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<document id="E2768172DF1DF17E7CDCAB84D813796C" ID-CLB-Dataset="62482" ID-DOI="10.5281/zenodo.6603807" ID-GBIF-Dataset="916bf600-5bc0-438d-9c6c-7fff4c5a388f" ID-ISBN="978-84-941892-3-4" ID-Zenodo-Dep="6603807" IM.illustrations_requiresApprovalFor="plazi" IM.metadata_requiresApprovalFor="plazi" IM.taxonomicNames_requiresApprovalFor="plazi" checkinTime="1654101401716" checkinUser="carolina" docAuthor="Don E. Wilson, Thomas E. Lacher, Jr &amp; Russell A. Mittermeier" docDate="2016" docId="0131878A073EFF91FF9AFD1C6BE74DDA" docLanguage="en" docName="hbmw_6_Geomyidae_0234.pdf.imf" docOrigin="Handbook of the Mammals of the World Volume 6 Lagomorphs and Rodents I, Barcelona: Lynx Edicions" docTitle="Cratogeomys goldman Merriam 1895" docType="treatment" docVersion="10" lastPageNumber="251" masterDocId="FD08FFF2072CFF83FFF3FFF96B0F4602" masterDocTitle="Geomyidae" masterLastPageNumber="269" masterPageNumber="234" pageNumber="251" updateTime="1699338600100" updateUser="ExternalLinkService">
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<mods:title id="A4F1175A65CD13F19536541B79E6A4E4">Geomyidae</mods:title>
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<mods:namePart id="302D06033E5D15756B12A8193169B359">Don E. Wilson</mods:namePart>
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<mods:namePart id="7B7E22029BD9D266B61A9CC992D5232C">Thomas E. Lacher, Jr</mods:namePart>
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<mods:namePart id="B8743150070CC794C80A5FB537EDF98B">Russell A. Mittermeier</mods:namePart>
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<mods:title id="B698012244F80BE2C25CB0118FA4BDFA">Handbook of the Mammals of the World Volume 6 Lagomorphs and Rodents I</mods:title>
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<treatment id="0131878A073EFF91FF9AFD1C6BE74DDA" ID-DOI="http://doi.org/10.5281/zenodo.6604066" ID-GBIF-Taxon="195708561" ID-Zenodo-Dep="6604066" LSID="urn:lsid:plazi:treatment:0131878A073EFF91FF9AFD1C6BE74DDA" httpUri="http://treatment.plazi.org/id/0131878A073EFF91FF9AFD1C6BE74DDA" lastPageNumber="251" pageId="18" pageNumber="251">
<subSubSection id="C1826517073EFF91FF9AFD1C6BAE4511" box="[105,161,741,787]" pageId="18" pageNumber="251" type="multiple">
<paragraph id="8927369C073EFF91FF9AFD1C6BAE4511" blockId="18.[102,1208,741,867]" box="[105,161,741,787]" pageId="18" pageNumber="251">
<heading id="D26F81F0073EFF91FF9AFD1C6BAE4511" box="[105,161,741,787]" pageId="18" pageNumber="251">
<figureCitation id="11A32A19073EFF91FF9AFD1C6BAE4511" box="[105,161,741,787]" captionStart="Plate 13: Geomyidae" captionStartId="14.[104,134,3318,3339]" captionTargetBox="[12,2731,15,3644]" captionTargetPageId="13" captionText="25. Michoacan Pocket Gopher (Zygogeomys trichopus), 26. Giant Pocket Gopher (Orthogeomys grandis), 27. Hispid Pocket Gopher (Heterogeomys hispidus), 28. Big Pocket Gopher (Heterogeomys lanius), 29. Chiriqui Pocket Gopher (Heterogeomys cavator), 30. Darien Pocket Gopher (Heterogeomys dariensis), 31. Cherries Pocket Gopher (Heterogeomys cherrier), 32. Variable Pocket Gopher (Heterogeomys heterodus), 33. Underwoods Pocket Gopher (Heterogeomys underwood), 34. Bullers Pocket Gopher (Pappogeomys bulleri), 35. Yellow-faced Pocket Gopher (Cratogeomys castanops), 36. Goldmans Pocket Gopher (Cratogeomys goldman), 37. Oriental Basin Pocket Gopher (Cratogeomys fulvescens), 38. Merriams Pocket Gopher (Cratogeomys merriami), 39. Cofre de Perote Pocket Gopher (Cratogeomys perotensis), 40. Volcan de Toluca Pocket Gopher (Cratogeomys planiceps), 41. Smoky Pocket Gopher (Cratogeomys fumosus)" figureDoi="http://doi.org/10.5281/zenodo.6603903" httpUri="https://zenodo.org/record/6603903/files/figure.png" pageId="18" pageNumber="251">36.</figureCitation>
