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<document id="B32AA3D293A7BED19E592DDF8105478D" ID-CLB-Dataset="46478" ID-DOI="10.11646/zootaxa.3764.3.6" ID-GBIF-Dataset="b7c3456f-0588-4d10-b263-062ca84f3fbb" ID-ISSN="1175-5326" ID-Zenodo-Dep="250851" ID-ZooBank="F96FB36D-49DA-4A1C-BBA6-B049531CAD8C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460382269943" checkinUser="plazi" docAuthor="Tan, Ming Kai &amp; Robillard, Tony" docDate="2014" docId="563E879E086AFFEC17F6F8EE0A99F9BF" docLanguage="en" docName="zt03764p376.pdf" docOrigin="Zootaxa 3764 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cardiodactylus admirabilis Tan &amp; Robillard, n. sp." docType="treatment" docVersion="13" lastPageNumber="371" masterDocId="AA07FFE6086BFFEB1761FF81083BFFD8" masterDocTitle="A new species of Cardiodactylus (Orthoptera: Grylloidea: Eneopterinae) from Singapore" masterLastPageNumber="376" masterPageNumber="364" pageNumber="365" updateTime="1698296948229" updateUser="ExternalLinkService">
<mods:mods id="6B4B5737A7442B6E49418590927BB410" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="028BB82BA95B16E1B9E6E33568C433F6">
<mods:title id="D99B468EB97DEBE2749BEDFCE7BE8F8D">A new species of Cardiodactylus (Orthoptera: Grylloidea: Eneopterinae) from Singapore</mods:title>
</mods:titleInfo>
<mods:name id="EB2BB5EEB3B6082B5EE7B77E8C751000" type="personal">
<mods:role id="A316F9F28A5AF270BAF5667227005201">
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</mods:role>
<mods:namePart id="AED8EA08BA145947D8DB4A5F6F4D6532">Tan, Ming Kai</mods:namePart>
</mods:name>
<mods:name id="85D8411FD8E026D4458E9C2799611020" type="personal">
<mods:role id="489D0150E5EE80E0CB008B7FA0E76D1C">
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<mods:namePart id="D9A041E22FAF7CC784382AF2D7A7DE41">Robillard, Tony</mods:namePart>
</mods:name>
<mods:typeOfResource id="D4F861AB670A100C35667C8953DD42EE">text</mods:typeOfResource>
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<mods:title id="E151920B41044B89F4D2E8EC8C69260C">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="7CF570A3FBC02C228D7CDBAB8ED68EE6">
<mods:date id="565BF833ACD25B69D07674B3CDA7C3D8">2014</mods:date>
<mods:detail id="ED258ED5DA301D5E3D402B0DD8189C43" type="volume">
<mods:number id="CB6A7AA0CF32DAF19AB9EF9537430D3D">3764</mods:number>
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<mods:number id="33BBF157E52A5982FC13BA96B0D7930F">3</mods:number>
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<mods:classification id="2A5C5D7E7660ECCB52DB5183B1ACC89F">journal article</mods:classification>
<mods:identifier id="EDA6F5416D54EA44570D22484CEC1382" type="CLB-Dataset">46478</mods:identifier>
<mods:identifier id="8C3224D65A8928B726CFDBA0A038C649" type="DOI">10.11646/zootaxa.3764.3.6</mods:identifier>
<mods:identifier id="AE68986FAE296428B68BB8D9745F92A3" type="GBIF-Dataset">b7c3456f-0588-4d10-b263-062ca84f3fbb</mods:identifier>
<mods:identifier id="C4AD08C8F1DB164B1AAF69ADF812D9AF" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="65F19996C2ED5529EBAB38AB82DF958D" type="Zenodo-Dep">250851</mods:identifier>
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</mods:mods>
<treatment id="563E879E086AFFEC17F6F8EE0A99F9BF" ID-DOI="http://doi.org/10.5281/zenodo.5621554" ID-GBIF-Taxon="119414372" ID-Zenodo-Dep="5621554" LSID="urn:lsid:plazi:treatment:563E879E086AFFEC17F6F8EE0A99F9BF" httpUri="http://treatment.plazi.org/id/563E879E086AFFEC17F6F8EE0A99F9BF" lastPageId="7" lastPageNumber="371" pageId="1" pageNumber="365">
<subSubSection id="968D6503086AFFEA17F6F8EE0979F873" pageId="1" pageNumber="365" type="nomenclature">
<paragraph id="DE283688086AFFEA17F6F8EE0B37F852" blockId="1.[151,780,1903,1963]" box="[151,780,1903,1930]" pageId="1" pageNumber="365">
<heading id="856081E4086AFFEA17F6F8EE0B37F852" bold="true" box="[151,780,1903,1930]" fontSize="11" level="1" pageId="1" pageNumber="365" reason="1">
<emphasis id="ECE3EA9A086AFFEA17F6F8EE0B37F852" bold="true" box="[151,780,1903,1930]" pageId="1" pageNumber="365">
<taxonomicName id="19974D0B086AFFEA17F6F8EE0A86F852" authority="Tan &amp; Robillard" authorityName="Tan &amp; Robillard" box="[151,701,1903,1930]" class="Insecta" family="Haglotettigoniidae" genus="Cardiodactylus" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="365" phylum="Arthropoda" rank="species" species="admirabilis" status="sp. nov.">
<emphasis id="ECE3EA9A086AFFEA17F6F8EE09E6F851" bold="true" box="[151,477,1903,1929]" italics="true" pageId="1" pageNumber="365">Cardiodactylus admirabilis</emphasis>
, Tan &amp; Robillard
</taxonomicName>
,
<taxonomicNameLabel id="F7D057E1086AFFEA15ABF8F10B37F852" box="[714,780,1904,1930]" pageId="1" pageNumber="365" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="DE283688086AFFEA17F6F8130979F873" blockId="1.[151,780,1903,1963]" box="[151,322,1938,1963]" pageId="1" pageNumber="365">
(
<figureCitation id="46AC2A0D086AFFEA17FEF81308D3F873" box="[159,232,1938,1963]" captionStart="FIGURE 1" captionStartId="4.[151,250,1928,1950]" captionTargetBox="[151,1436,193,1905]" captionTargetId="figure@4.[151,1436,193,1906]" captionTargetPageId="4" captionText="FIGURE 1. Cardiodactylus adults recorded in Singapore: Cardiodactylus admirabilis n. sp. (A) and Cardiodactylus singapura Robillard, 2011 (B) in dorsal view." httpUri="https://zenodo.org/record/250852/files/figure.png" pageId="1" pageNumber="365">Figs. 1</figureCitation>
A, 2–7)
</paragraph>
</subSubSection>
<subSubSection id="968D6503086AFFEA17F6F85B0C87F828" box="[151,1212,2010,2032]" pageId="1" pageNumber="365" type="reference_group">
<paragraph id="DE283688086AFFEA17F6F85B0C87F828" blockId="1.[151,1212,2010,2032]" box="[151,1212,2010,2032]" pageId="1" pageNumber="365">
<taxonomicName id="19974D0B086AFFEA17F6F85B0A79F828" authority="Robillard, 2011" authorityName="Robillard" authorityYear="2011" box="[151,578,2010,2032]" class="Insecta" family="Haglotettigoniidae" genus="Cardiodactylus" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="365" phylum="Arthropoda" rank="species" species="singapura">
<emphasis id="ECE3EA9A086AFFEA17F6F85B09A4F828" box="[151,415,2010,2032]" italics="true" pageId="1" pageNumber="365">Cardiodactylus singapura</emphasis>
<bibRefCitation id="BA064B79086AFFEA16C4F85B0A79F828" author="Robillard" box="[421,578,2010,2032]" pageId="1" pageNumber="365" refString="Robillard, T. (2011) New Cardiodactylus species from unsuspected places in Southeast Asia (Orthoptera, Grylloidea, Eneopterinae). Zootaxa, 2909, 14 - 26." type="journal article" year="2011">Robillard, 2011</bibRefCitation>
</taxonomicName>
—Tan
<emphasis id="ECE3EA9A086AFFEA15E9F85A0A87F828" box="[648,700,2010,2032]" italics="true" pageId="1" pageNumber="365">et al.</emphasis>
2012: 66 (juvenile specimens wrongly identified).
