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<mods:title id="5162735F7EFCE2E6AC70ECB403780383">Potamotrygon tigrina, a new species of freshwater stingray from the upper Amazon basin, closely related to Potamotrygon schroederi Fernandez-Yépez, 1958 (Chondrichthyes: Potamotrygonidae)</mods:title>
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<taxonomicName id="AFF24D43FFCAFFA2FF43FAEB8FC6FA14" ID-CoL="4M4RG" authority="Fernandez-Yepez, 1958" authorityName="Fernandez-Yepez" authorityYear="1958" box="[151,756,1403,1429]" class="Elasmobranchii" family="Potamotrygonidae" genus="Potamotrygon" kingdom="Animalia" order="Myliobatiformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="schroederi">
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<emphasis id="5A86EAD2FFCAFFA2FF43FAEB8CFAFA14" bold="true" box="[151,456,1403,1429]" italics="true" pageId="11" pageNumber="12">Potamotrygon schroederi</emphasis>
<bibRefCitation id="0C634B31FFCAFFA2FE1BFAEB8FC6FA14" author="Fernandez-Yepez" box="[463,756,1403,1429]" pageId="11" pageNumber="12" refString="Fernandez-Yepez, A. (1958) Nueva raya para la ciencia: Potomatrygon [sic] schroederi n. sp. Boletin Museo de Ciencia Natural, 2 - 3, 8 - 11." type="book chapter" year="1958">Fernandez-Yépez, 1958</bibRefCitation>
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<figureCitation id="F0C92A45FFCAFFA2FF43FA0E8C1FFA37" box="[151,301,1438,1462]" captionStart-0="FIGURE 9" captionStart-1="FIGURE 10" captionStart-10="FIGURE 19" captionStart-11="FIGURE 20" captionStart-2="FIGURE 11" captionStart-3="FIGURE 12" captionStart-4="FIGURE 13" captionStart-5="FIGURE 14" captionStart-6="FIGURE 15" captionStart-7="FIGURE 16" captionStart-8="FIGURE 17" captionStart-9="FIGURE 18" captionStartId-0="12.[151,250,969,992]" captionStartId-1="16.[151,250,1238,1261]" captionStartId-10="24.[151,250,1558,1581]" captionStartId-11="25.[151,250,1055,1078]" captionStartId-2="17.[151,250,1892,1915]" captionStartId-3="18.[151,250,1952,1975]" captionStartId-4="19.[151,250,907,930]" captionStartId-5="19.[151,250,1971,1994]" captionStartId-6="20.[151,250,1795,1818]" captionStartId-7="21.[151,250,1651,1674]" captionStartId-8="22.[151,250,1081,1104]" captionStartId-9="23.[151,250,1059,1082]" captionTargetBox-0="[498,1098,167,951]" captionTargetBox-1="[151,1436,176,1224]" captionTargetBox-10="[256,1327,406,1540]" captionTargetBox-11="[168,1424,198,1016]" captionTargetBox-2="[324,1264,581,1874]" captionTargetBox-3="[256,1326,183,1920]" captionTargetBox-4="[174,1417,177,889]" captionTargetBox-5="[172,1415,1004,1955]" captionTargetBox-6="[151,1436,193,1773]" captionTargetBox-7="[170,1413,185,1636]" captionTargetBox-8="[241,1342,195,1052]" captionTargetBox-9="[288,1298,182,1040]" captionTargetId-0="figure@12.[498,1099,167,951]" captionTargetId-1="figure@16.[151,1436,174,1225]" captionTargetId-10="figure@24.[256,1327,406,1540]" captionTargetId-11="figure@25.[151,1436,195,1034]" captionTargetId-2="figure@17.[324,1265,581,1874]" captionTargetId-3="figure@18.[256,1327,183,1922]" captionTargetId-4="figure@19.[174,1418,177,891]" captionTargetId-5="figure@19.[172,1416,1004,1955]" captionTargetId-6="figure@20.[151,1436,193,1774]" captionTargetId-7="figure@21.[170,1414,185,1636]" captionTargetId-8="figure@22.[241,1343,195,1053]" captionTargetId-9="figure@23.[288,1298,182,1041]" captionTargetPageId-0="12" captionTargetPageId-1="16" captionTargetPageId-10="24" captionTargetPageId-11="25" captionTargetPageId-2="17" captionTargetPageId-3="18" captionTargetPageId-4="19" captionTargetPageId-5="19" captionTargetPageId-6="20" captionTargetPageId-7="21" captionTargetPageId-8="22" captionTargetPageId-9="23" captionText-0="FIGURE 9. Remains of holotype of Potamotrygon schroederi (MHNLS 2504, Río Apure, Boca Apurito, Orinoco basin). A) Distal tail region with caudal stings. B) Nasoral region. C) Lateral portion of disc. Holotype was originally reported as a very large female (995 mm TL, 600 mm DL, 524 mm DW)." captionText-1="FIGURE 10. Freshly captured specimens of Potamotrygon schroederi from Venezuela (Orinoco basin). A) Dorsal view of AUM 44507 (Río Orinoco, island west of Puerto Venado, adult male, 780 mm TL, 470 mm DL, 420 mm DW). B) Dorsal view of ANSP 191999 (Río Ventuari, adult female, 605 mm TL, 395 mm DL, 358 mm DW)." captionText-10="FIGURE 19. Comparison of left anterolateral disc region in Potamotrygon schroederi among specimens from Orinoco (A and C, left column) and Negro basins (B and D, on right). A) AUM 44507. B) ANSP 191999. C) MZUSP 108452. D) MZUSP 108453. Scale bar = 5 cm." captionText-11="FIGURE 20. Ventral lateral line canals of Potamotrygon schroederi, highlighting specific features (on right) that may be derived for P. schroederi and P. tigrina (see text for details). Highlighted canals are as follows: 1: rostral loop of the infraorbital canal; 2: orbitonasal component of the supraorbital canal 3: suborbital loop of the infraorbital canal; 4: compressed posterior jugular loop formed between jugular and infraorbital canals. Illustration based on MZUSP 108455." captionText-2="FIGURE 11. Morphological details, in dorsal view, of freshly captured specimens of Potamotrygon schroederi from Venezuela (whole specimens shown in Figure 10). A) Orbital region of AUM 44507 (adult male, 780 mm TL). B) Orbital region of ANSP 191999 (adult female, 605 mm TL). C) Base of tail and posterior disc (AUM 44507), showing claspers. D) Base of tail and posterior disc (ANSP 191999). E) Ventral view of pelvic fins (ANSP 191999). F) Nasoral region (ANSP 191999). Scale bar = 5 cm, except for F (= 2 cm)." captionText-3="FIGURE 12. Dorsal views of freshly captured specimens of Potamotrygon schroederi from Rio Negro, near Barcelos, Brazil. A, B and D discarded after problems with temporary storage in Manaus. C) MZUSP 108453. Photographs by M. V. Domingues and N. Luchetti." captionText-4="FIGURE 13. Dorsal (A) and ventral (B) views of Potamotrygon schroederi from Barcelos, Rio Negro, Brazil (MZUSP 108452, 741 mm TL, 444 mm DL, 395 mm DW)." captionText-5="FIGURE 14. Potamotrygon schroederi from Barcelos, Rio Negro, Brazil, in dorsal view. A) MZUSP 108453 (689 mm TL, 444 mm DL, 409 mm DW). B) MZUSP 108445 (501 mm TL, 320 mm DL, 290 mm DW)." captionText-6="FIGURE 15. Morphological details of Potamotrygon schroederi from Rio Negro, Brazil. A) Dorsal anterior head and snout region (MZUSP 108453). B) Ventral snout and nasoral region (MZUSP 108452). C) Dorsal view of pelvic fin (MZUSP 108452). D) Ventral view of pelvis fins (MZUSP 108455). E) Dorsal view of posterior disc and tail base region in freshly captured adult female specimen (depicted Figure 12 A; specimen not preserved). F) Detail of lateral disc color in freshly captured adult female specimen (depicted in Figure 12 D; specimen not preserved). Scale bars in A, B, E and F = 5 cm; in C and D = 2.5 cm" captionText-7="FIGURE 16. Dorsal tail base region in specimens of Potamotrygon schroederi from Rio Negro, Brazil, showing distinct coloration and arrangement of tail spines and denticles. A-B) MZUSP 108453 (689 mm TL, 444 mm DL, 409 mm DW). C-D) MZUSP 108452 (741 mm TL, 444 mm DL, 395 mm DW). E) MZUSP 108445 (501 mm TL, 320 mm DL, 290 mm DW). F) MZUSP 108455 (319 mm TL, 202 mm DL, 188 mm DW). Scale bars = 5 cm." captionText-8="FIGURE 17. Skeletal features, dermal denticles and dorsal tail spines of Potamotrygon schroederi. A) Head, visceral arch, and scapular region. B) Close-up of angular cartilage. C) Base of tail region showing arrangement of dorsal spines and denticles. Based on AUM 44507 (adult male, 780 mm TL)." captionText-9="FIGURE 18. Morphological details of distal tail region in Potamotrygon schroederi, showing arrangement of dorsal tail spines, denticles, caudal stings, and vertebrae. A) AUM 44507 (adult male, 780 mm TL). B) ANSP 191999 (adult female, 605 mm TL)." httpUri-0="https://zenodo.org/record/319977/files/figure.png" httpUri-1="https://zenodo.org/record/3227628/files/figure.png" httpUri-10="https://zenodo.org/record/320012/files/figure.png" httpUri-11="https://zenodo.org/record/320016/files/figure.png" httpUri-2="https://zenodo.org/record/319980/files/figure.png" httpUri-3="https://zenodo.org/record/319985/files/figure.png" httpUri-4="https://zenodo.org/record/319991/files/figure.png" httpUri-5="https://zenodo.org/record/319995/files/figure.png" httpUri-6="https://zenodo.org/record/3227630/files/figure.png" httpUri-7="https://zenodo.org/record/3227632/files/figure.png" httpUri-8="https://zenodo.org/record/3227634/files/figure.png" httpUri-9="https://zenodo.org/record/320011/files/figure.png" pageId="11" pageNumber="12">Figures 9–20</figureCitation>
; Tables 3–4
</paragraph>
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<taxonomicName id="AFF24D43FFCAFFA2FF43FA708FABFA74" ID-CoL="4M4RG" authority="Fernandez-Yepez" authorityName="Fernandez-Yepez" authorityYear="1958" box="[151,665,1503,1525]" class="Elasmobranchii" family="Potamotrygonidae" genus="Potamotrygon" kingdom="Animalia" order="Myliobatiformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="schroederi">
<emphasis id="5A86EAD2FFCAFFA2FF43FA708CA9FA74" box="[151,411,1503,1525]" italics="true" pageId="11" pageNumber="12">Potamotrygon schroederi</emphasis>
<bibRefCitation id="0C634B31FFCAFFA2FE70FA4F8FABFA74" author="Fernandez-Yepez" box="[420,665,1503,1525]" pageId="11" pageNumber="12" refString="Fernandez-Yepez, A. (1958) Nueva raya para la ciencia: Potomatrygon [sic] schroederi n. sp. Boletin Museo de Ciencia Natural, 2 - 3, 8 - 11." type="book chapter" year="1958">Fernandez-Yépez (1958</bibRefCitation>
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[not 1957]: 7–10; original description, Boca Apurito, Río Apure, Orinoco drainage, figured); 
<bibRefCitation id="0C634B31FFCAFFA2FE5DFA6E8913F995" author="Mago-Leccia" box="[393,1057,1534,1556]" pageId="11" pageNumber="12" refString="Mago-Leccia, F. (1970) Lista de los peces de Venezuela, incluyendo un estudio preliminar de la ictiogeografia del pais. Ministerio de Agricultura y Cria, Caracas, 285 p." type="book" year="1970" yearSuffix="p">
Mago-Leccia (1970: 64; listed, 
<collectingCountry id="10E57650FFCAFFA2FD1DFA6E8E06F995" box="[713,820,1534,1556]" name="Venezuela" pageId="11" pageNumber="12">Venezuela</collectingCountry>
, photograph on p. 245)
</bibRefCitation>
; 
<bibRefCitation id="0C634B31FFCAFFA2FBFFFA6E8F5FF9B3" author="Castex" pageId="11" pageNumber="12" refString="Castex, M. N. &amp; Yagolkowski, D. R. (1970) Note on two fresh-water sting-rays species (Condrichthyes, Potamotrygonidae): a. Redescription of Potamotrygon schroederi, Yepez 1957. b. Designation of two paratypes of P. pauckei Castex 1963 and their description. Acta Scientifica, Universidad del Salvador, 9, 1 - 26." type="journal article" year="1970">Castex &amp; Yagolkowski (1970: 2–13; specimen from “Manaus”, redescription)</bibRefCitation>
; 
<bibRefCitation id="0C634B31FFCAFFA2FDACF98C8ED6F9B3" author="Mago-Leccia" box="[632,996,1564,1586]" pageId="11" pageNumber="12" refString="Mago-Leccia, F. (1978) Los Peces de Agua Dulce de Venezuela. Cuadernos Lagoven, Caracas. 36 p." type="book" year="1978">Mago-Leccia (1978: 5; name only)</bibRefCitation>
; 
<bibRefCitation id="0C634B31FFCAFFA2FC24F98D8C48F9D0" author="Rosa" pageId="11" pageNumber="12" refString="Rosa, R. S. (1985) A systematic revision of the South American freshwater stingrays (Chondrichthyes: Potamotrygonidae). Unpublished PhD diss., College of William and Mary, Williamsburg, xvi + 523 p." type="book" year="1985">Rosa (1985: 302–314; diagnosis, description, in part only)</bibRefCitation>
; 
<bibRefCitation id="0C634B31FFCAFFA2FE51F9AC8FA2F9D0" author="Compagno" box="[389,656,1595,1617]" pageId="11" pageNumber="12" refString="Compagno, L. J. V. &amp; Cook, S. F. (1995) The exploitation and conservation of freshwater elasmobranchs: status of taxa and prospects for the future. Pp. 62 - 90. In: Oetinger, M. I. &amp; Zorzi, G. D. (Eds.), The Biology of Freshwater Elasmobranchs, A Symposium to Honor Thomas B. Thorson. Journal of Aquariculture &amp; Aquatic Sciences, 7, 162 p." type="book chapter" year="1995">Compagno &amp; Cook (1995</bibRefCitation>
: 74; compilation, from 
<bibRefCitation id="0C634B31FFCAFFA2FCAAF9AC8EC1F9D0" author="Rosa" box="[894,1011,1595,1617]" pageId="11" pageNumber="12" refString="Rosa, R. S. (1985) A systematic revision of the South American freshwater stingrays (Chondrichthyes: Potamotrygonidae). Unpublished PhD diss., College of William and Mary, Williamsburg, xvi + 523 p." type="book" year="1985">Rosa, 1985</bibRefCitation>
); 
<bibRefCitation id="0C634B31FFCAFFA2FBD1F9AB89E4F9D0" author="Taphorn" box="[1029,1238,1595,1617]" pageId="11" pageNumber="12" refString="Taphorn, D., Royero, R., Machado-Allison, A. &amp; Mago Leccia, F. (1997) Lista actualizada de los peces de agua dulce de Venezuela. Pp. 55 - 100. In: La Marca, E. (Ed.), Catalogo zoologico de Venezuela. Vol. 1. Vertebrados actuales y fosiles de Venezuela. Museo de Ciencia y Tecnologia de Merida, Merida, 275 p." type="book chapter" year="1997">Taphorn et al. (1997</bibRefCitation>
: 67; listed, 
<collectingCountry id="10E57650FFCAFFA2FA9DF9AC8DDAF9F1" name="Venezuela" pageId="11" pageNumber="12">Venezuela</collectingCountry>
); 
<bibRefCitation id="0C634B31FFCAFFA2FF29F9CB8C89F9F1" author="Lasso" box="[253,443,1626,1648]" pageId="11" pageNumber="12" refString="Lasso, C., Ponte, V. &amp; Lasso-Alcala, O. (1998) Catalogo de la coleccion de tipos del Museo de Historia Natural La Salle, Caracas, Venezuela. Memoria Sociedad de Ciencias Naturales La Salle, 57 (147), 37 - 52." type="journal article" year="1998">Lasso et al. (1998</bibRefCitation>
: 41; 
<typeStatus id="B7498862FFCAFFA2FE24F9CA8F7BF9F1" box="[496,585,1626,1648]" pageId="11" pageNumber="12" type="holotype">holotype</typeStatus>
listed in catalog); 
<bibRefCitation id="0C634B31FFCAFFA2FCDBF9CB8990F9F1" author="Compagno" box="[783,1186,1626,1648]" pageId="11" pageNumber="12" refString="Compagno, L. J. V. (1999) Checklist of living Chondrichthyes. Pp. 471 - 498. In: Hamlett, W. C. (Ed.), Sharks, Skates and Rays, the Biology of Elasmobranch Fishes. Baltimore, The John Hopkins Univ. Press, 515 p." type="book chapter" year="1999">Compagno (1999: 495; listed as valid)</bibRefCitation>
; 
<bibRefCitation id="0C634B31FFCAFFA2FB7BF9CA8C37F90F" author="Machado-Allison" pageId="11" pageNumber="12" refString="Machado-Allison, A., Chernoff, B., Silvera, C., Bonilla, A., Lopez-Rojas, H., Lasso, C. A., Provenzano, F., Marcano, C. &amp; Machado-Aranda, D. (1999) Fish inventory of the Caura river basin, Bolivar state, Venezuela. Acta Biologica Venezuelica, 19 (4), 61 - 72." type="journal article" year="1999">Machado-Allison et al. (1999</bibRefCitation>
: 65; listed, 
<collectingCountry id="10E57650FFCAFFA2FE54F9E98CDFF90E" box="[384,493,1657,1679]" name="Venezuela" pageId="11" pageNumber="12">Venezuela</collectingCountry>
); 
<bibRefCitation id="0C634B31FFCAFFA2FDD4F9EA8FDEF90F" author="Ross" box="[512,748,1657,1679]" pageId="11" pageNumber="12" refString="Ross, R. A. &amp; Schafer, F. (2000) Aqualog Susswasser Rochen: Freshwater Rays. Verlag ACS, Morfelden-Walldorf, 192 p." type="book" year="2000">Ross &amp; Schäfer (2000</bibRefCitation>
: 45–46; brief account compiled from 
<bibRefCitation id="0C634B31FFCAFFA2FBA3F9EA89DCF90E" author="Rosa" box="[1143,1262,1657,1679]" pageId="11" pageNumber="12" refString="Rosa, R. S. (1985) A systematic revision of the South American freshwater stingrays (Chondrichthyes: Potamotrygonidae). Unpublished PhD diss., College of William and Mary, Williamsburg, xvi + 523 p." type="book" year="1985">Rosa, 1985</bibRefCitation>
); 
<bibRefCitation id="0C634B31FFCAFFA2FAD7F9E98C34F92C" author="Carvalho" pageId="11" pageNumber="12" refString="Carvalho, M. R. de, Lovejoy, N. R. &amp; Rosa, R. S. (2003) Family Potamotrygonidae. Pp. 22 - 29. In: Reis, R. E, Ferraris Jr, C. J. &amp; Kullander, S. O. (Eds.). Checklist of the Freshwater Fishes of South and Central America. Porto Alegre, EDIPUCRS, 729 p." type="book chapter" year="2003">Carvalho et al. (2003</bibRefCitation>
: 25; taxonomic account); 
<bibRefCitation id="0C634B31FFCAFFA2FDC1F9078E67F92C" author="Machado-Allison" box="[533,853,1687,1709]" pageId="11" pageNumber="12" refString="Machado-Allison, A., Chernoff, B., Provenzano, F., Willink, P. W., Marcano, A., Petry, P., Sidlauskas, B. &amp; Jones, T. (2003 a) Inventory, Relative Abundance and Importance of Fishes in the Caura River Basin, Bolivar State, Venezuela. Pp. 64 - 74. In: Chernoff, B., Machado-Allison, A., Riseng, K. &amp; Montambault, J. R. (Eds.), A Biological Assessment of the Aquatic Ecosystems of the Caura River Basin, Bolivar State, Venezuela. RAP Bulletin of Biological Assessment 28. Conservation International, Washington, DC, 284 p." type="book chapter" year="2003" yearSuffix="a">Machado-Allison et al. (2003a</bibRefCitation>
: 70; cited in text, 
<collectingCountry id="10E57650FFCAFFA2FBC0F907894DF92C" box="[1044,1151,1687,1709]" name="Venezuela" pageId="11" pageNumber="12">Venezuela</collectingCountry>
, as “ 
<taxonomicName id="AFF24D43FFCAFFA2FB60F9088DD1F94D" class="Elasmobranchii" family="Potamotrygonidae" genus="Potamotrygon" kingdom="Animalia" order="Myliobatiformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="schoederi">
<emphasis id="5A86EAD2FFCAFFA2FB60F9088DD1F94D" italics="true" pageId="11" pageNumber="12">Potamotrygon schoederi</emphasis>
</taxonomicName>
”); 
<bibRefCitation id="0C634B31FFCAFFA2FED2F9268F7AF94D" author="Machado-Allison" box="[262,584,1718,1740]" pageId="11" pageNumber="12" refString="Machado-Allison, A., Chernoff, B., Provenzano, F., Willink, P. W., Marcano, A., Petry, P., Jones, T. &amp; Sidlauskas, B. (2003 b) Fishes collected during the AquaRAP expedition to the Caura River Basin, Bolivar State, Venezuela. Pp. 267 - 275. In: Chernoff, B., Machado-Allison, A., Riseng, K. &amp; Montambault, J. R. (Eds.), A Biological Assessment of the Aquatic Ecosystems of the Caura River Basin, Bolivar State, Venezuela. RAP Bulletin of Biological Assessment 28. Conservation International, Washington, DC, 284 p." type="book chapter" year="2003" yearSuffix="b">Machado-Allison et al. (2003b</bibRefCitation>
: 267; listed in table, 
<collectingCountry id="10E57650FFCAFFA2FCFCF9268EA1F94D" box="[808,915,1718,1740]" name="Venezuela" pageId="11" pageNumber="12">Venezuela</collectingCountry>
, as “ 
<taxonomicName id="AFF24D43FFCAFFA2FC1CF92789D5F94D" box="[968,1255,1718,1740]" class="Elasmobranchii" family="Potamotrygonidae" genus="Potamotrygon" kingdom="Animalia" order="Myliobatiformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="schoederi">
<emphasis id="5A86EAD2FFCAFFA2FC1CF927896BF94D" box="[968,1113,1719,1740]" italics="true" pageId="11" pageNumber="12">Potamotrygon</emphasis>
cf. 
