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<document id="849BFA585C2E4A6A13C68601B00812B1" ID-CLB-Dataset="300549" ID-DOI="10.1590/S1984-4689.v39.e22042" ID-GBIF-Dataset="ec400598-f3ab-464c-9507-2e9039e7749a" ID-ISSN="1984-4689" ID-Zenodo-Dep="13173808" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722587011155" checkinUser="felipe" docAuthor="Li, Yan-Da, Huang, Di-Ying &amp; Cai, Chen-Yang" docDate="2022" docId="03F287ACE703126E92F9FB5CFC44B3C1" docLanguage="en" docName="Zoologia.39.e22042.pdf" docOrigin="Zoologia (e 22042) 39" docSource="http://dx.doi.org/10.1590/s1984-4689.v39.e22042" docStyle="DocumentStyle:C0DEAC3FDCE12EEF877F5538032D0DC0.6:Zoologia.2021-.journal_article" docStyleId="C0DEAC3FDCE12EEF877F5538032D0DC0" docStyleName="Zoologia.2021-.journal_article" docStyleVersion="6" docTitle="Delteredolaemus hei Li &amp; Huang &amp; Cai 2022, sp. nov." docType="treatment" docVersion="2" lastPageNumber="2" masterDocId="FFCBFFD4E702126F9215FFB9FFC5B754" masterDocTitle="Earliest teredid beetle from mid-Cretaceous amber of northern Myanmar (Coleoptera: Coccinelloidea: Teredidae): new genus and species" masterLastPageNumber="9" masterPageNumber="1" pageNumber="2" updateTime="1722619504406" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="3F63F5FCB7232B2DABBC425164C62457">Earliest teredid beetle from mid-Cretaceous amber of northern Myanmar (Coleoptera: Coccinelloidea: Teredidae): new genus and species</mods:title>
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<mods:namePart id="2AD0B588584EB42A4A21929DFB51DAA2">Huang, Di-Ying</mods:namePart>
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<taxonomicName id="4C5B4D39E703126E92F9FB5CFD83B254" authority="Li &amp; Cai" authorityName="Li &amp; Huang &amp; Cai" authorityYear="2022" box="[236,582,1253,1280]" class="Insecta" family="Teredidae" genus="Delteredolaemus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="hei" status="sp. nov.">
<emphasis id="B92FEAA8E703126E92F9FB5CFE1EB254" box="[236,475,1253,1280]" italics="true" pageId="1" pageNumber="2">Delteredolaemus hei</emphasis>
Li &amp; Cai
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<taxonomicNameLabel id="A21C57D3E703126E905BFB5CFD6BB254" box="[590,686,1253,1280]" pageId="1" pageNumber="2" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="8BE436BAE703126E93B4FAB6FE3CB277" blockId="1.[417,505,1295,1315]" box="[417,505,1295,1315]" pageId="1" pageNumber="2">
<figureCitation id="13602A3FE703126E93B4FAB6FE3CB277" box="[417,505,1295,1315]" captionStart-0="Figures 12" captionStart-1="Figures 34" captionStart-2="Figures 56" captionStart-3="Figures 712" captionStart-4="Figures 1317" captionStartId-0="2.[159,226,1730,1750]" captionStartId-1="3.[151,218,1733,1753]" captionStartId-2="4.[159,226,1733,1753]" captionStartId-3="5.[151,218,1068,1088]" captionStartId-4="6.[159,226,1698,1718]" captionTargetBox-0="[185,1396,228,1712]" captionTargetBox-1="[151,1428,197,1716]" captionTargetBox-2="[159,1436,197,1716]" captionTargetBox-3="[151,1428,197,1054]" captionTargetBox-4="[253,1342,197,1686]" captionTargetId-0="figure-27@2.[159,1436,194,1715]" captionTargetId-1="figure-27@3.[151,1428,197,1719]" captionTargetId-2="figure-31@4.[159,1436,194,1719]" captionTargetId-3="figure-383@5.[151,1428,197,1054]" captionTargetId-4="figure-43@6.[253,1342,197,1686]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionTargetPageId-4="6" captionText-0="Figures 12. General habitus of Delteredolaemus hei gen. et sp. nov., holotype, NIGP200000, under incident light: (1) dorsal view; (2) ventral view. Scale bars: 500 Μm." captionText-1="Figures 34. General habitus of Delteredolaemus hei gen. et sp. nov., holotype, NIGP200000, under confocal microscopy: (3) dorsal view; (4) ventral view. Scale bars: 500 Μm." captionText-2="Figures 56. General habitus of Delteredolaemus hei gen. et sp. nov., holotype, NIGP200000, under confocal microscopy, with depth color coding: (5) dorsal view; (6) ventral view. Scale bars: 500 Μm." captionText-3="Figures 712. Details of Delteredolaemus hei gen. et sp. nov., holotype, NIGP200000, under confocal microscopy: (7) head, anteroventral view; (8) prothorax, ventral view; (9) mesothorax, ventral view; (10) abdominal base, ventral view; (11) abdominal apex, ventral view; (12) pronotal and elytral bases, dorsal view, with arrowhead showing the medially produced posterior pronotal edge. (a1) antennomere 1, (el) elytron, (lbp) labial palp, (md) mandible, (msf) mesofemur, (mstb) mesotibia, (msv) mesoventrite, (mtf) metafemur, (mtv) metaventrite, (mxp) maxillary palp, (pc) procoxa, (pf) profemur, (pn) pronotum, (ptb) protibia, (ptc) protrochanter, (sc) scutellum, (ts) tibial spine, (v15) ventrites 15. Scale bars: 100 Μm." captionText-4="Figures 1317. X-ray microtomographic reconstruction of Delteredolaemus hei gen. et sp. nov., holotype, NIGP200000: (13) dorsal view; (14) ventral view; (15) lateral view; (16) anterodorsal view; (17) head, ventrolateral view. (a111) antennomeres 111. Scale bars: 500 Μm in (1316), 200 Μm in (17)." figureDoi-0="http://doi.org/10.5281/zenodo.13173810" figureDoi-1="http://doi.org/10.5281/zenodo.13173812" figureDoi-2="http://doi.org/10.5281/zenodo.13173814" figureDoi-3="http://doi.org/10.5281/zenodo.13173816" figureDoi-4="http://doi.org/10.5281/zenodo.13173818" httpUri-0="https://zenodo.org/record/13173810/files/figure.png" httpUri-1="https://zenodo.org/record/13173812/files/figure.png" httpUri-2="https://zenodo.org/record/13173814/files/figure.png" httpUri-3="https://zenodo.org/record/13173816/files/figure.png" httpUri-4="https://zenodo.org/record/13173818/files/figure.png" pageId="1" pageNumber="2">Figs 117</figureCitation>
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<paragraph id="8BE436BAE703126E928CFA87FCC7B206" blockId="1.[153,770,1342,1362]" box="[153,770,1342,1362]" pageId="1" pageNumber="2">
https://zoobank.org/
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<materialsCitation id="3B333CE7E703126E92C1FADAFDE4B223" box="[212,545,1379,1399]" collectionCode="NIGP" pageId="1" pageNumber="2" specimenCode="NIGP200000" specimenCount="1" typeStatus="holotype">
Material.
