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<mods:title id="1D138ED9319EF1FC0A406D9D66D87CC7">Taxonomic revision of Phascogale tapoatafa (Meyer, 1793) (Dasyuridae; Marsupialia), including descriptions of two new subspecies and confirmation of P. pirata Thomas, 1904 as a Top End endemic</mods:title>
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<treatment id="03A087E6FFE3831CD788F8D1D4DDFE2C" ID-DOI="http://doi.org/10.5281/zenodo.6097131" ID-GBIF-Taxon="119647150" ID-Zenodo-Dep="6097131" LSID="urn:lsid:plazi:treatment:03A087E6FFE3831CD788F8D1D4DDFE2C" httpUri="http://treatment.plazi.org/id/03A087E6FFE3831CD788F8D1D4DDFE2C" lastPageId="34" lastPageNumber="35" pageId="22" pageNumber="23">
<subSubSection id="C313657BFFE38328D788F8D1D4A2F8A7" box="[151,708,1804,1831]" pageId="22" pageNumber="23" type="nomenclature">
<paragraph id="8BB636F0FFE38328D788F8D1D4A2F8A7" blockId="22.[151,708,1804,1831]" box="[151,708,1804,1831]" pageId="22" pageNumber="23">
<heading id="D0FE819CFFE38328D788F8D1D4A2F8A7" bold="true" box="[151,708,1804,1831]" fontSize="11" level="1" pageId="22" pageNumber="23" reason="1">
<taxonomicName id="4C094D73FFE38328D788F8D1D4A2F8A7" ID-CoL="4FVN8" authority="Meyer, 1793" authorityName="Meyer" authorityYear="1793" box="[151,708,1804,1831]" class="Mammalia" family="Dasyuridae" genus="Phascogale" kingdom="Animalia" order="Dasyuromorphia" pageId="22" pageNumber="23" phylum="Chordata" rank="subSpecies" species="tapoatafa" subSpecies="tapoatafa">
<emphasis id="B97DEAE2FFE38328D788F8D1D4A2F8A7" bold="true" box="[151,708,1804,1831]" pageId="22" pageNumber="23">
<emphasis id="B97DEAE2FFE38328D788F8D1D474F8A6" bold="true" box="[151,530,1804,1830]" italics="true" pageId="22" pageNumber="23">Phascogale tapoatafa tapoatafa</emphasis>
(
<bibRefCitation id="EF984B01FFE38328D53EF8D0D4DAF8A7" author="Meyer" box="[545,700,1805,1831]" pageId="22" pageNumber="23" refString="Meyer, F. A. A. (1793) Systematische-summarische Uebersicht der neuesten zoologischen Entdeckungen in Neuholland und Afrika. Nebst zwey andern zoologischen Abhandlungen. im Verlage der Dykischen Buchhandlung, Leipzip, 208 pp." type="book" year="1793">Meyer, 1793</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C313657BFFE3831ED788F889D6A1FB4A" lastPageId="32" lastPageNumber="33" pageId="22" pageNumber="23" type="materials_examined">
<paragraph id="8BB636F0FFE38329D788F889D4A5FE08" blockId="22.[151,1436,1876,2009]" lastBlockId="23.[151,1437,152,392]" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23">
<emphasis id="B97DEAE2FFE38328D788F889D76BF8ED" bold="true" box="[151,269,1876,1901]" pageId="22" pageNumber="23">
<typeStatus id="54B28852FFE38328D788F889D76EF8ED" box="[151,264,1876,1901]" pageId="22" pageNumber="23" type="holotype">Holotype</typeStatus>
:
</emphasis>
As explained by
<bibRefCitation id="EF984B01FFE38328D6C0F889D549F8ED" author="Mahoney" box="[479,815,1876,1901]" pageId="22" pageNumber="23" refString="Mahoney, J. A. &amp; Ride, W. D. L. (1988) Dasyuridae. In: Walton, D. W. (Ed.), Zoological Catalogue of Australia. Vol. 5. Mammalia. Australian Government Printers, Canberra, pp. 14 - 33." type="book" year="1988">Mahoney &amp; Ride (1988: 25)</bibRefCitation>
, “the
<typeStatus id="54B28852FFE38328D468F889D5BFF8ED" box="[887,985,1876,1901]" pageId="22" pageNumber="23" type="holotype">holotype</typeStatus>
is the specimen described and figured under the name Tapoa Tafa by Anon. [Hunter, J.]
<emphasis id="B97DEAE2FFE38328D5DFF8A4D4B0F810" box="[704,726,1913,1936]" italics="true" pageId="22" pageNumber="23">in</emphasis>
<bibRefCitation id="EF984B01FFE38328D5C2F8A5D5F3F810" box="[733,917,1912,1937]" pageId="22" pageNumber="23" refString="White, J. (1790) Journal of a Voyage to New South Wales. J. Debrett, London. p. 281." type="book chapter">White, J. (1790)</bibRefCitation>
.
<emphasis id="B97DEAE2FFE38328D4BFF8A5D302F810" box="[928,1380,1912,1936]" italics="true" pageId="22" pageNumber="23">Journal of a Voyage to New South Wales</emphasis>
with sixty-five plates of non descript animals, birds, lizards, serpents, curious cones of trees and other natural productions. London: J. Debrett xvii 335 pp. (281284 pl 58); preparation 3758 of Owen, R. (1841).
<emphasis id="B97DEAE2FFE38329D204F81CD45DFF54" italics="true" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23">Descriptive and Illustrated Catalogue of the Physiological Series of Comparative Anatomy Contained in the Museum of the Royal College of Surgeons in London</emphasis>
. Products of generation. London: R. &amp; J.E. Taylor Vol.
<date id="FFB71030FFE28329D3AFFF61D2BDFF55" box="[1200,1243,188,213]" pageId="23" pageNumber="24">5 iv</date>
+ xxxvii 284 pp. (209) is part of the
<typeStatus id="54B28852FFE28329D670FF02D7B7FF78" box="[367,465,223,248]" pageId="23" pageNumber="24" type="holotype">holotype</typeStatus>
specimen)”. Although
<bibRefCitation id="EF984B01FFE28329D5C9FF02D5A1FF78" author="Troughton" box="[726,967,223,248]" pageId="23" pageNumber="24" refString="Troughton, E. (1967) Furred Animals of Australia. 9 th Edition. Angus and Roberston, Sydney, 376 pp." type="book" year="1967">Troughton (e.g. 1967</bibRefCitation>
: 31) notes that “the specimen figured was preserved in the Museum of the Royal College of Surgeons”, neither
<bibRefCitation id="EF984B01FFE28329D4ADFED9D206FE9C" author="Thomas" box="[946,1120,260,285]" pageId="23" pageNumber="24" refString="Thomas, O. (1888) Catalogue of the Marsupialia and Monotremata in the Collection of the British Museum (Natural History)." type="book" year="1888">Thomas (1888)</bibRefCitation>
nor Mahoney &amp; Ride were able to locate an appropriate candidate, either in this collection or in the Natural History Museum, London. However, the original plate in
<bibRefCitation id="EF984B01FFE28329D6F5FE91D4E6FEE4" author="White" box="[490,640,332,357]" pageId="23" pageNumber="24" refString="White, J. (1790) Journal of a Voyage to New South Wales. J. Debrett, London. p. 281." type="book chapter" year="1790">White (1790)</bibRefCitation>
is unambiguously diagnostic (provided locality data are also taken into account) and
<typeStatus id="54B28852FFE28329D631FEADD7E0FE08" box="[302,390,368,392]" pageId="23" pageNumber="24" type="neotype">neotype</typeStatus>
designation is unwarranted.
</paragraph>
<caption id="DF766678FFE28329D788FE65D54CFE6F" ID-Table-UUID="DF766678FFE28329D788FE65D54CFE6F" httpUri="http://table.plazi.org/id/DF766678FFE28329D788FE65D54CFE6F" pageId="23" pageNumber="24" targetBox="[159,1330,564,1991]" targetIsTable="true" targetPageId="23">
<paragraph id="8BB636F0FFE28329D788FE65D54CFE6F" blockId="23.[151,1435,439,495]" pageId="23" pageNumber="24">
<emphasis id="B97DEAE2FFE28329D788FE65D77EFE4F" bold="true" box="[151,280,440,463]" pageId="23" pageNumber="24">TABLE 4a.</emphasis>
Summary external measures of male
<taxonomicName id="4C094D73FFE28329D5ACFE6AD5F8FE4F" box="[691,926,439,463]" class="Mammalia" family="Dasyuridae" genus="Phascogale" kingdom="Animalia" order="Dasyuromorphia" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="tapoatafa">
<emphasis id="B97DEAE2FFE28329D5ACFE6AD5F8FE4F" box="[691,926,439,463]" italics="true" pageId="23" pageNumber="24">Phascogale tapoatafa</emphasis>
</taxonomicName>
group specimens used in this study (HV headvent, TV tail-vent, Wgt weight). Obvious subadults excluded.
</paragraph>
</caption>
<paragraph id="8BB636F0FFE28326D780FDD7D34CFEAA" blockId="23.[159,1330,522,544]" lastBlockId="24.[159,1363,234,1290]" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24">Geographic Region HV (mm) TV (mm) Wgt (g) Geographic Region HV (mm) TV (mm) Wgt (g) Southwest Western Australia Mean 179.95 199 190</paragraph>
<paragraph id="8BB636F0FFE28329D780FDE9D223F847" pageId="23" pageNumber="24">
<table id="F909C450FFE27CC1D780FDE9D354F847" box="[159,1330,564,1991]" gridcols="4" gridrows="13" pageId="23" pageNumber="24">
<tr id="353934B2FFE27CC1D780FDE9D354FD1C" box="[159,1330,564,668]" gridrow="0" pageId="23" pageNumber="24">
<th id="76E85DCEFFE27CC1D780FDE9D403FD1C" box="[159,613,564,668]" gridcol="0" gridrow="0" pageId="23" pageNumber="24">Southwest Western Australia Mean s.d. n</th>
<th id="76E85DCEFFE27CC1D42EFDE9D50CFD1C" box="[817,874,564,668]" gridcol="1" gridrow="0" pageId="23" pageNumber="24">200.5 17.95 10</th>
<th id="76E85DCEFFE27CC1D300FDE9D23EFD1C" box="[1055,1112,564,668]" gridcol="2" gridrow="0" pageId="23" pageNumber="24">208.5 10.63 10</th>
<th id="76E85DCEFFE27CC1D3FCFDE9D354FD1C" box="[1251,1330,564,668]" gridcol="3" gridrow="0" pageId="23" pageNumber="24">170 1</th>
</tr>
<tr id="353934B2FFE27CC1D780FD6DD354FC98" box="[159,1330,688,792]" gridrow="1" pageId="23" pageNumber="24" rowspan-3="1">
<th id="76E85DCEFFE27CC1D780FD6DD403FC98" box="[159,613,688,792]" gridcol="0" gridrow="1" pageId="23" pageNumber="24">Max Min South Australia Mean</th>
<td id="76E85DCEFFE27CC1D42EFD6DD50CFC98" box="[817,874,688,792]" gridcol="1" gridrow="1" pageId="23" pageNumber="24">228 175 230</td>
<td id="76E85DCEFFE27CC1D300FD6DD23EFC98" box="[1055,1112,688,792]" gridcol="2" gridrow="1" pageId="23" pageNumber="24">226 193 225</td>
</tr>
<tr id="353934B2FFE27CC1D780FCF1D354FCEB" box="[159,1330,812,875]" gridrow="2" pageId="23" pageNumber="24" rowspan-3="1">
<th id="76E85DCEFFE27CC1D780FCF1D403FCEB" box="[159,613,812,875]" gridcol="0" gridrow="2" pageId="23" pageNumber="24">s.d. n</th>
<td id="76E85DCEFFE27CC1D42EFCF1D50CFCEB" box="[817,874,812,875]" gridcol="1" gridrow="2" pageId="23" pageNumber="24">1</td>
<td id="76E85DCEFFE27CC1D300FCF1D23EFCEB" box="[1055,1112,812,875]" gridcol="2" gridrow="2" pageId="23" pageNumber="24">1</td>
</tr>
<tr id="353934B2FFE27CC1D780FCA2D354FC14" box="[159,1330,895,916]" gridrow="3" pageId="23" pageNumber="24" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th id="76E85DCEFFE27CC1D780FCA2D403FC14" box="[159,613,895,916]" gridcol="0" gridrow="3" pageId="23" pageNumber="24">Max</th>
</tr>
<tr id="353934B2FFE27CC1D780FC75D354FB90" box="[159,1330,936,1040]" gridrow="4" pageId="23" pageNumber="24">
<th id="76E85DCEFFE27CC1D780FC75D403FB90" box="[159,613,936,1040]" gridcol="0" gridrow="4" pageId="23" pageNumber="24">Min Victoria Mean s.d.</th>
<td id="76E85DCEFFE27CC1D42EFC75D50CFB90" box="[817,874,936,1040]" gridcol="1" gridrow="4" pageId="23" pageNumber="24">202.2 13.99</td>
<td id="76E85DCEFFE27CC1D300FC75D23EFB90" box="[1055,1112,936,1040]" gridcol="2" gridrow="4" pageId="23" pageNumber="24">203.2 16.18</td>
<td id="76E85DCEFFE27CC1D3FCFC75D354FB90" box="[1251,1330,936,1040]" gridcol="3" gridrow="4" pageId="23" pageNumber="24">185.33 15.95</td>
</tr>
<tr id="353934B2FFE27CC1D780FBF9D354FB0C" box="[159,1330,1060,1164]" gridrow="5" pageId="23" pageNumber="24">
<th id="76E85DCEFFE27CC1D780FBF9D403FB0C" box="[159,613,1060,1164]" gridcol="0" gridrow="5" pageId="23" pageNumber="24">n Max Min</th>
<td id="76E85DCEFFE27CC1D42EFBF9D50CFB0C" box="[817,874,1060,1164]" gridcol="1" gridrow="5" pageId="23" pageNumber="24">5 220 182</td>
<td id="76E85DCEFFE27CC1D300FBF9D23EFB0C" box="[1055,1112,1060,1164]" gridcol="2" gridrow="5" pageId="23" pageNumber="24">5 230 188</td>
<td id="76E85DCEFFE27CC1D3FCFBF9D354FB0C" box="[1251,1330,1060,1164]" gridcol="3" gridrow="5" pageId="23" pageNumber="24">3 196 167</td>
</tr>
<tr id="353934B2FFE27CC1D780FB7DD354FA88" box="[159,1330,1184,1288]" gridrow="6" pageId="23" pageNumber="24">
<th id="76E85DCEFFE27CC1D780FB7DD403FA88" box="[159,613,1184,1288]" gridcol="0" gridrow="6" pageId="23" pageNumber="24">New South Wales/Southern Mean Queensland s.d. n</th>
<td id="76E85DCEFFE27CC1D42EFB7DD50CFA88" box="[817,874,1184,1288]" gridcol="1" gridrow="6" pageId="23" pageNumber="24">196.4 26.17 5</td>
<td id="76E85DCEFFE27CC1D300FB7DD23EFA88" box="[1055,1112,1184,1288]" gridcol="2" gridrow="6" pageId="23" pageNumber="24">202.4 24.13 5</td>
<td id="76E85DCEFFE27CC1D3FCFB7DD354FA88" box="[1251,1330,1184,1288]" gridcol="3" gridrow="6" pageId="23" pageNumber="24">212.5 84.15 2</td>
</tr>
<tr id="353934B2FFE27CC1D780FAC1D354FA04" box="[159,1330,1308,1412]" gridrow="7" pageId="23" pageNumber="24">
<th id="76E85DCEFFE27CC1D780FAC1D403FA04" box="[159,613,1308,1412]" gridcol="0" gridrow="7" pageId="23" pageNumber="24">Max Min Northern Queensland Mean</th>
<td id="76E85DCEFFE27CC1D42EFAC1D50CFA04" box="[817,874,1308,1412]" gridcol="1" gridrow="7" pageId="23" pageNumber="24">230 170 201.5</td>
<td id="76E85DCEFFE27CC1D300FAC1D23EFA04" box="[1055,1112,1308,1412]" gridcol="2" gridrow="7" pageId="23" pageNumber="24">238 178 202.5</td>
<td id="76E85DCEFFE27CC1D3FCFAC1D354FA04" box="[1251,1330,1308,1412]" gridcol="3" gridrow="7" pageId="23" pageNumber="24">272 153 185</td>
</tr>
<tr id="353934B2FFE27CC1D780FA45D354F980" box="[159,1330,1432,1536]" gridrow="8" pageId="23" pageNumber="24">
<th id="76E85DCEFFE27CC1D780FA45D403F980" box="[159,613,1432,1536]" gridcol="0" gridrow="8" pageId="23" pageNumber="24">s.d. n Max</th>
<td id="76E85DCEFFE27CC1D42EFA45D50CF980" box="[817,874,1432,1536]" gridcol="1" gridrow="8" pageId="23" pageNumber="24">12.02 2 210</td>
<td id="76E85DCEFFE27CC1D300FA45D23EF980" box="[1055,1112,1432,1536]" gridcol="2" gridrow="8" pageId="23" pageNumber="24">3.54 2 205</td>
<td id="76E85DCEFFE27CC1D3FCFA45D354F980" box="[1251,1330,1432,1536]" gridcol="3" gridrow="8" pageId="23" pageNumber="24">1</td>
</tr>
<tr id="353934B2FFE27CC1D780F9C9D354F9FC" box="[159,1330,1556,1660]" gridrow="9" pageId="23" pageNumber="24">
<th id="76E85DCEFFE27CC1D780F9C9D403F9FC" box="[159,613,1556,1660]" gridcol="0" gridrow="9" pageId="23" pageNumber="24">Min 'Top End', Northern Territory Mean s.d.</th>
<td id="76E85DCEFFE27CC1D42EF9C9D50CF9FC" box="[817,874,1556,1660]" gridcol="1" gridrow="9" pageId="23" pageNumber="24">193 209.5 0.71</td>
<td id="76E85DCEFFE27CC1D300F9C9D23EF9FC" box="[1055,1112,1556,1660]" gridcol="2" gridrow="9" pageId="23" pageNumber="24">200 204 5.66</td>
<td id="76E85DCEFFE27CC1D3FCF9C9D354F9FC" box="[1251,1330,1556,1660]" gridcol="3" gridrow="9" pageId="23" pageNumber="24">234</td>
</tr>
<tr id="353934B2FFE27CC1D780F94DD354F978" box="[159,1330,1680,1784]" gridrow="10" pageId="23" pageNumber="24">
<th id="76E85DCEFFE27CC1D780F94DD403F978" box="[159,613,1680,1784]" gridcol="0" gridrow="10" pageId="23" pageNumber="24">n Max Min</th>
<td id="76E85DCEFFE27CC1D42EF94DD50CF978" box="[817,874,1680,1784]" gridcol="1" gridrow="10" pageId="23" pageNumber="24">2 210 209</td>
<td id="76E85DCEFFE27CC1D300F94DD23EF978" box="[1055,1112,1680,1784]" gridcol="2" gridrow="10" pageId="23" pageNumber="24">2 208 200</td>
<td id="76E85DCEFFE27CC1D3FCF94DD354F978" box="[1251,1330,1680,1784]" gridcol="3" gridrow="10" pageId="23" pageNumber="24">2 234 234</td>
</tr>
<tr id="353934B2FFE27CC1D780F8D1D354F8F4" box="[159,1330,1804,1908]" gridrow="11" pageId="23" pageNumber="24" rowspan-3="1">
<th id="76E85DCEFFE27CC1D780F8D1D403F8F4" box="[159,613,1804,1908]" gridcol="0" gridrow="11" pageId="23" pageNumber="24">Kimberley, Western Australia Mean s.d. n</th>
<td id="76E85DCEFFE27CC1D42EF8D1D50CF8F4" box="[817,874,1804,1908]" gridcol="1" gridrow="11" pageId="23" pageNumber="24">163 6.56 3</td>
<td id="76E85DCEFFE27CC1D300F8D1D23EF8F4" box="[1055,1112,1804,1908]" gridcol="2" gridrow="11" pageId="23" pageNumber="24">159 20.07 3</td>
</tr>
<tr id="353934B2FFE27CC1D780F855D354F847" box="[159,1330,1928,1991]" gridrow="12" pageId="23" pageNumber="24" rowspan-3="1">
<th id="76E85DCEFFE27CC1D780F855D403F847" box="[159,613,1928,1991]" gridcol="0" gridrow="12" pageId="23" pageNumber="24">Max Min</th>
<td id="76E85DCEFFE27CC1D42EF855D50CF847" box="[817,874,1928,1991]" gridcol="1" gridrow="12" pageId="23" pageNumber="24">170 157</td>
<td id="76E85DCEFFE27CC1D300F855D23EF847" box="[1055,1112,1928,1991]" gridcol="2" gridrow="12" pageId="23" pageNumber="24">180 140</td>
</tr>
</table>
</paragraph>
<caption id="DF766678FFED8326D788FF45D57AFF4F" pageId="24" pageNumber="25">
<paragraph id="8BB636F0FFED8326D788FF45D57AFF4F" blockId="24.[151,1435,151,207]" pageId="24" pageNumber="25">
<emphasis id="B97DEAE2FFED8326D788FF45D77EFF2F" bold="true" box="[151,280,152,175]" pageId="24" pageNumber="25">TABLE 4b.</emphasis>
Summary external measures of female
<taxonomicName id="4C094D73FFED8326D5DFFF4AD5CFFF2F" box="[704,937,151,175]" class="Mammalia" family="Dasyuridae" genus="Phascogale" kingdom="Animalia" order="Dasyuromorphia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="tapoatafa">
<emphasis id="B97DEAE2FFED8326D5DFFF4AD5CFFF2F" box="[704,937,151,175]" italics="true" pageId="24" pageNumber="25">Phascogale tapoatafa</emphasis>
</taxonomicName>
group specimens used in this study (HV headvent, TV tail-vent, Wt weight). Obvious subadults excluded.