</heading>
</paragraph>
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<subSubSection id="C1826517073EFF91FF41FD1C69D04511" box="[178,735,741,787]" pageId="18" pageNumber="251" type="vernacular_names">
<paragraph id="8927369C073EFF91FF41FD1C69D04511" blockId="18.[102,1208,741,867]" box="[178,735,741,787]" pageId="18" pageNumber="251">
<heading id="D26F81F0073EFF91FF41FD1C69D04511" box="[178,735,741,787]" pageId="18" pageNumber="251">
<vernacularName id="079B46B2073EFF91FF41FD1C69D04511" box="[178,735,741,787]" pageId="18" pageNumber="251">Goldmans Pocket Gopher</vernacularName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C1826517073EFF91FCDBFD1C6FB84511" box="[808,1207,741,787]" pageId="18" pageNumber="251" type="nomenclature">
<paragraph id="8927369C073EFF91FCDBFD1C6FB84511" blockId="18.[102,1208,741,867]" box="[808,1207,741,787]" pageId="18" pageNumber="251">
<heading id="D26F81F0073EFF91FCDBFD1C6FB84511" box="[808,1207,741,787]" pageId="18" pageNumber="251">
<taxonomicName id="4E984D1F073EFF91FCDBFD1C6FB84511" ID-CoL="b9605ca1-7a08-4dd4-a1ae-787258ad7bf4" authorityName="Merriam" authorityYear="1895" box="[808,1207,741,787]" class="Mammalia" family="Geomyidae" genus="Cratogeomys" kingdom="Animalia" order="Rodentia" pageId="18" pageNumber="251" phylum="Chordata" rank="species" species="goldman">
<emphasis id="BBECEA8E073EFF91FCDBFD1C6FB84511" box="[808,1207,741,787]" italics="true" pageId="18" pageNumber="251">Cratogeomys goldman</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C1826517073EFF91FF94FCDC69E54560" pageId="18" pageNumber="251" type="vernacular_names">
<paragraph id="8927369C073EFF91FF94FCDC68F74538" blockId="18.[102,1208,741,867]" box="[103,1016,805,826]" pageId="18" pageNumber="251">
<heading id="D26F81F0073EFF91FF94FCDC68F74538" box="[103,1016,805,826]" pageId="18" pageNumber="251">
<emphasis id="BBECEA8E073EFF91FF94FCDC6BBC4538" bold="true" box="[103,179,805,826]" pageId="18" pageNumber="251">French:</emphasis>
<vernacularName id="079B46B2073EFF91FF4FFCDC6A704538" box="[188,383,805,826]" pageId="18" pageNumber="251">Gaufre de Goldman</vernacularName>
/
<emphasis id="BBECEA8E073EFF91FE67FCDC6AE14538" bold="true" box="[404,494,805,826]" pageId="18" pageNumber="251">German:</emphasis>
<vernacularName id="079B46B2073EFF91FE0BFCDC69D94538" box="[504,726,805,826]" pageId="18" pageNumber="251">Goldman-Taschenratte</vernacularName>
/
<emphasis id="BBECEA8E073EFF91FD18FCDC68494538" bold="true" box="[747,838,805,826]" pageId="18" pageNumber="251">Spanish:</emphasis>
<vernacularName id="079B46B2073EFF91FCBFFCDC68F74538" box="[844,1016,805,826]" pageId="18" pageNumber="251">Tuza de Goldman</vernacularName>
</heading>
</paragraph>
<paragraph id="8927369C073EFF91FF94FCB469E54560" blockId="18.[102,1208,741,867]" box="[103,746,845,866]" pageId="18" pageNumber="251">
<heading id="D26F81F0073EFF91FF94FCB469E54560" box="[103,746,845,866]" pageId="18" pageNumber="251">
<emphasis id="BBECEA8E073EFF91FF94FCB46A514560" bold="true" box="[103,350,845,866]" pageId="18" pageNumber="251">Other common names:</emphasis>