<typeStatus id="54E08818E703126E9327FADAFE54B223" box="[306,401,1379,1399]" pageId="1" pageNumber="2" type="holotype">Holotype</typeStatus>
,
<specimenCode id="DBFD9EC1E703126E938FFADDFDD8B223" box="[410,541,1380,1399]" collectionCode="NIGP" country="China" httpUri="http://grbio.org/cool/j51y-avsb" name="Naking Institute of Geology and Palaeontology" pageId="1" pageNumber="2">NIGP200000</specimenCode>
.
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</paragraph>
<paragraph id="8BE436BAE703126E92C1FA38FD62B2BA" blockId="1.[150,773,1379,1787]" pageId="1" pageNumber="2">
Locality and horizon. Amber mine located near Noije Bum Village, Tanai Township, Myitkyina District,
<collectingRegion id="499FF858E703126E9097FA26FD3AB2E6" box="[642,767,1439,1458]" country="Myanmar" name="Kachin" pageId="1" pageNumber="2">Kachin State</collectingRegion>
,
<collectingCountry id="F34C762AE703126E9283FA04FF3CB284" box="[150,249,1469,1488]" name="Myanmar" pageId="1" pageNumber="2">Myanmar</collectingCountry>
; unnamed horizon, mid-Cretaceous (Upper Albian to Lower Cenomanian;
<bibRefCitation id="EFCA4B4BE703126E9373FA62FE30B2BA" author="Shi G &amp; Grimaldi DA &amp; Harlow GE &amp; Wang J &amp; Wang J &amp; Yang M &amp; Lei W &amp; Li Q &amp; Li X" box="[358,501,1499,1518]" pageId="1" pageNumber="2" pagination="155 - 163" refId="ref3730" refString="Shi G, Grimaldi DA, Harlow GE, Wang J, Wang J, Yang M, Lei W, Li Q, Li X (2012) Age constraint on Burmese amber based on U - Pb dating of zircons. Cretaceous Research 37: 155 - 163." type="journal article" year="2012">Shi et al. 2012</bibRefCitation>
,
<bibRefCitation id="EFCA4B4BE703126E93EAFA62FD58B2BA" author="Mao Y &amp; Liang K &amp; Su Y &amp; Li J &amp; Rao X &amp; Zhang H &amp; Xia F &amp; Fu Y &amp; Cai C &amp; Huang D" box="[511,669,1499,1518]" pageId="1" pageNumber="2" pagination="91 - 103" refId="ref3333" refString="Mao Y, Liang K, Su Y, Li J, Rao X, Zhang H, Xia F, Fu Y, Cai C, Huang D (2018) Various amberground marine animals on Burmese amber with discussions on its age. Palaeoentomology 1: 91 - 103." type="journal article" year="2018">Mao et al. 2018</bibRefCitation>
).
</paragraph>
<paragraph id="8BE436BAE703126E92C1FA41FE2DB158" blockId="1.[150,773,1379,1787]" box="[212,488,1528,1548]" pageId="1" pageNumber="2">Diagnosis. As for the genus.</paragraph>
<paragraph id="8BE436BAE703126E92C1F9AEFE9AB132" blockId="1.[150,773,1379,1787]" pageId="1" pageNumber="2">
Description. Body elongate, cylindrical, about
<quantity id="4CA39B5FE703126E90BDF9AEFCC6B17E" box="[680,771,1559,1578]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" pageId="1" pageNumber="2" unit="mm" value="1.75">1.75 mm</quantity>
long,
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wide, widest in anterior part of abdomen; surface with scattered setae.
