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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.271.4062" ID-GBIF-Dataset="c93c3a93-d2e1-4c85-8ece-bf3aa1a5109d" ID-PMC="PMC3652427" ID-Pensoft-Pub="1313-2970-271-1" ID-PubMed="23717185" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1313-2970-271-1" ModsDocOrigin="ZooKeys 271" ModsDocTitle="A systematic revision of Operclipygus Marseul (Coleoptera, Histeridae, Exosternini)" checkinTime="1451247655320" checkinUser="pensoft" docAuthor="Caterino, Michael S. &amp; Tishechkin, Alexey K." docDate="2013" docId="E0161EE25F2B1476DB0C9B7A97B558C1" docLanguage="en" docName="ZooKeys 271: 1-401" docOrigin="ZooKeys 271" docSource="http://dx.doi.org/10.3897/zookeys.271.4062" docTitle="Operclipygus impositus Caterino &amp; Tishechkin, 2013, sp. n." docType="treatment" docVersion="4" lastPageNumber="167" masterDocId="4403FF80FFE7FF95FFDA5D0E29259B2F" masterDocTitle="A systematic revision of Operclipygus Marseul (Coleoptera, Histeridae, Exosternini)" masterLastPageNumber="401" masterPageNumber="1" pageNumber="165" updateTime="1668155382655" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A systematic revision of Operclipygus Marseul (Coleoptera, Histeridae, Exosternini)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Caterino, Michael S.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tishechkin, Alexey K.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2013</mods:date>
<mods:detail type="volume">
<mods:number>271</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>401</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.271.4062</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.271.4062</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-271-1</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152040915" LSID="urn:lsid:zoobank.org:act:DECF9B78-D51C-4B51-9CDC-1FDBDC5EBE1A" httpUri="http://treatment.plazi.org/id/E0161EE25F2B1476DB0C9B7A97B558C1" lastPageId="166" lastPageNumber="167" pageId="164" pageNumber="165">
<subSubSection pageId="164" pageNumber="165" type="nomenclature">
<paragraph pageId="164" pageNumber="165">
<taxonomicName LSID="urn:lsid:zoobank.org:act:DECF9B78-D51C-4B51-9CDC-1FDBDC5EBE1A" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus impositus" order="Coleoptera" pageId="164" pageNumber="165" phylum="Arthropoda" rank="species" species="impositus">Operclipygus impositus</taxonomicName>
<taxonomicNameLabel pageId="164" pageNumber="165">sp. n.</taxonomicNameLabel>
Figs 44
<normalizedToken originalValue="AE45ABMap">A-E45A-BMap</normalizedToken>
16
</paragraph>
</subSubSection>
<subSubSection pageId="164" pageNumber="165" type="type locality">
<paragraph pageId="164" pageNumber="165">Type locality.</paragraph>
<paragraph pageId="164" pageNumber="165">
BRAZIL:
<normalizedToken originalValue="Pará">Para</normalizedToken>
: km 18 Altamira -
<normalizedToken originalValue="Marabá">Maraba</normalizedToken>
road [
<geoCoordinate direction="south" orientation="latitude" precision="925" value="-3.15">3°09'S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-52.05">52°03'W</geoCoordinate>
[
</paragraph>
</subSubSection>
<subSubSection pageId="164" pageNumber="165" type="type material">
<paragraph pageId="164" pageNumber="165">Type material.</paragraph>
<paragraph pageId="164" pageNumber="165">
Holotype male: &quot;BRASIL:
<normalizedToken originalValue="Pará">Para</normalizedToken>
: Altamira -
<normalizedToken originalValue="Marabá">Maraba</normalizedToken>
: km 18,
<geoCoordinate direction="south" orientation="latitude" precision="925" value="-3.15">3°09'S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-52.05">52°03'W</geoCoordinate>
.
