164 lines
18 KiB
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164 lines
18 KiB
XML
<document id="773AB88A5F65D6E744F4ACCF14EC3706" ID-CLB-Dataset="30062" ID-DOI="http://dx.doi.org/10.3897/zse.94.25460" ID-GBIF-Dataset="9391a8f6-116c-4bb3-9264-a360c659366f" ID-Pensoft-Pub="1860-0743-2-305" ID-ZooBank="C137A5DF3F684475A1B94A19AD684C6C" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1860-0743-2-305" ModsDocOrigin="Zoosystematics and Evolution 94 (2)" ModsDocTitle="Taxonomic study of the leafmining genus Liocrobyla Meyrick, 1916 from China (Lepidoptera, Gracillariidae, Ornixolinae) with a description of one new species" checkinTime="1526951388636" checkinUser="pensoft" docAuthor="Liu, Tengteng, Wang, Encui & Wang, Shuxia" docDate="2018" docId="9D5FCDDA4FFAF5CACE9A3CC236C9A5E4" docLanguage="en" docName="ZoosystEvol 94(2): 305-314" docOrigin="Zoosystematics and Evolution 94 (2)" docSource="http://dx.doi.org/10.3897/zse.94.25460" docTitle="Liocrobyla indigofera Liu, Wang & Wang, 2018, sp. n." docType="treatment" docUuid="2D79B9E2-C197-4377-B9F0-43F507168183" docUuidSource="ZooBank" docVersion="7" lastPageNumber="307" masterDocId="485BFFDFFFD5FF9BFFBAFFC8FFEC5736" masterDocTitle="Taxonomic study of the leafmining genus Liocrobyla Meyrick, 1916 from China (Lepidoptera, Gracillariidae, Ornixolinae) with a description of one new species" masterLastPageNumber="314" masterPageNumber="305" pageNumber="307" updateTime="1701381223587" updateUser="ExternalLinkService">
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<mods:titleInfo id="36E2D6182E9C6BC1D3B4E627EA8B34BF">
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<mods:title id="8671F88B91B19B8AEB1B2E445778E719">Taxonomic study of the leafmining genus Liocrobyla Meyrick, 1916 from China (Lepidoptera, Gracillariidae, Ornixolinae) with a description of one new species</mods:title>
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<mods:name id="19B6D3278186559FD49F208C9854BAC6" type="personal">
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<mods:roleTerm id="4377EF32D2F3C2DD7678BB6E74474F3C">Author</mods:roleTerm>
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<mods:namePart id="AF02D7C75029935B8F155134BA7CE6E6">Liu, Tengteng</mods:namePart>
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<mods:roleTerm id="EB186A7C3F331207140D6200A83A83C1">Author</mods:roleTerm>
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<mods:namePart id="E04613CDF6527D8A9402D8269615F0AF">Wang, Encui</mods:namePart>
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<mods:name id="A5B6C95E37C177A658065D1EB7049AE1" type="personal">
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<mods:roleTerm id="A45F61B747709EE094196D71F5131924">Author</mods:roleTerm>
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<mods:namePart id="9A84466289AB3068BB4A4F2DA0CC31B4">Wang, Shuxia</mods:namePart>
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<mods:typeOfResource id="DCF92F4E8AA17B8FA7E22E913FCEC928">text</mods:typeOfResource>
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<mods:title id="BFB014C05A021D1EE002DF6B56153B3A">Zoosystematics and Evolution</mods:title>
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<mods:part id="1C2B7004935C1936D61965B6435E9C81">
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<mods:date id="2A117CFBF5A64854A4B3E21DCBAD6448">2018</mods:date>
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<mods:detail id="4D452848570D2BB52871214EC20135E9" type="volume">
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<mods:number id="D2DD9F187B37734C7B7FED350A2E85CE">94</mods:number>
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</mods:detail>
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<mods:detail id="301F6F527F1DF4D1F2B2DB3E5261AB8F" type="issue">
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<mods:number id="891681E492E64DA609B4C5D231FE85B8">2</mods:number>
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<mods:start id="D8AC7D2347FE19A4DAE4134297071976">305</mods:start>
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<mods:end id="077A50BF2AF72C12F193AE8C95520748">314</mods:end>