</paragraph>
</subSubSection>
<subSubSection id="968D65030869FFE917F6FF160C8CFE74" pageId="2" pageNumber="366" type="materials_examined">
<paragraph id="DE2836880869FFE917F6FF16098FFF20" blockId="2.[151,1436,151,1981]" pageId="2" pageNumber="366">
<emphasis id="ECE3EA9A0869FFE917F6FF1609B7FF68" bold="true" box="[151,396,151,176]" pageId="2" pageNumber="366">Material examined.</emphasis>
<typeStatus id="012C882A0869FFE916FCFF160A3EFF68" box="[413,517,151,176]" pageId="2" pageNumber="366" type="holotype">Holotype</typeStatus>
(male):
<emphasis id="ECE3EA9A0869FFE9151AFF160ACFFF68" bold="true" box="[635,756,151,176]" pageId="2" pageNumber="366">
<collectingCountry id="A68076180869FFE9151AFF160ACFFF68" box="[635,756,151,176]" name="Singapore" pageId="2" pageNumber="366">Singapore</collectingCountry>
</emphasis>
, Admiralty Park (Nature Park), secondary forest near mangroves,
<geoCoordinate id="BBA3504F0869FFE91641FF3D099BFF0D" box="[288,416,188,213]" direction="north" orientation="latitude" pageId="2" pageNumber="366" precision="1" value="1.44639">N01.44639</geoCoordinate>
,
<geoCoordinate id="BBA3504F0869FFE916CDFF3D0A0CFF0C" box="[428,567,188,212]" direction="east" orientation="longitude" pageId="2" pageNumber="366" precision="1" value="103.7793">E103.77930</geoCoordinate>
, 11.2 ±
<quantity id="196F9B6D0869FFE915F0FF3D0AEFFF0C" box="[657,724,188,213]" metricMagnitude="0" metricUnit="m" metricValue="5.4" pageId="2" pageNumber="366" unit="m" value="5.4">5.4 m</quantity>
, on
<taxonomicName id="19974D0B0869FFE91462FF3D0B64FF0C" box="[771,863,188,212]" class="Magnoliopsida" family="Dilleniaceae" genus="Dillenia" kingdom="Plantae" order="Dilleniales" pageId="2" pageNumber="366" phylum="Tracheophyta" rank="genus">
<emphasis id="ECE3EA9A0869FFE91462FF3D0B64FF0C" box="[771,863,188,212]" italics="true" pageId="2" pageNumber="366">Dillenia</emphasis>
</taxonomicName>
leaf,
<date id="AA2910480869FFE914FEFF3D0C0CFF0C" box="[927,1079,188,213]" pageId="2" pageNumber="366" value="2013-06-25">25 June 2013</date>
, 2017 hours, coll. H. Yeo &amp; M. K. Tan (
<collectionCode id="B886AE4D0869FFE91790FF610917FF20" box="[241,300,224,248]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="366">ZRC</collectionCode>
.ORT.1020).
</paragraph>
<paragraph id="DE2836880869FFE917A6FE840C8CFE74" blockId="2.[151,1436,151,1981]" pageId="2" pageNumber="366">
<typeStatus id="012C882A0869FFE917A6FE840903FEC5" box="[199,312,261,285]" pageId="2" pageNumber="366" type="paratype">Paratypes</typeStatus>
.
<collectingCountry id="A68076180869FFE91620FE850981FEC5" box="[321,442,260,285]" name="Singapore" pageId="2" pageNumber="366">
<emphasis id="ECE3EA9A0869FFE91620FE850981FEC5" bold="true" box="[321,442,260,285]" pageId="2" pageNumber="366">Singapore</emphasis>
</collectingCountry>
, same locality as
<typeStatus id="012C882A0869FFE915E0FE850ADDFEC5" box="[641,742,260,285]" pageId="2" pageNumber="366" type="holotype">holotype</typeStatus>
, secondary forest near mangroves,
<specimenCount id="C891FD010869FFE9130EFE850CF3FEC5" box="[1135,1224,260,285]" pageId="2" pageNumber="366" type="male">3 males</specimenCount>
, coll. H. Yeo &amp; M. K. Tan: (1)
<geoCoordinate id="BBA3504F0869FFE91676FEA609ACFE98" box="[279,407,295,320]" direction="north" orientation="latitude" pageId="2" pageNumber="366" precision="1" value="1.44637">N01.44637</geoCoordinate>
,
<geoCoordinate id="BBA3504F0869FFE916C3FEA60A16FE98" box="[418,557,295,320]" direction="east" orientation="longitude" pageId="2" pageNumber="366" precision="1" value="103.77926">E103.77926</geoCoordinate>
, 27.4 ±
<quantity id="196F9B6D0869FFE915E5FEA60AFEFEE7" box="[644,709,295,320]" metricMagnitude="0" metricUnit="m" metricValue="5.6" pageId="2" pageNumber="366" unit="m" value="5.6">5.6 m</quantity>
, on
<taxonomicName id="19974D0B0869FFE91592FEA90B74FE98" box="[755,847,296,320]" class="Magnoliopsida" family="Dilleniaceae" genus="Dillenia" kingdom="Plantae" order="Dilleniales" pageId="2" pageNumber="366" phylum="Tracheophyta" rank="genus">
<emphasis id="ECE3EA9A0869FFE91592FEA90B74FE98" box="[755,847,296,320]" italics="true" pageId="2" pageNumber="366">Dillenia</emphasis>
</taxonomicName>
leaf,
<date id="AA2910480869FFE914EDFEA60C19FE98" box="[908,1058,295,320]" pageId="2" pageNumber="366" value="2013-06-15">15 June 2013</date>
, 2120 hours, collected as the final instar, moulted into adult on
<date id="AA2910480869FFE91685FECD0A45FEBC" box="[484,638,332,357]" pageId="2" pageNumber="366" value="2013-06-16">16 June 2013</date>
(
<collectionCode id="B886AE4D0869FFE915EEFECD0AF0FEBC" box="[655,715,332,356]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="366">ZRC</collectionCode>
.ORT.1021); (2)
<geoCoordinate id="BBA3504F0869FFE914E4FECD0C3EFEBD" box="[901,1029,332,357]" direction="north" orientation="latitude" pageId="2" pageNumber="366" precision="1" value="1.44676">N01.44676</geoCoordinate>
,
<geoCoordinate id="BBA3504F0869FFE91372FECD0CA5FEBC" box="[1043,1182,332,356]" direction="east" orientation="longitude" pageId="2" pageNumber="366" precision="1" value="103.77919">E103.77919</geoCoordinate>
, 10.7 ±
<quantity id="196F9B6D0869FFE9139FFECD0D79FEBC" box="[1278,1346,332,357]" metricMagnitude="0" metricUnit="m" metricValue="5.2" pageId="2" pageNumber="366" unit="m" value="5.2">5.2 m</quantity>
, among herbs,
<date id="AA2910480869FFE91782FEEE0944FE50" box="[227,383,367,392]" pageId="2" pageNumber="366" value="2013-06-25">25 June 2013</date>
, 2042 hours (
<collectionCode id="B886AE4D0869FFE9157AFEF10A6DFE50" box="[539,598,368,392]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="366">ZRC</collectionCode>
.ORT.1022); (3)
<geoCoordinate id="BBA3504F0869FFE91471FEEE0BAAFE50" box="[784,913,367,392]" direction="north" orientation="latitude" pageId="2" pageNumber="366" precision="1" value="1.44682">N01.44682</geoCoordinate>
,
<geoCoordinate id="BBA3504F0869FFE914FFFEF10C12FE50" box="[926,1065,368,392]" direction="east" orientation="longitude" pageId="2" pageNumber="366" precision="1" value="103.77908">E103.77908</geoCoordinate>
, 17.2 ±
<quantity id="196F9B6D0869FFE913E6FEEE0CF0FE5F" box="[1159,1227,367,392]" metricMagnitude="0" metricUnit="m" metricValue="6.4" pageId="2" pageNumber="366" unit="m" value="6.4">6.4 m</quantity>
, calling on leaf of tree sapling, calling song recorded in-situ,
<date id="AA2910480869FFE91515FE150AC1FE74" box="[628,762,404,429]" pageId="2" pageNumber="366" value="2013-07-06">6 July 2013</date>
, 2145 hours (MNHN-EO-ENSIF3441).
</paragraph>
</subSubSection>
<subSubSection id="968D65030869FFE917A6FE36097EFD15" pageId="2" pageNumber="366" type="diagnosis">
<paragraph id="DE2836880869FFE917A6FE36097EFD15" blockId="2.[151,1436,151,1981]" pageId="2" pageNumber="366">
<emphasis id="ECE3EA9A0869FFE917A6FE36097AFE08" bold="true" box="[199,321,439,464]" pageId="2" pageNumber="366">Diagnosis.</emphasis>
Species of average size for the species group, characterised by contrasted coloration dark brown and yellow, with a wide rounded yellow area posterior and male genitalia more or less triangular, with a little rounded posterior apex. Very close to
<taxonomicName id="19974D0B0869FFE9150CFD810C64FDC0" authority="Robillard, 2011" authorityName="Robillard" authorityYear="2011" box="[621,1119,511,536]" class="Insecta" family="Haglotettigoniidae" genus="Cardiodactylus" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="366" phylum="Arthropoda" rank="species" species="thailandia">
<emphasis id="ECE3EA9A0869FFE9150CFD810BA1FDC0" box="[621,922,512,536]" italics="true" pageId="2" pageNumber="366">Cardiodactylus thailandia</emphasis>
<bibRefCitation id="BA064B790869FFE914C9FE7E0C64FDC0" author="Robillard" box="[936,1119,511,536]" pageId="2" pageNumber="366" refString="Robillard, T. (2011) New Cardiodactylus species from unsuspected places in Southeast Asia (Orthoptera, Grylloidea, Eneopterinae). Zootaxa, 2909, 14 - 26." type="journal article" year="2011">Robillard, 2011</bibRefCitation>
</taxonomicName>
from which it differs the following characters: general colouration brown and whitish while
<taxonomicName id="19974D0B0869FFE914EAFDA50C24FDE4" box="[907,1055,548,572]" class="Insecta" family="Haglotettigoniidae" genus="Cardiodactylus" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="366" phylum="Arthropoda" rank="species" species="thailandia">
<emphasis id="ECE3EA9A0869FFE914EAFDA50C24FDE4" box="[907,1055,548,572]" italics="true" pageId="2" pageNumber="366">C. thailandia</emphasis>
</taxonomicName>
is almost black and vivid yellow; male FW: diagonal not bicolour; fore and middle legs: pale yellow brown with dark brown patterns (spots on femur, rings on tibiae) while legs of
<taxonomicName id="19974D0B0869FFE9152FFDED0ADDFD5C" box="[590,742,620,644]" class="Insecta" family="Haglotettigoniidae" genus="Cardiodactylus" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="366" phylum="Arthropoda" rank="species" species="thailandia">
<emphasis id="ECE3EA9A0869FFE9152FFDED0ADDFD5C" box="[590,742,620,644]" italics="true" pageId="2" pageNumber="366">C. thailandia</emphasis>
</taxonomicName>
are almost entirely orange brown; male genitalia differ by pseudephiphallus posterior region (posterior lobe wider than in
<taxonomicName id="19974D0B0869FFE9143DFD110BCBFD70" box="[860,1008,656,680]" class="Insecta" family="Haglotettigoniidae" genus="Cardiodactylus" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="366" phylum="Arthropoda" rank="species" species="thailandia">
<emphasis id="ECE3EA9A0869FFE9143DFD110BCBFD70" box="[860,1008,656,680]" italics="true" pageId="2" pageNumber="366">C. thailandia</emphasis>
</taxonomicName>
) and dorsal edge of dorsal ridges with different shape.
