265 lines
27 KiB
XML
265 lines
27 KiB
XML
<document id="43B4D2175ECD71E013AD75CC0A2D7CC2" ID-CLB-Dataset="45364" ID-DOI="10.11646/zootaxa.3826.1.3" ID-GBIF-Dataset="6f4b1535-6c6b-4446-aa26-b1ba83ad9126" ID-ISSN="1175-5326" ID-Zenodo-Dep="228537" ID-ZooBank="53549D58-8F38-47B5-879F-0245E900C131" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460444968431" checkinUser="plazi" docAuthor="Merle, Didier, Pacaud, Jean-Michel, Métais, Grégoire, Bartolini, Annachiara, Lashari, Rafiq A., Brohi, Imdad A., Solangi, Sarfraz H., Marivaux, Laurent & Welcomme, Jean-Loup" docDate="2014" docId="03A087E00E5EFFA5FF7FFDCAFCE1F8C5" docLanguage="en" docName="zt03826p138.pdf" docOrigin="Zootaxa 3826 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Athleta (Volutocorbis) lasharii Merle, Pacaud, Métais, Bartolini, Lashari, Brohi, Solangi, Marivaux & Welcomme, 2014, sp. nov." docType="treatment" docUuid="8E5B6D39-3B91-4CD8-AF72-1545807BB42A" docUuidSource="ZooBank" docVersion="10" lastPageNumber="124" masterDocId="FF99FF980E49FFB2FFE8FFEEFFDAFFAB" masterDocTitle="Volutidae (Mollusca: Gastropoda) of the Lakhra Formation (Earliest Eocene, Sindh, Pakistan): systematics, biostratigraphy and paleobiogeography" masterLastPageNumber="138" masterPageNumber="101" pageNumber="124" updateTime="1698333283847" updateUser="ExternalLinkService">
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<mods:mods id="085402123F4AECF1E615ED5798BA8D0D" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="B7D2A136C55A9603B92C41D8977337AD">
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<mods:title id="AF8F756AC040C515E305D11A031626F1">Volutidae (Mollusca: Gastropoda) of the Lakhra Formation (Earliest Eocene, Sindh, Pakistan): systematics, biostratigraphy and paleobiogeography</mods:title>
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<mods:roleTerm id="330430F9EC5F8E549017727AF95E7045">Author</mods:roleTerm>
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<mods:namePart id="988226A774AD0A1233814516BC30B72F">Merle, Didier</mods:namePart>
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<mods:namePart id="ED4DEBC35E02A5AE8E8FF1CC8731FD76">Pacaud, Jean-Michel</mods:namePart>
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</mods:name>
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<mods:name id="1AD424129EA1DA1019DEA32740EE1D6D" type="personal">
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<mods:roleTerm id="4838EA09EBFF5BDBC4DD4DF86936F91E">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="6E8B3866B78316886646DBCAC5B0834C">Métais, Grégoire</mods:namePart>
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</mods:name>
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<mods:name id="D0B3BB299E6D07703738F318F6A99761" type="personal">
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<mods:roleTerm id="40577747B4FF387E88BDCF0F97846C68">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="2CC63836617E94F1D394B6B616DC0103">Bartolini, Annachiara</mods:namePart>
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</mods:name>
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<mods:name id="8EC5AC28407A34831086D712D93DEE6D" type="personal">
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<mods:roleTerm id="AC4AB435A2B6BBA26F77C6497511FD50">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="3BD10821F535C992E53221BB7FFD9C74">Lashari, Rafiq A.</mods:namePart>
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</mods:name>
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<mods:roleTerm id="215F9442E4D2B103106723ADFD1AA9A4">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="1B2200B515CC7D0AF3A70502A203B0CB">Brohi, Imdad A.</mods:namePart>
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</mods:name>
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<mods:name id="21CAC34E6CC6D61B2BBD1FDDE972949C" type="personal">
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<mods:roleTerm id="6695EE62629CD875B7F670FB68302939">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="532F91717E30C0937A38901E7C119F40">Solangi, Sarfraz H.</mods:namePart>
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</mods:name>
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</mods:role>
