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<document ID-DOI="http://dx.doi.org/10.3897/mycokeys.46.32823" ID-GBIF-Dataset="e546c2a6-4668-4253-b2e2-b462f23dab8c" ID-PMC="PMC6379322" ID-Pensoft-Pub="1314-4049-46-1" ID-PubMed="30787668" ModsDocAuthor="" ModsDocDate="2019" ModsDocID="1314-4049-46-1" ModsDocOrigin="MycoKeys 46" ModsDocTitle="The genus Hebeloma in the Rocky Mountain Alpine Zone" checkinTime="1555333436509" checkinUser="pensoft" docAuthor="Cripps, Cathy L., Eberhardt, Ursula, Schuetz, Nicole, Beker, Henry J., Vera S. Evenson, &amp; Horak, Egon" docDate="2019" docId="13BA04743C27C6E5308D1C62D7220913" docLanguage="en" docName="MycoKeys 46: 1-54" docOrigin="MycoKeys 46" docSource="http://dx.doi.org/10.3897/mycokeys.46.32823" docTitle="Hebeloma subconcolor Bruchet, Bull. Mens. Soc. Linn. Lyon 39 (6, suppl.): 127 1970" docType="treatment" docVersion="4" lastPageNumber="17" masterDocId="AC18AA047D78FFBFD06BFFFDFF948577" masterDocTitle="The genus Hebeloma in the Rocky Mountain Alpine Zone" masterLastPageNumber="54" masterPageNumber="1" pageNumber="17" updateTime="1668136151891" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>The genus Hebeloma in the Rocky Mountain Alpine Zone</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Cripps, Cathy L.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Eberhardt, Ursula</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Schuetz, Nicole</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Beker, Henry J.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Vera S. Evenson,</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Horak, Egon</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>MycoKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2019</mods:date>
<mods:detail type="volume">
<mods:number>46</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>54</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/mycokeys.46.32823</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/mycokeys.46.32823</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-4049-46-1</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="156202112" LSID="urn:lsid:plazi:treatment:13BA04743C27C6E5308D1C62D7220913" httpUri="http://treatment.plazi.org/id/13BA04743C27C6E5308D1C62D7220913" lastPageNumber="17" pageId="16" pageNumber="17">
<subSubSection pageId="16" pageNumber="17" type="nomenclature">
<paragraph pageId="16" pageNumber="17">
3.
<taxonomicName authority="Bruchet, Bull. Mens. Soc. Linn. Lyon 39 (6, suppl.): 127 (1970)" class="Agaricomycetes" family="Hymenogastraceae" genus="Hebeloma" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Hebeloma subconcolor" order="Agaricales" pageId="16" pageNumber="17" phylum="Basidiomycota" rank="species" species="subconcolor">Hebeloma subconcolor Bruchet, Bull. Mens. Soc. Linn. Lyon 39 (6, suppl.): 127 (1970)</taxonomicName>
Figures 5, 9, 23 (3)
</paragraph>
</subSubSection>
<subSubSection pageId="16" pageNumber="17" type="etymology">
<paragraph pageId="16" pageNumber="17">Etymology.</paragraph>
<paragraph pageId="16" pageNumber="17">concolor for the similar coloration of pileus and stipe, which is not a consistent feature.</paragraph>
</subSubSection>
<subSubSection pageId="16" pageNumber="17" type="description">
<paragraph pageId="16" pageNumber="17">Description.</paragraph>
<paragraph pageId="16" pageNumber="17">
Cortina absent. Pileus 15-20 mm, convex, with or without a low broad umbo, becoming plane, smooth, moist, light to medium brown, pruinose with a grayish tint or sheen, lighter towards margin but not distinctly two-toned; margin turned down or not, entire. Lamellae adnexed, subdistant, well-separated, medium broad to broad, L = 25-32 plus lamellulae, dull brown, light brown; edges lighter. No beaded drops reported. Stipe 15-30
<normalizedToken originalValue="×">x</normalizedToken>
3-4 mm, equal, apex somewhat lighter tan and pruinose, below totally covered with longitudinal white fibers over a brownish ground base. Context buff. Odor astringent. Exsiccate: pileus medium brown, not two-toned, with grayish tint, dull; lamellae broad, warm cinnamon; stipe long, dull brown, narrow.
