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<mods:title id="FD41B09FB7F09B764A89B2A119987CAB">Illuminating a Dark Taxon: Revision of European Trichacis Förster (Hymenoptera Platygastridae) reveals a glut of synonyms</mods:title>
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<mods:affiliation id="FCBBD7DB23C4882896A1554D5EF6CA56">State Museum of Natural History Stuttgart, Rosenstein 1, 70191 Stuttgart; University of Hohenheim, Institute of Biology, Biological Systematics (190 w), 70599 Stuttgart, Germany.</mods:affiliation>
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<mods:affiliation id="09B95EB68E26ABC675D413379E841C12">State Museum of Natural History Stuttgart, Rosenstein 1, 70191 Stuttgart; University of Hohenheim, Institute of Biology, Biological Systematics (190 w), 70599 Stuttgart, Germany. &amp; lars. krogmann @ smns-bw. de; https: // orcid. org / 0000 - 0002 - 3724 - 1735</mods:affiliation>
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<mods:affiliation id="0A9E0606461597405F6D95A105C3DA79">Florida Department of Agriculture and Consumer Services, Division of Plant Industry, Gainesville, FL, USA. elijah. talamas @ fdacs. gov; https: // orcid. org / 0000 - 0002 - 1048 - 6345</mods:affiliation>
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<treatment id="15229365B057FFF6FF4DFB4AFC0BFA24" ID-DOI="http://doi.org/10.5281/zenodo.7906740" ID-GBIF-Taxon="209614746" ID-Zenodo-Dep="7906740" LSID="urn:lsid:plazi:treatment:15229365B057FFF6FF4DFB4AFC0BFA24" httpUri="http://treatment.plazi.org/id/15229365B057FFF6FF4DFB4AFC0BFA24" lastPageNumber="565" pageId="2" pageNumber="565">
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<paragraph id="9D342273B057FFF6FF4DFB4AFDAAFB56" blockId="2.[151,523,1194,1221]" box="[151,523,1194,1221]" pageId="2" pageNumber="565">
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<emphasis id="AFFFFE61B057FFF6FF4DFB4AFDAAFB56" bold="true" box="[151,523,1194,1221]" pageId="2" pageNumber="565">
Genus
<taxonomicName id="5A8B59F0B057FFF6FF34FB4AFDAAFB56" ID-CoL="7Z2X" ID-ENA="408505" authority="Forster, 1856" authorityName="Forster" authorityYear="1856" box="[238,523,1194,1221]" class="Insecta" family="Platygastridae" genus="Trichacis" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="565" phylum="Arthropoda" rank="genus">
<emphasis id="AFFFFE61B057FFF6FF34FB4AFEFEFB57" bold="true" box="[238,351,1194,1221]" italics="true" pageId="2" pageNumber="565">Trichacis</emphasis>
<bibRefCitation id="F91A5F82B057FFF6FEBDFB4BFDAAFB56" author="Forster, A." box="[359,523,1194,1221]" pageId="2" pageNumber="565" refId="ref7771" refString="Forster, A. (1856) Hymenopterologische Studien. II. Heft. Chalcidiae und Proctotrupii. Ernst ter Meer, Aachen, 152 pp. [in German]" type="book" year="1856">Förster, 1856</bibRefCitation>
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<treatmentCitation id="1C2A0462B057FFF6FF4DFB0FFE1AFA95" author="Forster, A." box="[151,443,1263,1287]" page="108" pageId="2" pageNumber="565" year="1856">
<taxonomicName id="5A8B59F0B057FFF6FF4DFB0FFE1AFA95" ID-CoL="7Z2X" authority="Forster, 1856: 108" authorityName="Forster" authorityPageNumber="108" authorityYear="1856" box="[151,443,1263,1287]" class="Insecta" family="Platygastridae" genus="Trichacis" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="565" phylum="Arthropoda" rank="genus">
<emphasis id="AFFFFE61B057FFF6FF4DFB0FFF54FA95" box="[151,245,1263,1287]" italics="true" pageId="2" pageNumber="565">Trichacis</emphasis>
<bibRefCitation id="F91A5F82B057FFF6FF27FB10FE1AFA95" author="Forster, A." box="[253,443,1263,1287]" pageId="2" pageNumber="565" refId="ref7771" refString="Forster, A. (1856) Hymenopterologische Studien. II. Heft. Chalcidiae und Proctotrupii. Ernst ter Meer, Aachen, 152 pp. [in German]" type="book" year="1856">Förster, 1856: 108</bibRefCitation>
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, 115.