</paragraph>
</subSubSection>
<subSubSection id="968D65030869FFE917A6FD590B3CFC35" pageId="2" pageNumber="366" type="reference_group">
<paragraph id="DE2836880869FFE917A6FD590B3CFC35" blockId="2.[151,1436,151,1981]" pageId="2" pageNumber="366">
<taxonomicName id="19974D0B0869FFE917A6FD590955FD28" box="[199,366,728,752]" class="Insecta" family="Haglotettigoniidae" genus="Cardiodactylus" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="366" phylum="Arthropoda" rank="species" species="admirabilis">
<emphasis id="ECE3EA9A0869FFE917A6FD590955FD28" box="[199,366,728,752]" italics="true" pageId="2" pageNumber="366">C. admirabilis</emphasis>
</taxonomicName>
differs strikingly from
<taxonomicName id="19974D0B0869FFE915ECFD590C42FD28" authority="Robillard, 2011" authorityName="Robillard" authorityYear="2011" box="[653,1145,727,752]" class="Insecta" family="Haglotettigoniidae" genus="Cardiodactylus" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="366" phylum="Arthropoda" rank="species" species="singapura">
<emphasis id="ECE3EA9A0869FFE915ECFD590B83FD37" box="[653,952,728,752]" italics="true" pageId="2" pageNumber="366">Cardiodactylus singapura</emphasis>
<bibRefCitation id="BA064B790869FFE914A5FD560C42FD28" author="Robillard" box="[964,1145,727,752]" pageId="2" pageNumber="366" refString="Robillard, T. (2011) New Cardiodactylus species from unsuspected places in Southeast Asia (Orthoptera, Grylloidea, Eneopterinae). Zootaxa, 2909, 14 - 26." type="journal article" year="2011">Robillard, 2011</bibRefCitation>
</taxonomicName>
by its colouration more contrasted on FW dorsal field, with a wide whitish area posterior to mirror (
<figureCitation id="46AC2A0D0869FFE91490FD7D0C08FCCC" box="[1009,1075,764,789]" captionStart="FIGURE 1" captionStartId="4.[151,250,1928,1950]" captionTargetBox="[151,1436,193,1905]" captionTargetId="figure@4.[151,1436,193,1906]" captionTargetPageId="4" captionText="FIGURE 1. Cardiodactylus adults recorded in Singapore: Cardiodactylus admirabilis n. sp. (A) and Cardiodactylus singapura Robillard, 2011 (B) in dorsal view." httpUri="https://zenodo.org/record/250852/files/figure.png" pageId="2" pageNumber="366">Fig. 1</figureCitation>
); and male genitalia: rami wide (narrow in
<taxonomicName id="19974D0B0869FFE91677FCA10990FCEF" box="[278,427,800,823]" class="Insecta" family="Haglotettigoniidae" genus="Cardiodactylus" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="366" phylum="Arthropoda" rank="species" species="singapura">
<emphasis id="ECE3EA9A0869FFE91677FCA10990FCEF" box="[278,427,800,823]" italics="true" pageId="2" pageNumber="366">C. singapura</emphasis>
</taxonomicName>
), with preapical plate more sclerotised along edge (equally sclerotised in
<taxonomicName id="19974D0B0869FFE91396FCA10DB1FCEF" box="[1271,1418,800,823]" class="Insecta" family="Haglotettigoniidae" genus="Cardiodactylus" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="366" phylum="Arthropoda" rank="species" species="singapura">
<emphasis id="ECE3EA9A0869FFE91396FCA10DB1FCEF" box="[1271,1418,800,823]" italics="true" pageId="2" pageNumber="366">C. singapura</emphasis>
</taxonomicName>
); ectophallic arc angulated (more broadly rounded in
<taxonomicName id="19974D0B0869FFE9159BFCC50BABFC84" box="[762,912,836,860]" class="Insecta" family="Haglotettigoniidae" genus="Cardiodactylus" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="366" phylum="Arthropoda" rank="species" species="singapura">
<emphasis id="ECE3EA9A0869FFE9159BFCC50BABFC84" box="[762,912,836,860]" italics="true" pageId="2" pageNumber="366">C. singapura</emphasis>
</taxonomicName>
); ectophallic apodemes lamellate (thin in
<taxonomicName id="19974D0B0869FFE912E3FCC50932FCA7" class="Insecta" family="Haglotettigoniidae" genus="Cardiodactylus" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="366" phylum="Arthropoda" rank="species" species="singapura">
<emphasis id="ECE3EA9A0869FFE912E3FCC50932FCA7" italics="true" pageId="2" pageNumber="366">C. singapura</emphasis>
</taxonomicName>
); endophallic sclerite sclerotized and rather wide, not tapering anteriorly but truncated with the apex acute (small and tapering anteriorly to an acute apex in
<taxonomicName id="19974D0B0869FFE91445FC0D0B81FC7C" box="[804,954,908,932]" class="Insecta" family="Haglotettigoniidae" genus="Cardiodactylus" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="366" phylum="Arthropoda" rank="species" species="singapura">
<emphasis id="ECE3EA9A0869FFE91445FC0D0B81FC7C" box="[804,954,908,932]" italics="true" pageId="2" pageNumber="366">C. singapura</emphasis>
</taxonomicName>
). Contrary to most species of the species group described from this geographic area (
<bibRefCitation id="BA064B790869FFE915F8FC2E0C4EFC10" author="Robillard" box="[665,1141,943,968]" pageId="2" pageNumber="366" refString="Robillard, T. (2011) New Cardiodactylus species from unsuspected places in Southeast Asia (Orthoptera, Grylloidea, Eneopterinae). Zootaxa, 2909, 14 - 26." type="journal article" year="2011">
Robillard, 2011; Robillard
<emphasis id="ECE3EA9A0869FFE914B7FC300C28FC10" box="[982,1043,944,968]" italics="true" pageId="2" pageNumber="366">et al.</emphasis>
in prep
</bibRefCitation>
),
<taxonomicName id="19974D0B0869FFE913EDFC310D08FC10" box="[1164,1331,944,968]" class="Insecta" family="Haglotettigoniidae" genus="Cardiodactylus" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="366" phylum="Arthropoda" rank="species" species="admirabilis">
<emphasis id="ECE3EA9A0869FFE913EDFC310D08FC10" box="[1164,1331,944,968]" italics="true" pageId="2" pageNumber="366">C. admirabilis</emphasis>
</taxonomicName>
shows a multisyllabic calling song and not a monosyllabic song.
