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<mods:title id="00A47F4919E9874757B79B8143407610">A revision of the genus Porteria and the phylogeny and biogeography of Porteriinae (Araneae: Desidae)</mods:title>
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<mods:namePart id="4AC3CE95341D6DDD7E260B1D78A2D632">Ramírez, Martín J.</mods:namePart>
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<taxonomicName id="5DCB4D049A29851FFED8FB1CFE20FABA" ID-CoL="63PPN" ID-ENA="1956907" authorityName="Simon" authorityYear="1904" box="[318,497,1272,1297]" class="Arachnida" family="Desidae" genus="Porteria" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="396" phylum="Arthropoda" rank="genus">
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This morphologically distinctive set of species is recognized by the common plesiomorphic genitalic characters of both males and females. Our phylogeny shows this set to be paraphyletic, so we refer to this as the
<taxonomicName id="5DCB4D049A29851FFF45FA78FE01FA1A" authority="Grade. Most" authorityName="Grade. Most" box="[163,464,1436,1458]" class="Arachnida" family="Desidae" genus="Porteria" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="396" phylum="Arthropoda" rank="species" species="albopunctata">Albopunctata Grade. Most</taxonomicName>
diagnostic are the tegulum without a median concavity (
<figureCitation id="02F02A029A29851FFE09FA5EFD85FA7B" box="[495,596,1466,1488]" captionStart="Figure 26" captionStartId="34.[163,242,1062,1084]" captionTargetBox="[179,1426,195,1032]" captionTargetId="figure-390@34.[179,1426,195,1032]" captionTargetPageId="34" captionText="Figure 26. Left male palp of P. albopunctata sp. nov. from Wellington Island, Puerto Edén, MCZ46441. A, prolateral view. B, ventral view. C, retrolateral view. D, dorsal view. Illustration by Rachel Diaz-Bastin." figureDoi="http://doi.org/10.5281/zenodo.8007469" httpUri="https://zenodo.org/record/8007469/files/figure.png" pageId="28" pageNumber="396">Figs 26B</figureCitation>
,
<figureCitation id="02F02A029A29851FFD87FA5EFD5CFA64" box="[609,653,1466,1487]" captionStart="Figure 29" captionStartId="37.[146,227,986,1008]" captionTargetBox="[171,1396,206,952]" captionTargetId="figure-424@37.[161,1408,195,956]" captionTargetPageId="37" captionText="Figure 29. Left male palp of P. fiura sp. nov. AMNH paratype, CASENT9044672, Chiloé Island. A, prolateral view. B, ventral view. C, retrolateral view. D, tibia, dorsal view. Illustration by Rachel Diaz-Bastin." figureDoi="http://doi.org/10.5281/zenodo.8007475" httpUri="https://zenodo.org/record/8007475/files/figure.png" pageId="28" pageNumber="396">29B</figureCitation>
,
<figureCitation id="02F02A029A29851FFD7CFA5EFD19FA64" box="[666,712,1466,1487]" captionStart="Figure 33" captionStartId="41.[145,223,1042,1064]" captionTargetBox="[165,1395,195,1013]" captionTargetId="figure-383@41.[161,1408,195,1013]" captionTargetPageId="41" captionText="Figure 33. Left male palp of P. eddardstarki sp. nov. from Valparaíso Province, Cuesta Pucalán (CASENT9032846), details of tibia and tarsus and outlines of whole tibia and tarsus. A, prolateral view. B, ventral view. C, retrolateral view. Illustration by Giovanni Maki." figureDoi="http://doi.org/10.5281/zenodo.8007483" httpUri="https://zenodo.org/record/8007483/files/figure.png" pageId="28" pageNumber="396">33B</figureCitation>
), outlined by the dark sperm duct; embolus base elongated and narrow, not shield like nor heavily sclerotized, hugging prolateral margin of conductor (
<figureCitation id="02F02A029A29851FFD99F9F2FCD4F987" box="[639,773,1558,1580]" captionStart="Figure 26" captionStartId="34.[163,242,1062,1084]" captionTargetBox="[179,1426,195,1032]" captionTargetId="figure-390@34.[179,1426,195,1032]" captionTargetPageId="34" captionText="Figure 26. Left male palp of P. albopunctata sp. nov. from Wellington Island, Puerto Edén, MCZ46441. A, prolateral view. B, ventral view. C, retrolateral view. D, dorsal view. Illustration by Rachel Diaz-Bastin." figureDoi="http://doi.org/10.5281/zenodo.8007469" httpUri="https://zenodo.org/record/8007469/files/figure.png" pageId="28" pageNumber="396">Figs 26A, B</figureCitation>
