treatments-xml/data/6E/6A/B6/6E6AB6166F52FFB1FF065A1AFE59CC33.xml
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<document id="A31B56EF676E2B6D5B7630705F0C6A83" ID-CLB-Dataset="26923" ID-DOI="10.11646/zootaxa.4594.1.1" ID-GBIF-Dataset="3fc46d02-9b74-434b-a777-46d70967d61f" ID-ISSN="1175-5326" ID-Zenodo-Dep="2667081" ID-ZooBank="4308D364-BCD7-473D-83BC-5B4130C32287" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1556963471166" checkinUser="plazi" docAuthor="Leong, Chi-Man, Guénard, Benoit, Shiao, Shiuh-Feng &amp; Lin, Chung-Chi" docDate="2019" docId="6E6AB6166F52FFB1FF065A1AFE59CC33" docLanguage="en" docName="zootaxa.4594.1.1.pdf.imf" docOrigin="Zootaxa 4594 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Ponera kohmoku Terayama 1996" docType="treatment" docVersion="11" lastPageNumber="39" masterDocId="9253CE6E6F71FF96FF915F65FFB6CD2E" masterDocTitle="Taxonomic revision of the genus Ponera Latreille, 1804 (Hymenoptera: Formicidae) of Taiwan and Japan, with a key to East Asian species" masterLastPageNumber="86" masterPageNumber="1" pageNumber="36" updateTime="1698725828403" updateUser="ExternalLinkService">
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<mods:title id="A8357BABCEDB5EC2D9834B0DD5E9593B">Taxonomic revision of the genus Ponera Latreille, 1804 (Hymenoptera: Formicidae) of Taiwan and Japan, with a key to East Asian species</mods:title>
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<mods:namePart id="377CAA1C0A3E461DD8F7862BDDB15310">Leong, Chi-Man</mods:namePart>
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<mods:namePart id="A60EBB049D91E20EE5343F6BC6F09241">Guénard, Benoit</mods:namePart>
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<mods:namePart id="8BC418A3E516DAB42A697CBFA3BBE72D">Shiao, Shiuh-Feng</mods:namePart>
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<mods:namePart id="B7CFF82DCBCD9549804EC4C95E276936">Lin, Chung-Chi</mods:namePart>
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<treatment id="6E6AB6166F52FFB1FF065A1AFE59CC33" ID-DOI="http://doi.org/10.5281/zenodo.5669050" ID-GBIF-Taxon="157059613" ID-Zenodo-Dep="5669050" LSID="urn:lsid:plazi:treatment:6E6AB6166F52FFB1FF065A1AFE59CC33" httpUri="http://treatment.plazi.org/id/6E6AB6166F52FFB1FF065A1AFE59CC33" lastPageId="39" lastPageNumber="39" pageId="35" pageNumber="36">
<subSubSection id="AED9548B6F52FFB5FF065A1AFD81C8B4" box="[151,567,1407,1434]" pageId="35" pageNumber="36" type="nomenclature">
<paragraph id="E67C07006F52FFB5FF065A1AFD81C8B4" blockId="35.[151,605,1407,1468]" box="[151,567,1407,1434]" pageId="35" pageNumber="36">
<heading id="BD34B06C6F52FFB5FF065A1AFD81C8B4" bold="true" box="[151,567,1407,1434]" fontSize="11" level="1" pageId="35" pageNumber="36" reason="1">
<taxonomicName id="21C37C836F52FFB5FF065A1AFD81C8B4" authority="Terayama, 1996" authorityName="Terayama" authorityYear="1996" box="[151,567,1407,1434]" class="Insecta" family="Formicidae" genus="Ponera" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="kohmoku">
<emphasis id="D4B7DB126F52FFB5FF065A1AFD81C8B4" bold="true" box="[151,567,1407,1434]" pageId="35" pageNumber="36">
<emphasis id="D4B7DB126F52FFB5FF065A1AFEDEC8B7" bold="true" box="[151,360,1407,1433]" italics="true" pageId="35" pageNumber="36">Ponera kohmoku</emphasis>
<bibRefCitation id="82527AF16F52FFB5FEFE5AE5FD81C8B4" author="Terayama, M." box="[367,567,1408,1434]" pageId="35" pageNumber="36" pagination="9 - 32" refId="ref36648" refString="Terayama, M. (1996) Taxonomic Studies on the Japanese Formicidae, Part 2 Seven Genera of Ponerinae, Cerapachyinae and Myrmicinae. Nature and Human Activities, 1, 9 - 32." type="journal article" year="1996">Terayama, 1996</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="AED9548B6F52FFB5FF065AC6FDEBC895" box="[151,605,1443,1468]" pageId="35" pageNumber="36" type="description">
<paragraph id="E67C07006F52FFB5FF065AC6FDEBC895" blockId="35.[151,605,1407,1468]" box="[151,605,1443,1468]" pageId="35" pageNumber="36">
(
<figureCitation id="7EF81B856F52FFB5FF0E5AC6FF42C895" box="[159,244,1443,1468]" captionStart="FIGURE 41" captionStartId="36.[151,250,1951,1973]" captionTargetBox="[180,1408,193,1929]" captionTargetId="figure@36.[180,1408,193,1929]" captionTargetPageId="36" captionText="FIGURE 41. P. kohmoku paratype worker (LCM_MT-Ponera-13b), A: body, lateral view, B: body, dorsal view, C: head, fullface view, D: specimen label." httpUri="https://zenodo.org/record/2667165/files/figure.png" pageId="35" pageNumber="36">Fig. 41</figureCitation>
: worker;
<figureCitation id="7EF81B856F52FFB5FECD5AC6FE06C895" box="[348,432,1443,1468]" captionStart="FIGURE 42" captionStartId="37.[151,250,1951,1973]" captionTargetBox="[180,1408,193,1929]" captionTargetId="figure@37.[180,1408,193,1929]" captionTargetPageId="37" captionText="FIGURE 42. P. kohmoku paratype dealate queen (LCM_MT-Ponera-14c), A: body, lateral view, B: body, dorsal view, C: head, full-face view, D: specimen label." httpUri="https://zenodo.org/record/2667167/files/figure.png" pageId="35" pageNumber="36">Fig. 42</figureCitation>
