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<document ID-DOI="http://doi.org/10.5281/zenodo.3788580" ID-GBIF-Dataset="8094be38-8ec7-4cd1-89b2-0173f551d554" ID-GBIF-Taxon="163750507" ID-ISSN="13132970" ID-Zenodo-Dep="3788580" ID-ZooBank="F4D24A8D-3EBF-4688-910D-413E328F92BF" LSID="urn:lsid:zoobank.org:act:3FCD51C4-A812-49D5-B653-56922696F08B" approvalRequired="22" approvalRequired_for_textStreams="22" checkinTime="1587982620088" checkinUser="plazi" docAuthor="Schmidt, Christian &amp; Anweiler, Gary" docDate="2010" docId="03D7D51E9C6AE61DFF0EFB01FC3409D7" docLanguage="en" docName="ZK_article_2174.pdf" docOrigin="ZooKeys 39 (39)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@258e5601" docStyleName="zookeys.2008.journal_article" docTitle="Charadra cakulha Schmidt &amp; Anweiler 2010, sp. n." docType="treatment" docUuid="3FCD51C4-A812-49D5-B653-56922696F08B" docUuidSource="ZooBank" docVersion="5" lastPageId="14" lastPageNumber="175" masterDocId="FFEEAD669C66E613FF82FFA2FFAF082A" masterDocTitle="The North American species of Charadra Walker, with a revision of the Charadra pata (Druce) group (Noctuidae, Pantheinae)" masterLastPageNumber="181" masterPageNumber="161" pageId="12" pageNumber="173" updateTime="1643563573501" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>The North American species of Charadra Walker, with a revision of the Charadra pata (Druce) group (Noctuidae, Pantheinae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Schmidt, Christian</mods:namePart>
<mods:affiliation>Canadian Food Inspection Agency, Ottawa ,, Canada</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Anweiler, Gary</mods:namePart>
<mods:affiliation>University of Alberta Strickland Entomology Museum ,, Canada</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2010</mods:date>
<mods:detail type="pubDate">
<mods:number>2010-03-18</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>39</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>39</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>161</mods:start>
<mods:end>181</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.3897/zookeys.39.432</mods:identifier>
<mods:identifier type="GBIF-Dataset">8094be38-8ec7-4cd1-89b2-0173f551d554</mods:identifier>
<mods:identifier type="ISSN">13132970</mods:identifier>
<mods:identifier type="Zenodo-Dep">576639</mods:identifier>
<mods:identifier type="ZooBank">F4D24A8D-3EBF-4688-910D-413E328F92BF</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3788580" ID-GBIF-Taxon="163750507" ID-Zenodo-Dep="3788580" LSID="urn:lsid:zoobank.org:act:3FCD51C4-A812-49D5-B653-56922696F08B" httpUri="http://treatment.plazi.org/id/03D7D51E9C6AE61DFF0EFB01FC3409D7" lastPageId="14" lastPageNumber="175" pageId="12" pageNumber="173">
<subSubSection pageId="12" pageNumber="173" type="nomenclature">
<paragraph blockId="12.[140,968,1187,1285]" box="[140,688,1187,1214]" pageId="12" pageNumber="173">
<heading box="[140,688,1187,1214]" fontSize="11" level="2" pageId="12" pageNumber="173" reason="3">
<emphasis bold="true" box="[140,688,1187,1214]" pageId="12" pageNumber="173">
<taxonomicName authority="Schmidt &amp; Anweiler" authorityName="Schmidt &amp; Anweiler" authorityYear="2010" box="[140,610,1187,1214]" class="Insecta" family="Pantheidae" genus="Charadra" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="173" phylum="Arthropoda" rank="species" species="cakulha" status="sp. n.">
<emphasis bold="true" box="[140,358,1187,1214]" italics="true" pageId="12" pageNumber="173">
<emphasis box="[140,257,1187,1214]" inLineHeading="true" pageId="12" pageNumber="173" reason="1">Charadra</emphasis>
cakulha
</emphasis>
Schmidt &amp; Anweiler
</taxonomicName>
,
<taxonomicNameLabel box="[623,688,1187,1213]" pageId="12" pageNumber="173" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph blockId="12.[140,968,1187,1285]" box="[140,968,1222,1249]" pageId="12" pageNumber="173">
<uri box="[140,968,1222,1249]" pageId="12" pageNumber="173">
urn:lsid:zoobank.org:act:
<uuid box="[423,968,1222,1249]" pageId="12" pageNumber="173">3FCD51C4-A812-49D5-B653-56922696F08B</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection box="[140,250,1258,1285]" pageId="12" pageNumber="173" type="description">
<paragraph blockId="12.[140,968,1187,1285]" box="[140,250,1258,1285]" pageId="12" pageNumber="173">Figs 7, 25</paragraph>
</subSubSection>
<subSubSection pageId="12" pageNumber="173" type="materials_examined">
<paragraph blockId="12.[140,1108,1328,1672]" pageId="12" pageNumber="173">
<emphasis bold="true" box="[140,317,1328,1354]" pageId="12" pageNumber="173">
<emphasis box="[140,198,1328,1354]" inLineHeading="true" pageId="12" pageNumber="173" reason="1">Type</emphasis>
material.
