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<mods:title id="6DA71C138BF3EE250A5136079C0632C5">Taxonomy of the freshwater crabs of Costa Rica, with a revision of the genus Ptychophallus Smalley, 1964 (Crustacea: Decapoda: Pseudothelphusidae)</mods:title>
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<treatment id="710D87BB2934162CD6BDF9C3FC89FA4E" ID-DOI="http://doi.org/10.5281/zenodo.5491544" ID-GBIF-Taxon="119590147" ID-Zenodo-Dep="5491544" LSID="urn:lsid:plazi:treatment:710D87BB2934162CD6BDF9C3FC89FA4E" httpUri="http://treatment.plazi.org/id/710D87BB2934162CD6BDF9C3FC89FA4E" lastPageId="36" lastPageNumber="337" pageId="34" pageNumber="335">
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<heading id="A25381C12934162AD6BDF9C3FD63F9AA" bold="true" box="[151,690,1632,1658]" fontSize="11" level="1" pageId="34" pageNumber="335" reason="1">
<taxonomicName id="3EA44D2E2934162AD6BDF9C3FD63F9AA" ID-CoL="4M3G8" authority="Rathbun, 1893" authorityName="Rathbun" authorityYear="1893" box="[151,690,1632,1658]" class="Malacostraca" family="Pseudothelphusidae" genus="Potamocarcinus" kingdom="Animalia" order="Decapoda" pageId="34" pageNumber="335" phylum="Arthropoda" rank="species" species="richmondi">
<emphasis id="CBD0EABF2934162AD6BDF9C3FD63F9AA" bold="true" box="[151,690,1632,1658]" pageId="34" pageNumber="335">
<emphasis id="CBD0EABF2934162AD6BDF9C3FE30F9AA" bold="true" box="[151,481,1632,1658]" italics="true" pageId="34" pageNumber="335">Potamocarcinus richmondi</emphasis>
(
<bibRefCitation id="9D354B5C2934162AD7C5F9C3FD7BF9AA" author="Rathbun" box="[495,682,1632,1658]" pageId="34" pageNumber="335" refString="Rathbun, M. J. (1893) Descriptions of new species of American fresh-water crabs. Proceedings of the United States National Museum, 16 (959), 649 - 661, pls. LXXIII - LXXVII. http: // dx. doi. org / 10.5479 / si. 00963801.16 - 959.649" type="journal article" year="1893">Rathbun, 1893</bibRefCitation>
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<paragraph id="F91B36AD2934162AD6BDF921FE86F94B" blockId="34.[151,690,1632,1691]" box="[151,343,1666,1691]" pageId="34" pageNumber="335">
(
<figureCitation id="619F2A282934162AD6B5F921FEF7F94B" box="[159,294,1666,1691]" captionStart="FIGURES 71 76" captionStartId="30.[151,264,1836,1859]" captionTargetBox="[169,1417,343,1804]" captionTargetId="figure@30.[165,1422,337,1815]" captionTargetPageId="30" captionText="FIGURES 71 76. Potamocarcinus magnus (Rathbun, 1896): 71, left first gonopod, holotype, USNM 19049, caudal-mesial view; 72, same data, latero-cephalic view of distal part. Potamocarcinus nicaraguensis Rathbun, 1893: 73, left first gonopod, INPA 1869, caudal-mesial view; 74, same data, lateral view of distal part. Potamocarcinus richmondi (Rathbun, 1893): 75, left first gonopod, TMNH Acc. Nr. 94 - 100, caudal-mesial view; 76, same data, lateral view of distal part (ce, cephalic process; cp, caudal process; cst, caudal setae; fas, field of apical spines; ls, lateral suture; me, mesial process; mp, marginal process; ms, marginal suture). Scale bars: 71, 75 = 1 mm; 72 74, 76 = 2 mm." httpUri="https://zenodo.org/record/235134/files/figure.png" pageId="34" pageNumber="335">Figs. 75, 76</figureCitation>
,
<figureCitation id="619F2A282934162AD718F921FE9EF94B" box="[306,335,1666,1691]" captionStart="FIGURE 86" captionStartId="41.[151,250,1130,1152]" captionTargetBox="[207,1378,199,1098]" captionTargetId="figure@41.[204,1378,193,1100]" captionTargetPageId="41" captionText="FIGURE 86. Distribution map of Potamocarcinus magnus (Rathbun, 1896), P. nicaraguensis Rathbun, 1893, and P. richmondi (Rathbun, 1893)." httpUri="https://zenodo.org/record/235144/files/figure.png" pageId="34" pageNumber="335">86</figureCitation>