: dealate queen)
</paragraph>
</subSubSection>
<subSubSection id="AED9548B6F52FFB5FF065A89FD63CB31" pageId="35" pageNumber="36" type="reference_group">
<paragraph id="E67C07006F52FFB5FF065A89FB07CB2F" blockId="35.[151,1201,1515,1567]" box="[151,1201,1515,1537]" pageId="35" pageNumber="36">
<treatmentCitationGroup id="C6D3202E6F52FFB5FF065A89FB07CB2F" box="[151,1201,1515,1537]" pageId="35" pageNumber="36">
<taxonomicName id="21C37C836F52FFB5FF065A89FDB0CB2F" authority="Terayama, 1996: 9" authorityName="Terayama" authorityPageNumber="9" authorityYear="1996" box="[151,518,1515,1537]" class="Insecta" family="Formicidae" genus="Ponera" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="kohmoku">
<emphasis id="D4B7DB126F52FFB5FF065A89FEF2CB2F" box="[151,324,1515,1537]" italics="true" pageId="35" pageNumber="36">Ponera kohmoku</emphasis>
<treatmentCitation id="676221116F52FFB5FED85A89FDB0CB2F" author="Terayama, M." box="[329,518,1515,1537]" page="9" pageId="35" pageNumber="36" year="1996">
<bibRefCitation id="82527AF16F52FFB5FED85A89FE46CB2F" author="Terayama, M." box="[329,496,1515,1537]" pageId="35" pageNumber="36" pagination="9 - 32" refId="ref36648" refString="Terayama, M. (1996) Taxonomic Studies on the Japanese Formicidae, Part 2 Seven Genera of Ponerinae, Cerapachyinae and Myrmicinae. Nature and Human Activities, 1, 9 - 32." type="journal article" year="1996">Terayama, 1996</bibRefCitation>
: 9
</treatmentCitation>
</taxonomicName>
(w. q.).
<materialsCitation id="56AB0D5D6F52FFB5FDCB5A89FB1BCB2F" ID-GBIF-Occurrence="2237845455" box="[602,1197,1515,1537]" country="Japan" location="Kurio" municipality="Yaku-shima" pageId="35" pageNumber="36" specimenCount="1" stateProvince="Kagoshima">
<typeStatus id="3978B9A26F52FFB5FDCB5A89FD3ACB2F" box="[602,652,1516,1537]" pageId="35" pageNumber="36">Type</typeStatus>
locality:
<collectingCountry id="9ED447906F52FFB5FD7D5A89FC9ECB2F" box="[748,808,1516,1537]" name="Japan" pageId="35" pageNumber="36">Japan</collectingCountry>
,
<collectingRegion id="2407C9E26F52FFB5FCA35A8EFC13CB2F" box="[818,933,1515,1537]" country="Japan" name="Kagoshima" pageId="35" pageNumber="36">Kagoshima</collectingRegion>
Pref.,
<collectingMunicipality id="06189D7A6F52FFB5FC785A8EFBD3CB2F" box="[1001,1125,1515,1537]" pageId="35" pageNumber="36">Yaku-shima</collectingMunicipality>
,
<location id="E31C51DB6F52FFB5FBFE5A8EFB1BCB2F" LSID="urn:lsid:plazi:treatment:6E6AB6166F52FFB1FF065A1AFE59CC33:E31C51DB6F52FFB5FBFE5A8EFB1BCB2F" box="[1135,1197,1515,1537]" country="Japan" municipality="Yaku-shima" name="Kurio" pageId="35" pageNumber="36" stateProvince="Kagoshima">Kurio</location>
</materialsCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="E67C07006F52FFB5FF06596FFD63CB31" blockId="35.[151,1201,1515,1567]" box="[151,725,1545,1567]" pageId="35" pageNumber="36">
<treatmentCitationGroup id="C6D3202E6F52FFB5FF06596FFD63CB31" box="[151,725,1545,1567]" pageId="35" pageNumber="36">
<taxonomicName id="21C37C836F52FFB5FF06596FFDB4CB31" ID-CoL="4LTNG" authority=": Santschi, 1937: 364" authorityName=": Santschi" authorityPageNumber="364" authorityYear="1937" box="[151,514,1545,1567]" class="Insecta" family="Formicidae" genus="Ponera" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="scabra">
<emphasis id="D4B7DB126F52FFB5FF06596FFE9BCB31" box="[151,301,1545,1567]" italics="true" pageId="35" pageNumber="36">Ponera scabra</emphasis>
:
<treatmentCitation id="676221116F52FFB5FEAB596CFDB4CB31" author="Santschi, F." box="[314,514,1545,1567]" page="364" pageId="35" pageNumber="36" year="1937">
<bibRefCitation id="82527AF16F52FFB5FEAB596CFE65CB31" author="Santschi, F." box="[314,467,1545,1567]" pageId="35" pageNumber="36" pagination="361 - 388" refId="ref36276" refString="Santschi, F. (1937) Fourmis du Japon et de Formose. Bulletin et Annales de la Societe Entomologique de Belgique, 361 - 388." type="book chapter" year="1937">Santschi, 1937</bibRefCitation>
: 364
</treatmentCitation>
</taxonomicName>
(
<emphasis id="D4B7DB126F52FFB5FD80596CFD63CB31" bold="true" box="[529,725,1545,1567]" pageId="35" pageNumber="36">misidentification).</emphasis>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="AED9548B6F52FFB5FF065929FB10CB83" pageId="35" pageNumber="36" type="materials_examined">
<paragraph id="E67C07006F52FFB5FF065929FB10CB83" blockId="35.[151,1437,1612,2033]" pageId="35" pageNumber="36">
<emphasis id="D4B7DB126F52FFB5FF065929FD85CB4B" bold="true" box="[151,563,1612,1637]" pageId="35" pageNumber="36">
Type material examined:
<materialsCitation id="56AB0D5D6F52FFB5FE435929FD85CB4B" ID-GBIF-Occurrence="2237845403" box="[466,563,1612,1637]" collectingDate="1981-03-23" collectionCode="MTC" collectorName="M Terayama" country="Japan" location="Yaku-shima" municipality="Kurio" pageId="35" pageNumber="36" specimenCount="6" specimenCount-worker="6" stateProvince="Kagoshima" typeStatus="paratype">
<collectingCountry id="9ED447906F52FFB5FE435929FD98CB4B" box="[466,558,1612,1637]" name="Japan" pageId="35" pageNumber="36">JAPAN</collectingCountry>
.