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<mods:namePart id="C7C22D154C010FA1F4EB59803B0228DF">Marivaux, Laurent</mods:namePart>
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</mods:role>
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<mods:namePart id="3F5BC6F5077EEA86437A1EEB95F7D147">Welcomme, Jean-Loup</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="403176F9FFB0C4C378DA587A0628432F">text</mods:typeOfResource>
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<mods:titleInfo id="185617F11095611580736DBCDA9C20F1">
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<mods:title id="9F816149E86C6A151150A6E237E1CA6E">Zootaxa</mods:title>
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<mods:part id="6D7F3615B0E632CBADE6C490F9A3B7F4">
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<mods:date id="F4C02C519CA43122DE41BF3E623FCE4E">2014</mods:date>
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<mods:detail id="3D3E81A6F2E1A08585FE5B603D7C343C" type="volume">
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<mods:number id="5E242C86D8054C42C284431FB609E961">3826</mods:number>
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<mods:detail id="FC7663446F52A156956C9ED3A44AC5C9" type="issue">
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<mods:number id="8A64D5BD4BE2938FF64A5FED453A0A07">1</mods:number>
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<mods:start id="A7F1B7A6AFDAF9C465D27400939B2407">101</mods:start>
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<mods:end id="7217735F7688C60E5FA5B86314C46FB1">138</mods:end>
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<mods:classification id="330C493E4221630E564316888C938EFB">journal article</mods:classification>
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<mods:identifier id="440AE29EA3B302349483109A8E2D3762" type="CLB-Dataset">45364</mods:identifier>
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<mods:identifier id="BBB2B18235C504B82B932CFCD23017E2" type="DOI">10.11646/zootaxa.3826.1.3</mods:identifier>
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<mods:identifier id="EF9E3118C5E5818FDBF09EBE95511A62" type="GBIF-Dataset">6f4b1535-6c6b-4446-aa26-b1ba83ad9126</mods:identifier>
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<mods:identifier id="405E666873E34EBDBC85770C1A5ED6D2" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="6EC73FE911FFE627B289D995A7B0B029" type="Zenodo-Dep">228537</mods:identifier>
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<treatment id="03A087E00E5EFFA5FF7FFDCAFCE1F8C5" ID-DOI="http://doi.org/10.5281/zenodo.6137852" ID-GBIF-Taxon="119509260" ID-Zenodo-Dep="6137852" LSID="urn:lsid:zoobank.org:act:8E5B6D39-3B91-4CD8-AF72-1545807BB42A" httpUri="http://treatment.plazi.org/id/03A087E00E5EFFA5FF7FFDCAFCE1F8C5" lastPageNumber="124" pageId="23" pageNumber="124">
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<subSubSection id="C313657D0E5EFFA5FF7FFDCAFEF5FDF4" pageId="23" pageNumber="124" type="nomenclature">
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<paragraph id="8BB636F60E5EFFA5FF7FFDCAFDB3FD95" blockId="23.[151,617,548,607]" box="[151,617,548,574]" pageId="23" pageNumber="124">
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<heading id="D0FE819A0E5EFFA5FF7FFDCAFDB3FD95" bold="true" box="[151,617,548,574]" fontSize="11" level="1" pageId="23" pageNumber="124" reason="1">
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||
<emphasis id="B97DEAE40E5EFFA5FF7FFDCAFDB3FD95" bold="true" box="[151,617,548,574]" pageId="23" pageNumber="124">
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||
<taxonomicName id="4C094D750E5EFFA5FF7FFDCAFDDFFD95" box="[151,517,548,574]" class="Gastropoda" family="Volutidae" genus="Athleta" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="124" phylum="Mollusca" rank="species" species="lasharii" status="sp. nov." subGenus="Volutocorbis">
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<emphasis id="B97DEAE40E5EFFA5FF7FFDCAFDDFFD95" bold="true" box="[151,517,548,574]" italics="true" pageId="23" pageNumber="124">Athleta (Volutocorbis) lasharii</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A24E579F0E5EFFA5FDE4FDCAFDB3FD95" box="[524,617,548,574]" pageId="23" pageNumber="124" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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<paragraph id="8BB636F60E5EFFA5FF7FFDA8FEF5FDF4" blockId="23.[151,617,548,607]" box="[151,303,582,607]" pageId="23" pageNumber="124">
