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<document id="EC33BBEB04F3AF00B6EEA8830C14B583" ID-CLB-Dataset="54267" ID-DOI="10.1093/zoolinnean/zlac093" ID-GBIF-Dataset="e327cc37-7927-4c15-98a9-dd956c03b3b7" ID-ISSN="0024-4082" ID-Zenodo-Dep="8007404" ID-ZooBank="76B9F689-4B90-433C-9837-92E49F1DDE80" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="diego" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="diego" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1685958949275" checkinUser="plazi" docAuthor="Morrill, Elizabeth, Crews, Sarah, Esposito, Lauren, Ramírez, Martín J. &amp; Griswold, Charles" docDate="2023" docId="126287919A178525FEDBF966FB56FA57" docLanguage="en" docName="zlac093.pdf" docOrigin="Zoological Journal of the Linnean Society 198 (2)" docSource="http://dx.doi.org/10.1093/zoolinnean/zlac093" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Porteria fiura Morrill &amp; Crews &amp; Esposito &amp; Ramírez &amp; Griswold 2023, SP. NOV." docType="treatment" docVersion="9" lastPageNumber="406" masterDocId="EE5BFFE99A358503FFE6FFE4FFD1FFAB" masterDocTitle="A revision of the genus Porteria and the phylogeny and biogeography of Porteriinae (Araneae: Desidae)" masterLastPageNumber="461" masterPageNumber="368" pageNumber="402" updateTime="1699516901368" updateUser="ExternalLinkService">
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<mods:title id="2F80C2D04F90ECEF584F69BFFFA31B8F">A revision of the genus Porteria and the phylogeny and biogeography of Porteriinae (Araneae: Desidae)</mods:title>
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<mods:namePart id="7FD563E33ED2E9A45F60A41AB7C11DDF">Morrill, Elizabeth</mods:namePart>
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<mods:namePart id="B5692551DF37005CB90C6ACF93510F80">Ramírez, Martín J.</mods:namePart>
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<treatment id="126287919A178525FEDBF966FB56FA57" ID-DOI="http://doi.org/10.5281/zenodo.8004928" ID-GBIF-Taxon="209669346" ID-Zenodo-Dep="8004928" LSID="urn:lsid:plazi:treatment:126287919A178525FEDBF966FB56FA57" httpUri="http://treatment.plazi.org/id/126287919A178525FEDBF966FB56FA57" lastPageId="38" lastPageNumber="406" pageId="34" pageNumber="402">
<subSubSection id="D2D1650C9A178521FEDBF966FDC9F96A" pageId="34" pageNumber="402" type="nomenclature">
<paragraph id="9A7436879A178521FEDBF966FDA1F932" blockId="34.[317,624,1665,1689]" box="[317,624,1665,1689]" pageId="34" pageNumber="402">
<heading id="C13C81EB9A178521FEDBF966FDA1F932" box="[317,624,1665,1689]" centered="true" fontSize="9" level="2" pageId="34" pageNumber="402" reason="2">
<taxonomicName id="5DCB4D049A178521FEDBF966FDD4F933" authority="Morrill &amp; Crews &amp; Esposito &amp; Ramírez &amp; Griswold, 2023" authorityName="Morrill &amp; Crews &amp; Esposito &amp; Ramírez &amp; Griswold" authorityYear="2023" box="[317,517,1666,1689]" class="Arachnida" family="Desidae" genus="Porteria" kingdom="Animalia" order="Araneae" pageId="34" pageNumber="456" phylum="Arthropoda" rank="species" species="fiura" status="sp. nov.">
<smallCapsWord id="9C92A05B9A178521FEDBF966FE66F932" baselines="1683,1683" box="[317,439,1666,1689]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Porteria" pageId="34" pageNumber="402">PORTERIA</smallCapsWord>
<smallCapsWord id="9C92A05B9A178521FE58F961FDD4F933" baselines="1683" box="[446,517,1669,1688]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="fiura" pageId="34" pageNumber="402">FIURA</smallCapsWord>
</taxonomicName>
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<emphasis id="A8BFEA959A178521FDEBF961FDA1F932" bold="true" box="[525,624,1665,1689]" pageId="34" pageNumber="402">
<smallCapsWord id="9C92A05B9A178521FDEBF961FDFBF933" baselines="1683" box="[525,554,1669,1688]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="sp" pageId="34" pageNumber="402">SP</smallCapsWord>
.
<smallCapsWord id="9C92A05B9A178521FDDEF961FDB8F933" baselines="1683" box="[568,617,1669,1688]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="nov" pageId="34" pageNumber="402">NOV</smallCapsWord>
.
</emphasis>
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</heading>
</paragraph>
<paragraph id="9A7436879A178521FE73F94CFDC9F96A" blockId="34.[405,536,1704,1729]" box="[405,536,1704,1729]" pageId="34" pageNumber="402">
<figureCitation id="02F02A029A178521FE73F94CFDC9F96A" box="[405,536,1704,1729]" captionStart-0="Figure 27" captionStart-1="Figure 28" captionStart-2="Figure 29" captionStart-3="Figure 30" captionStartId-0="35.[145,224,1559,1581]" captionStartId-1="36.[161,240,1111,1133]" captionStartId-2="37.[146,227,986,1008]" captionStartId-3="38.[163,246,1113,1135]" captionTargetBox-0="[164,1406,198,1526]" captionTargetBox-1="[182,1424,198,1078]" captionTargetBox-2="[171,1396,206,952]" captionTargetBox-3="[185,1420,198,1080]" captionTargetId-0="figure-165@35.[161,1409,195,1529]" captionTargetId-1="figure-360@36.[179,1427,195,1081]" captionTargetId-2="figure-424@37.[161,1408,195,956]" captionTargetId-3="figure-384@38.[182,1423,195,1083]" captionTargetPageId-0="35" captionTargetPageId-1="36" captionTargetPageId-2="37" captionTargetPageId-3="38" captionText-0="Figure 27. Habitus of P. fiura sp. nov. A-C, male from Chiloé Island CASENT9044672. A, dorsal view. B, ventral view. C, lateral view. D-F, female from Chiloé National Park CASENT9056299 and 9055645. D, dorsal view. E, ventral view. F, lateral view. Scale bars = 1 mm." captionText-1="Figure 28. Genitalia of P. fiura sp. nov. A-D, male from Chiloé Island CASENT9044672. A, left palp, retrolateral view. B, ventral view. C, prolateral view. D, dorsal view. E, F, female from Chiloé, CASENT9055645. E, ventral view. F, dorsal view, showing internal structures. Scale bars = 0.2 mm." captionText-2="Figure 29. Left male palp of P. fiura sp. nov. AMNH paratype, CASENT9044672, Chiloé Island. A, prolateral view. B, ventral view. C, retrolateral view. D, tibia, dorsal view. Illustration by Rachel Diaz-Bastin." captionText-3="Figure 30. Scanning electron micrograph of epigynum and vulva of P. fiura sp. nov. from Chiloé National Park, CASENT9055645. A, ventral view. B, dorsal view. C, close-up of scape and copulatory openings. D, head of spermatheca." figureDoi-0="http://doi.org/10.5281/zenodo.8007471" figureDoi-1="http://doi.org/10.5281/zenodo.8007473" figureDoi-2="http://doi.org/10.5281/zenodo.8007475" figureDoi-3="http://doi.org/10.5281/zenodo.8007477" httpUri-0="https://zenodo.org/record/8007471/files/figure.png" httpUri-1="https://zenodo.org/record/8007473/files/figure.png" httpUri-2="https://zenodo.org/record/8007475/files/figure.png" httpUri-3="https://zenodo.org/record/8007477/files/figure.png" pageId="34" pageNumber="402">
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2730
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</paragraph>
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<paragraph id="9A7436879A178521FF45F934FCD7F8AE" blockId="34.[163,779,1744,1797]" pageId="34" pageNumber="402">
<emphasis id="A8BFEA959A178521FF45F934FE3EF94D" box="[163,495,1744,1766]" italics="true" pageId="34" pageNumber="402">Zoobank registration:</emphasis>
urn: lsid: zoobank. org:act:
<uuid id="EE6D0C529A178521FF17F90BFCD2F8AE" box="[241,771,1775,1797]" pageId="34" pageNumber="402">D7676422-0EB3-461E-B74D-9CB5A7C44B99</uuid>
.