<emphasis id="5A86EAD2FFCAFFA2FB50F92689D5F94D" box="[1156,1255,1718,1740]" italics="true" pageId="11" pageNumber="12">schoederi</emphasis>
</taxonomicName>
”); 
<bibRefCitation id="0C634B31FFCAFFA2FADFF9268FF9F96B" author="Araujo" pageId="11" pageNumber="12" refString="Araujo, M. L. G. de (2004) Potamotrygon schroederi. In: IUCN 2010. IUCN Red List of Threatened Species. Version 2010.4. www. iucnredlist. org (downloaded on 20 March 2011)." type="journal article" year="2004">Araújo (2004; conservation status, biological data, no pagination)</bibRefCitation>
; 
<bibRefCitation id="0C634B31FFCAFFA2FD00F9458EB8F96B" author="Lasso" box="[724,906,1748,1770]" pageId="11" pageNumber="12" refString="Lasso, C., Lew, D., Taphorn, D., DoNascimiento, C., Lasso-Alcala, O., Provenzano, F. &amp; Machado-Allison, A. (2004) Biodiversidad ictiologica continental de Venezuela. Parte I. Lista de especies y distribucion por cuencas. Memoria de la Fundacion La Salle de Ciencias Naturales, 159 - 160, 105 - 195." type="book chapter" year="2004">Lasso et al. (2004</bibRefCitation>
: 144; listed, 
<collectingCountry id="10E57650FFCAFFA2FBDDF9448946F96B" box="[1033,1140,1748,1770]" name="Venezuela" pageId="11" pageNumber="12">Venezuela</collectingCountry>
, Orinoco); 
<bibRefCitation id="0C634B31FFCAFFA2FB31F9458CA3F888" author="Compagno" pageId="11" pageNumber="12" refString="Compagno, L. J. V. (2005) Checklist of living Chondrichthyes. Pp. 503 - 548. In: Hamlett, W. C. (Ed.), Reproductive Biology and Phylogeny of Chondrichthyes, Sharks, Batoids and Chimaeras. Enfield (NH), Science Publishers, Inc., 562 p." type="book chapter" year="2005">Compagno (2005: 541; listed as valid)</bibRefCitation>
; 
<bibRefCitation id="0C634B31FFCAFFA2FE4FF9638EE9F888" author="Gobbo" box="[411,987,1779,1801]" pageId="11" pageNumber="12" refString="Gobbo, R. G. (2006) Importancia sistematica da distribuicao dos canais ventrais da linha lateral em raias de agua doce neotropicais (Chondrichthyes: Myliobatiformes: Potamotrygonidae). Unpubl. Bachelor's monograph, Universidade de Sao Paulo, Sao Paulo, 55 p." type="book" year="2006">Gobbo (2006: 15–16; ventral lateral line canals, figured)</bibRefCitation>
; 
<bibRefCitation id="0C634B31FFCAFFA2FC31F96488AAF888" author="Rosa" box="[997,1432,1779,1801]" pageId="11" pageNumber="12" refString="Rosa, R. S. &amp; Carvalho, M. R. de (2007) Familia Potamotrygonidae. Pp. 17 - 18. In: Buckup, P. A., Menezes, N. A. &amp; Ghazzi, M. S. (Org.). Catalogo das especies de peixes de agua doce do Brasil. Rio de Janeiro, Museu Nacional, 195 p." type="book chapter" year="2007">
Rosa &amp; Carvalho (2007: 17; listed, 
<collectingCountry id="10E57650FFCAFFA2FA9AF96388A3F888" box="[1358,1425,1779,1801]" name="Brazil" pageId="11" pageNumber="12">Brazil</collectingCountry>
)
</bibRefCitation>
; 
<bibRefCitation id="0C634B31FFCAFFA2FF13F8828C8BF8A9" author="Mejia-Falla" box="[199,441,1810,1832]" pageId="11" pageNumber="12" refString="Mejia-Falla, P. A., Navia, A. F., Mejia-Ladino, L. M., Acero, A. &amp; Rubio, E. A. (2007) Tiburones y rayas de Colombia (Pisces Elasmobranchii): lista actualizada, revisada y comentada. Boletin de Investigaciones Marinas y Costeras, 36, 111 - 149." type="journal article" year="2007">Mejía-Falla et al. (2007</bibRefCitation>
: 126; listed, 
<collectingCountry id="10E57650FFCAFFA2FDECF8828F93F8A9" box="[568,673,1810,1832]" name="Colombia" pageId="11" pageNumber="12">Colombia</collectingCountry>
); 
<bibRefCitation id="0C634B31FFCAFFA2FD65F8828E47F8A9" author="Toffoli" box="[689,885,1810,1832]" pageId="11" pageNumber="12" refString="Toffoli, D., Hrbek, T., Araujo, M. L. G. de, Almeida, M. P. de, Charvet-Almeida, P. &amp; Farias, I. P. (2008) A test of the utility of DNA barcoding in the radiation of the freshwater stingray genus Potamotrygon (Potamotrygonidae, Myliobatiformes). Genetics and Molecular Biology, 31 (1) (suppl), 324 - 336." type="journal article" year="2008">Toffoli et al. (2008</bibRefCitation>
: 325-334; molecular phylogeny); 
<bibRefCitation id="0C634B31FFCAFFA2FB1FF8828C0EF8C7" author="Maldonado-Ocampo" pageId="11" pageNumber="12" refString="Maldonado-Ocampo, J. A., Vari, R. P. &amp; Usma, J. S. (2008) Checklist of the freshwater fishes of Colombia. Biota Colombiana 9 (2), 143 - 237." type="journal article" year="2008">Maldonado-Ocampo et al. (2008</bibRefCitation>
: 150; listed, 
<collectingCountry id="10E57650FFCAFFA2FE6FF8A18F16F8C6" box="[443,548,1841,1863]" name="Colombia" pageId="11" pageNumber="12">Colombia</collectingCountry>
); 
<bibRefCitation id="0C634B31FFCAFFA2FDE0F8A28FD3F8C7" author="Rosa" box="[564,737,1841,1863]" pageId="11" pageNumber="12" refString="Rosa, R. S., Charvet-Almeida, P. &amp; Quijada, C. C. D. (2010) Biology of the South American potamotrygonid stingrays. Pp. 241 - 281. In: Carrier, J. C., Musick, J. A. &amp; Heithaus, M. R. (Eds.), Sharks and Their Relativies II: Biodiversity, Adaptative Physiology, and Conservation. CRC Press, Florida, 736 p." type="book chapter" year="2010">Rosa et al. (2010</bibRefCitation>
: 252, 260; treated as valid); 
<bibRefCitation id="0C634B31FFCAFFA2FC29F8A289F5F8C7" author="Duncan" box="[1021,1223,1841,1863]" pageId="11" pageNumber="12" refString="Duncan, W. P., Inomata, S. O. &amp; Fernandes, M. N. (2010) Comercio de raias de agua doce na regiao do medio Rio Negro, Estado do Amazonas, Brasil. Revista Brasileira de Engenharia de Pesca, 5 (2), 13 - 22." type="journal article" year="2010">Duncan et al. (2010</bibRefCitation>
: 17–20; exploitation, photograph, Rio Negro).
</paragraph>
</subSubSection>
<subSubSection id="20E8654BFFCAFFA4FF43F8028859FEBE" lastPageId="13" lastPageNumber="14" pageId="11" pageNumber="12" type="materials_examined">
<paragraph id="684D36C0FFCAFFA2FF43F8028EB9F873" blockId="11.[151,1437,1938,2034]" pageId="11" pageNumber="12">
<emphasis id="5A86EAD2FFCAFFA2FF43F8028C36F82B" bold="true" box="[151,260,1938,1962]" pageId="11" pageNumber="12">
<typeStatus id="B7498862FFCAFFA2FF43F8028C36F82B" box="[151,260,1938,1962]" pageId="11" pageNumber="12" type="holotype">Holotype</typeStatus>
</emphasis>
. 
<collectionCode id="0EE3AE05FFCAFFA2FEC7F8028C47F82B" box="[275,373,1938,1962]" httpUri="http://grbio.org/cool/4kc6-e4ia" name="Coleccion de Mastozoologia, Museo de Historia Natural de La Salle" pageId="11" pageNumber="12">MHNLS</collectionCode>
2504, Río Apure, Boca Apurito, Orinoco basin, 
<collectingCountry id="10E57650FFCAFFA2FC70F8028928F82B" box="[932,1050,1938,1962]" name="Venezuela" pageId="11" pageNumber="12">Venezuela</collectingCountry>
, 
<date id="1C4C1000FFCAFFA2FBF2F80289EDF82B" box="[1062,1247,1938,1962]" pageId="11" pageNumber="12" value="1951-08-16">16 August 1951</date>
, coll. J. Marrero (only three fragments of the original specimen remain, comprising its caudal stings and distal tail region, mouth area, and a portion of the lateral disc; 
<bibRefCitation id="0C634B31FFCAFFA2FDEAF84B8E36F873" author="Lasso" box="[574,772,2010,2034]" pageId="11" pageNumber="12" refString="Lasso, C., Ponte, V. &amp; Lasso-Alcala, O. (1998) Catalogo de la coleccion de tipos del Museo de Historia Natural La Salle, Caracas, Venezuela. Memoria Sociedad de Ciencias Naturales La Salle, 57 (147), 37 - 52." type="journal article" year="1998">Lasso et al., 1998</bibRefCitation>
) (
<figureCitation id="F0C92A45FFCAFFA2FCCFF84A8E4FF873" box="[795,893,2010,2034]" captionStart="FIGURE 9" captionStartId="12.[151,250,969,992]" captionTargetBox="[498,1098,167,951]" captionTargetId="figure@12.[498,1099,167,951]" captionTargetPageId="12" captionText="FIGURE 9. Remains of holotype of Potamotrygon schroederi (MHNLS 2504, Río Apure, Boca Apurito, Orinoco basin). A) Distal tail region with caudal stings. B) Nasoral region. C) Lateral portion of disc. Holotype was originally reported as a very large female (995 mm TL, 600 mm DL, 524 mm DW)." httpUri="https://zenodo.org/record/319977/files/figure.png" pageId="11" pageNumber="12">Figure 9</figureCitation>
).
</paragraph>
<caption id="3C8D6648FFCDFFA5FF43FC598FF7FB9F" httpUri="https://zenodo.org/record/319977/files/figure.png" pageId="12" pageNumber="13" targetBox="[498,1098,167,951]" targetPageId="12">
<paragraph id="684D36C0FFCDFFA5FF43FC598FF7FB9F" blockId="12.[151,1436,969,1054]" pageId="12" pageNumber="13">
<emphasis id="5A86EAD2FFCDFFA5FF43FC598C3CFC5E" bold="true" box="[151,270,969,992]" pageId="12" pageNumber="13">FIGURE 9</emphasis>
. Remains of holotype of 
<taxonomicName id="AFF24D43FFCDFFA5FDC1FC5B8E2AFC61" box="[533,792,970,992]" class="Elasmobranchii" family="Potamotrygonidae" genus="Potamotrygon" kingdom="Animalia" order="Myliobatiformes" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="schroederi">
<emphasis id="5A86EAD2FFCDFFA5FDC1FC5B8E2AFC61" box="[533,792,970,992]" italics="true" pageId="12" pageNumber="13">Potamotrygon schroederi</emphasis>
</taxonomicName>
(MHNLS 2504, Río Apure, Boca Apurito, Orinoco basin). A) Distal tail region with caudal stings. B) Nasoral region. C) Lateral portion of disc. Holotype was originally reported as a very large female (995 mm TL, 600 mm DL, 524 mm DW).
</paragraph>
</caption>
<paragraph id="684D36C0FFCDFFA4FF43FBFC8859FEBE" blockId="12.[151,1436,1132,2021]" lastBlockId="13.[151,1436,151,319]" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
<emphasis id="5A86EAD2FFCDFFA5FF43FBFC8CF8FB04" bold="true" box="[151,458,1132,1157]" pageId="12" pageNumber="13">Other material examined</emphasis>
. (
<specimenCount id="7EF4FD49FFCDFFA5FE36FBFC8FB1FB05" box="[482,643,1132,1156]" pageId="12" pageNumber="13" type="generic">26 specimens</specimenCount>
). 
<collectingCountry id="10E57650FFCDFFA5FD4CFBFC8E39FB05" box="[664,779,1132,1156]" name="Venezuela" pageId="12" pageNumber="13">Venezuela</collectingCountry>
: 
<collectionCode id="0EE3AE05FFCDFFA5FCC8FBFC8E56FB05" LSID="urn:lsid:biocol.org:col:34784" box="[796,868,1132,1156]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="12" pageNumber="13">ANSP</collectionCode>
<date id="1C4C1000FFCDFFA5FCBAFBFC8EF7FB05" box="[878,965,1132,1156]" pageId="12" pageNumber="13" value="93-11-16">161193</date>
, preadult female (
<quantity id="AF0A9B25FFCDFFA5FB41FBFC89CAFB05" box="[1173,1272,1132,1156]" metricMagnitude="-1" metricUnit="m" metricValue="3.55" pageId="12" pageNumber="13" unit="mm" value="355.0">355 mm</quantity>
, 
<quantity id="AF0A9B25FFCDFFA5FAD3FBFC8854FB05" box="[1287,1382,1132,1156]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" pageId="12" pageNumber="13" unit="mm" value="200.0">200 mm</quantity>
DL, 
<quantity id="AF0A9B25FFCDFFA5FF43FB008DC6FB29" box="[151,244,1168,1192]" metricMagnitude="-1" metricUnit="m" metricValue="1.8" pageId="12" pageNumber="13" unit="mm" value="180.0">180 mm</quantity>
DW), Rio Caura at ferry crossing (E side) on Caicara-Ciudad Bolivar hwy., Bolivar state, 
<collectingCountry id="10E57650FFCDFFA5FB2AFB008845FB29" box="[1278,1399,1168,1192]" name="Venezuela" pageId="12" pageNumber="13">Venezuela</collectingCountry>
, 
<geoCoordinate id="0DC65007FFCDFFA5FA56FB008DE1FB4D" direction="north" orientation="latitude" pageId="12" pageNumber="13" precision="925" value="7.45">7° 27’N</geoCoordinate>
, 
<geoCoordinate id="0DC65007FFCDFFA5FF0AFB248C67FB4D" box="[222,341,1204,1228]" direction="west" orientation="longitude" pageId="12" pageNumber="13" precision="925" value="-65.2">065°12’W</geoCoordinate>
, 
<quantity id="AF0A9B25FFCDFFA5FEB4FB248C96FB4D" box="[352,420,1204,1228]" metricMagnitude="0" metricUnit="m" metricValue="1.0" metricValueMax="2.0" metricValueMin="0.0" pageId="12" pageNumber="13" unit="m" value="1.0" valueMax="2.0" valueMin="0.0">0-2 m</quantity>
, 
<quantity id="AF0A9B25FFCDFFA5FE7BFB248CD5FB4D" box="[431,487,1204,1228]" metricMagnitude="1" metricUnit="m" metricValue="1.524" pageId="12" pageNumber="13" unit="ft" value="50.0">50 ft</quantity>
. seine, 19:00-19:45, on bottom of sand and gravel, 
<date id="1C4C1000FFCDFFA5FBF4FB2489C8FB4D" box="[1056,1274,1204,1228]" pageId="12" pageNumber="13" value="1985-11-19">19 November 1985</date>
, coll. B. Chernoff, W.G. Saul, J.G. Lundberg, F. Provenzano, R. Royero (
<date id="1C4C1000FFCDFFA5FCE0FB488E5AFB71" box="[820,872,1240,1264]" pageId="12" pageNumber="13" value="1985-05">V85</date>
-35). 
<collectionCode id="0EE3AE05FFCDFFA5FC75FB488EDBFB71" LSID="urn:lsid:biocol.org:col:34784" box="[929,1001,1240,1264]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="12" pageNumber="13">ANSP</collectionCode>
191999, Río Ventuari (Orinoco basin), sandy beach and mouth of Caño Guayaje (right bank tributary), ca. 
<quantity id="AF0A9B25FFCDFFA5FC5FFB6C8ED2FA95" box="[907,992,1276,1300]" metricMagnitude="5" metricUnit="m" metricValue="2.1" pageId="12" pageNumber="13" unit="km" value="210.0">210 km</quantity>
northeast of San Fernando de Atabapo, Manapiare, 
<geoCoordinate id="0DC65007FFCDFFA5FECAFAB08CFBFAB9" box="[286,457,1312,1336]" direction="north" orientation="latitude" pageId="12" pageNumber="13" precision="1" value="5.1061597">5°6’22.176” N</geoCoordinate>
, 
<geoCoordinate id="0DC65007FFCDFFA5FE02FAB08F9CFAB9" box="[470,686,1312,1336]" direction="west" orientation="longitude" pageId="12" pageNumber="13" precision="1" value="-66.20916">066°12’32.976” W</geoCoordinate>
, 
<date id="1C4C1000FFCDFFA5FD6FFAB08E63FAB9" box="[699,849,1312,1336]" pageId="12" pageNumber="13" value="2010-04-04">4 April 2010</date>
, M. Sabaj Pérez and T. Carvalho (sampled at night with bag seine) (VEN 10–30); 
<collectionCode id="0EE3AE05FFCDFFA5FE3BFAD58F02FADC" box="[495,560,1349,1373]" httpUri="http://grbio.org/cool/8wk2-p21y" name="Auburn University Museum of Natural History" pageId="12" pageNumber="13">AUM</collectionCode>
44507, Río Orinoco, island W of Puerto Venado, 
<quantity id="AF0A9B25FFCDFFA5FB89FAD48999FADD" box="[1117,1195,1348,1372]" metricMagnitude="3" metricUnit="m" metricValue="4.5" pageId="12" pageNumber="13" unit="km" value="4.5">4.5 km</quantity>
S of Samariapo, 
<quantity id="AF0A9B25FFCDFFA5FABFFAD48D89FA01" metricMagnitude="4" metricUnit="m" metricValue="5.65" pageId="12" pageNumber="13" unit="km" value="56.5">56.5 km</quantity>
SW of Puerto Ayacucho, 
<geoCoordinate id="0DC65007FFCDFFA5FE34FAF88F57FA01" box="[480,613,1384,1408]" direction="north" orientation="latitude" pageId="12" pageNumber="13" precision="15" value="5.2069445">5°12’25” N</geoCoordinate>
, 
<geoCoordinate id="0DC65007FFCDFFA5FDA5FAF88E25FA01" box="[625,791,1384,1408]" direction="west" orientation="longitude" pageId="12" pageNumber="13" precision="15" value="-67.80889">067°48’32” W</geoCoordinate>
, 
<date id="1C4C1000FFCDFFA5FCF6FAF88E80FA01" box="[802,946,1384,1408]" pageId="12" pageNumber="13" value="2005-02-28">28 Feb 2005</date>
, coll. M. H. Sabaj, N. K. Lujan, D. C. Werneke, M. Arce et al. 