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<mods:url id="B2CC24EA0E17588ACD3977C86DD9B30F">http://dx.doi.org/10.3897/zse.94.25460</mods:url>
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<mods:classification id="58C3C7A66DA0FB7F85B9FAD4221DDAF3">journal article</mods:classification>
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<mods:identifier id="B06A684825EFFD0391503920AC9B3571" type="DOI">http://dx.doi.org/10.3897/zse.94.25460</mods:identifier>
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<mods:identifier id="5BBCD1B56D1C6F6732DAC92A16859C82" type="Pensoft-Pub">1860-0743-2-305</mods:identifier>
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</mods:mods>
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<treatment id="9D5FCDDA4FFAF5CACE9A3CC236C9A5E4" ID-GBIF-Taxon="144059764" LSID="urn:lsid:zoobank.org:act:2D79B9E2-C197-4377-B9F0-43F507168183" httpUri="http://treatment.plazi.org/id/9D5FCDDA4FFAF5CACE9A3CC236C9A5E4" lastPageNumber="307" pageId="2" pageNumber="307">
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<subSubSection id="BE2434227023FFBC557120B36B3A051B" pageId="2" pageNumber="307" type="nomenclature">
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<paragraph id="971F0DC30EB2A838AB018130928E6803" pageId="2" pageNumber="307">
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<taxonomicName id="02128969C70B5F57FA5013D77D64BF8D" ID-CoL="8VLYB" LSID="http://zoobank.org/2D79B9E2-C197-4377-B9F0-43F507168183" class="Insecta" family="Gracillariidae" genus="Liocrobyla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liocrobyla indigofera" order="Lepidoptera" pageId="2" pageNumber="307" phylum="Arthropoda" rank="species" species="indigofera">Liocrobyla indigofera</taxonomicName>
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<taxonomicNameLabel id="C6CE986FEAEA02F34FEEA34AD0753252" pageId="2" pageNumber="307">sp. n.</taxonomicNameLabel>
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Figures 3, 4, 7, 10, 12and 23-28
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</paragraph>
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</subSubSection>
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<subSubSection id="30783BA274F65757EECA3CCCE6EDA522" pageId="2" pageNumber="307" type="diagnosis">
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<paragraph id="6A579053FCF25A68924E7706A7E1913A" pageId="2" pageNumber="307">Diagnosis.</paragraph>
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<paragraph id="9C100BA4BF316AD6445A70844FA29ADE" pageId="2" pageNumber="307">
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The new species resembles
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<taxonomicName id="52613FAAA4819F9E884783133C1D473E" lsidName="L. lobata" pageId="2" pageNumber="307" rank="species" species="lobata">L. lobata</taxonomicName>
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in the male genitalia, but can be separated by the phallus longer than the valva and the minute cornutus; in
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<taxonomicName id="8622411BC8864263851B88BE6EBF9CCA" lsidName="L. lobata" pageId="2" pageNumber="307" rank="species" species="lobata">L. lobata</taxonomicName>
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, the phallus is shorter than valva and the cornutus is more than 1/12 length of the phallus. In
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<taxonomicName id="7B70572860F2C2C2D3FAF470788F2B81" lsidName="L. indigofera" pageId="2" pageNumber="307" rank="species" species="indigofera">L. indigofera</taxonomicName>
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, the head is dark grey and the forewing ground colour is blackish fuscous, while in
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<taxonomicName id="CED111EA22C3C1BDEE2A218EEC5FD1FE" lsidName="L. lobata" pageId="2" pageNumber="307" rank="species" species="lobata">L. lobata</taxonomicName>
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, the head is white and the forewing ground colour is brownish grey.