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<paragraph id="9A7436879A178521FF45F8CAFBA5FAFB" blockId="34.[163,779,1838,1890]" lastBlockId="34.[827,1443,1154,1360]" pageId="34" pageNumber="402">
<emphasis id="A8BFEA959A178521FF45F8CAFF3DF8E8" box="[163,236,1838,1859]" italics="true" pageId="34" pageNumber="402">Types:</emphasis>
<materialsCitation id="2AA33CDA9A178521FF1FF8CAFBCAFAB9" ID-GBIF-Occurrence="4126400329" collectingDate="1985-12-15" collectingDateMax="2013-01-16" collectingDateMin="1985-12-15" collectionCode="CASENT" collectorName="C. Griswold &amp; E. Morrill &amp; D. Faber &amp; Male &amp; L. E. Pena" country="Chile" county="Female" elevation="1" latitude="-42.617317" location="Chiloe Island" longLatPrecision="1" longitude="-74.10215" municipality="Chiloe Province" pageId="34" pageNumber="402" specimenCode="CASENT9055645, CASENT9044672" specimenCount="2" stateProvince="Los Lagos" typeStatus="holotype">
<collectingCounty id="73154E0B9A178521FF1FF8CAFE9EF8EF" box="[249,335,1838,1860]" pageId="34" pageNumber="402">Female</collectingCounty>
<typeStatus id="457088259A178521FEB1F8CAFE6AF8EF" box="[343,443,1838,1860]" pageId="34" pageNumber="402" type="holotype">holotype</typeStatus>
from
<collectingCountry id="E2DC76179A178521FDE2F8CAFD94F8EF" box="[516,581,1838,1860]" name="Chile" pageId="34" pageNumber="402">Chile</collectingCountry>
, X
<collectingRegion id="580FF8659A178521FD8AF8CAFF38F8C9" country="Chile" name="Los Lagos" pageId="34" pageNumber="402">Región de los Lagos</collectingRegion>
:
<collectingMunicipality id="7A10ACFD9A178521FF15F8A8FE76F8C9" box="[243,423,1868,1890]" pageId="34" pageNumber="402">Chiloé Province</collectingMunicipality>
,
<location id="9F14605C9A178521FE54F8A8FD9DF8C9" LSID="urn:lsid:plazi:treatment:126287919A178525FEDBF966FB56FA57:9F14605C9A178521FE54F8A8FD9DF8C9" box="[434,588,1868,1890]" country="Chile" county="Female" latitude="-42.617317" longLatPrecision="1" longitude="-74.10215" municipality="Chiloe Province" name="Chiloe Island" pageId="34" pageNumber="402" stateProvince="Los Lagos">Chiloé Island</location>
,
<location id="9F14605C9A178521FDB1F8A8FCA5FB33" LSID="urn:lsid:plazi:treatment:126287919A178525FEDBF966FB56FA57:9F14605C9A178521FDB1F8A8FCA5FB33" box="[599,884,1154,1890]" country="Chile" county="Female" latitude="-42.617317" longLatPrecision="1" longitude="-74.10215" municipality="Chiloe Province" name="Chiloe National Park" pageId="34" pageNumber="402" stateProvince="Los Lagos">Chiloé National Park</location>
,
<location id="9F14605C9A178521FC9BFB66FBFFFB33" LSID="urn:lsid:plazi:treatment:126287919A178525FEDBF966FB56FA57:9F14605C9A178521FC9BFB66FBFFFB33" box="[893,1070,1154,1176]" country="Chile" county="Female" latitude="-42.617317" longLatPrecision="1" longitude="-74.10215" municipality="Chiloe Province" name="Sendero Tepual" pageId="34" pageNumber="402" stateProvince="Los Lagos">Sendero Tepual</location>
,
<locationDeviation id="8BC62FA59A178521FBDEFB66FAC1FB3C" box="[1080,1296,1154,1176]" location="Cucao" pageId="34" pageNumber="402">
<quantity id="5D339B629A178521FBDEFB66FB54FB33" box="[1080,1157,1154,1176]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="34" pageNumber="402" unit="km" value="2.3">2.3 km</quantity>
NNE Cucao
</locationDeviation>
,
<geoCoordinate id="FFFF50409A178521FAFCFB66FA4EFB3C" box="[1306,1439,1154,1175]" degrees="42" direction="south" minutes="37.039" orientation="latitude" pageId="34" pageNumber="402" precision="1" value="-42.617317">S42º37.039</geoCoordinate>
,
<geoCoordinate id="FFFF50409A178521FCDDFB44FC18FB1D" box="[827,969,1184,1206]" degrees="74" direction="west" minutes="06.129" orientation="longitude" pageId="34" pageNumber="402" precision="1" value="-74.10215">W74º06.129</geoCoordinate>
, elev.
<quantity id="5D339B629A178521FBEAFB44FBE8FB1D" box="[1036,1081,1184,1206]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="34" pageNumber="402" unit="m" value="1.0">
<elevation id="11E6D1B49A178521FBEAFB44FBE8FB1D" box="[1036,1081,1184,1206]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="34" pageNumber="402" unit="m" value="1.0">1 m</elevation>
</quantity>
,
<date id="EE7510479A178521FBA2FB45FADDFB1E" box="[1092,1292,1184,1206]" pageId="34" pageNumber="402" value="2013-01-16">
<collectingDate id="FE31E9AF9A178521FBA2FB45FADDFB1E" box="[1092,1292,1184,1206]" pageId="34" pageNumber="402" value="2013-01-16">January 16, 2013</collectingDate>
</date>
,
<collectorName id="373E53519A178521FAFEFB44FA4EFB1D" box="[1304,1439,1184,1206]" pageId="34" pageNumber="402">C. Griswold</collectorName>
,
<collectorName id="373E53519A178521FCDDFB5BFC64FB7E" box="[827,949,1215,1237]" pageId="34" pageNumber="402">E. Morrill</collectorName>
and
<collectorName id="373E53519A178521FC10FB5BFBB1FB7E" box="[1014,1120,1215,1237]" pageId="34" pageNumber="402">D. Faber</collectorName>
, general collecting in bog forest dominated by
<taxonomicName id="5DCB4D049A178521FBDFFB3AFB72FB58" box="[1081,1187,1246,1267]" class="Magnoliopsida" family="Myrtaceae" genus="Tepualia" kingdom="Plantae" order="Myrtales" pageId="34" pageNumber="402" phylum="Tracheophyta" rank="genus">
<emphasis id="A8BFEA959A178521FBDFFB3AFB72FB58" box="[1081,1187,1246,1267]" italics="true" pageId="34" pageNumber="402">Tepualia</emphasis>
</taxonomicName>
deposited in MHNS (
<specimenCode id="CA6D9EFC9A178521FCA4FB18FBC1FAB9" box="[834,1040,1276,1298]" collectionCode="CASENT" pageId="34" pageNumber="402">CASENT9055645</specimenCode>
).