,
<figureCitation id="02F02A029A29851FFF45F9D1FF00F9E1" box="[163,209,1589,1610]" captionStart="Figure 29" captionStartId="37.[146,227,986,1008]" captionTargetBox="[171,1396,206,952]" captionTargetId="figure-424@37.[161,1408,195,956]" captionTargetPageId="37" captionText="Figure 29. Left male palp of P. fiura sp. nov. AMNH paratype, CASENT9044672, Chiloé Island. A, prolateral view. B, ventral view. C, retrolateral view. D, tibia, dorsal view. Illustration by Rachel Diaz-Bastin." figureDoi="http://doi.org/10.5281/zenodo.8007475" httpUri="https://zenodo.org/record/8007475/files/figure.png" pageId="28" pageNumber="396">29A</figureCitation>
,
<figureCitation id="02F02A029A29851FFF06F9D1FEDFF9E1" box="[224,270,1589,1610]" captionStart="Figure 33" captionStartId="41.[145,223,1042,1064]" captionTargetBox="[165,1395,195,1013]" captionTargetId="figure-383@41.[161,1408,195,1013]" captionTargetPageId="41" captionText="Figure 33. Left male palp of P. eddardstarki sp. nov. from Valparaíso Province, Cuesta Pucalán (CASENT9032846), details of tibia and tarsus and outlines of whole tibia and tarsus. A, prolateral view. B, ventral view. C, retrolateral view. Illustration by Giovanni Maki." figureDoi="http://doi.org/10.5281/zenodo.8007483" httpUri="https://zenodo.org/record/8007483/files/figure.png" pageId="28" pageNumber="396">33A</figureCitation>
,
<figureCitation id="02F02A029A29851FFEFBF9D1FEBDF9E1" box="[285,364,1589,1610]" captionStart="Figure 39" captionStartId="48.[164,242,899,921]" captionTargetBox="[185,1419,201,867]" captionTargetId="figure-493@48.[179,1426,195,870]" captionTargetPageId="48" captionText="Figure 39. Left male palp of P. ajimayo sp. nov., from Parque Pedro del Río Zañartu, (CASENT9053784). A, prolateral view. B, ventral view. C, retrolateral view. D, tibia, dorsal view. Illustration by Rachel Diaz-Bastin." figureDoi="http://doi.org/10.5281/zenodo.8007499" httpUri="https://zenodo.org/record/8007499/files/figure.png" pageId="28" pageNumber="396">39A, B</figureCitation>
). LRTA projected straight up from tibia, unbent, in line with frontal plane. Paracymbium absent (
<figureCitation id="02F02A029A29851FFF19F996FEB4F923" box="[255,357,1650,1672]" captionStart="Figure 26" captionStartId="34.[163,242,1062,1084]" captionTargetBox="[179,1426,195,1032]" captionTargetId="figure-390@34.[179,1426,195,1032]" captionTargetPageId="34" captionText="Figure 26. Left male palp of P. albopunctata sp. nov. from Wellington Island, Puerto Edén, MCZ46441. A, prolateral view. B, ventral view. C, retrolateral view. D, dorsal view. Illustration by Rachel Diaz-Bastin." figureDoi="http://doi.org/10.5281/zenodo.8007469" httpUri="https://zenodo.org/record/8007469/files/figure.png" pageId="28" pageNumber="396">Figs 26C</figureCitation>
,
<figureCitation id="02F02A029A29851FFE94F996FE4EF92C" box="[370,415,1650,1671]" captionStart="Figure 29" captionStartId="37.[146,227,986,1008]" captionTargetBox="[171,1396,206,952]" captionTargetId="figure-424@37.[161,1408,195,956]" captionTargetPageId="37" captionText="Figure 29. Left male palp of P. fiura sp. nov. AMNH paratype, CASENT9044672, Chiloé Island. A, prolateral view. B, ventral view. C, retrolateral view. D, tibia, dorsal view. Illustration by Rachel Diaz-Bastin." figureDoi="http://doi.org/10.5281/zenodo.8007475" httpUri="https://zenodo.org/record/8007475/files/figure.png" pageId="28" pageNumber="396">29C</figureCitation>
,