</paragraph>
</subSubSection>
<subSubSection id="968D65030869FFE817A6FC760A7FF9BF" lastPageId="3" lastPageNumber="367" pageId="2" pageNumber="366" type="description">
<paragraph id="DE2836880869FFE917A6FC760B6EFA3D" blockId="2.[151,1436,151,1981]" pageId="2" pageNumber="366">
<emphasis id="ECE3EA9A0869FFE917A6FC760962FBC8" bold="true" box="[199,345,1015,1040]" pageId="2" pageNumber="366">Description.</emphasis>
Size average for the species group. General colouration contrasted, light brown with dark brown patterns (
<figureCitation id="46AC2A0D0869FFE9179EFB9D097FFBEC" box="[255,324,1052,1077]" captionStart="FIGURE 2" captionStartId="5.[151,250,1975,1997]" captionTargetBox="[239,1332,197,1937]" captionTargetId="figure@5.[232,1333,193,1938]" captionTargetPageId="5" captionText="FIGURE 2. Cardiodactylus admirabilis n. sp. male habitus: dorsal (A) and lateral (B) views. Scale bars: 2 mm (A); 10 mm (B)." httpUri="https://zenodo.org/record/250853/files/figure.png" pageId="2" pageNumber="366">Fig. 2</figureCitation>
). Head dorsum yellow brown with 4 wide dark brown bands, more or less punctuated, bordered by dark brown triangles posterior to eyes and a yellow line more externally (Fig. 3A). Fastigium yellow laterally, median area dark brown, joint with the 2 outermost dark brown bands on vertex. Scapus light brown. Antennae heterogeneously brown. Face yellowish to light brown, diversely mottled with brown, with 2 parallel lines on the fastigium (Fig. 3B). Maxillary palpi yellowish. Head laterally sometimes with a dark brown area posterior to eyes and genae bordering the eyes, progressively lighter ventrally (Fig. 3C). Pronotum: Dorsal disk trapezoidal, anterior margin concave, posterior margin straight or slightly bisinuated; dark brown mottled with light brown or yellow spots. Lateral lobes dark brown, with a black longitudinal band on dorsal half, ventral edges and dorsal margins yellow (Fig. 3C). Legs light brown to yellow brown, knees dark brown, femora mottled with dark brown; tibiae darker, more or less banded; tarsomeres with dark brown ends (
<figureCitation id="46AC2A0D0869FFE9140FFADE0B8EFAA0" box="[878,949,1375,1400]" captionStart="FIGURE 2" captionStartId="5.[151,250,1975,1997]" captionTargetBox="[239,1332,197,1937]" captionTargetId="figure@5.[232,1333,193,1938]" captionTargetPageId="5" captionText="FIGURE 2. Cardiodactylus admirabilis n. sp. male habitus: dorsal (A) and lateral (B) views. Scale bars: 2 mm (A); 10 mm (B)." httpUri="https://zenodo.org/record/250853/files/figure.png" pageId="2" pageNumber="366">Fig. 2</figureCitation>
). Tarsomeres
<date id="AA2910480869FFE91335FAE10CBCFAA0" box="[1108,1159,1376,1400]" pageId="2" pageNumber="366">III-1</date>
with 3 spines on dorsoexternal edges (m=3, n=4).
<collectingCountry id="A68076180869FFE916B7FA050A35FA45" box="[470,526,1412,1437]" name="India" pageId="2" pageNumber="366">Hind</collectingCountry>
wings longer than FWs, the dark brown tail exceeding the forewings less than twice longer than the pronotum (
<figureCitation id="46AC2A0D0869FFE9156AFA260A6BFA18" box="[523,592,1447,1472]" captionStart="FIGURE 2" captionStartId="5.[151,250,1975,1997]" captionTargetBox="[239,1332,197,1937]" captionTargetId="figure@5.[232,1333,193,1938]" captionTargetPageId="5" captionText="FIGURE 2. Cardiodactylus admirabilis n. sp. male habitus: dorsal (A) and lateral (B) views. Scale bars: 2 mm (A); 10 mm (B)." httpUri="https://zenodo.org/record/250853/files/figure.png" pageId="2" pageNumber="366">Fig. 2</figureCitation>
A). Cerci light brown with faint black rings. Tergites dark brown or black; sternites light brown to brown. Subgenital plate yellow brown.
</paragraph>
<paragraph id="DE2836880869FFE917A6FA6E0AD5F864" blockId="2.[151,1436,151,1981]" pageId="2" pageNumber="366">
Male. FW as shown in Fig. 3D. Colouration: Dorsal field cells dark brown, veins mostly orange brown, some areas white with yellow veins, including base of FWs and anal veins, area near plectrum, part of the chords, and wide semi-circular area posterior to mirror. Base of chords with a dark brown sclerotization. Harp posterior angle and anterior area with a white and yellow sclerotized area. Lateral field with cells mostly dark brown, except areas near projections of Sc; veins M, R, Sc dark brown, transverse veins and projections of Sc pale yellow. FW venation: 1A slightly bisinuated; stridulatory vein with transverse part
<quantity id="196F9B6D0869FFE914A5F9250C25F964" box="[964,1054,1700,1724]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="2" pageNumber="366" unit="mm" value="3.3">3.3 mm</quantity>
, with 176 teeth on the transverse area of the file, 18 on the curved area, and 22 near base of 1A (n=1,
<typeStatus id="012C882A0869FFE914FEF9460C38F938" box="[927,1027,1735,1760]" pageId="2" pageNumber="366" type="holotype">holotype</typeStatus>
) (Fig. 3E). CuP absent. Harp longer than wide, with 2 w-shaped harp veins; posterior margin raised along diagonal vein. Mirror area variable, c1 very long and very narrow and fairly straight to slightly sinuous, c2 quite large; mirror (d1) longer than wide, not rounded, generally separated in four parts by transverse and oblique veins, separating vein more posterior than anterior, the posterior part triangular; e1 cell very elongated, along the mirror. Apical field long and triangular; with 3 wide cell alignments posterior to mirror, 2 apical most wide cells with dividing veins. Lateral field: Sc with 7–8 projections and 3 more ventral veins (m=7.8, 3, n=4).
</paragraph>
<paragraph id="DE2836880868FFE817A6FF160BAEFDC0" blockId="3.[151,1436,151,752]" pageId="3" pageNumber="367">
Male genitalia as shown in
<figureCitation id="46AC2A0D0868FFE81562FF160A71FF68" box="[515,586,151,176]" captionStart="FIGURE 4" captionStartId="7.[151,250,1077,1099]" captionTargetBox="[157,1433,200,1044]" captionTargetId="figure@7.[151,1436,193,1056]" captionTargetPageId="7" captionText="FIGURE 4. Cardiodactylus admirabilis n. sp. male genitalia: dorsal (A); ventral (B) and lateral (C) views. Scale bar: 1 mm." httpUri="https://zenodo.org/record/250854/files/figure.png" pageId="3" pageNumber="367">Fig. 4</figureCitation>
: Pseudepiphallus very sclerotized and little setose, more or less triangular, only slightly narrowed new mid-length. Dorsal ridges divergent posteriorly, their dorsal edge bean-like, slightly asymmetrical, carinated interiorly, the left one curved, the right one straight; pseudepiphallus bisinuated laterally. Pseudepiphallic sclerite with latero-anterior triangular expansions; the membrane between them with short strong setae. Apex of pseudepiphallus spoon-like, not rounded. Rami wide, with wide triangular preapical plate; plate with lateral edge rounded and more sclerotised; with apex lamellate and acute. Pseudepiphallic parameres trilobate, the posterior lobe wide. Ectophallic arc complete, with a short posterior expansion. Ectophallic apodemes lamellate, diverging, their apex curved dorso-interiorly. Apex of ectophallic fold trilobate, membranous. Base of ectophallic apodemes with thin sclerotized expansions oriented posteriorly. Endophallic sclerite small, with a medio posterior anpansion and 2 thin lateral arms sclerotised. Endophallic apodeme with both lateral lamellas, and a short medio-dorsal crest. Membrane of endophallic cavity smooth.
</paragraph>
<paragraph id="DE2836880868FFE817A6FDA509A6FDE5" blockId="3.[151,1436,151,752]" box="[199,413,548,573]" pageId="3" pageNumber="367">Female. Unknown.</paragraph>
<paragraph id="DE2836880868FFE817A6FDC60A21FD15" blockId="3.[151,1436,151,752]" pageId="3" pageNumber="367">
Juvenile (
<figureCitation id="46AC2A0D0868FFE81652FDC6094CFDB8" box="[307,375,583,608]" captionStart="FIGURE 5" captionStartId="8.[151,250,1981,2003]" captionTargetBox="[217,1366,197,1937]" captionTargetId="figure@8.[214,1368,195,1940]" captionTargetPageId="8" captionText="FIGURE 5. Cardiodactylus admirabilis n. sp. juvenile habitus: lateral (A) and dorsal (B) views of early instar and dorsal view of later instar (C)." httpUri="https://zenodo.org/record/250855/files/figure.png" pageId="3" pageNumber="367">Fig. 5</figureCitation>
). Early instars generally pale with irregular spots of different size; dorsum of head mostly dark brown with faint yellow bands; with black longitudinal bands from genae towards abdominal apex (
<figureCitation id="46AC2A0D0868FFE81271FDED0D6EFD5D" box="[1296,1365,620,645]" captionStart="FIGURE 5" captionStartId="8.[151,250,1981,2003]" captionTargetBox="[217,1366,197,1937]" captionTargetId="figure@8.[214,1368,195,1940]" captionTargetPageId="8" captionText="FIGURE 5. Cardiodactylus admirabilis n. sp. juvenile habitus: lateral (A) and dorsal (B) views of early instar and dorsal view of later instar (C)." httpUri="https://zenodo.org/record/250855/files/figure.png" pageId="3" pageNumber="367">Fig. 5</figureCitation>
A, B). Later instars with more contrasted colouration, body darker with spots denser; dorsum of head with more contrasted yellow bands (
<figureCitation id="46AC2A0D0868FFE816D4FD3509CCFD15" box="[437,503,692,717]" captionStart="FIGURE 5" captionStartId="8.[151,250,1981,2003]" captionTargetBox="[217,1366,197,1937]" captionTargetId="figure@8.[214,1368,195,1940]" captionTargetPageId="8" captionText="FIGURE 5. Cardiodactylus admirabilis n. sp. juvenile habitus: lateral (A) and dorsal (B) views of early instar and dorsal view of later instar (C)." httpUri="https://zenodo.org/record/250855/files/figure.png" pageId="3" pageNumber="367">Fig. 5</figureCitation>
C).