</materialsCitation>
<materialsCitation id="2AA33CDA9A178521FBC6FB18FBA1FAFB" ID-GBIF-Occurrence="4126400350" collectingDate="1985-12-15" collectingDateMax="2013-01-16" collectingDateMin="1985-12-15" collectionCode="CASENT" collectorName="L. E. Pena" country="Chile" county="Female" elevation="1" latitude="-42.617317" location="Chiloe Island" longLatPrecision="1" longitude="-74.10215" municipality="Chiloe Province" pageId="34" pageNumber="402" specimenCode="CASENT9055645, CASENT9044672" specimenCount="2" stateProvince="Los Lagos" typeStatus="paratype">
<collectorName id="373E53519A178521FBC6FB18FB88FAB9" box="[1056,1113,1276,1298]" pageId="34" pageNumber="402">Male</collectorName>
<typeStatus id="457088259A178521FBB9FB19FB14FAB9" box="[1119,1221,1277,1298]" pageId="34" pageNumber="402" type="paratype">paratype</typeStatus>
from
<location id="9F14605C9A178521FAE0FB18FA4EFAB9" LSID="urn:lsid:plazi:treatment:126287919A178525FEDBF966FB56FA57:9F14605C9A178521FAE0FB18FA4EFAB9" box="[1286,1439,1276,1298]" country="Chile" county="Female" latitude="-42.617317" longLatPrecision="1" longitude="-74.10215" municipality="Chiloe Province" name="Chiloe Island" pageId="34" pageNumber="402" stateProvince="Los Lagos">Chiloé Island</location>
, no specific locality,
<date id="EE7510479A178521FBFEFAFFFACEFA9A" box="[1048,1311,1307,1329]" pageId="34" pageNumber="402" value="1985-12-15" valueMax="1985-12-18" valueMin="1985-12-15">
<collectingDate id="FE31E9AF9A178521FBFEFAFFFACEFA9A" box="[1048,1311,1307,1329]" pageId="34" pageNumber="402" value="1985-12-15" valueMax="1985-12-18" valueMin="1985-12-15">December 1518, 1985</collectingDate>
</date>
,
<collectorName id="373E53519A178521FACAFAFFFA4FFA9B" box="[1324,1438,1307,1328]" pageId="34" pageNumber="402">L.E. Peña</collectorName>
, AMNH (
<specimenCode id="CA6D9EFC9A178521FC7DFADEFBB8FAFB" box="[923,1129,1338,1360]" collectionCode="CASENT" pageId="34" pageNumber="402">CASENT9044672</specimenCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="9A7436879A178521FCDDFA9CFC6AF9ED" blockId="34.[827,1443,1400,1606]" pageId="34" pageNumber="402">
<emphasis id="A8BFEA959A178521FCDDFA9CFC6AFA26" box="[827,955,1400,1421]" italics="true" pageId="34" pageNumber="402">Etymology:</emphasis>
Named after La Fiura, an ugly, villainous woman who lives in the forest and clothes herself in moss; according to Chilote mythology, she seduces men and then drives them insane. Chiloé Island is the
<typeStatus id="457088259A178521FC8FFA17FC4DF9A3" box="[873,924,1523,1544]" pageId="34" pageNumber="402">type</typeStatus>
locality, whose culture is rich in mythology. Sheet webs of
<taxonomicName id="5DCB4D049A178521FC1CF9F6FB88F98C" authorityName="Morrill &amp; Crews &amp; Esposito &amp; Ramírez &amp; Griswold" authorityYear="2023" box="[1018,1113,1554,1575]" class="Arachnida" family="Desidae" genus="Porteria" kingdom="Animalia" order="Araneae" pageId="34" pageNumber="402" phylum="Arthropoda" rank="species" species="fiura">
<emphasis id="A8BFEA959A178521FC1CF9F6FB88F98C" box="[1018,1113,1554,1575]" italics="true" pageId="34" pageNumber="402">P. fiura</emphasis>
</taxonomicName>
are often built on mossy substrates.
</paragraph>
<paragraph id="9A7436879A178520FCDDF994FE9CF8C0" blockId="34.[827,1444,1647,1884]" lastBlockId="35.[145,760,1694,1900]" lastPageId="35" lastPageNumber="403" pageId="34" pageNumber="402">
<emphasis id="A8BFEA959A178521FCDDF994FC65F92E" box="[827,948,1648,1669]" italics="true" pageId="34" pageNumber="402">Diagnosis:</emphasis>
Males can be distinguished from others in the
<taxonomicName id="5DCB4D049A178521FC75F96AFB5CF90F" authority="Grade" authorityName="Grade" box="[915,1165,1678,1700]" class="Arachnida" family="Desidae" genus="Porteria" kingdom="Animalia" order="Araneae" pageId="34" pageNumber="402" phylum="Arthropoda" rank="species" species="albopunctata">Albopunctata Grade</taxonomicName>
by the round, stubby RTA (
<figureCitation id="02F02A029A178521FC60F948FC24F96A" box="[902,1013,1708,1730]" captionStart="Figure 28" captionStartId="36.[161,240,1111,1133]" captionTargetBox="[182,1424,198,1078]" captionTargetId="figure-360@36.[179,1427,195,1081]" captionTargetPageId="36" captionText="Figure 28. Genitalia of P. fiura sp. nov. A-D, male from Chiloé Island CASENT9044672. A, left palp, retrolateral view. B, ventral view. C, prolateral view. D, dorsal view. E, F, female from Chiloé, CASENT9055645. E, ventral view. F, dorsal view, showing internal structures. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8007473" httpUri="https://zenodo.org/record/8007473/files/figure.png" pageId="34" pageNumber="402">Figs 28B</figureCitation>
,
<figureCitation id="02F02A029A178521FBEEF948FB8AF96A" box="[1032,1115,1708,1730]" captionStart="Figure 29" captionStartId="37.[146,227,986,1008]" captionTargetBox="[171,1396,206,952]" captionTargetId="figure-424@37.[161,1408,195,956]" captionTargetPageId="37" captionText="Figure 29. Left male palp of P. fiura sp. nov. AMNH paratype, CASENT9044672, Chiloé Island. A, prolateral view. B, ventral view. C, retrolateral view. D, tibia, dorsal view. Illustration by Rachel Diaz-Bastin." figureDoi="http://doi.org/10.5281/zenodo.8007475" httpUri="https://zenodo.org/record/8007475/files/figure.png" pageId="34" pageNumber="402">29B, C</figureCitation>
) and the small tooth-like DTA (
<figureCitation id="02F02A029A178521FC61F92FFC20F94B" box="[903,1009,1739,1761]" captionStart="Figure 29" captionStartId="37.[146,227,986,1008]" captionTargetBox="[171,1396,206,952]" captionTargetId="figure-424@37.[161,1408,195,956]" captionTargetPageId="37" captionText="Figure 29. Left male palp of P. fiura sp. nov. AMNH paratype, CASENT9044672, Chiloé Island. A, prolateral view. B, ventral view. C, retrolateral view. D, tibia, dorsal view. Illustration by Rachel Diaz-Bastin." figureDoi="http://doi.org/10.5281/zenodo.8007475" httpUri="https://zenodo.org/record/8007475/files/figure.png" pageId="34" pageNumber="402">Fig. 29D</figureCitation>
). The embolus base is wider (
<figureCitation id="02F02A029A178521FA90F92FFCB8F954" captionStart="Figure 29" captionStartId="37.[146,227,986,1008]" captionTargetBox="[171,1396,206,952]" captionTargetId="figure-424@37.[161,1408,195,956]" captionTargetPageId="37" captionText="Figure 29. Left male palp of P. fiura sp. nov. AMNH paratype, CASENT9044672, Chiloé Island. A, prolateral view. B, ventral view. C, retrolateral view. D, tibia, dorsal view. Illustration by Rachel Diaz-Bastin." figureDoi="http://doi.org/10.5281/zenodo.8007475" httpUri="https://zenodo.org/record/8007475/files/figure.png" pageId="34" pageNumber="402">Fig. 29B</figureCitation>
) than that of
<taxonomicName id="5DCB4D049A178521FBFFF90EFB00F954" box="[1049,1233,1770,1791]" class="Arachnida" family="Desidae" genus="Porteria" kingdom="Animalia" order="Araneae" pageId="34" pageNumber="402" phylum="Arthropoda" rank="species" species="albopunctata">
<emphasis id="A8BFEA959A178521FBFFF90EFB00F954" box="[1049,1233,1770,1791]" italics="true" pageId="34" pageNumber="402">P. albopunctata</emphasis>
</taxonomicName>
and the embolus tip more hook shaped (