</paragraph>
<paragraph pageId="16" pageNumber="17">
Basidiospores yellowish brown, amygdaliform, with a small apiculus, weakly ornamented (O2), loosening perispore observed in a few spores (P0, P1), distinctly dextrinoid (D2, D3), 10.5-12.5
<normalizedToken originalValue="×">x</normalizedToken>
6.5-7.5
<normalizedToken originalValue="µm">µm</normalizedToken>
, on average 11.6
<normalizedToken originalValue="×">x</normalizedToken>
7.1
<normalizedToken originalValue="µm">µm</normalizedToken>
, Q = 1.65. Basidia 25-34
<normalizedToken originalValue="×">x</normalizedToken>
8-10
<normalizedToken originalValue="µm">µm</normalizedToken>
, four-spored. Cheilocystidia gently clavate, some slightly swollen at apex and base, 40-60
<normalizedToken originalValue="×">x</normalizedToken>
6-11
<normalizedToken originalValue="µm">µm</normalizedToken>
at apex, 4.5-7
<normalizedToken originalValue="µm">µm</normalizedToken>
in middle, and 4-7 - (8)
<normalizedToken originalValue="µm">µm</normalizedToken>
at base. Pleurocystidia absent. Epicutis thickness 60-75
<normalizedToken originalValue="µm">µm</normalizedToken>
, with some encrusted hyphae.
</paragraph>
<caption pageId="16" pageNumber="17">
<paragraph pageId="16" pageNumber="17">
Figure 9.
<taxonomicName class="Agaricomycetes" family="Hymenogastraceae" genus="Hebeloma" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Hebeloma subconcolor" order="Agaricales" pageId="16" pageNumber="17" phylum="Basidiomycota" rank="species" species="subconcolor">Hebeloma subconcolor</taxonomicName>
, DBG-F-022785 and DBG-F-022786.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="16" pageNumber="17" type="rocky mountain ecology">
<paragraph pageId="16" pageNumber="17">Rocky Mountain ecology.</paragraph>
<paragraph pageId="16" pageNumber="17">Two collections reported under willow at alpine elevations of 4000 m in Colorado; noted as cespitose to gregarious.</paragraph>
</subSubSection>
<subSubSection pageId="16" pageNumber="17" type="materials_examined">
<paragraph pageId="16" pageNumber="17">Rocky Mountain specimens examined.</paragraph>
<paragraph pageId="16" pageNumber="17">
U.S.A. COLORADO: Clear Creek County, Summit Lake Park, under
<taxonomicName class="Ascidiacea" family="Polycitoridae" genus="Salix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Salix" order="Aplousobranchia" pageId="16" pageNumber="17" phylum="Chordata" rank="genus">Salix</taxonomicName>
, some in moss, at 4000 m, 22 Aug 2012, DBG-F-022785; DBG-F-022786, L. Gillman.
</paragraph>
</subSubSection>
<subSubSection pageId="16" pageNumber="17" type="discussion">
<paragraph pageId="16" pageNumber="17">Discussion.</paragraph>
<paragraph pageId="16" pageNumber="17">
The sequences of the two collections for
<taxonomicName lsidName="H. subconcolor" pageId="16" pageNumber="17" rank="species" species="subconcolor">H. subconcolor</taxonomicName>
from the Rocky Mountains are identical. The RM sequence differs by 1-4 [0] bp from the
<taxonomicName lsidName="H. subconcolor" pageId="16" pageNumber="17" rank="species" species="subconcolor">H. subconcolor</taxonomicName>
collections described in
<bibRefCitation pageId="16" pageNumber="17">Beker et al. (2016)</bibRefCitation>
and
<bibRefCitation author="Grilli, E" journalOrPublisher="Mycological Progress" pageId="44" pageNumber="45" pagination="1 - 46" title="Unexpected species diversity and contrasting evolutionary hypotheses in Hebeloma (Agaricales) sections Sinapizantia and Velutipes in Europe." url="https://doi.org/10.1007/s11557-015-1148-6" volume="15" year="2016">Grilli et al. (2016)</bibRefCitation>
, where the ITS ML results were also shown. The closest
<taxonomicName lsidName="H. velutipes" pageId="16" pageNumber="17" rank="species" species="velutipes">H. velutipes</taxonomicName>
sequence included in the dataset used in Fig. 5 differs in 3 [0] bp.