<vernacularName id="079B46B2073EFF91FE9BFCB469E54560" box="[360,746,845,866]" pageId="18" pageNumber="251">Goldman's Yellow-faced Pocket Gopher</vernacularName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C1826517073EFF91FD39FC6B68E945D5" pageId="18" pageNumber="251" type="reference_group">
<paragraph id="8927369C073EFF91FD39FC6B68E945D5" blockId="18.[714,1310,914,1337]" pageId="18" pageNumber="251">
<emphasis id="BBECEA8E073EFF91FD39FC6B686545AD" bold="true" box="[714,874,914,943]" pageId="18" pageNumber="251">Taxonomy.</emphasis>
<taxonomicName id="4E984D1F073EFF91FC70FC6B68EE45D5" authority="Merriam, 1895" authorityName="Merriam" authorityYear="1895" class="Mammalia" family="Geomyidae" genus="Cratogeomys" kingdom="Animalia" order="Rodentia" pageId="18" pageNumber="251" phylum="Chordata" rank="subSpecies" species="castanops" subSpecies="goldmant">Cratogeomys castanops goldmant Merriam, 1895</taxonomicName>
,
</paragraph>
</subSubSection>
<subSubSection id="C1826517073EFF91FC0BFC47684545FD" pageId="18" pageNumber="251" type="materials_examined">
<paragraph id="8927369C073EFF91FC0BFC47684545FD" blockId="18.[714,1310,914,1337]" pageId="18" pageNumber="251">
<materialsCitation id="39F03CC1073EFF91FC0BFC47684545FD" ID-GBIF-Occurrence="3799834322" pageId="18" pageNumber="251">“Canitas, Zacatecas, Mexico.”</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C1826517073EFF91FD3DFBFD6AAB43FD" pageId="18" pageNumber="251" type="discussion">
<paragraph id="8927369C073EFF91FD3DFBFD6AAB43FD" blockId="18.[714,1310,914,1337]" lastBlockId="18.[103,1311,1344,3032]" pageId="18" pageNumber="251">
<taxonomicName id="4E984D1F073EFF91FD3DFBFD68684227" authorityName="Merriam" authorityYear="1895" box="[718,871,1028,1061]" class="Mammalia" family="Geomyidae" genus="Cratogeomys" kingdom="Animalia" order="Rodentia" pageId="18" pageNumber="251" phylum="Chordata" rank="genus">Cratogeomys</taxonomicName>
was regarded as a subgenus of
<taxonomicName id="4E984D1F073EFF91FD38FBC9687F424F" authorityName="Merriam" authorityYear="1895" box="[715,880,1072,1101]" class="Mammalia" family="Geomyidae" genus="Pappogeomys" kingdom="Animalia" order="Rodentia" pageId="18" pageNumber="251" phylum="Chordata" rank="genus">Pappogeomys</taxonomicName>
by R. J. Russell in 1968 but was returned to generic status by R. L. Honeycutt and S. L. Willams in 1982. It was included in
<taxonomicName id="4E984D1F073EFF91FC84FB5B6F1C42C1" baseAuthorityName="Baird" baseAuthorityYear="1852" box="[887,1043,1186,1219]" class="Mammalia" family="Geomyidae" genus="Cratogeomys" kingdom="Animalia" order="Rodentia" pageId="18" pageNumber="267" phylum="Chordata" rank="species" species="castanops">C. castanops</taxonomicName>
by earlier workers, including Russell in 1968, but was recognized as a distinct species by numerous authors beginning with H. K. Lee and R. J. Baker in 1987. Molecular studies show
<taxonomicName id="4E984D1F073EFF91FD3CFAB968644363" authorityName="Merriam" authorityYear="1895" box="[719,875,1344,1377]" class="Mammalia" family="Geomyidae" genus="Cratogeomys" kingdom="Animalia" order="Rodentia" pageId="18" pageNumber="267" phylum="Chordata" rank="species" species="goldmani">C. goldmani</taxonomicName>
to be sister to
<taxonomicName id="4E984D1F073EFF91FBCAFAB96FD84363" baseAuthorityName="Baird" baseAuthorityYear="1852" box="[1081,1239,1344,1377]" class="Mammalia" family="Geomyidae" genus="Cratogeomys" kingdom="Animalia" order="Rodentia" pageId="18" pageNumber="267" phylum="Chordata" rank="species" species="castanops">C. castanops</taxonomicName>
, and these two species, together with C.