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<emphasis id="CBD0EABF2934162AD6BDF968FE6CF931" box="[151,445,1739,1761]" italics="true" pageId="34" pageNumber="335">Pseudothelphusa richmondi</emphasis>
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: 654
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, pl. 75, figs. 610.—
<treatmentCitation id="780510BC2934162AD587F968FBA7F931" author="Young" box="[941,1142,1739,1761]" page="218" pageId="34" pageNumber="335" year="1900">
<bibRefCitation id="9D354B5C2934162AD587F968FBEAF931" author="Young" box="[941,1083,1739,1761]" pageId="34" pageNumber="335" refString="Young, C. G. (1900) The stalk-eyed Crustacea of British Guiana, West Indies, and Bermuda. J. M. Watkins, London, 514 pp., 4 pls." type="book" year="1900">Young 1900</bibRefCitation>
: 218
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.—
<treatmentCitation id="780510BC2934162AD2B9F968FABFF931" author="Rathbun" box="[1171,1390,1739,1761]" page="242" pageId="34" pageNumber="335" year="1904">
<bibRefCitation id="9D354B5C2934162AD2B9F968FAE5F931" author="Rathbun" box="[1171,1332,1739,1761]" pageId="34" pageNumber="335" refString="Rathbun, M. J. (1904) Les crabes d'eau douce (Potamonidae). Nouvelles Archives du Museum d'Histoire naturelle Paris, 6 (4), 225 - 312, text-figs. 1 - 37, volume pls. 9 - 18 (pls. I - X for Rathbun's article)." type="journal article" year="1904">Rathbun 1904</bibRefCitation>
: 242
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(in list).—
<treatmentCitation id="780510BC2934162AD721F94AFE34F92F" author="Rathbun" box="[267,485,1769,1791]" page="303" pageId="34" pageNumber="335" year="1905">
<bibRefCitation id="9D354B5C2934162AD721F94AFE7DF92F" author="Rathbun" box="[267,428,1769,1791]" pageId="34" pageNumber="335" refString="Rathbun, M. J. (1905) Les crabes d'eau douce. Nouvelles Archives du Museum d'Histoire naturelle, Paris, 7 (4), 159 - 321, textfigs. 38 - 105, volume pls. 13 - 22. [pls. XI - X for Rathbun's article]" type="journal article" year="1905">Rathbun 1905</bibRefCitation>
: 303
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.—
<treatmentCitation id="780510BC2934162AD428F94AFD3CF92F" author="Coifmann" box="[514,749,1769,1791]" page="109" pageId="34" pageNumber="335" year="1939">
<bibRefCitation id="9D354B5C2934162AD428F94AFD63F92F" author="Coifmann" box="[514,690,1769,1791]" pageId="34" pageNumber="335" refString="Coifmann, I. (1939) Potamonidi della Guiana inglese raccolti dal Prof. Nello Beccari. Archivio Zoologico Italiano, 27, 93 - 116, pl. 3." type="journal article" year="1939">Coifmann 1939</bibRefCitation>
: 109
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(in list).—
<treatmentCitation id="780510BC2934162AD55AF94AFBB1F92F" author="Rodriguez" box="[880,1120,1769,1791]" page="436" pageId="34" pageNumber="335" year="2001">
<bibRefCitation id="9D354B5C2934162AD55AF94AFBF5F92F" author="Rodriguez" box="[880,1060,1769,1791]" pageId="34" pageNumber="335" refString="Rodriguez, G. (2001) New species and records of pseudothelphusid crabs (Crustacea: Brachyura) from Central America in the Museum of Natural History of Tulane University. Proceedings of the Biological Society of Washington, 114 (2), 435 - 443." type="journal article" year="2001">Rodríguez 2001</bibRefCitation>
: 436
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.—
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<bibRefCitation id="9D354B5C2934162AD254F94AFAE2F92F" author="Ng" box="[1150,1331,1769,1791]" pageId="34" pageNumber="335" refString="Ng, P. K. L., Guinot, D. &amp; Davie, P. J. F. (2008) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. The Raffles Bulletin of Zoology Supplement, 17, 1 - 286." type="journal article" year="2008">
