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<document id="79A37968072143870FE898B3D086CA81" ID-CLB-Dataset="29010" ID-DOI="10.11646/zootaxa.4459.3.7" ID-GBIF-Dataset="81db55ed-b19e-42e9-930e-f9697834cc1b" ID-ISSN="1175-5326" ID-Zenodo-Dep="1458879" ID-ZooBank="8CA301B7-A24A-4CB1-8231-FF0BAFF175DA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1539290198072" checkinUser="plazi" docAuthor="Semenyuk, Irina, Golovatch, Sergei I. &amp; Wesener, Thomas" docDate="2018" docId="03D7EC02FFDA2424CBBDE057DC7AFBC1" docLanguage="en" docName="zootaxa.4459.3.7.pdf" docOrigin="Zootaxa 4459 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Sphaerobelum cattiense Semenyuk &amp; Golovatch &amp; Wesener 2018, sp. nov." docType="treatment" docVersion="8" lastPageNumber="540" masterDocId="FFEE947AFFD82421CB2AE274D834FFAC" masterDocTitle="Four new species of giant pill-millipedes from Vietnam (Sphaerotheriida, Zephroniidae)" masterLastPageNumber="550" masterPageNumber="535" pageNumber="537" updateTime="1698578940799" updateUser="ExternalLinkService">
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<mods:title id="ADC8BB326354F2F13082556BB27B1E6E">Four new species of giant pill-millipedes from Vietnam (Sphaerotheriida, Zephroniidae)</mods:title>
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<mods:namePart id="9907E1B5C7BAB191E640A2AD3FD26966">Semenyuk, Irina</mods:namePart>
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<mods:namePart id="420D8144E1786D7336FAC69B8C182598">Golovatch, Sergei I.</mods:namePart>
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<mods:namePart id="238B91E4AE36A0958B416DD4C74BF2AF">Wesener, Thomas</mods:namePart>
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<treatment id="03D7EC02FFDA2424CBBDE057DC7AFBC1" ID-DOI="http://doi.org/10.5281/zenodo.6495776" ID-GBIF-Taxon="148692871" ID-Zenodo-Dep="6495776" LSID="urn:lsid:plazi:treatment:03D7EC02FFDA2424CBBDE057DC7AFBC1" httpUri="http://treatment.plazi.org/id/03D7EC02FFDA2424CBBDE057DC7AFBC1" lastPageId="5" lastPageNumber="540" pageId="2" pageNumber="537">
<subSubSection id="C3640E9FFFDA2423CBBDE057D96DFDF3" pageId="2" pageNumber="537" type="nomenclature">
<paragraph id="8BC15D14FFDA2423CBBDE057DBD0FD92" blockId="2.[151,996,547,607]" box="[151,996,547,574]" pageId="2" pageNumber="537">
<heading id="D089EA78FFDA2423CBBDE057DBD0FD92" bold="true" box="[151,996,547,574]" fontSize="11" level="1" pageId="2" pageNumber="537" reason="1">
<emphasis id="B90A8106FFDA2423CBBDE057DBD0FD92" bold="true" box="[151,996,547,574]" pageId="2" pageNumber="537">
<taxonomicName id="4C7E2697FFDA2423CBBDE057DB4EFD92" ID-CoL="84DDD" authority="Semenyuk, Golovatch and Wesener" authorityName="Semenyuk &amp; Golovatch &amp; Wesener" authorityYear="2018" box="[151,890,547,574]" class="Diplopoda" family="Zephroniidae" genus="Sphaerobelum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sphaerotheriida" pageId="2" pageNumber="537" phylum="Arthropoda" rank="species" species="cattiense" status="sp. nov.">
<emphasis id="B90A8106FFDA2423CBBDE057D983FD91" bold="true" box="[151,439,547,573]" italics="true" pageId="2" pageNumber="537">Sphaerobelum cattiense</emphasis>
Semenyuk, Golovatch and Wesener
</taxonomicName>
,
<taxonomicNameLabel id="A2393C7DFFDA2423C8ACE050DBD0FD92" box="[902,996,548,574]" pageId="2" pageNumber="537" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BC15D14FFDA2423CBBDE032D96DFDF3" blockId="2.[151,996,547,607]" box="[151,345,582,607]" pageId="2" pageNumber="537">
<figureCitation id="13454191FFDA2423CBBDE032D8C7FDF3" box="[151,243,582,607]" captionStart="FIGURE 1" captionStartId="3.[151,249,1736,1758]" captionTargetBox="[152,1435,447,1716]" captionTargetId="figure@3.[152,1435,447,1716]" captionTargetPageId="3" captionText="FIGURE 1. Distribution map with pictures of live specimens. A. Sphaerobelum cattiense sp. nov.; B. Sphaerobelum konkakinhense sp. nov.; C. Zephronia konkakinhensis sp. nov.; D. Zephronia montis sp. nov." httpUri="https://zenodo.org/record/1458881/files/figure.png" pageId="2" pageNumber="537">Figs 1A</figureCitation>
,
<figureCitation id="13454191FFDA2423CBD4E033D93AFDF3" box="[254,270,583,607]" captionStart="FIGURE 2" captionStartId="4.[151,249,1919,1941]" captionTargetBox="[191,1429,686,1892]" captionTargetId="figure@4.[176,1429,657,1899]" captionTargetPageId="4" captionText="FIGURE 2. Sphaerobelum cattiense sp. nov. AG = male paratypes, I = female paratype (all ZMUM). A. left anterior telopod, anterior view; B. left anterior telopod, lateral view; C. left anterior telopod, posterior view; D. leg 9, right, anterior view; E. right posterior telopod, posterior view; F. right posterior telopod, anterior view; G. telopoditomere 4, lateral view; H. right male antenna; I. left coxa and prefemur with vulva. Scale bar = 1 mm." httpUri="https://zenodo.org/record/1458883/files/figure.png" pageId="2" pageNumber="537">2</figureCitation>
,
<figureCitation id="13454191FFDA2423CA30E033D91EFDF3" box="[282,298,583,607]" captionStart="FIGURE 3" captionStartId="5.[151,249,662,684]" captionTargetBox="[166,1425,189,646]" captionTargetId="figure@5.[166,1426,187,647]" captionTargetPageId="5" captionText="FIGURE 3. Sphaerobelum cattiense sp. nov. Male paratype (ZFMK MYR6516), right antenna. A. lateral view; B. apical view of disc; C. details of apical cone. Abbreviations: sc = sclerotized scales; sb = sensilla basiconica. Scale bars: A = 1000 µm; B = 200 µm; C = 10 µm." httpUri="https://zenodo.org/record/1458885/files/figure.png" pageId="2" pageNumber="537">3</figureCitation>
,
<figureCitation id="13454191FFDA2423CA1CE033D96DFDF3" box="[310,345,583,607]" captionStart="FIGURE 4" captionStartId="6.[151,249,1534,1556]" captionTargetBox="[176,1427,837,1514]" captionTargetId="figure@6.[176,1428,837,1514]" captionTargetPageId="6" captionText="FIGURE 4. Endoterga of midbody rings. A. Sphaerobelum cattiense sp. nov. Male paratype (ZFMK MYR6516); B. Sphaerobelum konkakinhense sp. nov., male holotype (ZMUM). Scale bars: 100 µm." httpUri="https://zenodo.org/record/1458887/files/figure.png" pageId="2" pageNumber="537">4A</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3640E9FFFDA2423CBBDE0FAD8C9FCD3" pageId="2" pageNumber="537" type="materials_examined">
<paragraph id="8BC15D14FFDA2423CBBDE0FAD8C9FCD3" blockId="2.[151,1437,654,2011]" pageId="2" pageNumber="537">
<emphasis id="B90A8106FFDA2423CBBDE0FAD94EFD0B" bold="true" box="[151,378,654,679]" pageId="2" pageNumber="537">Material examined</emphasis>
: Type specimens. Holotype male (ZMUM), Vietnam, Dong Nai Province, Cat Tien National Park, monsoon lowland tropical forest (N 11°26' E 107°21',