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||
(
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<figureCitation id="13322A730E5EFFA5FF77FDA8FEF2FDF4" box="[159,296,582,607]" captionStart="FIGURE 10. A – D" captionStartId="24.[151,250,1920,1942]" captionTargetBox="[236,1350,195,1906]" captionTargetId="figure@24.[236,1350,195,1907]" captionTargetPageId="24" captionText="FIGURE 10. A – D. Athleta (Volutocorbis) lasharii sp. nov., A – C (holotype, CPAG. RAN. I. 37, dorsal (A) and ventral (B) views, enlarged view of the spire (C), stn 4), D (paratype, CPAG. RAN. I. 38, dorsal view, stn 4); E – H. Volutilithes welcommei sp. nov., E (paratype 1, CPAG. RAN. I. 40, dorsal view, stn 4), F (paratype 2, CPAG. RAN. I. 41, juvenile, dorsal view, stn 4), G – H (holotype, CPAG. RAN. I. 39, ventral (G) and (H) dorsal views. Scale bars: A – B, D – E, G – H = 10 mm; C, F = 5 mm." httpUri="https://zenodo.org/record/228547/files/figure.png" pageId="23" pageNumber="124">Fig. 10A–D</figureCitation>
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)
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C313657D0E5EFFA5FF7FFD60FE1CFD60" pageId="23" pageNumber="124" type="etymology">
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||
<paragraph id="8BB636F60E5EFFA5FF7FFD60FE1CFD60" blockId="23.[151,1437,654,1903]" pageId="23" pageNumber="124">
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||
<emphasis id="B97DEAE40E5EFFA5FF7FFD60FEC4FD0C" bold="true" box="[151,286,654,679]" pageId="23" pageNumber="124">Etymology.</emphasis>
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Dedicated to Doctor Rafique Ahmed Lashari, Centre for Pure and Applied Geology, University of Sindh, Jamshoro,
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<collectingCountry id="F31E76660E5EFFA5FEB7FD5CFE18FD60" box="[351,450,690,715]" name="Pakistan" pageId="23" pageNumber="124">Pakistan</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C313657D0E5EFFA5FF2FFD38FD5DFCF0" pageId="23" pageNumber="124" type="materials_examined">
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<paragraph id="8BB636F60E5EFFA5FF2FFD38FB6DFD44" blockId="23.[151,1437,654,1903]" box="[199,1207,726,751]" pageId="23" pageNumber="124">
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||
<emphasis id="B97DEAE40E5EFFA5FF2FFD38FEBFFD44" bold="true" box="[199,357,726,751]" pageId="23" pageNumber="124">
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||
<typeStatus id="54B288540E5EFFA5FF2FFD38FED8FD44" box="[199,258,726,751]" pageId="23" pageNumber="124">Type</typeStatus>
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locality.
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</emphasis>
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Stn 4: Lakhra Dome, Lakhra village section, base of the Lakhra Formation.
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</paragraph>
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<paragraph id="8BB636F60E5EFFA5FF2FFD14FDB4FC9C" blockId="23.[151,1437,654,1903]" pageId="23" pageNumber="124">
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<emphasis id="B97DEAE40E5EFFA5FF2FFD14FE5EFCB8" bold="true" box="[199,388,762,787]" pageId="23" pageNumber="124">
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||
<typeStatus id="54B288540E5EFFA5FF2FFD14FED8FCB8" box="[199,258,762,787]" pageId="23" pageNumber="124">Type</typeStatus>
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material.
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</emphasis>
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<typeStatus id="54B288540E5EFFA5FE72FD14FDD8FCB8" box="[410,514,762,787]" pageId="23" pageNumber="124" type="holotype">Holotype</typeStatus>
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(stn 4:
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<collectionCode id="ED18AE330E5EFFA5FD6DFD15FD0FFCB8" box="[645,725,763,787]" pageId="23" pageNumber="124">CPAG</collectionCode>
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.RAN.