</paragraph>
</caption>
<paragraph id="8BB636F0FFED8326D5F6FEE0D2FDFED3" blockId="24.[159,1363,234,1290]" box="[745,1179,317,339]" pageId="24" pageNumber="25">s.d. 19.61 16.16</paragraph>
<paragraph id="8BB636F0FFED8326D5F6FEBAD2EEFE26" blockId="24.[159,1363,234,1290]" pageId="24" pageNumber="25">
<geoCoordinate id="EE3D5037FFED8326D5F6FEBAD377FEFC" box="[745,1297,358,380]" degrees="11" direction="north" minutes="11" orientation="latitude" pageId="24" pageNumber="25" precision="15" seconds="1" value="11.183611">
n
<date id="FFB71030FFED8326D4AAFEBBD377FEFC" box="[949,1297,358,380]" pageId="24" pageNumber="25" value="2011-01-11">11 11 1</date>
</geoCoordinate>
Max 210 230
</paragraph>
<paragraph id="8BB636F0FFED8326D5F6FE64D2EEFE4F" blockId="24.[159,1363,234,1290]" box="[745,1160,441,463]" pageId="24" pageNumber="25">Min 153 182</paragraph>
<paragraph id="8BB636F0FFED8326D780FE3FD34CFD47" blockId="24.[159,1363,234,1290]" pageId="24" pageNumber="25">
Victoria Mean 193.25 206.25
<geoCoordinate id="EE3D5037FFED8326D21BFE3FD492FDA2" degrees="155.75" direction="south" orientation="latitude" pageId="24" pageNumber="25" precision="555" value="-155.75">155.75 s</geoCoordinate>
.d. 20.30 12.34
<geoCoordinate id="EE3D5037FFED8326D21BFDD1D490FDCB" degrees="41.44" direction="north" orientation="latitude" pageId="24" pageNumber="25" precision="555" value="41.44">
41.44
<geoCoordinate id="EE3D5037FFED8326D5F6FDEBD490FDCB" box="[745,758,566,587]" direction="north" orientation="latitude" pageId="24" pageNumber="25" precision="15" value="4.0677776">n</geoCoordinate>
</geoCoordinate>
4 4 4 Max 213 222 187 Min 165 195 96 New South Wales/Southern Queensland Mean 186 189.5 183
</paragraph>
<paragraph id="8BB636F0FFED8326D5F6FD07D2FDFD70" blockId="24.[159,1363,234,1290]" box="[745,1179,730,752]" pageId="24" pageNumber="25">s.d. 15.56 23.33</paragraph>
<paragraph id="8BB636F0FFED8326D5F6FCD8D2EEFCC3" blockId="24.[159,1363,234,1290]" pageId="24" pageNumber="25">
<geoCoordinate id="EE3D5037FFED8326D5F6FCD8D377FC9A" box="[745,1297,772,794]" degrees="2" direction="north" minutes="2" orientation="latitude" pageId="24" pageNumber="25" precision="15" seconds="1" value="2.033611">n 2 2 1</geoCoordinate>
Max 197 206
</paragraph>
<paragraph id="8BB636F0FFED8326D5F6FC8BD2EEFCEC" blockId="24.[159,1363,234,1290]" box="[745,1160,854,876]" pageId="24" pageNumber="25">Min 175 173</paragraph>
<paragraph id="8BB636F0FFED8326D780FC5DD34CFBBB" blockId="24.[159,1363,234,1290]" pageId="24" pageNumber="25">
'Top End', Northern Territory Mean 174.5 185.5
<geoCoordinate id="EE3D5037FFED8326D21BFC5DD492FC3F" degrees="125.75" direction="south" orientation="latitude" pageId="24" pageNumber="25" precision="555" value="-125.75">125.75 s</geoCoordinate>
.d. 11.90 5.20
<geoCoordinate id="EE3D5037FFED8326D21BFC74D490FC68" degrees="17.67" direction="north" orientation="latitude" pageId="24" pageNumber="25" precision="555" value="17.67">
17.67
<geoCoordinate id="EE3D5037FFED8326D5F6FC0ED490FC68" box="[745,758,979,1000]" direction="north" orientation="latitude" pageId="24" pageNumber="25" precision="15" value="4.0677776">n</geoCoordinate>
</geoCoordinate>
4 4 4 Max 188 192 150 Min 160 180 108
</paragraph>
<paragraph id="8BB636F0FFED8326D780FB93D2EEFBE4" blockId="24.[159,1363,234,1290]" box="[159,1160,1102,1124]" pageId="24" pageNumber="25">Kimberley, Western Australia Mean 145 144</paragraph>
<paragraph id="8BB636F0FFED8326D5F6FBA5D56AFB0E" blockId="24.[159,1363,234,1290]" box="[745,780,1144,1166]" pageId="24" pageNumber="25">s.d.</paragraph>
<paragraph id="8BB636F0FFED8326D5F6FB7FD209FB37" blockId="24.[159,1363,234,1290]" box="[745,1135,1185,1207]" pageId="24" pageNumber="25">
<geoCoordinate id="EE3D5037FFED8326D5F6FB7FD209FB37" box="[745,1135,1185,1207]" degrees="1" direction="north" minutes="1" orientation="latitude" pageId="24" pageNumber="25" precision="925" value="1.0166667">n 1 1</geoCoordinate>
</paragraph>
<paragraph id="8BB636F0FFED8326D5F6FB16D571FB60" blockId="24.[159,1363,234,1290]" box="[745,791,1227,1248]" pageId="24" pageNumber="25">Max</paragraph>
<paragraph id="8BB636F0FFED8326D5F6FB29D575FA8A" blockId="24.[159,1363,234,1290]" box="[745,787,1268,1290]" pageId="24" pageNumber="25">Min</paragraph>
<paragraph id="8BB636F0FFED8326D7D8FA93D407FA52" blockId="24.[151,1437,1358,2031]" pageId="24" pageNumber="25">
<emphasis id="B97DEAE2FFED8326D7D8FA93D7DEFAE7" bold="true" box="[199,440,1358,1383]" pageId="24" pageNumber="25">Revised description.</emphasis>
This account is primarily based on specimens from the central to north coastal region of New South Wales, including one topotypical specimen (
<collectionCode id="ED18AE35FFED8326D413FAAED55DFA0B" box="[780,827,1395,1419]" pageId="24" pageNumber="25">AM</collectionCode>
932) collected in 1894 at Kingswood near Sydney. An adult male specimen from Coraki in northern New South Wales (
<collectionCode id="ED18AE35FFED8326D484FA4AD5A1FA2F" box="[923,967,1431,1455]" pageId="24" pageNumber="25">CM</collectionCode>
20990) is illustrated in
<figureCitation id="13322A75FFED8326D3C0FA4BD34EFA2F" box="[1247,1320,1430,1455]" captionStart="FIGURE 4" captionStartId="8.[151,250,1565,1587]" captionTargetBox="[336,1237,723,1532]" captionTargetId="figure@8.[324,1262,715,1543]" captionTargetPageId="8" captionText="FIGURE 4. Representative puppet skins of nominotypical Phascogale tapoatafa (CM 20990, an adult male from Coraki, New South Wales, in a) dorsal view and b) ventral view and P. t. pirata (CM 7225, an adult male from Nourlangie Rock, Arnhem Land, Northern Territory, in c) dorsal view and d) ventral view), illustrating the extremes of variation in body proportions and colouration among the Brush-tailed Phascogales." httpUri="https://zenodo.org/record/240157/files/figure.png" pageId="24" pageNumber="25">Fig. 4</figureCitation>
. Regional variation is noted after the main account.
</paragraph>
<paragraph id="8BB636F0FFED8326D7D8FA03D2D6F99B" blockId="24.[151,1437,1358,2031]" pageId="24" pageNumber="25">
External measurements taken from museum specimen records are summarised in
<tableCitation id="C68B034BFFED8326D34BFA03D2CFFA77" box="[1108,1193,1502,1527]" captionStart="TABLE 4 a" captionStartId="23.[151,239,440,463]" captionTargetBox="[159,1330,564,1991]" captionTargetPageId="23" captionText="TABLE 4 a. Summary external measures of male Phascogale tapoatafa group specimens used in this study (HV headvent, TV tail-vent, Wgt weight). Obvious subadults excluded." httpUri="http://table.plazi.org/id/DF766678FFE28329D788FE65D54CFE6F" pageId="24" pageNumber="25" tableUuid="DF766678FFE28329D788FE65D54CFE6F">Table 4</tableCitation>
. Adults in a Victorian population attained body weights of
<quantity id="4CF19B15FFED8326D52EF9DED40EF99B" box="[561,616,1539,1563]" metricMagnitude="-1" metricUnit="kg" metricValue="3.11" pageId="24" pageNumber="25" unit="g" value="311.0">311g</quantity>
for males and
<quantity id="4CF19B15FFED8326D411F9DED520F99B" box="[782,838,1539,1563]" metricMagnitude="-1" metricUnit="kg" metricValue="2.12" pageId="24" pageNumber="25" unit="g" value="212.0">212g</quantity>
for females (
<bibRefCitation id="EF984B01FFED8326D4C5F9DFD2C2F99B" author="Soderquist" box="[986,1188,1538,1563]" pageId="24" pageNumber="25" refString="Soderquist, T. R. (1995 c) Ontogeny of sexual dimorphism in size among polytocous mammals: tests of two carnivorous marsupials. Journal of Mammalogy, 76, 376 - 390." type="journal article" year="1995" yearSuffix="c">Soderquist 1995c</bibRefCitation>
).
</paragraph>
<paragraph id="8BB636F0FFED8326D7D8F9FBD7ACF94F" blockId="24.[151,1437,1358,2031]" pageId="24" pageNumber="25">
The dorsal fur is relatively dense and lush, the contour hairs measuring
<quantity id="4CF19B15FFED8326D4E3F9FAD211F9BE" box="[1020,1143,1575,1599]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="24" pageNumber="25" unit="mm" value="12.5" valueMax="13.0" valueMin="12.0">1213 mm</quantity>
in the centre of the back. The overall colour of the back and flanks is grey with a silvery frosting, this effect imparted by narrow white tipping on grey-based contour hairs. Abundant black-tipped guard hairs are distributed evenly over the body and project as much as
<quantity id="4CF19B15FFED8326D670F94ED7D7F92A" box="[367,433,1683,1707]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="24" pageNumber="25" unit="mm" value="9.0">9 mm</quantity>
through the fur. The mid-dorsal zone is distinctly darker than the flanks owing to a more intense basal pigmentation.
</paragraph>
<paragraph id="8BB636F0FFED8326D7D8F907D43FF86F" blockId="24.[151,1437,1358,2031]" pageId="24" pageNumber="25">
Ventral fur colouration varies from grey with an extensive cream wash to predominantly cream (e.g.,
<collectionCode id="ED18AE35FFED8326D788F922D6A3F897" box="[151,197,1791,1815]" pageId="24" pageNumber="25">CM</collectionCode>
3740). In most specimens, the central strip at least consists of pure cream fur, merging laterally into grey-based fur with a progressively shorter cream tipping. In all specimens, the pale ventral colouration extends anteriorly to the chin, posteriorly to surround the cloaca, and laterally to clothe the inner surface of the fore- and hind-limbs. The scrotum is heavily pigmented and densely clothed in cream fur. A pigmented gular gland is evident on some adult specimens of both sexes but the location of this gland is not marked by any obvious external discolouration of the fur. In some specimens the fur in this region has a slight russet tinge basally. The ventral fur in juveniles tends to be duskier, with a diffuse pale cream wash.
</paragraph>
<caption id="DF766678FFEC8327D762FDD0D7E0FDD5" pageId="25" pageNumber="26">
<paragraph id="8BB636F0FFEC8327D762FDD0D7E0FDD5" blockId="25.[125,1808,524,623]" pageId="25" pageNumber="26">
<emphasis id="B97DEAE2FFEC8327D762FDD0D6BDFDA1" bold="true" box="[125,219,525,545]" pageId="25" pageNumber="26">TABLE 5</emphasis>
. Cranial and dental measurements of each taxon and major regional population within the
<taxonomicName id="4C094D73FFEC8327D306FDD1D2ECFDA1" box="[1049,1162,524,545]" class="Mammalia" family="Dasyuridae" genus="Phascogale" kingdom="Animalia" order="Dasyuromorphia" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="tapoatafa">
<emphasis id="B97DEAE2FFEC8327D306FDD1D2ECFDA1" box="[1049,1162,524,545]" italics="true" pageId="25" pageNumber="26">P. tapoatafa</emphasis>
</taxonomicName>
group, presented separately for males (5a) and females (5b). Data are presented separately for three regional populations within nominotypical
<taxonomicName id="4C094D73FFEC8327D405FDFBD5EAFDBB" box="[794,908,550,571]" class="Mammalia" family="Dasyuridae" genus="Phascogale" kingdom="Animalia" order="Dasyuromorphia" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="tapoatafa">
<emphasis id="B97DEAE2FFEC8327D405FDFBD5EAFDBB" box="[794,908,550,571]" italics="true" pageId="25" pageNumber="26">P. tapoatafa</emphasis>
</taxonomicName>
: 1) South Australia; 2) Victoria; 3) New South Wales and southern Queensland. Measurements of holotypes are also presented.
</paragraph>
</caption>
<paragraph id="8BB636F0FFEC8327D762FD86D77EFDEF" blockId="25.[125,1808,524,623]" box="[125,280,603,623]" pageId="25" pageNumber="26">
<emphasis id="B97DEAE2FFEC8327D762FD86D77EFDEF" bold="true" box="[125,280,603,623]" pageId="25" pageNumber="26">
<tableCitation id="C68B034BFFEC8327D762FD86D6A1FDEF" box="[125,199,603,623]" captionStart="TABLE 5" captionStartId="25.[125,203,525,545]" captionText="TABLE 5. Cranial and dental measurements of each taxon and major regional population within the P. tapoatafa group, presented separately for males (5 a) and females (5 b). Data are presented separately for three regional populations within nominotypical P. tapoatafa: 1) South Australia; 2) Victoria; 3) New South Wales and southern Queensland. Measurements of holotypes are also presented." pageId="25" pageNumber="26">Table 5</tableCitation>
a: Males
</emphasis>
</paragraph>
<paragraph id="8BB636F0FFEC8327D762FD52D0B2FD23" blockId="25.[125,1863,655,1664]" box="[125,1748,655,675]" pageId="25" pageNumber="26">Geographic Region SL ZB OBW BS FMW C1-M4 M1-4</paragraph>
<paragraph id="8BB636F0FFEC8327D762FD6CD16EFCA6" blockId="25.[125,1863,655,1664]" pageId="25" pageNumber="26">
SouthwestW.A. Mean 46.34 27.49 18.61 5.67 6.80 17.60
<geoCoordinate id="EE3D5037FFEC8327D1BEFD6CD71AFD65" degrees="10.28" direction="south" orientation="latitude" pageId="25" pageNumber="26" precision="555" value="-10.28">10.28 s</geoCoordinate>
.d. 1.10 1.01 0.43 0.42 0.18 0.51
<geoCoordinate id="EE3D5037FFEC8327D1BEFD0CD71BFC85" degrees="0.42" direction="north" orientation="latitude" pageId="25" pageNumber="26" precision="555" value="0.42">
0.42
<geoCoordinate id="EE3D5037FFEC8327D66DFD2DD71BFC85" box="[370,381,752,773]" direction="north" orientation="latitude" pageId="25" pageNumber="26" precision="15" value="10.169445">
<emphasis id="B97DEAE2FFEC8327D66DFD2DD71BFC85" box="[370,381,752,773]" italics="true" pageId="25" pageNumber="26">n</emphasis>
</geoCoordinate>
</geoCoordinate>
10 10
<date id="FFB71030FFEC8327D483FD2CD34CFC85" box="[924,1322,753,773]" pageId="25" pageNumber="26" value="2010-09-10">10 10 9</date>
10 10 Range 44.5747.79 26.1728.81 17.9719.09 4.946.32 6.407.02 16.8918.69 9.7011.23
</paragraph>
<paragraph id="8BB636F0FFEC8327D762FCEFD173FC26" blockId="25.[125,1863,655,1664]" pageId="25" pageNumber="26">
SouthAustralia Mean 50.47 29.98 18.92 6.13 6.85 18.83
<geoCoordinate id="EE3D5037FFEC8327D1BEFCEFD71AFCE6" degrees="10.94" direction="south" orientation="latitude" pageId="25" pageNumber="26" precision="555" value="-10.94">10.94 s</geoCoordinate>
.d. 0.86 0.66 0.58 0.87 0.31 0.67
<geoCoordinate id="EE3D5037FFEC8327D1BEFC8FD71BFC06" degrees="0.54" direction="north" orientation="latitude" pageId="25" pageNumber="26" precision="555" value="0.54">
0.54
<geoCoordinate id="EE3D5037FFEC8327D66DFCACD71BFC06" box="[370,381,881,902]" direction="north" orientation="latitude" pageId="25" pageNumber="26" precision="15" value="3.0508332">
<emphasis id="B97DEAE2FFEC8327D66DFCACD71BFC06" box="[370,381,881,902]" italics="true" pageId="25" pageNumber="26">n</emphasis>
</geoCoordinate>
</geoCoordinate>
3 3 3 3 3 3 3 Range 49.5651.28 29.4230.70 18.2519.30 5.427.10 6.507.05 18.2319.55 10.5411.56
</paragraph>
<paragraph id="8BB636F0FFEC8327D762FC6ED172FBA7" blockId="25.[125,1863,655,1664]" pageId="25" pageNumber="26">
Victoria Mean 47.29 27.75 18.76 6.01 6.86 18.33
<geoCoordinate id="EE3D5037FFEC8327D1BEFC6ED71AFC67" degrees="10.71" direction="south" orientation="latitude" pageId="25" pageNumber="26" precision="555" value="-10.71">10.71 s</geoCoordinate>
.d. 2.09 2.12 0.46 0.63 0.53 0.55
<geoCoordinate id="EE3D5037FFEC8327D1BEFC0ED71BFB87" degrees="0.25" direction="north" orientation="latitude" pageId="25" pageNumber="26" precision="555" value="0.25">
0.25
<geoCoordinate id="EE3D5037FFEC8327D66DFC2FD71BFB87" box="[370,381,1010,1031]" direction="north" orientation="latitude" pageId="25" pageNumber="26" precision="9" value="8.116667">
<emphasis id="B97DEAE2FFEC8327D66DFC2FD71BFB87" box="[370,381,1010,1031]" italics="true" pageId="25" pageNumber="26">n</emphasis>
</geoCoordinate>
</geoCoordinate>
8 7.00 7 7 7 8 8 Range 45.2451.39 24.7330.61 18.2019.45 5.227.00 6.437.98 17.7719.49 10.2911.17
</paragraph>
<paragraph id="8BB636F0FFEC8327D78FFBE9D0B3FBC8" blockId="25.[125,1863,655,1664]" box="[144,1749,1076,1096]" pageId="25" pageNumber="26">.S.W./ Mean 47.55 28.45 18.95 5.76 6.41 17.78 10.34</paragraph>
<paragraph id="8BB636F0FFEC8327D7BEFB90D76BFBE1" blockId="25.[125,1863,655,1664]" box="[161,269,1101,1121]" pageId="25" pageNumber="26">Queensland</paragraph>
<paragraph id="8BB636F0FFEC8327D66DFBBAD16FFB3C" blockId="25.[125,1863,655,1664]" pageId="25" pageNumber="26">
s.d. 2.15 1.65 0.86 0.39 0.20 0.44
<geoCoordinate id="EE3D5037FFEC8327D1BEFBBAD71BFB1C" degrees="0.23" direction="north" orientation="latitude" pageId="25" pageNumber="26" precision="555" value="0.23">
0.23
<geoCoordinate id="EE3D5037FFEC8327D66DFB5AD71BFB1C" box="[370,381,1159,1180]" direction="north" orientation="latitude" pageId="25" pageNumber="26" precision="15" value="15.288334">
<emphasis id="B97DEAE2FFEC8327D66DFB5AD71BFB1C" box="[370,381,1159,1180]" italics="true" pageId="25" pageNumber="26">n</emphasis>
</geoCoordinate>
</geoCoordinate>
15 17 18 18 17 17 18 Range 42.6051.73 24.0230.00 17.3119.95 4.956.35 6.046.74 16.7818.59 9.9110.70
</paragraph>
<paragraph id="8BB636F0FFEC8327D762FB15D7F4FB7C" blockId="25.[125,1863,655,1664]" pageId="25" pageNumber="26">
NorthQueensland Mean 47.50 29.33 18.61 6.51 6.39 17.25
<geoCoordinate id="EE3D5037FFEC8327D1BEFB15D71DFB7C" degrees="9.82" direction="south" orientation="latitude" pageId="25" pageNumber="26" precision="555" value="-9.82">9.82 s</geoCoordinate>
.d.