<collectingCountry id="10E57650FFCDFFA5FEA9FA1C8CF3FA25" box="[381,449,1420,1444]" name="Brazil" pageId="12" pageNumber="13">Brazil</collectingCountry>
: 
<collectionCode id="0EE3AE05FFCDFFA5FE04FA1C8F1FFA25" box="[464,557,1420,1444]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="12" pageNumber="13">MZUSP</collectionCode>
108443, Rio Negro, Amazonas, Barcelos, 
<geoCoordinate id="0DC65007FFCDFFA5FBC3FA1C89A4FA25" box="[1047,1174,1420,1444]" direction="south" orientation="latitude" pageId="12" pageNumber="13" precision="15" value="-0.9583333">0°57’30” S</geoCoordinate>
, 
<geoCoordinate id="0DC65007FFCDFFA5FB77FA1C887AFA25" box="[1187,1352,1420,1444]" direction="west" orientation="longitude" pageId="12" pageNumber="13" precision="15" value="-62.9375">062°56’15” W</geoCoordinate>
, 0 
<date id="1C4C1000FFCDFFA5FAB6FA1C8DE3FA49" pageId="12" pageNumber="13" value="2011-01-09">9 Jan 2011</date>
, coll. F. P. L. Marques, W. P. Duncan (RN11.14); 
<collectionCode id="0EE3AE05FFCDFFA5FCD7FA208E52FA49" box="[771,864,1456,1480]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="12" pageNumber="13">MZUSP</collectionCode>
108444, Rio Negro, Amazonas, Barcelos, coll. M. L. G. Araújo; 
<collectionCode id="0EE3AE05FFCDFFA5FEFBFA448CBEFA6D" box="[303,396,1492,1516]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="12" pageNumber="13">MZUSP</collectionCode>
108445, Rio Negro, Amazonas, Barcelos, 
<geoCoordinate id="0DC65007FFCDFFA5FCBEFA448EDBFA6D" box="[874,1001,1492,1516]" direction="south" orientation="latitude" pageId="12" pageNumber="13" precision="15" value="-0.9583333">0°57’30” S</geoCoordinate>
, 
<geoCoordinate id="0DC65007FFCDFFA5FC21FA4489ABFA6D" box="[1013,1177,1492,1516]" direction="west" orientation="longitude" pageId="12" pageNumber="13" precision="15" value="-62.9375">062°56’15” W</geoCoordinate>
, 
<date id="1C4C1000FFCDFFA5FB70FA44881EFA6D" box="[1188,1324,1492,1516]" pageId="12" pageNumber="13" value="2011-01-12">12 Jan 2011</date>
, coll. F. P. L. Marques, W. P. Duncan (RN11.34); 
<collectionCode id="0EE3AE05FFCDFFA5FD83FA688F86F991" box="[599,692,1528,1552]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="12" pageNumber="13">MZUSP</collectionCode>
108446, Rio Negro, Amazonas, Barcelos, coll. M. L. G. Araújo; 
<collectionCode id="0EE3AE05FFCDFFA5FF43F98C8DC6F9B5" box="[151,244,1564,1588]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="12" pageNumber="13">MZUSP</collectionCode>
108447, Rio Negro, Amazonas, Barcelos, Praia da Bananinha, Boca do Coibí, 
<geoCoordinate id="0DC65007FFCDFFA5FBBDF98C89D5F9B5" box="[1129,1255,1564,1588]" direction="south" orientation="latitude" pageId="12" pageNumber="13" precision="15" value="-0.6636111">0°39’49” S</geoCoordinate>
, 
<geoCoordinate id="0DC65007FFCDFFA5FB20F98C88AAF9B5" box="[1268,1432,1564,1588]" direction="west" orientation="longitude" pageId="12" pageNumber="13" precision="15" value="-63.016945">063°01’01” W</geoCoordinate>
, 
<date id="1C4C1000FFCDFFA5FF43F9D08C11F9D9" box="[151,291,1600,1624]" pageId="12" pageNumber="13" value="2011-01-14">14 Jan 2011</date>
, coll. F. P. L. Marques, W. P. Duncan (RN11.50); 
<collectionCode id="0EE3AE05FFCDFFA5FC8AF9D08E89F9D9" box="[862,955,1600,1624]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="12" pageNumber="13">MZUSP</collectionCode>
108448, Rio Negro, Amazonas, Barcelos, 
<geoCoordinate id="0DC65007FFCDFFA5FF43F9F48C2AF9FD" box="[151,280,1636,1660]" direction="south" orientation="latitude" pageId="12" pageNumber="13" precision="15" value="-0.9583333">0°57’30” S</geoCoordinate>
, 
<geoCoordinate id="0DC65007FFCDFFA5FEF2F9F48CFCF9FD" box="[294,462,1636,1660]" direction="west" orientation="longitude" pageId="12" pageNumber="13" precision="15" value="-62.9375">062°56’15” W</geoCoordinate>
, 
<date id="1C4C1000FFCDFFA5FE0EF9F48F5AF9FD" box="[474,616,1636,1660]" pageId="12" pageNumber="13" value="2011-01-16">16 Jan 2011</date>
, coll. F. P. L. Marques, W. P. Duncan (RN11.63); 
<collectionCode id="0EE3AE05FFCDFFA5FB7DF9F48834F9FD" box="[1193,1286,1636,1660]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="12" pageNumber="13">MZUSP</collectionCode>
108449, Rio Negro, Amazonas, Barcelos, coll. M. L. G. Araújo; 
<collectionCode id="0EE3AE05FFCDFFA5FD0FF9188E0AF921" box="[731,824,1672,1696]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="12" pageNumber="13">MZUSP</collectionCode>
108450, Rio Negro, Amazonas, Barcelos, Lago Cunimaru, 
<geoCoordinate id="0DC65007FFCDFFA5FF0BF93C8C6DF945" box="[223,351,1708,1732]" direction="south" orientation="latitude" pageId="12" pageNumber="13" precision="15" value="-0.83305556">0°49’59” S</geoCoordinate>
, 
<geoCoordinate id="0DC65007FFCDFFA5FEB8F93C8F20F945" box="[364,530,1708,1732]" direction="west" orientation="longitude" pageId="12" pageNumber="13" precision="15" value="-63.042778">063°02’34” W</geoCoordinate>
, 
<date id="1C4C1000FFCDFFA5FDC9F93C8F9AF945" box="[541,680,1708,1732]" pageId="12" pageNumber="13" value="2011-01-12">12 Jan 2011</date>
, coll. F. P. L. Marques, W. P. Duncan (RN11.42); 
<collectionCode id="0EE3AE05FFCDFFA5FB08F93C880BF945" box="[1244,1337,1708,1732]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="12" pageNumber="13">MZUSP</collectionCode>
108451, Rio Negro, Amazonas, Barcelos, 
<collectingCountry id="10E57650FFCDFFA5FDC7F9408F0DF969" box="[531,575,1744,1768]" name="Iceland" pageId="12" pageNumber="13">Ilha</collectingCountry>
do Jacami, 
<geoCoordinate id="0DC65007FFCDFFA5FD13F9408E75F969" box="[711,839,1744,1768]" direction="south" orientation="latitude" pageId="12" pageNumber="13" precision="15" value="-0.97999996">0°58’48” S</geoCoordinate>
, 
<geoCoordinate id="0DC65007FFCDFFA5FC81F9408EC9F969" box="[853,1019,1744,1768]" direction="west" orientation="longitude" pageId="12" pageNumber="13" precision="15" value="-62.920002">062°55’12” W</geoCoordinate>
, 0 
<date id="1C4C1000FFCDFFA5FBC1F94089AEF969" box="[1045,1180,1744,1768]" pageId="12" pageNumber="13" value="2004-03-06">6 Mar 2004</date>
, coll. F. P. L. Marques (RN04.94); 
<collectionCode id="0EE3AE05FFCDFFA5FECAF9648C49F88D" box="[286,379,1780,1804]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="12" pageNumber="13">MZUSP</collectionCode>
108452, Rio Negro, Amazonas, Barcelos, coll. M. L. G. Araújo; 
<collectionCode id="0EE3AE05FFCDFFA5FB8CF9648987F88D" box="[1112,1205,1780,1804]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="12" pageNumber="13">MZUSP</collectionCode>
108453, Rio Negro, Amazonas, Barcelos, 
<geoCoordinate id="0DC65007FFCDFFA5FE59F8888F3FF8B1" box="[397,525,1816,1840]" direction="south" orientation="latitude" pageId="12" pageNumber="13" precision="15" value="-0.97999996">0°58’48” S</geoCoordinate>
, 
<geoCoordinate id="0DC65007FFCDFFA5FDCFF8888FF3F8B1" box="[539,705,1816,1840]" direction="west" orientation="longitude" pageId="12" pageNumber="13" precision="15" value="-62.920002">062°55’12” W</geoCoordinate>
, 
<date id="1C4C1000FFCDFFA5FD19F8888E6BF8B1" box="[717,857,1816,1840]" pageId="12" pageNumber="13" value="2005-01-23">23 Jan 2005</date>
, coll. M. Domingues, N. Luchetti, M. L. G. Araújo (RN05.01); 
<collectionCode id="0EE3AE05FFCDFFA5FEF5F8AC8C4CF8D5" box="[289,382,1852,1876]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="12" pageNumber="13">MZUSP</collectionCode>
108454, Rio Negro, Amazonas, Barcelos, Rio Demeni, 
<geoCoordinate id="0DC65007FFCDFFA5FBDFF8AC89BFF8D5" box="[1035,1165,1852,1876]" direction="south" orientation="latitude" pageId="12" pageNumber="13" precision="15" value="-0.7336111">0°44’01” S</geoCoordinate>
, 
<geoCoordinate id="0DC65007FFCDFFA5FB48F8AC8876F8D5" box="[1180,1348,1852,1876]" direction="west" orientation="longitude" pageId="12" pageNumber="13" precision="15" value="-62.62722">062°37’38” W</geoCoordinate>
, 
<date id="1C4C1000FFCDFFA5FA85F8AC8DE3F8F9" pageId="12" pageNumber="13" value="2011-01-13">13 Jan 2011</date>
, coll. F. P. L. Marques, W. P. Duncan (RN11.48); 
<collectionCode id="0EE3AE05FFCDFFA5FCDCF8F08E57F8F9" box="[776,869,1888,1912]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="12" pageNumber="13">MZUSP</collectionCode>
108455, Rio Negro, Amazonas, Barcelos, 
<collectingCountry id="10E57650FFCDFFA5FA98F8F08845F8F9" box="[1356,1399,1888,1912]" name="Iceland" pageId="12" pageNumber="13">Ilha</collectingCountry>
do Jacami, 
<geoCoordinate id="0DC65007FFCDFFA5FF22F8148C4AF81D" box="[246,376,1924,1948]" direction="south" orientation="latitude" pageId="12" pageNumber="13" precision="15" value="-0.97999996">0°58’48” S</geoCoordinate>
, 
<geoCoordinate id="0DC65007FFCDFFA5FE5CF8148F02F81D" box="[392,560,1924,1948]" direction="west" orientation="longitude" pageId="12" pageNumber="13" precision="15" value="-62.920002">062°55’12” W</geoCoordinate>
, 
<date id="1C4C1000FFCDFFA5FDEAF8148FE1F81D" box="[574,723,1924,1948]" pageId="12" pageNumber="13" value="2004-02-24">24 Feb 2004</date>
, coll. F. P. L. Marques, M. L. G. Araújo (RN04.08); 
<collectionCode id="0EE3AE05FFCDFFA5FAEBF81488AEF81D" box="[1343,1436,1924,1948]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="12" pageNumber="13">MZUSP</collectionCode>
108456, Rio Negro, Amazonas, Barcelos, 
<collectingCountry id="10E57650FFCDFFA5FDADF8388F97F841" box="[633,677,1960,1984]" name="Iceland" pageId="12" pageNumber="13">Ilha</collectingCountry>
do Jacami, 
<geoCoordinate id="0DC65007FFCDFFA5FCFBF8388E82F841" box="[815,944,1960,1984]" direction="south" orientation="latitude" pageId="12" pageNumber="13" precision="15" value="-0.97999996">0°58’48” S</geoCoordinate>
, 
<geoCoordinate id="0DC65007FFCDFFA5FC6AF8388957F841" box="[958,1125,1960,1984]" direction="west" orientation="longitude" pageId="12" pageNumber="13" precision="15" value="-62.920002">062°55’12” W</geoCoordinate>
, 
<date id="1C4C1000FFCDFFA5FBA6F8388836F841" box="[1138,1284,1960,1984]" pageId="12" pageNumber="13" value="2004-02-29">29 Feb 2004</date>
, coll. F. P. L. Marques (RN04.54); 
<collectionCode id="0EE3AE05FFCDFFA5FE43F85C8CC6F865" box="[407,500,1996,2020]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="12" pageNumber="13">MZUSP</collectionCode>
108709 (cleared and stained), Rio Negro, Barcelos, Amazonas, 
<geoCoordinate id="0DC65007FFCDFFA5FADBF85C88A5F865" box="[1295,1431,1996,2020]" direction="south" orientation="latitude" pageId="12" pageNumber="13" precision="15" value="-0.97999996">0°58’48” S</geoCoordinate>
, 
<geoCoordinate id="0DC65007FFCCFFA4FF43FF078C09FF2E" box="[151,315,151,175]" direction="west" orientation="longitude" pageId="13" pageNumber="14" precision="15" value="-62.920002">062°55’12” W</geoCoordinate>
, 
<collectingCountry id="10E57650FFCCFFA4FE92FF078C40FF2E" box="[326,370,151,175]" name="Iceland" pageId="13" pageNumber="14">Ilha</collectingCountry>
Cuchara, 0 
<date id="1C4C1000FFCCFFA4FE26FF078FA3FF2E" box="[498,657,151,175]" pageId="13" pageNumber="14" value="2004-03-08">8 March 2004</date>
, coll. F. Marques (NR04.98); 
<collectionCode id="0EE3AE05FFCCFFA4FC0AFF078909FF2E" box="[990,1083,151,175]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="13" pageNumber="14">MZUSP</collectionCode>
108708 (dissected for muscles, central nervous system, and ventral lateral line), Rio Itu, Barcelos, Amazonas, 
<geoCoordinate id="0DC65007FFCCFFA4FBD2FF2C89B6FF55" box="[1030,1156,188,212]" direction="south" orientation="latitude" pageId="13" pageNumber="14" precision="15" value="-0.97999996">0°58’48” S</geoCoordinate>
, 
<geoCoordinate id="0DC65007FFCCFFA4FB44FF2C8807FF55" box="[1168,1333,188,212]" direction="west" orientation="longitude" pageId="13" pageNumber="14" precision="15" value="-62.920002">062°55’12” W</geoCoordinate>
, Boca do Igarapé Aduiá, 0 
<date id="1C4C1000FFCCFFA4FE8FFF4F8F33FF76" box="[347,513,223,247]" pageId="13" pageNumber="14" value="2004-03-08">8 March 2004</date>
, coll. A. da Silva Campos (NR04.93). [Seven specimens collected but not preserved, all from Rio Negro, Amazonas, Barcelos, 
<geoCoordinate id="0DC65007FFCCFFA4FD00FE948E67FE9D" box="[724,853,260,284]" direction="south" orientation="latitude" pageId="13" pageNumber="14" precision="15" value="-0.97999996">0°58’48” S</geoCoordinate>
, 
<geoCoordinate id="0DC65007FFCCFFA4FCB7FE948938FE9D" box="[867,1034,260,284]" direction="west" orientation="longitude" pageId="13" pageNumber="14" precision="15" value="-62.920002">062°55’12” W</geoCoordinate>
, 
<date id="1C4C1000FFCCFFA4FBC3FE9589B2FE9D" box="[1047,1152,260,284]" pageId="13" pageNumber="14" value="2005-01">Jan 2005</date>
, coll. M. Domingues, N. Luchetti, M. L. G. Araújo (field nos. RN05.12, RN05.21, RN05.22, RN05.23, RN05.25, RN05.36, RN05.43)].
</paragraph>
</subSubSection>
<subSubSection id="20E8654BFFCCFFA4FF43FEFF8C5CFCFE" pageId="13" pageNumber="14" type="diagnosis">
<paragraph id="684D36C0FFCCFFA4FF43FEFF8C5CFCFE" blockId="13.[151,1436,367,896]" pageId="13" pageNumber="14">
<emphasis id="5A86EAD2FFCCFFA4FF43FEFF8C38FE09" bold="true" box="[151,266,367,392]" pageId="13" pageNumber="14">Diagnosis</emphasis>
. A species of 
<taxonomicName id="AFF24D43FFCCFFA4FE65FEE18F63FE09" box="[433,593,369,392]" class="Elasmobranchii" family="Potamotrygonidae" genus="Potamotrygon" kingdom="Animalia" order="Myliobatiformes" pageId="13" pageNumber="14" phylum="Chordata" rank="genus">
<emphasis id="5A86EAD2FFCCFFA4FE65FEE18F63FE09" box="[433,593,369,392]" italics="true" pageId="13" pageNumber="14">Potamotrygon</emphasis>
</taxonomicName>
distinguished from all congeners by its unique dorsal color pattern, composed of a dark brown, black or olive background color forming a reticulate pattern, with numerous yellow, light brown or creamy colored cerebral or rosette-like designs over disc and base of tail region; these designs may contact each other but do not form a highly intricate vermiculate pattern. 