<figureCitation id="02F02A029A29851FFE4AF996FE08F92C" box="[428,473,1650,1671]" captionStart="Figure 33" captionStartId="41.[145,223,1042,1064]" captionTargetBox="[165,1395,195,1013]" captionTargetId="figure-383@41.[161,1408,195,1013]" captionTargetPageId="41" captionText="Figure 33. Left male palp of P. eddardstarki sp. nov. from Valparaíso Province, Cuesta Pucalán (CASENT9032846), details of tibia and tarsus and outlines of whole tibia and tarsus. A, prolateral view. B, ventral view. C, retrolateral view. Illustration by Giovanni Maki." figureDoi="http://doi.org/10.5281/zenodo.8007483" httpUri="https://zenodo.org/record/8007483/files/figure.png" pageId="28" pageNumber="396">33C</figureCitation>
,
<figureCitation id="02F02A029A29851FFE00F996FDC5F92C" box="[486,532,1650,1671]" captionStart="Figure 39" captionStartId="48.[164,242,899,921]" captionTargetBox="[185,1419,201,867]" captionTargetId="figure-493@48.[179,1426,195,870]" captionTargetPageId="48" captionText="Figure 39. Left male palp of P. ajimayo sp. nov., from Parque Pedro del Río Zañartu, (CASENT9053784). A, prolateral view. B, ventral view. C, retrolateral view. D, tibia, dorsal view. Illustration by Rachel Diaz-Bastin." figureDoi="http://doi.org/10.5281/zenodo.8007499" httpUri="https://zenodo.org/record/8007499/files/figure.png" pageId="28" pageNumber="396">39C</figureCitation>
). Epigyne atrium absent; copulatory openings flush with surface of epigynal plate (
<figureCitation id="02F02A029A29851FFF0DF954FE9DF96E" box="[235,332,1711,1734]" captionStart="Figure 25" captionStartId="33.[146,226,1571,1593]" captionTargetBox="[237,1332,196,1538]" captionTargetId="figure-93@33.[234,1335,193,1541]" captionTargetPageId="33" captionText="Figure 25. Genitalia of P. albopunctata sp. nov., A-E, lectotype from Punta Arenas, MNHN22323, CASENT9055543. A, ventral view. B, retrolateral view. C, dorsal view. D, prolateral view. E, ventral view with entire cymbium. F-H, female from Laguna San Rafael National Park, CASENT9044702. F, ventral view. G, lateral view showing small scape. H, dorsal view showing internal structures. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8007467" httpUri="https://zenodo.org/record/8007467/files/figure.png" pageId="28" pageNumber="396">Figs 25F</figureCitation>
,
<figureCitation id="02F02A029A29851FFEBFF954FE57F96E" box="[345,390,1712,1733]" captionStart="Figure 28" captionStartId="36.[161,240,1111,1133]" captionTargetBox="[182,1424,198,1078]" captionTargetId="figure-360@36.[179,1427,195,1081]" captionTargetPageId="36" captionText="Figure 28. Genitalia of P. fiura sp. nov. A-D, male from Chiloé Island CASENT9044672. A, left palp, retrolateral view. B, ventral view. C, prolateral view. D, dorsal view. E, F, female from Chiloé, CASENT9055645. E, ventral view. F, dorsal view, showing internal structures. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8007473" httpUri="https://zenodo.org/record/8007473/files/figure.png" pageId="28" pageNumber="396">28E</figureCitation>
,
<figureCitation id="02F02A029A29851FFE75F954FE11F96E" box="[403,448,1712,1733]" captionStart="Figure 30" captionStartId="38.[163,246,1113,1135]" captionTargetBox="[185,1420,198,1080]" captionTargetId="figure-384@38.[182,1423,195,1083]" captionTargetPageId="38" captionText="Figure 30. Scanning electron micrograph of epigynum and vulva of P. fiura sp. nov. from Chiloé National Park, CASENT9055645. A, ventral view. B, dorsal view. C, close-up of scape and copulatory openings. D, head of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.8007477" httpUri="https://zenodo.org/record/8007477/files/figure.png" pageId="28" pageNumber="396">30A</figureCitation>
,
<figureCitation id="02F02A029A29851FFE2BF94BFE34F96E" box="[461,485,1711,1733]" captionStart="Figure 35" captionStartId="43.[145,225,1140,1162]" captionTargetBox="[161,1409,198,1108]" captionTargetId="figure-343@43.[158,1412,195,1111]" captionTargetPageId="43" captionText="Figure 35. Genitalia of female P. eddardstarki sp. nov. A-C ventral view showing variability in scape form. A, from Los Molles CASENT9044704. B, C, from Pichicuy MACN-AR21060. D, dorsal view showing internal structures, from Zapallar CASENT9055699. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8007489" httpUri="https://zenodo.org/record/8007489/files/figure.png" pageId="28" pageNumber="396">35</figureCitation>