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<date id="FFB710360E5EFFA5FCF3FD15FC98FCB8" box="[795,834,763,787]" pageId="23" pageNumber="124" value="1937-01">I.37</date>
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, cast
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<collectionCode id="ED18AE330E5EFFA5FC77FD14FC20FCB8" LSID="urn:lsid:biocol.org:col:34988" box="[927,1018,762,787]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="23" pageNumber="124">MNHN</collectionCode>
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.F.
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<accessionNumber id="945AAB150E5EFFA5FBF8FD14FBB4FCB8" box="[1040,1134,762,787]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/A50375" pageId="23" pageNumber="124" type="EnaNcbi">A50375</accessionNumber>
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), one
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<typeStatus id="54B288540E5EFFA5FB27FD15FAF5FCB8" box="[1231,1327,763,787]" pageId="23" pageNumber="124" type="paratype">paratype</typeStatus>
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(stn 4:
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<collectionCode id="ED18AE330E5EFFA5FF7FFCF1FF3DFC9C" box="[151,231,799,823]" pageId="23" pageNumber="124">CPAG</collectionCode>
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.RAN.
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<date id="FFB710360E5EFFA5FEC4FCF1FE8EFC9C" box="[300,340,799,823]" pageId="23" pageNumber="124" value="1938-01">I.38</date>
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, cast
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<collectionCode id="ED18AE330E5EFFA5FE7AFCF0FE37FC9C" LSID="urn:lsid:biocol.org:col:34988" box="[402,493,798,823]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="23" pageNumber="124">MNHN</collectionCode>
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||
.F.A50376).
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||
</paragraph>
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||
<paragraph id="8BB636F60E5EFFA5FF2FFCACFD5DFCF0" blockId="23.[151,1437,654,1903]" box="[199,647,834,859]" pageId="23" pageNumber="124">
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<emphasis id="B97DEAE40E5EFFA5FF2FFCACFE5BFCF0" bold="true" box="[199,385,834,859]" pageId="23" pageNumber="124">Other material.</emphasis>
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||
2 spm (stn 4:
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<collectionCode id="ED18AE330E5EFFA5FDC9FCADFDA6FCF0" LSID="urn:lsid:biocol.org:col:34988" box="[545,636,835,859]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="23" pageNumber="124">MNHN</collectionCode>
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).
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C313657D0E5EFFA5FF2FFC88FCE1F8C5" pageId="23" pageNumber="124" type="description">
|
||
<paragraph id="8BB636F60E5EFFA5FF2FFC88FE45FADC" blockId="23.[151,1437,654,1903]" pageId="23" pageNumber="124">
|
||
<emphasis id="B97DEAE40E5EFFA5FF2FFC88FE83FCD4" bold="true" box="[199,345,870,895]" pageId="23" pageNumber="124">Description.</emphasis>
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||
Shell biconic, slightly fusiform, H 40–48, D ca.
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||
<quantity id="4CF19B130E5EFFA5FC6AFC89FC26FCD5" box="[898,1020,871,895]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" metricValueMax="2.2" metricValueMin="2.0" pageId="23" pageNumber="124" unit="mm" value="21.0" valueMax="22.0" valueMin="20.0">20–22 mm</quantity>
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||
(
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||
<typeStatus id="54B288540E5EFFA5FBE3FC88FBB5FCD4" box="[1035,1135,870,895]" pageId="23" pageNumber="124" type="holotype">holotype</typeStatus>
|
||
H 43.5, D
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||
<quantity id="4CF19B130E5EFFA5FB07FC88FA8CFCD5" box="[1263,1366,870,895]" metricMagnitude="-2" metricUnit="m" metricValue="1.96" pageId="23" pageNumber="124" unit="mm" value="19.6">19.6 mm</quantity>
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||