<normalizedToken originalValue="Piège">Piege</normalizedToken>
<normalizedToken originalValue="deinterception">de'interception</normalizedToken>
. v.1984&quot; / &quot;Caterino/Tishechkin
<taxonomicName lsidName="" pageId="164" pageNumber="165" rank="tribe" tribe="Exosternini">Exosternini</taxonomicName>
Voucher EXO-00307&quot; (UFPR). Paratypes (23): 1: same data as type(CHND); BRAZIL:
<normalizedToken originalValue="Amapá">Amapa</normalizedToken>
: 3: Serra do Navio,
<geoCoordinate direction="north" orientation="latitude" precision="925" value="0.98333335">0°59'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-52.0">52°00'W</geoCoordinate>
, 1-14.v.1991, FIT (CHND, MNHN); Mato Grosso: 1: Sinop,
<geoCoordinate direction="south" orientation="latitude" precision="925" value="-11.816667">11°49'S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-55.483334">55°29'W</geoCoordinate>
, 12-24.vi.1985, FIT (CHND);
<normalizedToken originalValue="Pará">Para</normalizedToken>
: 3:
<normalizedToken originalValue="Tucuruí">Tucurui</normalizedToken>
,
<geoCoordinate direction="south" orientation="latitude" precision="925" value="-3.75">3°45'S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-49.666668">49°40'W</geoCoordinate>
, 20.
<normalizedToken originalValue="v">v-</normalizedToken>
15.vi.1987, FIT (CHND, FMNH), 1: iv.1986, FIT (CHND), 1: IPEAN, Utinga,
<normalizedToken originalValue="Belém">Belem</normalizedToken>
,
<geoCoordinate direction="south" orientation="latitude" precision="925" value="-1.45">1°27'S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-48.433334">48°26'W</geoCoordinate>
, v.1985, FIT (CHND), 2: viii.1985, FIT (CHND), 1: x.1985, FIT (CHND), 2: xi.1984, FIT (MSCC, CHND); 3:
<normalizedToken originalValue="Carajás">Carajas</normalizedToken>
, Serra Norte,
<geoCoordinate direction="south" orientation="latitude" precision="925" value="-6.0666666">6°04'S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-50.2">50°12'W</geoCoordinate>
, 13.xi-2.xii.1987, FIT (CHND, UFPR, AKTC); 5: Marajo-Breves,
<geoCoordinate direction="south" orientation="latitude" precision="925" value="-0.8833333">0°53'S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-50.533333">50°32'W</geoCoordinate>
, 18.xi-5.xii.1987, FIT (CHND, UFPR).
</paragraph>
</subSubSection>
<subSubSection lastPageId="165" lastPageNumber="166" pageId="164" pageNumber="165" type="other material">
<paragraph pageId="164" pageNumber="165">Other material</paragraph>
<paragraph lastPageId="165" lastPageNumber="166" pageId="164" pageNumber="165">
. COLOMBIA:
<normalizedToken originalValue="Vaupés">Vaupes</normalizedToken>
: 1: Parque Nac. Mosiro-Itajura (
<normalizedToken originalValue="Caparú">Caparu</normalizedToken>
), Centro Ambiental,
<geoCoordinate direction="south" orientation="latitude" precision="925" value="-1.0666667">1°04'S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-69.51667">69°31'W</geoCoordinate>
, 60m, 20-30.i.2003, FIT, D. Arias &amp; M. Sharkey (IAVH). ECUADOR: Orellana: 1:Res. Ethnica Waorani, 1km S Onkone Gare Camp, Trans. Ent.,
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-0.65277773">0°39'10&quot;S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-76.433334">76°26'W</geoCoordinate>
, 220m, 16.i.1994, fogging, T.L. Erwin (USNM); 1:
<normalizedToken originalValue="Yasuní">Yasuni</normalizedToken>
Res. Stn., 215m, 5-10.ix.1999, FIT, primary forest, E.G. Riley (TAMU); 1: Tiputini Biodiversity Station,
<geoCoordinate direction="south" orientation="latitude" precision="5" value="-0.6376">0.6376°S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="5" value="-76.1499">76.1499°W</geoCoordinate>
, 4-9.vi.2011, FIT, M.S. Ca
<pageBreakToken pageId="165" pageNumber="166" start="start">terino</pageBreakToken>
&amp; A.K. Tishechkin, DNA Extract MSC-2190 (SBMNH); 1: 5-25.ix.2000, D.J. Inward &amp; K.A. Jackson (BMNH);
<normalizedToken originalValue="Sucumbíos">Sucumbios</normalizedToken>
: 2: Sacha Lodge,
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-0.47055554">0°28'14&quot;S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-76.45972">76°27'35&quot;W</geoCoordinate>
, 270m, 21-24.iii.1999, FIT, R. Brooks (SEMC). FRENCH GUIANA: 1: Roura, 18.4km SSE,
<geoCoordinate direction="north" orientation="latitude" precision="15" value="4.6105556">4°36'38&quot;N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-52.22361">52°13'25&quot;W</geoCoordinate>
, 240m, 29.