</emphasis>
<materialsCitation ID-GBIF-Occurrence="2608937997" collectingDate="1969-06-17" collectorName="A. Mutuura" country="Mexico" location="MEXICO" pageId="12" pageNumber="173" specimenCount="1" typeStatus="holotype">
<emphasis bold="true" box="[329,442,1328,1354]" pageId="12" pageNumber="173">
<typeStatus box="[329,442,1328,1354]" pageId="12" pageNumber="173">Holotype</typeStatus>
</emphasis>
<collectingCountry box="[513,633,1328,1354]" name="Mexico" pageId="12" pageNumber="173">MEXICO</collectingCountry>
: “MEX., San Cristobal / de Las Casas, Chi[apa]s. /
<date box="[279,430,1363,1390]" pageId="12" pageNumber="173" value="1969-06-17">
<collectingDate box="[279,430,1363,1390]" pageId="12" pageNumber="173" value="1969-06-17">June 17 1969</collectingDate>
</date>
/
<collectorName box="[451,587,1364,1390]" pageId="12" pageNumber="173">A. Mutuura</collectorName>
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2608937991" collectionCode="CNC" country="Mexico" pageId="12" pageNumber="173" specimenCount="1" typeStatus="holotype">
<typeStatus box="[617,774,1364,1390]" pageId="12" pageNumber="173">HOLOTYPE</typeStatus>
/
<taxonomicName authorityName="Schmidt &amp; Anweiler" authorityYear="2010" box="[795,990,1363,1389]" class="Insecta" family="Pantheidae" genus="Charadra" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="173" phylum="Arthropoda" rank="species" species="cakulha">
<emphasis box="[795,990,1363,1389]" italics="true" pageId="12" pageNumber="173">Charadra cakulha</emphasis>
</taxonomicName>
/ Schmidt &amp; Anweiler” [red label].
<collectionCode box="[420,487,1399,1425]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="12" pageNumber="173">CNC</collectionCode>
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="2608937994" box="[498,1028,1399,1426]" collectionCode="CNC" country="Mexico" pageId="12" pageNumber="173" specimenCount="1" typeStatus="paratype">
<typeStatus box="[498,604,1399,1425]" pageId="12" pageNumber="173">Paratypes</typeStatus>
same data as holotype, (1♁).
<collectionCode box="[961,1028,1399,1425]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="12" pageNumber="173">CNC</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="12" pageNumber="173" type="etymology">
<paragraph blockId="12.[140,1108,1328,1672]" pageId="12" pageNumber="173">
<emphasis bold="true" box="[185,320,1434,1460]" pageId="12" pageNumber="173">
<emphasis box="[185,315,1434,1460]" inLineHeading="true" pageId="12" pageNumber="173" reason="1">Etymology</emphasis>
.
</emphasis>
A deity from Mayan mythology, Cakulha is the ruler of the lesser lightning bolts, and brother of Coyopa. It is a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection lastPageId="13" lastPageNumber="174" pageId="12" pageNumber="173" type="diagnosis">
<paragraph blockId="12.[140,1108,1328,1672]" pageId="12" pageNumber="173">
<emphasis bold="true" box="[185,312,1504,1530]" pageId="12" pageNumber="173">
<emphasis box="[185,307,1504,1530]" inLineHeading="true" pageId="12" pageNumber="173" reason="1">Diagnosis</emphasis>
.