<typeStatus id="42309CD1B057FFF6FE20FB10FD8DFA9A" box="[506,556,1264,1288]" pageId="2" pageNumber="565">Type</typeStatus>
species
<taxonomicName id="5A8B59F0B057FFF6FD5EFB0FFC1EFA95" authority="Walker, 1835" authorityName="Walker" authorityYear="1835" box="[644,959,1263,1287]" class="Insecta" family="Platygastridae" genus="Platygaster" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="565" phylum="Arthropoda" rank="species" species="pisis">
<emphasis id="AFFFFE61B057FFF6FD5EFB0FFC8EFA95" box="[644,815,1263,1287]" italics="true" pageId="2" pageNumber="565">Platygaster pisis</emphasis>
<bibRefCitation id="F91A5F82B057FFF6FCECFB0FFC1EFA95" author="Walker, F." box="[822,959,1263,1287]" pageId="2" pageNumber="565" pagination="217 - 274" refId="ref9425" refString="Walker, F. (1835) On the species of Platygaster, &amp; c. The Entomological Magazine, 3, 217 - 274." type="journal article" year="1835">Walker, 1835</bibRefCitation>
</taxonomicName>
by subsequent designation by
<treatmentCitation id="1C2A0462B057FFF6FB21FB0FFF51FAB4" author="Ashmead, W. H." page="294" pageId="2" pageNumber="565" year="1893">
<bibRefCitation id="F91A5F82B057FFF6FB21FB0FFF51FAB4" author="Ashmead, W. H." pageId="2" pageNumber="565" pagination="1 - 472" refId="ref7103" refString="Ashmead, W. H. (1893) Monograph of the North American Proctotrypidae. Bulletin of the United States National Museum, 45, 1 - 472. https: // doi. org / 10.5479 / si. 03629236.45.1" type="journal article" year="1893">Ashmead 1893: 294</bibRefCitation>
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.
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<emphasis id="AFFFFE61B057FFF6FF4DFAB4FEB3FAFC" bold="true" box="[151,274,1364,1390]" pageId="2" pageNumber="565">Diagnosis.</emphasis>
Head smooth or finely sculptured with transverse striae above the toruli (
<figureCitation id="05B03EF6B057FFF6FBBEFAB4FB1AFAFC" box="[1124,1211,1364,1390]" captionStart="FIGURE 1" captionStartId="4.[151,250,1896,1920]" captionTargetBox="[165,1422,181,1873]" captionTargetId="figure-23@4.[165,1422,181,1873]" captionTargetPageId="4" captionText="FIGURE 1. Genus-level characters of Trichacis and Isocybus. AB. Trichacis tristis female, SMNS_Hym_Pla_000797; A. Dorsal head and mesosoma, showing broad aspect of clava; B. Dorsal head showing narrow aspect of clava; C. Isocybus female, dorsal habitus, SMNS_Hym_Pla_000880; D. Isocybus female, anterior head, SMNS_Hym_Pla_000851; E. Trichacis tristis female, anterior head, SMNS_Hym_Pla_000805." figureDoi="http://doi.org/10.5281/zenodo.7906724" httpUri="https://zenodo.org/record/7906724/files/figure.png" pageId="2" pageNumber="565">Fig. 1E</figureCitation>
); female with three to five clavomeres (
<figureCitation id="05B03EF6B057FFF6FEADFA98FE71FA00" box="[375,464,1400,1426]" captionStart="FIGURE 2" captionStartId="5.[151,250,1426,1450]" captionTargetBox="[157,1435,232,1401]" captionTargetId="figure-19@5.[151,1435,181,1401]" captionTargetPageId="5" captionText="FIGURE 2. Antennae of Trichacis and Isocybus. A. T. tristis female, SMNS_Hym_Pla_000217; B. T. tristis male, SMNS_ Hym_Pla_000220; C. Isocybus female (apical antennomere missing), SMNS_Hym_Pla_000870." figureDoi="http://doi.org/10.5281/zenodo.7906728" httpUri="https://zenodo.org/record/7906728/files/figure.png" pageId="2" pageNumber="565">Fig. 2A</figureCitation>