<taxonomicName id="AFF24D43FFCCFFA4FC14FE4D89D2FE75" box="[960,1248,476,500]" class="Elasmobranchii" family="Potamotrygonidae" genus="Potamotrygon" kingdom="Animalia" order="Myliobatiformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="schroederi">
<emphasis id="5A86EAD2FFCCFFA4FC14FE4D89D2FE75" box="[960,1248,476,500]" italics="true" pageId="13" pageNumber="14">Potamotrygon schroederi</emphasis>
</taxonomicName>
is distinguished from all species of 
<taxonomicName id="AFF24D43FFCCFFA4FEA0FD918F26FD99" box="[372,532,513,536]" class="Elasmobranchii" family="Potamotrygonidae" genus="Potamotrygon" kingdom="Animalia" order="Myliobatiformes" pageId="13" pageNumber="14" phylum="Chordata" rank="genus">
<emphasis id="5A86EAD2FFCCFFA4FEA0FD918F26FD99" box="[372,532,513,536]" italics="true" pageId="13" pageNumber="14">Potamotrygon</emphasis>
</taxonomicName>
, except 
<emphasis id="5A86EAD2FFCCFFA4FDA2FD918FE9FD99" box="[630,731,513,536]" italics="true" pageId="13" pageNumber="14">P. t i g r i n a</emphasis>
, by presenting the distal half of tail, posterior to caudal sting origin, with a sharply contrasted banding of five to six solid dark brown to blackish bars and uniformly cream colored interspaces of about equal width. Additionally, 
<taxonomicName id="AFF24D43FFCCFFA4FD3FFDD98E4DFDE1" box="[747,895,584,608]" class="Elasmobranchii" family="Potamotrygonidae" genus="Potamotrygon" kingdom="Animalia" order="Myliobatiformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="schroederi">
<emphasis id="5A86EAD2FFCCFFA4FD3FFDD98E4DFDE1" box="[747,895,584,608]" italics="true" pageId="13" pageNumber="14">P. schroederi</emphasis>
</taxonomicName>
can be distinguished from congeners, except 
<taxonomicName id="AFF24D43FFCCFFA4FA53FDD98DD1FD05" class="Elasmobranchii" family="Potamotrygonidae" genus="Potamotrygon" kingdom="Animalia" order="Myliobatiformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="tigrina">
<emphasis id="5A86EAD2FFCCFFA4FA53FDD98DD1FD05" italics="true" pageId="13" pageNumber="14">P. tigrina</emphasis>
</taxonomicName>
, 
<taxonomicName id="AFF24D43FFCCFFA4FF27FDFD8C5DFD05" box="[243,367,620,644]" class="Elasmobranchii" family="Potamotrygonidae" genus="Potamotrygon" kingdom="Animalia" order="Myliobatiformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="orbignyi">
<emphasis id="5A86EAD2FFCCFFA4FF27FDFD8C5DFD05" box="[243,367,620,644]" italics="true" pageId="13" pageNumber="14">P. orbignyi</emphasis>
</taxonomicName>
, 
<emphasis id="5A86EAD2FFCCFFA4FEAAFDFD8F34FD05" box="[382,518,620,644]" italics="true" pageId="13" pageNumber="14">P. hu m e ro s a</emphasis>
, 
<taxonomicName id="AFF24D43FFCCFFA4FDCDFDFD8F4CFD05" box="[537,638,620,644]" class="Elasmobranchii" family="Potamotrygonidae" genus="Potamotrygon" kingdom="Animalia" order="Myliobatiformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="histrix">
<emphasis id="5A86EAD2FFCCFFA4FDCDFDFD8F4CFD05" box="[537,638,620,644]" italics="true" pageId="13" pageNumber="14">P. histrix</emphasis>
</taxonomicName>
and 
<emphasis id="5A86EAD2FFCCFFA4FD63FDFD8E1DFD05" box="[695,815,621,644]" italics="true" pageId="13" pageNumber="14">P. m a r in ae</emphasis>
, by presenting a single developed and relatively short angular cartilage (more or less one-third length of hyomandibula; congeneric species with two more or less equally developed angulars, or with a single highly developed angular that is close to one-half hyomandibula length). The specific pattern of dorsal tail spines (usually low and well separated, occurring in one to three undefined rows, and originating relatively posteriorly on tail) further separates 
<taxonomicName id="AFF24D43FFCCFFA4FCF7FD6D8E87FC95" box="[803,949,764,788]" class="Elasmobranchii" family="Potamotrygonidae" genus="Potamotrygon" kingdom="Animalia" order="Myliobatiformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="schroederi">
<emphasis id="5A86EAD2FFCCFFA4FCF7FD6D8E87FC95" box="[803,949,764,788]" italics="true" pageId="13" pageNumber="14">P. schroederi</emphasis>
</taxonomicName>
from other 
<taxonomicName id="AFF24D43FFCCFFA4FBEEFD6D89E8FC95" box="[1082,1242,765,788]" class="Elasmobranchii" family="Potamotrygonidae" genus="Potamotrygon" kingdom="Animalia" order="Myliobatiformes" pageId="13" pageNumber="14" phylum="Chordata" rank="genus">
<emphasis id="5A86EAD2FFCCFFA4FBEEFD6D89E8FC95" box="[1082,1242,765,788]" italics="true" pageId="13" pageNumber="14">Potamotrygon</emphasis>
</taxonomicName>
species, with the exception of 
<taxonomicName id="AFF24D43FFCCFFA4FEFFFCB18CA6FCB9" box="[299,404,801,824]" class="Elasmobranchii" family="Potamotrygonidae" genus="Potamotrygon" kingdom="Animalia" order="Myliobatiformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="tigrina">
<emphasis id="5A86EAD2FFCCFFA4FEFFFCB18CA6FCB9" box="[299,404,801,824]" italics="true" pageId="13" pageNumber="14">P. tigrina</emphasis>
</taxonomicName>
(other congeners usually with a single or multiple row of dorsal tail spines, usually in well defined rows, with relatively taller tail and more closely packed spines, originating more anteriorly on dorsal tail region at tail base).
</paragraph>
</subSubSection>
<subSubSection id="20E8654BFFCCFFB8FF43FC3F8EBCFD9B" lastPageId="17" lastPageNumber="18" pageId="13" pageNumber="14" type="description">
<paragraph id="684D36C0FFCCFFA4FF43FC3F8CEEFC6D" blockId="13.[151,1437,943,1855]" pageId="13" pageNumber="14">
<emphasis id="5A86EAD2FFCCFFA4FF43FC3F8C13FC49" bold="true" box="[151,289,943,968]" pageId="13" pageNumber="14">Description</emphasis>
. Measurements are presented in Table 3 as both raw data in mm and transformed into % DW; meristic features are given in 
<tableCitation id="2570037BFFCCFFA4FE56FC448CEAFC6D" box="[386,472,980,1004]" captionStart="TABLE 4" captionStartId="15.[151,231,151,172]" captionText="TABLE 4. Meristic data for specimens of Potamotrygon schroederi. Specimens from Venezuela (Orinoco basin) in A – B, and speci-" pageId="13" pageNumber="14">Table 4</tableCitation>
.
</paragraph>
<paragraph id="684D36C0FFCCFFA4FF13FC688E52FAF9" blockId="13.[151,1437,943,1855]" pageId="13" pageNumber="14">
<emphasis id="5A86EAD2FFCCFFA4FF13FC688CF4FB91" bold="true" box="[199,454,1016,1040]" pageId="13" pageNumber="14">External morphology</emphasis>
. Disc clearly oval, longer than wide; disc length ranging from 106 to 116.4% DW (
<emphasis id="5A86EAD2FFCCFFA4FAACFC6988B6FB91" box="[1400,1412,1017,1040]" italics="true" pageId="13" pageNumber="14">x</emphasis>
= 110% DW), and widest close to its midlength. Snout oval, with small rostral knob protruding from anterior disc in smaller specimens, but very reduced and barely noticeable in larger individuals. Preorbital snout length more than twice interorbital distance, ranging from 18.7 to 29.8% DW (
<emphasis id="5A86EAD2FFCCFFA4FC83FBF58E51FBFD" box="[855,867,1125,1148]" italics="true" pageId="13" pageNumber="14">x</emphasis>
= 26.4% DW). Prenasal (14.1 to 19.9% DW, 
<emphasis id="5A86EAD2FFCCFFA4FAA2FBF588B0FBFD" box="[1398,1410,1125,1148]" italics="true" pageId="13" pageNumber="14">x</emphasis>
= 17% DW) and preoral (19.0 to 25.2% DW, 
<emphasis id="5A86EAD2FFCCFFA4FD5DFB198FA7FB21" box="[649,661,1161,1184]" italics="true" pageId="13" pageNumber="14">x</emphasis>
= 22.5% DW) snout lengths relatively short, shorter than preorbital snout length. Eyes somewhat small, protruding from top of head region; eyes smaller than spiracles in diameter. Spiracles trapezoidal in fresh material, greatly expanded, but more contracted and slit-like in preserved specimens. Spiracles closely adjacent to eyes, about as wide as long in fresh material, and without elevated spiracular rims or central knob posteriorly. Spiracles extend anteriorly to level of one-half to posterior two-thirds of eye length in preserved material, but spiracles extend anteriorly to beyond eye length in live specimens. Interorbital distance slightly smaller than, or about equal to, interspiracular distance.
</paragraph>
<paragraph id="684D36C0FFCCFFA4FF13FA148F4EF8BE" blockId="13.[151,1437,943,1855]" pageId="13" pageNumber="14">
Mouth somewhat small, its opening relatively straight across in larger specimens, but somewhat arched in smaller individuals. Mouth width slightly less than one-half of distance between first gill slits, and slightly greater than internarial distance. Faint grooves present on posterior margins of lower jaw and outer corners of jaws due to preservation; labial folds absent. Nostrils elongated anteroposteriorly, slit-like, similar in length to eye diameter, and about one-half of internarial distance. Nasal curtain widest posteriorly, with posterior margin fringed and with only slight medial notch. Teeth not visible externally with mouth closed. Tooth rows in adult females ranging from 45–49/53–60, from 38–41/45–52 rows in preadult females, and 38–41/ 
<quantity id="AF0A9B25FFCCFFA4FC00F9C5890EF9EC" box="[980,1084,1621,1645]" metricMagnitude="0" metricUnit="m" metricValue="1.1557" metricValueMax="1.1684" metricValueMin="1.143" pageId="13" pageNumber="14" unit="in" value="45.5" valueMax="46.0" valueMin="45.0">45–46 in</quantity>
juveniles (
<tableCitation id="2570037BFFCCFFA4FB14F9C5882EF9EC" box="[1216,1308,1621,1645]" captionStart="TABLE 4" captionStartId="15.[151,231,151,172]" captionText="TABLE 4. Meristic data for specimens of Potamotrygon schroederi. Specimens from Venezuela (Orinoco basin) in A – B, and speci-" pageId="13" pageNumber="14">Table 4</tableCitation>
; teeth also counted in specimens not in table). Teeth set in quincunx, relatively small, rhomboidal, and without conspicuous cusps. Five buccal papillae present inside mouth, lateralmost papillae much smaller in height. Branchial basket longer than wide, its length ranging from 16.5 to 18.6% DW (x = 17.6% DW). Ventral gill region relatively wide, distance between first and fifth gill slits about equal to distance between first pair of gill openings; distance between first gill slits slightly greater than distance between fifth gill slits. Gill flaps slightly undulated; fifth gill slit smallest, and slightly more obliquely positioned.
</paragraph>
<paragraph id="684D36C0FFCFFFA7FFA3FE698E5FFDF6" blockId="14.[119,1947,504,1716]" pageId="14" pageNumber="15">
<emphasis id="5A86EAD2FFCFFFA7FFA3FE698DEEFD92" bold="true" box="[119,220,505,531]" pageId="14" pageNumber="15">Table 3</emphasis>
. Measurements of specimens of 
<taxonomicName id="AFF24D43FFCFFFA7FD8EFE698EB6FD92" box="[602,900,504,531]" class="Elasmobranchii" family="Potamotrygonidae" genus="Potamotrygon" kingdom="Animalia" order="Myliobatiformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="schroederi">
<emphasis id="5A86EAD2FFCFFFA7FD8EFE698EB6FD92" box="[602,900,504,531]" italics="true" pageId="14" pageNumber="15">Potamotrygon schroederi</emphasis>
</taxonomicName>
from 
<collectingCountry id="10E57650FFCFFFA7FC1FFE698974FD92" box="[971,1094,505,531]" name="Venezuela" pageId="14" pageNumber="15">Venezuela</collectingCountry>
(Orinoco basin) in A–C and 
<collectingCountry id="10E57650FFCFFFA7FA4EFE6988D3FD92" box="[1434,1505,505,531]" name="Brazil" pageId="14" pageNumber="15">Brazil</collectingCountry>
(Negro basin) in D–F. A: 
<typeStatus id="B7498862FFCFFFA7F8C0FE698A4DFD92" box="[1812,1919,505,531]" pageId="14" pageNumber="15" type="holotype">Holotype</typeStatus>
from 
<bibRefCitation id="0C634B31FFCFFFA7FF15FD8A8CEEFDB5" author="Fernandez-Yepez" box="[193,476,538,564]" pageId="14" pageNumber="15" refString="Fernandez-Yepez, A. (1958) Nueva raya para la ciencia: Potomatrygon [sic] schroederi n. sp. Boletin Museo de Ciencia Natural, 2 - 3, 8 - 11." type="book chapter" year="1958">Fernandez-Yépez, 1958</bibRefCitation>
). B: AUM 44507 (adult male). C: ANSP 191999 (adult female). D: MZUSP 108453 (adult female). E: MZUSP 108452 (adult female). F: MZUSP 108456 (juvenile male). N: number of specimens measured; 
<emphasis id="5A86EAD2FFCFFFA7FBF3FDAC8906FDD4" box="[1063,1076,572,597]" italics="true" pageId="14" pageNumber="15">x</emphasis>
: mean; SD: standard deviation. Note that 
<typeStatus id="B7498862FFCFFFA7F9CAFDAC8BB1FDD7" box="[1566,1667,572,598]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
(A) is not included in range, mean or SD. * measurement not given in original account.
</paragraph>
<paragraph id="684D36C0FFCFFFA7FF28FD1C8A57FD1E" blockId="14.[119,1947,504,1716]" box="[252,1893,652,671]" pageId="14" pageNumber="15">
PARAMETER A B C D E F range 
<emphasis id="5A86EAD2FFCFFFA7F97BFD1C8B8BFD1F" box="[1711,1721,652,670]" italics="true" pageId="14" pageNumber="15">x</emphasis>
SD
</paragraph>
<paragraph id="684D36C0FFCFFFA7FE2BFD2E8ABFFCC5" blockId="14.[119,1947,504,1716]" pageId="14" pageNumber="15">Mm % DW mm % DW mm % DW mm % DW mm % DW mm % DW % DW N mm % DW mm % DW total length (TL) 995.0 – 780.0 – 605.0 – 689.0 – 741.0 – 320.0 – – 16 492.1 – 209.6 – disc length (DL) 600.0 114.5 470.0 111.9 395.0 110.3 444.0 108.6 444.0 112.4 181.0 106.5 106.0 – 116.4 16 301.8 110.4 128.2 3.0 disc width (DW) 524.0 100.0 420.0 100.0 358.0 100.0 409.0 100.0 395.0 100.0 170.0 100.0 – 16 273.1 100.0 113.3 0.0</paragraph>
<paragraph id="684D36C0FFCFFFA7F9E6FCDB8B78FCE1" blockId="14.[119,1947,504,1716]" box="[1586,1610,843,864]" pageId="14" pageNumber="15">16</paragraph>
<paragraph id="684D36C0FFCFFFA6FF4EFCC888AEFF4B" blockId="14.[119,1947,504,1716]" lastBlockId="15.[151,1436,151,754]" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
interorbital distance 67.0 12.8 47.5 11.3 42.5 11.9 48.0 11.7 48.4 12.3 18.0 10.6 8.2 – 15.3 32.9 12.0 14.4 2.0 interspiracular distance 65.0 12.4 61.5 14.6 51.7 14.4 49.0 12.0 50.6 12.8 30.0 17.6 12.0 – 17.6 16 37.1 13.9 13.9 1.6 eye length * – 13.4 3.2 12.5 3.5 13.0 3.2 10.6 2.7 12.0 7.1 2.7 – 7.1 16 9.9 4.1 2.4 1.1 spiracle length * – 37.0 8.8 28.0 7.8 33.8 8.3 36.0 9.1 11.0 6.5 6.0 – 9.1 16 20.3 7.7 9.0 0.9 preorbital length 160.0 30.5 113.2 27.0 101.0 28.2 106.3 26.0 105.0 26.6 41.0 24.1 18.7 – 29.8 16 73.9 26.8 34.1 2.7 prenasal length * – 60.8 14.5 56.3 15.7 61.8 15.1 61.6 15.6 24.0 14.1 14.1 – 19.9 16 42.2 16.8 13.0 1.7 preoral length * – 80.9 19.3 73.6 20.6 77.9 19.0 83.6 21.2 35.0 20.6 19.0 – 25.2 16 55.8 22.2 16.9 1.9 internarial distance * – 33.5 8.0 24.9 7.0 27.9 6.8 26.8 6.8 12.0 7.1 6.5 – 8.0 16 18.2 7.0 7.3 0.5 mouth width * – 32.0 7.6 24.6 6.9 36.0 8.8 38.2 9.7 11.0 6.5 5.0 – 9.7 16 19.1 7.2 9.5 1.2 distance between 1st gill slits * – 105.2 25.0 86.1 24.1 84.9 20.8 89.8 22.7 39.0 22.9 18.7 – 25.0 16 56.1 21.6 23.7 1.7 distance between 5th gill slits * – 67.8 16.1 57.5 16.1 63.3 15.5 60.5 15.3 27.0 15.9 13.3 – 16.5 16 38.9 15.0 15.6 1.0 branchial basket length * – 82.3 19.6 64.9 18.1 71.6 17.5 73.5 18.6 31.0 18.2 16.5 – 19.6 16 46.1 17.8 18.4 0.7 pelvic fin anterior margin length 135.0 25.8 92.6 22.0 85.0 23.7 74.1 18.1 90.9 23.0 35.0 20.6 18.1 – 24.7 16 61.1 22.2 27.9 2.2 pelvic fin width 300.0 57.3 238.0 56.7 215.0 60.1 166.0 40.6 208.0 52.7 88.0 51.8 40.6 – 60.1 16 143.2 52.2 64.8 5.3 clasper external length * – 50.9 12.1 – – – – – – 6.0 3.5 1.2 – 12.1 3 19.6 5.6 27.2 5.7 clasper internal length * – 98.7 23.5 – – – – – – 20.0 11.8 0.6 – 23.5 3 39.9 12.0 51.8 11.4 distance between cloaca and tail tip * – 386.0 91.9 273.0 76.3 300.0 73.3 379.0 95.9 166.0 97.6 59.6 – 98.3 16 218.9 85.7 83.6 10.5 tail width 68.0 13.0 66.5 15.8 50.6 14.1 53.7 13.1 49.9 12.6 31.0 18.2 9.0 – 18.2 16 35.0 12.5 17.6 2.3 snout to cloaca distance 502.0 95.8 403.0 96.0 316.0 88.3 354.0 86.6 368.0 93.2 142.0 83.5 83.5 – 96.0 16 246.4 89.5 109.4 3.5 pectoral to posterior pelvic length * – 55.8 13.3 37.6 10.5 56.0 13.7 55.0 13.9 16.0 9.4 8.2 – 18.8 16 31.8 12.1 14.6 2.7 distance from cloaca to sting origin * – 236.0 56.2 171.0 47.8 233.0 57.0 246.0 62.3 77.0 45.3 39.4 – 62.3 14 133.3 50.0 66.6 6.0 sting length 97.0 18.5 72.1 17.2 74.8 20.9 48.0 11.7 82.8 21.0 32.0 18.8 11.7 – 21.8 14 48.0 17.5 23.1 3.3 sting width * – 7.3 1.7 5.5 1.5 4.9 1.2 5.0 1.3 3.0 1.8 1.0 – 1.9 14 3.8 1.5 1.5 0.3 mens from 
<collectingCountry id="10E57650FFCEFFA6FED1FF258C72FF4B" box="[261,320,181,202]" name="Brazil" pageId="15" pageNumber="16">Brazil</collectingCountry>
(Negro basin) in C–J. A: AUM 44507 (adult male, 
<quantity id="AF0A9B25FFCEFFA6FCE3FF258EB4FF4B" box="[823,902,181,202]" metricMagnitude="-1" metricUnit="m" metricValue="7.8" pageId="15" pageNumber="16" unit="mm" value="780.0">780 mm</quantity>
TL). B) ANSP 191999 (adult female, 
<quantity id="AF0A9B25FFCEFFA6FB29FF25887EFF4B" box="[1277,1356,181,202]" metricMagnitude="-1" metricUnit="m" metricValue="6.05" pageId="15" pageNumber="16" unit="mm" value="605.0">605 mm</quantity>
TL). C:
</paragraph>
<caption id="3C8D6648FFCEFFA6FF43FF0788AEFF2D" box="[151,1436,151,173]" pageId="15" pageNumber="16">
<paragraph id="684D36C0FFCEFFA6FF43FF0788AEFF2D" blockId="15.[151,1436,151,754]" box="[151,1436,151,173]" pageId="15" pageNumber="16">
<emphasis id="5A86EAD2FFCEFFA6FF43FF078DCBFF2D" bold="true" box="[151,249,151,172]" pageId="15" pageNumber="16">TABLE 4</emphasis>
. Meristic data for specimens of 
<taxonomicName id="AFF24D43FFCEFFA6FDE1FF088E19FF2C" box="[565,811,152,173]" class="Elasmobranchii" family="Potamotrygonidae" genus="Potamotrygon" kingdom="Animalia" order="Myliobatiformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="schroederi">
<emphasis id="5A86EAD2FFCEFFA6FDE1FF088E19FF2C" box="[565,811,152,173]" italics="true" pageId="15" pageNumber="16">Potamotrygon schroederi</emphasis>
</taxonomicName>
. Specimens from Venezuela (Orinoco basin) in A–B, and speci-
</paragraph>
</caption>
<paragraph id="684D36C0FFCEFFA6FF43FF4288AEFF66" blockId="15.[151,1436,151,754]" box="[151,1436,210,231]" pageId="15" pageNumber="16">