<quantity id="4C86F0F1FFDA2423C878E0C7DBE3FD66" box="[850,983,691,715]" metricMagnitude="2" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="2" pageNumber="537" unit="m" value="150.0" valueMax="200.0" valueMin="100.0">100200 m</quantity>
. a.s.l.), litter, floor and under logs,
<date id="FFC07BD4FFDA2423CE58E0C7D92CFD43" pageId="2" pageNumber="537" value="2011-06-22" valueMax="2011-06-24" valueMin="2011-06-22">22 24.VI.2011</date>
, leg. I.I. Semenyuk. Paratypes: 1 males, 2 females (ZMUM), same data, together with holotype; 1 male, 1 female, 1 ex. of undetermined sex (ZFMK MYR 6516, 6519, 6565, same data as holotype, but collected in
<date id="FFC07BD4FFDA2423CBBDE16BD8C3FC9B" box="[151,247,799,823]" pageId="2" pageNumber="537" value="2017-06">VI.2017</date>
). Other material. 2 males, 2 females, 1 immature (topotypes), ZFMK MYR 6566, Vietnam, Dong Nai Province, Cat Tien National Park,
<taxonomicName id="4C7E2697FFDA2423C91DE137DB69FCF7" authorityName="Kurz" authorityYear="1872" box="[567,861,835,859]" class="Magnoliopsida" family="Lythraceae" genus="Lagerstroemia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Myrtales" pageId="2" pageNumber="537" phylum="Tracheophyta" rank="species" species="calyculata">
<emphasis id="B90A8106FFDA2423C91DE137DB69FCF7" box="[567,861,835,859]" italics="true" pageId="2" pageNumber="537">Lagerstroemia calyculata</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7E2697FFDA2423C8B5E137DC5EFCF7" authorityName="Craib" authorityYear="1912" box="[927,1130,835,859]" class="Insecta" family="Tachinidae" genus="Afzelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="537" phylum="Arthropoda" rank="species" species="xylocarpa">
<emphasis id="B90A8106FFDA2423C8B5E137DC5EFCF7" box="[927,1130,835,859]" italics="true" pageId="2" pageNumber="537">Afzelia xylocarpa</emphasis>
</taxonomicName>
forest,
<date id="FFC07BD4FFDA2423CFE1E137DD08FCF7" box="[1227,1340,835,859]" pageId="2" pageNumber="537" value="2011-08">VIII.2011</date>
, leg. E. Galoyan.
</paragraph>
</subSubSection>
<subSubSection id="C3640E9FFFDA2423CBEDE1FED8FFFBD7" pageId="2" pageNumber="537" type="diagnosis">
<paragraph id="8BC15D14FFDA2423CBEDE1FED8FFFBD7" blockId="2.[151,1437,654,2011]" pageId="2" pageNumber="537">
<emphasis id="B90A8106FFDA2423CBEDE1FED975FC0F" bold="true" box="[199,321,906,931]" pageId="2" pageNumber="537">Diagnosis.</emphasis>
Medium-sized,
<quantity id="4C86F0F1FFDA2423C928E1FEDAB4FC0E" box="[514,640,906,931]" metricMagnitude="-2" metricUnit="m" metricValue="2.75" metricValueMax="3.0" metricValueMin="2.5" pageId="2" pageNumber="537" unit="mm" value="27.5" valueMax="30.0" valueMin="25.0">2530 mm</quantity>
long and
<quantity id="4C86F0F1FFDA2423C9D5E1FEDB48FC0E" box="[767,892,906,931]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" metricValueMax="1.7" metricValueMin="1.5" pageId="2" pageNumber="537" unit="mm" value="16.0" valueMax="17.0" valueMin="15.0">1517 mm</quantity>
wide, dark olive-brown
<taxonomicName id="4C7E2697FFDA2423CF8EE1FFDD71FC0F" authorityName="Verhoeff" authorityYear="1924" box="[1188,1349,907,931]" class="Diplopoda" family="Zephroniidae" genus="Sphaerobelum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sphaerotheriida" pageId="2" pageNumber="537" phylum="Arthropoda" rank="genus">
<emphasis id="B90A8106FFDA2423CF8EE1FFDD71FC0F" box="[1188,1349,907,931]" italics="true" pageId="2" pageNumber="537">Sphaerobelum</emphasis>
</taxonomicName>
with a microsetose surface and orange antennae and appendages. Modified femur with a dentate mesal margin (
<figureCitation id="13454191FFDA2423CE1FE1DADDBBFC6B" box="[1333,1423,942,967]" captionStart="FIGURE 2" captionStartId="4.[151,249,1919,1941]" captionTargetBox="[191,1429,686,1892]" captionTargetId="figure@4.[176,1429,657,1899]" captionTargetPageId="4" captionText="FIGURE 2. Sphaerobelum cattiense sp. nov. AG = male paratypes, I = female paratype (all ZMUM). A. left anterior telopod, anterior view; B. left anterior telopod, lateral view; C. left anterior telopod, posterior view; D. leg 9, right, anterior view; E. right posterior telopod, posterior view; F. right posterior telopod, anterior view; G. telopoditomere 4, lateral view; H. right male antenna; I. left coxa and prefemur with vulva. Scale bar = 1 mm." httpUri="https://zenodo.org/record/1458883/files/figure.png" pageId="2" pageNumber="537">Fig. 2D</figureCitation>
), a character shared only with
<taxonomicName id="4C7E2697FFDA2423CAF5E1A7DA66FC47" authorityName="Attems" authorityYear="1938" box="[479,594,979,1003]" class="Diplopoda" family="Zephroniidae" genus="Sphaerobelum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sphaerotheriida" pageId="2" pageNumber="537" phylum="Arthropoda" rank="species" species="bicorne">
<emphasis id="B90A8106FFDA2423CAF5E1A7DA66FC47" box="[479,594,979,1003]" italics="true" pageId="2" pageNumber="537">S. bicorne</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7E2697FFDA2423C9A1E1A7DB79FC47" authority="Semenyuk &amp; Golovatch &amp; Wesener, 2018" authorityName="Semenyuk &amp; Golovatch &amp; Wesener" authorityYear="2018" box="[651,845,979,1003]" class="Diplopoda" family="Zephroniidae" genus="Sphaerobelum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sphaerotheriida" pageId="2" pageNumber="537" phylum="Arthropoda" rank="species" species="konkakinhense" status="sp. nov.">
<emphasis id="B90A8106FFDA2423C9A1E1A7DB79FC47" box="[651,845,979,1003]" italics="true" pageId="2" pageNumber="537">S. konkakinhense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2393C7DFFDA2423C87FE1A6DB84FC47" box="[853,944,978,1003]" pageId="2" pageNumber="537" rank="species">
<emphasis id="B90A8106FFDA2423C87FE1A6DB84FC47" bold="true" box="[853,944,978,1003]" pageId="2" pageNumber="537">sp. nov.</emphasis>
</taxonomicNameLabel>
Differing from both other congeners in the vulval operculum which lacks a recessed apical margin and projects in a single, well-rounded, mesal tip (
<figureCitation id="13454191FFDA2423CE15E182DDBBFBA3" box="[1343,1423,1014,1039]" captionStart="FIGURE 2" captionStartId="4.[151,249,1919,1941]" captionTargetBox="[191,1429,686,1892]" captionTargetId="figure@4.[176,1429,657,1899]" captionTargetPageId="4" captionText="FIGURE 2. Sphaerobelum cattiense sp. nov. AG = male paratypes, I = female paratype (all ZMUM). A. left anterior telopod, anterior view; B. left anterior telopod, lateral view; C. left anterior telopod, posterior view; D. leg 9, right, anterior view; E. right posterior telopod, posterior view; F. right posterior telopod, anterior view; G. telopoditomere 4, lateral view; H. right male antenna; I. left coxa and prefemur with vulva. Scale bar = 1 mm." httpUri="https://zenodo.org/record/1458883/files/figure.png" pageId="2" pageNumber="537">
Fig.