</materialsCitation>
</emphasis>
<typeStatus id="3978B9A26F52FFB5FDAC5928FD1BCB4B" box="[573,685,1613,1637]" pageId="35" pageNumber="36" type="paratype">Paratypes</typeStatus>
,
<specimenCount id="F0C5CC896F52FFB5FD2D5929FC98CB4B" box="[700,814,1612,1637]" pageId="35" pageNumber="36" type="worker">6 workers</specimenCount>
and 1 dealate queen,
<collectingMunicipality id="06189D7A6F52FFB5FBA35929FBC0CB4B" box="[1074,1142,1612,1637]" pageId="35" pageNumber="36">Kurio</collectingMunicipality>
,
<location id="E31C51DB6F52FFB5FB145929FAB8CB4B" LSID="urn:lsid:plazi:treatment:6E6AB6166F52FFB1FF065A1AFE59CC33:E31C51DB6F52FFB5FB145929FAB8CB4B" box="[1157,1294,1612,1637]" country="Japan" municipality="Kurio" name="Yaku-shima" pageId="35" pageNumber="36" stateProvince="Kagoshima">Yaku-shima</location>
,
<collectingRegion id="2407C9E26F52FFB5FA8C5929FA2ACB4B" box="[1309,1436,1612,1637]" country="Japan" name="Kagoshima" pageId="35" pageNumber="36">Kagoshima</collectingRegion>
Pref.,
<collectingDate id="8239D8286F52FFB5FF4D5914FED6CBA7" box="[220,352,1649,1673]" pageId="35" pageNumber="36" value="1981-03-23">
<date id="927D21C06F52FFB5FF4D5914FED6CBA7" box="[220,352,1649,1673]" pageId="35" pageNumber="36" value="1981-03-23">23 III 1981</date>
</collectingDate>
,
<collectorName id="4B3662D66F52FFB5FEFF5914FE48CBA7" box="[366,510,1649,1673]" pageId="35" pageNumber="36">M Terayama</collectorName>
leg (MTC: LCM_MT-Ponera-13, LCM_MT-Ponera-14, LCM_MT-Ponera-15).
<materialsCitation id="56AB0D5D6F52FFB5FA1F5914FB14CB83" ID-GBIF-Occurrence="2237845423" collectingDate="1981-03-10" collectionCode="LCM" collectorName="M Terayama" country="Japan" location="Sata-Misaki" pageId="35" pageNumber="36" specimenCount="1" specimenCount-worker="1" stateProvince="Kagoshima" typeStatus="paratype">
<specimenCount id="F0C5CC896F52FFB5FA1F5914FF5ECB83" pageId="35" pageNumber="36" type="worker">1 worker</specimenCount>
,
<location id="E31C51DB6F52FFB5FF6259F1FECBCB83" LSID="urn:lsid:plazi:treatment:6E6AB6166F52FFB1FF065A1AFE59CC33:E31C51DB6F52FFB5FF6259F1FECBCB83" box="[243,381,1684,1709]" country="Japan" name="Sata-Misaki" pageId="35" pageNumber="36" stateProvince="Kagoshima">Sata-Misaki</location>
,
<collectingRegion id="2407C9E26F52FFB5FE1959F1FDB1CB83" box="[392,519,1684,1709]" country="Japan" name="Kagoshima" pageId="35" pageNumber="36">Kagoshima</collectingRegion>
Pref.,
<collectingDate id="8239D8286F52FFB5FDC259F0FD64CB83" box="[595,722,1685,1709]" pageId="35" pageNumber="36" value="1981-03-10">
<date id="927D21C06F52FFB5FDC259F0FD64CB83" box="[595,722,1685,1709]" pageId="35" pageNumber="36" value="1981-03-10">10 III 1981</date>
</collectingDate>
,
<collectorName id="4B3662D66F52FFB5FD4F59F0FCDBCB83" box="[734,877,1685,1709]" pageId="35" pageNumber="36">M Terayama</collectorName>
leg (
<collectionCode id="80D29FC56F52FFB5FC3559F1FC55CB83" box="[932,995,1684,1709]" country="0" name="Universidad de Chile, Laboratorio de Citogenetica de Mamiferos" pageId="35" pageNumber="36">LCM</collectionCode>
_MT-Ponera-16)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="AED9548B6F52FFB5FF5659DDFF5BCADF" pageId="35" pageNumber="36" type="diagnosis">
<paragraph id="E67C07006F52FFB5FF5659DDFEF9CBDB" blockId="35.[151,1437,1612,2033]" pageId="35" pageNumber="36">
<emphasis id="D4B7DB126F52FFB5FF5659DDFE03CBFF" bold="true" box="[199,437,1720,1745]" pageId="35" pageNumber="36">Diagnosis (worker).</emphasis>
<taxonomicName id="21C37C836F52FFB5FE2E59DCFD37CBFF" authorityName="Terayama" authorityYear="1996" box="[447,641,1720,1745]" class="Insecta" family="Formicidae" genus="Ponera" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="kohmoku">
<emphasis id="D4B7DB126F52FFB5FE2E59DCFD37CBFF" box="[447,641,1720,1745]" italics="true" pageId="35" pageNumber="36">Ponera kohmoku</emphasis>
</taxonomicName>
can be easily distinguished by the large eye, consisting of about 20 indistinct facets.