<figureCitation id="02F02A029A178521FB9EF8ECFB39F8B6" box="[1144,1256,1800,1822]" captionStart="Figure 29" captionStartId="37.[146,227,986,1008]" captionTargetBox="[171,1396,206,952]" captionTargetId="figure-424@37.[161,1408,195,956]" captionTargetPageId="37" captionText="Figure 29. Left male palp of P. fiura sp. nov. AMNH paratype, CASENT9044672, Chiloé Island. A, prolateral view. B, ventral view. C, retrolateral view. D, tibia, dorsal view. Illustration by Rachel Diaz-Bastin." figureDoi="http://doi.org/10.5281/zenodo.8007475" httpUri="https://zenodo.org/record/8007475/files/figure.png" pageId="34" pageNumber="402">Fig. 29A</figureCitation>
); embolus tip much shorter than in
<taxonomicName id="5DCB4D049A178521FB84F8CCFAF6F897" authorityName="Morrill &amp; Crews &amp; Esposito &amp; Ramírez &amp; Griswold" authorityYear="2023" box="[1122,1319,1831,1853]" class="Arachnida" family="Desidae" genus="Porteria" kingdom="Animalia" order="Araneae" pageId="34" pageNumber="402" phylum="Arthropoda" rank="species" species="eddardstarki">
<emphasis id="A8BFEA959A178521FB84F8CCFAF6F897" box="[1122,1319,1831,1853]" italics="true" pageId="34" pageNumber="402">P. eddardstarki</emphasis>
</taxonomicName>
. Females most closely resemble
<taxonomicName id="5DCB4D049A178521FBB3F8A2FAC3F8F0" box="[1109,1298,1862,1883]" class="Arachnida" family="Desidae" genus="Porteria" kingdom="Animalia" order="Araneae" pageId="34" pageNumber="402" phylum="Arthropoda" rank="species" species="albopunctata">
<emphasis id="A8BFEA959A178521FBB3F8A2FAC3F8F0" box="[1109,1298,1862,1883]" italics="true" pageId="34" pageNumber="402">P. albopunctata</emphasis>
</taxonomicName>
but can be distinguished by the sclerotized median area being dilated anteriorly in
<taxonomicName id="5DCB4D049A168520FE6EF95AFE0CF979" authorityName="Morrill &amp; Crews &amp; Esposito &amp; Ramírez &amp; Griswold" authorityYear="2023" box="[392,477,1725,1747]" class="Arachnida" family="Desidae" genus="Porteria" kingdom="Animalia" order="Araneae" pageId="35" pageNumber="403" phylum="Arthropoda" rank="species" species="fiura">
<emphasis id="A8BFEA959A168520FE6EF95AFE0CF979" box="[392,477,1725,1747]" italics="true" pageId="35" pageNumber="403">P. fiura</emphasis>
</taxonomicName>
as well as the presence of a prominent scape (
<figureCitation id="02F02A029A168520FE7BF938FDD6F95A" box="[413,519,1756,1778]" captionStart="Figure 28" captionStartId="36.[161,240,1111,1133]" captionTargetBox="[182,1424,198,1078]" captionTargetId="figure-360@36.[179,1427,195,1081]" captionTargetPageId="36" captionText="Figure 28. Genitalia of P. fiura sp. nov. A-D, male from Chiloé Island CASENT9044672. A, left palp, retrolateral view. B, ventral view. C, prolateral view. D, dorsal view. E, F, female from Chiloé, CASENT9055645. E, ventral view. F, dorsal view, showing internal structures. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8007473" httpUri="https://zenodo.org/record/8007473/files/figure.png" pageId="35" pageNumber="403">Figs 28E</figureCitation>
,
<figureCitation id="02F02A029A168520FDF1F938FDB6F95A" box="[535,615,1756,1777]" captionStart="Figure 30" captionStartId="38.[163,246,1113,1135]" captionTargetBox="[185,1420,198,1080]" captionTargetId="figure-384@38.[182,1423,195,1083]" captionTargetPageId="38" captionText="Figure 30. Scanning electron micrograph of epigynum and vulva of P. fiura sp. nov. from Chiloé National Park, CASENT9055645. A, ventral view. B, dorsal view. C, close-up of scape and copulatory openings. D, head of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.8007477" httpUri="https://zenodo.org/record/8007477/files/figure.png" pageId="35" pageNumber="403">30A, B</figureCitation>
). Internally, the spermatheca stalks are situated much farther apart in
<taxonomicName id="5DCB4D049A168520FEE2F8FEFEB0F885" authorityName="Morrill &amp; Crews &amp; Esposito &amp; Ramírez &amp; Griswold" authorityYear="2023" box="[260,353,1817,1839]" class="Arachnida" family="Desidae" genus="Porteria" kingdom="Animalia" order="Araneae" pageId="35" pageNumber="403" phylum="Arthropoda" rank="species" species="fiura">
<emphasis id="A8BFEA959A168520FEE2F8FEFEB0F885" box="[260,353,1817,1839]" italics="true" pageId="35" pageNumber="403">P. fiura</emphasis>
</taxonomicName>
than
<taxonomicName id="5DCB4D049A168520FE55F8FEFDA3F885" box="[435,626,1817,1839]" class="Arachnida" family="Desidae" genus="Porteria" kingdom="Animalia" order="Araneae" pageId="35" pageNumber="403" phylum="Arthropoda" rank="species" species="albopunctata">
<emphasis id="A8BFEA959A168520FE55F8FEFDA3F885" box="[435,626,1817,1839]" italics="true" pageId="35" pageNumber="403">P. albopunctata</emphasis>
</taxonomicName>
; Bennetts gland pores large and conspicuous in dorsal view (
<figureCitation id="02F02A029A168520FF7FF8B2FF2FF8C0" box="[153,254,1878,1900]" captionStart="Figure 28" captionStartId="36.[161,240,1111,1133]" captionTargetBox="[182,1424,198,1078]" captionTargetId="figure-360@36.[179,1427,195,1081]" captionTargetPageId="36" captionText="Figure 28. Genitalia of P. fiura sp. nov. A-D, male from Chiloé Island CASENT9044672. A, left palp, retrolateral view. B, ventral view. C, prolateral view. D, dorsal view. E, F, female from Chiloé, CASENT9055645. E, ventral view. F, dorsal view, showing internal structures. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8007473" httpUri="https://zenodo.org/record/8007473/files/figure.png" pageId="35" pageNumber="403">Figs 28F</figureCitation>
,
<figureCitation id="02F02A029A168520FEE9F8B2FEEDF8C0" box="[271,316,1878,1899]" captionStart="Figure 30" captionStartId="38.[163,246,1113,1135]" captionTargetBox="[185,1420,198,1080]" captionTargetId="figure-384@38.[182,1423,195,1083]" captionTargetPageId="38" captionText="Figure 30. Scanning electron micrograph of epigynum and vulva of P. fiura sp. nov. from Chiloé National Park, CASENT9055645. A, ventral view. B, dorsal view. C, close-up of scape and copulatory openings. D, head of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.8007477" httpUri="https://zenodo.org/record/8007477/files/figure.png" pageId="35" pageNumber="403">30B</figureCitation>
).
</paragraph>
<caption id="CEB4660F9A168520FF77F9F3FE40F9C3" ID-DOI="http://doi.org/10.5281/zenodo.8007471" ID-Zenodo-Dep="8007471" httpUri="https://zenodo.org/record/8007471/files/figure.png" pageId="35" pageNumber="403" startId="35.[145,224,1559,1581]" targetBox="[164,1406,198,1526]" targetPageId="35" targetType="figure">
<paragraph id="9A7436879A168520FF77F9F3FE40F9C3" blockId="35.[145,1425,1559,1640]" pageId="35" pageNumber="403">
<emphasis id="A8BFEA959A168520FF77F9F3FED9F987" bold="true" box="[145,264,1559,1581]" pageId="35" pageNumber="403">Figure 27.</emphasis>
Habitus of
<taxonomicName id="5DCB4D049A168520FE6FF9FCFE04F986" authority="Morrill &amp; Crews &amp; Esposito &amp; Ramírez &amp; Griswold, 2023" authorityName="Morrill &amp; Crews &amp; Esposito &amp; Ramírez &amp; Griswold" authorityYear="2023" box="[393,469,1559,1581]" class="Arachnida" family="Desidae" genus="Porteria" kingdom="Animalia" order="Araneae" pageId="35" pageNumber="403" phylum="Arthropoda" rank="species" species="fiura" status="sp. nov.">
<emphasis id="A8BFEA959A168520FE6FF9FCFE04F986" box="[393,469,1559,1581]" italics="true" pageId="35" pageNumber="403">P. fiura</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B38C57EE9A168520FE3AF9F3FDFBF986" box="[476,554,1559,1581]" pageId="35" pageNumber="403" rank="species">sp. nov.</taxonomicNameLabel>
A-C, male from Chiloé Island CASENT9044672. A, dorsal view. B, ventral view. C, lateral view. D-F, female from Chiloé National Park CASENT9056299 and 9055645. D, dorsal view. E, ventral view. F, lateral view. Scale bars = 1 mm.