<date id="FFC07BD4FFDA2423CE5EE183DDBBFBA3" box="[1396,1423,1015,1039]" pageId="2" pageNumber="537">2I</date>
</figureCitation>
). Immovable finger of posterior telopod swollen apically, but extending only mesally (
<figureCitation id="13454191FFDA2423CF42E66EDCD2FB9F" box="[1128,1254,1050,1075]" captionStart="FIGURE 2" captionStartId="4.[151,249,1919,1941]" captionTargetBox="[191,1429,686,1892]" captionTargetId="figure@4.[176,1429,657,1899]" captionTargetPageId="4" captionText="FIGURE 2. Sphaerobelum cattiense sp. nov. AG = male paratypes, I = female paratype (all ZMUM). A. left anterior telopod, anterior view; B. left anterior telopod, lateral view; C. left anterior telopod, posterior view; D. leg 9, right, anterior view; E. right posterior telopod, posterior view; F. right posterior telopod, anterior view; G. telopoditomere 4, lateral view; H. right male antenna; I. left coxa and prefemur with vulva. Scale bar = 1 mm." httpUri="https://zenodo.org/record/1458883/files/figure.png" pageId="2" pageNumber="537">Figs 2E, F</figureCitation>
). Also differing from
<taxonomicName id="4C7E2697FFDA2423CBFEE64BD972FBFA" authorityName="Attems" authorityYear="1938" box="[212,326,1086,1110]" class="Diplopoda" family="Zephroniidae" genus="Sphaerobelum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sphaerotheriida" pageId="2" pageNumber="537" phylum="Arthropoda" rank="species" species="bicorne">
<emphasis id="B90A8106FFDA2423CBFEE64BD972FBFA" box="[212,326,1086,1110]" italics="true" pageId="2" pageNumber="537">S. bicorne</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7E2697FFDA2423CA56E64BDA09FBFA" authority="Semenyuk &amp; Golovatch &amp; Wesener, 2018" authorityName="Semenyuk &amp; Golovatch &amp; Wesener" authorityYear="2018" box="[380,573,1086,1110]" class="Diplopoda" family="Zephroniidae" genus="Sphaerobelum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sphaerotheriida" pageId="2" pageNumber="537" phylum="Arthropoda" rank="species" species="konkakinhense" status="sp. nov.">
<emphasis id="B90A8106FFDA2423CA56E64BDA09FBFA" box="[380,573,1086,1110]" italics="true" pageId="2" pageNumber="537">S. konkakinhense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2393C7DFFDA2423C96EE64ADAA9FBFB" box="[580,669,1086,1111]" pageId="2" pageNumber="537" rank="species">
<emphasis id="B90A8106FFDA2423C96EE64ADAA9FBFB" bold="true" box="[580,669,1086,1111]" pageId="2" pageNumber="537">sp. nov.</emphasis>
</taxonomicNameLabel>
in the absence of a lateral projection on the prefemur of the walking legs.
</paragraph>
</subSubSection>
<subSubSection id="C3640E9FFFDA2425CBEDE6F2DBCEFD29" lastPageId="4" lastPageNumber="539" pageId="2" pageNumber="537" type="description">
<paragraph id="8BC15D14FFDA2422CBEDE6F2DA14FE24" blockId="2.[151,1437,654,2011]" lastBlockId="3.[151,1436,151,392]" lastPageId="3" lastPageNumber="538" pageId="2" pageNumber="537">
<emphasis id="B90A8106FFDA2423CBEDE6F2DA3BFB33" bold="true" box="[199,527,1158,1183]" pageId="2" pageNumber="537">Description. Measurements</emphasis>
:
<typeStatus id="54C5E3B6FFDA2423C935E6F2DAB3FB33" box="[543,647,1158,1183]" pageId="2" pageNumber="537" type="holotype">Holotype</typeStatus>
male. Body length ca
<quantity id="4C86F0F1FFDA2423C8A2E6F3DBEEFB32" box="[904,986,1159,1183]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="537" unit="mm" value="30.0">30 mm</quantity>
, width of thoracic shield =
<quantity id="4C86F0F1FFDA2423CE38E6F2DD4DFB32" box="[1298,1401,1158,1183]" metricMagnitude="-2" metricUnit="m" metricValue="1.46" pageId="2" pageNumber="537" unit="mm" value="14.6">14.6 mm</quantity>
, of tergite 9 =
<quantity id="4C86F0F1FFDA2423CA30E6DED9B5FB6E" box="[282,385,1194,1219]" metricMagnitude="-2" metricUnit="m" metricValue="1.52" pageId="2" pageNumber="537" unit="mm" value="15.2">15.2 mm</quantity>
(= broadest), height of thoracic shield =
<quantity id="4C86F0F1FFDA2423C849E6DEDB8AFB6E" box="[867,958,1194,1219]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="2" pageNumber="537" unit="mm" value="8.5">8.5 mm</quantity>
, of tergite 9 =
<quantity id="4C86F0F1FFDA2423CF5BE6DFDCFEFB6E" box="[1137,1226,1195,1219]" metricMagnitude="-3" metricUnit="m" metricValue="9.3" pageId="2" pageNumber="537" unit="mm" value="9.3">9.3 mm</quantity>
(= highest). Male
<typeStatus id="54C5E3B6FFDA2423CBBDE6BBD936FB4B" box="[151,258,1231,1255]" pageId="2" pageNumber="537" type="paratype">paratypes</typeStatus>
ca 25 or
<quantity id="4C86F0F1FFDA2423CA5EE6BBD9F1FB4A" box="[372,453,1231,1255]" metricMagnitude="-2" metricUnit="m" metricValue="2.9" pageId="2" pageNumber="537" unit="mm" value="29.0">29 mm</quantity>
long, maximal width ca 17 or
<quantity id="4C86F0F1FFDA2423C804E6BADBB5FB4A" box="[814,897,1230,1255]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="537" unit="mm" value="15.0">15 mm</quantity>
, respectively. Females: length ca
<quantity id="4C86F0F1FFDA2423CE2FE6BADDACFB4A" box="[1285,1432,1230,1255]" metricMagnitude="-2" metricUnit="m" metricValue="2.875" metricValueMax="3.0" metricValueMin="2.75" pageId="2" pageNumber="537" unit="mm" value="28.75" valueMax="30.0" valueMin="27.5">27.530 mm</quantity>
, width of thoracic shield =
<quantity id="4C86F0F1FFDA2423CA95E686DA64FAA6" box="[447,592,1266,1291]" metricMagnitude="-2" metricUnit="m" metricValue="1.515" metricValueMax="1.6" metricValueMin="1.43" pageId="2" pageNumber="537" unit="mm" value="15.15" valueMax="16.0" valueMin="14.3">14.316 mm</quantity>
, of tergite 9 =
<quantity id="4C86F0F1FFDA2423C9DFE686DBB7FAA6" box="[757,899,1266,1291]" metricMagnitude="-2" metricUnit="m" metricValue="1.6149999999999998" metricValueMax="1.7" metricValueMin="1.53" pageId="2" pageNumber="537" unit="mm" value="16.15" valueMax="17.0" valueMin="15.3">15.317 mm</quantity>
(=broadest), height of thoracic shield = 8.69.0 mm, of tergite 9 =
<quantity id="4C86F0F1FFDA2423CA51E763D9CBFA82" box="[379,511,1303,1327]" metricMagnitude="-3" metricUnit="m" metricValue="9.65" metricValueMax="10.0" metricValueMin="9.3" pageId="2" pageNumber="537" unit="mm" value="9.65" valueMax="10.0" valueMin="9.3">9.310 mm</quantity>
(= highest).