</emphasis>
Externally recognizable by the very pale greyish-white outer third of the forewing and thorax, unique among the
<taxonomicName box="[637,685,1540,1565]" class="Insecta" family="Pantheidae" genus="Charadra" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="173" phylum="Arthropoda" rank="species" species="pata">
<emphasis box="[637,685,1540,1565]" italics="true" pageId="12" pageNumber="173">pata</emphasis>
</taxonomicName>
group. In addition to the wing markings,
<taxonomicName authorityName="Schmidt &amp; Anweiler" authorityYear="2010" box="[202,323,1575,1601]" class="Insecta" family="Pantheidae" genus="Charadra" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="173" phylum="Arthropoda" rank="species" species="cakulha">
<emphasis box="[202,221,1575,1601]" italics="true" pageId="12" pageNumber="173">C</emphasis>
.
<emphasis box="[238,323,1575,1601]" italics="true" pageId="12" pageNumber="173">cakulha</emphasis>
</taxonomicName>
is distinguishable from the similar
<taxonomicName authorityName="Dyar" authorityYear="1916" box="[734,876,1575,1601]" class="Insecta" family="Pantheidae" genus="Charadra" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="173" phylum="Arthropoda" rank="species" species="oligarchia">
<emphasis box="[734,876,1575,1601]" italics="true" pageId="12" pageNumber="173">C. oligarchia</emphasis>
</taxonomicName>
by the lateral, symmetrical cornuti that are fused to the aedeagus (in
<taxonomicName authorityName="Dyar" authorityYear="1916" box="[712,852,1610,1636]" class="Insecta" family="Pantheidae" genus="Charadra" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="173" phylum="Arthropoda" rank="species" species="oligarchia">
<emphasis box="[712,852,1610,1636]" italics="true" pageId="12" pageNumber="173">C. oligarchia</emphasis>
</taxonomicName>
they are at the base of the vesica with one cornutus massive and one small).
</paragraph>
<paragraph blockId="13.[140,1108,1184,1270]" pageId="13" pageNumber="174">
<emphasis bold="true" box="[140,300,1184,1206]" pageId="13" pageNumber="174">Figures 3034.</emphasis>
Female genitalia of
<taxonomicName authorityName="Walker" authorityYear="1865" box="[487,572,1185,1206]" class="Insecta" family="Pantheidae" genus="Charadra" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="174" phylum="Arthropoda" rank="genus">
<emphasis box="[487,572,1185,1206]" italics="true" pageId="13" pageNumber="174">Charadra</emphasis>
</taxonomicName>
species.
<emphasis bold="true" box="[653,679,1184,1206]" pageId="13" pageNumber="174">30</emphasis>
<taxonomicName authorityName="Anweiler &amp; Schmidt" authorityYear="2010" box="[685,824,1185,1206]" class="Insecta" family="Pantheidae" genus="Charadra" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="174" phylum="Arthropoda" rank="species" species="franclemonti">
<emphasis box="[685,824,1185,1206]" italics="true" pageId="13" pageNumber="174">C. franclemonti</emphasis>
</taxonomicName>
<typeStatus box="[831,913,1185,1206]" pageId="13" pageNumber="174">paratype</typeStatus>
, Madera Cyn., Santa Rita Mtns., AZ
<emphasis bold="true" box="[287,314,1216,1238]" pageId="13" pageNumber="174">3Ι</emphasis>
<taxonomicName authorityName="Schmidt &amp; Anweiler" authorityYear="2010" box="[319,385,1217,1238]" class="Insecta" family="Pantheidae" genus="Charadra" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="174" phylum="Arthropoda" rank="species" species="tapa">
<emphasis box="[319,385,1217,1238]" italics="true" pageId="13" pageNumber="174">C. tapa</emphasis>
</taxonomicName>
<typeStatus box="[391,473,1217,1238]" pageId="13" pageNumber="174">paratype</typeStatus>
, Ramsey Cyn., Huachuca Mtns., AZ
<emphasis bold="true" box="[819,845,1216,1238]" pageId="13" pageNumber="174">32</emphasis>
<taxonomicName authorityName="Schmidt &amp; Anweiler" authorityYear="2010" box="[851,943,1217,1238]" class="Insecta" family="Pantheidae" genus="Charadra" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="174" phylum="Arthropoda" rank="species" species="moneta">
<emphasis box="[851,943,1217,1238]" italics="true" pageId="13" pageNumber="174">C. moneta</emphasis>
</taxonomicName>
<typeStatus box="[948,1030,1217,1238]" pageId="13" pageNumber="174">paratype</typeStatus>
, Walnut Cyn., Coconino Co., AZ
<emphasis bold="true" box="[378,404,1248,1270]" pageId="13" pageNumber="174">33</emphasis>
<taxonomicName authority=", Morehead" authorityName="Morehead" box="[411,620,1248,1270]" class="Insecta" family="Pantheidae" genus="Charadra" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="174" phylum="Arthropoda" rank="species" species="deridens">
<emphasis box="[411,511,1249,1270]" italics="true" pageId="13" pageNumber="174">C. deridens</emphasis>
, Morehead
</taxonomicName>
, KY
<emphasis bold="true" box="[665,691,1248,1270]" pageId="13" pageNumber="174">34</emphasis>
<taxonomicName authority=", Sinton" authorityName="Sinton" box="[697,870,1248,1270]" class="Insecta" family="Pantheidae" genus="Charadra" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="174" phylum="Arthropoda" rank="species" species="dispulsa">
<emphasis box="[697,795,1249,1270]" italics="true" pageId="13" pageNumber="174">C. dispulsa</emphasis>
, Sinton
</taxonomicName>
, TX.