Ng
<emphasis id="CBD0EABF2934162AD284F949FB3CF92F" box="[1198,1261,1769,1791]" italics="true" pageId="34" pageNumber="335">et al.</emphasis>
2008
</bibRefCitation>
: 176
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(in list).—
<treatmentCitation id="780510BC2934162AD721F8ABFD4DF8CE" author="Villalobos" box="[267,668,1800,1822]" page="297" pageId="34" pageNumber="335" year="2008">
<bibRefCitation id="9D354B5C2934162AD721F8ABFDBDF8CE" author="Villalobos" box="[267,620,1800,1822]" pageId="34" pageNumber="335" refString="Villalobos Hiriart, J. L. &amp; Alvarez, F. (2008) Los cangrejos de la familia Pseudothelphusidae (Decapoda: Brachyura: Eubrachyura) de Mexico, con un apendice de las especies citadas para America hasta el 2006. In: Alvarez, F. &amp; Rodriguez- Almaraz, G. (Eds.), Crustaceos de Mexico: Estado actual de su Conocimiento. Universidad Autonoma de Nuevo Leon, Monterrey, Nuevo Leon, pp. 239 - 300. [Mexico]" type="book chapter" year="2008">Villalobos Hiriart &amp; Álvarez 2008</bibRefCitation>
: 297
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(in list).
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</paragraph>
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<taxonomicName id="3EA44D2E2934162AD6BDF885FDAFF8EC" ID-CoL="6WHWN" box="[151,638,1830,1852]" class="Malacostraca" family="Pseudothelphusidae" genus="Pseudothelphusa" kingdom="Animalia" order="Decapoda" pageId="34" pageNumber="335" phylum="Arthropoda" rank="species" species="richmondi" subGenus="Megathelphusa">
<emphasis id="CBD0EABF2934162AD6BDF885FE95F8EC" box="[151,324,1830,1852]" italics="true" pageId="34" pageNumber="335">Pseudothelphusa</emphasis>
(
<emphasis id="CBD0EABF2934162AD775F885FE2AF8EC" box="[351,507,1830,1852]" italics="true" pageId="34" pageNumber="335">Megathelphusa</emphasis>
)
<emphasis id="CBD0EABF2934162AD43CF885FDAFF8EC" box="[534,638,1830,1852]" italics="true" pageId="34" pageNumber="335">richmondi</emphasis>
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<bibRefCitation id="9D354B5C2934162AD4B2F885FC91F8EC" author="Smalley" box="[664,832,1830,1852]" pageId="34" pageNumber="335" refString="Smalley, A. E. (1964 b) The river crabs of Costa Rica, and the subfamilies of the Pseudothelphusidae. Tulane Studies in Zoology, 12 (1), 5 - 13." type="journal article" year="1964" yearSuffix="b">Smalley 1964b</bibRefCitation>
: 9
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, figs. 46.—
<treatmentCitation id="780510BC2934162AD5D7F885FB36F8EC" author="Pretzmann" box="[1021,1255,1830,1852]" page="70" pageId="34" pageNumber="335" year="1972">
<bibRefCitation id="9D354B5C2934162AD5D7F885FB64F8EC" author="Pretzmann" box="[1021,1205,1830,1852]" pageId="34" pageNumber="335" refString="Pretzmann, G. (1972) Die Pseudothelphusidae (Crustacea Brachyura). Zoologica, 42 (120), pt. 1, 1 - 182." type="journal article" year="1972">Pretzmann 1972</bibRefCitation>
: 70
</treatmentCitation>
, figs. 373376, 366368.—
<treatmentCitation id="780510BC2934162AD716F8E6FDCAF88B" author="Pretzmann" box="[316,539,1861,1883]" page="653" pageId="34" pageNumber="335" year="1980">
<bibRefCitation id="9D354B5C2934162AD716F8E6FE39F88B" author="Pretzmann" box="[316,488,1861,1883]" pageId="34" pageNumber="335" refString="Pretzmann, G. (1980) Von Dr. Ivo Poglayen-Neuwall 1975 in Mittelamerika gesammelte Krabben. Annalen des Naturhistorischen Museums, 83, 651 - 666, pls. 1 - 18. [Wien]" type="journal article" year="1980">Pretzmann 1980</bibRefCitation>
: 653
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.