<emphasis id="B90A8106FFDA2423C9B3E762DB2EFA83" bold="true" box="[665,794,1302,1327]" pageId="2" pageNumber="537">Coloration</emphasis>
: in live specimens, ventral side including head and all tergites blackish brown; collum, tergites and posterior part of anal shield brown. Thoracic shield brown at margins. Legs and antennae dark orange. In preserved specimens, dark olive to olive brown with contrasting, narrow, darkish, orange bands at caudal margins of all tergites (
<figureCitation id="13454191FFDA2423C82DE7F6DB55FA37" box="[775,865,1410,1435]" captionStart="FIGURE 1" captionStartId="3.[151,249,1736,1758]" captionTargetBox="[152,1435,447,1716]" captionTargetId="figure@3.[152,1435,447,1716]" captionTargetPageId="3" captionText="FIGURE 1. Distribution map with pictures of live specimens. A. Sphaerobelum cattiense sp. nov.; B. Sphaerobelum konkakinhense sp. nov.; C. Zephronia konkakinhensis sp. nov.; D. Zephronia montis sp. nov." httpUri="https://zenodo.org/record/1458881/files/figure.png" pageId="2" pageNumber="537">Fig. 1A</figureCitation>
). Head dark brown to dark olive-brown. Antennae and legs contrasting bright orange.
<emphasis id="B90A8106FFDA2423C90DE7D2DA53FA13" bold="true" box="[551,615,1446,1471]" pageId="2" pageNumber="537">Head</emphasis>
: wide and short, partly micropunctate. Clypeolabral region very densely setose. Occipital region finely and confusedly striolate and with a row of longer hairs near caudal edge. Eyes with&gt;65 ocelli. Aberrant ocellus located inside antennal groove. Antennae short, with rounded joints, extending posteriorly to leg-pair 2. First antennomere with cuticular scales (
<figureCitation id="13454191FFDA2423C8F7E466DC76F987" box="[989,1090,1554,1579]" captionStart="FIGURE 3" captionStartId="5.[151,249,662,684]" captionTargetBox="[166,1425,189,646]" captionTargetId="figure@5.[166,1426,187,647]" captionTargetPageId="5" captionText="FIGURE 3. Sphaerobelum cattiense sp. nov. Male paratype (ZFMK MYR6516), right antenna. A. lateral view; B. apical view of disc; C. details of apical cone. Abbreviations: sc = sclerotized scales; sb = sensilla basiconica. Scale bars: A = 1000 µm; B = 200 µm; C = 10 µm." httpUri="https://zenodo.org/record/1458885/files/figure.png" pageId="2" pageNumber="537">Fig. 3A</figureCitation>
). Lengths of antennomeres: 1&gt;2=3=4=5&lt;&lt;6 (
<figureCitation id="13454191FFDA2423CA72E442D985F9E3" box="[344,433,1590,1615]" captionStart="FIGURE 3" captionStartId="5.[151,249,662,684]" captionTargetBox="[166,1425,189,646]" captionTargetId="figure@5.[166,1426,187,647]" captionTargetPageId="5" captionText="FIGURE 3. Sphaerobelum cattiense sp. nov. Male paratype (ZFMK MYR6516), right antenna. A. lateral view; B. apical view of disc; C. details of apical cone. Abbreviations: sc = sclerotized scales; sb = sensilla basiconica. Scale bars: A = 1000 µm; B = 200 µm; C = 10 µm." httpUri="https://zenodo.org/record/1458885/files/figure.png" pageId="2" pageNumber="537">Fig. 3A</figureCitation>
). Antennomere 6 densely pubescent, sensilla basiconica surrounding apical disc. Shape of antennae sexually dimorphic: cylindrical in female; thickened, almost axe-shaped in male (
<figureCitation id="13454191FFDA2423CF81E42EDD30F9DF" box="[1195,1284,1626,1651]" captionStart="FIGURE 3" captionStartId="5.[151,249,662,684]" captionTargetBox="[166,1425,189,646]" captionTargetId="figure@5.[166,1426,187,647]" captionTargetPageId="5" captionText="FIGURE 3. Sphaerobelum cattiense sp. nov. Male paratype (ZFMK MYR6516), right antenna. A. lateral view; B. apical view of disc; C. details of apical cone. Abbreviations: sc = sclerotized scales; sb = sensilla basiconica. Scale bars: A = 1000 µm; B = 200 µm; C = 10 µm." httpUri="https://zenodo.org/record/1458885/files/figure.png" pageId="2" pageNumber="537">Fig. 3B</figureCitation>
). Apical disc with ca 105/109 (male) (
<figureCitation id="13454191FFDA2423CA83E40ADA32F93B" box="[425,518,1662,1687]" captionStart="FIGURE 2" captionStartId="4.[151,249,1919,1941]" captionTargetBox="[191,1429,686,1892]" captionTargetId="figure@4.[176,1429,657,1899]" captionTargetPageId="4" captionText="FIGURE 2. Sphaerobelum cattiense sp. nov. AG = male paratypes, I = female paratype (all ZMUM). A. left anterior telopod, anterior view; B. left anterior telopod, lateral view; C. left anterior telopod, posterior view; D. leg 9, right, anterior view; E. right posterior telopod, posterior view; F. right posterior telopod, anterior view; G. telopoditomere 4, lateral view; H. right male antenna; I. left coxa and prefemur with vulva. Scale bar = 1 mm." httpUri="https://zenodo.org/record/1458883/files/figure.png" pageId="2" pageNumber="537">Figs 2H</figureCitation>
,
<figureCitation id="13454191FFDA2423C93BE40BDA07F93B" box="[529,563,1663,1687]" captionStart="FIGURE 3" captionStartId="5.[151,249,662,684]" captionTargetBox="[166,1425,189,646]" captionTargetId="figure@5.[166,1426,187,647]" captionTargetPageId="5" captionText="FIGURE 3. Sphaerobelum cattiense sp. nov. Male paratype (ZFMK MYR6516), right antenna. A. lateral view; B. apical view of disc; C. details of apical cone. Abbreviations: sc = sclerotized scales; sb = sensilla basiconica. Scale bars: A = 1000 µm; B = 200 µm; C = 10 µm." httpUri="https://zenodo.org/record/1458885/files/figure.png" pageId="2" pageNumber="537">3B</figureCitation>
) or 5661 (female) apical cones. Apical cones typical of Diplopoda (
<figureCitation id="13454191FFDA2423CE1DE40ADDBBF93B" box="[1335,1423,1662,1687]" captionStart="FIGURE 3" captionStartId="5.[151,249,662,684]" captionTargetBox="[166,1425,189,646]" captionTargetId="figure@5.[166,1426,187,647]" captionTargetPageId="5" captionText="FIGURE 3. Sphaerobelum cattiense sp. nov. Male paratype (ZFMK MYR6516), right antenna. A. lateral view; B. apical view of disc; C. details of apical cone. Abbreviations: sc = sclerotized scales; sb = sensilla basiconica. Scale bars: A = 1000 µm; B = 200 µm; C = 10 µm." httpUri="https://zenodo.org/record/1458885/files/figure.png" pageId="2" pageNumber="537">Fig. 3C</figureCitation>
). Organ of Tömösváry located inside antennal groove.