</paragraph>
<paragraph id="E67C07006F52FFB5FF565864FEA1CAAB" blockId="35.[151,1437,1612,2033]" pageId="35" pageNumber="36">Measurements (mm) and indices. Workers (n=6): HL 0.830.89; HW 0.700.77; SL 0.610.68; A06L 0.05; A07L 0.05; A08L 0.08; A09L 0.10; A10L 0.11; PrW 0.570.65; WL 1.061.29; PeH 0.620.65; PeNL 0.340.37; PeW 0.440.51; ATL 0.630.71; ATW 0.760.80; CI 8087, SI 8291, PeI 7382, LPeI 5458, DPeI 124143, ATI 8189.</paragraph>
<paragraph id="E67C07006F52FFB5FF5658F4FF5BCADF" blockId="35.[151,1437,1612,2033]" pageId="35" pageNumber="36">Queen (n=1): HL 0.94; HW 0.81; SL 0.71; A06L 0.06; A07L 0.06; A08L 0.09; A09L 0.11; A10L 0.11; PrW 0.75; WL 1.52; PeH 0.74; PeNL 0.37; PeW 0.55; ATL 0.80; ATW 0.90; CI 86, SI 88, PeI 74, LPeI 50, DPeI 150, ATI 88.</paragraph>
</subSubSection>
<caption id="B2BC57886F55FFB2FF0658FAFE77CAFD" httpUri="https://zenodo.org/record/2667165/files/figure.png" pageId="36" pageNumber="37" startId="36.[151,250,1951,1973]" targetBox="[180,1408,193,1929]" targetPageId="36">
<paragraph id="E67C07006F55FFB2FF0658FAFE77CAFD" blockId="36.[151,1436,1951,2003]" pageId="36" pageNumber="37">
<emphasis id="D4B7DB126F55FFB2FF0658FAFE96CA9B" bold="true" box="[151,288,1951,1973]" pageId="36" pageNumber="37">FIGURE 41.</emphasis>
<taxonomicName id="21C37C836F55FFB2FEB658C5FE2ACA9B" authorityName="Terayama" authorityYear="1996" box="[295,412,1951,1973]" class="Insecta" family="Formicidae" genus="Ponera" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="kohmoku">
<emphasis id="D4B7DB126F55FFB2FEB658C5FE85CA9B" box="[295,307,1952,1973]" italics="true" pageId="36" pageNumber="37">P</emphasis>
.
<emphasis id="D4B7DB126F55FFB2FED058FAFE2ACA9B" box="[321,412,1951,1973]" italics="true" pageId="36" pageNumber="37">kohmoku</emphasis>
</taxonomicName>
paratype worker (LCM_MT-Ponera-13b), A: body, lateral view, B: body, dorsal view, C: head, fullface view, D: specimen label.
</paragraph>
</caption>
<caption id="B2BC57886F54FFB3FF0658FAFE5BCAFD" httpUri="https://zenodo.org/record/2667167/files/figure.png" pageId="37" pageNumber="38" startId="37.[151,250,1951,1973]" targetBox="[180,1408,193,1929]" targetPageId="37">
<paragraph id="E67C07006F54FFB3FF0658FAFE5BCAFD" blockId="37.[151,1436,1951,2003]" pageId="37" pageNumber="38">
<emphasis id="D4B7DB126F54FFB3FF0658FAFEA9CA9B" bold="true" box="[151,287,1951,1973]" pageId="37" pageNumber="38">FIGURE 42.</emphasis>
<taxonomicName id="21C37C836F54FFB3FEB458C5FE2CCA9B" authorityName="Terayama" authorityYear="1996" box="[293,410,1951,1973]" class="Insecta" family="Formicidae" genus="Ponera" kingdom="Animalia" order="Hymenoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="kohmoku">
<emphasis id="D4B7DB126F54FFB3FEB458C5FE84CA9B" box="[293,306,1952,1973]" italics="true" pageId="37" pageNumber="38">P</emphasis>
.
<emphasis id="D4B7DB126F54FFB3FEAF58FAFE2CCA9B" box="[318,410,1951,1973]" italics="true" pageId="37" pageNumber="38">kohmoku</emphasis>
</taxonomicName>
paratype dealate queen (LCM_MT-Ponera-14c), A: body, lateral view, B: body, dorsal view, C: head, full-face view, D: specimen label.