</paragraph>
<paragraph id="DE2836880868FFE817A6FD590A3AFD28" blockId="3.[151,1436,151,752]" box="[199,513,727,752]" pageId="3" pageNumber="367">
Measurements. See
<tableCitation id="931503330868FFE816C6FD5609C0FD28" box="[423,507,727,752]" captionStart="TABLE 1" captionStartId="3.[151,239,800,823]" captionTargetBox="[159,1412,851,1130]" captionTargetPageId="3" captionText="TABLE 1. Measurements of Cardiodactylus admirabilis n. sp. (in mm, mean values in brackets)." httpUri="http://table.plazi.org/id/8AE866000868FFE817F6FCA10C93FCEF" pageId="3" pageNumber="367" tableUuid="8AE866000868FFE817F6FCA10C93FCEF">Table 1</tableCitation>
.
</paragraph>
<caption id="8AE866000868FFE817F6FCA10C93FCEF" ID-Table-UUID="8AE866000868FFE817F6FCA10C93FCEF" box="[151,1192,799,823]" httpUri="http://table.plazi.org/id/8AE866000868FFE817F6FCA10C93FCEF" pageId="3" pageNumber="367" targetBox="[159,1412,851,1130]" targetIsTable="true" targetPageId="3">
<paragraph id="DE2836880868FFE817F6FCA10C93FCEF" blockId="3.[151,1192,799,823]" box="[151,1192,799,823]" pageId="3" pageNumber="367">
<emphasis id="ECE3EA9A0868FFE817F6FCA10932FCEF" bold="true" box="[151,265,800,823]" pageId="3" pageNumber="367">TABLE 1.</emphasis>
Measurements of
<taxonomicName id="19974D0B0868FFE816AEFC9E0ACEFCEF" box="[463,757,799,823]" class="Insecta" family="Haglotettigoniidae" genus="Cardiodactylus" kingdom="Animalia" order="Orthoptera" pageId="3" pageNumber="367" phylum="Arthropoda" rank="species" species="admirabilis" status="sp. nov.">
<emphasis id="ECE3EA9A0868FFE816AEFC9E0ACEFCEF" box="[463,757,799,823]" italics="true" pageId="3" pageNumber="367">Cardiodactylus admirabilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F7D057E10868FFE8159AFCA10B0CFCEF" box="[763,823,800,823]" pageId="3" pageNumber="367" rank="species">
<emphasis id="ECE3EA9A0868FFE8159AFCA10B0CFCEF" bold="true" box="[763,823,800,823]" pageId="3" pageNumber="367">n. sp.</emphasis>
</taxonomicNameLabel>
(in mm, mean values in brackets).
</paragraph>
</caption>
<paragraph id="DE2836880868FFE81609FCD20D71FB93" pageId="3" pageNumber="367">
<table id="AC97C4280868001417FEFCD20DBFFBB2" box="[159,1412,851,1130]" gridcols="8" gridrows="5" pageId="3" pageNumber="367">
<tr id="60A734CA0868001417FEFCD20DBFFCB0" box="[159,1412,851,872]" gridrow="0" pageId="3" pageNumber="367">
<th id="23765DB60868001417FEFCD2099CFCB0" box="[159,423,851,872]" gridcol="0" gridrow="0" pageId="3" pageNumber="367">PronL</th>
<th id="23765DB6086800141691FCD20A03FCB0" box="[496,568,851,872]" gridcol="1" gridrow="0" pageId="3" pageNumber="367">PronW</th>
<th id="23765DB6086800141519FCD20A96FCB0" box="[632,685,851,872]" gridcol="2" gridrow="0" pageId="3" pageNumber="367">FWL</th>
<th id="23765DB608680014159EFCD20B06FCB0" box="[767,829,851,872]" gridcol="3" gridrow="0" pageId="3" pageNumber="367">FWW</th>
<th id="23765DB60868001414E6FCD20BFAFCB0" box="[903,961,851,872]" gridcol="4" gridrow="0" pageId="3" pageNumber="367">HWT</th>
<th id="23765DB608680014136EFCD20C7DFCB0" box="[1039,1094,851,872]" gridcol="5" gridrow="0" pageId="3" pageNumber="367">FIIIL</th>
<th id="23765DB60868001413F6FCD20CEDFCB0" box="[1175,1238,851,872]" gridcol="6" gridrow="0" pageId="3" pageNumber="367">FIIIW</th>
<th id="23765DB608680014127FFCD20DBFFCB0" box="[1310,1412,851,872]" gridcol="7" gridrow="0" pageId="3" pageNumber="367">TIIIL</th>
</tr>
<tr id="60A734CA0868001417FEFCFD0DBFFC4A" box="[159,1412,892,914]" gridrow="1" pageId="3" pageNumber="367">
<th id="23765DB60868001417FEFCFD099CFC4A" box="[159,423,892,914]" gridcol="0" gridrow="1" pageId="3" pageNumber="367">Holotype 3.9</th>
<td id="23765DB6086800141691FCFD0A03FC4A" box="[496,568,892,914]" gridcol="1" gridrow="1" pageId="3" pageNumber="367">4.8</td>
<td id="23765DB6086800141519FCFD0A96FC4A" box="[632,685,892,914]" gridcol="2" gridrow="1" pageId="3" pageNumber="367">15.2</td>
<td id="23765DB608680014159EFCFD0B06FC4A" box="[767,829,892,914]" gridcol="3" gridrow="1" pageId="3" pageNumber="367">4.6</td>
<td id="23765DB60868001414E6FCFD0BFAFC4A" box="[903,961,892,914]" gridcol="4" gridrow="1" pageId="3" pageNumber="367">5.0</td>
<td id="23765DB608680014136EFCFD0C7DFC4A" box="[1039,1094,892,914]" gridcol="5" gridrow="1" pageId="3" pageNumber="367">15.9</td>
<td id="23765DB60868001413F6FCFD0CEDFC4A" box="[1175,1238,892,914]" gridcol="6" gridrow="1" pageId="3" pageNumber="367">4.1</td>
<td id="23765DB608680014127FFCFD0DBFFC4A" box="[1310,1412,892,914]" gridcol="7" gridrow="1" pageId="3" pageNumber="367">13.6</td>
</tr>
<tr id="60A734CA0868001417FEFC240DBFFC02" box="[159,1412,933,986]" gridrow="2" pageId="3" pageNumber="367">
<th id="23765DB60868001417FEFC24099CFC02" box="[159,423,933,986]" gridcol="0" gridrow="2" pageId="3" pageNumber="367">Male paratype 3.6 (15 Jun.2013)</th>
<td id="23765DB6086800141691FC240A03FC02" box="[496,568,933,986]" gridcol="1" gridrow="2" pageId="3" pageNumber="367">4.8</td>
<td id="23765DB6086800141519FC240A96FC02" box="[632,685,933,986]" gridcol="2" gridrow="2" pageId="3" pageNumber="367">14.9</td>
<td id="23765DB608680014159EFC240B06FC02" box="[767,829,933,986]" gridcol="3" gridrow="2" pageId="3" pageNumber="367">4.5</td>
<td id="23765DB60868001414E6FC240BFAFC02" box="[903,961,933,986]" gridcol="4" gridrow="2" pageId="3" pageNumber="367">5.0</td>
<td id="23765DB608680014136EFC240C7DFC02" box="[1039,1094,933,986]" gridcol="5" gridrow="2" pageId="3" pageNumber="367">15.8</td>
<td id="23765DB60868001413F6FC240CEDFC02" box="[1175,1238,933,986]" gridcol="6" gridrow="2" pageId="3" pageNumber="367">3.5*</td>
<td id="23765DB608680014127FFC240DBFFC02" box="[1310,1412,933,986]" gridcol="7" gridrow="2" pageId="3" pageNumber="367">14.6</td>
</tr>
<tr id="60A734CA0868001417FEFC6C0DBFFBFA" box="[159,1412,1005,1058]" gridrow="3" pageId="3" pageNumber="367">
<th id="23765DB60868001417FEFC6C099CFBFA" box="[159,423,1005,1058]" gridcol="0" gridrow="3" pageId="3" pageNumber="367">Male paratype 3.5 (25 Jun.2013)</th>
<td id="23765DB6086800141691FC6C0A03FBFA" box="[496,568,1005,1058]" gridcol="1" gridrow="3" pageId="3" pageNumber="367">4.6</td>
<td id="23765DB6086800141519FC6C0A96FBFA" box="[632,685,1005,1058]" gridcol="2" gridrow="3" pageId="3" pageNumber="367">15.0</td>
<td id="23765DB608680014159EFC6C0B06FBFA" box="[767,829,1005,1058]" gridcol="3" gridrow="3" pageId="3" pageNumber="367">4.5</td>
<td id="23765DB60868001414E6FC6C0BFAFBFA" box="[903,961,1005,1058]" gridcol="4" gridrow="3" pageId="3" pageNumber="367">5.0</td>