<emphasis id="B90A8106FFDA2423C9C4E4D6DB89F917" bold="true" box="[750,957,1698,1723]" pageId="2" pageNumber="537">Gnathochilarium</emphasis>
: structure typical of the order. Stipites and paramentum bearing several long setae.
<emphasis id="B90A8106FFDA2423C9A8E4B2DB34F973" bold="true" box="[642,768,1734,1759]" pageId="2" pageNumber="537">Mandibles</emphasis>
(not examined).
<emphasis id="B90A8106FFDA2423C8FDE4B2DCA9F973" bold="true" box="[983,1181,1734,1759]" pageId="2" pageNumber="537">Stigmatic plates</emphasis>
: first stigmatic plate triangular, apex well-rounded, weakly curved towards coxa 1.
<emphasis id="B90A8106FFDA2423C84EE49EDC39F8AF" bold="true" box="[868,1037,1770,1795]" pageId="2" pageNumber="537">Laterotergites</emphasis>
: laterotergite 1 slightly projecting, well-rounded and slender at tip.
<emphasis id="B90A8106FFDA2423C92AE57ADA6CF88B" bold="true" box="[512,600,1806,1831]" pageId="2" pageNumber="537">Collum</emphasis>
with 23 rows of sparse setae near caudal margin and a few short, isolated setae on surface.
<emphasis id="B90A8106FFDA2423CA77E546DA23F8E7" bold="true" box="[349,535,1842,1867]" pageId="2" pageNumber="537">Thoracic shield</emphasis>
: surface like that of tergites. Shallow grooves beset with numerous long setae, sloping towards groove with 5 keels.
<emphasis id="B90A8106FFDA2423C912E522DAA1F8C3" bold="true" box="[568,661,1878,1903]" pageId="2" pageNumber="537">Tergites</emphasis>
: surface very densely micropunctate, many of the punctures carrying a tiny seta. Tips of paratergites of midbody tergites projecting posteriorly.
<emphasis id="B90A8106FFDA2423C8F1E50EDC46F83F" bold="true" box="[987,1138,1914,1939]" pageId="2" pageNumber="537">Endotergum</emphasis>
: inner section lacking any large spines or setae. Middle area with a single row of large, sparse, elliptical, cuticular impressions. Apically, 23 dense rows of short marginal bristles, a few with their tips slightly protruding beyond tergal margin (
<figureCitation id="13454191FFDA2423CE18E5B6DDA4F877" box="[1330,1424,1986,2011]" captionStart="FIGURE 4" captionStartId="6.[151,249,1534,1556]" captionTargetBox="[176,1427,837,1514]" captionTargetId="figure@6.[176,1428,837,1514]" captionTargetPageId="6" captionText="FIGURE 4. Endoterga of midbody rings. A. Sphaerobelum cattiense sp. nov. Male paratype (ZFMK MYR6516); B. Sphaerobelum konkakinhense sp. nov., male holotype (ZMUM). Scale bars: 100 µm." httpUri="https://zenodo.org/record/1458887/files/figure.png" pageId="2" pageNumber="537">Fig. 4A</figureCitation>
). Bristles not smooth, but with numerous small spicules (
<figureCitation id="13454191FFDB2422C828E2E3DB68FF1C" box="[770,860,151,176]" captionStart="FIGURE 4" captionStartId="6.[151,249,1534,1556]" captionTargetBox="[176,1427,837,1514]" captionTargetId="figure@6.[176,1428,837,1514]" captionTargetPageId="6" captionText="FIGURE 4. Endoterga of midbody rings. A. Sphaerobelum cattiense sp. nov. Male paratype (ZFMK MYR6516); B. Sphaerobelum konkakinhense sp. nov., male holotype (ZMUM). Scale bars: 100 µm." httpUri="https://zenodo.org/record/1458887/files/figure.png" pageId="3" pageNumber="538">Fig. 4A</figureCitation>
).
<emphasis id="B90A8106FFDB2422C845E2E3DBC2FF1C" bold="true" box="[879,1014,151,176]" pageId="3" pageNumber="538">Anal shield</emphasis>
: large and regularly rounded. Surface similar to that of tergites. Underside with a single, short, black, locking carina, the latter located close to last laterotergite.
<emphasis id="B90A8106FFDB2422CA06E2ABD950FF54" bold="true" box="[300,356,223,248]" pageId="3" pageNumber="538">Legs</emphasis>
: leg-pair 1 with 2 or 3, leg-pair 2 with 3 or 4, leg-pair 3 with 46 ventral spines. First two legpairs without an apical spine. Leg-pairs 421 with 710 ventral spines and one apical spine. On leg-pair 9, femur 1.5X, tarsus 3.4X longer than wide (
<figureCitation id="13454191FFDB2422C91FE353DAA5FEEC" box="[565,657,295,320]" captionStart="FIGURE 2" captionStartId="4.[151,249,1919,1941]" captionTargetBox="[191,1429,686,1892]" captionTargetId="figure@4.[176,1429,657,1899]" captionTargetPageId="4" captionText="FIGURE 2. Sphaerobelum cattiense sp. nov. AG = male paratypes, I = female paratype (all ZMUM). A. left anterior telopod, anterior view; B. left anterior telopod, lateral view; C. left anterior telopod, posterior view; D. leg 9, right, anterior view; E. right posterior telopod, posterior view; F. right posterior telopod, anterior view; G. telopoditomere 4, lateral view; H. right male antenna; I. left coxa and prefemur with vulva. Scale bar = 1 mm." httpUri="https://zenodo.org/record/1458883/files/figure.png" pageId="3" pageNumber="538">Fig. 2D</figureCitation>
). All podomeres densely setose. Coxa with a large and well-rounded process. Prefemur of usual shape, but at apical margin with a mesal projection. Femur extended mesally into a dentate margin featuring 68 teeth.