MZUSP 108452 (adult female, 
<quantity id="AF0A9B25FFCEFFA6FE1EFF428F28FF66" box="[458,538,210,231]" metricMagnitude="-1" metricUnit="m" metricValue="4.4399999999999995" pageId="15" pageNumber="16" unit="mm" value="444.0">444 mm</quantity>
DL). D: MZUSP 108453 (adult female, 
<quantity id="AF0A9B25FFCEFFA6FC7CFF428ECBFF66" box="[936,1017,210,231]" metricMagnitude="-1" metricUnit="m" metricValue="4.4399999999999995" pageId="15" pageNumber="16" unit="mm" value="444.0">444 mm</quantity>
DL). E: MZUSP 108445 (preadult female,
</paragraph>
<paragraph id="684D36C0FFCEFFA6FF43FF7F88AEFE85" blockId="15.[151,1436,151,754]" box="[151,1436,239,260]" pageId="15" pageNumber="16">
<quantity id="AF0A9B25FFCEFFA6FF43FF7F8DDAFE85" box="[151,232,239,260]" metricMagnitude="-1" metricUnit="m" metricValue="3.2" pageId="15" pageNumber="16" unit="mm" value="320.0">320 mm</quantity>
DL). F: MZUSP 108447 (preadult female, 
<quantity id="AF0A9B25FFCEFFA6FD4CFF7F8FDBFE85" box="[664,745,239,260]" metricMagnitude="-1" metricUnit="m" metricValue="3.05" pageId="15" pageNumber="16" unit="mm" value="305.0">305 mm</quantity>
DL). G: MZUSP 108448 (preadult female, 
<quantity id="AF0A9B25FFCEFFA6FB49FF7F89DCFE85" box="[1181,1262,239,260]" metricMagnitude="-1" metricUnit="m" metricValue="2.4899999999999998" pageId="15" pageNumber="16" unit="mm" value="249.0">249 mm</quantity>
DL). H: MZUSP
</paragraph>
<paragraph id="684D36C0FFCEFFA6FF43FE9D88AEFEA3" blockId="15.[151,1436,151,754]" box="[151,1436,269,290]" pageId="15" pageNumber="16">
108455 (juvenile male, 
<quantity id="AF0A9B25FFCEFFA6FEA9FE9D8CFEFEA3" box="[381,460,269,290]" metricMagnitude="-1" metricUnit="m" metricValue="2.02" pageId="15" pageNumber="16" unit="mm" value="202.0">202 mm</quantity>
DL). I: MZUSP 108451 (juvenile female, 
<quantity id="AF0A9B25FFCEFFA6FCB9FE9D8E8FFEA3" box="[877,957,269,290]" metricMagnitude="-1" metricUnit="m" metricValue="1.93" pageId="15" pageNumber="16" unit="mm" value="193.0">193 mm</quantity>
DL). J: MZUSP 108456 (juvenile male, 
<quantity id="AF0A9B25FFCEFFA6FA98FE9D88AEFEA3" box="[1356,1436,269,290]" metricMagnitude="-1" metricUnit="m" metricValue="1.81" pageId="15" pageNumber="16" unit="mm" value="181.0">181 mm</quantity>
</paragraph>
<paragraph id="684D36C0FFCEFFA6FF43FEBA8DF4FEBE" blockId="15.[151,1436,151,754]" box="[151,198,298,319]" pageId="15" pageNumber="16">DL).</paragraph>
<paragraph id="684D36C0FFCEFFA6FECEFEDF89C8FD73" blockId="15.[151,1436,151,754]" pageId="15" pageNumber="16">CHARACTER A B C D E F G H I J range mode precaudal vertebrae 28 27 31 27 29 30 29 25 29 31 25–31 29 caudal vertebrae 124 122 99 100 96 100 88 81 104 98 81–124 100 total vertebrae 152 149 130 127 125 130 117 106 133 129 106–152 130 diplospondylous vertebrae 120 119 96 98 95 96 84 77 101 92 77–120 96 upper tooth rows 43 44 50 55 41 38 47 39 40 38 38–55 38 lower tooth rows 50 51 60 59 52 45 52 48 49 45 45–60 52 propterygial radials 40 40 42 41 38 40 39 40 41 45 38–45 40 mesopterygial radials 17 18 17 16 17 17 16 16 17 16 16–18 17 metapterygial radials 38 35 35 35 34 36 34 37 34 34 34–38 34 total pectoral radials 95 93 94 92 89 93 89 93 92 95 89–95 93 pelvic radials 24 26 24 22 20 19 24 21 19–26 24</paragraph>
<paragraph id="684D36C0FFCEFFA6FF13FCB18F75FB5C" blockId="15.[151,1437,801,2036]" pageId="15" pageNumber="16">
Pelvic fins somewhat short, much wider than long (ranging from 40.6 to 58.0% DW, 
<emphasis id="5A86EAD2FFCEFFA6FB7DFCB38987FCBB" box="[1193,1205,803,826]" italics="true" pageId="15" pageNumber="16">x</emphasis>
= 50.9% DW), and broadest at more or less their anterior third. Apices of pelvic fins broadly oval; posterior margins of pelvics undulated. Pelvic fins mostly concealed in dorsal view. Tail relatively wide at base, its width about equal to interorbital distance (tail width ranging from 9.0 to 18.2% DW, 
<emphasis id="5A86EAD2FFCEFFA6FD33FC1C8FC1FC22" box="[743,755,908,931]" italics="true" pageId="15" pageNumber="16">x</emphasis>
= 12.1% DW). Tail tapering posteriorly, but not resembling an elongated whip distally. Ventral tail flat in cross-section. Lateral tail folds slender, ridge-like in adults, wider in juveniles, and originating at about base of tail, extending to more or less level of caudal stings; ridge-like dorsal and ventral tail folds present, originating posterior to caudal stings and extending caudally for a short distance. Dorsal and ventral tail folds anteriorly wider, extending posteriorly as a very shallow ridge. Length from cloaca to distal tail variable, ranging between 59.6 and 98.3% DW (
<emphasis id="5A86EAD2FFCEFFA6FCF7FBAA8E1DFBD0" box="[803,815,1082,1105]" italics="true" pageId="15" pageNumber="16">x</emphasis>
= 85.9% DW), much smaller than disc length, but similar to snout to cloaca length (varying from 83.5 to 94.0% DW, 
<emphasis id="5A86EAD2FFCEFFA6FCA7FBCD8E4DFBF5" box="[883,895,1117,1140]" italics="true" pageId="15" pageNumber="16">x</emphasis>
= 88.7% DW). Caudal sting varying greatly in length, ranging from 11.7 to 21.8% DW (
<emphasis id="5A86EAD2FFCEFFA6FDBDFB108F47FB16" box="[617,629,1152,1175]" italics="true" pageId="15" pageNumber="16">x</emphasis>
= 17.2% DW), and much greater than interorbital distance or tail width at base. Caudal sting with posteriorly directed, sharp serrations on each side, terminating in a very acute point, and positioned posterior to tail mid-length.
</paragraph>
<paragraph id="684D36C0FFCEFFA6FF13FB778863F875" blockId="15.[151,1437,801,2036]" pageId="15" pageNumber="16">
<emphasis id="5A86EAD2FFCEFFA6FF13FB778C7AFA81" bold="true" box="[199,328,1255,1280]" pageId="15" pageNumber="16">Coloration</emphasis>
. Dorsal surface of disc with a gray, olive, dark brown to purplish-black background color, covered by numerous conspicuous tan, beige, golden yellow or orangish rosettes and brain-shaped figures over disc and anterior dorsal tail region (
<figureCitation id="F0C92A45FFCEFFA6FE12FAA08F5FFAC9" box="[454,621,1328,1352]" captionStart-0="FIGURE 10" captionStart-1="FIGURE 11" captionStart-2="FIGURE 12" captionStart-3="FIGURE 13" captionStart-4="FIGURE 14" captionStart-5="FIGURE 15" captionStart-6="FIGURE 16" captionStartId-0="16.[151,250,1238,1261]" captionStartId-1="17.[151,250,1892,1915]" captionStartId-2="18.[151,250,1952,1975]" captionStartId-3="19.[151,250,907,930]" captionStartId-4="19.[151,250,1971,1994]" captionStartId-5="20.[151,250,1795,1818]" captionStartId-6="21.[151,250,1651,1674]" captionTargetBox-0="[151,1436,176,1224]" captionTargetBox-1="[324,1264,581,1874]" captionTargetBox-2="[256,1326,183,1920]" captionTargetBox-3="[174,1417,177,889]" captionTargetBox-4="[172,1415,1004,1955]" captionTargetBox-5="[151,1436,193,1773]" captionTargetBox-6="[170,1413,185,1636]" captionTargetId-0="figure@16.[151,1436,174,1225]" captionTargetId-1="figure@17.[324,1265,581,1874]" captionTargetId-2="figure@18.[256,1327,183,1922]" captionTargetId-3="figure@19.[174,1418,177,891]" captionTargetId-4="figure@19.[172,1416,1004,1955]" captionTargetId-5="figure@20.[151,1436,193,1774]" captionTargetId-6="figure@21.[170,1414,185,1636]" captionTargetPageId-0="16" captionTargetPageId-1="17" captionTargetPageId-2="18" captionTargetPageId-3="19" captionTargetPageId-4="19" captionTargetPageId-5="20" captionTargetPageId-6="21" captionText-0="FIGURE 10. Freshly captured specimens of Potamotrygon schroederi from Venezuela (Orinoco basin). A) Dorsal view of AUM 44507 (Río Orinoco, island west of Puerto Venado, adult male, 780 mm TL, 470 mm DL, 420 mm DW). B) Dorsal view of ANSP 191999 (Río Ventuari, adult female, 605 mm TL, 395 mm DL, 358 mm DW)." captionText-1="FIGURE 11. Morphological details, in dorsal view, of freshly captured specimens of Potamotrygon schroederi from Venezuela (whole specimens shown in Figure 10). A) Orbital region of AUM 44507 (adult male, 780 mm TL). B) Orbital region of ANSP 191999 (adult female, 605 mm TL). C) Base of tail and posterior disc (AUM 44507), showing claspers. D) Base of tail and posterior disc (ANSP 191999). E) Ventral view of pelvic fins (ANSP 191999). F) Nasoral region (ANSP 191999). Scale bar = 5 cm, except for F (= 2 cm)." captionText-2="FIGURE 12. Dorsal views of freshly captured specimens of Potamotrygon schroederi from Rio Negro, near Barcelos, Brazil. A, B and D discarded after problems with temporary storage in Manaus. C) MZUSP 108453. Photographs by M. V. Domingues and N. Luchetti." captionText-3="FIGURE 13. Dorsal (A) and ventral (B) views of Potamotrygon schroederi from Barcelos, Rio Negro, Brazil (MZUSP 108452, 741 mm TL, 444 mm DL, 395 mm DW)." captionText-4="FIGURE 14. Potamotrygon schroederi from Barcelos, Rio Negro, Brazil, in dorsal view. A) MZUSP 108453 (689 mm TL, 444 mm DL, 409 mm DW). B) MZUSP 108445 (501 mm TL, 320 mm DL, 290 mm DW)." captionText-5="FIGURE 15. Morphological details of Potamotrygon schroederi from Rio Negro, Brazil. A) Dorsal anterior head and snout region (MZUSP 108453). B) Ventral snout and nasoral region (MZUSP 108452). C) Dorsal view of pelvic fin (MZUSP 108452). D) Ventral view of pelvis fins (MZUSP 108455). E) Dorsal view of posterior disc and tail base region in freshly captured adult female specimen (depicted Figure 12 A; specimen not preserved). F) Detail of lateral disc color in freshly captured adult female specimen (depicted in Figure 12 D; specimen not preserved). Scale bars in A, B, E and F = 5 cm; in C and D = 2.5 cm" captionText-6="FIGURE 16. Dorsal tail base region in specimens of Potamotrygon schroederi from Rio Negro, Brazil, showing distinct coloration and arrangement of tail spines and denticles. A-B) MZUSP 108453 (689 mm TL, 444 mm DL, 409 mm DW). C-D) MZUSP 108452 (741 mm TL, 444 mm DL, 395 mm DW). E) MZUSP 108445 (501 mm TL, 320 mm DL, 290 mm DW). F) MZUSP 108455 (319 mm TL, 202 mm DL, 188 mm DW). Scale bars = 5 cm." httpUri-0="https://zenodo.org/record/3227628/files/figure.png" httpUri-1="https://zenodo.org/record/319980/files/figure.png" httpUri-2="https://zenodo.org/record/319985/files/figure.png" httpUri-3="https://zenodo.org/record/319991/files/figure.png" httpUri-4="https://zenodo.org/record/319995/files/figure.png" httpUri-5="https://zenodo.org/record/3227630/files/figure.png" httpUri-6="https://zenodo.org/record/3227632/files/figure.png" pageId="15" pageNumber="16">Figures 10–16</figureCitation>
, 
<figureCitation id="F0C92A45FFCEFFA6FDAEFAA08FABFAC9" box="[634,665,1328,1352]" captionStart="FIGURE 19" captionStartId="24.[151,250,1558,1581]" captionTargetBox="[256,1327,406,1540]" captionTargetId="figure@24.[256,1327,406,1540]" captionTargetPageId="24" captionText="FIGURE 19. Comparison of left anterolateral disc region in Potamotrygon schroederi among specimens from Orinoco (A and C, left column) and Negro basins (B and D, on right). A) AUM 44507. B) ANSP 191999. C) MZUSP 108452. D) MZUSP 108453. Scale bar = 5 cm." httpUri="https://zenodo.org/record/320012/files/figure.png" pageId="15" pageNumber="16">19</figureCitation>
). Background color becomes slightly lighter toward outer disc margin. Outer disc with numerous spots much smaller than rosettes, more or less equal to eye diameter. These spots may be elongated, oval or appear as streaks of lighter color. Dorsal aspect of pelvic fins with beige to golden spots, smaller toward disc margins and larger at central pelvic fins region. Elaborate rosette-like figures on dorsal disc and anterior tail region formed by numerous smaller, lighter spots grouped into irregular circular, oval, clover or brain-shaped figures with lighter color in center. Size and particular pattern of elaborate rosettes vary from larger and more convoluted in disc center to smaller and less invaginated toward outer disc; markings very tightly associated, with more elaborate and finest convoluted pattern on interorbital region and posterior to spiracles on middisc. Dorsal rosettes and brain-shaped figures do not merge to form more conspicuous vermicular patterns, except over head and interorbital regions where finer, more slender rosette outlines usually occur. In specimens from the Orinoco basin, rosette outlines more elaborate. Smaller individuals with rosettes and brain-shaped figures proportionally smaller and more simple, with fewer convolutions, resembling ocelli with darker centers. Rosettes and brainshaped figures may form ocelli-like markings with darker central color in adults, but more restricted to outer and posterolateral disc surface. Anterior to mid-dorsal tail region with saddles of lighter color formed from somewhat incomplete or simple rosettes. Dorsal and lateral tail with strong banded pattern of alternating darker and lighter bars from caudal sting origin to posterior tail extremity. Ventral coloration uniform white to creamy-white, without conspicuous markings, but darker posterior margins of disc and pelvic fins sometimes present, especially in smaller individuals. Ventral tail with laterally appearing darker bands continuous from dorsal and lateral tail; these bands do not connect at ventral tail midline, more noticeable in smaller specimens. Posterior tail region as of caudal stings with alternating dark and lighter bands present; tail extremity dark. Live specimens with more intense color.
</paragraph>
<caption id="3C8D6648FFD1FFB9FF43FB46893CFAAB" httpUri="https://zenodo.org/record/3227628/files/figure.png" pageId="16" pageNumber="17" targetBox="[151,1436,176,1224]" targetPageId="16">
<paragraph id="684D36C0FFD1FFB9FF43FB46893CFAAB" blockId="16.[151,1436,1238,1322]" pageId="16" pageNumber="17">
<emphasis id="5A86EAD2FFD1FFB9FF43FB468C29FB6D" bold="true" box="[151,283,1238,1261]" pageId="16" pageNumber="17">FIGURE 10</emphasis>
. Freshly captured specimens of 
<taxonomicName id="AFF24D43FFD1FFB9FDA5FB488E44FB6C" box="[625,886,1239,1261]" class="Elasmobranchii" family="Potamotrygonidae" genus="Potamotrygon" kingdom="Animalia" order="Myliobatiformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="schroederi">
<emphasis id="5A86EAD2FFD1FFB9FDA5FB488E44FB6C" box="[625,886,1239,1261]" italics="true" pageId="16" pageNumber="17">Potamotrygon schroederi</emphasis>
</taxonomicName>
from Venezuela (Orinoco basin). A) Dorsal view of AUM 44507 (Río Orinoco, island west of Puerto Venado, adult male, 780 mm TL, 470 mm DL, 420 mm DW). B) Dorsal view of ANSP 191999 (Río Ventuari, adult female, 605 mm TL, 395 mm DL, 358 mm DW).
</paragraph>
</caption>
<paragraph id="684D36C0FFD1FFB9FF13FAC28F5BF92C" blockId="16.[151,1436,1361,2033]" pageId="16" pageNumber="17">
<emphasis id="5A86EAD2FFD1FFB9FF13FAC28CA2FAEB" bold="true" box="[199,400,1361,1386]" pageId="16" pageNumber="17">Dermal denticles</emphasis>
. The dermal covering in 
<taxonomicName id="AFF24D43FFD1FFB9FD7DFAC28E09FAE8" box="[681,827,1361,1385]" class="Elasmobranchii" family="Potamotrygonidae" genus="Potamotrygon" kingdom="Animalia" order="Myliobatiformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="schroederi">
<emphasis id="5A86EAD2FFD1FFB9FD7DFAC28E09FAE8" box="[681,827,1361,1385]" italics="true" pageId="16" pageNumber="17">P. schroederi</emphasis>
</taxonomicName>
is very similar to 
<emphasis id="5A86EAD2FFD1FFB9FBD0FAC2895BFAE8" box="[1028,1129,1362,1385]" italics="true" pageId="16" pageNumber="17">P. t ig r in a</emphasis>
(see above). Dermal denticles present over central disc and dorsal tail base regions, composed of small denticles more or less evenly distributed. Denticles slightly greater and more spaced apart over middisc area, continuing posteriorly to base of tail. Base of dorsal tail with slightly enlarged, flattened, asterisk-shaped denticles. Denticles over midline of disc and tail with three or more crown dichotomies converging on crown apex. Dorsal tail spines in one to four very irregular rows. Dorsal tail spines smaller in juveniles, but increasingly taller, very straight in lateral view, and with robust bases in adults. Dorsal tail spines widely spaced apart (
<figureCitation id="F0C92A45FFD1FFB9FD78F9B98E15F9C0" box="[684,807,1577,1601]" captionStart="FIGURE 16" captionStartId="21.[151,250,1651,1674]" captionTargetBox="[170,1413,185,1636]" captionTargetId="figure@21.[170,1414,185,1636]" captionTargetPageId="21" captionText="FIGURE 16. Dorsal tail base region in specimens of Potamotrygon schroederi from Rio Negro, Brazil, showing distinct coloration and arrangement of tail spines and denticles. A-B) MZUSP 108453 (689 mm TL, 444 mm DL, 409 mm DW). C-D) MZUSP 108452 (741 mm TL, 444 mm DL, 395 mm DW). E) MZUSP 108445 (501 mm TL, 320 mm DL, 290 mm DW). F) MZUSP 108455 (319 mm TL, 202 mm DL, 188 mm DW). Scale bars = 5 cm." httpUri="https://zenodo.org/record/3227632/files/figure.png" pageId="16" pageNumber="17">Figures 16</figureCitation>
, 
<figureCitation id="F0C92A45FFD1FFB9FCE3F9B98E64F9C0" box="[823,854,1577,1601]" captionStart="FIGURE 18" captionStartId="23.[151,250,1059,1082]" captionTargetBox="[288,1298,182,1040]" captionTargetId="figure@23.[288,1298,182,1041]" captionTargetPageId="23" captionText="FIGURE 18. Morphological details of distal tail region in Potamotrygon schroederi, showing arrangement of dorsal tail spines, denticles, caudal stings, and vertebrae. A) AUM 44507 (adult male, 780 mm TL). B) ANSP 191999 (adult female, 605 mm TL)." httpUri="https://zenodo.org/record/320011/files/figure.png" pageId="16" pageNumber="17">18</figureCitation>
). Smaller spines occur in between taller dorsal tail spines. Dorsal tail spines originate relatively posteriorly on dorsal tail, posterior to, or more or less at level of, disc apices. Lateral tail region, posterior to or level with caudal stings and extending to distal tail extremity, with small, sharp denticles, especially in large adults.