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<emphasis id="CBD0EABF2934162AD6BDF8C6FEEDF8AA" box="[151,316,1893,1914]" italics="true" pageId="34" pageNumber="335">Potamocarcinus</emphasis>
(
<emphasis id="CBD0EABF2934162AD761F8C6FE21F8AA" box="[331,496,1893,1914]" italics="true" pageId="34" pageNumber="335">Potamocarcinus</emphasis>
)
<emphasis id="CBD0EABF2934162AD7D4F8C7FDB7F8AA" box="[510,614,1892,1914]" italics="true" pageId="34" pageNumber="335">richmondi</emphasis>
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: 2
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.—
<treatmentCitation id="780510BC2934162AD571F8C7FBFCF8AA" author="Pretzmann" box="[859,1069,1892,1914]" page="20" pageId="34" pageNumber="335" year="1971">
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: 20
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<subSubSection id="B1BE65262934162BD6BDF805FF10FF05" lastPageId="35" lastPageNumber="336" pageId="34" pageNumber="335" type="materials_examined">
<paragraph id="F91B36AD2934162BD6BDF805FF10FF05" blockId="34.[151,1436,1958,2019]" lastBlockId="35.[151,1437,151,752]" lastPageId="35" lastPageNumber="336" pageId="34" pageNumber="335">
<emphasis id="CBD0EABF2934162AD6BDF805FEA8F86F" bold="true" box="[151,377,1958,1983]" pageId="34" pageNumber="335">Material examined</emphasis>
.
<collectingCountry id="81B3763D2934162AD7ACF804FDFFF86F" box="[390,558,1959,1983]" name="Costa Rica" pageId="34" pageNumber="335">COSTA RICA</collectingCountry>
. Alajuela Province, Pacific drainage:
<specimenCount id="EFA2FD242934162AD5FBF804FBF6F86F" box="[977,1063,1958,1983]" pageId="34" pageNumber="335" type="male">3 males</specimenCount>
immat.,
<specimenCount id="EFA2FD242934162AD2A0F80BFB27F86F" box="[1162,1270,1958,1983]" pageId="34" pageNumber="335" type="female">7 females</specimenCount>
immat., UCR- MZ 2499, Canton de Orotina,
<date id="8D1A106D2934162AD7DEF869FDBFF833" box="[500,622,1994,2019]" pageId="34" pageNumber="335" value="1966-09-29">29.ix.1966</date>
, D.C. Robinson leg.—Cartago Province, Atlantic drainage:
<specimenCount id="EFA2FD242934162AD336F868FAB8F833" box="[1308,1385,1994,2019]" pageId="34" pageNumber="335" type="male">1 male</specimenCount>
(cw 78.4, cl 51.8),
<collectionCode id="9FB5AE682935162BD710FF3BFE5CFF7F" box="[314,397,152,175]" httpUri="http://grbio.org/cool/1h73-pg9e" name="Tianjin Museum of Natural History" pageId="35" pageNumber="336">TMNH</collectionCode>
(Acc. nr 98-101),
<quantity id="3E5C9B482935162BD474FF34FD65FF60" box="[606,692,151,176]" metricMagnitude="4" metricUnit="m" metricValue="2.62323072" pageId="35" pageNumber="336" unit="mi" value="16.3">16.3 mi</quantity>
east of Turrialba,
<date id="8D1A106D2935162BD5ABFF34FC2DFF60" box="[897,1020,151,176]" pageId="35" pageNumber="336" value="1973-06-28">28.vi.1973</date>
, R.D. Suttkus &amp; S. Jiménez-Canossa leg.