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Head (
<figureCitation id="13602A3FE703126E9301F9C9FE8EB1D7" box="[276,331,1648,1668]" captionStart="Figures 712" captionStartId="5.[151,218,1068,1088]" captionTargetBox="[151,1428,197,1054]" captionTargetId="figure-383@5.[151,1428,197,1054]" captionTargetPageId="5" captionText="Figures 712. Details of Delteredolaemus hei gen. et sp. nov., holotype, NIGP200000, under confocal microscopy: (7) head, anteroventral view; (8) prothorax, ventral view; (9) mesothorax, ventral view; (10) abdominal base, ventral view; (11) abdominal apex, ventral view; (12) pronotal and elytral bases, dorsal view, with arrowhead showing the medially produced posterior pronotal edge. (a1) antennomere 1, (el) elytron, (lbp) labial palp, (md) mandible, (msf) mesofemur, (mstb) mesotibia, (msv) mesoventrite, (mtf) metafemur, (mtv) metaventrite, (mxp) maxillary palp, (pc) procoxa, (pf) profemur, (pn) pronotum, (ptb) protibia, (ptc) protrochanter, (sc) scutellum, (ts) tibial spine, (v15) ventrites 15. Scale bars: 100 Μm." figureDoi="http://doi.org/10.5281/zenodo.13173816" httpUri="https://zenodo.org/record/13173816/files/figure.png" pageId="1" pageNumber="2">Fig. 7</figureCitation>
) prognathous, widest across the eyes. Compound eyes (
<figureCitation id="13602A3FE703126E930CF937FE94B1F5" box="[281,337,1678,1698]" captionStart="Figures 712" captionStartId="5.[151,218,1068,1088]" captionTargetBox="[151,1428,197,1054]" captionTargetId="figure-383@5.[151,1428,197,1054]" captionTargetPageId="5" captionText="Figures 712. Details of Delteredolaemus hei gen. et sp. nov., holotype, NIGP200000, under confocal microscopy: (7) head, anteroventral view; (8) prothorax, ventral view; (9) mesothorax, ventral view; (10) abdominal base, ventral view; (11) abdominal apex, ventral view; (12) pronotal and elytral bases, dorsal view, with arrowhead showing the medially produced posterior pronotal edge. (a1) antennomere 1, (el) elytron, (lbp) labial palp, (md) mandible, (msf) mesofemur, (mstb) mesotibia, (msv) mesoventrite, (mtf) metafemur, (mtv) metaventrite, (mxp) maxillary palp, (pc) procoxa, (pf) profemur, (pn) pronotum, (ptb) protibia, (ptc) protrochanter, (sc) scutellum, (ts) tibial spine, (v15) ventrites 15. Scale bars: 100 Μm." figureDoi="http://doi.org/10.5281/zenodo.13173816" httpUri="https://zenodo.org/record/13173816/files/figure.png" pageId="1" pageNumber="2">Fig. 7</figureCitation>
) finely facetted, without interfacetal setae. Antennal insertions (
<figureCitation id="13602A3FE703126E937AF915FE70B1EB" box="[367,437,1708,1728]" captionStart="Figures 1317" captionStartId="6.[159,226,1698,1718]" captionTargetBox="[253,1342,197,1686]" captionTargetId="figure-43@6.[253,1342,197,1686]" captionTargetPageId="6" captionText="Figures 1317. X-ray microtomographic reconstruction of Delteredolaemus hei gen. et sp. nov., holotype, NIGP200000: (13) dorsal view; (14) ventral view; (15) lateral view; (16) anterodorsal view; (17) head, ventrolateral view. (a111) antennomeres 111. Scale bars: 500 Μm in (1316), 200 Μm in (17)." figureDoi="http://doi.org/10.5281/zenodo.13173818" httpUri="https://zenodo.org/record/13173818/files/figure.png" pageId="1" pageNumber="2">Fig. 16</figureCitation>
) exposed from above. Antennae (
<figureCitation id="13602A3FE703126E9288F970FF19B189" box="[157,220,1737,1757]" captionStart="Figures 712" captionStartId="5.[151,218,1068,1088]" captionTargetBox="[151,1428,197,1054]" captionTargetId="figure-383@5.[151,1428,197,1054]" captionTargetPageId="5" captionText="Figures 712. Details of Delteredolaemus hei gen. et sp. nov., holotype, NIGP200000, under confocal microscopy: (7) head, anteroventral view; (8) prothorax, ventral view; (9) mesothorax, ventral view; (10) abdominal base, ventral view; (11) abdominal apex, ventral view; (12) pronotal and elytral bases, dorsal view, with arrowhead showing the medially produced posterior pronotal edge. (a1) antennomere 1, (el) elytron, (lbp) labial palp, (md) mandible, (msf) mesofemur, (mstb) mesotibia, (msv) mesoventrite, (mtf) metafemur, (mtv) metaventrite, (mxp) maxillary palp, (pc) procoxa, (pf) profemur, (pn) pronotum, (ptb) protibia, (ptc) protrochanter, (sc) scutellum, (ts) tibial spine, (v15) ventrites 15. Scale bars: 100 Μm." figureDoi="http://doi.org/10.5281/zenodo.13173816" httpUri="https://zenodo.org/record/13173816/files/figure.png" pageId="1" pageNumber="2">Figs 7</figureCitation>
,
<figureCitation id="13602A3FE703126E92FFF973FEC6B189" box="[234,259,1738,1757]" captionStart="Figures 1317" captionStartId="6.[159,226,1698,1718]" captionTargetBox="[253,1342,197,1686]" captionTargetId="figure-43@6.[253,1342,197,1686]" captionTargetPageId="6" captionText="Figures 1317. X-ray microtomographic reconstruction of Delteredolaemus hei gen. et sp. nov., holotype, NIGP200000: (13) dorsal view; (14) ventral view; (15) lateral view; (16) anterodorsal view; (17) head, ventrolateral view. (a111) antennomeres 111. Scale bars: 500 Μm in (1316), 200 Μm in (17)." figureDoi="http://doi.org/10.5281/zenodo.13173818" httpUri="https://zenodo.org/record/13173818/files/figure.png" pageId="1" pageNumber="2">17</figureCitation>