, outer lip not complete. Protoconch and two first teleoconch whorls unknown; at least 5 teleoconch whorls. Spire moderately high, occupying 20% of total shell height. Spire whorls weakly convex, last whorl rather oval. Spiral sculpture on spire of a first row of rounded subsutural nodules, appearing on third whorl and becoming more developed aperturally; abapically, three rows of rounded nodules including shoulder row, which is less distinct than others (
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<figureCitation id="13322A730E5EFFA5FF02FBF4FE8BFB98" box="[234,337,1050,1075]" captionStart="FIGURE 10. A – D" captionStartId="24.[151,250,1920,1942]" captionTargetBox="[236,1350,195,1906]" captionTargetId="figure@24.[236,1350,195,1907]" captionTargetPageId="24" captionText="FIGURE 10. A – D. Athleta (Volutocorbis) lasharii sp. nov., A – C (holotype, CPAG. RAN. I. 37, dorsal (A) and ventral (B) views, enlarged view of the spire (C), stn 4), D (paratype, CPAG. RAN. I. 38, dorsal view, stn 4); E – H. Volutilithes welcommei sp. nov., E (paratype 1, CPAG. RAN. I. 40, dorsal view, stn 4), F (paratype 2, CPAG. RAN. I. 41, juvenile, dorsal view, stn 4), G – H (holotype, CPAG. RAN. I. 39, ventral (G) and (H) dorsal views. Scale bars: A – B, D – E, G – H = 10 mm; C, F = 5 mm." httpUri="https://zenodo.org/record/228547/files/figure.png" pageId="23" pageNumber="124">Fig. 10C</figureCitation>
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). On last whorl, spiral sculpture displaying: 1) subsutural row of spiny nodules; 2) shoulder row of spiny nodules; 3) between shoulder and mid-whorl, 9 to 10 rows of almost flat cords, separated by narrow spiral grooves; 4) from mid-whorl to base, nodules and cords produce imbricate effect. From second to penultimate whorl, costae interrupted by rows of nodules. On last whorl, costae obsolete between subsutural and shoulder nodules, narrow but relatively high below shoulder angle, extending across base, becoming almost obsolete towards end of last whorl. Second whorl: 12 costae; third whorl: 14 costae; fourth whorl: 12–14 costae; fifth and last whorl: 9–13 costae. Aperture ovate, rather narrow, occupying 72% of total height, 61% of diameter. Inner lip straight, columella with one fold, succeeded posteriorly by several weaker and smaller folds; inner lip callus very narrow, spreading a little anteriorly. Outer lip thin, not thickened externally. Siphonal canal slightly elongate; siphonal notch shallow.
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</paragraph>
|
||
<paragraph id="8BB636F60E5EFFA5FF2FFA6CFD76F8E1" blockId="23.[151,1437,654,1903]" pageId="23" pageNumber="124">
|
||
<emphasis id="B97DEAE40E5EFFA5FF2FFA6CFEB6FA30" bold="true" box="[199,364,1410,1435]" pageId="23" pageNumber="124">Comparisons.</emphasis>
|
||
<taxonomicName id="4C094D750E5EFFA5FE9BFA6CFD1DFA30" box="[371,711,1410,1435]" class="Gastropoda" family="Volutidae" genus="Athleta" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="124" phylum="Mollusca" rank="species" species="lasharii" status="sp. nov." subGenus="Volutocorbis">
|
||
<emphasis id="B97DEAE40E5EFFA5FE9BFA6CFD1DFA30" box="[371,711,1410,1435]" italics="true" pageId="23" pageNumber="124">Athleta (Volutocorbis) lasharii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24E579F0E5EFFA5FD26FA6CFCFDFA30" box="[718,807,1410,1435]" pageId="23" pageNumber="124" rank="species">
|
||
<emphasis id="B97DEAE40E5EFFA5FD26FA6CFCFDFA30" bold="true" box="[718,807,1410,1435]" pageId="23" pageNumber="124">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
differs from the other Sindh species of
|
||
<emphasis id="B97DEAE40E5EFFA5FB35FA6DFA46FA30" box="[1245,1436,1410,1435]" italics="true" pageId="23" pageNumber="124">
|
||
A. (
|
||
<taxonomicName id="4C094D750E5EFFA5FAECFA6CFA4EFA30" box="[1284,1428,1410,1435]" class="Gastropoda" family="Volutidae" genus="Volutocorbis" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="124" phylum="Mollusca" rank="genus">Volutocorbis</taxonomicName>
|
||
)
|
||
</emphasis>
|
||
in its larger size, its spiral sculpture on the last whorl characterized by the loss of crenulations or nodules between the shoulder and the mid-whorl and by its axial sculpture of higher, narrow costae. Such sculpture distinguishes it from the
|
||
<emphasis id="B97DEAE40E5EFFA5FF14FA01FE10F9AC" box="[252,458,1518,1543]" italics="true" pageId="23" pageNumber="124">A. (V.) digitalinus-</emphasis>
|
||
<typeStatus id="54B288540E5EFFA5FE22FA01FE20F9AC" box="[458,506,1519,1543]" pageId="23" pageNumber="124">type</typeStatus>
|
||
species bearing strongly nodular sculpture on the last whorl (e.g.