A-C, 41A, 43E). Our phylogeny (
<figureCitation id="02F02A029A29851FFF1CF92AFE9CF948" box="[250,333,1742,1764]" captionStart="Figure 73" captionStartId="86.[163,241,1523,1545]" captionTargetBox="[179,1426,196,1492]" captionTargetId="figure-200@86.[798,1428,582,973]" captionTargetPageId="86" captionText="Figure 73. Maximum clade credibility tree of the Bayesian phylogenetic analysis." figureDoi="http://doi.org/10.5281/zenodo.8007570" httpUri="https://zenodo.org/record/8007570/files/figure.png" pageId="28" pageNumber="396">Fig. 73</figureCitation>
) implies those are all plesiomorphic traits for
<taxonomicName id="5DCB4D049A29851FFEE9F90AFEBBF8A8" authorityName="Simon" authorityYear="1904" box="[271,362,1774,1795]" class="Arachnida" family="Desidae" genus="Porteria" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="396" phylum="Arthropoda" rank="genus">
<emphasis id="A8BFEA959A29851FFEE9F90AFEBBF8A8" box="[271,362,1774,1795]" italics="true" pageId="28" pageNumber="396">Porteria</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
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<paragraph id="9A7436879A29851FFF45F8C8FD39F8CB" blockId="28.[163,780,1836,1888]" pageId="28" pageNumber="396">
<emphasis id="A8BFEA959A29851FFF45F8C8FD7DF8EA" box="[163,684,1836,1857]" italics="true" pageId="28" pageNumber="396">
Included species:
<taxonomicName id="5DCB4D049A29851FFE76F8C8FD7DF8EA" box="[400,684,1836,1857]" class="Arachnida" family="Desidae" genus="Porteria" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="396" phylum="Arthropoda" rank="species" species="albopunctata">Porteria albopunctata</taxonomicName>
</emphasis>
and
<taxonomicName id="5DCB4D049A29851FFD13F8C8FF2FF8CB" authorityName="Morrill &amp; Crews &amp; Esposito &amp; Ramírez &amp; Griswold" authorityYear="2023" class="Arachnida" family="Desidae" genus="Porteria" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="396" phylum="Arthropoda" rank="species" species="ajimayo">
<emphasis id="A8BFEA959A29851FFD13F8C8FF2FF8CB" italics="true" pageId="28" pageNumber="396">P. ajimayo</emphasis>
</taxonomicName>
,
<taxonomicName id="5DCB4D049A29851FFEECF8AFFE68F8F4" authorityName="Morrill &amp; Crews &amp; Esposito &amp; Ramírez &amp; Griswold" authorityYear="2023" box="[266,441,1866,1888]" class="Arachnida" family="Desidae" genus="Porteria" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="396" phylum="Arthropoda" rank="species" species="eddardstarki">
<emphasis id="A8BFEA959A29851FFEECF8AFFE68F8F4" box="[266,441,1866,1888]" italics="true" pageId="28" pageNumber="396">P. eddardstarki</emphasis>
</taxonomicName>
,
<taxonomicName id="5DCB4D049A29851FFE23F8AFFDC7F8F4" authorityName="Morrill &amp; Crews &amp; Esposito &amp; Ramírez &amp; Griswold" authorityYear="2023" box="[453,534,1866,1888]" class="Arachnida" family="Desidae" genus="Porteria" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="396" phylum="Arthropoda" rank="species" species="fiura">
<emphasis id="A8BFEA959A29851FFE23F8AFFDC7F8F4" box="[453,534,1866,1888]" italics="true" pageId="28" pageNumber="396">P. fiura</emphasis>
</taxonomicName>
and
<taxonomicName id="5DCB4D049A29851FFDA9F8AFFD30F8F4" box="[591,737,1866,1888]" class="Arachnida" family="Desidae" genus="Porteria" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="396" phylum="Arthropoda" rank="species" species="misbianka">
<emphasis id="A8BFEA959A29851FFDA9F8AFFD30F8F4" box="[591,737,1866,1888]" italics="true" pageId="28" pageNumber="396">P. misbianka</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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