<td id="23765DB608680014136EFC6C0C7DFBFA" box="[1039,1094,1005,1058]" gridcol="5" gridrow="3" pageId="3" pageNumber="367">15.9</td>
<td id="23765DB60868001413F6FC6C0CEDFBFA" box="[1175,1238,1005,1058]" gridcol="6" gridrow="3" pageId="3" pageNumber="367">4.2</td>
<td id="23765DB608680014127FFC6C0DBFFBFA" box="[1310,1412,1005,1058]" gridcol="7" gridrow="3" pageId="3" pageNumber="367">14.4</td>
</tr>
<tr id="60A734CA0868001417FEFBB40DBFFBB2" box="[159,1412,1077,1130]" gridrow="4" pageId="3" pageNumber="367">
<th id="23765DB60868001417FEFBB4099CFBB2" box="[159,423,1077,1130]" gridcol="0" gridrow="4" pageId="3" pageNumber="367">Male paratype 3.1 (6 Jul.2013)</th>
<td id="23765DB6086800141691FBB40A03FBB2" box="[496,568,1077,1130]" gridcol="1" gridrow="4" pageId="3" pageNumber="367">4.9</td>
<td id="23765DB6086800141519FBB40A96FBB2" box="[632,685,1077,1130]" gridcol="2" gridrow="4" pageId="3" pageNumber="367">16.3</td>
<td id="23765DB608680014159EFBB40B06FBB2" box="[767,829,1077,1130]" gridcol="3" gridrow="4" pageId="3" pageNumber="367">4.7</td>
<td id="23765DB60868001414E6FBB40BFAFBB2" box="[903,961,1077,1130]" gridcol="4" gridrow="4" pageId="3" pageNumber="367">5.5</td>
<td id="23765DB608680014136EFBB40C7DFBB2" box="[1039,1094,1077,1130]" gridcol="5" gridrow="4" pageId="3" pageNumber="367">15.8</td>
<td id="23765DB60868001413F6FBB40CEDFBB2" box="[1175,1238,1077,1130]" gridcol="6" gridrow="4" pageId="3" pageNumber="367">4.3</td>
<td id="23765DB608680014127FFBB40DBFFBB2" box="[1310,1412,1077,1130]" gridcol="7" gridrow="4" pageId="3" pageNumber="367">13.9</td>
</tr>
</table>
</paragraph>
<paragraph id="DE2836880868FFE817FEFBFC0B12FADC" blockId="3.[159,1412,1149,1326]" pageId="3" pageNumber="367">Males (n = 4) 3.1–3.9 4.6–4.9 14.9–16.3 4.5–4.7 5.0–5.5 15.8–15.9 - 13.6–14.6 (3.5) (4.8) (15.4) (4.6) (5.1) (15.9) (14.1) TIIIs TaIIIs Ias Ibs Oas Obs</paragraph>
<paragraph id="DE2836880868FFE817FEFA990BAFFAF6" blockId="3.[159,1412,1149,1326]" box="[159,916,1304,1326]" pageId="3" pageNumber="367">
<typeStatus id="012C882A0868FFE817FEFA9908C6FAF6" box="[159,253,1304,1326]" pageId="3" pageNumber="367" type="holotype">Holotype</typeStatus>
<date id="AA2910480868FFE81609FA990AA9FAF6" box="[360,658,1304,1326]" pageId="3" pageNumber="367" value="2013-02-09">9 2 13</date>
2–3 5
</paragraph>
<paragraph id="DE2836880868FFE817FEFAC00BAFFA3F" pageId="3" pageNumber="367">
<table id="AC97C4280868001417FEFAC00DBFF9DE" box="[159,1412,1345,1542]" gridcols="5" gridrows="3" pageId="3" pageNumber="367">
<tr id="60A734CA0868001417FEFAC00DBFFAAE" box="[159,1412,1345,1398]" gridrow="0" pageId="3" pageNumber="367">
<th id="23765DB60868001417FEFAC009B9FAAE" box="[159,386,1345,1398]" gridcol="0" gridrow="0" pageId="3" pageNumber="367">Male paratype 8 (15 Jun.2013)</th>
<th id="23765DB6086800141691FAC009C6FAAE" box="[496,509,1345,1398]" gridcol="1" gridrow="0" pageId="3" pageNumber="367">2</th>
<th id="23765DB6086800141519FAC00AA9FAAE" box="[632,658,1345,1398]" gridcol="2" gridrow="0" pageId="3" pageNumber="367">12</th>
<th id="23765DB608680014159EFAC00B1EFAAE" box="[767,805,1345,1398]" gridcol="3" gridrow="0" pageId="3" pageNumber="367">2–3</th>
<th id="23765DB60868001414E6FAC00DBFFAAE" box="[903,1412,1345,1398]" gridcol="4" gridrow="0" pageId="3" pageNumber="367">4</th>
</tr>
<tr id="60A734CA0868001417FEFA080DBFFA66" box="[159,1412,1417,1470]" gridrow="1" pageId="3" pageNumber="367">
<th id="23765DB60868001417FEFA0809B9FA66" box="[159,386,1417,1470]" gridcol="0" gridrow="1" pageId="3" pageNumber="367">Male paratype 8 (25 Jun.2013)</th>
<td id="23765DB6086800141691FA0809C6FA66" box="[496,509,1417,1470]" gridcol="1" gridrow="1" pageId="3" pageNumber="367">2</td>
<td id="23765DB6086800141519FA080AA9FA66" box="[632,658,1417,1470]" gridcol="2" gridrow="1" pageId="3" pageNumber="367">10</td>
<td id="23765DB608680014159EFA080B1EFA66" box="[767,805,1417,1470]" gridcol="3" gridrow="1" pageId="3" pageNumber="367">2–3</td>
<td id="23765DB60868001414E6FA080DBFFA66" box="[903,1412,1417,1470]" gridcol="4" gridrow="1" pageId="3" pageNumber="367">3</td>
</tr>
<tr id="60A734CA0868001417FEFA500DBFF9DE" box="[159,1412,1489,1542]" gridrow="2" pageId="3" pageNumber="367">
<th id="23765DB60868001417FEFA5009B9F9DE" box="[159,386,1489,1542]" gridcol="0" gridrow="2" pageId="3" pageNumber="367">Male paratype 10 (6 Jul.2013)</th>
<td id="23765DB6086800141691FA5009C6F9DE" box="[496,509,1489,1542]" gridcol="1" gridrow="2" pageId="3" pageNumber="367">2</td>
<td id="23765DB6086800141519FA500AA9F9DE" box="[632,658,1489,1542]" gridcol="2" gridrow="2" pageId="3" pageNumber="367">13</td>
<td id="23765DB608680014159EFA500B1EF9DE" box="[767,805,1489,1542]" gridcol="3" gridrow="2" pageId="3" pageNumber="367">2</td>
<td id="23765DB60868001414E6FA500DBFF9DE" box="[903,1412,1489,1542]" gridcol="4" gridrow="2" pageId="3" pageNumber="367">3</td>
</tr>
</table>
</paragraph>
<paragraph id="DE2836880868FFE817FEF9980BD8F9F7" blockId="3.[159,995,1561,1583]" box="[159,995,1561,1583]" pageId="3" pageNumber="367">Males (n = 4) 8–10 (8.8) 2 (2) 10–13 (12) - 3–5 (3.8)</paragraph>
<paragraph id="DE2836880868FFE817F6F9D10A7FF9BF" blockId="3.[151,580,1616,1639]" box="[151,580,1616,1639]" pageId="3" pageNumber="367">
*
<collectingCountry id="A68076180868FFE817CAF9D108DBF9BF" box="[171,224,1616,1639]" name="India" pageId="3" pageNumber="367">Hind</collectingCountry>
femur shrunk upon preservation.
</paragraph>
</subSubSection>
<subSubSection id="968D65030868FFE817A6F90E0991F928" pageId="3" pageNumber="367" type="etymology">
<paragraph id="DE2836880868FFE817A6F90E0991F928" blockId="3.[151,1436,1679,1885]" pageId="3" pageNumber="367">
<emphasis id="ECE3EA9A0868FFE817A6F90E0975F970" bold="true" box="[199,334,1679,1704]" pageId="3" pageNumber="367">Etymology.</emphasis>
The name reflects the surprise to discover a new species from the non-reserve parkland in a highly urbanised
<collectingCountry id="A68076180868FFE8166DF93509BAF915" box="[268,385,1716,1741]" name="Singapore" pageId="3" pageNumber="367">Singapore</collectingCountry>
. The name also coincide with part of the name the
<typeStatus id="012C882A0868FFE814A4F9340BCEF915" box="[965,1013,1717,1741]" pageId="3" pageNumber="367">type</typeStatus>
locality, Admiralty Park; from Latin,
<taxonomicName id="19974D0B0868FFE817F6F956092EF928" box="[151,277,1751,1776]" class="Insecta" family="Haglotettigoniidae" genus="Cardiodactylus" kingdom="Animalia" order="Orthoptera" pageId="3" pageNumber="367" phylum="Arthropoda" rank="species" species="admirabilis">admirabilis</taxonomicName>
= surprising.