</paragraph>
<paragraph id="8BB636F0FFEC8327D66DFAD5D7CAFABD" blockId="25.[125,1863,655,1664]" pageId="25" pageNumber="26">
<geoCoordinate id="EE3D5037FFEC8327D66DFAD5D5CEFA9C" box="[370,936,1288,1309]" degrees="1" direction="north" minutes="1" orientation="latitude" pageId="25" pageNumber="26" precision="15" seconds="1" value="1.0169444">
<emphasis id="B97DEAE2FFEC8327D66DFAD5D71BFA9D" box="[370,381,1288,1309]" italics="true" pageId="25" pageNumber="26">n</emphasis>
1 1 1
</geoCoordinate>
1 1 1 1 Range
</paragraph>
<paragraph id="8BB636F0FFEC8327D79EFA94D098FA3E" blockId="25.[125,1863,655,1664]" pageId="25" pageNumber="26">
TopEnd',N.T. Mean 46.55 27.37 17.21 5.31 5.84 17.13
<geoCoordinate id="EE3D5037FFEC8327D1BEFA94D71AFAFD" degrees="9.46" direction="south" orientation="latitude" pageId="25" pageNumber="26" precision="555" value="-9.46">9.46 s</geoCoordinate>
.d. 0.98 1.84 0.25 0.38 0.04 0.31
<geoCoordinate id="EE3D5037FFEC8327D1BEFAB4D71BFA1D" degrees="0.31" direction="north" orientation="latitude" pageId="25" pageNumber="26" precision="555" value="0.31">
0.31
<geoCoordinate id="EE3D5037FFEC8327D66DFA55D71BFA1D" box="[370,381,1416,1437]" direction="north" orientation="latitude" pageId="25" pageNumber="26" precision="15" value="3.0677776">
<emphasis id="B97DEAE2FFEC8327D66DFA55D71BFA1D" box="[370,381,1416,1437]" italics="true" pageId="25" pageNumber="26">n</emphasis>
</geoCoordinate>
</geoCoordinate>
3 4 4 4 3 4 5 Range 45.9147.67 25.5829.94 16.9717.50 5.025.87 5.815.89 16.8317.55 9.079.92
</paragraph>
<paragraph id="8BB636F0FFEC8327D762FA17D16FF9BE" blockId="25.[125,1863,655,1664]" pageId="25" pageNumber="26">
Kimberley,W.A. Mean 44.16 25.28 17.75 5.01 6.01 16.79
<geoCoordinate id="EE3D5037FFEC8327D1BEFA17D71AFA7E" degrees="9.91" direction="south" orientation="latitude" pageId="25" pageNumber="26" precision="555" value="-9.91">9.91 s</geoCoordinate>
.d. 0.85 1.35 1.09 0.33 0.24 0.15
<geoCoordinate id="EE3D5037FFEC8327D1BEFA37D71BF99E" degrees="0.26" direction="north" orientation="latitude" pageId="25" pageNumber="26" precision="555" value="0.26">
0.26
<geoCoordinate id="EE3D5037FFEC8327D66DF9D4D71BF99E" box="[370,381,1545,1566]" direction="north" orientation="latitude" pageId="25" pageNumber="26" precision="15" value="2.0338888">
<emphasis id="B97DEAE2FFEC8327D66DF9D4D71BF99E" box="[370,381,1545,1566]" italics="true" pageId="25" pageNumber="26">n</emphasis>
</geoCoordinate>
</geoCoordinate>
2 2 2 2 2 2 3 Range 43.5644.76 24.3226.23 16.9818.52 4.775.24 5.846.18 16.6816.89 9.7410.21
</paragraph>
<paragraph id="8BB636F0FFEC8327D762F996D0DEF9DF" blockId="25.[125,1863,655,1664]" box="[125,1720,1611,1631]" pageId="25" pageNumber="26">
<collectionCode id="ED18AE35FFEC8327D762F996D6A1F9DF" box="[125,199,1611,1631]" pageId="25" pageNumber="26">TOTAL</collectionCode>
42 44 45 45 42 45 48
</paragraph>
<paragraph id="8BB636F0FFEC8325D139F9B6D16FFCA5" blockId="25.[125,1863,655,1664]" lastBlockId="27.[125,1863,785,1611]" lastPageId="27" lastPageNumber="28" pageId="25" pageNumber="26">
<emphasis id="B97DEAE2FFEC8327D139F9B6D121F900" box="[1574,1863,1643,1664]" italics="true" pageId="25" pageNumber="26">……continued on the next page</emphasis>
<emphasis id="B97DEAE2FFEF8324D768FDE6D680FDCF" bold="true" box="[119,230,571,591]" pageId="26" pageNumber="27">
<tableCitation id="C68B034BFFEF8324D768FDE6D6B0FDCF" box="[119,214,571,591]" captionStart="TABLE 5" captionStartId="25.[125,203,525,545]" captionText="TABLE 5. Cranial and dental measurements of each taxon and major regional population within the P. tapoatafa group, presented separately for males (5 a) and females (5 b). Data are presented separately for three regional populations within nominotypical P. tapoatafa: 1) South Australia; 2) Victoria; 3) New South Wales and southern Queensland. Measurements of holotypes are also presented." pageId="26" pageNumber="27">
<collectionCode id="ED18AE35FFEF8324D768FDE6D6A3FDCF" box="[119,197,571,591]" pageId="26" pageNumber="27">TABLE</collectionCode>
5
</tableCitation>
a.
</emphasis>
(Continued)
<emphasis id="B97DEAE2FFEE8325D762FD91D724FDE0" bold="true" box="[125,322,588,608]" pageId="27" pageNumber="28">
<tableCitation id="C68B034BFFEE8325D762FD91D6BAFDE0" box="[125,220,588,608]" captionStart="TABLE 5" captionStartId="25.[125,203,525,545]" captionText="TABLE 5. Cranial and dental measurements of each taxon and major regional population within the P. tapoatafa group, presented separately for males (5 a) and females (5 b). Data are presented separately for three regional populations within nominotypical P. tapoatafa: 1) South Australia; 2) Victoria; 3) New South Wales and southern Queensland. Measurements of holotypes are also presented." pageId="27" pageNumber="28">
<collectionCode id="ED18AE35FFEE8325D762FD91D6ADFDE0" box="[125,203,588,608]" pageId="27" pageNumber="28">TABLE</collectionCode>
5
</tableCitation>
b. Females
</emphasis>
Range 42.9946.84 24.9828.17 18.2318.78 4.925.83 6.576.92 16.8617.79 9.9910.72
</paragraph>
<paragraph id="8BB636F0FFEF8324D768FDADD0D2F925" pageId="26" pageNumber="27">
<table id="F909C450FFEF7CC1D768FDADD09FF925" box="[119,1785,624,1701]" gridcols="8" gridrows="18" pageId="26" pageNumber="27">
<tr id="353934B2FFEF7CC1D768FDADD09FFD04" box="[119,1785,624,644]" gridrow="0" pageId="26" pageNumber="27">
<th id="76E85DCEFFEF7CC1D768FDADD7C1FD04" box="[119,423,624,644]" gridcol="0" gridrow="0" pageId="26" pageNumber="27">Geographic Region</th>
<th id="76E85DCEFFEF7CC1D50BFDADD4EEFD04" box="[532,648,624,644]" gridcol="1" gridrow="0" pageId="26" pageNumber="27">PALW</th>
<th id="76E85DCEFFEF7CC1D5C9FDADD558FD04" box="[726,830,624,644]" gridcol="2" gridrow="0" pageId="26" pageNumber="27">ARW</th>
<th id="76E85DCEFFEF7CC1D488FDADD26DFD04" box="[919,1035,624,644]" gridcol="3" gridrow="0" pageId="26" pageNumber="27">PRW</th>
<th id="76E85DCEFFEF7CC1D346FDADD2D3FD04" box="[1113,1205,624,644]" gridcol="4" gridrow="0" pageId="26" pageNumber="27">IOW</th>
<th id="76E85DCEFFEF7CC1D205FDADD311FD04" box="[1306,1399,624,644]" gridcol="5" gridrow="0" pageId="26" pageNumber="27">RD</th>
<th id="76E85DCEFFEF7CC1D2C4FDADD05EFD04" box="[1499,1592,624,644]" gridcol="6" gridrow="0" pageId="26" pageNumber="27">LCL</th>
<th id="76E85DCEFFEF7CC1D182FDADD09FFD04" box="[1693,1785,624,644]" gridcol="7" gridrow="0" pageId="26" pageNumber="27">LCW</th>
</tr>
<tr id="353934B2FFEF7CC1D768FD48D09FFD29" box="[119,1785,661,681]" gridrow="1" pageId="26" pageNumber="27">
<th id="76E85DCEFFEF7CC1D768FD48D7C1FD29" box="[119,423,661,681]" gridcol="0" gridrow="1" pageId="26" pageNumber="27">SouthwestW.A. Mean</th>
<td id="76E85DCEFFEF7CC1D50BFD48D4EEFD29" box="[532,648,661,681]" gridcol="1" gridrow="1" pageId="26" pageNumber="27">14.66</td>
<td id="76E85DCEFFEF7CC1D5C9FD48D558FD29" box="[726,830,661,681]" gridcol="2" gridrow="1" pageId="26" pageNumber="27">8.55</td>
<td id="76E85DCEFFEF7CC1D488FD48D26DFD29" box="[919,1035,661,681]" gridcol="3" gridrow="1" pageId="26" pageNumber="27">18.71</td>
<td id="76E85DCEFFEF7CC1D346FD48D2D3FD29" box="[1113,1205,661,681]" gridcol="4" gridrow="1" pageId="26" pageNumber="27">8.45</td>
<td id="76E85DCEFFEF7CC1D205FD48D311FD29" box="[1306,1399,661,681]" gridcol="5" gridrow="1" pageId="26" pageNumber="27">5.65</td>
<td id="76E85DCEFFEF7CC1D2C4FD48D05EFD29" box="[1499,1592,661,681]" gridcol="6" gridrow="1" pageId="26" pageNumber="27">3.04</td>
<td id="76E85DCEFFEF7CC1D182FD48D09FFD29" box="[1693,1785,661,681]" gridcol="7" gridrow="1" pageId="26" pageNumber="27">1.95</td>
</tr>
<tr id="353934B2FFEF7CC1D768FD64D09FFD71" box="[119,1785,697,753]" gridrow="2" pageId="26" pageNumber="27">
<th id="76E85DCEFFEF7CC1D768FD64D7C1FD71" box="[119,423,697,753]" gridcol="0" gridrow="2" pageId="26" pageNumber="27">
s.d.
<emphasis id="B97DEAE2FFEF8324D672FD01D71EFD71" box="[365,376,732,753]" italics="true" pageId="26" pageNumber="27">n</emphasis>
</th>
<td id="76E85DCEFFEF7CC1D50BFD64D4EEFD71" box="[532,648,697,753]" gridcol="1" gridrow="2" pageId="26" pageNumber="27">0.33 10</td>
<td id="76E85DCEFFEF7CC1D5C9FD64D558FD71" box="[726,830,697,753]" gridcol="2" gridrow="2" pageId="26" pageNumber="27">0.50 10</td>
<td id="76E85DCEFFEF7CC1D488FD64D26DFD71" box="[919,1035,697,753]" gridcol="3" gridrow="2" pageId="26" pageNumber="27">0.51 10</td>
<td id="76E85DCEFFEF7CC1D346FD64D2D3FD71" box="[1113,1205,697,753]" gridcol="4" gridrow="2" pageId="26" pageNumber="27">0.19 10</td>
<td id="76E85DCEFFEF7CC1D205FD64D311FD71" box="[1306,1399,697,753]" gridcol="5" gridrow="2" pageId="26" pageNumber="27">0.37 10</td>
<td id="76E85DCEFFEF7CC1D2C4FD64D05EFD71" box="[1499,1592,697,753]" gridcol="6" gridrow="2" pageId="26" pageNumber="27">0.22 7</td>
<td id="76E85DCEFFEF7CC1D182FD64D09FFD71" box="[1693,1785,697,753]" gridcol="7" gridrow="2" pageId="26" pageNumber="27">0.10 7</td>
</tr>
<tr id="353934B2FFEF7CC1D768FCDCD09FFCB8" box="[119,1785,769,824]" gridrow="3" pageId="26" pageNumber="27">
<th id="76E85DCEFFEF7CC1D768FCDCD7C1FCB8" box="[119,423,769,824]" gridcol="0" gridrow="3" pageId="26" pageNumber="27">Range SouthAustralia Mean</th>
<td id="76E85DCEFFEF7CC1D50BFCDCD4EEFCB8" box="[532,648,769,824]" gridcol="1" gridrow="3" pageId="26" pageNumber="27">14.2715.08 16.37</td>
<td id="76E85DCEFFEF7CC1D5C9FCDCD558FCB8" box="[726,830,769,824]" gridcol="2" gridrow="3" pageId="26" pageNumber="27">7.619.24 9.83</td>
<td id="76E85DCEFFEF7CC1D488FCDCD26DFCB8" box="[919,1035,769,824]" gridcol="3" gridrow="3" pageId="26" pageNumber="27">17.6119.26 18.91</td>
<td id="76E85DCEFFEF7CC1D346FCDCD2D3FCB8" box="[1113,1205,769,824]" gridcol="4" gridrow="3" pageId="26" pageNumber="27">8.198.89 8.25</td>
<td id="76E85DCEFFEF7CC1D205FCDCD311FCB8" box="[1306,1399,769,824]" gridcol="5" gridrow="3" pageId="26" pageNumber="27">5.286.44 6.17</td>
<td id="76E85DCEFFEF7CC1D2C4FCDCD05EFCB8" box="[1499,1592,769,824]" gridcol="6" gridrow="3" pageId="26" pageNumber="27">2.583.24 3.45</td>
<td id="76E85DCEFFEF7CC1D182FCDCD09FFCB8" box="[1693,1785,769,824]" gridcol="7" gridrow="3" pageId="26" pageNumber="27">1.822.10 2.16</td>
</tr>
<tr id="353934B2FFEF7CC1D768FC95D09FFC00" box="[119,1785,840,896]" gridrow="4" pageId="26" pageNumber="27">
<th id="76E85DCEFFEF7CC1D768FC95D7C1FC00" box="[119,423,840,896]" gridcol="0" gridrow="4" pageId="26" pageNumber="27">
s.d.
<emphasis id="B97DEAE2FFEF8324D672FCB6D71EFC00" box="[365,376,875,896]" italics="true" pageId="26" pageNumber="27">n</emphasis>
</th>
<td id="76E85DCEFFEF7CC1D50BFC95D4EEFC00" box="[532,648,840,896]" gridcol="1" gridrow="4" pageId="26" pageNumber="27">0.56 3</td>
<td id="76E85DCEFFEF7CC1D5C9FC95D558FC00" box="[726,830,840,896]" gridcol="2" gridrow="4" pageId="26" pageNumber="27">0.37 3</td>
<td id="76E85DCEFFEF7CC1D488FC95D26DFC00" box="[919,1035,840,896]" gridcol="3" gridrow="4" pageId="26" pageNumber="27">0.53 3</td>
<td id="76E85DCEFFEF7CC1D346FC95D2D3FC00" box="[1113,1205,840,896]" gridcol="4" gridrow="4" pageId="26" pageNumber="27">0.24 3</td>
<td id="76E85DCEFFEF7CC1D205FC95D311FC00" box="[1306,1399,840,896]" gridcol="5" gridrow="4" pageId="26" pageNumber="27">0.05 3</td>
<td id="76E85DCEFFEF7CC1D2C4FC95D05EFC00" box="[1499,1592,840,896]" gridcol="6" gridrow="4" pageId="26" pageNumber="27">0.27 2</td>
<td id="76E85DCEFFEF7CC1D182FC95D09FFC00" box="[1693,1785,840,896]" gridcol="7" gridrow="4" pageId="26" pageNumber="27">0.03 2</td>
</tr>
<tr id="353934B2FFEF7CC1D768FC4DD09FFC24" box="[119,1785,912,932]" gridrow="5" pageId="26" pageNumber="27">
<th id="76E85DCEFFEF7CC1D768FC4DD7C1FC24" box="[119,423,912,932]" gridcol="0" gridrow="5" pageId="26" pageNumber="27">Range</th>
<td id="76E85DCEFFEF7CC1D50BFC4DD4EEFC24" box="[532,648,912,932]" gridcol="1" gridrow="5" pageId="26" pageNumber="27">15.9917.01</td>
<td id="76E85DCEFFEF7CC1D5C9FC4DD558FC24" box="[726,830,912,932]" gridcol="2" gridrow="5" pageId="26" pageNumber="27">9.5410.25</td>
<td id="76E85DCEFFEF7CC1D488FC4DD26DFC24" box="[919,1035,912,932]" gridcol="3" gridrow="5" pageId="26" pageNumber="27">18.3519.41</td>
<td id="76E85DCEFFEF7CC1D346FC4DD2D3FC24" box="[1113,1205,912,932]" gridcol="4" gridrow="5" pageId="26" pageNumber="27">8.058.51</td>
<td id="76E85DCEFFEF7CC1D205FC4DD311FC24" box="[1306,1399,912,932]" gridcol="5" gridrow="5" pageId="26" pageNumber="27">6.086.26</td>
<td id="76E85DCEFFEF7CC1D2C4FC4DD05EFC24" box="[1499,1592,912,932]" gridcol="6" gridrow="5" pageId="26" pageNumber="27">3.263.64</td>
<td id="76E85DCEFFEF7CC1D182FC4DD09FFC24" box="[1693,1785,912,932]" gridcol="7" gridrow="5" pageId="26" pageNumber="27">2.142.18</td>
</tr>
<tr id="353934B2FFEF7CC1D768FC69D09FFC48" box="[119,1785,948,968]" gridrow="6" pageId="26" pageNumber="27">
<th id="76E85DCEFFEF7CC1D768FC69D7C1FC48" box="[119,423,948,968]" gridcol="0" gridrow="6" pageId="26" pageNumber="27">Victoria Mean</th>
<td id="76E85DCEFFEF7CC1D50BFC69D4EEFC48" box="[532,648,948,968]" gridcol="1" gridrow="6" pageId="26" pageNumber="27">15.25</td>
<td id="76E85DCEFFEF7CC1D5C9FC69D558FC48" box="[726,830,948,968]" gridcol="2" gridrow="6" pageId="26" pageNumber="27">9.05</td>
<td id="76E85DCEFFEF7CC1D488FC69D26DFC48" box="[919,1035,948,968]" gridcol="3" gridrow="6" pageId="26" pageNumber="27">18.11</td>
<td id="76E85DCEFFEF7CC1D346FC69D2D3FC48" box="[1113,1205,948,968]" gridcol="4" gridrow="6" pageId="26" pageNumber="27">8.49</td>
<td id="76E85DCEFFEF7CC1D205FC69D311FC48" box="[1306,1399,948,968]" gridcol="5" gridrow="6" pageId="26" pageNumber="27">5.62</td>
<td id="76E85DCEFFEF7CC1D2C4FC69D05EFC48" box="[1499,1592,948,968]" gridcol="6" gridrow="6" pageId="26" pageNumber="27">3.35</td>
<td id="76E85DCEFFEF7CC1D182FC69D09FFC48" box="[1693,1785,948,968]" gridcol="7" gridrow="6" pageId="26" pageNumber="27">2.11</td>
</tr>
<tr id="353934B2FFEF7CC1D768FC05D09FFB90" box="[119,1785,984,1040]" gridrow="7" pageId="26" pageNumber="27">
<th id="76E85DCEFFEF7CC1D768FC05D7C1FB90" box="[119,423,984,1040]" gridcol="0" gridrow="7" pageId="26" pageNumber="27">
s.d.
<emphasis id="B97DEAE2FFEF8324D672FC26D71EFB90" box="[365,376,1019,1040]" italics="true" pageId="26" pageNumber="27">n</emphasis>
</th>
<td id="76E85DCEFFEF7CC1D50BFC05D4EEFB90" box="[532,648,984,1040]" gridcol="1" gridrow="7" pageId="26" pageNumber="27">0.64 8</td>
<td id="76E85DCEFFEF7CC1D5C9FC05D558FB90" box="[726,830,984,1040]" gridcol="2" gridrow="7" pageId="26" pageNumber="27">0.86 8</td>
<td id="76E85DCEFFEF7CC1D488FC05D26DFB90" box="[919,1035,984,1040]" gridcol="3" gridrow="7" pageId="26" pageNumber="27">1.16 8</td>
<td id="76E85DCEFFEF7CC1D346FC05D2D3FB90" box="[1113,1205,984,1040]" gridcol="4" gridrow="7" pageId="26" pageNumber="27">0.38 8</td>
<td id="76E85DCEFFEF7CC1D205FC05D311FB90" box="[1306,1399,984,1040]" gridcol="5" gridrow="7" pageId="26" pageNumber="27">0.21 8</td>
<td id="76E85DCEFFEF7CC1D2C4FC05D05EFB90" box="[1499,1592,984,1040]" gridcol="6" gridrow="7" pageId="26" pageNumber="27">0.23 3</td>
<td id="76E85DCEFFEF7CC1D182FC05D09FFB90" box="[1693,1785,984,1040]" gridcol="7" gridrow="7" pageId="26" pageNumber="27">0.11 3</td>
</tr>
<tr id="353934B2FFEF7CC1D768FBFDD09FFB2F" box="[119,1785,1056,1199]" gridrow="8" pageId="26" pageNumber="27">
<th id="76E85DCEFFEF7CC1D768FBFDD7C1FB2F" box="[119,423,1056,1199]" gridcol="0" gridrow="8" pageId="26" pageNumber="27">
Range N.S.W./ Mean Sth Queensland s.d.