<taxonomicName id="4E984D1F073EFF91FDBFFA8969B1438B" baseAuthorityName="Thomas" baseAuthorityYear="1893" box="[588,702,1392,1417]" class="Mammalia" family="Geomyidae" genus="Cratogeomys" kingdom="Animalia" order="Rodentia" pageId="18" pageNumber="251" phylum="Chordata" rank="species" species="merriami">merriami</taxonomicName>
, C.
<taxonomicName id="4E984D1F073EFF91FD1FFA896861438B" authorityName="Merriam" authorityYear="1895" box="[748,878,1392,1417]" class="Mammalia" family="Geomyidae" genus="Cratogeomys" kingdom="Animalia" order="Rodentia" pageId="18" pageNumber="251" phylum="Chordata" rank="species" species="fulvescens">fulvescens</taxonomicName>
, and
<taxonomicName id="4E984D1F073EFF91FC4EFA896F58438B" authorityName="Merriam" authorityYear="1895" box="[957,1111,1392,1417]" class="Mammalia" family="Geomyidae" genus="Cratogeomys" kingdom="Animalia" order="Rodentia" pageId="18" pageNumber="268" phylum="Chordata" rank="species" species="perotensis">C. perotensis</taxonomicName>
, comprise the
<taxonomicName id="4E984D1F073EFF91FF98FA6A6A0843B2" baseAuthorityName="Baird" baseAuthorityYear="1852" box="[107,263,1427,1456]" class="Mammalia" family="Geomyidae" genus="Cratogeomys" kingdom="Animalia" order="Rodentia" pageId="18" pageNumber="267" phylum="Chordata" rank="species" species="castanops">C. castanops</taxonomicName>
species group. J. L. Patton in 2005 recognized six subspecies of
<taxonomicName id="4E984D1F073EFF91FB8FFA6A6E1A43B2" authorityName="Merriam" authorityYear="1895" box="[1148,1301,1427,1456]" class="Mammalia" family="Geomyidae" genus="Cratogeomys" kingdom="Animalia" order="Rodentia" pageId="18" pageNumber="267" phylum="Chordata" rank="species" species="goldmani">C. goldmani</taxonomicName>
, subsequently reduced to two subspecies by D. J. Hafner and colleagues in 2008. Two subspecies recognized.