|
||
<taxonomicName id="4C094D750E5EFFA5FB3FFA01FAACF9AC" box="[1239,1398,1518,1543]" class="Gastropoda" family="Volutidae" genus="Athleta" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="124" phylum="Mollusca" rank="species" species="burtoni" subGenus="Volutocorbis">
|
||
<emphasis id="B97DEAE40E5EFFA5FB3FFA01FAACF9AC" box="[1239,1398,1518,1543]" italics="true" pageId="23" pageNumber="124">A. (V.) burtoni</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C094D750E5EFFA5FA6CFA01FEC9F980" class="Gastropoda" family="Volutidae" genus="Athleta" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="124" phylum="Mollusca" rank="species" species="daviesi" subGenus="Volutocorbis">
|
||
<emphasis id="B97DEAE40E5EFFA5FA6CFA01FEC9F980" italics="true" pageId="23" pageNumber="124">A. (V.) daviesi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C094D750E5EFFA5FEA2F9FDFE20F980" box="[330,506,1555,1579]" class="Gastropoda" family="Volutidae" genus="Volutocorbis" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="124" phylum="Mollusca" rank="species" species="soriensis" subGenus="Volutocorbis">
|
||
<emphasis id="B97DEAE40E5EFFA5FEA2F9FDFE20F980" box="[330,506,1555,1579]" italics="true" pageId="23" pageNumber="124">A. (V.) soriensis</emphasis>
|
||
</taxonomicName>
|
||
from the Early Paleogene of
|
||
<collectingCountry id="F31E76660E5EFFA5FCA9F9FCFC78F980" box="[833,930,1554,1579]" name="Pakistan" pageId="23" pageNumber="124">Pakistan</collectingCountry>
|
||
) and the
|
||
<taxonomicName id="4C094D750E5EFFA5FBE0F9FDFB6AF980" box="[1032,1200,1554,1579]" class="Gastropoda" family="Volutidae" genus="Athleta" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="124" phylum="Mollusca" rank="species" species="elevatus" subGenus="Volutocorbis">
|
||
<emphasis id="B97DEAE40E5EFFA5FBE0F9FDFB6AF980" box="[1032,1200,1554,1579]" italics="true" pageId="23" pageNumber="124">A. (V.) elevatus</emphasis>
|
||
</taxonomicName>
|
||
-
|
||
<typeStatus id="54B288540E5EFFA5FB50F9FDFB30F980" box="[1208,1258,1555,1579]" pageId="23" pageNumber="124">type</typeStatus>
|
||
species bearing crenulate sculpture (e.g.