</paragraph>
</subSubSection>
<subSubSection id="968D65030868FFEC17A6F97D09DBF9C7" lastPageId="7" lastPageNumber="371" pageId="3" pageNumber="367" type="distribution">
<paragraph id="DE2836880868FFE817A6F97D0D6FF885" blockId="3.[151,1436,1679,1885]" pageId="3" pageNumber="367">
<emphasis id="ECE3EA9A0868FFE817A6F97D095BF8CD" bold="true" box="[199,352,1788,1813]" pageId="3" pageNumber="367">Distribution.</emphasis>
This species is so far known from Admiralty Park in the north of
<collectingCountry id="A68076180868FFE81303F97D0CE8F8CD" box="[1122,1235,1788,1813]" name="Singapore" pageId="3" pageNumber="367">Singapore</collectingCountry>
and not found in central areas (Bukit Timah Nature Reserve and Central Catchment Nature Reserve) in which
<taxonomicName id="19974D0B0868FFE81391F8A10932F884" class="Insecta" family="Haglotettigoniidae" genus="Cardiodactylus" kingdom="Animalia" order="Orthoptera" pageId="3" pageNumber="367" phylum="Arthropoda" rank="species" species="singapura">
<emphasis id="ECE3EA9A0868FFE81391F8A10932F884" italics="true" pageId="3" pageNumber="367">Cardiodactylus singapura</emphasis>
</taxonomicName>
exists. Conversely,
<taxonomicName id="19974D0B0868FFE81688F8C50B36F884" box="[489,781,1860,1884]" class="Insecta" family="Haglotettigoniidae" genus="Cardiodactylus" kingdom="Animalia" order="Orthoptera" pageId="3" pageNumber="367" phylum="Arthropoda" rank="species" species="singapura">
<emphasis id="ECE3EA9A0868FFE81688F8C50B36F884" box="[489,781,1860,1884]" italics="true" pageId="3" pageNumber="367">Cardiodactylus singapura</emphasis>
</taxonomicName>
was not sighted and recorded from Admiralty Park.
</paragraph>
<caption id="8AE86600086FFFEF17F6F80909C3F864" httpUri="https://zenodo.org/record/250852/files/figure.png" pageId="4" pageNumber="368" targetBox="[151,1436,193,1905]" targetPageId="4">
<paragraph id="DE283688086FFFEF17F6F80909C3F864" blockId="4.[151,1436,1928,1980]" pageId="4" pageNumber="368">
<emphasis id="ECE3EA9A086FFFEF17F6F8090929F845" bold="true" box="[151,274,1928,1950]" pageId="4" pageNumber="368">FIGURE 1.</emphasis>
<taxonomicName id="19974D0B086FFFEF1679F8090988F846" box="[280,435,1928,1950]" class="Insecta" family="Haglotettigoniidae" genus="Cardiodactylus" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="368" phylum="Arthropoda" rank="genus">
<emphasis id="ECE3EA9A086FFFEF1679F8090988F846" box="[280,435,1928,1950]" italics="true" pageId="4" pageNumber="368">Cardiodactylus</emphasis>
</taxonomicName>
adults recorded in Singapore:
<taxonomicName id="19974D0B086FFFEF1586F8090BC6F846" box="[743,1021,1928,1950]" class="Insecta" family="Haglotettigoniidae" genus="Cardiodactylus" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="368" phylum="Arthropoda" rank="species" species="admirabilis" status="sp. nov.">
<emphasis id="ECE3EA9A086FFFEF1586F8090BC6F846" box="[743,1021,1928,1950]" italics="true" pageId="4" pageNumber="368">Cardiodactylus admirabilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F7D057E1086FFFEF1363F8090C01F845" box="[1026,1082,1928,1949]" pageId="4" pageNumber="368" rank="species">
<emphasis id="ECE3EA9A086FFFEF1363F8090C01F845" bold="true" box="[1026,1082,1928,1949]" pageId="4" pageNumber="368">n. sp.</emphasis>
</taxonomicNameLabel>
(A) and
<taxonomicName id="19974D0B086FFFEF13F5F809090EF864" authority="Robillard, 2011" authorityName="Robillard" authorityYear="2011" class="Insecta" family="Haglotettigoniidae" genus="Cardiodactylus" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="368" phylum="Arthropoda" rank="species" species="singapura">
<emphasis id="ECE3EA9A086FFFEF13F5F8090DA7F846" box="[1172,1436,1928,1950]" italics="true" pageId="4" pageNumber="368">Cardiodactylus singapura</emphasis>
Robillard, 2011
</taxonomicName>
(B) in dorsal view.
</paragraph>
</caption>
<caption id="8AE86600086EFFEE17F6F83608FBF833" httpUri="https://zenodo.org/record/250853/files/figure.png" pageId="5" pageNumber="369" targetBox="[239,1332,197,1937]" targetPageId="5">
<paragraph id="DE283688086EFFEE17F6F83608FBF833" blockId="5.[151,1435,1975,2027]" pageId="5" pageNumber="369">
<emphasis id="ECE3EA9A086EFFEE17F6F836092FF814" bold="true" box="[151,276,1975,1997]" pageId="5" pageNumber="369">FIGURE 2.</emphasis>
<taxonomicName id="19974D0B086EFFEE167DF8360A08F815" box="[284,563,1975,1997]" class="Insecta" family="Haglotettigoniidae" genus="Cardiodactylus" kingdom="Animalia" order="Orthoptera" pageId="5" pageNumber="369" phylum="Arthropoda" rank="species" species="admirabilis" status="sp. nov.">
<emphasis id="ECE3EA9A086EFFEE167DF8360A08F815" box="[284,563,1975,1997]" italics="true" pageId="5" pageNumber="369">Cardiodactylus admirabilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F7D057E1086EFFEE155DF8360A4DF814" box="[572,630,1975,1996]" pageId="5" pageNumber="369" rank="species">
<emphasis id="ECE3EA9A086EFFEE155DF8360A4DF814" bold="true" box="[572,630,1975,1996]" pageId="5" pageNumber="369">n. sp.</emphasis>
</taxonomicNameLabel>
male habitus: dorsal (A) and lateral (B) views. Scale bars: 2 mm (A); 10 mm (B).
</paragraph>
</caption>
<paragraph id="DE283688086DFFED17F6F8FB0C18F86C" blockId="6.[151,1435,1914,1973]" pageId="6" pageNumber="370">
<emphasis id="ECE3EA9A086DFFED17F6F8FB092FF857" bold="true" box="[151,276,1914,1936]" pageId="6" pageNumber="370">FIGURE 3.</emphasis>
<taxonomicName id="19974D0B086DFFED167AF8FB0A08F848" box="[283,563,1914,1936]" class="Insecta" family="Haglotettigoniidae" genus="Cardiodactylus" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="370" phylum="Arthropoda" rank="species" species="admirabilis" status="sp. nov.">
<emphasis id="ECE3EA9A086DFFED167AF8FB0A08F848" box="[283,563,1914,1936]" italics="true" pageId="6" pageNumber="370">Cardiodactylus admirabilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F7D057E1086DFFED155BF8FA0A4EF848" box="[570,629,1915,1936]" pageId="6" pageNumber="370" rank="species">
<emphasis id="ECE3EA9A086DFFED155BF8FA0A4EF848" bold="true" box="[570,629,1915,1936]" pageId="6" pageNumber="370">n. sp.</emphasis>
</taxonomicNameLabel>
male: head in dorsal (A) and anterior (B) views; head and pronotum in lateral view (C); FW in dorsal view (D); stridulatory file (E). Scale bar: A–D,
<quantity id="196F9B6D086DFFED140AF81F0B92F86C" box="[875,937,1950,1972]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="370" unit="mm" value="1.0">1 mm</quantity>
; E,
<quantity id="196F9B6D086DFFED14AFF81F0C24F86C" box="[974,1055,1950,1972]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="6" pageNumber="370" unit="mm" value="0.5">0.5 mm</quantity>
.
</paragraph>
<caption id="8AE86600086CFFEC17F6FBB40DBCFB93" box="[151,1415,1077,1099]" httpUri="https://zenodo.org/record/250854/files/figure.png" pageId="7" pageNumber="371" targetBox="[157,1433,200,1044]" targetPageId="7">
<paragraph id="DE283688086CFFEC17F6FBB40DBCFB93" blockId="7.[151,1415,1077,1099]" box="[151,1415,1077,1099]" pageId="7" pageNumber="371">
<emphasis id="ECE3EA9A086CFFEC17F6FBB40928FB92" bold="true" box="[151,275,1077,1099]" pageId="7" pageNumber="371">FIGURE 4.</emphasis>
<taxonomicName id="19974D0B086CFFEC1678FBB40A14FB93" box="[281,559,1077,1099]" class="Insecta" family="Haglotettigoniidae" genus="Cardiodactylus" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="371" phylum="Arthropoda" rank="species" species="admirabilis" status="sp. nov.">
<emphasis id="ECE3EA9A086CFFEC1678FBB40A14FB93" box="[281,559,1077,1099]" italics="true" pageId="7" pageNumber="371">Cardiodactylus admirabilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F7D057E1086CFFEC1554FBB70A54FB93" box="[565,623,1078,1099]" pageId="7" pageNumber="371" rank="species">
<emphasis id="ECE3EA9A086CFFEC1554FBB70A54FB93" bold="true" box="[565,623,1078,1099]" pageId="7" pageNumber="371">n. sp.</emphasis>
</taxonomicNameLabel>
male genitalia: dorsal (A); ventral (B) and lateral (C) views. Scale bar: 1 mm.