) likely 11-segmented; antennomere 1 robust; antennomeres 14 progressively narrower; antennomeres 10 and 11 forming a relatively compact club; antennomeres 19 with scattered long setae only; antennomere 11 with dense sensorial setae. Mandibles (
<figureCitation id="13602A3FE703126E91C6FEB8FBC2B641" box="[979,1031,257,277]" captionStart="Figures 712" captionStartId="5.[151,218,1068,1088]" captionTargetBox="[151,1428,197,1054]" captionTargetId="figure-383@5.[151,1428,197,1054]" captionTargetPageId="5" captionText="Figures 712. Details of Delteredolaemus hei gen. et sp. nov., holotype, NIGP200000, under confocal microscopy: (7) head, anteroventral view; (8) prothorax, ventral view; (9) mesothorax, ventral view; (10) abdominal base, ventral view; (11) abdominal apex, ventral view; (12) pronotal and elytral bases, dorsal view, with arrowhead showing the medially produced posterior pronotal edge. (a1) antennomere 1, (el) elytron, (lbp) labial palp, (md) mandible, (msf) mesofemur, (mstb) mesotibia, (msv) mesoventrite, (mtf) metafemur, (mtv) metaventrite, (mxp) maxillary palp, (pc) procoxa, (pf) profemur, (pn) pronotum, (ptb) protibia, (ptc) protrochanter, (sc) scutellum, (ts) tibial spine, (v15) ventrites 15. Scale bars: 100 Μm." figureDoi="http://doi.org/10.5281/zenodo.13173816" httpUri="https://zenodo.org/record/13173816/files/figure.png" pageId="1" pageNumber="2">Fig. 7</figureCitation>
) (at least) bidentate. Maxillary palps (
<figureCitation id="13602A3FE703126E9765FEB8FCF6B667" captionStart="Figures 712" captionStartId="5.[151,218,1068,1088]" captionTargetBox="[151,1428,197,1054]" captionTargetId="figure-383@5.[151,1428,197,1054]" captionTargetPageId="5" captionText="Figures 712. Details of Delteredolaemus hei gen. et sp. nov., holotype, NIGP200000, under confocal microscopy: (7) head, anteroventral view; (8) prothorax, ventral view; (9) mesothorax, ventral view; (10) abdominal base, ventral view; (11) abdominal apex, ventral view; (12) pronotal and elytral bases, dorsal view, with arrowhead showing the medially produced posterior pronotal edge. (a1) antennomere 1, (el) elytron, (lbp) labial palp, (md) mandible, (msf) mesofemur, (mstb) mesotibia, (msv) mesoventrite, (mtf) metafemur, (mtv) metaventrite, (mxp) maxillary palp, (pc) procoxa, (pf) profemur, (pn) pronotum, (ptb) protibia, (ptc) protrochanter, (sc) scutellum, (ts) tibial spine, (v15) ventrites 15. Scale bars: 100 Μm." figureDoi="http://doi.org/10.5281/zenodo.13173816" httpUri="https://zenodo.org/record/13173816/files/figure.png" pageId="1" pageNumber="2">Fig. 7</figureCitation>
) 4-segmented, with apical palpomere subconical. Labial palps (
<figureCitation id="13602A3FE703126E913BFE84FCA0B605" box="[814,869,317,337]" captionStart="Figures 712" captionStartId="5.[151,218,1068,1088]" captionTargetBox="[151,1428,197,1054]" captionTargetId="figure-383@5.[151,1428,197,1054]" captionTargetPageId="5" captionText="Figures 712. Details of Delteredolaemus hei gen. et sp. nov., holotype, NIGP200000, under confocal microscopy: (7) head, anteroventral view; (8) prothorax, ventral view; (9) mesothorax, ventral view; (10) abdominal base, ventral view; (11) abdominal apex, ventral view; (12) pronotal and elytral bases, dorsal view, with arrowhead showing the medially produced posterior pronotal edge. (a1) antennomere 1, (el) elytron, (lbp) labial palp, (md) mandible, (msf) mesofemur, (mstb) mesotibia, (msv) mesoventrite, (mtf) metafemur, (mtv) metaventrite, (mxp) maxillary palp, (pc) procoxa, (pf) profemur, (pn) pronotum, (ptb) protibia, (ptc) protrochanter, (sc) scutellum, (ts) tibial spine, (v15) ventrites 15. Scale bars: 100 Μm." figureDoi="http://doi.org/10.5281/zenodo.13173816" httpUri="https://zenodo.org/record/13173816/files/figure.png" pageId="1" pageNumber="2">Fig. 7</figureCitation>
) with apical palpomere likely cylindrical.
</paragraph>
<paragraph id="8BE436BAE703126E9170FEE2FB33B56B" blockId="1.[807,1430,198,1173]" pageId="1" pageNumber="2">
Pronotal disc about 1.4 times as long as wide; surface without special modifications (carinae, cavities, grooves, or paired impressions); posterior pronotal edge posteriorly produced at middle, forming rather distinct angle (