</paragraph>
</caption>
<paragraph id="9A7436879A168527FCCFF97BFB3DFA02" blockId="35.[809,1426,1694,1900]" lastBlockId="36.[827,1442,1243,1572]" lastPageId="36" lastPageNumber="404" pageId="35" pageNumber="403">
<emphasis id="A8BFEA959A168520FCCFF97BFB9DF91F" box="[809,1100,1694,1716]" italics="true" pageId="35" pageNumber="403">Description: Male</emphasis>
: based on
<typeStatus id="457088259A168520FAEBF97BFA5DF91F" box="[1293,1420,1695,1716]" pageId="35" pageNumber="403" type="paratype">paratype</typeStatus>
CASENT9044672. Markings as in
<figureCitation id="02F02A029A168520FB06F959FA88F979" box="[1248,1369,1725,1747]" captionStart="Figure 27" captionStartId="35.[145,224,1559,1581]" captionTargetBox="[164,1406,198,1526]" captionTargetId="figure-165@35.[161,1409,195,1529]" captionTargetPageId="35" captionText="Figure 27. Habitus of P. fiura sp. nov. A-C, male from Chiloé Island CASENT9044672. A, dorsal view. B, ventral view. C, lateral view. D-F, female from Chiloé National Park CASENT9056299 and 9055645. D, dorsal view. E, ventral view. F, lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8007471" httpUri="https://zenodo.org/record/8007471/files/figure.png" pageId="35" pageNumber="403">Figure 27</figureCitation>
A-C, dorsum of abdomen with yellow anterolateral lines followed by two posterior spots and two pairs of median spots; sternum with pale centre surrounded by dark grey border. Total length 5.63. Carapace length 1.37 times width. Clypeus height 2.29 times AME diameter. ALE diameter 1.71 times AME diameter. Chelicera length 6.28 times clypeus height. Sternum as long as wide. Femur I 1.37 times carapace length. Leg formula 4123. Cymbium length 3.25 times length of bulb. Cheliceral promargin with four teeth and one denticle. Leg spination as follows: palp: femur d1-1-3, patella d1-1, tarsus p0-0-1, v0-0-1(r); leg I: d1-1-1(p)-2, patella d1-1, tibia d1(r)-0-0, v0-0-2, metatarsus v0-2-2; leg II: femur d1-1-1(p)-2, patella d1-1, tibia d1(r)-0-0, p0-0-1-0, v0-1(r)-2, metatarsus d0-0-1, p1-0-1, r0-0-1, v0-2-1; leg III: femur d1-1(r)-2, patella d1-1, tibia d1(r)- 0-0, p0-1-1-0, r0-0-1-0, v0-1(r)-2, metatarsus d1(p)-0-1, p0-1-0-1, r0-1-0-1, v0-2-2, tarsus r0-1-1(v), v0-1; leg IV: femur d1-1(p)-2, patella d1-1, tibia d1(r)-0-0, p0-1-1- 0, v0-0-2, metatarsus d2-0-1, p0-1-1, r0-1-1-1, v0-0-2, tarsus r0-1. Palp as in
<figureCitation id="02F02A029A118527FE53F943FDE5F917" box="[437,564,1703,1725]" captionStart="Figure 28" captionStartId="36.[161,240,1111,1133]" captionTargetBox="[182,1424,198,1078]" captionTargetId="figure-360@36.[179,1427,195,1081]" captionTargetPageId="36" captionText="Figure 28. Genitalia of P. fiura sp. nov. A-D, male from Chiloé Island CASENT9044672. A, left palp, retrolateral view. B, ventral view. C, prolateral view. D, dorsal view. E, F, female from Chiloé, CASENT9055645. E, ventral view. F, dorsal view, showing internal structures. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8007473" httpUri="https://zenodo.org/record/8007473/files/figure.png" pageId="36" pageNumber="404">Figures 28</figureCitation>
and
<figureCitation id="02F02A029A118527FD94F943FD5FF917" box="[626,654,1703,1724]" captionStart="Figure 29" captionStartId="37.[146,227,986,1008]" captionTargetBox="[171,1396,206,952]" captionTargetId="figure-424@37.[161,1408,195,956]" captionTargetPageId="37" captionText="Figure 29. Left male palp of P. fiura sp. nov. AMNH paratype, CASENT9044672, Chiloé Island. A, prolateral view. B, ventral view. C, retrolateral view. D, tibia, dorsal view. Illustration by Rachel Diaz-Bastin." figureDoi="http://doi.org/10.5281/zenodo.8007475" httpUri="https://zenodo.org/record/8007475/files/figure.png" pageId="36" pageNumber="404">29</figureCitation>
, with four tibial apophyses: VTA triangular, flattened plate in ventral view, outline finger like in retrolateral view; RTA short and apex round, curved, directed ventrally; LRTA projected straight out of tibia, parallel to frontal plane. DTA small and tooth like; DTA and LRTA appear connected at fleshy base (
<figureCitation id="02F02A029A118527FB25FB3FFAF7FB5B" box="[1219,1318,1243,1265]" captionStart="Figure 28" captionStartId="36.[161,240,1111,1133]" captionTargetBox="[182,1424,198,1078]" captionTargetId="figure-360@36.[179,1427,195,1081]" captionTargetPageId="36" captionText="Figure 28. Genitalia of P. fiura sp. nov. A-D, male from Chiloé Island CASENT9044672. A, left palp, retrolateral view. B, ventral view. C, prolateral view. D, dorsal view. E, F, female from Chiloé, CASENT9055645. E, ventral view. F, dorsal view, showing internal structures. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8007473" httpUri="https://zenodo.org/record/8007473/files/figure.png" pageId="36" pageNumber="404">Fig. 28D</figureCitation>
), like that of
<taxonomicName id="5DCB4D049A118527FCBEFB1EFC23FAA4" box="[856,1010,1274,1295]" class="Arachnida" family="Desidae" genus="Porteria" kingdom="Animalia" order="Araneae" pageId="36" pageNumber="404" phylum="Arthropoda" rank="species" species="albopunctata">
<emphasis id="A8BFEA959A118527FCBEFB1EFC23FAA4" box="[856,1010,1274,1295]" italics="true" pageId="36" pageNumber="404">albopunctata</emphasis>
</taxonomicName>
. Tegulum without median concavity, outlined by dark sperm duct. Conductor originating on the midline of bulb, short and simple projection. Embolus base elongated, nearly the length of the bulb; embolus tip short, spiniform, curved like a hook to meet conductor. Paracymbium absent.
</paragraph>
<caption id="CEB4660F9A118527FF47FBB3FD7AFB03" ID-DOI="http://doi.org/10.5281/zenodo.8007473" ID-Zenodo-Dep="8007473" httpUri="https://zenodo.org/record/8007473/files/figure.png" pageId="36" pageNumber="404" startId="36.[161,240,1111,1133]" targetBox="[182,1424,198,1078]" targetPageId="36" targetType="figure">
<paragraph id="9A7436879A118527FF47FBB3FD7AFB03" blockId="36.[161,1444,1111,1192]" pageId="36" pageNumber="404">
<emphasis id="A8BFEA959A118527FF47FBB3FEC6FBC7" bold="true" box="[161,279,1111,1133]" pageId="36" pageNumber="404">Figure 28.</emphasis>
Genitalia of
<taxonomicName id="5DCB4D049A118527FE4FFBBCFE24FBC6" authority="Morrill &amp; Crews &amp; Esposito &amp; Ramírez &amp; Griswold, 2023" authorityName="Morrill &amp; Crews &amp; Esposito &amp; Ramírez &amp; Griswold" authorityYear="2023" box="[425,501,1111,1133]" class="Arachnida" family="Desidae" genus="Porteria" kingdom="Animalia" order="Araneae" pageId="36" pageNumber="404" phylum="Arthropoda" rank="species" species="fiura" status="sp. nov.">
<emphasis id="A8BFEA959A118527FE4FFBBCFE24FBC6" box="[425,501,1111,1133]" italics="true" pageId="36" pageNumber="404">P. fiura</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B38C57EE9A118527FE1AFBB3FD98FBC6" box="[508,585,1111,1133]" pageId="36" pageNumber="404" rank="species">sp. nov.</taxonomicNameLabel>
A-D, male from Chiloé Island CASENT9044672. A, left palp, retrolateral view. B, ventral view. C, prolateral view. D, dorsal view. E, F, female from Chiloé, CASENT9055645. E, ventral view. F, dorsal view, showing internal structures. Scale bars = 0.2 mm.