</paragraph>
</subSubSection>
<subSubSection id="B1BE65262935162BD6EDFF7CFD38FD78" pageId="35" pageNumber="336" type="description">
<paragraph id="F91B36AD2935162BD6EDFF7CFD38FD78" blockId="35.[151,1437,151,752]" pageId="35" pageNumber="336">
<emphasis id="CBD0EABF2935162BD6EDFF7CFE3BFF28" bold="true" box="[199,490,223,248]" pageId="35" pageNumber="336">Description of gonopod</emphasis>
. Straight in both caudal, mesial views, moderately compressed mesio-laterally; caudal border concave, narrower on its distal third, smooth; mesio-caudal surface smooth, patch of caudal setae absent along its proximal half; mesial border nearly straight in mesial view; mesio-cephalic surface with subdistal patch of short setae increasing size distally. Spermatic channel opening on apex cavity, situated towards lateral end. Marginal suture on mesial side, straight along stem, short row of marginal setae proximally. Marginal process (caudal process,
<emphasis id="CBD0EABF2935162BD77EFE36FE43FE7C" box="[340,402,405,428]" italics="true" pageId="35" pageNumber="336">sensu</emphasis>
<bibRefCitation id="9D354B5C2935162BD7B1FE37FD95FE7D" author="Smalley" box="[411,580,404,429]" pageId="35" pageNumber="336" refString="Smalley, A. (1964 a) A terminology for the gonopods of the American river crabs. Systematic Zoology, 13, 28 - 31." type="journal article" year="1964" yearSuffix="a">Smalley 1964a</bibRefCitation>
) broad, rounded, much shorter than mesial lobe. Lateral suture incomplete, marked by conspicuously weak sulcus on the first two-thirds of caudal surface. Mesial process as short, broadly triangular apical tooth directed cephalad on distal portion of mesial border. Apex compressed in mesio-lateral direction, slightly distorted in lateral direction; mesial lobe enlarged, rounded in mesio-caudal view, slightly folded over the apex cavity. Apex cavity narrow, slit-like, lateral border crested; field of apical spines weakly developed, with small patch of few minute spines. Cephalic process as two well separated teeth on cephalic surface, apical tooth smaller, blunt, subapical tooth larger, rather sharp. Field of apical spines weakly developed, very narrow path of few minute spines along caudo-cephalic direction.
</paragraph>
</subSubSection>
<subSubSection id="B1BE65262935162BD6EDFD17FE3CFD20" pageId="35" pageNumber="336" type="materials_examined">
<paragraph id="F91B36AD2935162BD6EDFD17FE3CFD20" blockId="35.[151,1437,151,752]" pageId="35" pageNumber="336">
<emphasis id="CBD0EABF2935162BD6EDFD17FDDAFD1D" bold="true" box="[199,523,692,717]" pageId="35" pageNumber="336">
<typeStatus id="261F880F2935162BD6EDFD17FEE5FD1D" box="[199,308,692,717]" pageId="35" pageNumber="336" type="holotype">Holotype</typeStatus>
and
<typeStatus id="261F880F2935162BD752FD17FE7AFD1D" box="[376,427,692,717]" pageId="35" pageNumber="336">type</typeStatus>
locality
</emphasis>
. Male, cw 49.8, cl 32.5.5,
<collectionCode id="9FB5AE682935162BD57BFD17FC73FD1C" LSID="urn:lsid:biocol.org:col:34871" box="[849,930,692,716]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="35" pageNumber="336">USNM</collectionCode>
17725.
<collectingCountry id="81B3763D2935162BD22CFD17FBADFD1D" box="[1030,1148,692,717]" name="Nicaragua" pageId="35" pageNumber="336">Nicaragua</collectingCountry>
, Atlántico Sur Province, Bluefields (Atlantic drainage).
</paragraph>
</subSubSection>
<caption id="ADDB66252935162BD6BDF88DFA97F894" box="[151,1350,1838,1860]" httpUri="https://zenodo.org/record/235139/files/figure.png" pageId="35" pageNumber="336" targetBox="[154,1431,813,1813]" targetPageId="35">
<paragraph id="F91B36AD2935162BD6BDF88DFA97F894" blockId="35.[151,1350,1838,1860]" box="[151,1350,1838,1860]" pageId="35" pageNumber="336">
<emphasis id="CBD0EABF2935162BD6BDF88DFEC8F893" bold="true" box="[151,281,1838,1860]" pageId="35" pageNumber="336">FIGURE 81</emphasis>
. Distribution map of
<taxonomicName id="3EA44D2E2935162BD7DEF88DFC7AF894" authority="Pretzmann, 1965" authorityName="Pretzmann" authorityYear="1965" box="[500,939,1838,1860]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="35" pageNumber="336" phylum="Arthropoda" rank="species" species="goldmanni">
<emphasis id="CBD0EABF2935162BD7DEF88DFD29F894" box="[500,760,1838,1860]" italics="true" pageId="35" pageNumber="336">Ptychophallus goldmanni</emphasis>
Pretzmann, 1965
</taxonomicName>
, and
<taxonomicName id="3EA44D2E2935162BD5CAF88CFA93F894" authority="Campos &amp; Lemaitre, 1999" authorityName="Campos &amp; Lemaitre" authorityYear="1999" box="[992,1346,1838,1860]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="35" pageNumber="336" phylum="Arthropoda" rank="species" species="kuna">
<emphasis id="CBD0EABF2935162BD5CAF88CFBFBF894" box="[992,1066,1838,1860]" italics="true" pageId="35" pageNumber="336">P. kuna</emphasis>
Campos &amp; Lemaitre, 1999
</taxonomicName>
.