<emphasis id="B97DEAE2FFEF8324D672FB47D71EFB2F" box="[365,376,1178,1199]" italics="true" pageId="26" pageNumber="27">n</emphasis>
</th>
<td id="76E85DCEFFEF7CC1D50BFBFDD4EEFB2F" box="[532,648,1056,1199]" gridcol="1" gridrow="8" pageId="26" pageNumber="27">14.3516.41 15.17 0.41 16</td>
<td id="76E85DCEFFEF7CC1D5C9FBFDD558FB2F" box="[726,830,1056,1199]" gridcol="2" gridrow="8" pageId="26" pageNumber="27">7.6710.42 8.99 0.71 15</td>
<td id="76E85DCEFFEF7CC1D488FBFDD26DFB2F" box="[919,1035,1056,1199]" gridcol="3" gridrow="8" pageId="26" pageNumber="27">16.6819.98 18.23 1.10 17</td>
<td id="76E85DCEFFEF7CC1D346FBFDD2D3FB2F" box="[1113,1205,1056,1199]" gridcol="4" gridrow="8" pageId="26" pageNumber="27">7.748.94 8.14 0.31 18</td>
<td id="76E85DCEFFEF7CC1D205FBFDD311FB2F" box="[1306,1399,1056,1199]" gridcol="5" gridrow="8" pageId="26" pageNumber="27">4.826.87 6.00 0.73 17</td>
<td id="76E85DCEFFEF7CC1D2C4FBFDD05EFB2F" box="[1499,1592,1056,1199]" gridcol="6" gridrow="8" pageId="26" pageNumber="27">3.203.62 3.21 0.18 14</td>
<td id="76E85DCEFFEF7CC1D182FBFDD09FFB2F" box="[1693,1785,1056,1199]" gridcol="7" gridrow="8" pageId="26" pageNumber="27">2.002.22 2.17 0.12 14</td>
</tr>
<tr id="353934B2FFEF7CC1D768FB62D09FFB53" box="[119,1785,1215,1235]" gridrow="9" pageId="26" pageNumber="27">
<th id="76E85DCEFFEF7CC1D768FB62D7C1FB53" box="[119,423,1215,1235]" gridcol="0" gridrow="9" pageId="26" pageNumber="27">Range</th>
<td id="76E85DCEFFEF7CC1D50BFB62D4EEFB53" box="[532,648,1215,1235]" gridcol="1" gridrow="9" pageId="26" pageNumber="27">14.5916.07</td>
<td id="76E85DCEFFEF7CC1D5C9FB62D558FB53" box="[726,830,1215,1235]" gridcol="2" gridrow="9" pageId="26" pageNumber="27">7.6210.20</td>
<td id="76E85DCEFFEF7CC1D488FB62D26DFB53" box="[919,1035,1215,1235]" gridcol="3" gridrow="9" pageId="26" pageNumber="27">15.1719.67</td>
<td id="76E85DCEFFEF7CC1D346FB62D2D3FB53" box="[1113,1205,1215,1235]" gridcol="4" gridrow="9" pageId="26" pageNumber="27">7.518.74</td>
<td id="76E85DCEFFEF7CC1D205FB62D311FB53" box="[1306,1399,1215,1235]" gridcol="5" gridrow="9" pageId="26" pageNumber="27">4.857.30</td>
<td id="76E85DCEFFEF7CC1D2C4FB62D05EFB53" box="[1499,1592,1215,1235]" gridcol="6" gridrow="9" pageId="26" pageNumber="27">3.023.66</td>
<td id="76E85DCEFFEF7CC1D182FB62D09FFB53" box="[1693,1785,1215,1235]" gridcol="7" gridrow="9" pageId="26" pageNumber="27">1.962.36</td>
</tr>
<tr id="353934B2FFEF7CC1D768FB3ED09FFB77" box="[119,1785,1251,1271]" gridrow="10" pageId="26" pageNumber="27">
<th id="76E85DCEFFEF7CC1D768FB3ED7C1FB77" box="[119,423,1251,1271]" gridcol="0" gridrow="10" pageId="26" pageNumber="27">NorthQueensland Mean</th>
<td id="76E85DCEFFEF7CC1D50BFB3ED4EEFB77" box="[532,648,1251,1271]" gridcol="1" gridrow="10" pageId="26" pageNumber="27">14.46</td>
<td id="76E85DCEFFEF7CC1D5C9FB3ED558FB77" box="[726,830,1251,1271]" gridcol="2" gridrow="10" pageId="26" pageNumber="27">8.95</td>
<td id="76E85DCEFFEF7CC1D488FB3ED26DFB77" box="[919,1035,1251,1271]" gridcol="3" gridrow="10" pageId="26" pageNumber="27">18.08</td>
<td id="76E85DCEFFEF7CC1D346FB3ED2D3FB77" box="[1113,1205,1251,1271]" gridcol="4" gridrow="10" pageId="26" pageNumber="27">7.60</td>
<td id="76E85DCEFFEF7CC1D205FB3ED311FB77" box="[1306,1399,1251,1271]" gridcol="5" gridrow="10" pageId="26" pageNumber="27">6.45</td>
<td id="76E85DCEFFEF7CC1D2C4FB3ED05EFB77" box="[1499,1592,1251,1271]" gridcol="6" gridrow="10" pageId="26" pageNumber="27">2.84</td>
<td id="76E85DCEFFEF7CC1D182FB3ED09FFB77" box="[1693,1785,1251,1271]" gridcol="7" gridrow="10" pageId="26" pageNumber="27">1.82</td>
</tr>
<tr id="353934B2FFEF7CC1D768FADAD09FFAE2" box="[119,1785,1287,1378]" gridrow="11" pageId="26" pageNumber="27">
<th id="76E85DCEFFEF7CC1D768FADAD7C1FAE2" box="[119,423,1287,1378]" gridcol="0" gridrow="11" pageId="26" pageNumber="27">
s.d.
<emphasis id="B97DEAE2FFEF8324D672FAF7D71EFABF" box="[365,376,1322,1343]" italics="true" pageId="26" pageNumber="27">n</emphasis>
Range
</th>
<td id="76E85DCEFFEF7CC1D50BFADAD4EEFAE2" box="[532,648,1287,1378]" gridcol="1" gridrow="11" pageId="26" pageNumber="27">1</td>
<td id="76E85DCEFFEF7CC1D5C9FADAD558FAE2" box="[726,830,1287,1378]" gridcol="2" gridrow="11" pageId="26" pageNumber="27">1</td>
<td id="76E85DCEFFEF7CC1D488FADAD26DFAE2" box="[919,1035,1287,1378]" gridcol="3" gridrow="11" pageId="26" pageNumber="27">1</td>
<td id="76E85DCEFFEF7CC1D346FADAD2D3FAE2" box="[1113,1205,1287,1378]" gridcol="4" gridrow="11" pageId="26" pageNumber="27">1</td>
<td id="76E85DCEFFEF7CC1D205FADAD311FAE2" box="[1306,1399,1287,1378]" gridcol="5" gridrow="11" pageId="26" pageNumber="27">1</td>
<td id="76E85DCEFFEF7CC1D2C4FADAD05EFAE2" box="[1499,1592,1287,1378]" gridcol="6" gridrow="11" pageId="26" pageNumber="27">1</td>
<td id="76E85DCEFFEF7CC1D182FADAD09FFAE2" box="[1693,1785,1287,1378]" gridcol="7" gridrow="11" pageId="26" pageNumber="27">1</td>
</tr>
<tr id="353934B2FFEF7CC1D768FAAFD09FFA06" box="[119,1785,1394,1414]" gridrow="12" pageId="26" pageNumber="27">
<th id="76E85DCEFFEF7CC1D768FAAFD7C1FA06" box="[119,423,1394,1414]" gridcol="0" gridrow="12" pageId="26" pageNumber="27">TopEnd',N.T. Mean</th>
<td id="76E85DCEFFEF7CC1D50BFAAFD4EEFA06" box="[532,648,1394,1414]" gridcol="1" gridrow="12" pageId="26" pageNumber="27">13.12</td>
<td id="76E85DCEFFEF7CC1D5C9FAAFD558FA06" box="[726,830,1394,1414]" gridcol="2" gridrow="12" pageId="26" pageNumber="27">8.73</td>
<td id="76E85DCEFFEF7CC1D488FAAFD26DFA06" box="[919,1035,1394,1414]" gridcol="3" gridrow="12" pageId="26" pageNumber="27">16.10</td>
<td id="76E85DCEFFEF7CC1D346FAAFD2D3FA06" box="[1113,1205,1394,1414]" gridcol="4" gridrow="12" pageId="26" pageNumber="27">7.07</td>
<td id="76E85DCEFFEF7CC1D205FAAFD311FA06" box="[1306,1399,1394,1414]" gridcol="5" gridrow="12" pageId="26" pageNumber="27">5.78</td>
<td id="76E85DCEFFEF7CC1D2C4FAAFD05EFA06" box="[1499,1592,1394,1414]" gridcol="6" gridrow="12" pageId="26" pageNumber="27">2.85</td>
<td id="76E85DCEFFEF7CC1D182FAAFD09FFA06" box="[1693,1785,1394,1414]" gridcol="7" gridrow="12" pageId="26" pageNumber="27">1.82</td>
</tr>
<tr id="353934B2FFEF7CC1D768FA4BD09FFA4E" box="[119,1785,1430,1486]" gridrow="13" pageId="26" pageNumber="27">
<th id="76E85DCEFFEF7CC1D768FA4BD7C1FA4E" box="[119,423,1430,1486]" gridcol="0" gridrow="13" pageId="26" pageNumber="27">
s.d.
<emphasis id="B97DEAE2FFEF8324D672FA64D71EFA4E" box="[365,376,1465,1486]" italics="true" pageId="26" pageNumber="27">n</emphasis>
</th>
<td id="76E85DCEFFEF7CC1D50BFA4BD4EEFA4E" box="[532,648,1430,1486]" gridcol="1" gridrow="13" pageId="26" pageNumber="27">0.47 4</td>
<td id="76E85DCEFFEF7CC1D5C9FA4BD558FA4E" box="[726,830,1430,1486]" gridcol="2" gridrow="13" pageId="26" pageNumber="27">0.51 4</td>
<td id="76E85DCEFFEF7CC1D488FA4BD26DFA4E" box="[919,1035,1430,1486]" gridcol="3" gridrow="13" pageId="26" pageNumber="27">1.00 4</td>
<td id="76E85DCEFFEF7CC1D346FA4BD2D3FA4E" box="[1113,1205,1430,1486]" gridcol="4" gridrow="13" pageId="26" pageNumber="27">0.73 4</td>
<td id="76E85DCEFFEF7CC1D205FA4BD311FA4E" box="[1306,1399,1430,1486]" gridcol="5" gridrow="13" pageId="26" pageNumber="27">0.84 5</td>
<td id="76E85DCEFFEF7CC1D2C4FA4BD05EFA4E" box="[1499,1592,1430,1486]" gridcol="6" gridrow="13" pageId="26" pageNumber="27">0.21 3</td>
<td id="76E85DCEFFEF7CC1D182FA4BD09FFA4E" box="[1693,1785,1430,1486]" gridcol="7" gridrow="13" pageId="26" pageNumber="27">0.09 3</td>
</tr>
<tr id="353934B2FFEF7CC1D768FA03D09FFA72" box="[119,1785,1502,1522]" gridrow="14" pageId="26" pageNumber="27">
<th id="76E85DCEFFEF7CC1D768FA03D7C1FA72" box="[119,423,1502,1522]" gridcol="0" gridrow="14" pageId="26" pageNumber="27">Range</th>
<td id="76E85DCEFFEF7CC1D50BFA03D4EEFA72" box="[532,648,1502,1522]" gridcol="1" gridrow="14" pageId="26" pageNumber="27">12.7113.62</td>
<td id="76E85DCEFFEF7CC1D5C9FA03D558FA72" box="[726,830,1502,1522]" gridcol="2" gridrow="14" pageId="26" pageNumber="27">7.989.12</td>
<td id="76E85DCEFFEF7CC1D488FA03D26DFA72" box="[919,1035,1502,1522]" gridcol="3" gridrow="14" pageId="26" pageNumber="27">14.7016.94</td>
<td id="76E85DCEFFEF7CC1D346FA03D2D3FA72" box="[1113,1205,1502,1522]" gridcol="4" gridrow="14" pageId="26" pageNumber="27">6.407.71</td>
<td id="76E85DCEFFEF7CC1D205FA03D311FA72" box="[1306,1399,1502,1522]" gridcol="5" gridrow="14" pageId="26" pageNumber="27">4.616.37</td>
<td id="76E85DCEFFEF7CC1D2C4FA03D05EFA72" box="[1499,1592,1502,1522]" gridcol="6" gridrow="14" pageId="26" pageNumber="27">2.623.02</td>
<td id="76E85DCEFFEF7CC1D182FA03D09FFA72" box="[1693,1785,1502,1522]" gridcol="7" gridrow="14" pageId="26" pageNumber="27">1.761.92</td>
</tr>
<tr id="353934B2FFEF7CC1D768F9DFD09FF996" box="[119,1785,1538,1558]" gridrow="15" pageId="26" pageNumber="27">
<th id="76E85DCEFFEF7CC1D768F9DFD7C1F996" box="[119,423,1538,1558]" gridcol="0" gridrow="15" pageId="26" pageNumber="27">Kimberley,W.A. Mean</th>
<td id="76E85DCEFFEF7CC1D50BF9DFD4EEF996" box="[532,648,1538,1558]" gridcol="1" gridrow="15" pageId="26" pageNumber="27">14.04</td>
<td id="76E85DCEFFEF7CC1D5C9F9DFD558F996" box="[726,830,1538,1558]" gridcol="2" gridrow="15" pageId="26" pageNumber="27">8.64</td>
<td id="76E85DCEFFEF7CC1D488F9DFD26DF996" box="[919,1035,1538,1558]" gridcol="3" gridrow="15" pageId="26" pageNumber="27">17.14</td>
<td id="76E85DCEFFEF7CC1D346F9DFD2D3F996" box="[1113,1205,1538,1558]" gridcol="4" gridrow="15" pageId="26" pageNumber="27">7.88</td>
<td id="76E85DCEFFEF7CC1D205F9DFD311F996" box="[1306,1399,1538,1558]" gridcol="5" gridrow="15" pageId="26" pageNumber="27">5.58</td>
<td id="76E85DCEFFEF7CC1D2C4F9DFD05EF996" box="[1499,1592,1538,1558]" gridcol="6" gridrow="15" pageId="26" pageNumber="27">2.84</td>
<td id="76E85DCEFFEF7CC1D182F9DFD09FF996" box="[1693,1785,1538,1558]" gridcol="7" gridrow="15" pageId="26" pageNumber="27">1.84</td>
</tr>
<tr id="353934B2FFEF7CC1D768F9FBD09FF901" box="[119,1785,1574,1665]" gridrow="16" pageId="26" pageNumber="27">
<th id="76E85DCEFFEF7CC1D768F9FBD7C1F901" box="[119,423,1574,1665]" gridcol="0" gridrow="16" pageId="26" pageNumber="27">
s.d.
<emphasis id="B97DEAE2FFEF8324D672F994D71EF9DE" box="[365,376,1609,1630]" italics="true" pageId="26" pageNumber="27">n</emphasis>
Range
</th>
<td id="76E85DCEFFEF7CC1D50BF9FBD4EEF901" box="[532,648,1574,1665]" gridcol="1" gridrow="16" pageId="26" pageNumber="27">0.76 2 13.514.57</td>
<td id="76E85DCEFFEF7CC1D5C9F9FBD558F901" box="[726,830,1574,1665]" gridcol="2" gridrow="16" pageId="26" pageNumber="27">0.45 2 8.328.96</td>
<td id="76E85DCEFFEF7CC1D488F9FBD26DF901" box="[919,1035,1574,1665]" gridcol="3" gridrow="16" pageId="26" pageNumber="27">0.43 2 16.8317.44</td>
<td id="76E85DCEFFEF7CC1D346F9FBD2D3F901" box="[1113,1205,1574,1665]" gridcol="4" gridrow="16" pageId="26" pageNumber="27">0.04 2 7.857.90</td>
<td id="76E85DCEFFEF7CC1D205F9FBD311F901" box="[1306,1399,1574,1665]" gridcol="5" gridrow="16" pageId="26" pageNumber="27">0.60 3 4.436.42</td>
<td id="76E85DCEFFEF7CC1D2C4F9FBD05EF901" box="[1499,1592,1574,1665]" gridcol="6" gridrow="16" pageId="26" pageNumber="27">1</td>
<td id="76E85DCEFFEF7CC1D182F9FBD09FF901" box="[1693,1785,1574,1665]" gridcol="7" gridrow="16" pageId="26" pageNumber="27">1</td>
</tr>
<tr id="353934B2FFEF7CC1D768F94CD09FF925" box="[119,1785,1681,1701]" gridrow="17" pageId="26" pageNumber="27">
<th id="76E85DCEFFEF7CC1D768F94CD7C1F925" box="[119,423,1681,1701]" gridcol="0" gridrow="17" pageId="26" pageNumber="27">TOTAL</th>
<td id="76E85DCEFFEF7CC1D50BF94CD4EEF925" box="[532,648,1681,1701]" gridcol="1" gridrow="17" pageId="26" pageNumber="27">44</td>
<td id="76E85DCEFFEF7CC1D5C9F94CD558F925" box="[726,830,1681,1701]" gridcol="2" gridrow="17" pageId="26" pageNumber="27">43</td>
<td id="76E85DCEFFEF7CC1D488F94CD26DF925" box="[919,1035,1681,1701]" gridcol="3" gridrow="17" pageId="26" pageNumber="27">45</td>
<td id="76E85DCEFFEF7CC1D346F94CD2D3F925" box="[1113,1205,1681,1701]" gridcol="4" gridrow="17" pageId="26" pageNumber="27">46</td>
<td id="76E85DCEFFEF7CC1D205F94CD311F925" box="[1306,1399,1681,1701]" gridcol="5" gridrow="17" pageId="26" pageNumber="27">47</td>
<td id="76E85DCEFFEF7CC1D2C4F94CD05EF925" box="[1499,1592,1681,1701]" gridcol="6" gridrow="17" pageId="26" pageNumber="27">31</td>
<td id="76E85DCEFFEF7CC1D182F94CD09FF925" box="[1693,1785,1681,1701]" gridcol="7" gridrow="17" pageId="26" pageNumber="27">31</td>
</tr>
</table>
</paragraph>
<paragraph id="8BB636F0FFEE8325D762FD5DD0DFFC81" pageId="27" pageNumber="28">
<table id="F909C450FFEE7CC1D762FD5DD121FC81" box="[125,1863,640,769]" gridcols="8" gridrows="3" pageId="27" pageNumber="28" rowsContinueIn="28.[119,1785,700,829]">
<tr id="353934B2FFEE7CC1D762FD5DD121FD14" box="[125,1863,640,660]" gridrow="0" pageId="27" pageNumber="28">
<th id="76E85DCEFFEE7CC1D762FD5DD7C3FD14" box="[125,421,640,660]" gridcol="0" gridrow="0" pageId="27" pageNumber="28">Geographic region</th>
<th id="76E85DCEFFEE7CC1D506FD5DD42BFD14" box="[537,589,640,660]" gridcol="1" gridrow="0" pageId="27" pageNumber="28">SL</th>
<th id="76E85DCEFFEE7CC1D5C5FD5DD569FD14" box="[730,783,640,660]" gridcol="2" gridrow="0" pageId="27" pageNumber="28">ZW</th>
<th id="76E85DCEFFEE7CC1D483FD5DD5B7FD14" box="[924,977,640,660]" gridcol="3" gridrow="0" pageId="27" pageNumber="28">OBW</th>
<th id="76E85DCEFFEE7CC1D342FD5DD2E3FD14" box="[1117,1157,640,660]" gridcol="4" gridrow="0" pageId="27" pageNumber="28">BS</th>
<th id="76E85DCEFFEE7CC1D200FD5DD333FD14" box="[1311,1365,640,660]" gridcol="5" gridrow="0" pageId="27" pageNumber="28">FMW</th>
<th id="76E85DCEFFEE7CC1D2FFFD5DD043FD14" box="[1504,1573,640,660]" gridcol="6" gridrow="0" pageId="27" pageNumber="28">C-M4L</th>
<th id="76E85DCEFFEE7CC1D1BDFD5DD121FD14" box="[1698,1863,640,660]" gridcol="7" gridrow="0" pageId="27" pageNumber="28">M1-4L</th>
</tr>
<tr id="353934B2FFEE7CC1D762FD78D121FD39" box="[125,1863,677,697]" gridrow="1" pageId="27" pageNumber="28">
<th id="76E85DCEFFEE7CC1D762FD78D7C3FD39" box="[125,421,677,697]" gridcol="0" gridrow="1" pageId="27" pageNumber="28">SouthwestW.A. Mean</th>
<td id="76E85DCEFFEE7CC1D506FD78D42BFD39" box="[537,589,677,697]" gridcol="1" gridrow="1" pageId="27" pageNumber="28">44.84</td>
<td id="76E85DCEFFEE7CC1D5C5FD78D569FD39" box="[730,783,677,697]" gridcol="2" gridrow="1" pageId="27" pageNumber="28">26.34</td>
<td id="76E85DCEFFEE7CC1D483FD78D5B7FD39" box="[924,977,677,697]" gridcol="3" gridrow="1" pageId="27" pageNumber="28">18.54</td>
<td id="76E85DCEFFEE7CC1D342FD78D2E3FD39" box="[1117,1157,677,697]" gridcol="4" gridrow="1" pageId="27" pageNumber="28">5.46</td>
<td id="76E85DCEFFEE7CC1D200FD78D333FD39" box="[1311,1365,677,697]" gridcol="5" gridrow="1" pageId="27" pageNumber="28">6.73</td>
<td id="76E85DCEFFEE7CC1D2FFFD78D043FD39" box="[1504,1573,677,697]" gridcol="6" gridrow="1" pageId="27" pageNumber="28">17.29</td>
<td id="76E85DCEFFEE7CC1D1BDFD78D121FD39" box="[1698,1863,677,697]" gridcol="7" gridrow="1" pageId="27" pageNumber="28">10.26</td>
</tr>
<tr id="353934B2FFEE7CC1D762FD14D121FC81" box="[125,1863,713,769]" gridrow="2" pageId="27" pageNumber="28">
<th id="76E85DCEFFEE7CC1D762FD14D7C3FC81" box="[125,421,713,769]" gridcol="0" gridrow="2" pageId="27" pageNumber="28">
s.d.