</paragraph>
<caption id="DF010D9CFFDB2422CBBDE4BCDC7CF950" httpUri="https://zenodo.org/record/1458881/files/figure.png" pageId="3" pageNumber="538" startId="3.[151,249,1736,1758]" targetBox="[152,1435,447,1716]" targetPageId="3">
<paragraph id="8BC15D14FFDB2422CBBDE4BCDC7CF950" blockId="3.[151,1436,1736,1789]" pageId="3" pageNumber="538">
<emphasis id="B90A8106FFDB2422CBBDE4BCD92DF972" bold="true" box="[151,281,1736,1758]" pageId="3" pageNumber="538">FIGURE 1.</emphasis>
Distribution map with pictures of live specimens.
<emphasis id="B90A8106FFDB2422C864E4BCDB54F972" bold="true" box="[846,864,1736,1758]" pageId="3" pageNumber="538">A</emphasis>
.
<taxonomicName id="4C7E2697FFDB2422C85EE4BCDC59F972" authority="Semenyuk &amp; Golovatch &amp; Wesener, 2018" authorityName="Semenyuk &amp; Golovatch &amp; Wesener" authorityYear="2018" box="[884,1133,1736,1758]" class="Diplopoda" family="Zephroniidae" genus="Sphaerobelum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sphaerotheriida" pageId="3" pageNumber="538" phylum="Arthropoda" rank="species" species="cattiense" status="sp. nov.">
<emphasis id="B90A8106FFDB2422C85EE4BCDC59F972" box="[884,1133,1736,1758]" italics="true" pageId="3" pageNumber="538">Sphaerobelum cattiense</emphasis>
</taxonomicName>
<emphasis id="B90A8106FFDB2422CF53E4BCDCC3F972" bold="true" box="[1145,1271,1736,1758]" pageId="3" pageNumber="538">
<taxonomicNameLabel id="A2393C7DFFDB2422CF53E4BCDCE0F972" box="[1145,1236,1736,1758]" pageId="3" pageNumber="538" rank="species">sp. nov.</taxonomicNameLabel>
; B
</emphasis>
.
<taxonomicName id="4C7E2697FFDB2422CE21E4BCD918F950" authority="Semenyuk &amp; Golovatch &amp; Wesener, 2018" authorityName="Semenyuk &amp; Golovatch &amp; Wesener" authorityYear="2018" class="Diplopoda" family="Zephroniidae" genus="Sphaerobelum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sphaerotheriida" pageId="3" pageNumber="538" phylum="Arthropoda" rank="species" species="konkakinhense" status="sp. nov.">
<emphasis id="B90A8106FFDB2422CE21E4BCD918F950" italics="true" pageId="3" pageNumber="538">Sphaerobelum konkakinhense</emphasis>
</taxonomicName>
<emphasis id="B90A8106FFDB2422CA18E492D990F950" bold="true" box="[306,420,1766,1788]" pageId="3" pageNumber="538">
<taxonomicNameLabel id="A2393C7DFFDB2422CA18E492D9B2F950" box="[306,390,1766,1788]" pageId="3" pageNumber="538" rank="species">sp. nov.</taxonomicNameLabel>
; C
</emphasis>
.
<taxonomicName id="4C7E2697FFDB2422CA9BE492DA8FF950" authority="Semenyuk &amp; Golovatch &amp; Wesener, 2018" authorityName="Semenyuk &amp; Golovatch &amp; Wesener" authorityYear="2018" box="[433,699,1766,1788]" class="Diplopoda" family="Zephroniidae" genus="Zephronia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sphaerotheriida" pageId="3" pageNumber="538" phylum="Arthropoda" rank="species" species="konkakinhensis" status="sp. nov.">
<emphasis id="B90A8106FFDB2422CA9BE492DA8FF950" box="[433,699,1766,1788]" italics="true" pageId="3" pageNumber="538">Zephronia konkakinhensis</emphasis>
</taxonomicName>
<emphasis id="B90A8106FFDB2422C9EAE492DB06F950" bold="true" box="[704,818,1766,1788]" pageId="3" pageNumber="538">
<taxonomicNameLabel id="A2393C7DFFDB2422C9EAE492DB21F950" box="[704,789,1766,1788]" pageId="3" pageNumber="538" rank="species">sp. nov.</taxonomicNameLabel>
; D
</emphasis>
.
<taxonomicName id="4C7E2697FFDB2422C815E492DBC6F951" authority="Semenyuk &amp; Golovatch &amp; Wesener, 2018" authorityName="Semenyuk &amp; Golovatch &amp; Wesener" authorityYear="2018" box="[831,1010,1766,1789]" class="Diplopoda" family="Zephroniidae" genus="Zephronia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sphaerotheriida" pageId="3" pageNumber="538" phylum="Arthropoda" rank="species" species="montis" status="sp. nov.">
<emphasis id="B90A8106FFDB2422C815E492DBC6F951" box="[831,1010,1766,1789]" italics="true" pageId="3" pageNumber="538">Zephronia montis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2393C7DFFDB2422C8D2E492DC7CF950" box="[1016,1096,1766,1788]" pageId="3" pageNumber="538" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8BC15D14FFDB2422CBEDE55DDD45F802" blockId="3.[151,1436,1833,2038]" pageId="3" pageNumber="538">
<emphasis id="B90A8106FFDB2422CBEDE55DD9C1F8EE" bold="true" box="[199,501,1833,1858]" pageId="3" pageNumber="538">Female sexual characters</emphasis>
: vulva large, covering 2/3 of coxa, extending mesally to half of prefemur (
<figureCitation id="13454191FFDB2422CE6AE55DDDBBF8EE" box="[1344,1423,1833,1858]" captionStart="FIGURE 2" captionStartId="4.[151,249,1919,1941]" captionTargetBox="[191,1429,686,1892]" captionTargetId="figure@4.[176,1429,657,1899]" captionTargetPageId="4" captionText="FIGURE 2. Sphaerobelum cattiense sp. nov. AG = male paratypes, I = female paratype (all ZMUM). A. left anterior telopod, anterior view; B. left anterior telopod, lateral view; C. left anterior telopod, posterior view; D. leg 9, right, anterior view; E. right posterior telopod, posterior view; F. right posterior telopod, anterior view; G. telopoditomere 4, lateral view; H. right male antenna; I. left coxa and prefemur with vulva. Scale bar = 1 mm." httpUri="https://zenodo.org/record/1458883/files/figure.png" pageId="3" pageNumber="538">
Fig.
<date id="FFC07BD4FFDB2422CE5EE55EDDBBF8EE" box="[1396,1423,1834,1858]" pageId="3" pageNumber="538">2I</date>
</figureCitation>
). Operculum rounded, very slightly invaginated medially, mesal margin strongly projecting into a well-rounded lobe twice as high as remaining operculum.