</paragraph>
</caption>
<subSubSection id="AED9548B6F57FFB0FF565FF2FD82CA19" pageId="38" pageNumber="39" type="description">
<paragraph id="E67C07006F57FFB0FF565FF2FB7DCCAE" blockId="38.[151,1436,151,320]" lastBlockId="38.[151,1227,359,384]" pageId="38" pageNumber="39">
<emphasis id="D4B7DB126F57FFB0FF565FF2FD9CCD9E" bold="true" box="[199,554,151,176]" pageId="38" pageNumber="39">Description of worker. Head.</emphasis>
In full-face view, head subrectangular and distinctly longer than broad (CI: 8087), with straight posterior margin, moderately convex lateral margins, and bluntly rounded posterolateral corners (
<figureCitation id="7EF81B856F57FFB0FF6B5FBAFED4CDD6" box="[250,354,223,248]" captionStart="FIGURE 41" captionStartId="36.[151,250,1951,1973]" captionTargetBox="[180,1408,193,1929]" captionTargetId="figure@36.[180,1408,193,1929]" captionTargetPageId="36" captionText="FIGURE 41. P. kohmoku paratype worker (LCM_MT-Ponera-13b), A: body, lateral view, B: body, dorsal view, C: head, fullface view, D: specimen label." httpUri="https://zenodo.org/record/2667165/files/figure.png" pageId="38" pageNumber="39">Fig. 41C</figureCitation>
). Eye large; composed of a total of 1822 indistinct facets. Anterior clypeal margin with strong and blunt medial tooth. Masticatory margin of mandible with a series of about ten indistinct denticles, and three large teeth on the apical part. Antennal scape, when laid backward, exactly reaching the posterolateral corner; average ratio of the length of antennomeres = 1.16: 1.68: 1.98: 2.24 (n=6).
</paragraph>
<paragraph id="E67C07006F57FFB0FF565EFEFEC7CF6A" blockId="38.[151,1437,411,1228]" pageId="38" pageNumber="39">
<emphasis id="D4B7DB126F57FFB0FF565EFEFEFACC9A" bold="true" box="[199,332,411,436]" pageId="38" pageNumber="39">Mesosoma.</emphasis>
Mesosomal dorsum in lateral view slightly convex (
<figureCitation id="7EF81B856F57FFB0FC2B5EFEFB91CC9D" box="[954,1063,411,436]" captionStart="FIGURE 41" captionStartId="36.[151,250,1951,1973]" captionTargetBox="[180,1408,193,1929]" captionTargetId="figure@36.[180,1408,193,1929]" captionTargetPageId="36" captionText="FIGURE 41. P. kohmoku paratype worker (LCM_MT-Ponera-13b), A: body, lateral view, B: body, dorsal view, C: head, fullface view, D: specimen label." httpUri="https://zenodo.org/record/2667165/files/figure.png" pageId="38" pageNumber="39">Fig. 41A</figureCitation>
). Pronotum in dorsal view with broadly convex anterior margin, and moderately convex lateral margins (
<figureCitation id="7EF81B856F57FFB0FC455EDBFB8ACCF9" box="[980,1084,446,471]" captionStart="FIGURE 41" captionStartId="36.[151,250,1951,1973]" captionTargetBox="[180,1408,193,1929]" captionTargetId="figure@36.[180,1408,193,1929]" captionTargetPageId="36" captionText="FIGURE 41. P. kohmoku paratype worker (LCM_MT-Ponera-13b), A: body, lateral view, B: body, dorsal view, C: head, fullface view, D: specimen label." httpUri="https://zenodo.org/record/2667165/files/figure.png" pageId="38" pageNumber="39">Fig. 41B</figureCitation>
). Metanotal groove deeply and broadly incised. Lateral mesopleural suture clearly incised. Propodeal dorsum in dorsal view narrow, with straight lateral margins. Propodeal corner in lateral view angular; propodeal dorsum and declivity forming approximatively a 130 degree angle.
</paragraph>
<paragraph id="E67C07006F57FFB0FF565D2BFBA2CE32" blockId="38.[151,1437,411,1228]" pageId="38" pageNumber="39">
<emphasis id="D4B7DB126F57FFB0FF565D2BFEFCCF49" bold="true" box="[199,330,590,615]" pageId="38" pageNumber="39">Metasoma.</emphasis>
Petiolar node in dorsal view subcircular and thick, with well convex anterior margin, and moderately convex posterior margin (
<figureCitation id="7EF81B856F57FFB0FDD85D16FD05CFA5" box="[585,691,627,652]" captionStart="FIGURE 41" captionStartId="36.[151,250,1951,1973]" captionTargetBox="[180,1408,193,1929]" captionTargetId="figure@36.[180,1408,193,1929]" captionTargetPageId="36" captionText="FIGURE 41. P. kohmoku paratype worker (LCM_MT-Ponera-13b), A: body, lateral view, B: body, dorsal view, C: head, fullface view, D: specimen label." httpUri="https://zenodo.org/record/2667165/files/figure.png" pageId="38" pageNumber="39">Fig. 41B</figureCitation>
). Petiolar node in dorsal view well thick and subtrapezoid, with straight anterior and posterior margins, and slightly convex dorsal margin (
<figureCitation id="7EF81B856F57FFB0FC6B5DF3FBD3CF81" box="[1018,1125,662,687]" captionStart="FIGURE 41" captionStartId="36.[151,250,1951,1973]" captionTargetBox="[180,1408,193,1929]" captionTargetId="figure@36.[180,1408,193,1929]" captionTargetPageId="36" captionText="FIGURE 41. P. kohmoku paratype worker (LCM_MT-Ponera-13b), A: body, lateral view, B: body, dorsal view, C: head, fullface view, D: specimen label." httpUri="https://zenodo.org/record/2667165/files/figure.png" pageId="38" pageNumber="39">Fig. 41A</figureCitation>
). Subpetiolar process thick with medium-sized and oval fenestra, anteroventral corner blunt, straight ventral margin, and posteroventral corner concave with a pair of medium-sized teeth. Third abdominal tergum broader than long (ATI: 8189) and narrow anteriad, with broadly convex anterior margin, and well convex lateral margins.