<emphasis id="B97DEAE2FFEE8325D66EFD31D71AFC81" box="[369,380,748,769]" italics="true" pageId="27" pageNumber="28">n</emphasis>
</th>
<td id="76E85DCEFFEE7CC1D506FD14D42BFC81" box="[537,589,713,769]" gridcol="1" gridrow="2" pageId="27" pageNumber="28">1.15 9</td>
<td id="76E85DCEFFEE7CC1D5C5FD14D569FC81" box="[730,783,713,769]" gridcol="2" gridrow="2" pageId="27" pageNumber="28">1.18 9</td>
<td id="76E85DCEFFEE7CC1D483FD14D5B7FC81" box="[924,977,713,769]" gridcol="3" gridrow="2" pageId="27" pageNumber="28">0.20 9</td>
<td id="76E85DCEFFEE7CC1D342FD14D2E3FC81" box="[1117,1157,713,769]" gridcol="4" gridrow="2" pageId="27" pageNumber="28">0.35 9</td>
<td id="76E85DCEFFEE7CC1D200FD14D333FC81" box="[1311,1365,713,769]" gridcol="5" gridrow="2" pageId="27" pageNumber="28">0.13 8</td>
<td id="76E85DCEFFEE7CC1D2FFFD14D043FC81" box="[1504,1573,713,769]" gridcol="6" gridrow="2" pageId="27" pageNumber="28">0.28 9</td>
<td id="76E85DCEFFEE7CC1D1BDFD14D121FC81" box="[1698,1863,713,769]" gridcol="7" gridrow="2" pageId="27" pageNumber="28">0.20 10</td>
</tr>
</table>
</paragraph>
<paragraph id="8BB636F0FFEE8325D762FCE8D173FC35" blockId="27.[125,1863,785,1611]" pageId="27" pageNumber="28">
SouthAustralia Mean 47.33 26.96 19.00 5.37 6.95 18.66
<geoCoordinate id="EE3D5037FFEE8325D1BDFCE8D71DFCED" degrees="11.01" direction="south" orientation="latitude" pageId="27" pageNumber="28" precision="555" value="-11.01">11.01 s</geoCoordinate>
.d. 1.14 1.22 0.76 0.22
<geoCoordinate id="EE3D5037FFEE8325D1BDFC84D7E6FC11" degrees="0.08" direction="north" orientation="latitude" pageId="27" pageNumber="28" precision="555" value="0.08">
0.08
<geoCoordinate id="EE3D5037FFEE8325D66EFCA1D7E6FC11" box="[369,384,892,913]" direction="north" orientation="latitude" pageId="27" pageNumber="28" precision="15" value="1.0338888">
<emphasis id="B97DEAE2FFEE8325D66EFCA1D7E6FC11" box="[369,384,892,913]" italics="true" pageId="27" pageNumber="28">N</emphasis>
</geoCoordinate>
</geoCoordinate>
1 2 2 2 1 2 2 Range 26.1527.76 18.1419.86 4.835.90 18.518.81 10.9511.07
</paragraph>
<paragraph id="8BB636F0FFEE8325D762FC19D173FBC4" blockId="27.[125,1863,785,1611]" pageId="27" pageNumber="28">
Victoria Mean 46.64 27.20 18.42 5.70 6.65 18.10
<geoCoordinate id="EE3D5037FFEE8325D1BDFC19D71DFC7C" degrees="10.69" direction="south" orientation="latitude" pageId="27" pageNumber="28" precision="555" value="-10.69">10.69 s</geoCoordinate>
.d. 1.16 1.19 0.74 0.48 0.39 0.34
<geoCoordinate id="EE3D5037FFEE8325D1BDFC35D7E6FBA0" degrees="0.35" direction="north" orientation="latitude" pageId="27" pageNumber="28" precision="555" value="0.35">
0.35
<geoCoordinate id="EE3D5037FFEE8325D66EFBD6D7E6FBA0" box="[369,384,1035,1056]" direction="north" orientation="latitude" pageId="27" pageNumber="28" precision="15" value="6.1016665">
<emphasis id="B97DEAE2FFEE8325D66EFBD6D7E6FBA0" box="[369,384,1035,1056]" italics="true" pageId="27" pageNumber="28">N</emphasis>
</geoCoordinate>
</geoCoordinate>
6 6 6 6 6 6 7 Range 45.0647.75 25.1418.11 17.5119.42 5.226.37 6.067.23 17.6118.52 10.1911.07
</paragraph>
<paragraph id="8BB636F0FFEE8325D790FB89D0B0FBE8" blockId="27.[125,1863,785,1611]" box="[143,1750,1108,1128]" pageId="27" pageNumber="28">.S.W./Sth Mean 44.29 26.24 18.52 5.35 6.51 16.75 10.24</paragraph>
<paragraph id="8BB636F0FFEE8325D762FBB3D68FFB02" blockId="27.[125,1863,785,1611]" box="[125,233,1134,1154]" pageId="27" pageNumber="28">Queensland</paragraph>
<paragraph id="8BB636F0FFEE8325D66EFB55D16FFB63" blockId="27.[125,1863,785,1611]" pageId="27" pageNumber="28">
s.d. 1.07 0.45 0.32 0.44 0.22 1.41
<geoCoordinate id="EE3D5037FFEE8325D1BDFB55D7E6FB40" degrees="0.30" direction="north" orientation="latitude" pageId="27" pageNumber="28" precision="555" value="0.3">
0.30
<geoCoordinate id="EE3D5037FFEE8325D66EFB76D7E6FB40" box="[369,384,1195,1216]" direction="north" orientation="latitude" pageId="27" pageNumber="28" precision="15" value="8.152499">
<emphasis id="B97DEAE2FFEE8325D66EFB76D7E6FB40" box="[369,384,1195,1216]" italics="true" pageId="27" pageNumber="28">N</emphasis>
</geoCoordinate>
</geoCoordinate>
8 9 9 9 9 9 9 Range 42.0545.5 25.6327.10 18.018.97 4.906.32 6.106.80 16.3918.20 9.7910.70
</paragraph>
<paragraph id="8BB636F0FFEE8325D79EFB2ED098FAF3" blockId="27.[125,1863,785,1611]" pageId="27" pageNumber="28">
TopEnd',N.T. Mean 41.29 23.80 16.14 4.80 5.98 16.08
<geoCoordinate id="EE3D5037FFEE8325D1BDFB2ED71DFAAB" degrees="9.28" direction="south" orientation="latitude" pageId="27" pageNumber="28" precision="555" value="-9.28">9.28 s</geoCoordinate>
.d. 1.10 0.85 0.71 0.40 0.26 0.34
<geoCoordinate id="EE3D5037FFEE8325D1BDFACAD71AFACF" degrees="0.17" direction="north" orientation="latitude" pageId="27" pageNumber="28" precision="555" value="0.17">
0.17
<geoCoordinate id="EE3D5037FFEE8325D66EFAE7D71AFACF" box="[369,380,1338,1359]" direction="north" orientation="latitude" pageId="27" pageNumber="28" precision="15" value="6.1016665">
<emphasis id="B97DEAE2FFEE8325D66EFAE7D71AFACF" box="[369,380,1338,1359]" italics="true" pageId="27" pageNumber="28">n</emphasis>
</geoCoordinate>
</geoCoordinate>
6 6 6 6 6 6 6 Range 40.0942.66 22.4324.67 15.0516.98 4.285.24 5.686.46 15.4916.38 9.049.50
</paragraph>
<paragraph id="8BB636F0FFEE8325D762FA5ED7F7FA3B" blockId="27.[125,1863,785,1611]" pageId="27" pageNumber="28">
Kimberley,W.A. Mean 41.51 16.66 5.42 6.70 16.58
<geoCoordinate id="EE3D5037FFEE8325D1BDFA5ED71CFA3B" degrees="10.19" direction="south" orientation="latitude" pageId="27" pageNumber="28" precision="555" value="-10.19">10.19 s</geoCoordinate>
.d.
</paragraph>
<paragraph id="8BB636F0FFEE8325D66EFA17D7CDF982" blockId="27.[125,1863,785,1611]" pageId="27" pageNumber="28">
<geoCoordinate id="EE3D5037FFEE8325D66EFA17D20FFA5E" box="[369,1129,1482,1503]" degrees="1" direction="north" minutes="1" orientation="latitude" pageId="27" pageNumber="28" precision="15" seconds="1" value="1.0169444">
<emphasis id="B97DEAE2FFEE8325D66EFA17D71AFA5F" box="[369,380,1482,1503]" italics="true" pageId="27" pageNumber="28">n</emphasis>
1 1 1
</geoCoordinate>
1 1 1 Range
</paragraph>
<paragraph id="8BB636F0FFEE8325D762F9CFD0DFF9A6" blockId="27.[125,1863,785,1611]" box="[125,1721,1554,1574]" pageId="27" pageNumber="28">
<collectionCode id="ED18AE35FFEE8325D762F9CFD6A1F9A6" box="[125,199,1554,1574]" pageId="27" pageNumber="28">TOTAL</collectionCode>
31 32 33 33 31 33 35
</paragraph>
<paragraph id="8BB636F0FFEE8322D13EF9EBD09EFCE1" blockId="27.[125,1863,785,1611]" lastBlockId="28.[119,1785,845,1634]" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28">
<emphasis id="B97DEAE2FFEE8325D13EF9EBD121F9CB" box="[1569,1863,1590,1611]" italics="true" pageId="27" pageNumber="28">…….continued on the next page</emphasis>
<emphasis id="B97DEAE2FFE98322D768FD5AD68EFD1B" bold="true" box="[119,232,647,667]" pageId="28" pageNumber="29">
<tableCitation id="C68B034BFFE98322D768FD5AD6B0FD1B" box="[119,214,647,667]" captionStart="TABLE 5" captionStartId="25.[125,203,525,545]" captionText="TABLE 5. Cranial and dental measurements of each taxon and major regional population within the P. tapoatafa group, presented separately for males (5 a) and females (5 b). Data are presented separately for three regional populations within nominotypical P. tapoatafa: 1) South Australia; 2) Victoria; 3) New South Wales and southern Queensland. Measurements of holotypes are also presented." pageId="28" pageNumber="29">
<collectionCode id="ED18AE35FFE98322D768FD5AD6A3FD1B" box="[119,197,647,667]" pageId="28" pageNumber="29">TABLE</collectionCode>
5
</tableCitation>
b.
</emphasis>
(Continued) Range 13.7815.46 7.698.90 16.1218.81 7.948.94 4.756.03 2.803.10 1.762.02
</paragraph>
<paragraph id="8BB636F0FFE98322D768FD61D0CEFCBD" pageId="28" pageNumber="29">
<table id="F909C450FFE97CC1D768FD61D09FFCBD" box="[119,1785,700,829]" gridcols="8" gridrows="3" pageId="28" pageNumber="29" rowsContinueFrom="27.[125,1863,640,769]">
<tr id="353934B2FFE97CC1D768FD61D09FFD50" box="[119,1785,700,720]" gridrow="0" pageId="28" pageNumber="29">
<th id="76E85DCEFFE97CC1D768FD61D7C6FD50" box="[119,416,700,720]" gridcol="0" gridrow="0" pageId="28" pageNumber="29">Geographic region</th>
<th id="76E85DCEFFE97CC1D50CFD61D435FD50" box="[531,595,700,720]" gridcol="1" gridrow="0" pageId="28" pageNumber="29">PALW</th>
<th id="76E85DCEFFE97CC1D5CAFD61D56CFD50" box="[725,778,700,720]" gridcol="2" gridrow="0" pageId="28" pageNumber="29">ARW</th>
<th id="76E85DCEFFE97CC1D489FD61D5ACFD50" box="[918,970,700,720]" gridcol="3" gridrow="0" pageId="28" pageNumber="29">PRW</th>
<th id="76E85DCEFFE97CC1D347FD61D2E3FD50" box="[1112,1157,700,720]" gridcol="4" gridrow="0" pageId="28" pageNumber="29">IOW</th>
<th id="76E85DCEFFE97CC1D206FD61D327FD50" box="[1305,1345,700,720]" gridcol="5" gridrow="0" pageId="28" pageNumber="29">RD</th>
<th id="76E85DCEFFE97CC1D2C4FD61D060FD50" box="[1499,1542,700,720]" gridcol="6" gridrow="0" pageId="28" pageNumber="29">LCL</th>
<th id="76E85DCEFFE97CC1D183FD61D09FFD50" box="[1692,1785,700,720]" gridcol="7" gridrow="0" pageId="28" pageNumber="29">LCW</th>
</tr>
<tr id="353934B2FFE97CC1D768FD3CD09FFD75" box="[119,1785,737,757]" gridrow="1" pageId="28" pageNumber="29">
<th id="76E85DCEFFE97CC1D768FD3CD7C6FD75" box="[119,416,737,757]" gridcol="0" gridrow="1" pageId="28" pageNumber="29">SouthwestW.A. Mean</th>
<td id="76E85DCEFFE97CC1D50CFD3CD435FD75" box="[531,595,737,757]" gridcol="1" gridrow="1" pageId="28" pageNumber="29">14.62</td>
<td id="76E85DCEFFE97CC1D5CAFD3CD56CFD75" box="[725,778,737,757]" gridcol="2" gridrow="1" pageId="28" pageNumber="29">8.36</td>
<td id="76E85DCEFFE97CC1D489FD3CD5ACFD75" box="[918,970,737,757]" gridcol="3" gridrow="1" pageId="28" pageNumber="29">17.91</td>
<td id="76E85DCEFFE97CC1D347FD3CD2E3FD75" box="[1112,1157,737,757]" gridcol="4" gridrow="1" pageId="28" pageNumber="29">8.24</td>
<td id="76E85DCEFFE97CC1D206FD3CD327FD75" box="[1305,1345,737,757]" gridcol="5" gridrow="1" pageId="28" pageNumber="29">5.16</td>
<td id="76E85DCEFFE97CC1D2C4FD3CD060FD75" box="[1499,1542,737,757]" gridcol="6" gridrow="1" pageId="28" pageNumber="29">2.93</td>
<td id="76E85DCEFFE97CC1D183FD3CD09FFD75" box="[1692,1785,737,757]" gridcol="7" gridrow="1" pageId="28" pageNumber="29">1.91</td>
</tr>
<tr id="353934B2FFE97CC1D768FCD8D09FFCBD" box="[119,1785,773,829]" gridrow="2" pageId="28" pageNumber="29">
<th id="76E85DCEFFE97CC1D768FCD8D7C6FCBD" box="[119,416,773,829]" gridcol="0" gridrow="2" pageId="28" pageNumber="29">
s.d.
