathecae provided with low and wide base carrying two thin and weakly diverging branches (
<figureCitation id="133A1AB05505FF95FDE6E269FDC2CB05" box="[521,624,1347,1374]" captionStart="Figs 522–536" captionStartId="158.[189,241,1728,1754]" captionTargetBox="[189,1398,265,1688]" captionTargetId="figure-119@158.[189,1398,265,1688]" captionTargetPageId="158" captionText="Figs 522–536. Raveniola spp., ♀♀, spermathecae, dorsal views. 522–523. R. pamira sp. nov., paratype and female from Shahob, respectively (both SMNH). 524. R. sororcula sp. nov., holotype (SMNH). 525. R. sororcula, paratype (SMNH). 526–527. R. diluta sp. nov., paratype from Kondara (SMNH). 528–529. R. fedotovi (Charitonov, 1946), from Hojakurgan (SMNH). 530–531. R. pallens sp. nov., paratype (SMNH). 532–533. R. zyuzini sp. nov., paratype (SMNH). 534–536. R. ferghanensis (Zonstein, 1984), from Andijan (SMNH), non-type specimen from environs of Jalal-Abad (SMNH), and paratype from Jalal-Abad (SMNH), respectively. Abbreviations: ld = lateral diverticulum; sb = spermathecal base; st = spermathecal trunk. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990945" httpUri="https://zenodo.org/record/13990945/files/figure.png" pageId="47" pageNumber="48">Fig. 524</figureCitation>
).
</paragraph>
<paragraph id="8BBE06355505FF95FF52E2AFFCD3CBBC" blockId="47.[188,1398,1413,1511]" pageId="47" pageNumber="48">
<smallCapsWord id="8D5890E95505FF95FF52E2AFFE8ECBC5" baselines="1432,1433" box="[189,316,1413,1439]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Spinnerets" pageId="47" pageNumber="48">SPINNERETS</smallCapsWord>
. See
<figureCitation id="133A1AB05505FF95FE94E2AFFDACCBFB" box="[379,542,1413,1440]" captionStart="Figs 584–592" captionStartId="163.[189,241,1821,1847]" captionTargetBox="[189,1398,265,1789]" captionTargetId="figure-92@163.[189,1398,265,1789]" captionText="Figs 584–592. Raveniola spp., ♂♂ (584, 589, 591) and ♀♀ (585–588, 590, 592), holotypes (except 585–586, 590, 592). Spinnerets, ventral (584–585, 587, 589, 591–592) and lateral (586, 588, 590) views. 584. R. pamira sp. nov. (SMNH). 585–586. R. pamira, paratype (SMNH). 587–588. R. sororcula sp. nov. (SMNH). 589. R. diluta sp. nov. (SMNH). 590. R. diluta, paratype (SMNH). 591. R. fedotovi (Charitonov, 1946), paralectotype (PSU). 592. R. fedotovi, ♀ from Hojakurgan (SMNH). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14018408" httpUri="https://zenodo.org/record/14018408/files/figure.png" pageId="47" pageNumber="48">Figs 587–588</figureCitation>
. PMS: length 0.64, diameter 0.22. PLS: maximal diameter 0,74; length of basal, medial and apical segments 1.22, 0.63, 0.30; total length 2.15; apical segment shortly triangular and obliquely truncated from base to apex in lateral view.
</paragraph>
<paragraph id="8BBE06355505FF95FF05E125FE98C872" blockId="47.[189,313,1551,1613]" box="[234,298,1551,1577]" pageId="47" pageNumber="48">
<emphasis id="B975DA275505FF95FF05E125FE98C872" bold="true" box="[234,298,1551,1577]" pageId="47" pageNumber="48">Male</emphasis>
</paragraph>
<paragraph id="8BBE06355505FF95FF52E119FE8BC816" blockId="47.[189,313,1551,1613]" box="[189,313,1587,1613]" pageId="47" pageNumber="48">Unknown.</paragraph>
<paragraph id="8BBE06355505FF95FF52E15FFE81C8D4" blockId="47.[189,307,1653,1679]" box="[189,307,1653,1679]" pageId="47" pageNumber="48">
<heading id="D0F6B1595505FF95FF52E15FFE81C8D4" bold="true" box="[189,307,1653,1679]" fontSize="11" level="3" pageId="47" pageNumber="48" reason="3">
<emphasis id="B975DA275505FF95FF52E15FFE81C8D4" bold="true" box="[189,307,1653,1679]" pageId="47" pageNumber="48">Variation</emphasis>
</heading>
</paragraph>
<paragraph id="8BBE06355505FF95FF52E18EFDEDC8B8" blockId="47.[189,1399,1700,1763]" pageId="47" pageNumber="48">
Carapace length in
<specimenCount id="9D07CDBC5505FF95FE46E18EFD0AC8E5" box="[425,696,1700,1727]" count="3" pageId="47" pageNumber="48" type="female" typeStatus="paratype">three paratype females</specimenCount>
varies from 6.72 to 7.79. Variation in the structure of the spermathecae as shown in