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</paragraph>
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</subSubSection>
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<subSubSection id="5764047A005C7961D80FB999578D4F79" pageId="2" pageNumber="307" type="type material">
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<paragraph id="1D4575DD3CBE09A1A0820230C2FC8914" pageId="2" pageNumber="307">Type material.</paragraph>
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<paragraph id="6C0B6545D3545B021786AA4D00FDEF27" pageId="2" pageNumber="307">
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Holotype, ♂, China: Shandong Province: Mt. Laoshan,
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<geoCoordinate id="8703F12C1C0BC4C24FCB9D39DEB3FCFA" direction="north" orientation="latitude" precision="55" value="36.204">36.204°N</geoCoordinate>
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,
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<geoCoordinate id="479CBC2BCACFD05319A593BBCA8E48DD" direction="east" orientation="longitude" precision="55" value="120.609">120.609°E</geoCoordinate>
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, 400 m, Qingdao City, 2017.vii.01, leafmine collected on
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<taxonomicName id="628F7BEA3772F4567ACAC70E4D3DA135" class="Magnoliopsida" family="Fabaceae" genus="Indigofera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Indigofera kirilowii" order="Fabales" pageId="2" pageNumber="307" phylum="Tracheophyta" rank="species" species="kirilowii">Indigofera kirilowii</taxonomicName>
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, pupated vii.05, emerged vii.18, leg. Tengteng Liu & Zhenquan Gao, genitalia slide no. LIU0030♂, registered no. SDNU.QD170710.2 (SDNU); Paratype: 1♀, genitalia slide no. LIU0029♀, registered no. SDNU.QD170710.1, other data same as holotype (SDNU).
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</paragraph>
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</subSubSection>
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<subSubSection id="AF9AD4E868A11642A8445207356633B5" pageId="2" pageNumber="307" type="other material">
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<paragraph id="E09875FCD0F365CABFA8F8697B20B62F" pageId="2" pageNumber="307">Other material.</paragraph>
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<paragraph id="543DFA9ACAC34FF86CDCB6C1F1DDD8F6" pageId="2" pageNumber="307">
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Leaf mines collected on
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<taxonomicName id="EDCEC77661F95376FB9F304C1C553B43" class="Magnoliopsida" family="Fabaceae" genus="Indigofera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Indigofera tinctoria" order="Fabales" pageId="2" pageNumber="307" phylum="Tracheophyta" rank="species" species="tinctoria">Indigofera tinctoria</taxonomicName>
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Linn., Mt. Laoshan, 120.609°E, 36.204°N, 400 m, Qingdao City, 2017.vii.01, leg. Tengteng Liu.
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</paragraph>
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<paragraph id="346D19C592D6CEACCB486FA4FEBA234F" pageId="2" pageNumber="307">
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Adult (Figs 3 and 4). Forewing length 3.0 mm. Head with frons white, vertex dark grey with a black median line. Maxillary palpus black, pointed apically, about 1/4 length of labial palpus. Labial palpus white, with black rings at middle and before apex. Antennae with scape dark dorsally, white ventrally, other segments dark with black rings. Thorax yellowish gray, with a dark central line, tegula blackish-fuscous; legs white with blackish-fuscous rings. Forewing blackish-fuscous, fuscous along dorsum, stripes and spots white; a sinuous stripe from costal 1/3 to apex of disc divided into three or occasionally two parts (which is continuous in
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<taxonomicName id="68F07A4433DB8876FA0077208C23D030" lsidName="L. lobata" pageId="2" pageNumber="307" rank="species" species="lobata">L. lobata</taxonomicName>
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); one stria at costal 2/3; a transverse outwards arched stripe at 4/5, separated at middle, with violet reflection; two spots above the fuscous stripe on dorsum, with outer one larger; two minute spots near apex; cilia white, with three black lines. Hind wing blackish-grey. Abdomen: blackish-grey dorsally, silvery white ventrally.
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</paragraph>
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<paragraph id="036A0C2733E102B88B29E48E3F2411D7" pageId="2" pageNumber="307">Male genitalia (Figs 7 and 10). Tegumen weakly sclerotized. Valva more or less rectangular, divided by a sclerotized ridge, dorsal part more sclerotized than costal one; costal part densely covered with setae apically, dorsal part with two long digital processes apically and one or two minute processes below apex. Vinculum narrowly triangular. Phallus longer than valva, curved beyond middle, pointed apically, vesica roughened, with a minute inversed cornutus. Ninth tergite more or less oval (Fig. 10).</paragraph>
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<paragraph id="90A05C6F1946689001B6C88370C9DB8C" pageId="2" pageNumber="307">Female genitalia (Fig. 12). Posterior and anterior apophyses triangular, about as long as eighth tergite. Antrum sclerotized. Ductus bursae about twice the length of seventh segment, sinuous at middle, strongly sclerotized ventrally, membranous dorsally; ductus seminalis originated from ductus bursae near antrum, sclerotized and sinuous basally. Corpus bursae oval, membranous.</paragraph>
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<caption id="F645381967609968BE0547203599BC90" pageId="2" pageNumber="307">
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<paragraph id="84EEDB20A5A63AA8C96FA0E7A5DC615F" pageId="2" pageNumber="307">
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Figures 11-12. Female genitalia of
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<taxonomicName id="1C3C56F4F3A76ADA259CAED168181607" class="Insecta" family="Gracillariidae" genus="Liocrobyla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liocrobyla" order="Lepidoptera" pageId="2" pageNumber="307" phylum="Arthropoda" rank="genus">Liocrobyla</taxonomicName>
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spp. 11.