<emphasis id="B97DEAE2FFE98322D673FCF5D711FCBD" box="[364,375,808,829]" italics="true" pageId="28" pageNumber="29">n</emphasis>
</th>
<td id="76E85DCEFFE97CC1D50CFCD8D435FCBD" box="[531,595,773,829]" gridcol="1" gridrow="2" pageId="28" pageNumber="29">0.55 9</td>
<td id="76E85DCEFFE97CC1D5CAFCD8D56CFCBD" box="[725,778,773,829]" gridcol="2" gridrow="2" pageId="28" pageNumber="29">0.34 9</td>
<td id="76E85DCEFFE97CC1D489FCD8D5ACFCBD" box="[918,970,773,829]" gridcol="3" gridrow="2" pageId="28" pageNumber="29">0.84 9</td>
<td id="76E85DCEFFE97CC1D347FCD8D2E3FCBD" box="[1112,1157,773,829]" gridcol="4" gridrow="2" pageId="28" pageNumber="29">0.31 9</td>
<td id="76E85DCEFFE97CC1D206FCD8D327FCBD" box="[1305,1345,773,829]" gridcol="5" gridrow="2" pageId="28" pageNumber="29">0.47 10</td>
<td id="76E85DCEFFE97CC1D2C4FCD8D060FCBD" box="[1499,1542,773,829]" gridcol="6" gridrow="2" pageId="28" pageNumber="29">0.12 5</td>
<td id="76E85DCEFFE97CC1D183FCD8D09FFCBD" box="[1692,1785,773,829]" gridcol="7" gridrow="2" pageId="28" pageNumber="29">0.11 5</td>
</tr>
</table>
</paragraph>
<paragraph id="8BB636F0FFE98322D768FCACD327FC05" blockId="28.[119,1785,845,1634]" box="[119,1345,881,901]" pageId="28" pageNumber="29">SouthAustralia Mean 15.39 8.39 17.85 8.32 5.25</paragraph>
<paragraph id="8BB636F0FFE98322D673FC48D327FC29" blockId="28.[119,1785,845,1634]" box="[364,1345,917,937]" pageId="28" pageNumber="29">s.d. 0.40 0.09 0.19 0.55</paragraph>
<paragraph id="8BB636F0FFE98322D673FC65D343FC4D" blockId="28.[119,1785,845,1634]" box="[364,1317,952,973]" pageId="28" pageNumber="29">
<geoCoordinate id="EE3D5037FFE98322D673FC65D5C4FC4D" box="[364,930,952,973]" degrees="1" direction="north" minutes="2" orientation="latitude" pageId="28" pageNumber="29" precision="15" seconds="2" value="1.0338888">
<emphasis id="B97DEAE2FFE98322D673FC65D71DFC4D" box="[364,379,952,973]" italics="true" pageId="28" pageNumber="29">N</emphasis>
1 2 2
</geoCoordinate>
2 2
</paragraph>
<paragraph id="8BB636F0FFE98322D673FC00D310FC71" blockId="28.[119,1785,845,1634]" box="[364,1398,989,1009]" pageId="28" pageNumber="29">Range 8.118.67 17.7017.91 8.188.45 4.865.64</paragraph>
<paragraph id="8BB636F0FFE98322D768FBDDD09FFB00" blockId="28.[119,1785,845,1634]" pageId="28" pageNumber="29">
Victoria Mean 15.16 8.81 18.00 8.61 5.41 2.72
<geoCoordinate id="EE3D5037FFE98322D183FBDDD710FBB8" degrees="1.84" direction="south" orientation="latitude" pageId="28" pageNumber="29" precision="555" value="-1.84">1.84 s</geoCoordinate>
.d. 0.66 0.55 0.62 0.17 0.11 0.11
<geoCoordinate id="EE3D5037FFE98322D183FBF9D71DFBDC" degrees="0.03" direction="north" orientation="latitude" pageId="28" pageNumber="29" precision="555" value="0.03">
0.03
<geoCoordinate id="EE3D5037FFE98322D673FB9AD71DFBDC" box="[364,379,1095,1116]" direction="north" orientation="latitude" pageId="28" pageNumber="29" precision="15" value="6.1016665">
<emphasis id="B97DEAE2FFE98322D673FB9AD71DFBDC" box="[364,379,1095,1116]" italics="true" pageId="28" pageNumber="29">N</emphasis>
</geoCoordinate>
</geoCoordinate>
6 6 6 6 7 2 2 Range 14.3015.81 7.799.30 16.8218.63 8.448.80 4.935.84 2.622.80 1.821.86
</paragraph>
<paragraph id="8BB636F0FFE98322D793FB4DD0A2FB24" blockId="28.[119,1785,845,1634]" box="[140,1732,1168,1188]" pageId="28" pageNumber="29">S.W./Sth Mean 14.53 8.34 16.91 8.07 5.37 2.64 1.92</paragraph>
<paragraph id="8BB636F0FFE98322D768FB77D685FB3E" blockId="28.[119,1785,845,1634]" box="[119,227,1194,1214]" pageId="28" pageNumber="29">Queensland</paragraph>
<paragraph id="8BB636F0FFE98322D673FB19D09EFA9F" blockId="28.[119,1785,845,1634]" pageId="28" pageNumber="29">
s.d. 0.28 0.18 0.36 0.44 0.37 0.13
<geoCoordinate id="EE3D5037FFE98322D183FB19D71DFB7C" degrees="0.05" direction="north" orientation="latitude" pageId="28" pageNumber="29" precision="555" value="0.05">
0.05
<geoCoordinate id="EE3D5037FFE98322D673FB3AD71DFB7C" box="[364,379,1255,1276]" direction="north" orientation="latitude" pageId="28" pageNumber="29" precision="15" value="9.152499">
<emphasis id="B97DEAE2FFE98322D673FB3AD71DFB7C" box="[364,379,1255,1276]" italics="true" pageId="28" pageNumber="29">N</emphasis>
</geoCoordinate>
</geoCoordinate>
9 9 9 9 6 9 9 Range 13.9514.91 8.038.54 16.3117.50 7.429.00 4.815.78 2.502.86 1.822.00
</paragraph>
<paragraph id="8BB636F0FFE98322D764FAF2D09FFA2F" blockId="28.[119,1785,845,1634]" pageId="28" pageNumber="29">
TopEnd',N.T. Mean 12.64 7.38 14.57 6.82 5.18 2.23
<geoCoordinate id="EE3D5037FFE98322D183FAF2D710FAE7" degrees="1.54" direction="south" orientation="latitude" pageId="28" pageNumber="29" precision="555" value="-1.54">1.54 s</geoCoordinate>
.d. 0.43 0.54 0.50 0.17 0.32 0.12
<geoCoordinate id="EE3D5037FFE98322D183FA8ED711FA0B" degrees="0.05" direction="north" orientation="latitude" pageId="28" pageNumber="29" precision="555" value="0.05">
0.05
<geoCoordinate id="EE3D5037FFE98322D673FAABD711FA0B" box="[364,375,1398,1419]" direction="north" orientation="latitude" pageId="28" pageNumber="29" precision="15" value="6.1016665">
<emphasis id="B97DEAE2FFE98322D673FAABD711FA0B" box="[364,375,1398,1419]" italics="true" pageId="28" pageNumber="29">n</emphasis>
</geoCoordinate>
</geoCoordinate>
6 6 6 6 6 4 4 Range 11.8212.97 6.718.08 16.3117.50 6.617.05 4.685.60 2.122.40 1.501.58
</paragraph>
<paragraph id="8BB636F0FFE98322D768FA62D0A2FA53" blockId="28.[119,1785,845,1634]" box="[119,1732,1471,1491]" pageId="28" pageNumber="29">Kimberley,W.A. Mean 13.10 7.60 7.08 6.06 2.40 1.60</paragraph>
<paragraph id="8BB636F0FFE98322D673FA3ED7EAFA77" blockId="28.[119,1785,845,1634]" box="[364,396,1507,1527]" pageId="28" pageNumber="29">s.d.</paragraph>
<paragraph id="8BB636F0FFE98322D673F9DBD7C0F9BE" blockId="28.[119,1785,845,1634]" pageId="28" pageNumber="29">
<geoCoordinate id="EE3D5037FFE98322D673F9DBD202F99A" box="[364,1124,1542,1563]" degrees="1" direction="north" minutes="1" orientation="latitude" pageId="28" pageNumber="29" precision="15" seconds="1" value="1.0169444">
<emphasis id="B97DEAE2FFE98322D673F9DBD711F99B" box="[364,375,1542,1563]" italics="true" pageId="28" pageNumber="29">n</emphasis>
1 1 1
</geoCoordinate>
1 1 1 Range
</paragraph>
<paragraph id="8BB636F0FFE98323D768F993D161FD3C" blockId="28.[119,1785,845,1634]" lastBlockId="29.[125,1799,680,1169]" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29">
<collectionCode id="ED18AE35FFE98322D768F993D6A7F9E2" box="[119,193,1614,1634]" pageId="28" pageNumber="29">TOTAL</collectionCode>
32 33 32 33 32 21 21 Geographic region SL ZW OBW BS FMW C1-M4 M1-4
</paragraph>
<caption id="DF766678FFE88323D762FDAED3D5FD07" box="[125,1459,627,648]" pageId="29" pageNumber="30">
<paragraph id="8BB636F0FFE88323D762FDAED3D5FD07" blockId="29.[125,1459,627,648]" box="[125,1459,627,648]" pageId="29" pageNumber="30">
<emphasis id="B97DEAE2FFE88323D762FDAED687FD07" bold="true" box="[125,225,627,647]" pageId="29" pageNumber="30">TABLE 6.</emphasis>
Statistical results for 1-factor ANOVA tests for sexual dimorphism in cranio-dental measurements of the
<taxonomicName id="4C094D73FFE88323D3B7FDAED316FD08" box="[1192,1392,627,648]" class="Mammalia" family="Dasyuridae" genus="Phascogale" kingdom="Animalia" order="Dasyuromorphia" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="tapoatafa">
<emphasis id="B97DEAE2FFE88323D3B7FDAED316FD08" box="[1192,1392,627,648]" italics="true" pageId="29" pageNumber="30">Phascogale tapoatafa</emphasis>
</taxonomicName>
group.
</paragraph>
</caption>
<paragraph id="8BB636F0FFE88323D762FD10D097FD61" blockId="29.[125,1799,680,1169]" box="[125,1777,717,737]" pageId="29" pageNumber="30">Southwest W.A. F2 8.42 5.24 NS NS NS NS NS</paragraph>
<paragraph id="8BB636F0FFE88323D56EFD2CD5CBFC85" blockId="29.[125,1799,680,1169]" box="[625,941,753,773]" pageId="29" pageNumber="30">d.f. 1 1</paragraph>
<paragraph id="8BB636F0FFE88323D56EFCC8D5B3FCA9" blockId="29.[125,1799,680,1169]" box="[625,981,789,809]" pageId="29" pageNumber="30">
P
<geoCoordinate id="EE3D5037FFE88323D41BFCC8D55EFCA9" box="[772,824,789,809]" degrees="0.010" direction="north" orientation="latitude" pageId="29" pageNumber="30" precision="55" value="0.01">0.010</geoCoordinate>
<geoCoordinate id="EE3D5037FFE88323D4BEFCC8D5B3FCA9" box="[929,981,789,809]" degrees="0.035" direction="east" orientation="longitude" pageId="29" pageNumber="30" precision="55" value="0.035">0.035</geoCoordinate>
</paragraph>
<paragraph id="8BB636F0FFE88323D762FCE4D097FCCD" blockId="29.[125,1799,680,1169]" box="[125,1777,825,845]" pageId="29" pageNumber="30">South Australia / Victoria F2 NS NS NS NS NS NS NS</paragraph>
<paragraph id="8BB636F0FFE88323D56EFC80D4F6FCF1" blockId="29.[125,1799,680,1169]" box="[625,656,861,881]" pageId="29" pageNumber="30">d.f.</paragraph>
<paragraph id="8BB636F0FFE88323D56EFC5CD418FC15" blockId="29.[125,1799,680,1169]" box="[625,638,897,917]" pageId="29" pageNumber="30">P</paragraph>
<paragraph id="8BB636F0FFE88323D791FC78D097FC39" blockId="29.[125,1799,680,1169]" box="[142,1777,933,953]" pageId="29" pageNumber="30">.S.W./Southeast Queensland F2 15.99 15.23 NS 6.13 NS 7.93 NS</paragraph>
<paragraph id="8BB636F0FFE88323D56EFC15D00CFB80" blockId="29.[125,1799,680,1169]" pageId="29" pageNumber="30">d.f. 1 1 1 1 P 0.001 0.001 0.020 0.010</paragraph>
<paragraph id="8BB636F0FFE88323D79FFBCDD097FBA4" blockId="29.[125,1799,680,1169]" box="[128,1777,1040,1060]" pageId="29" pageNumber="30">Top End', Northern Territory F2 48.79 NS 8.08 NS NS 24.17 NS</paragraph>
<paragraph id="8BB636F0FFE88323D56EFBE9D00CFBEC" blockId="29.[125,1799,680,1169]" pageId="29" pageNumber="30">d.f. 1 1 1 P 0.000 0.022 0.001</paragraph>
<paragraph id="8BB636F0FFE88323D762FBA0D685FB11" blockId="29.[125,1799,680,1169]" box="[125,227,1149,1169]" pageId="29" pageNumber="30">Continued.</paragraph>
<paragraph id="8BB636F0FFE88323D762FB6FD09AFB46" blockId="29.[125,1790,1202,1654]" box="[125,1788,1202,1222]" pageId="29" pageNumber="30">
Geographic region
<collectionCode id="ED18AE35FFE88323D5F2FB6FD54BFB46" box="[749,813,1202,1222]" pageId="29" pageNumber="30">PALW</collectionCode>
ARW PRW IOW RD LCL LCW
</paragraph>
<paragraph id="8BB636F0FFE88323D762FB0AD081FB6B" blockId="29.[125,1790,1202,1654]" box="[125,1767,1239,1259]" pageId="29" pageNumber="30">Southwest W.A. F2 NS NS 6.36 NS 7.78 NS NS</paragraph>
<paragraph id="8BB636F0FFE88323D540FB26D3F8FA8F" blockId="29.[125,1790,1202,1654]" box="[607,1438,1275,1295]" pageId="29" pageNumber="30">d.f. 1 1</paragraph>
<paragraph id="8BB636F0FFE88323D540FAC2D3A0FAB3" blockId="29.[125,1790,1202,1654]" box="[607,1478,1311,1331]" pageId="29" pageNumber="30">
P
<geoCoordinate id="EE3D5037FFE88323D345FAC2D2E8FAB3" box="[1114,1166,1311,1331]" degrees="0.022" direction="north" orientation="latitude" pageId="29" pageNumber="30" precision="55" value="0.022">0.022</geoCoordinate>
<geoCoordinate id="EE3D5037FFE88323D28DFAC2D3A0FAB3" box="[1426,1478,1311,1331]" degrees="0.012" direction="east" orientation="longitude" pageId="29" pageNumber="30" precision="55" value="0.012">0.012</geoCoordinate>
</paragraph>
<paragraph id="8BB636F0FFE88323D762FA9ED098FA1E" blockId="29.[125,1790,1202,1654]" pageId="29" pageNumber="30">
South Australia / Victoria F2 NS NS NS NS NS 12.01 10.73 d.f. 1 1.00 P
<geoCoordinate id="EE3D5037FFE88323D12DFA57D000FA1E" box="[1586,1638,1418,1438]" degrees="0.041" direction="north" orientation="latitude" pageId="29" pageNumber="30" precision="55" value="0.041">0.041</geoCoordinate>
<geoCoordinate id="EE3D5037FFE88323D1D5FA57D098FA1E" box="[1738,1790,1418,1438]" degrees="0.047" direction="east" orientation="longitude" pageId="29" pageNumber="30" precision="55" value="0.047">0.047</geoCoordinate>
</paragraph>
<paragraph id="8BB636F0FFE88323D791FA73D098F98A" blockId="29.[125,1790,1202,1654]" pageId="29" pageNumber="30">.S.W./Southeast Queensland F2 17.25 7.34 12.09 NS NS 68.43 33.19 d.f. 1 1 1 1 1 P 0.000 0.013 0.002 0.000 0.000</paragraph>
<paragraph id="8BB636F0FFE88320D79FF9C7D4E5F8D6" blockId="29.[125,1790,1202,1654]" lastBlockId="30.[151,1437,1529,2022]" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30">Top End', Northern Territory F2 NS 15.74 10.69 NS NS 25.49 31.11 d.f. 1 1 1 1 P 0.004 0.011 0.004 0.003 The fur on the head is pale grey on the sides of the face and dark grey with diffuse frosting on the crown. Additional facial patterning includes a well-developed, dark russet mid-rostral stripe, a broad and indistinct, dark grey loreal stripe and small, black postorbital spot, distinct off-white infra- and supra-orbital crescents, and a variably expressed cream-tipped preauricular tuft. Preauricular patches are indistinct in around 30 % of specimens and seem to be less conspicuous in juveniles than in adults. In most specimens, the cream fur of the chin and throat extends around the corner of the mouth and merges into a supralabial bar, while the cheek patch is cream peppered with black. The rostral vibrissal pads are densely clothed in short grey fur and the basal papillae are unpigmented and not visible. All facial vibrissae are thick and black, sometimes with gingery tipping. The pinnae are moderately elongate and broad, weakly to moderately pigmented and sparsely furred. The anterior border of each pinna supports a narrow band of short black hairs.</paragraph>
<caption id="DF766678FFEB8320D788FAA4D5BEFA4D" httpUri="https://zenodo.org/record/240169/files/figure.png" pageId="30" pageNumber="31" targetBox="[151,1436,207,1382]" targetPageId="30">
<paragraph id="8BB636F0FFEB8320D788FAA4D5BEFA4D" blockId="30.[151,1436,1401,1485]" pageId="30" pageNumber="31">
<emphasis id="B97DEAE2FFEB8320D788FAA4D77FFA0E" bold="true" box="[151,281,1401,1423]" pageId="30" pageNumber="31">FIGURE 16</emphasis>
. Crania and dentaries of
<taxonomicName id="4C094D73FFEB8320D504FAA7D49BFA0F" box="[539,765,1401,1423]" class="Mammalia" family="Dasyuridae" genus="Phascogale" kingdom="Animalia" order="Dasyuromorphia" pageId="30" pageNumber="31" phylum="Chordata" rank="subSpecies" species="tapoatafa" subSpecies="tapoatafa">
<emphasis id="B97DEAE2FFEB8320D504FAA7D49BFA0F" box="[539,765,1401,1423]" italics="true" pageId="30" pageNumber="31">P. tapoatafa tapoatafa</emphasis>
</taxonomicName>
. a) CM24425, an adult male from Coraki, New South Wales, and b) CM1316, a subadult male from Wallaby Creek, near Urbanville, New South Wales. Each series consists of (left to right) dorsal view of cranium, ventral view of cranium, and lateral view of right dentary.
</paragraph>
</caption>
<paragraph id="8BB636F0FFEB8321D7D8F8BCD4EEFF77" blockId="30.[151,1437,1529,2022]" lastBlockId="31.[151,1437,151,608]" lastPageId="31" lastPageNumber="32" pageId="30" pageNumber="31">
The basal portion of the tail is thickly furred above and on the sides, and supports numerous projecting guard hairs, continuing the colour and texture of the rump (
<figureCitation id="13322A75FFEB8320D41CF858D52BF81E" box="[771,845,1925,1950]" captionStart="FIGURE 4" captionStartId="8.[151,250,1565,1587]" captionTargetBox="[336,1237,723,1532]" captionTargetId="figure@8.[324,1262,715,1543]" captionTargetPageId="8" captionText="FIGURE 4. Representative puppet skins of nominotypical Phascogale tapoatafa (CM 20990, an adult male from Coraki, New South Wales, in a) dorsal view and b) ventral view and P. t. pirata (CM 7225, an adult male from Nourlangie Rock, Arnhem Land, Northern Territory, in c) dorsal view and d) ventral view), illustrating the extremes of variation in body proportions and colouration among the Brush-tailed Phascogales." httpUri="https://zenodo.org/record/240157/files/figure.png" pageId="30" pageNumber="31">Figs 4</figureCitation>
and
<figureCitation id="13322A75FFEB8320D496F858D5FFF81E" box="[905,921,1925,1950]" captionStart="FIGURE 6" captionStartId="10.[151,250,971,993]" captionTargetBox="[151,1436,445,949]" captionTargetId="figure@10.[151,1436,445,949]" captionTargetPageId="10" captionText="FIGURE 6. Ventral surface of the tail in representatives of four populations of the Phascogale tapoatafa group; a) nominotypical P. tapoatafa (CM 20990); b) P. t. pirata (CM 7225); c) P. taoatafa southwestern W. A. (WAM 5006, from Balingup Brook); d) P. tapoatafa Kimblerley, W. A. (WAM 16028, from Pago Mission)." httpUri="https://zenodo.org/record/240159/files/figure.png" pageId="30" pageNumber="31">6</figureCitation>
). The tail narrows rapidly to form a distinct stem, which is clothed above and below by long, adpressed hairs. Dorsally and around the sides of the tail, these hairs are white with short black tips, while those on the ventral surface are black with short cream tips grading to dark russet at the base of the brush. The tail brush is lush and measures up to
<quantity id="4CF19B15FFEA8321D31CFF4AD204FF2F" box="[1027,1122,151,176]" metricMagnitude="-1" metricUnit="m" metricValue="1.65" pageId="31" pageNumber="32" unit="mm" value="165.0">165 mm</quantity>
; it consists of jet black hairs except for a short section of dark russet hairs ventrally, at the base of the brush. The basal portion of the tail, from cloaca to base of brush, measures to
<quantity id="4CF19B15FFEA8321D52EFF3DD4E5FF77" box="[561,643,224,248]" metricMagnitude="-2" metricUnit="m" metricValue="9.0" pageId="31" pageNumber="32" unit="mm" value="90.0">90 mm</quantity>
.
</paragraph>
<paragraph id="8BB636F0FFEA8321D7D8FED9D481FD98" blockId="31.[151,1437,151,608]" pageId="31" pageNumber="32">
The upper surface of the pes supports a mixture of white, dark grey and russet hairs, giving it an overall grizzled appearance. This merges into white fur on the sides of the foot and around the heel. The digits are densely clothed in white hairs. The plantar surface of the pes is granular and supports a total of five conspicuously striated pads. The inner metatarsal and first interdigital pads are fused, usually with no obvious line of demarcation; their combined length is
<quantity id="4CF19B15FFEA8321D669FE49D7B9FE2C" box="[374,479,404,428]" metricMagnitude="-2" metricUnit="m" metricValue="1.31" pageId="31" pageNumber="32" unit="mm" value="13.1">13.1 mm</quantity>
. The outer metatarsal pad is moderately long; on AM932 it measures
<quantity id="4CF19B15FFEA8321D3E5FE49D337FE2C" box="[1274,1361,404,428]" metricMagnitude="-3" metricUnit="m" metricValue="7.4" pageId="31" pageNumber="32" unit="mm" value="7.4">7.4 mm</quantity>
and is separated by a gap of
<quantity id="4CF19B15FFEA8321D68EFE65D78FFE4F" box="[401,489,440,464]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="31" pageNumber="32" unit="mm" value="3.9">3.9 mm</quantity>
from the fourth interdigital pad. The second to fourth interdigital pads are elongate and of subequal length. Both feet on one specimen (
<collectionCode id="ED18AE35FFEA8321D418FE01D553FE75" box="[775,821,476,501]" pageId="31" pageNumber="32">CM</collectionCode>
1316) show a clear line of demarcation between the conjoined inner metatarsal and first interdigital pads.
</paragraph>
<paragraph id="8BB636F0FFEA8321D7D8FDF9D433FDDF" blockId="31.[151,1437,151,608]" pageId="31" pageNumber="32">
Females in all populations usually have eight teats in the pouch (e.g.
<bibRefCitation id="EF984B01FFEA8321D33FFDF9D2C8FDBC" author="Cuttle" box="[1056,1198,548,573]" pageId="31" pageNumber="32" refString="Cuttle, P. (1982) Life history strategy of the dasyurid marsupial Phascogale tapoatafa. In: Archer, M (Ed.), Carnivorous Marsupials. Royal Zoological Society of New South Wales. Surrey Beatty, NSW, pp. 13 - 22." type="book chapter" year="1982">Cuttle 1982</bibRefCitation>
;
<bibRefCitation id="EF984B01FFEA8321D3DEFDF9D3E9FDBC" author="Soderquist" box="[1217,1423,548,573]" pageId="31" pageNumber="32" refString="Soderquist, T. R. (1993 a) Maternal strategies of Phascogale tapoatafa (Marsupialia: Dasyuridae). I. Breeding seasonality and maternal investment. Australian Journal of Zoology, 41, 549 - 566." type="journal article" year="1993" yearSuffix="a">Soderquist 1993a</bibRefCitation>
); occasional individuals have seven teats.