<figureCitation id="133A1AB05505FF95FE19E1E2FDEEC8B8" box="[502,604,1736,1763]" captionStart="Figs 522–536" captionStartId="158.[189,241,1728,1754]" captionTargetBox="[189,1398,265,1688]" captionTargetId="figure-119@158.[189,1398,265,1688]" captionTargetPageId="158" captionText="Figs 522–536. Raveniola spp., ♀♀, spermathecae, dorsal views. 522–523. R. pamira sp. nov., paratype and female from Shahob, respectively (both SMNH). 524. R. sororcula sp. nov., holotype (SMNH). 525. R. sororcula, paratype (SMNH). 526–527. R. diluta sp. nov., paratype from Kondara (SMNH). 528–529. R. fedotovi (Charitonov, 1946), from Hojakurgan (SMNH). 530–531. R. pallens sp. nov., paratype (SMNH). 532–533. R. zyuzini sp. nov., paratype (SMNH). 534–536. R. ferghanensis (Zonstein, 1984), from Andijan (SMNH), non-type specimen from environs of Jalal-Abad (SMNH), and paratype from Jalal-Abad (SMNH), respectively. Abbreviations: ld = lateral diverticulum; sb = spermathecal base; st = spermathecal trunk. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990945" httpUri="https://zenodo.org/record/13990945/files/figure.png" pageId="47" pageNumber="48">Fig. 525</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31B55BE5505FF95FF52E020FB3EC92C" pageId="47" pageNumber="48" type="biology_ecology">
<paragraph id="8BBE06355505FF95FF52E020FE92C97F" blockId="47.[189,288,1802,1828]" box="[189,288,1802,1828]" pageId="47" pageNumber="48">
<heading id="D0F6B1595505FF95FF52E020FE92C97F" bold="true" box="[189,288,1802,1828]" fontSize="11" level="3" pageId="47" pageNumber="48" reason="3">
<emphasis id="B975DA275505FF95FF52E020FE92C97F" bold="true" box="[189,288,1802,1828]" pageId="47" pageNumber="48">Ecology</emphasis>
</heading>
</paragraph>
<paragraph id="8BBE06355505FF95FF52E013FB3EC92C" blockId="47.[189,1398,1849,1912]" pageId="47" pageNumber="48">
The spiders were found under stones along the stream in the riverside gallery woodland dominated by
<taxonomicName id="4C017DB65505FF95FF51E077FEEFC92C" box="[190,349,1885,1911]" class="Magnoliopsida" family="Juglandaceae" genus="Juglans" kingdom="Plantae" order="Fagales" pageId="47" pageNumber="48" phylum="Tracheophyta" rank="species" species="regia">
<emphasis id="B975DA275505FF95FF51E077FEEFC92C" box="[190,349,1885,1911]" italics="true" pageId="47" pageNumber="48">Juglans regia</emphasis>
</taxonomicName>
. Their retreats and females with cocoons are shown in
<figureCitation id="133A1AB05505FF95FC0CE077FB3AC92C" box="[995,1160,1885,1911]" captionStart="Figs 691–698" captionStartId="176.[189,241,1818,1844]" captionTargetBox="[189,1398,265,1783]" captionTargetId="figure-27@176.[189,1398,265,1784]" captionTargetPageId="176" captionText="Figs 691–698. Retreats of Raveniola sororcula sp. nov. and live females protecting their egg sacs. Tajikistan, Darvaz Mts, Obiviskharvi Canyon, surroundings of Hurk Village (1700 m a.s.l.)." figureDoi="http://doi.org/10.5281/zenodo.13990969" httpUri="https://zenodo.org/record/13990969/files/figure.png" pageId="47" pageNumber="48">Figs 691–698</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31B55BE5505FF95FF52E0B5FD49C9B2" box="[189,763,1951,2025]" pageId="47" pageNumber="48" type="distribution">
<paragraph id="8BBE06355505FF95FF52E0B5FEE4C9E2" blockId="47.[189,342,1951,1977]" box="[189,342,1951,1977]" pageId="47" pageNumber="48">
<heading id="D0F6B1595505FF95FF52E0B5FEE4C9E2" bold="true" box="[189,342,1951,1977]" fontSize="11" level="3" pageId="47" pageNumber="48" reason="3">
<emphasis id="B975DA275505FF95FF52E0B5FEE4C9E2" bold="true" box="[189,342,1951,1977]" pageId="47" pageNumber="48">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BBE06355505FF95FF52E0E5FD49C9B2" blockId="47.[189,763,1998,2025]" box="[189,763,1998,2025]" pageId="47" pageNumber="48">
Known only from the
<typeStatus id="54BAB8975505FF95FE2DE0E5FE47C9B2" box="[450,501,1999,2025]" pageId="47" pageNumber="48">type</typeStatus>
locality. See
<figureCitation id="133A1AB05505FF95FD7DE0E4FD46C9B2" box="[658,756,1998,2025]" captionStart="Figs 751–760" captionStartId="184.[189,241,1662,1688]" captionTargetBox="[258,1323,265,1629]" captionTargetId="figure-247@184.[258,1323,265,1629]" captionTargetPageId="184" captionText="Figs 751–760. Distribution records of Raveniola spp. 751. R. alajensis sp. nov. (), R. cucullata sp. nov. () and R. dolosa sp. nov. (). 752. R. hirta sp. nov. (), R. ignobilis sp. nov. () and R. insolita sp. nov. (). 753. R. karategensis sp. nov. (), R. ornata sp. nov. () and R. ornatula sp. nov. (). 754. R. pamira sp. nov. (), R. sororcula sp. nov. () and R. diluta sp. nov. (). 755. R. fedotovi (Charitonov, 1946) (), R. pallens sp. nov. () and R. zyuzini sp. nov. (). 756. R. ferghanensis (Zonstein, 1984) () and R. kirgizica sp. nov. (). 757. R. kopetdaghensis (Fet, 1984) (). 758. R. mikhailovi Zonstein, 2021 () and R. nenilini sp. nov. (). 759. R. ovchinnikovi sp. nov. () and R. tarabaevi sp. nov. (). 760. R. virgata (Simon, 1891) (;? = the questionable type locality) and R. vulpina sp. nov. (). Country abbreviations as in Figs 747–750." figureDoi="http://doi.org/10.5281/zenodo.14018423" httpUri="https://zenodo.org/record/14018423/files/figure.png" pageId="47" pageNumber="48">Fig. 754</figureCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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