<figureCitation id="13602A3FE703126E96B0FE0CFB2CB69C" box="[1189,1257,437,457]" captionStart="Figures 712" captionStartId="5.[151,218,1068,1088]" captionTargetBox="[151,1428,197,1054]" captionTargetId="figure-383@5.[151,1428,197,1054]" captionTargetPageId="5" captionText="Figures 712. Details of Delteredolaemus hei gen. et sp. nov., holotype, NIGP200000, under confocal microscopy: (7) head, anteroventral view; (8) prothorax, ventral view; (9) mesothorax, ventral view; (10) abdominal base, ventral view; (11) abdominal apex, ventral view; (12) pronotal and elytral bases, dorsal view, with arrowhead showing the medially produced posterior pronotal edge. (a1) antennomere 1, (el) elytron, (lbp) labial palp, (md) mandible, (msf) mesofemur, (mstb) mesotibia, (msv) mesoventrite, (mtf) metafemur, (mtv) metaventrite, (mxp) maxillary palp, (pc) procoxa, (pf) profemur, (pn) pronotum, (ptb) protibia, (ptc) protrochanter, (sc) scutellum, (ts) tibial spine, (v15) ventrites 15. Scale bars: 100 Μm." figureDoi="http://doi.org/10.5281/zenodo.13173816" httpUri="https://zenodo.org/record/13173816/files/figure.png" pageId="1" pageNumber="2">Fig. 12</figureCitation>
); lateral pronotal carinae complete (
<figureCitation id="13602A3FE703126E91F1FE6BFBEEB6B2" box="[996,1067,466,486]" captionStart="Figures 1317" captionStartId="6.[159,226,1698,1718]" captionTargetBox="[253,1342,197,1686]" captionTargetId="figure-43@6.[253,1342,197,1686]" captionTargetPageId="6" captionText="Figures 1317. X-ray microtomographic reconstruction of Delteredolaemus hei gen. et sp. nov., holotype, NIGP200000: (13) dorsal view; (14) ventral view; (15) lateral view; (16) anterodorsal view; (17) head, ventrolateral view. (a111) antennomeres 111. Scale bars: 500 Μm in (1316), 200 Μm in (17)." figureDoi="http://doi.org/10.5281/zenodo.13173818" httpUri="https://zenodo.org/record/13173818/files/figure.png" pageId="1" pageNumber="2">Fig. 15</figureCitation>
). Notosternal suture complete (
<figureCitation id="13602A3FE703126E9765FE6BFC86B550" captionStart="Figures 1317" captionStartId="6.[159,226,1698,1718]" captionTargetBox="[253,1342,197,1686]" captionTargetId="figure-43@6.[253,1342,197,1686]" captionTargetPageId="6" captionText="Figures 1317. X-ray microtomographic reconstruction of Delteredolaemus hei gen. et sp. nov., holotype, NIGP200000: (13) dorsal view; (14) ventral view; (15) lateral view; (16) anterodorsal view; (17) head, ventrolateral view. (a111) antennomeres 111. Scale bars: 500 Μm in (1316), 200 Μm in (17)." figureDoi="http://doi.org/10.5281/zenodo.13173818" httpUri="https://zenodo.org/record/13173818/files/figure.png" pageId="1" pageNumber="2">Fig. 15</figureCitation>
). Prosternal process either absent or completely concealed by projecting procoxae (
<figureCitation id="13602A3FE703126E9603FDB7FB89B576" box="[1046,1100,526,546]" captionStart="Figures 712" captionStartId="5.[151,218,1068,1088]" captionTargetBox="[151,1428,197,1054]" captionTargetId="figure-383@5.[151,1428,197,1054]" captionTargetPageId="5" captionText="Figures 712. Details of Delteredolaemus hei gen. et sp. nov., holotype, NIGP200000, under confocal microscopy: (7) head, anteroventral view; (8) prothorax, ventral view; (9) mesothorax, ventral view; (10) abdominal base, ventral view; (11) abdominal apex, ventral view; (12) pronotal and elytral bases, dorsal view, with arrowhead showing the medially produced posterior pronotal edge. (a1) antennomere 1, (el) elytron, (lbp) labial palp, (md) mandible, (msf) mesofemur, (mstb) mesotibia, (msv) mesoventrite, (mtf) metafemur, (mtv) metaventrite, (mxp) maxillary palp, (pc) procoxa, (pf) profemur, (pn) pronotum, (ptb) protibia, (ptc) protrochanter, (sc) scutellum, (ts) tibial spine, (v15) ventrites 15. Scale bars: 100 Μm." figureDoi="http://doi.org/10.5281/zenodo.13173816" httpUri="https://zenodo.org/record/13173816/files/figure.png" pageId="1" pageNumber="2">Fig. 8</figureCitation>
). Procoxae (
<figureCitation id="13602A3FE703126E96D6FDB7FB3DB576" box="[1219,1272,526,546]" captionStart="Figures 712" captionStartId="5.[151,218,1068,1088]" captionTargetBox="[151,1428,197,1054]" captionTargetId="figure-383@5.[151,1428,197,1054]" captionTargetPageId="5" captionText="Figures 712. Details of Delteredolaemus hei gen. et sp. nov., holotype, NIGP200000, under confocal microscopy: (7) head, anteroventral view; (8) prothorax, ventral view; (9) mesothorax, ventral view; (10) abdominal base, ventral view; (11) abdominal apex, ventral view; (12) pronotal and elytral bases, dorsal view, with arrowhead showing the medially produced posterior pronotal edge. (a1) antennomere 1, (el) elytron, (lbp) labial palp, (md) mandible, (msf) mesofemur, (mstb) mesotibia, (msv) mesoventrite, (mtf) metafemur, (mtv) metaventrite, (mxp) maxillary palp, (pc) procoxa, (pf) profemur, (pn) pronotum, (ptb) protibia, (ptc) protrochanter, (sc) scutellum, (ts) tibial spine, (v15) ventrites 15. Scale bars: 100 Μm." figureDoi="http://doi.org/10.5281/zenodo.13173816" httpUri="https://zenodo.org/record/13173816/files/figure.png" pageId="1" pageNumber="2">Fig. 8</figureCitation>
) projecting and contiguous (status of coxal cavities unknown).