</paragraph>
<paragraph id="E67C07006F57FFB0FF565C43FABFCE85" blockId="38.[151,1437,411,1228]" pageId="38" pageNumber="39">
<emphasis id="D4B7DB126F57FFB0FF565C43FEF4CE11" bold="true" box="[199,322,806,831]" pageId="38" pageNumber="39">Sculpture.</emphasis>
Head densely punctate. Mandible sparsely punctate. Mesosomal dorsum in dorsal view evenly punctate. Mesopleuron, metapleuron and propodeum evenly striate. Propodeal declivity smooth and shining. Lateral face of petiole evenly punctate, posterior face smooth, dorsum rough and evenly punctate. The third and fourth abdominal segments densely punctate, other segments smooth and shining with few punctures.
</paragraph>
<paragraph id="E67C07006F57FFB0FF565CD3FD1DC912" blockId="38.[151,1437,411,1228]" pageId="38" pageNumber="39">
<emphasis id="D4B7DB126F57FFB0FF565CD3FEE1CEE1" bold="true" box="[199,343,950,975]" pageId="38" pageNumber="39">Pubescence.</emphasis>
Head, antennae, mesosoma, petiole, and gaster with evenly distributed short hairs. Mesopleuron, metapleuron, propodeum and lateral face of petiole with scattered short hairs. Dorsal and ventral faces of head, anterior margin of clypeus, sides of mandibles, dorsum of petiolar node and gaster with many long erect hairs. Subpetiolar process with a few long erect hairs.
</paragraph>
<paragraph id="E67C07006F57FFB0FF565B23FB3DC94E" blockId="38.[151,1437,411,1228]" box="[199,1163,1094,1120]" pageId="38" pageNumber="39">
<emphasis id="D4B7DB126F57FFB0FF565B23FEA6C971" bold="true" box="[199,272,1094,1119]" pageId="38" pageNumber="39">Color.</emphasis>
Body color dark. Mandible, clypeus, antennae, legs, and apex of gaster orange.
</paragraph>
<paragraph id="E67C07006F57FFB0FF565B0EFCC9C825" blockId="38.[151,1437,411,1228]" lastBlockId="38.[151,895,1266,1291]" pageId="38" pageNumber="39">
<emphasis id="D4B7DB126F57FFB0FF565B0EFDA9C9AA" bold="true" box="[199,543,1131,1156]" pageId="38" pageNumber="39">Description of queen. Head.</emphasis>
(
<figureCitation id="7EF81B856F57FFB0FDA35B0EFD2DC9AD" box="[562,667,1131,1156]" captionStart="FIGURE 42" captionStartId="37.[151,250,1951,1973]" captionTargetBox="[180,1408,193,1929]" captionTargetId="figure@37.[180,1408,193,1929]" captionTargetPageId="37" captionText="FIGURE 42. P. kohmoku paratype dealate queen (LCM_MT-Ponera-14c), A: body, lateral view, B: body, dorsal view, C: head, full-face view, D: specimen label." httpUri="https://zenodo.org/record/2667167/files/figure.png" pageId="38" pageNumber="39">Fig. 42C</figureCitation>
) Similar to worker caste, but with large and oval Eye, maximum diameter of each eye about
<quantity id="213BAAE56F57FFB0FE445BEAFD8DC989" box="[469,571,1167,1191]" metricMagnitude="-4" metricUnit="m" metricValue="2.1" pageId="38" pageNumber="39" unit="mm" value="0.21">0.21 mm</quantity>
with 15 ommatidia along the maximum diameter. Three ocelli present, and forming a subequilateral triangle. In full-face view, scape similar to worker; average ratio of the length of antennomeres = 1.07: 1.53: 1.80: 1.85 (n=1).
</paragraph>
<paragraph id="E67C07006F57FFB0FF565A43FD4CC8DD" blockId="38.[151,1437,1318,2027]" pageId="38" pageNumber="39">
<emphasis id="D4B7DB126F57FFB0FF565A43FEFAC811" bold="true" box="[199,332,1318,1343]" pageId="38" pageNumber="39">Mesosoma.</emphasis>
Mesosomal dorsum in lateral view broadly convex (
<figureCitation id="7EF81B856F57FFB0FC2A5A43FB9EC811" box="[955,1064,1318,1343]" captionStart="FIGURE 42" captionStartId="37.[151,250,1951,1973]" captionTargetBox="[180,1408,193,1929]" captionTargetId="figure@37.[180,1408,193,1929]" captionTargetPageId="37" captionText="FIGURE 42. P. kohmoku paratype dealate queen (LCM_MT-Ponera-14c), A: body, lateral view, B: body, dorsal view, C: head, full-face view, D: specimen label." httpUri="https://zenodo.org/record/2667167/files/figure.png" pageId="38" pageNumber="39">Fig. 42A</figureCitation>
). Pronotum in dorsal view with convex anterior margin, and broadly convex lateral margins (
<figureCitation id="7EF81B856F57FFB0FCAE5A2FFC10C84C" box="[831,934,1354,1379]" captionStart="FIGURE 42" captionStartId="37.[151,250,1951,1973]" captionTargetBox="[180,1408,193,1929]" captionTargetId="figure@37.[180,1408,193,1929]" captionTargetPageId="37" captionText="FIGURE 42. P. kohmoku paratype dealate queen (LCM_MT-Ponera-14c), A: body, lateral view, B: body, dorsal view, C: head, full-face view, D: specimen label." httpUri="https://zenodo.org/record/2667167/files/figure.png" pageId="38" pageNumber="39">Fig. 42B</figureCitation>
). Scutum suboval and narrow posteriad, with moderately convex anterior margin, and slightly convex posterior margin. Transcutal suture distinct and broad. Scutellum suboval. Anapleural sulcus clearly incised. Propodeal-metapleural suture weakly incised. Propodeal dorsum in dorsal view broad, with straight lateral margins. Propodeal corner rounded; propodeal dorsum and declivity forming approximatively a 120 degree angle.