<emphasis id="B90A8106FFDB2422C970E505DB34F826" bold="true" box="[602,768,1905,1930]" pageId="3" pageNumber="538">Subanal plate</emphasis>
: large, shaped like a wide, well-rounded triangle, in some females with a slight invagination at the highest point of apical margin, creating two wide, well-rounded lobes.
</paragraph>
<paragraph id="8BC15D14FFDB2422CBEDE5CDDDA8F859" blockId="3.[151,1436,1833,2038]" pageId="3" pageNumber="538">
<emphasis id="B90A8106FFDB2422CBEDE5CDD9D4F87E" bold="true" box="[199,480,1977,2002]" pageId="3" pageNumber="538">Male sexual characters</emphasis>
: gonopore covered with a single, undivided, triangular, sclerotized plate.
<emphasis id="B90A8106FFDB2422CE1EE5CDD8CEF85A" bold="true" pageId="3" pageNumber="538">Anterior telopods</emphasis>
(
<figureCitation id="13454191FFDB2422CA20E5A9D9B9F85A" box="[266,397,2013,2038]" captionStart="FIGURE 2" captionStartId="4.[151,249,1919,1941]" captionTargetBox="[191,1429,686,1892]" captionTargetId="figure@4.[176,1429,657,1899]" captionTargetPageId="4" captionText="FIGURE 2. Sphaerobelum cattiense sp. nov. AG = male paratypes, I = female paratype (all ZMUM). A. left anterior telopod, anterior view; B. left anterior telopod, lateral view; C. left anterior telopod, posterior view; D. leg 9, right, anterior view; E. right posterior telopod, posterior view; F. right posterior telopod, anterior view; G. telopoditomere 4, lateral view; H. right male antenna; I. left coxa and prefemur with vulva. Scale bar = 1 mm." httpUri="https://zenodo.org/record/1458883/files/figure.png" pageId="3" pageNumber="538">Figs 2AC</figureCitation>
): consisting of a syncoxite plus 4 telopoditomeres. Telopoditomere 1 rectangular, as long as
</paragraph>
<paragraph id="8BC15D14FFDC2425CBBDE2E3DBCEFD29" blockId="4.[151,1437,151,645]" pageId="4" pageNumber="539">
wide. Telopoditomere 2 large, as long as telopoditomeres 3 and 4 combined. Process of telopoditomere 2 located posteriorly, not visible in anterior view. Process slender, projecting as far as base of telopoditomere 4, apical part well-rounded. Telopoditomere 3 shortest, as long as wide, anterior side with a few small, triangular spines, posterior side smooth. Telopoditomere 4 well-rounded, twice as long as wide, slightly tapering apically. Posterior aspect with 5 or 6 black spines, apical-most being the largest. All podomeres covered with long setae.
<emphasis id="B90A8106FFDC2425CE07E353D8CEFEC9" bold="true" pageId="4" pageNumber="539">Posterior telopods</emphasis>
(
<figureCitation id="13454191FFDC2425CA39E338D9A8FEC8" box="[275,412,332,357]" captionStart="FIGURE 2" captionStartId="4.[151,249,1919,1941]" captionTargetBox="[191,1429,686,1892]" captionTargetId="figure@4.[176,1429,657,1899]" captionTargetPageId="4" captionText="FIGURE 2. Sphaerobelum cattiense sp. nov. AG = male paratypes, I = female paratype (all ZMUM). A. left anterior telopod, anterior view; B. left anterior telopod, lateral view; C. left anterior telopod, posterior view; D. leg 9, right, anterior view; E. right posterior telopod, posterior view; F. right posterior telopod, anterior view; G. telopoditomere 4, lateral view; H. right male antenna; I. left coxa and prefemur with vulva. Scale bar = 1 mm." httpUri="https://zenodo.org/record/1458883/files/figure.png" pageId="4" pageNumber="539">Figs 2EG</figureCitation>
): telopoditomere 1 elongate, twice as long as wide. Immovable finger (process of telopoditomere 2) shorter than movable finger, consisting of telopoditomeres 3 and 4. Immovable finger with a characteristic, distally swollen apex, extending only mesally, apex therefore only slightly broader than base. Telopoditomere 3 elongate, towards immovable finger in apical half with a large extension carrying at least two large spines. Telopoditomere 4, 2.8X longer than wide, tapering apically, slightly curved towards immovable finger. At inner margin, telopoditomere 4 with a large spine at midpoint and 3 or 4 smaller spines apically. Entire telopodite on anterior side covered with long setae, except for apical part of telopoditomere 3 and telopoditomere 4. In posterior view, telopoditomere 1 clothed with numerous long setae, telopoditomere 2 only basally and at lateral margin clothed with setae, telopoditomere 3 with setae only at lateral margin.
</paragraph>
</subSubSection>
<caption id="DF010D9CFFDC2425CBBDE50BDB1BF85D" httpUri="https://zenodo.org/record/1458883/files/figure.png" pageId="4" pageNumber="539" startId="4.[151,249,1919,1941]" targetBox="[191,1429,686,1892]" targetPageId="4">
<paragraph id="8BC15D14FFDC2425CBBDE50BDB1BF85D" blockId="4.[151,1436,1919,2033]" pageId="4" pageNumber="539">
<emphasis id="B90A8106FFDC2425CBBDE50BD925F839" bold="true" box="[151,273,1919,1941]" pageId="4" pageNumber="539">FIGURE 2.</emphasis>
<taxonomicName id="4C7E2697FFDC2425CA32E50BDA3DF839" authority="Semenyuk &amp; Golovatch &amp; Wesener, 2018" authorityName="Semenyuk &amp; Golovatch &amp; Wesener" authorityYear="2018" box="[280,521,1919,1941]" class="Diplopoda" family="Zephroniidae" genus="Sphaerobelum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sphaerotheriida" pageId="4" pageNumber="539" phylum="Arthropoda" rank="species" species="cattiense" status="sp. nov.">
<emphasis id="B90A8106FFDC2425CA32E50BDA3DF839" box="[280,521,1919,1941]" italics="true" pageId="4" pageNumber="539">Sphaerobelum cattiense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2393C7DFFDC2425C93AE50BDA54F839" box="[528,608,1919,1941]" pageId="4" pageNumber="539" rank="species">
<emphasis id="B90A8106FFDC2425C93AE50BDA54F839" bold="true" box="[528,608,1919,1941]" pageId="4" pageNumber="539">sp. nov.</emphasis>
</taxonomicNameLabel>
AG = male paratypes, I = female paratype (all ZMUM).