</paragraph>
</caption>
<caption id="ADDB66252932162CD6BDFB1BFC48FB1E" box="[151,921,1208,1230]" httpUri="https://zenodo.org/record/235140/files/figure.png" pageId="36" pageNumber="337" targetBox="[158,1431,200,1182]" targetPageId="36">
<paragraph id="F91B36AD2932162CD6BDFB1BFC48FB1E" blockId="36.[151,921,1208,1230]" box="[151,921,1208,1230]" pageId="36" pageNumber="337">
<emphasis id="CBD0EABF2932162CD6BDFB1BFEC8FB1D" bold="true" box="[151,281,1208,1230]" pageId="36" pageNumber="337">FIGURE 82</emphasis>
. Distribution map of
<taxonomicName id="3EA44D2E2932162CD7DEFB1BFC44FB1E" authority="(Bott, 1968)" authorityName="Bott" authorityYear="1968" baseAuthorityName="Bott" baseAuthorityYear="1968" box="[500,917,1208,1230]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="36" pageNumber="337" phylum="Arthropoda" rank="species" species="paraxanthusi">
<emphasis id="CBD0EABF2932162CD7DEFB1BFCC0FB1E" box="[500,785,1208,1230]" italics="true" pageId="36" pageNumber="337">Ptychophallus paraxanthusi</emphasis>
(Bott, 1968)
</taxonomicName>
.
</paragraph>
</caption>
<subSubSection id="B1BE65262932162CD6EDFB56FC89FA4E" pageId="36" pageNumber="337" type="distribution">
<paragraph id="F91B36AD2932162CD6EDFB56FC89FA4E" blockId="36.[151,1436,1269,1438]" pageId="36" pageNumber="337">
<emphasis id="CBD0EABF2932162CD6EDFB56FE88FADE" bold="true" box="[199,345,1269,1294]" pageId="36" pageNumber="337">Distribution</emphasis>
. Central
<collectingCountry id="81B3763D2932162CD7EAFB55FDECFADE" box="[448,573,1269,1294]" name="Costa Rica" pageId="36" pageNumber="337">Costa Rica</collectingCountry>
, in both Atlantic and Pacific drainages (
<figureCitation id="619F2A282932162CD5D4FB56FB83FADE" box="[1022,1106,1269,1294]" captionStart="FIGURE 86" captionStartId="41.[151,250,1130,1152]" captionTargetBox="[207,1378,199,1098]" captionTargetId="figure@41.[204,1378,193,1100]" captionTargetPageId="41" captionText="FIGURE 86. Distribution map of Potamocarcinus magnus (Rathbun, 1896), P. nicaraguensis Rathbun, 1893, and P. richmondi (Rathbun, 1893)." httpUri="https://zenodo.org/record/235144/files/figure.png" pageId="36" pageNumber="337">Fig. 86</figureCitation>
). The species is also found in
<collectingCountry id="81B3763D2932162CD6BDFABAFEDBFAE2" box="[151,266,1305,1330]" name="Nicaragua" pageId="36" pageNumber="337">Nicaragua</collectingCountry>
(
<bibRefCitation id="9D354B5C2932162CD732FABAFE6EFAE2" author="Rathbun" box="[280,447,1305,1330]" pageId="36" pageNumber="337" refString="Rathbun, M. J. (1893) Descriptions of new species of American fresh-water crabs. Proceedings of the United States National Museum, 16 (959), 649 - 661, pls. LXXIII - LXXVII. http: // dx. doi. org / 10.5479 / si. 00963801.16 - 959.649" type="journal article" year="1893">Rathbun, 1893</bibRefCitation>
) and
<collectingCountry id="81B3763D2932162CD7D7FAB9FD87FAE1" box="[509,598,1306,1329]" name="Panama" pageId="36" pageNumber="337">Panama</collectingCountry>
(