</paragraph>
</subSubSection>
<subSubSection id="C1826517073EFF91FF94F9FC699940C6" pageId="18" pageNumber="251" type="distribution">
<caption id="DDE76614073EFF91FF94F9FC699940C6" ID-DOI="http://doi.org/10.5281/zenodo.6603885" ID-Zenodo-Dep="6603885" httpUri="https://zenodo.org/record/6603885/files/figure.png" inLine="true" pageId="18" pageNumber="251" targetBox="[100,690,922,1336]" targetPageId="18">
<paragraph id="8927369C073EFF91FF94F9FC6AF84024" blockId="18.[103,1311,1344,3032]" box="[103,503,1541,1574]" pageId="18" pageNumber="251">
<emphasis id="BBECEA8E073EFF91FF94F9FC6AF84024" bold="true" box="[103,503,1541,1574]" pageId="18" pageNumber="251">Subspecies and Distribution.</emphasis>
</paragraph>
<paragraph id="8927369C073EFF91FF98F9D468344077" blockId="18.[103,1311,1344,3032]" pageId="18" pageNumber="251">
<taxonomicName id="4E984D1F073EFF91FF98F9D468344077" authority="Merriam, 1895" authorityName="Merriam" authorityYear="1895" class="Mammalia" family="Geomyidae" genus="Cratogeomys" kingdom="Animalia" order="Rodentia" pageId="18" pageNumber="251" phylum="Chordata" rank="subSpecies" species="goldman" subSpecies="goldmani">C.g.goldmaniMerriam,1895—drainageoftheRioAguanaval(NEDurango,SCoahuila,andZacatecas)toWSanLuisPotosi,Mexico.</taxonomicName>
</paragraph>
<paragraph id="8927369C073EFF91FF98F982699940C6" blockId="18.[103,1311,1344,3032]" pageId="18" pageNumber="251">
<taxonomicName id="4E984D1F073EFF91FF98F98269A7409E" authority="Nelson &amp; Goldman, 1934" authorityName="Nelson &amp; Goldman" authorityYear="1934" box="[107,680,1659,1692]" class="Mammalia" family="Geomyidae" genus="Cratogeomys" kingdom="Animalia" order="Rodentia" pageId="18" pageNumber="251" phylum="Chordata" rank="subSpecies" species="goldman" subSpecies="subnubilus">C. g. subnubilus Nelson &amp; Goldman, 1934</taxonomicName>
— SE Coahuila and SW Nuevo Leon S to the Rio Verde in C San Luis Potosi, Mexico.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="C1826517073EFF91FF94F9336FF74E4F" pageId="18" pageNumber="251" type="description">
<paragraph id="8927369C073EFF91FF94F9336FF74E4F" blockId="18.[103,1311,1344,3032]" pageId="18" pageNumber="251">
<emphasis id="BBECEA8E073EFF91FF94F9336A6D40E9" bold="true" box="[103,354,1738,1771]" pageId="18" pageNumber="251">Descriptive notes.</emphasis>
Head—body 160-210 mm (males) and 145-190 mm (females), tail 60-95 mm (males) and 55-90 mm (females); weight 140-350 g (males) and 130-215 ¢g (females). Male Goldmans Pocket Gophers are usually larger than females, and this is especially pronounced in populations with larger average body size. It is relatively small for the genus. It has a fusiform body shape typical of all pocket gophers, and it possesses fur-lined cheek pouches that open external to the mouth. Most individuals are ochreous-buff to dark brown dorsally and pale ochreous-buff to white ventrally. Up to 89% of individuals in some populations have white spots on belly, sides, or rump. Anterior surface of each upper incisor has a single medial groove slightly displaced to the innerside of the tooth. Chromosomal complement has 2n = 42 and FN = 78.
</paragraph>
</subSubSection>
<subSubSection id="C1826517073EFF91FF9BF7AD6AEC4EC6" pageId="18" pageNumber="251" type="biology_ecology">
<paragraph id="8927369C073EFF91FF9BF7AD6AEC4EC6" blockId="18.[103,1311,1344,3032]" pageId="18" pageNumber="251">
<emphasis id="BBECEA8E073EFF91FF9BF7AD6BD84E77" bold="true" box="[104,215,2132,2165]" pageId="18" pageNumber="251">Habitat.</emphasis>
Usually in arid and semiarid areas with deep sandy or silty soils that support xeric-adapted shrubs and grasses. Goldman's Pocket Gopher is known from elevations of 750 m to almost 2700 m.
</paragraph>
</subSubSection>
<subSubSection id="C1826517073EFF91FF9BF7336E154FD5" pageId="18" pageNumber="251" type="food_feeding">
<paragraph id="8927369C073EFF91FF9BF7336E154FD5" blockId="18.[103,1311,1344,3032]" pageId="18" pageNumber="251">
<emphasis id="BBECEA8E073EFF91FF9BF7336A7E4EE9" bold="true" box="[104,369,2250,2283]" pageId="18" pageNumber="251">Food and Feeding.</emphasis>
There is no specific information available for this species, but all pocket gophers that have been studied eat predominately underground roots and tubers and a limited amount of surface vegetation. Goldmans Pocket Gopher readily invades cultivated fields and is considered an agricultural pest wherever it occurs in contact with humans. As in all other pocket gophers, the burrow system is a series of shallow feeding tunnels radiating spoke-like from a deeper, central network that contains one or more nest chambers and several smaller chambers for storage of food or fecal pellets.