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<taxonomicName id="AC41C8C2EC314779F96D9E0B2E44C570" lsidName="L. desmodiella" pageId="2" pageNumber="307" rank="species" species="desmodiella">L. desmodiella</taxonomicName>
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, slide no. LTT12612 (NKU); 12.
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<taxonomicName id="8AAFDCC64C216AE75602676AB78D905F" lsidName="L. indigofera" pageId="2" pageNumber="307" rank="species" species="indigofera">L. indigofera</taxonomicName>
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sp. n., slide no. LIU0029, paratype. Scale bars: 0.5 mm.
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</paragraph>
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</caption>
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<paragraph id="64680928945EEA11DAF0B8DAC373C120" pageId="2" pageNumber="307">Biology (Figs 23-28). Larval mine is a white blotch, with several digital galleries, always located in the middle of the upper side of a leaflet (Figs 23 and 27). A frass opening on the lower side of the leaflet always locates close to the midrib, covered by a piece of white dry leaf tissue (Fig. 28). A single mine per leaflet (Figs 23-25).</paragraph>
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</subSubSection>
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<subSubSection id="270ADD7020CD0232673538842998BEF6" pageId="2" pageNumber="307" type="host plants">
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<paragraph id="74C956DBB486D18B0CF1D849E7A16D18" pageId="2" pageNumber="307">Host plants.</paragraph>
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<paragraph id="57C569CA97E316244EE9FD6FF9324885" pageId="2" pageNumber="307">
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<taxonomicName id="8178C23F058B2B1BF716DC354A74D39D" family="Fabaceae" lsidName="" pageId="2" pageNumber="307" rank="family">Fabaceae</taxonomicName>
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:
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<taxonomicName id="0E6F7A99EBAFAFC42FFE9158CD1DBDBE" class="Magnoliopsida" family="Fabaceae" genus="Indigofera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Indigofera kirilowii" order="Fabales" pageId="2" pageNumber="307" phylum="Tracheophyta" rank="species" species="kirilowii">Indigofera kirilowii</taxonomicName>
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Palib.,
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<taxonomicName id="FB6DFF3F9BE2103D656DE1BF5FDD4D1D" lsidName="I. tinctoria" pageId="2" pageNumber="307" rank="species" species="tinctoria">I. tinctoria</taxonomicName>
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L.
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</paragraph>
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</subSubSection>
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<subSubSection id="E90E9748CAF2C36ED9DAC5B2E850EC1D" pageId="2" pageNumber="307" type="distribution">
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<paragraph id="4C8AF35880B80E589FDB371C0FE7304A" pageId="2" pageNumber="307">Distribution.</paragraph>
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<paragraph id="40EDC0DA1A5DDC8B1FCE7BBD2DCD5DFC" pageId="2" pageNumber="307">China (Shandong).</paragraph>
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</subSubSection>
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<subSubSection id="D9C539DF6E0C7D8760E4AB093B0C1DB0" pageId="2" pageNumber="307" type="etymology">
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<paragraph id="A4841680BCB9EA1AC2A50E2E29C73351" pageId="2" pageNumber="307">Etymology.</paragraph>
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<paragraph id="966617A86ECCF9990BF895CC73261B82" pageId="2" pageNumber="307">The specific name is derived from the genus name of the host plants.</paragraph>
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<caption id="7ED914009996977F8B2A7254428DD1D0" pageId="2" pageNumber="307">
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<paragraph id="276D58000687DA6B9005C00D1710C0C2" pageId="2" pageNumber="307">
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Figures 13-20. Biology of
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<taxonomicName id="8A436A20B5281D88C8DB77B4F4381CD4" class="Insecta" family="Gracillariidae" genus="Liocrobyla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liocrobyla" order="Lepidoptera" pageId="2" pageNumber="307" phylum="Arthropoda" rank="genus">Liocrobyla</taxonomicName>
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spp. 13-17.