</paragraph>
<paragraph id="8BE436BAE703126E9170FDF3FB30B47A" blockId="1.[807,1430,198,1173]" pageId="1" pageNumber="2">
Scutellar shield (
<figureCitation id="13602A3FE703126E961EFDF3FB8AB509" box="[1035,1103,586,606]" captionStart="Figures 712" captionStartId="5.[151,218,1068,1088]" captionTargetBox="[151,1428,197,1054]" captionTargetId="figure-383@5.[151,1428,197,1054]" captionTargetPageId="5" captionText="Figures 712. Details of Delteredolaemus hei gen. et sp. nov., holotype, NIGP200000, under confocal microscopy: (7) head, anteroventral view; (8) prothorax, ventral view; (9) mesothorax, ventral view; (10) abdominal base, ventral view; (11) abdominal apex, ventral view; (12) pronotal and elytral bases, dorsal view, with arrowhead showing the medially produced posterior pronotal edge. (a1) antennomere 1, (el) elytron, (lbp) labial palp, (md) mandible, (msf) mesofemur, (mstb) mesotibia, (msv) mesoventrite, (mtf) metafemur, (mtv) metaventrite, (mxp) maxillary palp, (pc) procoxa, (pf) profemur, (pn) pronotum, (ptb) protibia, (ptc) protrochanter, (sc) scutellum, (ts) tibial spine, (v15) ventrites 15. Scale bars: 100 Μm." figureDoi="http://doi.org/10.5281/zenodo.13173816" httpUri="https://zenodo.org/record/13173816/files/figure.png" pageId="1" pageNumber="2">Fig. 12</figureCitation>
) small and rounded. Elytra (
<figureCitation id="13602A3FE703126E9765FDF3FCF3B52F" captionStart="Figures 34" captionStartId="3.[151,218,1733,1753]" captionTargetBox="[151,1428,197,1716]" captionTargetId="figure-27@3.[151,1428,197,1719]" captionTargetPageId="3" captionText="Figures 34. General habitus of Delteredolaemus hei gen. et sp. nov., holotype, NIGP200000, under confocal microscopy: (3) dorsal view; (4) ventral view. Scale bars: 500 Μm." figureDoi="http://doi.org/10.5281/zenodo.13173812" httpUri="https://zenodo.org/record/13173812/files/figure.png" pageId="1" pageNumber="2">Fig. 3</figureCitation>
) elongate; surface with clear puncture rows, without costae; scutellar striole absent; epipleura narrow. Mesoventrite (
<figureCitation id="13602A3FE703126E974DFD3FFA4BB5CD" box="[1368,1422,646,666]" captionStart="Figures 712" captionStartId="5.[151,218,1068,1088]" captionTargetBox="[151,1428,197,1054]" captionTargetId="figure-383@5.[151,1428,197,1054]" captionTargetPageId="5" captionText="Figures 712. Details of Delteredolaemus hei gen. et sp. nov., holotype, NIGP200000, under confocal microscopy: (7) head, anteroventral view; (8) prothorax, ventral view; (9) mesothorax, ventral view; (10) abdominal base, ventral view; (11) abdominal apex, ventral view; (12) pronotal and elytral bases, dorsal view, with arrowhead showing the medially produced posterior pronotal edge. (a1) antennomere 1, (el) elytron, (lbp) labial palp, (md) mandible, (msf) mesofemur, (mstb) mesotibia, (msv) mesoventrite, (mtf) metafemur, (mtv) metaventrite, (mxp) maxillary palp, (pc) procoxa, (pf) profemur, (pn) pronotum, (ptb) protibia, (ptc) protrochanter, (sc) scutellum, (ts) tibial spine, (v15) ventrites 15. Scale bars: 100 Μm." figureDoi="http://doi.org/10.5281/zenodo.13173816" httpUri="https://zenodo.org/record/13173816/files/figure.png" pageId="1" pageNumber="2">Fig. 9</figureCitation>
) without median carina; mesoventral process well-developed, slightly broadened at middle, apically narrowly rounded. Mesocoxal cavities narrowly separated (
<figureCitation id="13602A3FE703126E9668FD66FB73B5A7" box="[1149,1206,735,755]" captionStart="Figures 712" captionStartId="5.[151,218,1068,1088]" captionTargetBox="[151,1428,197,1054]" captionTargetId="figure-383@5.[151,1428,197,1054]" captionTargetPageId="5" captionText="Figures 712. Details of Delteredolaemus hei gen. et sp. nov., holotype, NIGP200000, under confocal microscopy: (7) head, anteroventral view; (8) prothorax, ventral view; (9) mesothorax, ventral view; (10) abdominal base, ventral view; (11) abdominal apex, ventral view; (12) pronotal and elytral bases, dorsal view, with arrowhead showing the medially produced posterior pronotal edge. (a1) antennomere 1, (el) elytron, (lbp) labial palp, (md) mandible, (msf) mesofemur, (mstb) mesotibia, (msv) mesoventrite, (mtf) metafemur, (mtv) metaventrite, (mxp) maxillary palp, (pc) procoxa, (pf) profemur, (pn) pronotum, (ptb) protibia, (ptc) protrochanter, (sc) scutellum, (ts) tibial spine, (v15) ventrites 15. Scale bars: 100 Μm." figureDoi="http://doi.org/10.5281/zenodo.13173816" httpUri="https://zenodo.org/record/13173816/files/figure.png" pageId="1" pageNumber="2">Fig. 9</figureCitation>
). Metaventrite anteromedially tumid (
<figureCitation id="13602A3FE703126E91DFFD44FBC6B444" box="[970,1027,765,785]" captionStart="Figures 56" captionStartId="4.[159,226,1733,1753]" captionTargetBox="[159,1436,197,1716]" captionTargetId="figure-31@4.[159,1436,194,1719]" captionTargetPageId="4" captionText="Figures 56. General habitus of Delteredolaemus hei gen. et sp. nov., holotype, NIGP200000, under confocal microscopy, with depth color coding: (5) dorsal view; (6) ventral view. Scale bars: 500 Μm." figureDoi="http://doi.org/10.5281/zenodo.13173814" httpUri="https://zenodo.org/record/13173814/files/figure.png" pageId="1" pageNumber="2">Figs 6</figureCitation>
,