</paragraph>
<paragraph id="E67C07006F57FFB0FF565A9BFC14CB89" blockId="38.[151,1437,1318,2027]" pageId="38" pageNumber="39">
<emphasis id="D4B7DB126F57FFB0FF565A9BFEFCCB39" bold="true" box="[199,330,1534,1559]" pageId="38" pageNumber="39">Metasoma.</emphasis>
Petiolar node in dorsal view thick and arched, broader than long, with broadly convex anterior margin, and slightly convex posterior margin (
<figureCitation id="7EF81B856F57FFB0FD5C5947FC8ACB14" box="[717,828,1570,1595]" captionStart="FIGURE 42" captionStartId="37.[151,250,1951,1973]" captionTargetBox="[180,1408,193,1929]" captionTargetId="figure@37.[180,1408,193,1929]" captionTargetPageId="37" captionText="FIGURE 42. P. kohmoku paratype dealate queen (LCM_MT-Ponera-14c), A: body, lateral view, B: body, dorsal view, C: head, full-face view, D: specimen label." httpUri="https://zenodo.org/record/2667167/files/figure.png" pageId="38" pageNumber="39">Fig. 42B</figureCitation>
). Petiolar node in lateral view subrectangular and moderately thick, with straight anterior and posterior margins (
<figureCitation id="7EF81B856F57FFB0FCCE5923FC7FCB71" box="[863,969,1606,1631]" captionStart="FIGURE 42" captionStartId="37.[151,250,1951,1973]" captionTargetBox="[180,1408,193,1929]" captionTargetId="figure@37.[180,1408,193,1929]" captionTargetPageId="37" captionText="FIGURE 42. P. kohmoku paratype dealate queen (LCM_MT-Ponera-14c), A: body, lateral view, B: body, dorsal view, C: head, full-face view, D: specimen label." httpUri="https://zenodo.org/record/2667167/files/figure.png" pageId="38" pageNumber="39">Fig. 42A</figureCitation>
). Subpetiolar process with large and oval fenestra, anteroventral corner blunt, ventral margin straight, and posteroventral corner concave with a pair of small teeth. The third abdominal segment similar to worker caste (ATI: 88).
</paragraph>
<paragraph id="E67C07006F57FFB0FF5659D7FDE3CBE4" blockId="38.[151,1437,1318,2027]" box="[199,597,1714,1739]" pageId="38" pageNumber="39">
<emphasis id="D4B7DB126F57FFB0FF5659D7FEF4CBE5" bold="true" box="[199,322,1714,1739]" pageId="38" pageNumber="39">Sculpture.</emphasis>
Similar to worker caste.
</paragraph>
<paragraph id="E67C07006F57FFB0FF5659B3FDDCCBC0" blockId="38.[151,1437,1318,2027]" box="[199,618,1750,1775]" pageId="38" pageNumber="39">
<emphasis id="D4B7DB126F57FFB0FF5659B3FEE1CBC1" bold="true" box="[199,343,1750,1775]" pageId="38" pageNumber="39">Pubescence.</emphasis>
Similar to worker caste.
</paragraph>
<paragraph id="E67C07006F57FFB0FF56599FFD95CA3C" blockId="38.[151,1437,1318,2027]" box="[199,547,1786,1811]" pageId="38" pageNumber="39">
<emphasis id="D4B7DB126F57FFB0FF56599FFEA6CA3D" bold="true" box="[199,272,1786,1811]" pageId="38" pageNumber="39">Color.</emphasis>
Similar to worker caste.
</paragraph>
<paragraph id="E67C07006F57FFB0FF56587BFD82CA19" blockId="38.[151,1437,1318,2027]" box="[199,564,1822,1847]" pageId="38" pageNumber="39">
<emphasis id="D4B7DB126F57FFB0FF56587BFE01CA19" bold="true" box="[199,439,1822,1847]" pageId="38" pageNumber="39">Description of male.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="AED9548B6F57FFB0FF565827FD32CA75" box="[199,644,1858,1883]" pageId="38" pageNumber="39" type="distribution">
<paragraph id="E67C07006F57FFB0FF565827FD32CA75" blockId="38.[151,1437,1318,2027]" box="[199,644,1858,1883]" pageId="38" pageNumber="39">
<emphasis id="D4B7DB126F57FFB0FF565827FED6CA75" bold="true" box="[199,352,1858,1883]" pageId="38" pageNumber="39">Distribution.</emphasis>
<collectingCountry id="9ED447906F57FFB0FEF65826FE11CA74" box="[359,423,1859,1882]" name="Japan" pageId="38" pageNumber="39">Japan</collectingCountry>
(
<collectingRegion id="2407C9E26F57FFB0FE245827FD81CA75" box="[437,567,1858,1883]" country="Japan" name="Kagoshima" pageId="38" pageNumber="39">Kagoshima</collectingRegion>
Pref.).