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<taxonomicName id="C65C52E4535358CA5420A28984053604" lsidName="L. desmodiella" pageId="2" pageNumber="307" rank="species" species="desmodiella">L. desmodiella</taxonomicName>
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, 13. host plant,
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<taxonomicName id="3B1078E01A40645B03B73881F635B1C1" class="Magnoliopsida" family="Fabaceae" genus="Lespedeza" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Lespedeza bicolor" order="Fabales" pageId="2" pageNumber="307" phylum="Tracheophyta" rank="species" species="bicolor">Lespedeza bicolor</taxonomicName>
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; 14. leafmine; 15. frass opening on upper side of leaf; 16. frass opening on lower side of leaf; 17. cocoon and pupal exuviae; 18-20.
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<taxonomicName id="729AB141DC64129B8B172D86772056AC" lsidName="L. lobata" pageId="2" pageNumber="307" rank="species" species="lobata">L. lobata</taxonomicName>
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, 18. host plant,
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<taxonomicName id="5AF46F62D9E50BD47876B5866C76AE91" class="Magnoliopsida" family="Fabaceae" genus="Pueraria" higherTaxonomySource="CoL" infraspecific-rank="var." kingdom="Plantae" lsidName="Pueraria montana var. lobata" order="Fabales" pageId="2" pageNumber="307" phylum="Tracheophyta" rank="variety" species="montana" variety="lobata">Pueraria montana var. lobata</taxonomicName>
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; 19. leafmines; 20. an opened mine with a larva exposed.
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</paragraph>
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</caption>
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<caption id="64DCE4099E6226FD814FDBD96BFFF62E" pageId="2" pageNumber="307">
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<paragraph id="D841E9D09598538BBD580A0441436E14" pageId="2" pageNumber="307">
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Figures 21-28. Host plants, leafmines and biology of
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<taxonomicName id="B1A654FD86FE09C100C6044BAFE5175B" class="Insecta" family="Gracillariidae" genus="Liocrobyla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liocrobyla" order="Lepidoptera" pageId="2" pageNumber="307" phylum="Arthropoda" rank="genus">Liocrobyla</taxonomicName>
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spp. 21-22,
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<taxonomicName id="EC5CF84F216F318AD36899530E7DC7E2" lsidName="L. lobata" pageId="2" pageNumber="307" rank="species" species="lobata">L. lobata</taxonomicName>
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, 21. leafmine with an alive larva; 22. frass opening on lower side of leaf; 23-28.
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<taxonomicName id="AEFCD274DA2814CA50FFECABCBB29103" lsidName="L. indigofera" pageId="2" pageNumber="307" rank="species" species="indigofera">L. indigofera</taxonomicName>
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sp. n., 23. leafmines each on a leaf of
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<taxonomicName id="F578F19FF5F1CC59F9C10B3847038FB8" class="Magnoliopsida" family="Fabaceae" genus="Indigofera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Indigofera kirilowii" order="Fabales" pageId="2" pageNumber="307" phylum="Tracheophyta" rank="species" species="kirilowii">Indigofera kirilowii</taxonomicName>
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; 24-25. leafmines on
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<taxonomicName id="AC3D17C304B0E3EED88C5ED022D6D139" lsidName="I. tinctoria" pageId="2" pageNumber="307" rank="species" species="tinctoria">I. tinctoria</taxonomicName>
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; 26. raceme of
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<taxonomicName id="AC961E76752646B73D84C3B50195229A" lsidName="I. tinctoria" pageId="2" pageNumber="307" rank="species" species="tinctoria">I. tinctoria</taxonomicName>
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; 27. leafmine with a living larva, identical mine to the lower-right mine in Fig. 23; 28. frass opening on lower side of leaf, identical mine to Fig. 27.
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</paragraph>
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</caption>
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</subSubSection>
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</treatment>
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</document> |