</paragraph>
</subSubSection>
<subSubSection id="C1826517073EFF91FF9BF6246A124C24" pageId="18" pageNumber="251" type="breeding">
<paragraph id="8927369C073EFF91FF9BF6246A124C24" blockId="18.[103,1311,1344,3032]" pageId="18" pageNumber="251">
<emphasis id="BBECEA8E073EFF91FF9BF6246BE04FFC" bold="true" box="[104,239,2525,2558]" pageId="18" pageNumber="251">Breeding.</emphasis>
Pregnant female Goldmans Pocket Gophers have been collected in spring and summer.
</paragraph>
</subSubSection>
<subSubSection id="C1826517073EFF91FF94F5C96FA54C9E" pageId="18" pageNumber="251" type="activity">
<paragraph id="8927369C073EFF91FF94F5C96FA54C9E" blockId="18.[103,1311,1344,3032]" pageId="18" pageNumber="251">
<emphasis id="BBECEA8E073EFF91FF94F5C96A5C4C4F" bold="true" box="[103,339,2608,2637]" pageId="18" pageNumber="251">Activity patterns.</emphasis>
There is no specific information available for this species, but Goldmans Pocket Gopher is probably active at any hour of the day, with periods of peak activity around dawn and dusk. It does not hibernate and is active year-round.
</paragraph>
</subSubSection>
<subSubSection id="C1826517073EFF91FF9AF55F6F704D3B" pageId="18" pageNumber="251" type="biology_ecology">
<paragraph id="8927369C073EFF91FF9AF55F6F704D3B" blockId="18.[103,1311,1344,3032]" pageId="18" pageNumber="251">
<emphasis id="BBECEA8E073EFF91FF9AF55F68374CC1" bold="true" box="[105,824,2726,2755]" pageId="18" pageNumber="251">Movements, Home range and Social organization.</emphasis>
There is no specific information available for this species, but Goldmans Pocket Gopher is likely solitary and aggressively territorial. Individuals probably leave their burrow systems only rarely, meaning that their home range is defined by size and extent of their burrow system.
</paragraph>
</subSubSection>
<subSubSection id="C1826517073EFF91FF99F4BA6F8C4D62" box="[106,1155,2883,2912]" pageId="18" pageNumber="251" type="conservation">
<paragraph id="8927369C073EFF91FF99F4BA6F8C4D62" blockId="18.[103,1311,1344,3032]" box="[106,1155,2883,2912]" pageId="18" pageNumber="251">
<emphasis id="BBECEA8E073EFF91FF99F4BA6AC94D62" bold="true" box="[106,454,2883,2912]" pageId="18" pageNumber="251">Status and Conservation.</emphasis>
Classified as Least Concern on The IUCN Red Lust.
</paragraph>
</subSubSection>
<subSubSection id="C1826517073EFF91FF99F4896BE74DDA" pageId="18" pageNumber="251" type="bibRefCitation_list">
<paragraph id="8927369C073EFF91FF99F4896BE74DDA" blockId="18.[103,1311,1344,3032]" pageId="18" pageNumber="251">
<emphasis id="BBECEA8E073EFF91FF99F4896BF24D8B" bold="true" box="[106,253,2928,2953]" pageId="18" pageNumber="251">Bibliography.</emphasis>
Alvarez-Castafeda, Castro-Arellano, Lacher &amp; Vazquez (2008e), Ceballos (2014), Hafner, Hafner et al. (2008), Honeycutt &amp; Williams (1982), Lee Hae-Kyung &amp; Baker (1987), Patton (2005b), Russell (1968b), Spradling et al. (1993).
</paragraph>
</subSubSection>
</treatment>
</document>