</paragraph>
<caption id="DF766678FFEA8321D788F8E5D584F84E" httpUri="https://zenodo.org/record/240170/files/figure.png" pageId="31" pageNumber="32" targetBox="[155,1436,677,1804]" targetPageId="31">
<paragraph id="8BB636F0FFEA8321D788F8E5D584F84E" blockId="31.[151,1436,1848,1998]" pageId="31" pageNumber="32">
<emphasis id="B97DEAE2FFEA8321D788F8E5D77FF8CD" bold="true" box="[151,281,1848,1870]" pageId="31" pageNumber="32">FIGURE 17</emphasis>
. Graphical representation of geographic variation within
<taxonomicName id="4C094D73FFEA8321D479F8E7D593F8CE" box="[870,1013,1848,1871]" class="Mammalia" family="Dasyuridae" genus="Phascogale" kingdom="Animalia" order="Dasyuromorphia" pageId="31" pageNumber="32" phylum="Chordata" rank="subSpecies" species="tapoatafa" subSpecies="tapoatafa">
<emphasis id="B97DEAE2FFEA8321D479F8E7D593F8CE" box="[870,1013,1848,1871]" italics="true" pageId="31" pageNumber="32">P. t. tapoatafa</emphasis>
</taxonomicName>
in selected craniodental dimensions (SL; M1-4; FMW). Values for individual specimens are plotted in geographic sequence (Geo Seq) from west to east for South Australia and Victoria, and then from south to north for New South Wales and southeast Queensland.
<emphasis id="B97DEAE2FFEA8321D3DAF8A5D31FF80E" box="[1221,1401,1912,1934]" italics="true" pageId="31" pageNumber="32">Population codes</emphasis>
: 2: South Australia; 3: Victoria; 4: New South Wales and southeast Queensland. Plotted variables are a) Skull Length; b) M1-4; c) Foramen Magnum Width. Left hand series are females; right hand series are males.
</paragraph>
</caption>
<paragraph id="8BB636F0FFD5831ED7D8FF4AD2A2FF55" blockId="32.[151,1437,151,2018]" pageId="32" pageNumber="33">
Cranial and mandibular anatomy of a fully adult male (
<collectionCode id="ED18AE35FFD5831ED42FFF4AD53BFF30" box="[816,861,151,176]" pageId="32" pageNumber="33">CM</collectionCode>
24425) and subadult male (
<collectionCode id="ED18AE35FFD5831ED391FF4AD2DAFF30" box="[1166,1212,151,176]" pageId="32" pageNumber="33">CM</collectionCode>
1316) are illustrated in
<figureCitation id="13322A75FFD5831ED7ABFF61D745FF55" box="[180,291,188,213]" captionStart="FIGURE 16" captionStartId="30.[151,250,1401,1423]" captionTargetBox="[151,1436,207,1382]" captionTargetId="figure@30.[151,1436,207,1382]" captionTargetPageId="30" captionText="FIGURE 16. Crania and dentaries of P. tapoatafa tapoatafa. a) CM 24425, an adult male from Coraki, New South Wales, and b) CM 1316, a subadult male from Wallaby Creek, near Urbanville, New South Wales. Each series consists of (left to right) dorsal view of cranium, ventral view of cranium, and lateral view of right dentary." httpUri="https://zenodo.org/record/240169/files/figure.png" pageId="32" pageNumber="33">Figure 16</figureCitation>
. Cranio-mandibular and dental measurements for each sex are provided in
<tableCitation id="C68B034BFFD5831ED374FF61D2A6FF55" box="[1131,1216,188,213]" captionStart="TABLE 5" captionStartId="25.[125,203,525,545]" captionText="TABLE 5. Cranial and dental measurements of each taxon and major regional population within the P. tapoatafa group, presented separately for males (5 a) and females (5 b). Data are presented separately for three regional populations within nominotypical P. tapoatafa: 1) South Australia; 2) Victoria; 3) New South Wales and southern Queensland. Measurements of holotypes are also presented." pageId="32" pageNumber="33">Table 5</tableCitation>
.
</paragraph>
<paragraph id="8BB636F0FFD5831ED7D8FF02D449FD0A" blockId="32.[151,1437,151,2018]" pageId="32" pageNumber="33">
The New South Wales to southeast Queensland populations of
<taxonomicName id="4C094D73FFD5831ED487FF3CD25FFF78" box="[920,1081,224,248]" class="Mammalia" family="Dasyuridae" genus="Phascogale" kingdom="Animalia" order="Dasyuromorphia" pageId="32" pageNumber="33" phylum="Chordata" rank="subSpecies" species="tapoatafa" subSpecies="tapoatafa">
<emphasis id="B97DEAE2FFD5831ED487FF3CD25FFF78" box="[920,1081,224,248]" italics="true" pageId="32" pageNumber="33">P. t. tapoatafa</emphasis>
</taxonomicName>
show clear male-biased sexual dimorphism in cranio-dental measurements (
<tableCitation id="C68B034BFFD5831ED596FED9D4BBFE9D" box="[649,733,260,285]" captionStart="TABLE 6" captionStartId="29.[125,203,627,647]" captionText="TABLE 6. Statistical results for 1 - factor ANOVA tests for sexual dimorphism in cranio-dental measurements of the Phascogale tapoatafa group." pageId="32" pageNumber="33">Table 6</tableCitation>
&amp;
<figureCitation id="13322A75FFD5831ED41FFED9D524FE9C" box="[768,834,260,285]" captionStart="FIGURE 9" captionStartId="14.[151,250,1875,1897]" captionTargetBox="[341,1221,226,1853]" captionTargetId="figure@14.[341,1232,194,1855]" captionTargetPageId="14" captionText="FIGURE 9. Bivariate scatter plots of cranio-dental dimensions, illustrating varying degrees of inter-populational variability separately for male and female samples. Variables: a) SL vs ZW; b) SL vs C 1 - M 4; c) SL vs OBW; d) SL vs M 1 - 4; e) SL vs FMW; f) SL vs IOW; g) SL vs PRW. Population codes: 1: southwest Western Australia; 2: South Australia; 3: Victoria; 4: New South Wales and southeast Queensland; 5: northeast Queensland; 6: Top End of the Northern Territory; 7: Kimberley region of Western Australia. Left hand series are females; right hand series are males." httpUri="https://zenodo.org/record/240162/files/figure.png" pageId="32" pageNumber="33">Fig. 9</figureCitation>
). For all dimensions bar one (
<collectionCode id="ED18AE35FFD5831ED391FED9D2B1FE9C" box="[1166,1239,260,284]" pageId="32" pageNumber="33">FMW</collectionCode>
), mean values are higher in males than females and for nine dimensions the contrast is statistically significant. This latter subset includes measures of overall cranial size (
<collectionCode id="ED18AE35FFD5831ED56AFE91D4FFFEE4" box="[629,665,332,356]" pageId="32" pageNumber="33">SL</collectionCode>
,
<collectionCode id="ED18AE35FFD5831ED5B9FE90D4B4FEE4" box="[678,722,333,356]" pageId="32" pageNumber="33">ZW</collectionCode>
,
<collectionCode id="ED18AE35FFD5831ED5C0FE91D54CFEE4" box="[735,810,332,356]" pageId="32" pageNumber="33">PALW</collectionCode>
,
<collectionCode id="ED18AE35FFD5831ED428FE91D512FEE4" box="[823,884,332,356]" pageId="32" pageNumber="33">PRW</collectionCode>
) and of the lower canine (
<collectionCode id="ED18AE35FFD5831ED3BEFE91D2D2FEE4" box="[1185,1204,332,356]" pageId="32" pageNumber="33">C</collectionCode>
1
<collectionCode id="ED18AE35FFD5831ED3A2FE90D2B7FEE4" box="[1213,1233,333,356]" pageId="32" pageNumber="33">L</collectionCode>
;
<collectionCode id="ED18AE35FFD5831ED3C1FE91D297FEE4" box="[1246,1265,332,356]" pageId="32" pageNumber="33">C</collectionCode>
1
<collectionCode id="ED18AE35FFD5831ED3E6FE91D371FEE4" box="[1273,1303,332,356]" pageId="32" pageNumber="33">W</collectionCode>
), as well as measures that are sensitive to canine enlargement (
<collectionCode id="ED18AE35FFD5831ED5C9FEABD48FFE0E" box="[726,745,374,398]" pageId="32" pageNumber="33">C</collectionCode>
1-
<collectionCode id="ED18AE35FFD5831ED5E6FEABD572FE0D" box="[761,788,374,397]" pageId="32" pageNumber="33">M</collectionCode>
4
<collectionCode id="ED18AE35FFD5831ED404FEABD549FE0D" box="[795,815,374,397]" pageId="32" pageNumber="33">L</collectionCode>
,
<collectionCode id="ED18AE35FFD5831ED423FEABD5E7FE0E" box="[828,897,374,398]" pageId="32" pageNumber="33">ARW</collectionCode>
). The combined South Australian and Victorian sample shows a less pronounced male-biased dimorphism in which statistical significance is attained only for measures of the lower canine. In this population, the contrast in PRW also approaches significance. These observations on the Victorian population are consistent with Soderquists (1995c) finding that the marked sexual dimorphism in body weight among Victorian
<taxonomicName id="4C094D73FFD5831ED5BBFDDBD54BFD9E" box="[676,813,518,542]" class="Mammalia" family="Dasyuridae" genus="Phascogale" kingdom="Animalia" order="Dasyuromorphia" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="tapoatafa">
<emphasis id="B97DEAE2FFD5831ED5BBFDDBD54BFD9E" box="[676,813,518,542]" italics="true" pageId="32" pageNumber="33">P. tapoatafa</emphasis>
</taxonomicName>
is due primarily to somatic energy stores that are laid down in preparation for the short breeding season when foraging effort is curtailed. In the Victorian population studied by Soderquist the ratio of average adult male to female body weights is 1.62 (Soderquist unpublished data, reported by
<bibRefCitation id="EF984B01FFD5831ED601FDACD445FD0A" author="Rhind" box="[286,547,625,650]" pageId="32" pageNumber="33" refString="Rhind, S. G. &amp; Bradley, J. S. (2002) The effect of drought on body size, growth and abundance of wild brush-tailed phascogales (Phascogale tapoatafa) in south-western Australia. Wildlife Research, 29, 235 - 245. http: // dx. doi. org / 10.1071 / WR 01014" type="journal article" year="2002">Rhind &amp; Bradley 2002</bibRefCitation>
).
</paragraph>
<paragraph id="8BB636F0FFD5831ED7D8FD48D32DFC06" blockId="32.[151,1437,151,2018]" pageId="32" pageNumber="33">
<emphasis id="B97DEAE2FFD5831ED7D8FD48D7A8FD2E" bold="true" box="[199,462,661,686]" pageId="32" pageNumber="33">Geographic variation.</emphasis>
Possible clinal variation within
<taxonomicName id="4C094D73FFD5831ED429FD4BD5B5FD2E" box="[822,979,662,686]" class="Mammalia" family="Dasyuridae" genus="Phascogale" kingdom="Animalia" order="Dasyuromorphia" pageId="32" pageNumber="33" phylum="Chordata" rank="subSpecies" species="tapoatafa" subSpecies="tapoatafa">
<emphasis id="B97DEAE2FFD5831ED429FD4BD5B5FD2E" box="[822,979,662,686]" italics="true" pageId="32" pageNumber="33">P. t. tapoatafa</emphasis>
</taxonomicName>
is observed in various cranial and dental dimensions but with somewhat different expression in each sex. Among males there is a very slight and gradual decrease in cranial length (
<figureCitation id="13322A75FFD5831ED6C4FD00D457FD76" box="[475,561,733,758]" captionStart="FIGURE 17" captionStartId="31.[151,250,1848,1870]" captionTargetBox="[155,1436,677,1804]" captionTargetId="figure@31.[151,1436,661,1827]" captionTargetPageId="31" captionText="FIGURE 17. Graphical representation of geographic variation within P. t. tapoatafa in selected craniodental dimensions (SL; M 1 - 4; FMW). Values for individual specimens are plotted in geographic sequence (Geo Seq) from west to east for South Australia and Victoria, and then from south to north for New South Wales and southeast Queensland. Population codes: 2: South Australia; 3: Victoria; 4: New South Wales and southeast Queensland. Plotted variables are a) Skull Length; b) M 1 - 4; c) Foramen Magnum Width. Left hand series are females; right hand series are males." httpUri="https://zenodo.org/record/240170/files/figure.png" pageId="32" pageNumber="33">Fig. 17</figureCitation>
a) from South Australia to Queensland, coupled with a more pronounced shortening of the upper molar row that also shows a more abrupt break between southern and northeastern populations (
<figureCitation id="13322A75FFD5831ED633FCF8D7E6FCBE" box="[300,384,805,830]" captionStart="FIGURE 17" captionStartId="31.[151,250,1848,1870]" captionTargetBox="[155,1436,677,1804]" captionTargetId="figure@31.[151,1436,661,1827]" captionTargetPageId="31" captionText="FIGURE 17. Graphical representation of geographic variation within P. t. tapoatafa in selected craniodental dimensions (SL; M 1 - 4; FMW). Values for individual specimens are plotted in geographic sequence (Geo Seq) from west to east for South Australia and Victoria, and then from south to north for New South Wales and southeast Queensland. Population codes: 2: South Australia; 3: Victoria; 4: New South Wales and southeast Queensland. Plotted variables are a) Skull Length; b) M 1 - 4; c) Foramen Magnum Width. Left hand series are females; right hand series are males." httpUri="https://zenodo.org/record/240170/files/figure.png" pageId="32" pageNumber="33">Fig. 17</figureCitation>
b). Females show a more pronounced decrease in both cranial and dental dimensions (
<figureCitation id="13322A75FFD5831ED275FCF8D6D5FCE2" captionStart="FIGURE 17" captionStartId="31.[151,250,1848,1870]" captionTargetBox="[155,1436,677,1804]" captionTargetId="figure@31.[151,1436,661,1827]" captionTargetPageId="31" captionText="FIGURE 17. Graphical representation of geographic variation within P. t. tapoatafa in selected craniodental dimensions (SL; M 1 - 4; FMW). Values for individual specimens are plotted in geographic sequence (Geo Seq) from west to east for South Australia and Victoria, and then from south to north for New South Wales and southeast Queensland. Population codes: 2: South Australia; 3: Victoria; 4: New South Wales and southeast Queensland. Plotted variables are a) Skull Length; b) M 1 - 4; c) Foramen Magnum Width. Left hand series are females; right hand series are males." httpUri="https://zenodo.org/record/240170/files/figure.png" pageId="32" pageNumber="33">Figs 17</figureCitation>
de). Both sexes show an abrupt and significant contrast between southern and northeastern populations in the breadth of the foramen magnum (
<figureCitation id="13322A75FFD5831ED511FCB0D43FFC06" box="[526,601,877,902]" captionStart="FIGURE 17" captionStartId="31.[151,250,1848,1870]" captionTargetBox="[155,1436,677,1804]" captionTargetId="figure@31.[151,1436,661,1827]" captionTargetPageId="31" captionText="FIGURE 17. Graphical representation of geographic variation within P. t. tapoatafa in selected craniodental dimensions (SL; M 1 - 4; FMW). Values for individual specimens are plotted in geographic sequence (Geo Seq) from west to east for South Australia and Victoria, and then from south to north for New South Wales and southeast Queensland. Population codes: 2: South Australia; 3: Victoria; 4: New South Wales and southeast Queensland. Plotted variables are a) Skull Length; b) M 1 - 4; c) Foramen Magnum Width. Left hand series are females; right hand series are males." httpUri="https://zenodo.org/record/240170/files/figure.png" pageId="32" pageNumber="33">Fig 17</figureCitation>
c,f); this variable is little affected by overall growth of the cranium.
</paragraph>
<paragraph id="8BB636F0FFD5831ED7D8FC4CD6A1FB4A" blockId="32.[151,1437,151,2018]" pageId="32" pageNumber="33">
The different pattern of geographic variation between the sexes of
<taxonomicName id="4C094D73FFD5831ED4D8FC4FD20FFC29" box="[967,1129,913,937]" class="Mammalia" family="Dasyuridae" genus="Phascogale" kingdom="Animalia" order="Dasyuromorphia" pageId="32" pageNumber="33" phylum="Chordata" rank="subSpecies" species="tapoatafa" subSpecies="tapoatafa">
<emphasis id="B97DEAE2FFD5831ED4D8FC4FD20FFC29" box="[967,1129,913,937]" italics="true" pageId="32" pageNumber="33">P. t. tapoatafa</emphasis>
</taxonomicName>
is largely a product of the variable expression of sexual dimorphism between the southern and southeastern Australian populations. Essentially, males of
<taxonomicName id="4C094D73FFD5831ED68CFC07D45FFC71" box="[403,569,985,1009]" class="Mammalia" family="Dasyuridae" genus="Phascogale" kingdom="Animalia" order="Dasyuromorphia" pageId="32" pageNumber="33" phylum="Chordata" rank="subSpecies" species="tapoatafa" subSpecies="tapoatafa">
<emphasis id="B97DEAE2FFD5831ED68CFC07D45FFC71" box="[403,569,985,1009]" italics="true" pageId="32" pageNumber="33">P. t. tapoatafa</emphasis>
</taxonomicName>
vary only slightly in cranial size throughout their range, while the more pronounced clinal variation in female cranial size, from large in the south to smaller in the north, creates a widening contrast between the two sexes. Overlaid on this pattern is a more pronounced geographic variation in molar size, with southern animals of both sexes having larger teeth on average than those from New South Wales and Queensland. On present evidence, this variation appears to be manifested in different ways between the sexes—step-like in males
<emphasis id="B97DEAE2FFD5831ED6DEFB53D7BEFB25" box="[449,472,1166,1189]" italics="true" pageId="32" pageNumber="33">vs</emphasis>
clinal in females—but larger samples from critical areas are needed to be certain of this.
</paragraph>
</subSubSection>
<subSubSection id="C313657BFFD5831FD7D8FB08D58AF836" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33" type="distribution">
<paragraph id="8BB636F0FFD5831ED7D8FB08D715FA6A" blockId="32.[151,1437,151,2018]" pageId="32" pageNumber="33">
<emphasis id="B97DEAE2FFD5831ED7D8FB08D4E3FB6E" bold="true" box="[199,645,1237,1262]" pageId="32" pageNumber="33">Distribution and conservation status.</emphasis>
The historical distribution of
<taxonomicName id="4C094D73FFD5831ED4C5FB0BD215FB6E" box="[986,1139,1238,1262]" class="Mammalia" family="Dasyuridae" genus="Phascogale" kingdom="Animalia" order="Dasyuromorphia" pageId="32" pageNumber="33" phylum="Chordata" rank="subSpecies" species="tapoatafa" subSpecies="tapoatafa">
<emphasis id="B97DEAE2FFD5831ED4C5FB0BD215FB6E" box="[986,1139,1238,1262]" italics="true" pageId="32" pageNumber="33">P. t tapoatafa</emphasis>
</taxonomicName>
corresponded to the more or less continuous tract of forested country that extended from the southeastern corner of South Australia, extending across most of Victoria (probably excluding only the northwestern mallee region and country above the treeline) and then running north along the coastal strip and mountainous hinterland of New South Wales and Queensland, north at least to Mackay (
<figureCitation id="13322A75FFD5831ED55BFAB8D4F0FAFE" box="[580,662,1381,1406]" captionStart="FIGURE 18" captionStartId="33.[151,250,1467,1489]" captionTargetBox="[429,1157,267,1446]" captionTargetId="figure@33.[429,1158,267,1446]" captionTargetPageId="33" captionText="FIGURE 18. Map of eastern Australia, showing the contemporary (dark grey) and historic (light grey) distribution of P. tapoatafa tapoatafa, based on museum specimens and reliable sightings. Star symbols are sub-fossil records judged to be of late Holocene age. Square symbols represent the location of two sight records that might pertain to either P. pi r at a or P. tapoatafa." httpUri="https://zenodo.org/record/240171/files/figure.png" pageId="32" pageNumber="33">Fig. 18</figureCitation>
). A recently collected sub-fossil specimen from Ukerbarley station,
<quantity id="4CF19B15FFD5831ED291FABBD6DDFA22" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="32" pageNumber="33" unit="km" value="7.0">7 km</quantity>
NW of Coonabarabran (
<figureCitation id="13322A75FFD5831ED6FEFA54D453FA22" box="[481,565,1417,1442]" captionStart="FIGURE 18" captionStartId="33.[151,250,1467,1489]" captionTargetBox="[429,1157,267,1446]" captionTargetId="figure@33.[429,1158,267,1446]" captionTargetPageId="33" captionText="FIGURE 18. Map of eastern Australia, showing the contemporary (dark grey) and historic (light grey) distribution of P. tapoatafa tapoatafa, based on museum specimens and reliable sightings. Star symbols are sub-fossil records judged to be of late Holocene age. Square symbols represent the location of two sight records that might pertain to either P. pi r at a or P. tapoatafa." httpUri="https://zenodo.org/record/240171/files/figure.png" pageId="32" pageNumber="33">Fig. 18</figureCitation>
&amp;
<tableCitation id="C68B034BFFD5831ED57EFA54D4DCFA22" box="[609,698,1417,1442]" captionStart="TABLE 9" captionStartId="47.[151,239,152,175]" captionTargetBox="[159,1304,266,1074]" captionTargetPageId="47" captionText="TABLE 9. Location information, including State [NSW: New South Wales, Qld: Queensland, SA: South Australia, VIC: Victoria, WA: Western Australia), decimal latitude and longitude for fossil and sub-fossil records of members of the Phascogale tapoatafa group." httpUri="http://table.plazi.org/id/DF766678FFDA8311D788FF45D7AAFF6F" pageId="32" pageNumber="33" tableUuid="DF766678FFDA8311D788FF45D7AAFF6F">Table 9</tableCitation>
), indicates that populations of this taxon once occurred on the western slopes of the Great Dividing Range in New South Wales. Whether any of these survived into historical times is not known.