<normalizedToken originalValue="v">v-</normalizedToken>
10.vi.1997, FIT, J. Ashe, R. Brooks (SEMC); 2:
<normalizedToken originalValue="Saül">Sauel</normalizedToken>
, 7km N, Les Eaux Claires,
<geoCoordinate direction="north" orientation="latitude" precision="15" value="3.662778">3°39'46&quot;N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-53.221947">53°13'19&quot;W</geoCoordinate>
, 220m, 31.
<normalizedToken originalValue="v">v-</normalizedToken>
3.vi.1997, FIT, J. Ashe, R. Brooks (SEMC); 1:
<normalizedToken originalValue="Belvèdére">Belvedere</normalizedToken>
de
<normalizedToken originalValue="Saül">Sauel</normalizedToken>
, point de vue,
<geoCoordinate direction="north" orientation="latitude" precision="15" value="3.0227778">3°1'22&quot;N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-53.209446">53°12'34&quot;W</geoCoordinate>
, 20.xii.2010, Window trap, SEAG (MNHN); 1:
<normalizedToken originalValue="Rés">Res</normalizedToken>
. Natur. des Nouragues, Camp Inselberg,
<geoCoordinate direction="north" orientation="latitude" precision="925" value="4.0833335">4°05'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-52.683334">52°41'W</geoCoordinate>
, 25.i.2011, Window trap, SEAG (MNHN), 1: 9.xi.2010, Window trap, SEAG (MNHN). GUYANA: Mazaruni Potaro: 1: Takutu Mountains,
<geoCoordinate direction="north" orientation="latitude" precision="925" value="6.25">6°15'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-59.083332">59°5'W</geoCoordinate>
, 11.xii.1983, Window trap, montane rainforest near logging area, P.J. Spangler, R.A. Faitoute &amp; W.E. Steiner (USNM); Region 8: 1: Iwokrama Forest, Turtle Mt. summit,
<geoCoordinate direction="north" orientation="latitude" precision="15" value="4.7325">4°43'57&quot;N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-58.733612">58°44'1&quot;W</geoCoordinate>
, 290m, 30.
<normalizedToken originalValue="v">v-</normalizedToken>
1.vi.2001, FIT, R. Brooks &amp; Z. Falin (SEMC). PERU:
<normalizedToken originalValue="Junín">Junin</normalizedToken>
: 1: 11km NE Puerto Ocopa, Los Olivos,
<geoCoordinate direction="south" orientation="latitude" precision="9" value="-11.116667">11°7.00'S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="9" value="-74.25867">74°15.52'W</geoCoordinate>
, 1200m, 24-26.iii.2009, Window trap, A.V. Petrov (AKTC), 1: 26-28.iii.2009, Window trap, A.V. Petrov (AKTC), 1: 30-31.iii.2009, Window trap, A.V. Petrov (MUSM); Loreto: 1: 68km SW Iquitos to Nauta, Rio Itaya,
<geoCoordinate direction="south" orientation="latitude" precision="925" value="-4.1833334">4°11'S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-73.433334">73°26'W</geoCoordinate>
, 110m, 28.ii.2008, A.V. Petrov (AKTC), 1: 1-3.iii.2008, A.V. Petrov (MUSM). SURINAME: Brokopondo: 1: Brownsberg Nature Preserve, Witi Creek Trail,
<geoCoordinate direction="north" orientation="latitude" precision="15" value="4.9486113">4°56'55&quot;N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-55.18139">55°10'53&quot;W</geoCoordinate>
, 480m, 23-25.vi.1999, FIT, Z. Falin, A. Gangadin, H. Hiwat (SEMC); Marowijne: 1: Nassau Mountain,
<geoCoordinate direction="north" orientation="latitude" precision="15" value="4.8100004">4°48'36&quot;N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-54.52111">54°31'16&quot;W</geoCoordinate>
, 500m, 2-4.vi.1999, FIT, Z. Falin, B. DeDijn (SEMC); Para: 1: nr. Overbridge River Resort,
<geoCoordinate direction="north" orientation="latitude" precision="92" value="5.5299997">5°31.8'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-55.058334">55°3.5'W</geoCoordinate>
, 15-18.ii.2010, FIT, C. Gillet, P. Skelley, W. Warner (FSCA).