</paragraph>
</caption>
<paragraph id="DE283688086CFFEC17A6FBF90981FB01" blockId="7.[151,1436,1144,1639]" pageId="7" pageNumber="371">
<emphasis id="ECE3EA9A086CFFEC17A6FBF90A20FB49" bold="true" box="[199,539,1144,1169]" pageId="7" pageNumber="371">Habit and life history traits.</emphasis>
<taxonomicName id="19974D0B086CFFEC1545FBF90B62FB48" box="[548,857,1144,1168]" class="Insecta" family="Haglotettigoniidae" genus="Cardiodactylus" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="371" phylum="Arthropoda" rank="species" species="admirabilis" status="sp. nov.">
<emphasis id="ECE3EA9A086CFFEC1545FBF90B62FB48" box="[548,857,1144,1168]" italics="true" pageId="7" pageNumber="371">Cardiodactylus admirabilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F7D057E1086CFFEC1400FBF90B99FB49" box="[865,930,1144,1169]" pageId="7" pageNumber="371" rank="species">
<emphasis id="ECE3EA9A086CFFEC1400FBF90B99FB49" bold="true" box="[865,930,1144,1169]" pageId="7" pageNumber="371">n. sp.</emphasis>
</taxonomicNameLabel>
is a nocturnal species, although the nymphs are sometimes sighted in the day. In captivity, the adults were fed with
<taxonomicName id="19974D0B086CFFEC14D4FB1D0C2AFB6C" box="[949,1041,1180,1204]" class="Magnoliopsida" family="Dilleniaceae" genus="Dillenia" kingdom="Plantae" order="Dilleniales" pageId="7" pageNumber="371" phylum="Tracheophyta" rank="genus">
<emphasis id="ECE3EA9A086CFFEC14D4FB1D0C2AFB6C" box="[949,1041,1180,1204]" italics="true" pageId="7" pageNumber="371">Dillenia</emphasis>
</taxonomicName>
fruits, flowers and leaves in which the species may be found.
</paragraph>
<paragraph id="DE283688086CFFEC17A6FB6509DBF9C7" blockId="7.[151,1436,1144,1639]" pageId="7" pageNumber="371">
<emphasis id="ECE3EA9A086CFFEC17A6FB650973FB25" bold="true" box="[199,328,1252,1277]" pageId="7" pageNumber="371">Behaviour.</emphasis>
Calling song (
<figureCitation id="46AC2A0D086CFFEC168FFB650A05FB25" box="[494,574,1252,1277]" captionStart="FIGURE 6" captionStartId="9.[151,250,1655,1677]" captionTargetBox="[157,1431,219,1618]" captionTargetId="figure@9.[151,1436,207,1626]" captionTargetPageId="9" captionText="FIGURE 6. Cardiodactylus admirabilis n. sp. calling song: oscillogram of four echemes (A); sonogram of four echemes (B); oscillogram of one echeme (C). Symbols: f 1, f 2, f 3 correspond to the first, second and third harmonic frequencies in the spectrum." httpUri="https://zenodo.org/record/250856/files/figure.png" pageId="7" pageNumber="371">Figs. 6</figureCitation>
,
<figureCitation id="46AC2A0D086CFFEC152AFB650A67FB24" box="[587,604,1252,1276]" captionStart="FIGURE 7" captionStartId="10.[151,250,1916,1938]" captionTargetBox="[159,1428,198,1891]" captionTargetId="figure@10.[150,1436,193,1895]" captionTargetPageId="10" captionText="FIGURE 7. Cardiodactylus admirabilis n. sp. calling song: oscillograms (A, C, E) and power spectrum (B, D, F) of syllable 1 (A, B), syllable 2 (C, D) and syllable 3 (E, F). Symbols: f 1, f 2, f 3 correspond to the first, second and third harmonic frequencies in the spectrum." httpUri="https://zenodo.org/record/250857/files/figure.png" pageId="7" pageNumber="371">7</figureCitation>
): The calling song of
<taxonomicName id="19974D0B086CFFEC1435FB650CB2FB24" box="[852,1161,1252,1276]" class="Insecta" family="Haglotettigoniidae" genus="Cardiodactylus" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="371" phylum="Arthropoda" rank="species" species="admirabilis" status="sp. nov.">
<emphasis id="ECE3EA9A086CFFEC1435FB650CB2FB24" box="[852,1161,1252,1276]" italics="true" pageId="7" pageNumber="371">Cardiodactylus admirabilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F7D057E1086CFFEC13F0FB650CEBFB25" box="[1169,1232,1252,1277]" pageId="7" pageNumber="371" rank="species">
<emphasis id="ECE3EA9A086CFFEC13F0FB650CEBFB25" bold="true" box="[1169,1232,1252,1277]" pageId="7" pageNumber="371">n. sp.</emphasis>
</taxonomicNameLabel>
is a short echeme made of a triplet of syllables emitted by groups of 3 or 4, unlike the other
<taxonomicName id="19974D0B086CFFEC1484FA890CAAFAF8" box="[997,1169,1288,1312]" class="Insecta" family="Haglotettigoniidae" genus="Cardiodactylus" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="371" phylum="Arthropoda" rank="genus">
<emphasis id="ECE3EA9A086CFFEC1484FA890CAAFAF8" box="[997,1169,1288,1312]" italics="true" pageId="7" pageNumber="371">Cardiodactylus</emphasis>
</taxonomicName>
species described from the region, including
<taxonomicName id="19974D0B086CFFEC16ECFAAD0A19FA9C" box="[397,546,1324,1348]" class="Insecta" family="Haglotettigoniidae" genus="Cardiodactylus" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="371" phylum="Arthropoda" rank="species" species="singapura">
<emphasis id="ECE3EA9A086CFFEC16ECFAAD0A19FA9C" box="[397,546,1324,1348]" italics="true" pageId="7" pageNumber="371">C. singapura</emphasis>
</taxonomicName>
, which have mono-syllabic calling songs (
<bibRefCitation id="BA064B79086CFFEC136DFAAD08F5FAB0" author="Robillard" pageId="7" pageNumber="371" refString="Robillard, T. (2011) New Cardiodactylus species from unsuspected places in Southeast Asia (Orthoptera, Grylloidea, Eneopterinae). Zootaxa, 2909, 14 - 26." type="journal article" year="2011">
Robillard, 2011; Robillard
<emphasis id="ECE3EA9A086CFFEC1222FAAC0D46FA9C" box="[1347,1405,1324,1348]" italics="true" pageId="7" pageNumber="371">et al.</emphasis>
in prep
</bibRefCitation>
). The calling song has the following features: echeme duration = 205.6±7 ms, echeme period = 2.11±1.3 s, echeme duty cycle = 11.7±4.4 %; syllable duration = 36.9±3.9 ms, syllable period = 84.8±0.8 ms, syllable duty cycle = 43.5 %; dominant frequency (3rd peak of the spectrum) = 14.26 0.74 kHz. The syllables within the echeme tend to be slightly different, as shown in
<tableCitation id="93150333086CFFEC1500FA3F0A8CFA0F" box="[609,695,1470,1495]" captionStart="TABLE 2" captionStartId="7.[151,239,1687,1710]" captionTargetBox="[159,1300,1771,1947]" captionTargetPageId="7" captionText="TABLE 2. Measurements of syllable features within the calling song of Cardiodactylus admirabilis n. sp. (based on measurement of 5 echemes)." httpUri="http://table.plazi.org/id/8AE86600086CFFEC17F6F91609F2F917" pageId="7" pageNumber="371" tableUuid="8AE86600086CFFEC17F6F91609F2F917">Table 2</tableCitation>
: the frequency increases by more than 1.5 kHz between syllable 1 an d syllable 3; syllable 2 tend to be longer then the others, and syllables 1 and 2 include gaps, while syllable 3 is continuous and more intense.
</paragraph>
</subSubSection>
<subSubSection id="968D6503086CFFEC17A6F9AB0A99F9BF" pageId="7" pageNumber="371" type="discussion">
<paragraph id="DE283688086CFFEC17A6F9AB0A99F9BF" blockId="7.[151,1436,1144,1639]" pageId="7" pageNumber="371">
<emphasis id="ECE3EA9A086CFFEC17A6F9AB0977F99B" bold="true" box="[199,332,1578,1603]" pageId="7" pageNumber="371">Discussion.</emphasis>
The nymphs of this species were previously sighted in Admiralty Park and wrongly recorded in Tan
<emphasis id="ECE3EA9A086CFFEC17A6F9D1093BF9BF" box="[199,256,1615,1639]" italics="true" pageId="7" pageNumber="371">et al.</emphasis>
(2012) as
<taxonomicName id="19974D0B086CFFEC1617F9CE0AA0F9BF" box="[374,667,1615,1639]" class="Insecta" family="Haglotettigoniidae" genus="Cardiodactylus" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="371" phylum="Arthropoda" rank="species" species="singapura">
<emphasis id="ECE3EA9A086CFFEC1617F9CE0AA0F9BF" box="[374,667,1615,1639]" italics="true" pageId="7" pageNumber="371">Cardiodactylus singapura</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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