</paragraph>
<paragraph id="684D36C0FFD1FFB8FF13F9298EBCFD9B" blockId="16.[151,1436,1361,2033]" lastBlockId="17.[151,1436,151,539]" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
<emphasis id="5A86EAD2FFD1FFB9FF13F9298CA2F953" bold="true" box="[199,400,1721,1746]" pageId="16" pageNumber="17">Skeletal features</emphasis>
. Skeleton very similar to 
<taxonomicName id="AFF24D43FFD1FFB9FD6BF92A8E18F950" box="[703,810,1722,1745]" class="Elasmobranchii" family="Potamotrygonidae" genus="Potamotrygon" kingdom="Animalia" order="Myliobatiformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="tigrina">
<emphasis id="5A86EAD2FFD1FFB9FD6BF92A8E18F950" box="[703,810,1722,1745]" italics="true" pageId="16" pageNumber="17">P. tigrina</emphasis>
</taxonomicName>
. Neurocranium elongate, widest at nasal capsules and postorbital processes (
<figureCitation id="F0C92A45FFD1FFB9FE46F94D8F36F974" box="[402,516,1757,1781]" captionStart="FIGURE 17" captionStartId="22.[151,250,1081,1104]" captionTargetBox="[241,1342,195,1052]" captionTargetId="figure@22.[241,1343,195,1053]" captionTargetPageId="22" captionText="FIGURE 17. Skeletal features, dermal denticles and dorsal tail spines of Potamotrygon schroederi. A) Head, visceral arch, and scapular region. B) Close-up of angular cartilage. C) Base of tail region showing arrangement of dorsal spines and denticles. Based on AUM 44507 (adult male, 780 mm TL)." httpUri="https://zenodo.org/record/3227634/files/figure.png" pageId="16" pageNumber="17">Figure 17</figureCitation>
). Nasal capsules much wider than long, broadly rounded; internasal septum slen- der. Preorbital processes slightly wider than postorbital processes. Precerebral and frontoparietal fontenellae about fourth-fifths of neurocranial length; very short otic roof posteriorly at parietal fossa. Supraorbital process sharply triangular. Preorbital processes slender and acute distally. Antorbital cartilage very elongate and triangular, extending posteriorly to close to angular cartilage. Stout Meckel’s cartilage with dorsally projecting lateral process well developed; palatoquadrate much more slender. Hyomandibulae relatively slender and only slightly curved anteriorly toward midline. Anterior angular cartilage well developed, about one-third to one-fourth length of hyomandibula, with concave anterior margin; posterior angular apparently absent. Elongate cervicothoracic synarcual, about as wide as neurocranium, with about 30 spinal nerve foramina. Thoracolumbar synarcual slender. Individual vertebral centra occurring posterior to level of caudal sting origin (
<figureCitation id="F0C92A45FFD0FFB8FC8EFF2C8EFFFF55" box="[858,973,188,212]" captionStart="FIGURE 18" captionStartId="23.[151,250,1059,1082]" captionTargetBox="[288,1298,182,1040]" captionTargetId="figure@23.[288,1298,182,1041]" captionTargetPageId="23" captionText="FIGURE 18. Morphological details of distal tail region in Potamotrygon schroederi, showing arrangement of dorsal tail spines, denticles, caudal stings, and vertebrae. A) AUM 44507 (adult male, 780 mm TL). B) ANSP 191999 (adult female, 605 mm TL)." httpUri="https://zenodo.org/record/320011/files/figure.png" pageId="17" pageNumber="18">Figure 18</figureCitation>
); somewhat calcified notochordal extension (cartilaginous rod) present distally. Transition from mono- to diplospondyly occurs at about fourth centra posterior to pelvic girdle. Propterygium widest posteriorly, much more stout than meso- and metapterygium. Metapterygium with four posterior segments. Anterior segment of propterygium elongate, about equal in length to nasal aperture. Mesopterygium slightly convex externally. Pectoral radial elements sometimes fused at base, especially at posterior propterygium. Pectoral radials slender close to pectoral basals, slightly widening and shortening toward middisc, and slender again distally; pectoral radials subdivided at 9th or 10th segment. Pelvic girdle concave anteriorly lateral to low prepelvic process. Iliac processes extending caudally just beyond triangular ischial processes. Three to four obturator foramina present. Basipterygium stout, short, but longer than posterior metapterygial segments. Enlarged pelvic radial articulating with iliac process.
</paragraph>
</subSubSection>
<caption id="3C8D6648FFD0FFB8FF43F8F48C93F874" httpUri="https://zenodo.org/record/319980/files/figure.png" pageId="17" pageNumber="18" targetBox="[324,1264,581,1874]" targetPageId="17">
<paragraph id="684D36C0FFD0FFB8FF43F8F48C93F874" blockId="17.[151,1436,1892,2037]" pageId="17" pageNumber="18">
<emphasis id="5A86EAD2FFD0FFB8FF43F8F48C25F8FB" bold="true" box="[151,279,1892,1915]" pageId="17" pageNumber="18">FIGURE 11</emphasis>
. Morphological details, in dorsal view, of freshly captured specimens of 
<taxonomicName id="AFF24D43FFD0FFB8FC21F8F689C5F8FA" box="[1013,1271,1893,1915]" class="Elasmobranchii" family="Potamotrygonidae" genus="Potamotrygon" kingdom="Animalia" order="Myliobatiformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="schroederi">
<emphasis id="5A86EAD2FFD0FFB8FC21F8F689C5F8FA" box="[1013,1271,1893,1915]" italics="true" pageId="17" pageNumber="18">Potamotrygon schroederi</emphasis>
</taxonomicName>
from Venezuela (whole specimens shown in Figure 10). A) Orbital region of AUM 44507 (adult male, 780 mm TL). B) Orbital region of ANSP 191999 (adult female, 605 mm TL). C) Base of tail and posterior disc (AUM 44507), showing claspers. D) Base of tail and posterior disc (ANSP 191999). E) Ventral view of pelvic fins (ANSP 191999). F) Nasoral region (ANSP 191999). Scale bar = 5 cm, except for F (= 2 cm).
</paragraph>
</caption>
<caption id="3C8D6648FFD3FFBBFF43F8308C0EF875" httpUri="https://zenodo.org/record/319985/files/figure.png" pageId="18" pageNumber="19" targetBox="[256,1326,183,1920]" targetPageId="18">
<paragraph id="684D36C0FFD3FFBBFF43F8308C0EF875" blockId="18.[151,1437,1952,2036]" pageId="18" pageNumber="19">
<emphasis id="5A86EAD2FFD3FFBBFF43F8308C2BF837" bold="true" box="[151,281,1952,1975]" pageId="18" pageNumber="19">FIGURE 12</emphasis>
. Dorsal views of freshly captured specimens of 
<taxonomicName id="AFF24D43FFD3FFBBFCD2F832893BF836" box="[774,1033,1953,1975]" class="Elasmobranchii" family="Potamotrygonidae" genus="Potamotrygon" kingdom="Animalia" order="Myliobatiformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="schroederi">
<emphasis id="5A86EAD2FFD3FFBBFCD2F832893BF836" box="[774,1033,1953,1975]" italics="true" pageId="18" pageNumber="19">Potamotrygon schroederi</emphasis>
</taxonomicName>
from Rio Negro, near Barcelos, Brazil. A, B and D discarded after problems with temporary storage in Manaus. C) MZUSP 108453. Photographs by M. V. Domingues and N. Luchetti.
</paragraph>
</caption>
<caption id="3C8D6648FFD2FFBAFF43FC1B8FA3FC41" httpUri="https://zenodo.org/record/319991/files/figure.png" pageId="19" pageNumber="20" targetBox="[174,1417,177,889]" targetPageId="19">
<paragraph id="684D36C0FFD2FFBAFF43FC1B8FA3FC41" blockId="19.[151,1436,907,960]" pageId="19" pageNumber="20">
<emphasis id="5A86EAD2FFD2FFBAFF43FC1B8C2FFC20" bold="true" box="[151,285,907,930]" pageId="19" pageNumber="20">FIGURE 13</emphasis>
. Dorsal (A) and ventral (B) views of 
<taxonomicName id="AFF24D43FFD2FFBAFD6AFC1D8EF7FC23" box="[702,965,908,930]" class="Elasmobranchii" family="Potamotrygonidae" genus="Potamotrygon" kingdom="Animalia" order="Myliobatiformes" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="schroederi">
<emphasis id="5A86EAD2FFD2FFBAFD6AFC1D8EF7FC23" box="[702,965,908,930]" italics="true" pageId="19" pageNumber="20">Potamotrygon schroederi</emphasis>
</taxonomicName>
from Barcelos, Rio Negro, Brazil (MZUSP 108452, 741 mm TL, 444 mm DL, 395 mm DW).
</paragraph>
</caption>
<caption id="3C8D6648FFD2FFBAFF43F823891DF868" httpUri="https://zenodo.org/record/319995/files/figure.png" pageId="19" pageNumber="20" targetBox="[172,1415,1004,1955]" targetPageId="19">
<paragraph id="684D36C0FFD2FFBAFF43F823891DF868" blockId="19.[151,1436,1971,2025]" pageId="19" pageNumber="20">
<emphasis id="5A86EAD2FFD2FFBAFF43F8238C28F848" bold="true" box="[151,282,1971,1994]" pageId="19" pageNumber="20">FIGURE 14</emphasis>
. 
<taxonomicName id="AFF24D43FFD2FFBAFEFDF8258F1FF84B" box="[297,557,1972,1994]" class="Elasmobranchii" family="Potamotrygonidae" genus="Potamotrygon" kingdom="Animalia" order="Myliobatiformes" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="schroederi">
<emphasis id="5A86EAD2FFD2FFBAFEFDF8258F1FF84B" box="[297,557,1972,1994]" italics="true" pageId="19" pageNumber="20">Potamotrygon schroederi</emphasis>
</taxonomicName>
from Barcelos, Rio Negro, Brazil, in dorsal view. A) MZUSP 108453 (689 mm TL, 444 mm DL, 409 mm DW). B) MZUSP 108445 (501 mm TL, 320 mm DL, 290 mm DW).
</paragraph>
</caption>
<caption id="3C8D6648FFD5FFBDFF43F8938D84F833" httpUri="https://zenodo.org/record/3227630/files/figure.png" pageId="20" pageNumber="21" targetBox="[151,1436,193,1773]" targetPageId="20">
<paragraph id="684D36C0FFD5FFBDFF43F8938D84F833" blockId="20.[151,1437,1795,1970]" pageId="20" pageNumber="21">
<emphasis id="5A86EAD2FFD5FFBDFF43F8938C28F899" bold="true" box="[151,282,1795,1818]" pageId="20" pageNumber="21">FIGURE 15</emphasis>
. Morphological details of 
<taxonomicName id="AFF24D43FFD5FFBDFDFBF8948E01F898" box="[559,819,1795,1817]" class="Elasmobranchii" family="Potamotrygonidae" genus="Potamotrygon" kingdom="Animalia" order="Myliobatiformes" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="schroederi">
<emphasis id="5A86EAD2FFD5FFBDFDFBF8948E01F898" box="[559,819,1795,1817]" italics="true" pageId="20" pageNumber="21">Potamotrygon schroederi</emphasis>
</taxonomicName>
from Rio Negro, Brazil. A) Dorsal anterior head and snout region (MZUSP 108453). B) Ventral snout and nasoral region (MZUSP 108452). C) Dorsal view of pelvic fin (MZUSP 108452). D) Ventral view of pelvis fins (MZUSP 108455). E) Dorsal view of posterior disc and tail base region in freshly captured adult female specimen (depicted Figure 12A; specimen not preserved). F) Detail of lateral disc color in freshly captured adult female specimen (depicted in Figure 12D; specimen not preserved). Scale bars in A, B, E and F = 5 cm; in C and D = 2.5 cm
</paragraph>
</caption>
<caption id="3C8D6648FFD4FFBCFF43F9E38E9DF95B" httpUri="https://zenodo.org/record/3227632/files/figure.png" pageId="21" pageNumber="22" targetBox="[170,1413,185,1636]" targetPageId="21">
<paragraph id="684D36C0FFD4FFBCFF43F9E38E9DF95B" blockId="21.[151,1436,1651,1754]" pageId="21" pageNumber="22">
<emphasis id="5A86EAD2FFD4FFBCFF43F9E38C2AF908" bold="true" box="[151,280,1651,1674]" pageId="21" pageNumber="22">FIGURE 16</emphasis>
. Dorsal tail base region in specimens of 
<taxonomicName id="AFF24D43FFD4FFBCFD67F9E58E87F90B" box="[691,949,1652,1674]" class="Elasmobranchii" family="Potamotrygonidae" genus="Potamotrygon" kingdom="Animalia" order="Myliobatiformes" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="schroederi">
<emphasis id="5A86EAD2FFD4FFBCFD67F9E58E87F90B" box="[691,949,1652,1674]" italics="true" pageId="21" pageNumber="22">Potamotrygon schroederi</emphasis>
</taxonomicName>
from Rio Negro, Brazil, showing distinct coloration and arrangement of tail spines and denticles. A-B) MZUSP 108453 (689 mm TL, 444 mm DL, 409 mm DW). C-D) MZUSP 108452 (741 mm TL, 444 mm DL, 395 mm DW). E) MZUSP 108445 (501 mm TL, 320 mm DL, 290 mm DW). F) MZUSP 108455 (319 mm TL, 202 mm DL, 188 mm DW). Scale bars = 5 cm.
</paragraph>
</caption>
<subSubSection id="20E8654BFFD4FFB1FF43F8938E38F873" lastPageId="24" lastPageNumber="25" pageId="21" pageNumber="22" type="discussion">
<paragraph id="684D36C0FFD4FFBCFF43F8938827F873" blockId="21.[151,1436,1794,2034]" pageId="21" pageNumber="22">
<emphasis id="5A86EAD2FFD4FFBCFF43F8938C36F89A" bold="true" box="[151,260,1795,1819]" pageId="21" pageNumber="22">Remarks</emphasis>
. There is some confusion regarding the year of publication of this species. The particular issue of the Boletin del Museo de Ciencia Naturales where 
<taxonomicName id="AFF24D43FFD4FFBCFD70F8B88E04F8BE" box="[676,822,1831,1855]" class="Elasmobranchii" family="Potamotrygonidae" genus="Potamotrygon" kingdom="Animalia" order="Myliobatiformes" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="schroederi">
<emphasis id="5A86EAD2FFD4FFBCFD70F8B88E04F8BE" box="[676,822,1831,1855]" italics="true" pageId="21" pageNumber="22">P. schroederi</emphasis>
</taxonomicName>
was described refers to the years 
<date id="1C4C1000FFD4FFBCFB78F8B78863F8BE" box="[1196,1361,1831,1855]" pageId="21" pageNumber="22" value="1956" valueMax="1957">1956 and 1957</date>
on the inside and outside covers, but only in terms of accumulating papers accepted from those years. This same issue has printed on the inside cover “Junio de 1958” (June of 1958), and “Noviembre de 1958” (November of 1958) on the outside cover. In any case, this is indication that this particular issue was published in 1958 and not in 1957. 
<taxonomicName id="AFF24D43FFD4FFBCFA8AF8048CB3F84E" class="Elasmobranchii" family="Potamotrygonidae" genus="Potamotrygon" kingdom="Animalia" order="Myliobatiformes" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="schroederi">
<emphasis id="5A86EAD2FFD4FFBCFA8AF8048CB3F84E" italics="true" pageId="21" pageNumber="22">Potamotrygon schroederi</emphasis>
</taxonomicName>
has been dated from 1957 by almost all previous authors (e.g. 
<bibRefCitation id="0C634B31FFD4FFBCFB9DF82788A5F84E" author="Castex" box="[1097,1431,1975,1999]" pageId="21" pageNumber="22" refString="Castex, M. N. &amp; Yagolkowski, D. R. (1970) Note on two fresh-water sting-rays species (Condrichthyes, Potamotrygonidae): a. Redescription of Potamotrygon schroederi, Yepez 1957. b. Designation of two paratypes of P. pauckei Castex 1963 and their description. Acta Scientifica, Universidad del Salvador, 9, 1 - 26." type="journal article" year="1970">Castex &amp; Yagolkowski, 1970</bibRefCitation>
; 
<bibRefCitation id="0C634B31FFD4FFBCFF43F84B8C2AF873" author="Rosa" box="[151,280,2010,2034]" pageId="21" pageNumber="22" refString="Rosa, R. S. (1985) A systematic revision of the South American freshwater stingrays (Chondrichthyes: Potamotrygonidae). Unpublished PhD diss., College of William and Mary, Williamsburg, xvi + 523 p." type="book" year="1985">Rosa, 1985</bibRefCitation>
; 
<bibRefCitation id="0C634B31FFD4FFBCFEF0F84A8CDAF873" author="Compagno" box="[292,488,2010,2034]" pageId="21" pageNumber="22" refString="Compagno, L. J. V. (1999) Checklist of living Chondrichthyes. Pp. 471 - 498. In: Hamlett, W. C. (Ed.), Sharks, Skates and Rays, the Biology of Elasmobranch Fishes. Baltimore, The John Hopkins Univ. Press, 515 p." type="book chapter" year="1999">Compagno, 1999</bibRefCitation>
, 
<bibRefCitation id="0C634B31FFD4FFBCFE27F84A8F1CF873" author="Compagno" box="[499,558,2010,2034]" pageId="21" pageNumber="22" refString="Compagno, L. J. V. (2005) Checklist of living Chondrichthyes. Pp. 503 - 548. In: Hamlett, W. C. (Ed.), Reproductive Biology and Phylogeny of Chondrichthyes, Sharks, Batoids and Chimaeras. Enfield (NH), Science Publishers, Inc., 562 p." type="book chapter" year="2005">2005</bibRefCitation>
; 
<bibRefCitation id="0C634B31FFD4FFBCFDEDF84A8E14F873" author="Carvalho" box="[569,806,2010,2034]" pageId="21" pageNumber="22" refString="Carvalho, M. R. de, Lovejoy, N. R. &amp; Rosa, R. S. (2003) Family Potamotrygonidae. Pp. 22 - 29. In: Reis, R. E, Ferraris Jr, C. J. &amp; Kullander, S. O. (Eds.). Checklist of the Freshwater Fishes of South and Central America. Porto Alegre, EDIPUCRS, 729 p." type="book chapter" year="2003">Carvalho et al., 2003</bibRefCitation>
; 
<bibRefCitation id="0C634B31FFD4FFBCFCE6F84B890FF873" author="Rosa" box="[818,1085,2010,2034]" pageId="21" pageNumber="22" refString="Rosa, R. S. &amp; Carvalho, M. R. de (2007) Familia Potamotrygonidae. Pp. 17 - 18. In: Buckup, P. A., Menezes, N. A. &amp; Ghazzi, M. S. (Org.). Catalogo das especies de peixes de agua doce do Brasil. Rio de Janeiro, Museu Nacional, 195 p." type="book chapter" year="2007">Rosa &amp; Carvalho, 2007</bibRefCitation>
; 
<bibRefCitation id="0C634B31FFD4FFBCFB9DF84B883BF873" author="Rosa" box="[1097,1289,2010,2034]" pageId="21" pageNumber="22" refString="Rosa, R. S., Charvet-Almeida, P. &amp; Quijada, C. C. D. (2010) Biology of the South American potamotrygonid stingrays. Pp. 241 - 281. In: Carrier, J. C., Musick, J. A. &amp; Heithaus, M. R. (Eds.), Sharks and Their Relativies II: Biodiversity, Adaptative Physiology, and Conservation. CRC Press, Florida, 736 p." type="book chapter" year="2010">Rosa et al., 2010</bibRefCitation>
).