</paragraph>
<paragraph id="8BB636F0FFD5831ED7D8FA2BD7C2F8AE" blockId="32.[151,1437,151,2018]" pageId="32" pageNumber="33">
At the western end of its range, a population of
<taxonomicName id="4C094D73FFD5831ED5C7FA2BD512F98E" box="[728,884,1526,1550]" class="Mammalia" family="Dasyuridae" genus="Phascogale" kingdom="Animalia" order="Dasyuromorphia" pageId="32" pageNumber="33" phylum="Chordata" rank="subSpecies" species="tapoatafa" subSpecies="tapoatafa">
<emphasis id="B97DEAE2FFD5831ED5C7FA2BD512F98E" box="[728,884,1526,1550]" italics="true" pageId="32" pageNumber="33">P. t. tapoatafa</emphasis>
</taxonomicName>
was recorded historically in the Mt Lofty Ranges of South Australia (
<figureCitation id="13322A75FFD5831ED66DF9C4D7A3F9B2" box="[370,453,1561,1586]" captionStart="FIGURE 18" captionStartId="33.[151,250,1467,1489]" captionTargetBox="[429,1157,267,1446]" captionTargetId="figure@33.[429,1158,267,1446]" captionTargetPageId="33" captionText="FIGURE 18. Map of eastern Australia, showing the contemporary (dark grey) and historic (light grey) distribution of P. tapoatafa tapoatafa, based on museum specimens and reliable sightings. Star symbols are sub-fossil records judged to be of late Holocene age. Square symbols represent the location of two sight records that might pertain to either P. pi r at a or P. tapoatafa." httpUri="https://zenodo.org/record/240171/files/figure.png" pageId="32" pageNumber="33">Fig. 18</figureCitation>
). Although
<bibRefCitation id="EF984B01FFD5831ED556F9C4D547F9B2" box="[585,801,1561,1586]" pageId="32" pageNumber="33" refString="Wood Jones, F. (1923) The Mammals of South Australia. Part I. R. E. E. Rogers, Government Printer, Adelaide, pp. 98 - 101." type="book chapter">Wood Jones (1923)</bibRefCitation>
understood the range of this species (as
<taxonomicName id="4C094D73FFD5831ED3FBF9C7D310F9B1" box="[1252,1398,1561,1585]" class="Mammalia" family="Dasyuridae" genus="Dasyurus" kingdom="Animalia" order="Dasyuromorphia" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="penicillata">
<emphasis id="B97DEAE2FFD5831ED3FBF9C7D310F9B1" box="[1252,1398,1561,1585]" italics="true" pageId="32" pageNumber="33">P. penicillata</emphasis>
</taxonomicName>
) to extend to the Murray River, sub-fossil collections from several archaeological sites in this area have failed to confirm its former presence (
<bibRefCitation id="EF984B01FFD5831ED6FBF9BCD4F8F9FA" author="Wakefield" box="[484,670,1633,1658]" pageId="32" pageNumber="33" refString="Wakefield, N. A. (1964) Mammal remains. Supplementary Table (pp. 494 - 498) to: Mulvaney, D. J. (1964). Archeological excavations at Fromm's Landing on the lower Murray River, South Australia. Proceedings of the Royal Society Victoria, 77, 479 - 516." type="journal article" year="1964">Wakefield 1964</bibRefCitation>
). Morphologically, the few available specimens from the Mt Lofty population appear to be indistinguishable from Victorian specimens. However, in the absence of genetic data, it is not possible to decide whether the Mt Lofty population was genuinely isolated or not. Caution in this regard is further indicated by a subfossil record of
<taxonomicName id="4C094D73FFD5831ED57EF913D48EF966" box="[609,744,1742,1766]" class="Mammalia" family="Dasyuridae" genus="Phascogale" kingdom="Animalia" order="Dasyuromorphia" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="tapoatafa">
<emphasis id="B97DEAE2FFD5831ED57EF913D48EF966" box="[609,744,1742,1766]" italics="true" pageId="32" pageNumber="33">P. tapoatafa</emphasis>
</taxonomicName>
from a cave deposit of mid-Holocene age on Kangaroo Island (
<figureCitation id="13322A75FFD5831ED780F92CD692F88A" box="[159,244,1777,1802]" captionStart="FIGURE 18" captionStartId="33.[151,250,1467,1489]" captionTargetBox="[429,1157,267,1446]" captionTargetId="figure@33.[429,1158,267,1446]" captionTargetPageId="33" captionText="FIGURE 18. Map of eastern Australia, showing the contemporary (dark grey) and historic (light grey) distribution of P. tapoatafa tapoatafa, based on museum specimens and reliable sightings. Star symbols are sub-fossil records judged to be of late Holocene age. Square symbols represent the location of two sight records that might pertain to either P. pi r at a or P. tapoatafa." httpUri="https://zenodo.org/record/240171/files/figure.png" pageId="32" pageNumber="33">Fig. 18</figureCitation>
&amp;
<tableCitation id="C68B034BFFD5831ED63DF92CD71AF88A" box="[290,380,1777,1802]" captionStart="TABLE 9" captionStartId="47.[151,239,152,175]" captionTargetBox="[159,1304,266,1074]" captionTargetPageId="47" captionText="TABLE 9. Location information, including State [NSW: New South Wales, Qld: Queensland, SA: South Australia, VIC: Victoria, WA: Western Australia), decimal latitude and longitude for fossil and sub-fossil records of members of the Phascogale tapoatafa group." httpUri="http://table.plazi.org/id/DF766678FFDA8311D788FF45D7AAFF6F" pageId="32" pageNumber="33" tableUuid="DF766678FFDA8311D788FF45D7AAFF6F">Table 9</tableCitation>
), indicating a wider distributional and habitat range than might be inferred from the few historical records alone.
</paragraph>
<paragraph id="8BB636F0FFD5831FD7D8F8E7D6B3FF55" blockId="32.[151,1437,151,2018]" lastBlockId="33.[151,1436,151,213]" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33">
As reported in a later section, populations of
<taxonomicName id="4C094D73FFD5831ED5D0F8E4D5A0F8D1" box="[719,966,1849,1873]" class="Mammalia" family="Dasyuridae" genus="Phascogale" kingdom="Animalia" order="Dasyuromorphia" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="tapoatafa">
<emphasis id="B97DEAE2FFD5831ED5D0F8E4D5A0F8D1" box="[719,966,1849,1873]" italics="true" pageId="32" pageNumber="33">Phascogale tapoatafa</emphasis>
</taxonomicName>
are also present in northern Queensland. These resemble the nominate race in most morphological features but show a moderate level of genetic differentiation (albeit based on a single individual from the northern population). Today these populations appear to be separated from those in the south by a gap of more than
<quantity id="4CF19B15FFD5831ED425F87BD5F6F83E" box="[826,912,1957,1982]" metricMagnitude="5" metricUnit="m" metricValue="4.0" pageId="32" pageNumber="33" unit="km" value="400.0">400 km</quantity>
of significantly drier habitat. However, a subfossil record of
<taxonomicName id="4C094D73FFD5831ED656F817D7B6F862" box="[329,464,1994,2018]" class="Mammalia" family="Dasyuridae" genus="Phascogale" kingdom="Animalia" order="Dasyuromorphia" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="tapoatafa">
<emphasis id="B97DEAE2FFD5831ED656F817D7B6F862" box="[329,464,1994,2018]" italics="true" pageId="32" pageNumber="33">P. tapoatafa</emphasis>
</taxonomicName>
from Christmas Creek, inland of Townsville (
<tableCitation id="C68B034BFFD5831ED4C8F814D24AF862" box="[983,1068,1993,2018]" captionStart="TABLE 9" captionStartId="47.[151,239,152,175]" captionTargetBox="[159,1304,266,1074]" captionTargetPageId="47" captionText="TABLE 9. Location information, including State [NSW: New South Wales, Qld: Queensland, SA: South Australia, VIC: Victoria, WA: Western Australia), decimal latitude and longitude for fossil and sub-fossil records of members of the Phascogale tapoatafa group." httpUri="http://table.plazi.org/id/DF766678FFDA8311D788FF45D7AAFF6F" pageId="32" pageNumber="33" tableUuid="DF766678FFDA8311D788FF45D7AAFF6F">Table 9</tableCitation>
) signals the possibility of recent continuity between these populations. The northern Queensland population is herein left unallocated at subspecific level.
</paragraph>
<caption id="DF766678FFD4831FD788FA66D698F9AD" httpUri="https://zenodo.org/record/240171/files/figure.png" pageId="33" pageNumber="34" targetBox="[429,1157,267,1446]" targetPageId="33">
<paragraph id="8BB636F0FFD4831FD788FA66D698F9AD" blockId="33.[151,1436,1467,1581]" pageId="33" pageNumber="34">
<emphasis id="B97DEAE2FFD4831FD788FA66D77AFA50" bold="true" box="[151,284,1467,1489]" pageId="33" pageNumber="34">FIGURE 18</emphasis>
. Map of eastern Australia, showing the contemporary (dark grey) and historic (light grey) distribution of
<taxonomicName id="4C094D73FFD4831FD296FA61D707FA70" class="Mammalia" family="Dasyuridae" genus="Phascogale" kingdom="Animalia" order="Dasyuromorphia" pageId="33" pageNumber="34" phylum="Chordata" rank="subSpecies" species="tapoatafa" subSpecies="tapoatafa">
<emphasis id="B97DEAE2FFD4831FD296FA61D707FA70" italics="true" pageId="33" pageNumber="34">P. tapoatafa tapoatafa</emphasis>
</taxonomicName>
, based on museum specimens and reliable sightings. Star symbols are sub-fossil records judged to be of late Holocene age. Square symbols represent the location of two sight records that might pertain to either
<emphasis id="B97DEAE2FFD4831FD21AFA27D33BF98F" box="[1285,1373,1530,1551]" italics="true" pageId="33" pageNumber="34">P. pi r at a</emphasis>
or
<taxonomicName id="4C094D73FFD4831FD296FA27D69EF9AD" class="Mammalia" family="Dasyuridae" genus="Phascogale" kingdom="Animalia" order="Dasyuromorphia" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="tapoatafa">
<emphasis id="B97DEAE2FFD4831FD296FA27D69EF9AD" italics="true" pageId="33" pageNumber="34">P. tapoatafa</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BB636F0FFD4831FD7D8F987D58AF836" blockId="33.[151,1436,1626,1975]" pageId="33" pageNumber="34">
Although Victoria, New South Wales and Queensland still host populations of
<taxonomicName id="4C094D73FFD4831FD353F986D28AF9F2" box="[1100,1260,1626,1650]" class="Mammalia" family="Dasyuridae" genus="Phascogale" kingdom="Animalia" order="Dasyuromorphia" pageId="33" pageNumber="34" phylum="Chordata" rank="subSpecies" species="tapoatafa" subSpecies="tapoatafa">
<emphasis id="B97DEAE2FFD4831FD353F986D28AF9F2" box="[1100,1260,1626,1650]" italics="true" pageId="33" pageNumber="34">P. t. tapoatafa</emphasis>
</taxonomicName>
, its geographic range is now extremely fragmented and many regional populations are in decline. Indeed, the only region in which the taxon might be said to be secure is southeastern Queensland.
<taxonomicName id="4C094D73FFD4831FD473F97FD207F93A" box="[876,1121,1698,1722]" class="Mammalia" family="Dasyuridae" genus="Phascogale" kingdom="Animalia" order="Dasyuromorphia" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="tapoatafa">
<emphasis id="B97DEAE2FFD4831FD473F97FD207F93A" box="[876,1121,1698,1722]" italics="true" pageId="33" pageNumber="34">Phascogale tapoatafa</emphasis>
</taxonomicName>
is probably extinct in South Australia, with the last vouchered record dating to 1969.
<bibRefCitation id="EF984B01FFD4831FD430F91BD26EF95F" author="Wood" box="[815,1032,1734,1759]" pageId="33" pageNumber="34" refString="Wood Jones, F. (1923) The Mammals of South Australia. Part I. R. E. E. Rogers, Government Printer, Adelaide, pp. 98 - 101." type="book chapter" year="1923">
Wood
<bibRefCitation id="EF984B01FFD4831FD465F91AD26EF95F" author="Jones" box="[890,1032,1735,1759]" pageId="33" pageNumber="34" refString="Wood Jones, F. (1923) The Mammals of South Australia. Part I. R. E. E. Rogers, Government Printer, Adelaide, pp. 98 - 101." type="book chapter" year="1923">Jones (1923</bibRefCitation>
</bibRefCitation>
: 101102) remarked on the former abundance of the species in South Australia and noted that it was known to “the South Australian Murray River natives as “
<emphasis id="B97DEAE2FFD4831FD63CF8D3D705F8A6" box="[291,355,1806,1830]" italics="true" pageId="33" pageNumber="34">pundi</emphasis>
”. He also speculated on the possible existence of populations of this taxon “over a wide area in the Centre”. It is possible that these reports referred instead to
<taxonomicName id="4C094D73FFD4831FD44BF8EED5DFF8CA" box="[852,953,1842,1866]" class="Mammalia" family="Dasyuridae" genus="Phascogale" kingdom="Animalia" order="Dasyuromorphia" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="calura">
<emphasis id="B97DEAE2FFD4831FD44BF8EED5DFF8CA" box="[852,953,1842,1866]" italics="true" pageId="33" pageNumber="34">P. calura</emphasis>
</taxonomicName>
, which Wood Jones tentatively included in the South Australian fauna on the basis of an early record from Adelaide but otherwise knew little about. In fact, Spencer and Gillen had obtained several specimens of
<emphasis id="B97DEAE2FFD4831FD415F8A6D50BF812" box="[778,877,1914,1938]" italics="true" pageId="33" pageNumber="34">P. c a l u r a</emphasis>
from Central Australia prior to 1901 (
<bibRefCitation id="EF984B01FFD4831FD236F8A7D750F837" author="Baynes" pageId="33" pageNumber="34" refString="Baynes, A. &amp; Johnson, K. A. (1996) The contributions of the Horn Expedition and cave deposits to knowledge of the original mammal fauna of central Australia. In: Morton, S. R. &amp; Mulvaney, D. J. (Eds.), Exploring Central Australia: Society, the Environment and the 1894 Horn Expedition. Surrey Beatty &amp; Sons Pty Ltd, Chipping Norton, pp. 168 - 186." type="book chapter" year="1996">Baynes &amp; Johnson 1996</bibRefCitation>
), although no further specimens were obtained from this area.
</paragraph>
</subSubSection>
<subSubSection id="C313657BFFD7831CD7D8FF45D4DDFE2C" pageId="34" pageNumber="35" type="reference_group">
<paragraph id="8BB636F0FFD7831CD7D8FF45D4DDFE2C" blockId="34.[151,1436,151,429]" pageId="34" pageNumber="35">
<taxonomicName id="4C094D73FFD7831CD7D8FF45D7B3FF30" ID-CoL="4FVN8" box="[199,469,152,176]" class="Mammalia" family="Dasyuridae" genus="Phascogale" kingdom="Animalia" order="Dasyuromorphia" pageId="34" pageNumber="35" phylum="Chordata" rank="subSpecies" species="tapoatafa" subSpecies="tapoatafa">
<emphasis id="B97DEAE2FFD7831CD7D8FF45D7B3FF30" box="[199,469,152,176]" italics="true" pageId="34" pageNumber="35">Phascogale t. tapoatafa</emphasis>
</taxonomicName>
has been the focus of several field studies in Victoria (
<bibRefCitation id="EF984B01FFD7831CD358FF4AD2B6FF30" author="Cuttle" box="[1095,1232,151,176]" pageId="34" pageNumber="35" refString="Cuttle, P. (1978) The Behaviour in Captivity of the Dasyurid Marsupial Phascogale tapoatafa (Meyer). M. Sc. Thesis, Monash University, Melbourne, 241 pp." type="book" year="1978">Cuttle 1978</bibRefCitation>
,
<bibRefCitation id="EF984B01FFD7831CD3C3FF45D371FF30" author="Cuttle" box="[1244,1303,152,176]" pageId="34" pageNumber="35" refString="Cuttle, P. (1982) Life history strategy of the dasyurid marsupial Phascogale tapoatafa. In: Archer, M (Ed.), Carnivorous Marsupials. Royal Zoological Society of New South Wales. Surrey Beatty, NSW, pp. 13 - 22." type="book chapter" year="1982">1982</bibRefCitation>
;
<bibRefCitation id="EF984B01FFD7831CD23BFF4AD6B8FF54" author="Soderquist" pageId="34" pageNumber="35" refString="Soderquist, T. R. (1993 a) Maternal strategies of Phascogale tapoatafa (Marsupialia: Dasyuridae). I. Breeding seasonality and maternal investment. Australian Journal of Zoology, 41, 549 - 566." type="journal article" year="1993" yearSuffix="a">Soderquist 1993a</bibRefCitation>
,
<bibRefCitation id="EF984B01FFD7831CD7F4FF61D755FF55" author="Soderquist" box="[235,307,188,213]" pageId="34" pageNumber="35" refString="Soderquist, T. R. (1993 b) Maternal strategies of Phascogale tapoatafa (Marsupialia: Dasyuridae). II. Juvenile thermoregulation and maternal attendance. Australian Journal of Zoology, 41, 567 - 576." type="journal article" year="1993" yearSuffix="b">1993b</bibRefCitation>
,
<bibRefCitation id="EF984B01FFD7831CD65FFF61D7E1FF54" author="Soderquist" box="[320,391,188,212]" pageId="34" pageNumber="35" refString="Soderquist, T. R. (1994 a) Reproductive Strategies of Phascogale tapoatafa (Marsupialia: Dasyuridae). Ph. D. Thesis, Monash University, Melbourne. [unkown pagination]" type="book" year="1994" yearSuffix="a">1994a</bibRefCitation>
,
<bibRefCitation id="EF984B01FFD7831CD68CFF61D7BAFF55" author="Soderquist" box="[403,476,188,213]" pageId="34" pageNumber="35" refString="Soderquist, T. R. (1994 b) Anti-predator behaviour of the brushtailed phascogale (Phascogale tapoatafa). Victorian Naturalist, 111, 22 - 24." type="journal article" year="1994" yearSuffix="b">1994b</bibRefCitation>
,
<bibRefCitation id="EF984B01FFD7831CD6F6FF61D456FF55" author="Soderquist" box="[489,560,188,213]" pageId="34" pageNumber="35" refString="Soderquist, T. R. (1995 a) Spatial organisation of the arboreal carnivorous marsupial Phascogale tapoatafa. Australian Journal of Zoology, 237, 385 - 398." type="journal article" year="1995" yearSuffix="a">1995a</bibRefCitation>
;
<bibRefCitation id="EF984B01FFD7831CD522FF61D539FF54" author="Soderquist" box="[573,863,188,213]" pageId="34" pageNumber="35" refString="Soderquist, T. R. &amp; Ealey, L. (1994) Social interactions and mating strategies of a solitary carnivorous marsupial Phascogale tapoatafa, in the wild. Wildlife Research, 21, 527 - 542. http: // dx. doi. org / 10.1071 / WR 9940527" type="journal article" year="1994">Soderquist &amp; Ealey 1994</bibRefCitation>
;
<bibRefCitation id="EF984B01FFD7831CD474FF61D215FF55" author="Soderquist" box="[875,1139,188,213]" pageId="34" pageNumber="35" refString="Soderquist, T. R. &amp; Lill, A. (1995) Natal dispersal and philopatry in the carnivorous marsupial Phascogale tapoatafa (Dasyuridae). Ethology, 99, 297 - 312. http: // dx. doi. org / 10.1111 / j. 1439 - 0310.1995. tb 00904. x" type="journal article" year="1995">Soderquist &amp; Lill 1995</bibRefCitation>
) but little is known of the species ecology in New South Wales or Queensland. The difference between these populations in the degree of sexual dimorphism suggests caution in assuming a uniformity of all aspects of ecology among populations of this widespread subspecies.
<bibRefCitation id="EF984B01FFD7831CD6B2FEFAD4ACFEC0" author="Dixon" box="[429,714,295,320]" pageId="34" pageNumber="35" refString="Dixon, J. M. &amp; Huxley, L. M. (1989) Mammals of Victoria from the collection of Donald F. Thomson. Victorian Naturalist, 106, 4 - 25." type="journal article" year="1989">Dixon &amp; Huxley (1989)</bibRefCitation>
reported extracts concerning Victorian
<taxonomicName id="4C094D73FFD7831CD380FEF4D34AFEC0" ID-CoL="4FVN8" box="[1183,1324,296,320]" class="Mammalia" family="Dasyuridae" genus="Phascogale" kingdom="Animalia" order="Dasyuromorphia" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="tapoatafa">
<emphasis id="B97DEAE2FFD7831CD380FEF4D34AFEC0" box="[1183,1324,296,320]" italics="true" pageId="34" pageNumber="35">P. tapoatafa</emphasis>
</taxonomicName>
from the natural history notes of Donald F. Thomson, drawn primarily from sources on the Bellarine Peninsula where the species was so common in the 1940s1960s that it invaded local residences and was regarded as a nuisance. There are no modern records of the species in this area.
</paragraph>
</subSubSection>
</treatment>
</document>