</paragraph>
</subSubSection>
<subSubSection id="AED9548B6F57FFB1FF565803FE59CC33" lastPageId="39" lastPageNumber="40" pageId="38" pageNumber="39" type="discussion">
<paragraph id="E67C07006F57FFB1FF565803FE59CC33" blockId="38.[151,1437,1318,2027]" lastBlockId="39.[151,1436,151,285]" lastPageId="39" lastPageNumber="40" pageId="38" pageNumber="39">
<emphasis id="D4B7DB126F57FFB0FF565803FE86CA51" bold="true" box="[199,304,1894,1919]" pageId="38" pageNumber="39">Remark.</emphasis>
A specimen collected from KiuShiu, Shiroyama,
<collectingCountry id="9ED447906F57FFB0FCFE5802FC04CA50" box="[879,946,1895,1918]" name="Japan" pageId="38" pageNumber="39">Japan</collectingCountry>
, is mistakenly labeled as a
<typeStatus id="3978B9A26F57FFB0FB7E5802FAA9CA51" box="[1263,1311,1895,1919]" pageId="38" pageNumber="39">type</typeStatus>
of
<taxonomicName id="21C37C836F57FFB0FAD85802FF55CA8D" authorityName=": Santschi" authorityYear="1937" class="Insecta" family="Formicidae" genus="Ponera" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="scabra">
<emphasis id="D4B7DB126F57FFB0FAD85802FF55CA8D" italics="true" pageId="38" pageNumber="39">Ponera scabra</emphasis>
</taxonomicName>
at NHMB (AntWeb images, specimen CASENT0915299).
<taxonomicName id="21C37C836F57FFB0FC1C58EEFB83CA8D" authorityName=": Santschi" authorityYear="1937" box="[909,1077,1930,1955]" class="Insecta" family="Formicidae" genus="Ponera" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="scabra">
<emphasis id="D4B7DB126F57FFB0FC1C58EEFB83CA8D" box="[909,1077,1930,1955]" italics="true" pageId="38" pageNumber="39">Ponera scabra</emphasis>
</taxonomicName>
was described from Mt. Maya,
<collectingRegion id="2407C9E26F57FFB0FF0658CAFED6CAE9" box="[151,352,1966,1991]" country="Japan" name="Hyogo" pageId="38" pageNumber="39">Hyogo Prefecture</collectingRegion>
,
<collectingCountry id="9ED447906F57FFB0FEFA58CAFE18CAE8" box="[363,430,1967,1990]" name="Japan" pageId="38" pageNumber="39">Japan</collectingCountry>
.
<bibRefCitation id="82527AF16F57FFB0FE2858CBFDD2CAE9" author="Santschi, F." box="[441,612,1966,1991]" pageId="38" pageNumber="39" pagination="361 - 388" refId="ref36276" refString="Santschi, F. (1937) Fourmis du Japon et de Formose. Bulletin et Annales de la Societe Entomologique de Belgique, 361 - 388." type="book chapter" year="1937">Santschi (1937</bibRefCitation>
: 364) identified the KiuShiu specimen as a gynecoid worker of
<taxonomicName id="21C37C836F57FFB0FABC58CAFA23CAE9" authorityName=": Santschi" authorityYear="1937" box="[1325,1429,1966,1991]" class="Insecta" family="Formicidae" genus="Ponera" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="scabra">
<emphasis id="D4B7DB126F57FFB0FABC58CAFA8DCAE8" box="[1325,1339,1967,1990]" italics="true" pageId="38" pageNumber="39">P</emphasis>
.
<emphasis id="D4B7DB126F57FFB0FAD858CBFA23CAE9" box="[1353,1429,1966,1991]" italics="true" pageId="38" pageNumber="39">scabra</emphasis>
</taxonomicName>
. However, this specimen can be easily identified as
<taxonomicName id="21C37C836F57FFB0FC9758B6FBE3CAC5" authority="Terayama, 1996" authorityName="Terayama" authorityYear="1996" box="[774,1109,2002,2027]" class="Insecta" family="Formicidae" genus="Ponera" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="kohmoku">
<emphasis id="D4B7DB126F57FFB0FC9758B6FC3BCAC5" box="[774,909,2002,2027]" italics="true" pageId="38" pageNumber="39">P. kohmoku</emphasis>
<bibRefCitation id="82527AF16F57FFB0FC0A58B6FBE3CAC5" author="Terayama, M." box="[923,1109,2002,2027]" pageId="38" pageNumber="39" pagination="9 - 32" refId="ref36648" refString="Terayama, M. (1996) Taxonomic Studies on the Japanese Formicidae, Part 2 Seven Genera of Ponerinae, Cerapachyinae and Myrmicinae. Nature and Human Activities, 1, 9 - 32." type="journal article" year="1996">Terayama, 1996</bibRefCitation>
</taxonomicName>
by the combination of the following characters: large compound eye (ca. 30 facets), distinct metanotal groove and subcircular petiolar node in dorsal view; in contrast to
<taxonomicName id="21C37C836F56FFB1FE555FD8FD9BCDFB" authorityName=": Santschi" authorityYear="1937" box="[452,557,188,213]" class="Insecta" family="Formicidae" genus="Ponera" kingdom="Animalia" order="Hymenoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="scabra">
<emphasis id="D4B7DB126F56FFB1FE555FD8FE64CDFA" box="[452,466,189,212]" italics="true" pageId="39" pageNumber="40">P</emphasis>
.
<emphasis id="D4B7DB126F56FFB1FE705FD9FD9BCDFB" box="[481,557,188,213]" italics="true" pageId="39" pageNumber="40">scabra</emphasis>
</taxonomicName>
, which has small compound eye (ca. 5 facets), an indistinct metanotal groove and an arched petiolar node in dorsal view. Thus, there are no records of gynecoid workers in
<taxonomicName id="21C37C836F56FFB1FB705F84FA85CDD6" authorityName="Latreille" authorityYear="1804" box="[1249,1331,225,248]" class="Insecta" family="Formicidae" genus="Ponera" kingdom="Animalia" order="Hymenoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="genus">
<emphasis id="D4B7DB126F56FFB1FB705F84FA85CDD6" box="[1249,1331,225,248]" italics="true" pageId="39" pageNumber="40">Ponera</emphasis>
</taxonomicName>
after the exclusion of Santschis record.
</paragraph>
</subSubSection>
</treatment>
</document>