) as opposed to six in
<taxonomicName id="5A8B59F0B057FFF6FD1CFA98FC89FA00" authority="Forster" box="[710,808,1400,1426]" class="Insecta" family="Platygastridae" genus="Isocybus" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="565" phylum="Arthropoda" rank="genus">
<emphasis id="AFFFFE61B057FFF6FD1CFA98FC89FA00" box="[710,808,1400,1426]" italics="true" pageId="2" pageNumber="565">Isocybus</emphasis>
</taxonomicName>
(
<figureCitation id="05B03EF6B057FFF6FCE2FA98FC30FA00" box="[824,913,1400,1426]" captionStart="FIGURE 2" captionStartId="5.[151,250,1426,1450]" captionTargetBox="[157,1435,232,1401]" captionTargetId="figure-19@5.[151,1435,181,1401]" captionTargetPageId="5" captionText="FIGURE 2. Antennae of Trichacis and Isocybus. A. T. tristis female, SMNS_Hym_Pla_000217; B. T. tristis male, SMNS_ Hym_Pla_000220; C. Isocybus female (apical antennomere missing), SMNS_Hym_Pla_000870." figureDoi="http://doi.org/10.5281/zenodo.7906728" httpUri="https://zenodo.org/record/7906728/files/figure.png" pageId="2" pageNumber="565">Fig. 2C</figureCitation>
); OOL less than or equal to LOL (
<figureCitation id="05B03EF6B057FFF6FACCFA98FA2FFA00" box="[1302,1422,1400,1426]" captionStart="FIGURE 1" captionStartId="4.[151,250,1896,1920]" captionTargetBox="[165,1422,181,1873]" captionTargetId="figure-23@4.[165,1422,181,1873]" captionTargetPageId="4" captionText="FIGURE 1. Genus-level characters of Trichacis and Isocybus. AB. Trichacis tristis female, SMNS_Hym_Pla_000797; A. Dorsal head and mesosoma, showing broad aspect of clava; B. Dorsal head showing narrow aspect of clava; C. Isocybus female, dorsal habitus, SMNS_Hym_Pla_000880; D. Isocybus female, anterior head, SMNS_Hym_Pla_000851; E. Trichacis tristis female, anterior head, SMNS_Hym_Pla_000805." figureDoi="http://doi.org/10.5281/zenodo.7906724" httpUri="https://zenodo.org/record/7906724/files/figure.png" pageId="2" pageNumber="565">Fig. 1B, E</figureCitation>
); mesoscutellum with tuft of setae originating in a depression (
<figureCitation id="05B03EF6B057FFF6FC99FA7CFC3DFA24" box="[835,924,1436,1462]" captionStart="FIGURE 1" captionStartId="4.[151,250,1896,1920]" captionTargetBox="[165,1422,181,1873]" captionTargetId="figure-23@4.[165,1422,181,1873]" captionTargetPageId="4" captionText="FIGURE 1. Genus-level characters of Trichacis and Isocybus. AB. Trichacis tristis female, SMNS_Hym_Pla_000797; A. Dorsal head and mesosoma, showing broad aspect of clava; B. Dorsal head showing narrow aspect of clava; C. Isocybus female, dorsal habitus, SMNS_Hym_Pla_000880; D. Isocybus female, anterior head, SMNS_Hym_Pla_000851; E. Trichacis tristis female, anterior head, SMNS_Hym_Pla_000805." figureDoi="http://doi.org/10.5281/zenodo.7906724" httpUri="https://zenodo.org/record/7906724/files/figure.png" pageId="2" pageNumber="565">Fig. 1A</figureCitation>
).
</paragraph>
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