</paragraph>
</subSubSection>
<subSubSection lastPageId="14" lastPageNumber="175" pageId="13" pageNumber="174" type="description">
<paragraph blockId="13.[140,1108,1328,1672]" lastBlockId="14.[140,1108,167,510]" lastPageId="14" lastPageNumber="175" pageId="13" pageNumber="174">
<emphasis bold="true" box="[185,335,1328,1354]" pageId="13" pageNumber="174">
<emphasis box="[185,330,1328,1354]" inLineHeading="true" pageId="13" pageNumber="174" reason="1">Description</emphasis>
.
</emphasis>
Female unknown. Forewing length 19.7 mm.
<emphasis italics="true" pageId="13" pageNumber="174">Head, thorax and abdomen</emphasis>
as in
<taxonomicName authorityName="Anweiler &amp; Schmidt" authorityYear="2010" box="[313,486,1363,1390]" class="Insecta" family="Pantheidae" genus="Charadra" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="174" phylum="Arthropoda" rank="species" species="franclemonti">
<emphasis box="[313,486,1363,1390]" italics="true" pageId="13" pageNumber="174">C. franclemonti</emphasis>
</taxonomicName>
, differing in the following characters: vestiture much paler grey, nearly white; thorax with margin of tegulae and patagia bordered with black scales. Forewing light grey, distal third powdery whitish grey; reniform spot not discernible, reniform area entirely whitish; orbicular broadly oval, whitish with brown diffuse central area; subterminal line poorly defined medially by irregular white line, black scales at costal and anal margin, expanded to a thick, prominent line at anal margin.
<emphasis box="[140,296,1575,1601]" italics="true" pageId="13" pageNumber="174">Male genitalia</emphasis>
(Fig. 25) Valves simple, subquadrate apically with a strongly incurved, scoop-shaped dorsocaudal apex; costal process absent; clasper a simple scoop-shaped process about 1/6 length of dorsal valve margin, with broad surface directed dorso- cephalad; sacculus unmodified; uncus strongly constricted medially and apex bifid, with an overall Y-shaped appearance; tegumen expanded dorsally; saccus V-shaped; juxta triangular; subscaphium strongly sclerotized; aedeagus 3 × longer than wide, with a very wide, dorsally directed opening to vesica, opening about 1/3 total length of aedeagus; vesica directed at about 90 degrees to aedeagus; cornuti usually placed on base of vesica and fused laterally to aedeagus, so aedeagus armed distolaterally with two stout, laterally projecting spines, left one placed slightly more apically; one medial and one distal diverticulum of similar size, oriented dorsally.
</paragraph>
</subSubSection>
<subSubSection pageId="14" pageNumber="175" type="distribution">
<paragraph blockId="14.[140,1108,167,510]" pageId="14" pageNumber="175">
<emphasis bold="true" box="[185,489,448,474]" pageId="14" pageNumber="175">
<emphasis box="[185,336,448,474]" inLineHeading="true" pageId="14" pageNumber="175" reason="1">Distribution</emphasis>
and biology.
</emphasis>
Known only from the two specimens of the
<typeStatus box="[986,1034,449,475]" pageId="14" pageNumber="175">type</typeStatus>
series, collected in mid-June at San Cristobal de las Casas,
<collectingRegion box="[726,820,483,509]" country="Mexico" name="Chiapas" pageId="14" pageNumber="175">Chiapas</collectingRegion>
,
<collectingCountry box="[832,920,483,509]" name="Mexico" pageId="14" pageNumber="175">Mexico</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>