<emphasis id="B90A8106FFDC2425CF84E50BDCF4F839" bold="true" box="[1198,1216,1919,1941]" pageId="4" pageNumber="539">A</emphasis>
. left anterior telopod, anterior view;
<emphasis id="B90A8106FFDC2425CA04E5E9D90BF81F" bold="true" box="[302,319,1949,1971]" pageId="4" pageNumber="539">B</emphasis>
. left anterior telopod, lateral view;
<emphasis id="B90A8106FFDC2425C985E5E9DAF5F81F" bold="true" box="[687,705,1949,1971]" pageId="4" pageNumber="539">C</emphasis>
. left anterior telopod, posterior view;
<emphasis id="B90A8106FFDC2425CF67E5E9DC6BF81F" bold="true" box="[1101,1119,1949,1971]" pageId="4" pageNumber="539">D</emphasis>
. leg 9, right, anterior view;
<emphasis id="B90A8106FFDC2425CEAEE5E9DDA1F81F" bold="true" box="[1412,1429,1949,1971]" pageId="4" pageNumber="539">E</emphasis>
. right posterior telopod, posterior view;
<emphasis id="B90A8106FFDC2425C909E5C8DA04F87E" bold="true" box="[547,560,1980,2002]" pageId="4" pageNumber="539">F</emphasis>
. right posterior telopod, anterior view;
<emphasis id="B90A8106FFDC2425C897E5C8DBFAF87E" bold="true" box="[957,974,1980,2002]" pageId="4" pageNumber="539">G</emphasis>
. telopoditomere 4, lateral view;
<emphasis id="B90A8106FFDC2425CE3FE5C8DD1DF87E" bold="true" box="[1301,1321,1980,2002]" pageId="4" pageNumber="539">H</emphasis>
. right male antenna;
<emphasis id="B90A8106FFDC2425CBD9E5AFD8C9F85D" bold="true" box="[243,253,2011,2033]" pageId="4" pageNumber="539">I</emphasis>
. left coxa and prefemur with vulva.
<emphasis id="B90A8106FFDC2425C944E5AFDAE6F85D" bold="true" box="[622,722,2011,2033]" pageId="4" pageNumber="539">Scale bar</emphasis>
= 1 mm.
</paragraph>
</caption>
<caption id="DF010D9CFFDD2424CBBDE0E2D948FD45" httpUri="https://zenodo.org/record/1458885/files/figure.png" pageId="5" pageNumber="540" startId="5.[151,249,662,684]" targetBox="[166,1425,189,646]" targetPageId="5">
<paragraph id="8BC15D14FFDD2424CBBDE0E2D948FD45" blockId="5.[151,1436,662,745]" pageId="5" pageNumber="540">
<emphasis id="B90A8106FFDD2424CBBDE0E2D926FD00" bold="true" box="[151,274,662,684]" pageId="5" pageNumber="540">FIGURE 3.</emphasis>
<taxonomicName id="4C7E2697FFDD2424CA32E0E2DA3EFD00" authority="Semenyuk &amp; Golovatch &amp; Wesener, 2018" authorityName="Semenyuk &amp; Golovatch &amp; Wesener" authorityYear="2018" box="[280,522,662,684]" class="Diplopoda" family="Zephroniidae" genus="Sphaerobelum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sphaerotheriida" pageId="5" pageNumber="540" phylum="Arthropoda" rank="species" species="cattiense" status="sp. nov.">
<emphasis id="B90A8106FFDD2424CA32E0E2DA3EFD00" box="[280,522,662,684]" italics="true" pageId="5" pageNumber="540">Sphaerobelum cattiense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2393C7DFFDD2424C93AE0E2DA55FD00" box="[528,609,662,684]" pageId="5" pageNumber="540" rank="species">
<emphasis id="B90A8106FFDD2424C93AE0E2DA55FD00" bold="true" box="[528,609,662,684]" pageId="5" pageNumber="540">sp. nov.</emphasis>
</taxonomicNameLabel>
Male paratype (ZFMK MYR6516), right antenna.
<emphasis id="B90A8106FFDD2424CF4DE0E2DC4DFD00" bold="true" box="[1127,1145,662,684]" pageId="5" pageNumber="540">A</emphasis>
. lateral view;
<emphasis id="B90A8106FFDD2424CE20E0E2DD2FFD00" bold="true" box="[1290,1307,662,684]" pageId="5" pageNumber="540">B</emphasis>
. apical view of disc;
<emphasis id="B90A8106FFDD2424CBC2E0C0D8CEFD66" bold="true" box="[232,250,692,714]" pageId="5" pageNumber="540">C</emphasis>
. details of apical cone.
<emphasis id="B90A8106FFDD2424CAC0E0C0DAB7FD66" bold="true" box="[490,643,692,714]" pageId="5" pageNumber="540">Abbreviations</emphasis>
: sc = sclerotized scales; sb = sensilla basiconica.
<emphasis id="B90A8106FFDD2424CF53E0C0DCDCFD66" bold="true" box="[1145,1256,692,714]" pageId="5" pageNumber="540">Scale bars</emphasis>
: A = 1000 µm; B = 200 µm; C = 10 µm.
</paragraph>
</caption>
<subSubSection id="C3640E9FFFDD2424CBEDE165DC7AFBC1" pageId="5" pageNumber="540" type="distribution">
<paragraph id="8BC15D14FFDD2424CBEDE165DA0AFBE6" blockId="5.[151,1437,785,1134]" pageId="5" pageNumber="540">
<emphasis id="B90A8106FFDD2424CBEDE165D9C3FC86" bold="true" box="[199,503,785,810]" pageId="5" pageNumber="540">Distribution and Ecology</emphasis>
. This species is widespread exclusively in patches of mature, lowland, monsoon forest at elevations of about
<specimenCount id="9D78969DFFDD2424CAFFE141DA10FCE1" box="[469,548,821,845]" pageId="5" pageNumber="540" type="generic">
<quantity id="4C86F0F1FFDD2424CAFFE141DA2BFCE1" box="[469,543,821,845]" metricMagnitude="2" metricUnit="m" metricValue="1.2" pageId="5" pageNumber="540" unit="m" value="120.0">120 m</quantity>
.
</specimenCount>
a.s.l. Active animals prefer patches with less leaf litter and usually walk on the forest floor, sometimes digging into the leaf litter layer. Resting individuals were found in leaf litter, and very rarely in open places. Millipedes roll up at rest, being active both in daylight and in the dark. When disturbed, and having rolled up, animals unroll a few minutes later and typically do not roll up again from subsequent slight disturbances. The peak of abundance of adult animals is in June-July, at the onset of the rainy season, forming local swarming sites in some forest areas. Mating was noted during swarming time, and was observed infrequently. During the remaining parts of the year, individuals are more scarce, and show low population densities. Juveniles or oviposition places were not observed.
</paragraph>
<paragraph id="8BC15D14FFDD2424CBEDE621DC7AFBC1" blockId="5.[151,1437,785,1134]" box="[199,1102,1109,1134]" pageId="5" pageNumber="540">
<emphasis id="B90A8106FFDD2424CBEDE621D920FBC2" bold="true" box="[199,276,1109,1134]" pageId="5" pageNumber="540">Name.</emphasis>
Adjective, to emphasize Cat Tien National Park, its
<typeStatus id="54C5E3B6FFDD2424C84BE622DBA5FBC2" box="[865,913,1110,1134]" pageId="5" pageNumber="540">type</typeStatus>
locality (
<figureCitation id="13454191FFDD2424C8D1E621DC09FBC1" box="[1019,1085,1109,1134]" captionStart="FIGURE 1" captionStartId="3.[151,249,1736,1758]" captionTargetBox="[152,1435,447,1716]" captionTargetId="figure@3.[152,1435,447,1716]" captionTargetPageId="3" captionText="FIGURE 1. Distribution map with pictures of live specimens. A. Sphaerobelum cattiense sp. nov.; B. Sphaerobelum konkakinhense sp. nov.; C. Zephronia konkakinhensis sp. nov.; D. Zephronia montis sp. nov." httpUri="https://zenodo.org/record/1458881/files/figure.png" pageId="5" pageNumber="540">Fig. 1</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>