<taxonomicName class="Agaricomycetes" family="Hymenogastraceae" genus="Hebeloma" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Hebeloma velutipes" order="Agaricales" pageId="16" pageNumber="17" phylum="Basidiomycota" rank="species" species="velutipes">Hebeloma velutipes</taxonomicName>
is the only species of
<taxonomicName class="Agaricomycetes" family="Hymenogastraceae" genus="Hebeloma" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Hebeloma" order="Agaricales" pageId="16" pageNumber="17" phylum="Basidiomycota" rank="genus">Hebeloma</taxonomicName>
that cannot be distinguished from
<taxonomicName lsidName="H. subconcolor" pageId="16" pageNumber="17" rank="species" species="subconcolor">H. subconcolor</taxonomicName>
by ITS sequence (
<bibRefCitation pageId="16" pageNumber="17">Beker et al. 2016</bibRefCitation>
;
<bibRefCitation author="Grilli, E" journalOrPublisher="Mycological Progress" pageId="44" pageNumber="45" pagination="1 - 46" title="Unexpected species diversity and contrasting evolutionary hypotheses in Hebeloma (Agaricales) sections Sinapizantia and Velutipes in Europe." url="https://doi.org/10.1007/s11557-015-1148-6" volume="15" year="2016">Grilli et al. 2016</bibRefCitation>
; Fig. 5). However, morphologically these two species are very different and can be easily separated.
</paragraph>
<paragraph pageId="16" pageNumber="17">
This small species has a grayish cast not found in other taxa in sections
<taxonomicName class="Magnoliopsida" family="Cactaceae" genus="Denudata" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Denudata" order="Caryophyllales" pageId="16" pageNumber="17" phylum="Tracheophyta" rank="genus">Denudata</taxonomicName>
and
<taxonomicName genus="Velutipes" lsidName="Velutipes" pageId="16" pageNumber="17" rank="genus">Velutipes</taxonomicName>
that we report from the Rocky Mountains; also, the lamellae are well separated and few in number. It should be compared to the other non-veiled, small species such as
<taxonomicName lsidName="H. aurantioumbrinum" pageId="16" pageNumber="17" rank="species" species="aurantioumbrinum">H. aurantioumbrinum</taxonomicName>
and
<taxonomicName lsidName="H. vaccinum" pageId="16" pageNumber="17" rank="species" species="vaccinum">H. vaccinum</taxonomicName>
.
<taxonomicName class="Agaricomycetes" family="Hymenogastraceae" genus="Hebeloma" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Hebeloma velutipes" order="Agaricales" pageId="16" pageNumber="17" phylum="Basidiomycota" rank="species" species="velutipes">Hebeloma velutipes</taxonomicName>
has a different coloration and is larger with many more full length lamellae.
<taxonomicName class="Agaricomycetes" family="Hymenogastraceae" genus="Hebeloma" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Hebeloma subconcolor" order="Agaricales" pageId="16" pageNumber="17" phylum="Basidiomycota" rank="species" species="subconcolor">Hebeloma subconcolor</taxonomicName>
is known from arctic and alpine locations in the European Alps, Greenland, Iceland and Scandinavia (
<bibRefCitation pageId="16" pageNumber="17">Beker et al. 2016</bibRefCitation>
,
<bibRefCitation pageId="16" pageNumber="17">2018</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>