<figureCitation id="13602A3FE703126E9619FD47FBDCB445" box="[1036,1049,766,785]" captionStart="Figures 712" captionStartId="5.[151,218,1068,1088]" captionTargetBox="[151,1428,197,1054]" captionTargetId="figure-383@5.[151,1428,197,1054]" captionTargetPageId="5" captionText="Figures 712. Details of Delteredolaemus hei gen. et sp. nov., holotype, NIGP200000, under confocal microscopy: (7) head, anteroventral view; (8) prothorax, ventral view; (9) mesothorax, ventral view; (10) abdominal base, ventral view; (11) abdominal apex, ventral view; (12) pronotal and elytral bases, dorsal view, with arrowhead showing the medially produced posterior pronotal edge. (a1) antennomere 1, (el) elytron, (lbp) labial palp, (md) mandible, (msf) mesofemur, (mstb) mesotibia, (msv) mesoventrite, (mtf) metafemur, (mtv) metaventrite, (mxp) maxillary palp, (pc) procoxa, (pf) profemur, (pn) pronotum, (ptb) protibia, (ptc) protrochanter, (sc) scutellum, (ts) tibial spine, (v15) ventrites 15. Scale bars: 100 Μm." figureDoi="http://doi.org/10.5281/zenodo.13173816" httpUri="https://zenodo.org/record/13173816/files/figure.png" pageId="1" pageNumber="2">9</figureCitation>
); median discrimen absent. Metacoxae (
<figureCitation id="13602A3FE703126E913BFCA2FCB4B47A" box="[814,881,795,815]" captionStart="Figures 712" captionStartId="5.[151,218,1068,1088]" captionTargetBox="[151,1428,197,1054]" captionTargetId="figure-383@5.[151,1428,197,1054]" captionTargetPageId="5" captionText="Figures 712. Details of Delteredolaemus hei gen. et sp. nov., holotype, NIGP200000, under confocal microscopy: (7) head, anteroventral view; (8) prothorax, ventral view; (9) mesothorax, ventral view; (10) abdominal base, ventral view; (11) abdominal apex, ventral view; (12) pronotal and elytral bases, dorsal view, with arrowhead showing the medially produced posterior pronotal edge. (a1) antennomere 1, (el) elytron, (lbp) labial palp, (md) mandible, (msf) mesofemur, (mstb) mesotibia, (msv) mesoventrite, (mtf) metafemur, (mtv) metaventrite, (mxp) maxillary palp, (pc) procoxa, (pf) profemur, (pn) pronotum, (ptb) protibia, (ptc) protrochanter, (sc) scutellum, (ts) tibial spine, (v15) ventrites 15. Scale bars: 100 Μm." figureDoi="http://doi.org/10.5281/zenodo.13173816" httpUri="https://zenodo.org/record/13173816/files/figure.png" pageId="1" pageNumber="2">Fig. 10</figureCitation>
) transversely oval, narrowly separated.
</paragraph>
<paragraph id="8BE436BAE703126E9170FC83FB15B4B5" blockId="1.[807,1430,198,1173]" pageId="1" pageNumber="2">
Legs well-developed, slender. Trochanters large and well visible, with strongly oblique trochanterofemoral joint (
<figureCitation id="13602A3FE703126E977BFCEEFC9EB4DC" captionStart="Figures 712" captionStartId="5.[151,218,1068,1088]" captionTargetBox="[151,1428,197,1054]" captionTargetId="figure-383@5.[151,1428,197,1054]" captionTargetPageId="5" captionText="Figures 712. Details of Delteredolaemus hei gen. et sp. nov., holotype, NIGP200000, under confocal microscopy: (7) head, anteroventral view; (8) prothorax, ventral view; (9) mesothorax, ventral view; (10) abdominal base, ventral view; (11) abdominal apex, ventral view; (12) pronotal and elytral bases, dorsal view, with arrowhead showing the medially produced posterior pronotal edge. (a1) antennomere 1, (el) elytron, (lbp) labial palp, (md) mandible, (msf) mesofemur, (mstb) mesotibia, (msv) mesoventrite, (mtf) metafemur, (mtv) metaventrite, (mxp) maxillary palp, (pc) procoxa, (pf) profemur, (pn) pronotum, (ptb) protibia, (ptc) protrochanter, (sc) scutellum, (ts) tibial spine, (v15) ventrites 15. Scale bars: 100 Μm." figureDoi="http://doi.org/10.5281/zenodo.13173816" httpUri="https://zenodo.org/record/13173816/files/figure.png" pageId="1" pageNumber="2">Figs 810</figureCitation>
). Tibiae with spines present at outer apical angle only (
<figureCitation id="13602A3FE703126E977BFCCCFC9FB4F2" captionStart="Figures 712" captionStartId="5.[151,218,1068,1088]" captionTargetBox="[151,1428,197,1054]" captionTargetId="figure-383@5.[151,1428,197,1054]" captionTargetPageId="5" captionText="Figures 712. Details of Delteredolaemus hei gen. et sp. nov., holotype, NIGP200000, under confocal microscopy: (7) head, anteroventral view; (8) prothorax, ventral view; (9) mesothorax, ventral view; (10) abdominal base, ventral view; (11) abdominal apex, ventral view; (12) pronotal and elytral bases, dorsal view, with arrowhead showing the medially produced posterior pronotal edge. (a1) antennomere 1, (el) elytron, (lbp) labial palp, (md) mandible, (msf) mesofemur, (mstb) mesotibia, (msv) mesoventrite, (mtf) metafemur, (mtv) metaventrite, (mxp) maxillary palp, (pc) procoxa, (pf) profemur, (pn) pronotum, (ptb) protibia, (ptc) protrochanter, (sc) scutellum, (ts) tibial spine, (v15) ventrites 15. Scale bars: 100 Μm." figureDoi="http://doi.org/10.5281/zenodo.13173816" httpUri="https://zenodo.org/record/13173816/files/figure.png" pageId="1" pageNumber="2">Figs 810</figureCitation>
); tibial spurs 2-2-2, well-developed (