</paragraph>
</subSubSection>
<subSubSection lastPageId="166" lastPageNumber="167" pageId="165" pageNumber="166" type="description">
<paragraph pageId="165" pageNumber="166">Diagnostic description.</paragraph>
<paragraph lastPageId="166" lastPageNumber="167" pageId="165" pageNumber="166">
Length: 1.22-1.40 mm, width: 0.84-0.97 mm; body rufobrunneus, elongate, sides subparallel, depressed; frons weakly depressed at middle, frontal stria complete, sinuate, slightly recurved at middle; supraorbital stria complete, meeting sides of frontal stria; epistoma broad, sides rounded, emarginate across anterior margin; labrum half as long as wide, straight across apical margin; left and right mandibles both with acute basal tooth; pronotal disk with linear prescutellar impression about 1.5
<normalizedToken originalValue="×">x</normalizedToken>
as long as scutellum; lateral submarginal pronotal stria absent, anterior marginal stria interrupted behind head; anterior submarginal stria strongly recurved posterad, reaching pronotal midpoint; median pronotal gland openings along side base of recurved arms of anterior stria, about 4 puncture widths from anterior margin; elytra with 1 complete epipleural stria, outer subhumeral stria present in apical half, inner subhumeral stria absent, striae 1-4 complete, 5th stria present in apical half, sutural stria present in apical three-fourths; prosternal keel with base produced, rounded, carinal striae long, close and subparallel in anterior three-fourths, united in narrow anterior arch, separate basally; mesoventrite emarginate anteriorly, marginal stria interrupted; mesometaventral stria narrowly arched forward to near margin, sinuate laterally, continued by lateral metaventral stria to near middle of metacoxa; 1st abdominal ventrite with two complete lateral striae; propygidium (Fig. 45B) with sparse fine ground punctation, small round coarser punctures uniformly separated by about their diameters; pygidium with fine, dense ground punctation, with distinct coarser
<pageBreakToken pageId="166" pageNumber="167" start="start">punctures</pageBreakToken>
densely interspersed, especially near basal margin; marginal pygidial sulcus complete, deep, strongly crenulate on inner and outer edges. Male genitalia (Figs 44
<normalizedToken originalValue="AE">A-E</normalizedToken>
): accessory sclerites present; T8 elongate, sides slightly convergent to apex, basal emargination shallow, narrow, basal membrane attachment line distad emargination by nearly its depth, apical emargination narrow, shallow, ventrolateral apodemes most strongly developed at base, nearly meeting at midline, narrowed to apex; S8 short, widest at base, sides weakly rounded, apical guides developed throughout length, widened gradually to apex, apices subacute in outer corners, halves separate, strongly divergent in apical half; T9 with sides subparallel in basal two-thirds, convergent to narrow, obliquely truncate apices; T10 with halves separate; S9 broad, weakly desclerotized along midline, narrowest at middle, strongly widened to broadly rounded base, lateral flanges wide, apex with small median emargination, apical flanges short, separate; tegmen widest just beyond midpoint, narrowing to subacute base, becoming quite flat dorsoventrally, curving downward toward apex, with wide
<normalizedToken originalValue="V">'V'</normalizedToken>
-shaped medioventral process strongly projecting beneath about one-third from base; basal piece nearly half tegmen length; median lobe over half tegmen length, with proximal apodemes not strongly differentiated, proximal portions conspicuous.