</paragraph>
<caption id="3C8D6648FFD7FFBFFF43FBA98FBBFB0C" httpUri="https://zenodo.org/record/3227634/files/figure.png" pageId="22" pageNumber="23" targetBox="[241,1342,195,1052]" targetPageId="22">
<paragraph id="684D36C0FFD7FFBFFF43FBA98FBBFB0C" blockId="22.[151,1437,1081,1165]" pageId="22" pageNumber="23">
<emphasis id="5A86EAD2FFD7FFBFFF43FBA98C2AFBCE" bold="true" box="[151,280,1081,1104]" pageId="22" pageNumber="23">FIGURE 17</emphasis>
. Skeletal features, dermal denticles and dorsal tail spines of 
<taxonomicName id="AFF24D43FFD7FFBFFCAEFBAB894EFBCE" box="[890,1148,1081,1104]" class="Elasmobranchii" family="Potamotrygonidae" genus="Potamotrygon" kingdom="Animalia" order="Myliobatiformes" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="schroederi">
<emphasis id="5A86EAD2FFD7FFBFFCAEFBAB894EFBCE" box="[890,1148,1081,1104]" italics="true" pageId="22" pageNumber="23">Potamotrygon schroederi</emphasis>
</taxonomicName>
. A) Head, visceral arch, and scapular region. B) Close-up of angular cartilage. C) Base of tail region showing arrangement of dorsal spines and denticles. Based on AUM 44507 (adult male, 780 mm TL).
</paragraph>
</caption>
<paragraph id="684D36C0FFD7FFBFFF13FB29886CFA4A" blockId="22.[151,1437,1209,2020]" pageId="22" pageNumber="23">
The original description of 
<taxonomicName id="AFF24D43FFD7FFBFFE2AFB2B8FA3FB53" box="[510,657,1210,1234]" class="Elasmobranchii" family="Potamotrygonidae" genus="Potamotrygon" kingdom="Animalia" order="Myliobatiformes" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="schroederi">
<emphasis id="5A86EAD2FFD7FFBFFE2AFB2B8FA3FB53" box="[510,657,1210,1234]" italics="true" pageId="22" pageNumber="23">P. schroederi</emphasis>
</taxonomicName>
is very brief and does not provide many morphological details (
<bibRefCitation id="0C634B31FFD7FFBFFAB8FB298CB1FB74" author="Fernandez-Yepez" pageId="22" pageNumber="23" refString="Fernandez-Yepez, A. (1958) Nueva raya para la ciencia: Potomatrygon [sic] schroederi n. sp. Boletin Museo de Ciencia Natural, 2 - 3, 8 - 11." type="book chapter" year="1958">Fernandez-Yépez, 1958</bibRefCitation>
). The description was based on a single very large female specimen (
<quantity id="AF0A9B25FFD7FFBFFB76FB4D8831FB74" box="[1186,1283,1245,1269]" metricMagnitude="-1" metricUnit="m" metricValue="9.95" pageId="22" pageNumber="23" unit="mm" value="995.0">995 mm</quantity>
TL, 
<quantity id="AF0A9B25FFD7FFBFFAE9FB4D88AEFB74" box="[1341,1436,1245,1269]" metricMagnitude="-1" metricUnit="m" metricValue="6.0" pageId="22" pageNumber="23" unit="mm" value="600.0">600 mm</quantity>
DL, 
<quantity id="AF0A9B25FFD7FFBFFF19FA918C1EFA98" box="[205,300,1281,1305]" metricMagnitude="-1" metricUnit="m" metricValue="5.24" pageId="22" pageNumber="23" unit="mm" value="524.0">524 mm</quantity>
DW; 
<tableCitation id="2570037BFFD7FFBFFEA0FA918CFEFA98" box="[372,460,1281,1305]" captionStart="TABLE 2" captionStartId="7.[151,231,655,676]" captionText="TABLE 2. Meristic data for specimens of Potamotrygon tigrina, n. sp. A: MUSM 39978 (holotype), juvenile male, 495 mm TL. B: ANSP 191988 (paratype), preadult female, 560 mm TL." pageId="22" pageNumber="23">Table 2</tableCitation>
), of which three small fragments remain (
<typeStatus id="B7498862FFD7FFBFFC67FA918928FA98" box="[947,1050,1281,1305]" pageId="22" pageNumber="23" type="holotype">holotype</typeStatus>
, MHNLS 2504; 
<figureCitation id="F0C92A45FFD7FFBFFB08FA91880CFA98" box="[1244,1342,1281,1305]" captionStart="FIGURE 9" captionStartId="12.[151,250,969,992]" captionTargetBox="[498,1098,167,951]" captionTargetId="figure@12.[498,1099,167,951]" captionTargetPageId="12" captionText="FIGURE 9. Remains of holotype of Potamotrygon schroederi (MHNLS 2504, Río Apure, Boca Apurito, Orinoco basin). A) Distal tail region with caudal stings. B) Nasoral region. C) Lateral portion of disc. Holotype was originally reported as a very large female (995 mm TL, 600 mm DL, 524 mm DW)." httpUri="https://zenodo.org/record/319977/files/figure.png" pageId="22" pageNumber="23">Figure 9</figureCitation>
). Fortunately, 
<taxonomicName id="AFF24D43FFD7FFBFFF3DFAB68C4CFABC" box="[233,382,1317,1341]" class="Elasmobranchii" family="Potamotrygonidae" genus="Potamotrygon" kingdom="Animalia" order="Myliobatiformes" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="schroederi">
<emphasis id="5A86EAD2FFD7FFBFFF3DFAB68C4CFABC" box="[233,382,1317,1341]" italics="true" pageId="22" pageNumber="23">P. schroederi</emphasis>
</taxonomicName>
is very distinctive, and identifying these remains as 
<taxonomicName id="AFF24D43FFD7FFBFFC0FFAB6895DFABC" box="[987,1135,1317,1341]" class="Elasmobranchii" family="Potamotrygonidae" genus="Potamotrygon" kingdom="Animalia" order="Myliobatiformes" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="schroederi">
<emphasis id="5A86EAD2FFD7FFBFFC0FFAB6895DFABC" box="[987,1135,1317,1341]" italics="true" pageId="22" pageNumber="23">P. schroederi</emphasis>
</taxonomicName>
is unequivocal, especially the distal caudal extremity and outer disc fragments (both have the typical coloration of 
<taxonomicName id="AFF24D43FFD7FFBFFB51FAD9882BFAE1" box="[1157,1305,1352,1376]" class="Elasmobranchii" family="Potamotrygonidae" genus="Potamotrygon" kingdom="Animalia" order="Myliobatiformes" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="schroederi">
<emphasis id="5A86EAD2FFD7FFBFFB51FAD9882BFAE1" box="[1157,1305,1352,1376]" italics="true" pageId="22" pageNumber="23">P. schroederi</emphasis>
</taxonomicName>
). The fragment representing the outer margin of the disc reveals the coloration in the 
<typeStatus id="B7498862FFD7FFBFFC27FAFC8967FA05" box="[1011,1109,1388,1412]" pageId="22" pageNumber="23" type="holotype">holotype</typeStatus>
to be typical of larger, older individuals; a large examined specimen, smaller than the 
<typeStatus id="B7498862FFD7FFBFFCC9FA008EB0FA29" box="[797,898,1424,1448]" pageId="22" pageNumber="23" type="holotype">holotype</typeStatus>
, has a dorsal pattern with rosettes slightly more separated and with less defined margins, with rosettes containing numerous spots (cf. 
<figureCitation id="F0C92A45FFD7FFBFFB8DFA2389F1FA4A" box="[1113,1219,1459,1483]" captionStart="FIGURE 9" captionStartId="12.[151,250,969,992]" captionTargetBox="[498,1098,167,951]" captionTargetId="figure@12.[498,1099,167,951]" captionTargetPageId="12" captionText="FIGURE 9. Remains of holotype of Potamotrygon schroederi (MHNLS 2504, Río Apure, Boca Apurito, Orinoco basin). A) Distal tail region with caudal stings. B) Nasoral region. C) Lateral portion of disc. Holotype was originally reported as a very large female (995 mm TL, 600 mm DL, 524 mm DW)." httpUri="https://zenodo.org/record/319977/files/figure.png" pageId="22" pageNumber="23">Figures 9</figureCitation>
C, 12A, 15E).
</paragraph>
<paragraph id="684D36C0FFD7FFBFFF13FA478FA4F968" blockId="22.[151,1437,1209,2020]" pageId="22" pageNumber="23">
Fernandez-Yépez had briefly characterized 
<taxonomicName id="AFF24D43FFD7FFBFFD16FA488E6BFA6E" box="[706,857,1495,1519]" class="Elasmobranchii" family="Potamotrygonidae" genus="Potamotrygon" kingdom="Animalia" order="Myliobatiformes" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="schroederi">
<emphasis id="5A86EAD2FFD7FFBFFD16FA488E6BFA6E" box="[706,857,1495,1519]" italics="true" pageId="22" pageNumber="23">P. schroederi</emphasis>
</taxonomicName>
, without naming it, in a previous publication (as “raya guacamaya”; 
<bibRefCitation id="0C634B31FFD7FFBFFEA1FA6B8FB0F992" author="Fernandez-Yepez" box="[373,642,1531,1555]" pageId="22" pageNumber="23" refString="Fernandez-Yepez, A. (1949) Uma interesante raya del Rio Orinoco, Venezuela. Contribuiciones Ocasionales de La Coleccion Ictiologica del Laboratorio de Pesqueria Caiguire, 1, 1 - 12." type="journal article" year="1949">Fernandez-Yépez, 1949</bibRefCitation>
) based on a different specimen from the 
<typeStatus id="B7498862FFD7FFBFFB86FA6B8986F992" box="[1106,1204,1531,1555]" pageId="22" pageNumber="23" type="holotype">holotype</typeStatus>
(from Puerto Carreño, Río Orinoco), which constitutes the first record and photograph of this species. In the original description, 
<taxonomicName id="AFF24D43FFD7FFBFFA53F9B08C3CF9DA" class="Elasmobranchii" family="Potamotrygonidae" genus="Potamotrygon" kingdom="Animalia" order="Myliobatiformes" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="schroederi">
<emphasis id="5A86EAD2FFD7FFBFFA53F9B08C3CF9DA" italics="true" pageId="22" pageNumber="23">P. schroederi</emphasis>
</taxonomicName>
is considered as being most similar to 
<emphasis id="5A86EAD2FFD7FFBFFD0EF9D48E77F9DA" box="[730,837,1604,1627]" italics="true" pageId="22" pageNumber="23">P. m o to ro</emphasis>
, contrasting it to 
<taxonomicName id="AFF24D43FFD7FFBFFBCDF9D4884BF9DB" authority="Garman, 1913" authorityName="Garman" authorityYear="1913" box="[1049,1401,1602,1627]" class="Elasmobranchii" family="Potamotrygonidae" genus="Disceus" higherTaxonomySource="CoL" kingdom="Animalia" order="Myliobatiformes" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="thayeri">
<emphasis id="5A86EAD2FFD7FFBFFBCDF9D489F9F9DA" box="[1049,1227,1603,1627]" italics="true" pageId="22" pageNumber="23">Disceus thayeri</emphasis>
<bibRefCitation id="0C634B31FFD7FFBFFB01F9D2884BF9DB" author="Garman" box="[1237,1401,1602,1626]" pageId="22" pageNumber="23" refString="Garman, S. (1913) The Plagiostomia (sharks, skates and rays). Memoirs of the Museum of Comparative Zoology, 36, xiii + 515 p., 77 pls." type="book" year="1913">
<collectingCountry id="10E57650FFD7FFBFFB01F9D28800F9DB" box="[1237,1330,1602,1626]" name="Germany" pageId="22" pageNumber="23">Garman</collectingCountry>
, 1913
</bibRefCitation>
</taxonomicName>
[= 
<taxonomicName id="AFF24D43FFD7FFBFFF43F9F88FB5F9FF" authority="Muller &amp; Henle, 1841" authorityName="Muller &amp; Henle" authorityYear="1841" box="[151,647,1638,1663]" class="Elasmobranchii" family="Potamotrygonidae" genus="Paratrygon" kingdom="Animalia" order="Myliobatiformes" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="aiereba">
<emphasis id="5A86EAD2FFD7FFBFFF43F9F88C46F9FF" box="[151,372,1638,1663]" italics="true" pageId="22" pageNumber="23">Paratrygon aiereba</emphasis>
(Müller &amp; Henle, 1841)
</taxonomicName>
] and “ 
<taxonomicName id="AFF24D43FFD7FFBFFD05F9F88E0AF9FF" box="[721,824,1638,1663]" class="Elasmobranchii" family="Potamotrygonidae" genus="Paratrygon" kingdom="Animalia" order="Myliobatiformes" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="hystrix">
<emphasis id="5A86EAD2FFD7FFBFFD05F9F88E0AF9FF" box="[721,824,1638,1663]" italics="true" pageId="22" pageNumber="23">P. hystrix</emphasis>
</taxonomicName>
” [= 
<taxonomicName id="AFF24D43FFD7FFBFFCB8F9F889D1F9FF" authority="Muller &amp; Henle, 1834" authorityName="Muller &amp; Henle" authorityYear="1834" box="[876,1251,1638,1663]" class="Elasmobranchii" family="Potamotrygonidae" genus="Potamotrygon" kingdom="Animalia" order="Myliobatiformes" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="histrix">
<emphasis id="5A86EAD2FFD7FFBFFCB8F9F88EFCF9FF" box="[876,974,1638,1663]" italics="true" pageId="22" pageNumber="23">P. histrix</emphasis>
(Müller &amp; Henle, 1834)
</taxonomicName>
, a species previously thought to be more widespread in the Amazon basin, but which is restricted to the Paraná-Paraguay system; 
<bibRefCitation id="0C634B31FFD7FFBFFF43F93E8CA2F947" author="Carvalho" box="[151,400,1710,1734]" pageId="22" pageNumber="23" refString="Carvalho, M. R. de, Lovejoy, N. R. &amp; Rosa, R. S. (2003) Family Potamotrygonidae. Pp. 22 - 29. In: Reis, R. E, Ferraris Jr, C. J. &amp; Kullander, S. O. (Eds.). Checklist of the Freshwater Fishes of South and Central America. Porto Alegre, EDIPUCRS, 729 p." type="book chapter" year="2003">Carvalho et al., 2003</bibRefCitation>
; Carvalho, unpubl.]. 
<bibRefCitation id="0C634B31FFD7FFBFFD5DF93E8E96F947" author="Fernandez-Yepez" box="[649,932,1710,1734]" pageId="22" pageNumber="23" refString="Fernandez-Yepez, A. (1958) Nueva raya para la ciencia: Potomatrygon [sic] schroederi n. sp. Boletin Museo de Ciencia Natural, 2 - 3, 8 - 11." type="book chapter" year="1958">Fernandez-Yépez (1958)</bibRefCitation>
characterized 
<taxonomicName id="AFF24D43FFD7FFBFFB9BF93F89D4F947" box="[1103,1254,1710,1734]" class="Elasmobranchii" family="Potamotrygonidae" genus="Potamotrygon" kingdom="Animalia" order="Myliobatiformes" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="schroederi">
<emphasis id="5A86EAD2FFD7FFBFFB9BF93F89D4F947" box="[1103,1254,1710,1734]" italics="true" pageId="22" pageNumber="23">P. schroederi</emphasis>
</taxonomicName>
as lacking oral papillae, but these are present in our material.
</paragraph>
<paragraph id="684D36C0FFD7FFBFFF13F965887FF862" blockId="22.[151,1437,1209,2020]" pageId="22" pageNumber="23">
<bibRefCitation id="0C634B31FFD7FFBFFF13F9658F2EF88C" author="Castex" box="[199,540,1781,1805]" pageId="22" pageNumber="23" refString="Castex, M. N. &amp; Yagolkowski, D. R. (1970) Note on two fresh-water sting-rays species (Condrichthyes, Potamotrygonidae): a. Redescription of Potamotrygon schroederi, Yepez 1957. b. Designation of two paratypes of P. pauckei Castex 1963 and their description. Acta Scientifica, Universidad del Salvador, 9, 1 - 26." type="journal article" year="1970">Castex &amp; Yagolkowski (1970)</bibRefCitation>
extended the distribution of 
<taxonomicName id="AFF24D43FFD7FFBFFCB4F9668EC0F88C" box="[864,1010,1781,1805]" class="Elasmobranchii" family="Potamotrygonidae" genus="Potamotrygon" kingdom="Animalia" order="Myliobatiformes" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="schroederi">
<emphasis id="5A86EAD2FFD7FFBFFCB4F9668EC0F88C" box="[864,1010,1781,1805]" italics="true" pageId="22" pageNumber="23">P. schroederi</emphasis>
</taxonomicName>
to 
<collectingCountry id="10E57650FFD7FFBFFBC3F965896CF88C" box="[1047,1118,1781,1805]" name="Brazil" pageId="22" pageNumber="23">Brazil</collectingCountry>
, citing a large (
<quantity id="AF0A9B25FFD7FFBFFADFF965885BF88C" box="[1291,1385,1781,1805]" metricMagnitude="-1" metricUnit="m" metricValue="7.2" pageId="22" pageNumber="23" unit="mm" value="720.0">720 mm</quantity>
TL) female specimen from Manaus. This specimen, however, most probably arrived in Manaus from the mid to upper Rio Negro. 
<bibRefCitation id="0C634B31FFD7FFBFFEF4F8AD8F4EF8D4" author="Castex" box="[288,636,1853,1877]" pageId="22" pageNumber="23" refString="Castex, M. N. &amp; Yagolkowski, D. R. (1970) Note on two fresh-water sting-rays species (Condrichthyes, Potamotrygonidae): a. Redescription of Potamotrygon schroederi, Yepez 1957. b. Designation of two paratypes of P. pauckei Castex 1963 and their description. Acta Scientifica, Universidad del Salvador, 9, 1 - 26." type="journal article" year="1970">Castex &amp; Yagolkowski (1970)</bibRefCitation>
also cite another specimen from the Orinoco from close to the typelocality, and publish the same figure present in Mago-Leccia’s (1970) account. They stated that 
<taxonomicName id="AFF24D43FFD7FFBFFB2BF8F188A7F8F9" box="[1279,1429,1888,1912]" class="Elasmobranchii" family="Potamotrygonidae" genus="Potamotrygon" kingdom="Animalia" order="Myliobatiformes" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="schroederi">
<emphasis id="5A86EAD2FFD7FFBFFB2BF8F188A7F8F9" box="[1279,1429,1888,1912]" italics="true" pageId="22" pageNumber="23">P. schroederi</emphasis>
</taxonomicName>
, based on coloration, “low disc” and tooth pattern (among other ambiguous features), can be placed in a group with 
<taxonomicName id="AFF24D43FFD7FFBFFF43F8398CD7F841" authority="Gunther, 1880" authorityName="Gunther" authorityYear="1880" box="[151,485,1960,1984]" class="Elasmobranchii" family="Potamotrygonidae" genus="Paratrygon" kingdom="Animalia" order="Myliobatiformes" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="brachyura">
<emphasis id="5A86EAD2FFD7FFBFFF43F8398C15F841" box="[151,295,1960,1984]" italics="true" pageId="22" pageNumber="23">P. brachyura</emphasis>
(Günther, 1880)
</taxonomicName>
, 
<taxonomicName id="AFF24D43FFD7FFBFFE25F8398F56F841" box="[497,612,1960,1984]" class="Elasmobranchii" family="Potamotrygonidae" genus="Potamotrygon" kingdom="Animalia" order="Myliobatiformes" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="falkneri">
<emphasis id="5A86EAD2FFD7FFBFFE25F8398F56F841" box="[497,612,1960,1984]" italics="true" pageId="22" pageNumber="23">P. falkneri</emphasis>
</taxonomicName>
and 
<taxonomicName id="AFF24D43FFD7FFBFFD48F8398E03F841" box="[668,817,1960,1984]" class="Elasmobranchii" family="Potamotrygonidae" genus="Potamotrygon" kingdom="Animalia" order="Myliobatiformes" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="menchacai">
<emphasis id="5A86EAD2FFD7FFBFFD48F8398E03F841" box="[668,817,1960,1984]" italics="true" pageId="22" pageNumber="23">P. menchacai</emphasis>
</taxonomicName>
. Even though their redescription did not provide many informative details, at least specimens from the Orinoco and Negro basins were compared for the first time.