<figureCitation id="13602A3FE703126E96D8FC2BFAF5B4F2" box="[1229,1328,914,934]" captionStart="Figures 712" captionStartId="5.[151,218,1068,1088]" captionTargetBox="[151,1428,197,1054]" captionTargetId="figure-383@5.[151,1428,197,1054]" captionTargetPageId="5" captionText="Figures 712. Details of Delteredolaemus hei gen. et sp. nov., holotype, NIGP200000, under confocal microscopy: (7) head, anteroventral view; (8) prothorax, ventral view; (9) mesothorax, ventral view; (10) abdominal base, ventral view; (11) abdominal apex, ventral view; (12) pronotal and elytral bases, dorsal view, with arrowhead showing the medially produced posterior pronotal edge. (a1) antennomere 1, (el) elytron, (lbp) labial palp, (md) mandible, (msf) mesofemur, (mstb) mesotibia, (msv) mesoventrite, (mtf) metafemur, (mtv) metaventrite, (mxp) maxillary palp, (pc) procoxa, (pf) profemur, (pn) pronotum, (ptb) protibia, (ptc) protrochanter, (sc) scutellum, (ts) tibial spine, (v15) ventrites 15. Scale bars: 100 Μm." figureDoi="http://doi.org/10.5281/zenodo.13173816" httpUri="https://zenodo.org/record/13173816/files/figure.png" pageId="1" pageNumber="2">Figs 810</figureCitation>
). Tarsi all 4-segmented; tarsomeres simple, unlobed (
<figureCitation id="13602A3FE703126E96C5FC09FAF5B490" box="[1232,1328,944,964]" captionStart="Figures 712" captionStartId="5.[151,218,1068,1088]" captionTargetBox="[151,1428,197,1054]" captionTargetId="figure-383@5.[151,1428,197,1054]" captionTargetPageId="5" captionText="Figures 712. Details of Delteredolaemus hei gen. et sp. nov., holotype, NIGP200000, under confocal microscopy: (7) head, anteroventral view; (8) prothorax, ventral view; (9) mesothorax, ventral view; (10) abdominal base, ventral view; (11) abdominal apex, ventral view; (12) pronotal and elytral bases, dorsal view, with arrowhead showing the medially produced posterior pronotal edge. (a1) antennomere 1, (el) elytron, (lbp) labial palp, (md) mandible, (msf) mesofemur, (mstb) mesotibia, (msv) mesoventrite, (mtf) metafemur, (mtv) metaventrite, (mxp) maxillary palp, (pc) procoxa, (pf) profemur, (pn) pronotum, (ptb) protibia, (ptc) protrochanter, (sc) scutellum, (ts) tibial spine, (v15) ventrites 15. Scale bars: 100 Μm." figureDoi="http://doi.org/10.5281/zenodo.13173816" httpUri="https://zenodo.org/record/13173816/files/figure.png" pageId="1" pageNumber="2">Figs 810</figureCitation>
). Pretarsal claws simple; empodium present, bisetose.
</paragraph>
<paragraph id="8BE436BAE703126E9170FC55FA81B36F" blockId="1.[807,1430,198,1173]" pageId="1" pageNumber="2">
Abdomen with five ventrites (
<figureCitation id="13602A3FE703126E9697FC55FB2FB354" box="[1154,1258,1004,1024]" captionStart="Figures 712" captionStartId="5.[151,218,1068,1088]" captionTargetBox="[151,1428,197,1054]" captionTargetId="figure-383@5.[151,1428,197,1054]" captionTargetPageId="5" captionText="Figures 712. Details of Delteredolaemus hei gen. et sp. nov., holotype, NIGP200000, under confocal microscopy: (7) head, anteroventral view; (8) prothorax, ventral view; (9) mesothorax, ventral view; (10) abdominal base, ventral view; (11) abdominal apex, ventral view; (12) pronotal and elytral bases, dorsal view, with arrowhead showing the medially produced posterior pronotal edge. (a1) antennomere 1, (el) elytron, (lbp) labial palp, (md) mandible, (msf) mesofemur, (mstb) mesotibia, (msv) mesoventrite, (mtf) metafemur, (mtv) metaventrite, (mxp) maxillary palp, (pc) procoxa, (pf) profemur, (pn) pronotum, (ptb) protibia, (ptc) protrochanter, (sc) scutellum, (ts) tibial spine, (v15) ventrites 15. Scale bars: 100 Μm." figureDoi="http://doi.org/10.5281/zenodo.13173816" httpUri="https://zenodo.org/record/13173816/files/figure.png" pageId="1" pageNumber="2">Figs 10, 11</figureCitation>
). Ratio of ventrite lengths along middle: 3.8:1.3:1.1:1.0:1.5. Ventrite 1 with intercoxal process narrow and apically acuminate (
<figureCitation id="13602A3FE703126E96E6FB91FAFDB36F" box="[1267,1336,1064,1084]" captionStart="Figures 712" captionStartId="5.[151,218,1068,1088]" captionTargetBox="[151,1428,197,1054]" captionTargetId="figure-383@5.[151,1428,197,1054]" captionTargetPageId="5" captionText="Figures 712. Details of Delteredolaemus hei gen. et sp. nov., holotype, NIGP200000, under confocal microscopy: (7) head, anteroventral view; (8) prothorax, ventral view; (9) mesothorax, ventral view; (10) abdominal base, ventral view; (11) abdominal apex, ventral view; (12) pronotal and elytral bases, dorsal view, with arrowhead showing the medially produced posterior pronotal edge. (a1) antennomere 1, (el) elytron, (lbp) labial palp, (md) mandible, (msf) mesofemur, (mstb) mesotibia, (msv) mesoventrite, (mtf) metafemur, (mtv) metaventrite, (mxp) maxillary palp, (pc) procoxa, (pf) profemur, (pn) pronotum, (ptb) protibia, (ptc) protrochanter, (sc) scutellum, (ts) tibial spine, (v15) ventrites 15. Scale bars: 100 Μm." figureDoi="http://doi.org/10.5281/zenodo.13173816" httpUri="https://zenodo.org/record/13173816/files/figure.png" pageId="1" pageNumber="2">Fig. 10</figureCitation>
).
</paragraph>
<paragraph id="8BE436BAE703126E9170FBFFFC44B3C1" blockId="1.[807,1430,198,1173]" pageId="1" pageNumber="2">
Etymology. The species is named after Mr. Hai-Kun He (
<collectingRegion id="499FF858E703126E9125FBDAFC6DB323" box="[816,936,1123,1143]" country="China" name="Guangdong" pageId="1" pageNumber="2">Guangdong</collectingRegion>
,
<collectingCountry id="F34C762AE703126E91A7FBDDFC37B323" box="[946,1010,1124,1143]" name="China" pageId="1" pageNumber="2">China</collectingCountry>
), who kindly donated many fossils for our research.
</paragraph>
</subSubSection>
</treatment>
</document>