<bibRefCitation id="9D354B5C2932162CD44EFABAFCF3FAE2" author="Rodriguez" box="[612,802,1305,1330]" pageId="36" pageNumber="337" refString="Rodriguez, G. (1982) Les crabes d'eau douce d'Amerique. Familie des Pseudothelphusidae. ORSTOM, Paris, 223 pp. [Faune Tropicale 22]" type="book" year="1982">Rodriguez, 1982</bibRefCitation>
;
<bibRefCitation id="9D354B5C2932162CD504FAB9FC3AFAE2" author="Pretzmann" box="[814,1003,1306,1330]" pageId="36" pageNumber="337" refString="Pretzmann, G. (1972) Die Pseudothelphusidae (Crustacea Brachyura). Zoologica, 42 (120), pt. 1, 1 - 182." type="journal article" year="1972">Pretzmann, 1972</bibRefCitation>
[as
<taxonomicName id="3EA44D2E2932162CD232FAB9FB32FAE2" box="[1048,1251,1305,1330]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="36" pageNumber="337" phylum="Arthropoda" rank="species" species="masimbari" subGenus="Microptychophallus">
<emphasis id="CBD0EABF2932162CD232FAB9FBF7FAE1" box="[1048,1062,1306,1329]" italics="true" pageId="36" pageNumber="337">P</emphasis>
. (
<emphasis id="CBD0EABF2932162CD217FAB9FB85FAE1" box="[1085,1108,1306,1329]" italics="true" pageId="36" pageNumber="337">M</emphasis>
.)
<emphasis id="CBD0EABF2932162CD241FABAFB32FAE2" box="[1131,1251,1305,1330]" italics="true" pageId="36" pageNumber="337">masimbari</emphasis>
</taxonomicName>
]). Other records from
<collectingCountry id="81B3763D2932162CD6FEFA9DFE9CFA86" box="[212,333,1341,1366]" name="Costa Rica" pageId="36" pageNumber="337">Costa Rica</collectingCountry>
were given by
<bibRefCitation id="9D354B5C2932162CD7D2FA9EFD62FA86" author="Smalley" box="[504,691,1341,1366]" pageId="36" pageNumber="337" refString="Smalley, A. E. (1964 b) The river crabs of Costa Rica, and the subfamilies of the Pseudothelphusidae. Tulane Studies in Zoology, 12 (1), 5 - 13." type="journal article" year="1964" yearSuffix="b">Smalley (1964b)</bibRefCitation>
(Cartago Province,
<quantity id="3E5C9B482932162CD5B9FA9DFC0BFA86" box="[915,986,1341,1366]" metricMagnitude="3" metricUnit="m" metricValue="1.7702784" pageId="36" pageNumber="337" unit="mi" value="1.1">1.1 mi</quantity>
N Turrialba),
<bibRefCitation id="9D354B5C2932162CD253FA9DFA90FA86" author="Pretzmann" box="[1145,1345,1342,1366]" pageId="36" pageNumber="337" refString="Pretzmann, G. (1972) Die Pseudothelphusidae (Crustacea Brachyura). Zoologica, 42 (120), pt. 1, 1 - 182." type="journal article" year="1972">Pretzmann (1972)</bibRefCitation>
(Limón Province, small creek near a farm), and
<bibRefCitation id="9D354B5C2932162CD44BFAC1FCFDFAAA" author="Pretzmann" box="[609,812,1378,1402]" pageId="36" pageNumber="337" refString="Pretzmann, G. (1980) Von Dr. Ivo Poglayen-Neuwall 1975 in Mittelamerika gesammelte Krabben. Annalen des Naturhistorischen Museums, 83, 651 - 666, pls. 1 - 18. [Wien]" type="journal article" year="1980">Pretzmann (1980)</bibRefCitation>
(Guanacaste Province: Piedras River, W Bagaces, and Alajuela Province: tributary of Arenal River,
<quantity id="3E5C9B482932162CD4B9FA26FD1DFA4E" box="[659,716,1413,1438]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="36" pageNumber="337" unit="km" value="5.0">5 km</quantity>
SE Arenal).
</paragraph>
</subSubSection>
</treatment>
</document>