</paragraph>
</subSubSection>
<subSubSection pageId="166" pageNumber="167" type="remarks">
<paragraph pageId="166" pageNumber="167">Remarks.</paragraph>
<paragraph pageId="166" pageNumber="167">
In general body shape (Fig. 45A) this species strongly resembles members of the
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus dubius" order="Coleoptera" pageId="166" pageNumber="167" phylum="Arthropoda" rank="species" species="dubius">Operclipygus dubius</taxonomicName>
group. However, its male genitalia absolutely preclude placement there. This species can be distinguished by its lack of lateral submarginal pronotal stria, and by its elongate, subcylindrical body shape. It is very similar to the sympatric
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus minutus" order="Coleoptera" pageId="166" pageNumber="167" phylum="Arthropoda" rank="species" species="minutus">Operclipygus minutus</taxonomicName>
, but is more distinctly parallel-sided and subdepressed, and has entirely different genitalia, lacking the truncate medioventral process and subapical tegmenal cleft characteristic of that species and several close relatives. The type series is limited to those specimens from northeastern Brazil, as populations from other areas do exhibit some variation, some with anterior fragments of the lateral submarginal pronotal stria.
</paragraph>
<caption pageId="166" pageNumber="167">
<paragraph pageId="166" pageNumber="167">
Figure 44. Male genitalia of
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus impositus" order="Coleoptera" pageId="166" pageNumber="167" phylum="Arthropoda" rank="species" species="impositus">Operclipygus impositus</taxonomicName>
(hospes group). A T8 B S8 C T9 &amp; T10 D S9 E Aedeagus, dorsal and lateral views.
</paragraph>
</caption>
<caption pageId="166" pageNumber="167">
<paragraph pageId="166" pageNumber="167">
Figure 45.
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus hospes" order="Coleoptera" pageId="166" pageNumber="167" phylum="Arthropoda" rank="species" species="hospes">Operclipygus hospes</taxonomicName>
group. A Dorsal habitus of
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus impositus" order="Coleoptera" pageId="166" pageNumber="167" phylum="Arthropoda" rank="species" species="impositus">Operclipygus impositus</taxonomicName>
B Pygidia of
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus impositus" order="Coleoptera" pageId="166" pageNumber="167" phylum="Arthropoda" rank="species" species="impositus">Operclipygus impositus</taxonomicName>
C Dorsal habitus of
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus curtistrius" order="Coleoptera" pageId="166" pageNumber="167" phylum="Arthropoda" rank="species" species="curtistrius">Operclipygus curtistrius</taxonomicName>
D Frons of
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus curtistrius" order="Coleoptera" pageId="166" pageNumber="167" phylum="Arthropoda" rank="species" species="curtistrius">Operclipygus curtistrius</taxonomicName>
E Pygidia of
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus curtistrius" order="Coleoptera" pageId="166" pageNumber="167" phylum="Arthropoda" rank="species" species="curtistrius">Operclipygus curtistrius</taxonomicName>
F Dorsal habitus of
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus bulbistoma" order="Coleoptera" pageId="166" pageNumber="167" phylum="Arthropoda" rank="species" species="bulbistoma">Operclipygus bulbistoma</taxonomicName>
G Frons of
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus bulbistoma" order="Coleoptera" pageId="166" pageNumber="167" phylum="Arthropoda" rank="species" species="bulbistoma">Operclipygus bulbistoma</taxonomicName>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="166" pageNumber="167" type="etymology">
<paragraph pageId="166" pageNumber="167">Etymology.</paragraph>
<paragraph pageId="166" pageNumber="167">
The Latin name impositus is related to the English impostor, as this species looks very much like members of the
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus dubius" order="Coleoptera" pageId="166" pageNumber="167" phylum="Arthropoda" rank="species" species="dubius">Operclipygus dubius</taxonomicName>
species group.
</paragraph>
</subSubSection>
</treatment>
</document>