</paragraph>
<caption id="3C8D6648FFD6FFBEFF43FBB38DC1FBF6" httpUri="https://zenodo.org/record/320011/files/figure.png" pageId="23" pageNumber="24" targetBox="[288,1298,182,1040]" targetPageId="23">
<paragraph id="684D36C0FFD6FFBEFF43FBB38DC1FBF6" blockId="23.[151,1436,1059,1143]" pageId="23" pageNumber="24">
<emphasis id="5A86EAD2FFD6FFBEFF43FBB38C2FFBB9" bold="true" box="[151,285,1059,1082]" pageId="23" pageNumber="24">FIGURE 18</emphasis>
. Morphological details of distal tail region in 
<taxonomicName id="AFF24D43FFD6FFBEFCC6FBB4892AFBB8" box="[786,1048,1059,1081]" class="Elasmobranchii" family="Potamotrygonidae" genus="Potamotrygon" kingdom="Animalia" order="Myliobatiformes" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="schroederi">
<emphasis id="5A86EAD2FFD6FFBEFCC6FBB4892AFBB8" box="[786,1048,1059,1081]" italics="true" pageId="23" pageNumber="24">Potamotrygon schroederi</emphasis>
</taxonomicName>
, showing arrangement of dorsal tail spines, denticles, caudal stings, and vertebrae. A) AUM 44507 (adult male, 780 mm TL). B) ANSP 191999 (adult female, 605 mm TL).
</paragraph>
</caption>
<paragraph id="684D36C0FFD6FFBEFF13FB338803FAEF" blockId="23.[151,1437,1187,2034]" pageId="23" pageNumber="24">
The only more or less consistent distinction, based on our material, between populations of 
<taxonomicName id="AFF24D43FFD6FFBEFB18FB35886CFB3D" box="[1228,1374,1188,1212]" class="Elasmobranchii" family="Potamotrygonidae" genus="Potamotrygon" kingdom="Animalia" order="Myliobatiformes" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="schroederi">
<emphasis id="5A86EAD2FFD6FFBEFB18FB35886CFB3D" box="[1228,1374,1188,1212]" italics="true" pageId="23" pageNumber="24">P. schroederi</emphasis>
</taxonomicName>
from the Rio Negro and Río Orinoco is in numbers of vertebral centra. Total vertebral centra in the two specimens from the Orinoco that were radiographed are clearly higher (149–152) than in the Rio Negro material (106–133), reflecting a much higher number of caudal centra (122–124 vs. 81–104, respectively). But our radiographs of material from the Orinoco were much more clear and easy to interpret, which could be partially responsible for such a significant difference. Also more material from 
<collectingCountry id="10E57650FFD6FFBEFD5BFAC68E30FAEF" box="[655,770,1366,1390]" name="Venezuela" pageId="23" pageNumber="24">Venezuela</collectingCountry>
should be examined to corroborate this character.
</paragraph>
<paragraph id="684D36C0FFD6FFBEFF13FAEA8C12F8E3" blockId="23.[151,1437,1187,2034]" pageId="23" pageNumber="24">
Another difference, but more slight, occurs in dorsal coloration. Freshly collected specimens from both Rio Negro and Río Orinoco have a strong blackish-brown or purplish-black background color, and the rosettes and other dorsal ornamentation are also similar in intensity and brightness. But material from Rio Negro have relatively smaller and more numerous rosettes and brain-shaped vermiculate figures on dorsal disc region, especially on outermost rows closer to disc margin and on middisc. The specific patterns formed by these vermiculate figures also appear slightly distinct. In material from the Orinoco, the rosettes, as well as being slightly larger and fewer (especially on middisc region, close to midline), are more complex and invaginated (
<figureCitation id="F0C92A45FFD6FFBEFBC6F9C089BFF9E9" box="[1042,1165,1616,1640]" captionStart="FIGURE 10" captionStartId="16.[151,250,1238,1261]" captionTargetBox="[151,1436,176,1224]" captionTargetId="figure@16.[151,1436,174,1225]" captionTargetPageId="16" captionText="FIGURE 10. Freshly captured specimens of Potamotrygon schroederi from Venezuela (Orinoco basin). A) Dorsal view of AUM 44507 (Río Orinoco, island west of Puerto Venado, adult male, 780 mm TL, 470 mm DL, 420 mm DW). B) Dorsal view of ANSP 191999 (Río Ventuari, adult female, 605 mm TL, 395 mm DL, 358 mm DW)." httpUri="https://zenodo.org/record/3227628/files/figure.png" pageId="23" pageNumber="24">Figures 10</figureCitation>
, 
<figureCitation id="F0C92A45FFD6FFBEFB4EF9C0898AF9E9" box="[1178,1208,1616,1640]" captionStart="FIGURE 12" captionStartId="18.[151,250,1952,1975]" captionTargetBox="[256,1326,183,1920]" captionTargetId="figure@18.[256,1327,183,1922]" captionTargetPageId="18" captionText="FIGURE 12. Dorsal views of freshly captured specimens of Potamotrygon schroederi from Rio Negro, near Barcelos, Brazil. A, B and D discarded after problems with temporary storage in Manaus. C) MZUSP 108453. Photographs by M. V. Domingues and N. Luchetti." httpUri="https://zenodo.org/record/319985/files/figure.png" pageId="23" pageNumber="24">12</figureCitation>
, 
<figureCitation id="F0C92A45FFD6FFBEFB10F9C089D3F9E9" box="[1220,1249,1616,1640]" captionStart="FIGURE 19" captionStartId="24.[151,250,1558,1581]" captionTargetBox="[256,1327,406,1540]" captionTargetId="figure@24.[256,1327,406,1540]" captionTargetPageId="24" captionText="FIGURE 19. Comparison of left anterolateral disc region in Potamotrygon schroederi among specimens from Orinoco (A and C, left column) and Negro basins (B and D, on right). A) AUM 44507. B) ANSP 191999. C) MZUSP 108452. D) MZUSP 108453. Scale bar = 5 cm." httpUri="https://zenodo.org/record/320012/files/figure.png" pageId="23" pageNumber="24">19</figureCitation>
) (note that these distinctions are difficult to quantify as counting individual rosettes is not straightforward). In material from the Orinoco, many rosettes and brain-shaped figures are clearly greater in diameter than interorbital space, whereas in specimens from Rio Negro, usually just a few of these figures, at the most, will surpass interorbital space in width. We have verified these patterns on specimens of similar dimensions and sex. Even though we have included here only two collected specimens from the Orinoco, we have seen others (also 
<bibRefCitation id="0C634B31FFD6FFBEFC27F8938876F89A" author="Castex" box="[1011,1348,1795,1819]" pageId="23" pageNumber="24" refString="Castex, M. N. &amp; Yagolkowski, D. R. (1970) Note on two fresh-water sting-rays species (Condrichthyes, Potamotrygonidae): a. Redescription of Potamotrygon schroederi, Yepez 1957. b. Designation of two paratypes of P. pauckei Castex 1963 and their description. Acta Scientifica, Universidad del Salvador, 9, 1 - 26." type="journal article" year="1970">Castex &amp; Yagolkowski, 1970</bibRefCitation>
; Mago- 
<bibRefCitation id="0C634B31FFD6FFBEFF43F8B78C18F8BE" author="Leccia" box="[151,298,1831,1855]" pageId="23" pageNumber="24" refString="Mago-Leccia, F. (1970) Lista de los peces de Venezuela, incluyendo un estudio preliminar de la ictiogeografia del pais. Ministerio de Agricultura y Cria, Caracas, 285 p." type="book" year="1970">Leccia, 1970</bibRefCitation>
; numerous specimens in aquarium literature). Clearly, though, more material is needed to verify these distinctions.
</paragraph>
<paragraph id="684D36C0FFD6FFB1FF13F8FF894AFF76" blockId="23.[151,1437,1187,2034]" lastBlockId="24.[151,1436,151,355]" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24">
Even though 
<taxonomicName id="AFF24D43FFD6FFBEFE8BF8FF8CC1F807" box="[351,499,1902,1926]" class="Elasmobranchii" family="Potamotrygonidae" genus="Potamotrygon" kingdom="Animalia" order="Myliobatiformes" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="schroederi">
<emphasis id="5A86EAD2FFD6FFBEFE8BF8FF8CC1F807" box="[351,499,1902,1926]" italics="true" pageId="23" pageNumber="24">P. schroederi</emphasis>
</taxonomicName>
is known from two major river basins (Negro and Orinoco) spanning a large geographical area, this species can be easily identified from congeners; however, the distinctions among populations of both basins outlined above are either relatively minor or require further corroboration, and at present do not warrant specific separation. Both rivers are interconnected by the Río Casiquiare in 
<collectingCountry id="10E57650FFD6FFBEFC20F8498955F870" box="[1012,1127,2009,2033]" name="Venezuela" pageId="23" pageNumber="24">Venezuela</collectingCountry>
(for a recent summary, see 
<bibRefCitation id="0C634B31FFD9FFB1FF43FF078CAFFF2E" author="Winemiller" box="[151,413,151,175]" pageId="24" pageNumber="25" refString="Winemiller, K. O., Lopez-Fernandez, H., Taphorn, D. C., Nico, L. G. &amp; Duque, A. B. (2008) Fish assemblages of the Casiquiare River, a corridor and zoogeographical filter for dispersal between the Orinoco and Amazon basins. Journal of Biogeography, 35 (9), 1551 - 1563." type="journal article" year="2008">Winemiller et al., 2008</bibRefCitation>
), but tagging studies are needed to verify if specimens traverse from one basin to the other, or if 
<taxonomicName id="AFF24D43FFD9FFB1FF1AFF2D8C53FF55" box="[206,353,188,212]" class="Elasmobranchii" family="Potamotrygonidae" genus="Potamotrygon" kingdom="Animalia" order="Myliobatiformes" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="schroederi">
<emphasis id="5A86EAD2FFD9FFB1FF1AFF2D8C53FF55" box="[206,353,188,212]" italics="true" pageId="24" pageNumber="25">P. schroederi</emphasis>
</taxonomicName>
occurs without significant discontinuities between both basins; in any case this biogeographical link explains the historical distribution of 
<taxonomicName id="AFF24D43FFD9FFB1FDB8FF708FCDFF76" box="[620,767,223,247]" class="Elasmobranchii" family="Potamotrygonidae" genus="Potamotrygon" kingdom="Animalia" order="Myliobatiformes" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="schroederi">
<emphasis id="5A86EAD2FFD9FFB1FDB8FF708FCDFF76" box="[620,767,223,247]" italics="true" pageId="24" pageNumber="25">P. schroederi</emphasis>
</taxonomicName>
in the Negro and Orinoco basins.
</paragraph>
<paragraph id="684D36C0FFD9FFB1FF13FE938914FEE3" blockId="24.[151,1436,151,355]" pageId="24" pageNumber="25">
According to 
<bibRefCitation id="0C634B31FFD9FFB1FEB6FE938F37FE9A" author="Araujo" box="[354,517,259,283]" pageId="24" pageNumber="25" refString="Araujo, M. L. G. de (2004) Potamotrygon schroederi. In: IUCN 2010. IUCN Red List of Threatened Species. Version 2010.4. www. iucnredlist. org (downloaded on 20 March 2011)." type="journal article" year="2004">Araújo (2004)</bibRefCitation>
, specimens of 
<taxonomicName id="AFF24D43FFD9FFB1FD78FE948E72FE9A" box="[684,832,259,283]" class="Elasmobranchii" family="Potamotrygonidae" genus="Potamotrygon" kingdom="Animalia" order="Myliobatiformes" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="schroederi">
<emphasis id="5A86EAD2FFD9FFB1FD78FE948E72FE9A" box="[684,832,259,283]" italics="true" pageId="24" pageNumber="25">P. schroederi</emphasis>
</taxonomicName>
are born at 
<quantity id="AF0A9B25FFD9FFB1FC1CFE938914FE9A" box="[968,1062,259,283]" metricMagnitude="-1" metricUnit="m" metricValue="1.4" pageId="24" pageNumber="25" unit="mm" value="140.0">140 mm</quantity>
DW, mature sexually at 
<quantity id="AF0A9B25FFD9FFB1FAEBFE9388AEFE9A" box="[1343,1436,259,283]" metricMagnitude="-1" metricUnit="m" metricValue="4.2" pageId="24" pageNumber="25" unit="mm" value="420.0">420 mm</quantity>
DW (males) and 
<quantity id="AF0A9B25FFD9FFB1FE8BFEB78C8DFEBE" box="[351,447,295,319]" metricMagnitude="-1" metricUnit="m" metricValue="4.4" pageId="24" pageNumber="25" unit="mm" value="440.0">440 mm</quantity>
DW (females), and attain a maximum size of 
<quantity id="AF0A9B25FFD9FFB1FC08FEB7890EFEBE" box="[988,1084,295,319]" metricMagnitude="-1" metricUnit="m" metricValue="5.4" pageId="24" pageNumber="25" unit="mm" value="540.0">540 mm</quantity>
DW (dimensions similar to 
<taxonomicName id="AFF24D43FFD9FFB1FA53FEB88DD1FEE2" class="Elasmobranchii" family="Potamotrygonidae" genus="Potamotrygon" kingdom="Animalia" order="Myliobatiformes" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="tigrina">
<emphasis id="5A86EAD2FFD9FFB1FA53FEB88DD1FEE2" italics="true" pageId="24" pageNumber="25">P. tigrina</emphasis>
</taxonomicName>
). This species is commonly exported by the aquarium trade from Manaus.
</paragraph>
<caption id="3C8D6648FFD9FFB1FF43F9868C93F9E8" httpUri="https://zenodo.org/record/320012/files/figure.png" pageId="24" pageNumber="25" targetBox="[256,1327,406,1540]" targetPageId="24">
<paragraph id="684D36C0FFD9FFB1FF43F9868C93F9E8" blockId="24.[151,1436,1558,1641]" pageId="24" pageNumber="25">
<emphasis id="5A86EAD2FFD9FFB1FF43F9868C2BF9AA" bold="true" box="[151,281,1558,1581]" pageId="24" pageNumber="25">FIGURE 19</emphasis>
. Comparison of left anterolateral disc region in 
<taxonomicName id="AFF24D43FFD9FFB1FCD5F9878931F9AD" box="[769,1027,1558,1580]" class="Elasmobranchii" family="Potamotrygonidae" genus="Potamotrygon" kingdom="Animalia" order="Myliobatiformes" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="schroederi">
<emphasis id="5A86EAD2FFD9FFB1FCD5F9878931F9AD" box="[769,1027,1558,1580]" italics="true" pageId="24" pageNumber="25">Potamotrygon schroederi</emphasis>
</taxonomicName>
among specimens from Orinoco (A and C, left column) and Negro basins (B and D, on right). A) AUM 44507. B) ANSP 191999. C) MZUSP 108452. D) MZUSP 108453. Scale bar = 5 cm.
</paragraph>
</caption>
<paragraph id="684D36C0FFD9FFB1FF43F9058C3BF82B" blockId="24.[151,1436,1685,1962]" pageId="24" pageNumber="25">
<emphasis id="5A86EAD2FFD9FFB1FF43F9058C85F92F" bold="true" box="[151,439,1685,1710]" pageId="24" pageNumber="25">Geographic distribution</emphasis>
. 
<taxonomicName id="AFF24D43FFD9FFB1FE11F9078FD1F92F" box="[453,739,1686,1710]" class="Elasmobranchii" family="Potamotrygonidae" genus="Potamotrygon" kingdom="Animalia" order="Myliobatiformes" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="schroederi">
<emphasis id="5A86EAD2FFD9FFB1FE11F9078FD1F92F" box="[453,739,1686,1710]" italics="true" pageId="24" pageNumber="25">Potamotrygon schroederi</emphasis>
</taxonomicName>
is distributed in the mid to upper Rio Negro and mid to upper Río Orinoco basins (
<figureCitation id="F0C92A45FFD9FFB1FE56F92A8CD7F953" box="[386,485,1722,1746]" captionStart="FIGURE 8" captionStartId="11.[151,250,1068,1091]" captionTargetBox="[159,1431,159,1057]" captionTargetId="figure@11.[159,1432,159,1058]" captionTargetPageId="11" captionText="FIGURE 8. Río Nanay at Pampa Chica, large sandy beach along left bank of hairpin curve in river (flowing towards right), approximately 11 river km upstream from confluence with Río Amazonas and 4 km west of Iquitos center. Photo taken August 18, 2003. Large sandy beach where ANSP 182452 was collected in 2005 is largely submerged. Photograph by M. Sabaj Pérez." httpUri="https://zenodo.org/record/3227626/files/figure.png" pageId="24" pageNumber="25">Figure 8</figureCitation>
). This species appears to prefer faster flowing waters of central river channels, and has not been captured in the lower Rio Negro (M. L. G. de Araújo, pers. comm.). Specimens from the upper Río Ventuari (ANSP 191999) and Río Orinoco (AUM 44507) were collected at night in a swift current (under 
<quantity id="AF0A9B25FFD9FFB1FABAF89288AEF89B" box="[1390,1436,1794,1818]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="24" pageNumber="25" unit="m" value="2.0">2 m</quantity>
deep), over a large beach of coarse sand. This species probably also penetrates the region of the Río Casiquiare interchange between the Negro and Orinoco basins. Reported also from the Orinoco basin of 
<collectingCountry id="10E57650FFD9FFB1FB65F8DA8810F8E3" box="[1201,1314,1866,1890]" name="Colombia" pageId="24" pageNumber="25">Colombia</collectingCountry>
, but specimens were not examined by us and are not illustrated in accounts (
<bibRefCitation id="0C634B31FFD9FFB1FC55F8FE89BFF807" author="Mejia-Falla" box="[897,1165,1902,1926]" pageId="24" pageNumber="25" refString="Mejia-Falla, P. A., Navia, A. F., Mejia-Ladino, L. M., Acero, A. &amp; Rubio, E. A. (2007) Tiburones y rayas de Colombia (Pisces Elasmobranchii): lista actualizada, revisada y comentada. Boletin de Investigaciones Marinas y Costeras, 36, 111 - 149." type="journal article" year="2007">Mejía-Falla et al., 2007</bibRefCitation>
; 
<bibRefCitation id="0C634B31FFD9FFB1FB4DF8FE8DCFF82B" author="Maldonado-Ocampo" pageId="24" pageNumber="25" refString="Maldonado-Ocampo, J. A., Vari, R. P. &amp; Usma, J. S. (2008) Checklist of the freshwater fishes of Colombia. Biota Colombiana 9 (2), 143 - 237." type="journal article" year="2008">Maldonado-Ocampo et al., 2008</bibRefCitation>
).
</paragraph>
<paragraph id="684D36C0FFD9FFB1FF43F8498E38F873" blockId="24.[151,778,2009,2035]" box="[151,778,2009,2035]" pageId="24" pageNumber="25">
<emphasis id="5A86EAD2FFD9FFB1FF43F8498C7FF873" bold="true" box="[151,333,2009,2034]" pageId="24" pageNumber="25">Common name</emphasis>
. Flower ray (from aquarium literature).
</paragraph>
</subSubSection>
</treatment>
</document>