</paragraph>
</caption>
<paragraph id="9A7436879A118527FCB5FA56FAD9F98F" blockId="36.[827,1442,1243,1572]" pageId="36" pageNumber="404">Leg measurements (right): leg I 13.50 (3.75, 4.50, 3.25, 2.00); leg II 11.15 (3.15, 3.50, 2.85, 1.65); leg III 10.55 (2.85, 3.25, 3.00, 1.45); leg IV 13.80 (3.75, 4.30, 4.20, 1.55); palp 4.85 (1.80, 1.10, , 1.95).</paragraph>
<paragraph id="9A7436879A118526FCDDF9A9FAB3FB4E" blockId="36.[827,1443,1612,1880]" lastBlockId="37.[809,1425,1109,1376]" lastPageId="37" lastPageNumber="405" pageId="36" pageNumber="404">
<emphasis id="A8BFEA959A118527FCDDF9A9FBECF9CA" box="[827,1085,1612,1634]" italics="true" pageId="36" pageNumber="404">Description: Female</emphasis>
: based on CASENT9055645, from Chiloé National Park. Markings as in
<figureCitation id="02F02A029A118527FAB5F98FFC84F934" captionStart="Figure 27" captionStartId="35.[145,224,1559,1581]" captionTargetBox="[164,1406,198,1526]" captionTargetId="figure-165@35.[161,1409,195,1529]" captionTargetPageId="35" captionText="Figure 27. Habitus of P. fiura sp. nov. A-C, male from Chiloé Island CASENT9044672. A, dorsal view. B, ventral view. C, lateral view. D-F, female from Chiloé National Park CASENT9056299 and 9055645. D, dorsal view. E, ventral view. F, lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8007471" httpUri="https://zenodo.org/record/8007471/files/figure.png" pageId="36" pageNumber="404">Figure 27</figureCitation>
D-F. Total length 5.80. Carapace length 1.34 times width. Clypeus height 1.86 times AME diameter. ALE diameter 1.71 times AME diameter. Chelicera length 6.15 times clypeus height. Sternum as long as wide. Femur I 1.24 times carapace length. Leg formula 4123. Cheliceral promargin with five teeth and one denticle. Leg spination as follows: palp: d1-1-3, patella d1-1, tibia d2-0-1(r), tarsus d2-0, p0-1-1-0, r0-1-1-0, v2-1(r)-3; leg I: femur d1-2-1(p)-2, patella d1-1, tibia d1(r)-0-0, v0-2- 2, metatarsus p0-0-1, r0-0-1, v2-2-1; leg II: femur d1-1- 1(p)-2, patella d1-1, tibia d1(r)-0-0, p0-1-1-0, v0-2-2, metatarsus p0-1-0-1, r0-1-0-1, v1(r)-2-2; leg III: femur d2-1-1(p)-2, patella d1-1, tibia d1(r)-0-0, p0-1-1-0, r0-0- 1-0, v0-1(r)-2, metatarsus d1(p)-0-2, p0-1-0-1, r0-1-0- 1, v2-2-1, tarsus r0-1, v0-2; leg IV: femur d1-0-1(p)-2, patella d1-1, tibia d1(r)-0-0, p0-1-1-0, r0-1-1-0, v0-2-2, metatarsus d2-1(r)-2, p0-1-0-1, r0-1-0-1, v2-2-1, tarsus r0-2, v0-2. Epigyne as in
<figureCitation id="02F02A029A108526FE5CFA6CFDB9FA36" box="[442,616,1416,1438]" captionStart="Figure 28" captionStartId="36.[161,240,1111,1133]" captionTargetBox="[182,1424,198,1078]" captionTargetId="figure-360@36.[179,1427,195,1081]" captionTargetPageId="36" captionText="Figure 28. Genitalia of P. fiura sp. nov. A-D, male from Chiloé Island CASENT9044672. A, left palp, retrolateral view. B, ventral view. C, prolateral view. D, dorsal view. E, F, female from Chiloé, CASENT9055645. E, ventral view. F, dorsal view, showing internal structures. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8007473" httpUri="https://zenodo.org/record/8007473/files/figure.png" pageId="37" pageNumber="405">Figures 28E, F</figureCitation>
and
<figureCitation id="02F02A029A108526FD43FA6CFD22FA36" box="[677,755,1416,1437]" captionStart="Figure 30" captionStartId="38.[163,246,1113,1135]" captionTargetBox="[185,1420,198,1080]" captionTargetId="figure-384@38.[182,1423,195,1083]" captionTargetPageId="38" captionText="Figure 30. Scanning electron micrograph of epigynum and vulva of P. fiura sp. nov. from Chiloé National Park, CASENT9055645. A, ventral view. B, dorsal view. C, close-up of scape and copulatory openings. D, head of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.8007477" httpUri="https://zenodo.org/record/8007477/files/figure.png" pageId="37" pageNumber="405">30A, C</figureCitation>
, lightly sclerotized except for narrow median area which is dilated anteriorly where the scape originates; scape small and finger like, pointed posteriad; copulatory openings inconspicuous, located just anterior to scape, divided by fleshy septum; internal structures visible through cuticle, most prominently two large circular patches on either side of sclerotized area. Vulva (
<figureCitation id="02F02A029A108526FD2EF9BAFF6DF939" captionStart="Figure 28" captionStartId="36.[161,240,1111,1133]" captionTargetBox="[182,1424,198,1078]" captionTargetId="figure-360@36.[179,1427,195,1081]" captionTargetPageId="36" captionText="Figure 28. Genitalia of P. fiura sp. nov. A-D, male from Chiloé Island CASENT9044672. A, left palp, retrolateral view. B, ventral view. C, prolateral view. D, dorsal view. E, F, female from Chiloé, CASENT9055645. E, ventral view. F, dorsal view, showing internal structures. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8007473" httpUri="https://zenodo.org/record/8007473/files/figure.png" pageId="37" pageNumber="405">Figs 28F</figureCitation>
,
<figureCitation id="02F02A029A108526FF2AF999FECEF939" box="[204,287,1661,1682]" captionStart="Figure 30" captionStartId="38.[163,246,1113,1135]" captionTargetBox="[185,1420,198,1080]" captionTargetId="figure-384@38.[182,1423,195,1083]" captionTargetPageId="38" captionText="Figure 30. Scanning electron micrograph of epigynum and vulva of P. fiura sp. nov. from Chiloé National Park, CASENT9055645. A, ventral view. B, dorsal view. C, close-up of scape and copulatory openings. D, head of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.8007477" httpUri="https://zenodo.org/record/8007477/files/figure.png" pageId="37" pageNumber="405">30B, D</figureCitation>
) with anterior, short copulatory ducts joining the spermathecal stalk just posterior to head of spermatheca. Head situated at the anterior apex of stalk, pores present. Stalks banana shaped, bowed out laterally, pores present. Base 1 large circular structure with large Bennetts gland pore in its centre when viewed dorsally; fertilization duct attached to Base 1 on median posterior corner. Base 2 joining the anterior margin of Base 1, somewhat crescent shaped; copulatory duct resting on anterior surface of Base 2. In total, the two receptacles look like one large, ovoid receptacle; scanning electron micrograph shows the division of the two bases in more detail (
<figureCitation id="02F02A029A108526FB12FB34FA82FB4E" box="[1268,1363,1232,1254]" captionStart="Figure 30" captionStartId="38.[163,246,1113,1135]" captionTargetBox="[185,1420,198,1080]" captionTargetId="figure-384@38.[182,1423,195,1083]" captionTargetPageId="38" captionText="Figure 30. Scanning electron micrograph of epigynum and vulva of P. fiura sp. nov. from Chiloé National Park, CASENT9055645. A, ventral view. B, dorsal view. C, close-up of scape and copulatory openings. D, head of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.8007477" httpUri="https://zenodo.org/record/8007477/files/figure.png" pageId="37" pageNumber="405">Fig. 30B</figureCitation>
).
</paragraph>
<caption id="CEB4660F9A108526FF74FC3EFBE8FBA6" ID-DOI="http://doi.org/10.5281/zenodo.8007475" ID-Zenodo-Dep="8007475" httpUri="https://zenodo.org/record/8007475/files/figure.png" pageId="37" pageNumber="405" startId="37.[146,227,986,1008]" targetBox="[171,1396,206,952]" targetPageId="37" targetType="figure">
<paragraph id="9A7436879A108526FF74FC3EFBE8FBA6" blockId="37.[145,1425,986,1037]" pageId="37" pageNumber="405">
<emphasis id="A8BFEA959A108526FF74FC3EFEDDFC44" bold="true" box="[146,268,986,1008]" pageId="37" pageNumber="405">Figure 29.</emphasis>
Left male palp of
<taxonomicName id="5DCB4D049A108526FE3EFC3FFDF7FC5B" authority="Morrill &amp; Crews &amp; Esposito &amp; Ramírez &amp; Griswold, 2023" authorityName="Morrill &amp; Crews &amp; Esposito &amp; Ramírez &amp; Griswold" authorityYear="2023" box="[472,550,986,1008]" class="Arachnida" family="Desidae" genus="Porteria" kingdom="Animalia" order="Araneae" pageId="37" pageNumber="405" phylum="Arthropoda" rank="species" species="fiura" status="sp. nov.">
<emphasis id="A8BFEA959A108526FE3EFC3FFDF7FC5B" box="[472,550,986,1008]" italics="true" pageId="37" pageNumber="405">P. fiura</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B38C57EE9A108526FDC8FC3EFDACFC5B" box="[558,637,986,1008]" pageId="37" pageNumber="405" rank="species">sp. nov.</taxonomicNameLabel>
AMNH paratype, CASENT9044672, Chiloé Island. A, prolateral view. B, ventral view. C, retrolateral view. D, tibia, dorsal view. Illustration by Rachel Diaz-Bastin.
</paragraph>
</caption>
<paragraph id="9A7436879A108526FCA7FB0BFB68FAF4" blockId="37.[809,1425,1109,1376]" pageId="37" pageNumber="405">Leg measurements: leg I 12.17 (3.32, 4.20, 2.85, 1.80); leg II 10.25 (2.95, 3.40, 2.45, 1.45); leg III 9.70 (2.80, 3.00, 2.55, 1.35); leg IV 12.65 (3.45, 4.00, 3.55, 1.65); palp 3.92 (1.32, 1.32, , 1.28).</paragraph>
<paragraph id="9A7436879A108526FCCFFA6EFAAEF9FC" blockId="37.[809,1425,1417,1623]" pageId="37" pageNumber="405">
<emphasis id="A8BFEA959A108526FCCFFA6EFC4EFA34" box="[809,927,1418,1439]" italics="true" pageId="37" pageNumber="405">Variation:</emphasis>
(
<emphasis id="A8BFEA959A108526FC5BFA6EFC00FA34" box="[957,977,1418,1439]" italics="true" pageId="37" pageNumber="405">N</emphasis>
= 3). Total length 3.95.76. Carapace length 1.271.37 times width. Clypeus height 1.672.2 times AME diameter. ALE diameter 1.672.2 times AME diameter. Chelicera length 56.18 times clypeus height. Sternum length 11.04 times width. Femur I 1.081.26 times carapace length. Cheliceral promargin with four teeth and one denticle left, five teeth right.
</paragraph>
<paragraph id="9A7436879A108526FCCFF99BFCB4F979" blockId="37.[809,1426,1663,1747]" pageId="37" pageNumber="405">
<emphasis id="A8BFEA959A108526FCCFF99BFC6DF93F" box="[809,956,1663,1684]" italics="true" pageId="37" pageNumber="405">Distribution:</emphasis>
Only known from Chiloé Island and on the mainland in Palena,
<quantity id="5D339B629A108526FBA0F97AFB5FF91F" box="[1094,1166,1694,1716]" metricMagnitude="4" metricUnit="m" metricValue="7.0" pageId="37" pageNumber="405" unit="km" value="70.0">70 km</quantity>
south of Chaitén (
<figureCitation id="02F02A029A108526FA83F97AFC87F978" captionStart="Figure 76" captionStartId="89.[145,224,1353,1375]" captionTargetBox="[147,1425,197,1322]" captionTargetId="figure-260@89.[145,1425,195,1323]" captionTargetPageId="89" captionText="Figure 76. A, records of Porteria species in Chile. B, distribution of P. eddardstarki sp. nov.—the northernmost record is Fray Jorge National Park. C, records of species with restricted distributions, and of P. albopunctata sp. nov. in southern Chile. D, distribution of P. ajimayo sp. nov.—the southernmost records are in Reserva Costera Valdiviana and Alerce Costero National Park. E, distribution of P. bunnyanasp. nov.—the southernmost record is in Chonchi, Chiloé Island." figureDoi="http://doi.org/10.5281/zenodo.8007576" httpUri="https://zenodo.org/record/8007576/files/figure.png" pageId="37" pageNumber="405">Fig. 76C</figureCitation>
).
</paragraph>
<paragraph id="9A7436879A108526FCCFF91FFA40F8C6" blockId="37.[809,1425,1787,1901]" pageId="37" pageNumber="405">
<materialsCitation id="2AA33CDA9A108526FCCFF91FFA40F8C6" collectingDate="1986-01-16" collectingDateMax="2013-01-16" collectingDateMin="1985-12-15" collectionCode="CAS" collectorName="C. Griswold &amp; E. Morrill &amp; D. Faber &amp; Male &amp; L. E. Pena" country="Chile" county="Female" elevation="500" latitude="-42.617317" location="Palena Province" longLatPrecision="1" longitude="-74.10215" municipality="Chiloe Province" pageId="37" pageNumber="405" specimenCode="CASENT9055645" specimenCount="1" specimenCount-female="1" stateProvince="Los Lagos">
<emphasis id="A8BFEA959A108526FCCFF91FFBB6F8BB" box="[809,1127,1787,1808]" italics="true" pageId="37" pageNumber="405">Other material examined:</emphasis>
X
<collectingRegion id="580FF8659A108526FB77F91FFA5DF8BA" box="[1169,1420,1787,1809]" country="Chile" name="Los Lagos" pageId="37" pageNumber="405">Región de los Lagos</collectingRegion>
:
<collectingMunicipality id="7A10ACFD9A108526FCCFF8FEFC22F89B" box="[809,1011,1818,1840]" pageId="37" pageNumber="405">Chiloé Province</collectingMunicipality>
, same locality as holotype,
<specimenCount id="8CCDFD0E9A108526FABDF8FEFC53F8E4" pageId="37" pageNumber="405" type="female">nine females</specimenCount>
,
<collectionCode id="FCDAAE429A108526FC6BF8DDFC11F8E5" box="[909,960,1849,1870]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="37" pageNumber="405" type="Museum">CAS</collectionCode>
(
<specimenCode id="CA6D9EFC9A108526FC29F8DDFB4DF8E4" box="[975,1180,1849,1871]" collectionCode="CASENT" pageId="37" pageNumber="405">CASENT9055645</specimenCode>
);
<location id="9F14605C9A108526FB49F8DDFABBF8E4" LSID="urn:lsid:plazi:treatment:126287919A178525FEDBF966FB56FA57:9F14605C9A108526FB49F8DDFABBF8E4" box="[1199,1386,1849,1871]" municipality="Chiloe Province" name="Palena Province" pageId="37" pageNumber="405" stateProvince="Los Lagos">Palena Province</location>
,
<locationDeviation id="8BC62FA59A108526FA93F8DDFC3DF8C6" location="Chaiten" pageId="37" pageNumber="405">
<quantity id="5D339B629A108526FA93F8DDFC9FF8C6" metricMagnitude="4" metricUnit="m" metricValue="7.0" pageId="37" pageNumber="405" unit="km" value="70.0">70 km</quantity>
S of Chaitén
</locationDeviation>
, elev.
<quantity id="5D339B629A108526FBD5F8B3FBAFF8C6" box="[1075,1150,1879,1901]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="37" pageNumber="405" unit="m" value="500.0">
<elevation id="11E6D1B49A108526FBD5F8B3FBAFF8C6" box="[1075,1150,1879,1901]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="37" pageNumber="405" unit="m" value="500.0">500 m</elevation>
</quantity>
,
<date id="EE7510479A108526FB6CF8BCFA88F8C6" box="[1162,1369,1879,1901]" pageId="37" pageNumber="405" value="1986-01-16">
<collectingDate id="FE31E9AF9A108526FB6CF8BCFA88F8C6" box="[1162,1369,1879,1901]" pageId="37" pageNumber="405" value="1986-01-16">January 16, 1986</collectingDate>
</date>
, N.I.
</materialsCitation>
</paragraph>
<caption id="CEB4660F9A138525FF45FBBDFA51FB26" ID-DOI="http://doi.org/10.5281/zenodo.8007477" ID-Zenodo-Dep="8007477" httpUri="https://zenodo.org/record/8007477/files/figure.png" pageId="38" pageNumber="406" startId="38.[163,246,1113,1135]" targetBox="[185,1420,198,1080]" targetPageId="38" targetType="figure">
<paragraph id="9A7436879A138525FF45FBBDFA51FB26" blockId="38.[163,1441,1113,1165]" pageId="38" pageNumber="406">
<emphasis id="A8BFEA959A138525FF45FBBDFEF3FBC5" bold="true" box="[163,290,1113,1135]" pageId="38" pageNumber="406">Figure 30.</emphasis>
Scanning electron micrograph of epigynum and vulva of
<taxonomicName id="5DCB4D049A138525FC50FBBEFBD8FBC4" authority="Morrill &amp; Crews &amp; Esposito &amp; Ramírez &amp; Griswold, 2023" authorityName="Morrill &amp; Crews &amp; Esposito &amp; Ramírez &amp; Griswold" authorityYear="2023" box="[950,1033,1113,1135]" class="Arachnida" family="Desidae" genus="Porteria" kingdom="Animalia" order="Araneae" pageId="38" pageNumber="406" phylum="Arthropoda" rank="species" species="fiura" status="sp. nov.">
<emphasis id="A8BFEA959A138525FC50FBBEFBD8FBC4" box="[950,1033,1113,1135]" italics="true" pageId="38" pageNumber="406">P. fiura</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B38C57EE9A138525FBF5FBBDFBB7FBC4" box="[1043,1126,1113,1135]" pageId="38" pageNumber="406" rank="species">sp. nov.</taxonomicNameLabel>
from Chiloé National Park, CASENT9055645. A, ventral view. B, dorsal view. C, close-up of scape and copulatory openings. D, head of spermatheca.
</paragraph>
</caption>
<paragraph id="9A7436879A138525FF45FB36FDB2FAAD" blockId="38.[163,778,1234,1286]" pageId="38" pageNumber="406">
<collectorName id="373E53519A138525FF45FB36FEEAFB4C" box="[163,315,1234,1256]" pageId="38" pageNumber="406">Platnick, P.A</collectorName>
.
<collectorName id="373E53519A138525FEA1FB36FE0CFB4C" box="[327,477,1234,1256]" pageId="38" pageNumber="406">Goloboff, R.T</collectorName>
. Schuh, wet streambank,
<specimenCount id="8CCDFD0E9A138525FF45FB15FECEFAAD" box="[163,287,1264,1286]" pageId="38" pageNumber="406" type="female">one female</specimenCount>
, AMNH (
<specimenCode id="CA6D9EFC9A138525FE6FFB14FD86FAAD" box="[393,599,1264,1286]" collectionCode="CASENT" pageId="38" pageNumber="406">CASENT9044706</specimenCode>
).
</paragraph>
<paragraph id="9A7436879A138525FF45FAD4FB56FA57" blockId="38.[163,780,1327,1901]" lastBlockId="38.[827,1443,1234,1532]" pageId="38" pageNumber="406">
<emphasis id="A8BFEA959A138525FF45FAD4FF3BFAEE" box="[163,234,1328,1349]" italics="true" pageId="38" pageNumber="406">Notes:</emphasis>
Webs were found near ground level on fallen logs and low branches with retreats leading into the hollows of the trees, which were wet.
<taxonomicName id="5DCB4D049A138525FD4CFA89FCDBFA29" authorityName="Simon" authorityYear="1904" box="[682,778,1389,1410]" class="Arachnida" family="Desidae" genus="Porteria" kingdom="Animalia" order="Araneae" pageId="38" pageNumber="406" phylum="Arthropoda" rank="genus">
<emphasis id="A8BFEA959A138525FD4CFA89FCDBFA29" box="[682,778,1389,1410]" italics="true" pageId="38" pageNumber="406">Porteria</emphasis>
</taxonomicName>
was the predominant web along with austrochilids, orb weavers and linyphiids. Only females were collected on the 2013 expedition and were found in forest near the southern shore of Lago Huillinco; they were not found on the northern shore. In the southern sector of Chiloé National Park, they were found in high density along the Tepual Trail, and it is likely that males would have been found if we had been able to night collect. Male and female were tentatively matched by locality information. The only other species that has been found on Chiloé Island is
<taxonomicName id="5DCB4D049A138525FE8BF939FDD3F95A" authorityName="Morrill &amp; Crews &amp; Esposito &amp; Ramírez &amp; Griswold" authorityYear="2023" box="[365,514,1756,1778]" class="Arachnida" family="Desidae" genus="Porteria" kingdom="Animalia" order="Araneae" pageId="38" pageNumber="406" phylum="Arthropoda" rank="species" species="bunnyana">
<emphasis id="A8BFEA959A138525FE8BF939FDD3F95A" box="[365,514,1756,1778]" italics="true" pageId="38" pageNumber="406">P. bunnyana</emphasis>
</taxonomicName>
. Because this species has been collected with other males outside of the island, and never together with females of
<taxonomicName id="5DCB4D049A138525FD4BF8FEFCD5F884" authorityName="Morrill &amp; Crews &amp; Esposito &amp; Ramírez &amp; Griswold" authorityYear="2023" box="[685,772,1818,1839]" class="Arachnida" family="Desidae" genus="Porteria" kingdom="Animalia" order="Araneae" pageId="38" pageNumber="406" phylum="Arthropoda" rank="species" species="fiura">
<emphasis id="A8BFEA959A138525FD4BF8FEFCD5F884" box="[685,772,1818,1839]" italics="true" pageId="38" pageNumber="406">P. fiura</emphasis>
</taxonomicName>
, the unique male from Chiloé is hypothesized to be the same species as the female
<typeStatus id="457088259A138525FDCFF8B3FD40F8C6" box="[553,657,1879,1901]" pageId="38" pageNumber="406" type="holotype">holotype</typeStatus>
. It is also noteworthy that
<specimenCount id="8CCDFD0E9A138525FBE7FB36FB51FB43" box="[1025,1152,1234,1256]" pageId="38" pageNumber="406" type="female">one female</specimenCount>
specimen of
<taxonomicName id="5DCB4D049A138525FAFFFB36FABFFB4C" authorityName="Morrill &amp; Crews &amp; Esposito &amp; Ramírez &amp; Griswold" authorityYear="2023" box="[1305,1390,1234,1255]" class="Arachnida" family="Desidae" genus="Porteria" kingdom="Animalia" order="Araneae" pageId="38" pageNumber="406" phylum="Arthropoda" rank="species" species="fiura">
<emphasis id="A8BFEA959A138525FAFFFB36FABFFB4C" box="[1305,1390,1234,1255]" italics="true" pageId="38" pageNumber="406">P. fiura</emphasis>
</taxonomicName>
was found on the mainland, across the bay, south of El Chaitén; a lack of sampling in the area between Alerce Andino National Park and Torres del Paine could be obscuring the true range of this species, making it seem nearly endemic to Chiloé when this could merely be a sampling bias. This inadequate sampling applies to
<taxonomicName id="5DCB4D049A138525FBDBFA4DFB2FFA16" box="[1085,1278,1448,1470]" class="Arachnida" family="Desidae" genus="Porteria" kingdom="Animalia" order="Araneae" pageId="38" pageNumber="406" phylum="Arthropoda" rank="species" species="albopunctata">
<emphasis id="A8BFEA959A138525FBDBFA4DFB2FFA16" box="[1085,1278,1448,1470]" italics="true" pageId="38" pageNumber="406">P. albopunctata</emphasis>
</taxonomicName>
as well, and possibly more undescribed species are awaiting discovery in southern
<collectingCountry id="E2DC76179A138525FBA4FA02FB52FA57" box="[1090,1155,1510,1532]" name="Chile" pageId="38" pageNumber="406">Chile</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>