|
||
<taxonomicName id="4C094D750E5EFFA5FE5AF9D9FDB3F9E5" box="[434,617,1591,1614]" class="Gastropoda" family="Volutidae" genus="Athleta" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="124" phylum="Mollusca" rank="species" species="eugeniae" subGenus="Volutocorbis">
|
||
<emphasis id="B97DEAE40E5EFFA5FE5AF9D9FDB3F9E5" box="[434,617,1591,1614]" italics="true" pageId="23" pageNumber="124">A. (V.) eugeniae</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C094D750E5EFFA5FD4CF9D9FC9CF9E5" box="[676,838,1591,1614]" class="Gastropoda" family="Volutidae" genus="Athleta" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="124" phylum="Mollusca" rank="species" species="wynnei" subGenus="Volutocorbis">
|
||
<emphasis id="B97DEAE40E5EFFA5FD4CF9D9FC9CF9E5" box="[676,838,1591,1614]" italics="true" pageId="23" pageNumber="124">A. (V.) wynnei</emphasis>
|
||
</taxonomicName>
|
||
from the Early Paleogene of
|
||
<collectingCountry id="F31E76660E5EFFA5FB75F9D8FADBF9E4" box="[1181,1281,1590,1615]" name="Pakistan" pageId="23" pageNumber="124">Pakistan</collectingCountry>
|
||
). The
|
||
<typeStatus id="54B288540E5EFFA5FAA3F9D9FAA1F9E4" box="[1355,1403,1591,1615]" pageId="23" pageNumber="124">type</typeStatus>
|
||
of sculpture displayed in
|
||
<taxonomicName id="4C094D750E5EFFA5FE70F9B5FDE5F9D8" box="[408,575,1626,1651]" class="Gastropoda" family="Volutidae" genus="Athleta" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="124" phylum="Mollusca" rank="species" species="lasharii" status="sp. nov." subGenus="Volutocorbis">
|
||
<emphasis id="B97DEAE40E5EFFA5FE70F9B5FDE5F9D8" box="[408,575,1626,1651]" italics="true" pageId="23" pageNumber="124">A. (V.) lasharii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24E579F0E5EFFA5FDA0F9B4FD79F9D8" box="[584,675,1626,1651]" pageId="23" pageNumber="124" rank="species">
|
||
<emphasis id="B97DEAE40E5EFFA5FDA0F9B4FD79F9D8" bold="true" box="[584,675,1626,1651]" pageId="23" pageNumber="124">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
more resembles that of
|
||
<taxonomicName id="4C094D750E5EFFA5FC50F9B5FAE8F9D8" authority="Lamarck, 1803" authorityName="Lamarck" authorityYear="1803" box="[952,1330,1626,1651]" class="Gastropoda" family="Volutidae" genus="Athleta" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="124" phylum="Mollusca" rank="species" species="bicorona" subGenus="Volutocorbis">
|
||
<emphasis id="B97DEAE40E5EFFA5FC50F9B5FBB6F9D8" box="[952,1132,1626,1651]" italics="true" pageId="23" pageNumber="124">A. (V.) bicorona</emphasis>
|
||
(
|
||
<bibRefCitation id="EF984B070E5EFFA5FB94F9B4FAF0F9D8" author="Lamarck" box="[1148,1322,1626,1651]" pageId="23" pageNumber="124" refString="Lamarck, J. - B. (1803) Memoires sur les fossiles des environs de Paris comprenant la determination des especes qui appartiennent aux animaux marins sans vertebres, et dont la plupart sont figures dans la collection des velins du Museum. Annales du Museum d'Histoire naturelle, 1, 474 - 479." type="journal article" year="1803">Lamarck, 1803</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
from the Lutetian of the Paris Basin.
|
||
<taxonomicName id="4C094D750E5EFFA5FE27F991FD5AF93C" box="[463,640,1662,1687]" class="Gastropoda" family="Volutidae" genus="Athleta" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="124" phylum="Mollusca" rank="species" species="bicorona" subGenus="Volutocorbis">
|
||
<emphasis id="B97DEAE40E5EFFA5FE27F991FD5AF93C" box="[463,640,1662,1687]" italics="true" pageId="23" pageNumber="124">A. (V.) bicorona</emphasis>
|
||
</taxonomicName>
|
||
shares a last whorl characterized by the loss of crenulations or nodules between the shoulder and the mid-whorl. Moreover, both species have two rows of spiny nodules developed adapically (the subsutural and shoulder rows).
|
||
<taxonomicName id="4C094D750E5EFFA5FD49F929FC91F974" box="[673,843,1734,1759]" class="Gastropoda" family="Volutidae" genus="Athleta" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="124" phylum="Mollusca" rank="species" species="bicorona" subGenus="Volutocorbis">
|
||
<emphasis id="B97DEAE40E5EFFA5FD49F929FC91F974" box="[673,843,1734,1759]" italics="true" pageId="23" pageNumber="124">A (V.) bicorona</emphasis>
|
||
</taxonomicName>
|
||
differs, however, from
|
||
<taxonomicName id="4C094D750E5EFFA5FBBBF929FB2DF974" box="[1107,1271,1734,1759]" class="Gastropoda" family="Volutidae" genus="Athleta" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="124" phylum="Mollusca" rank="species" species="lasharii" status="sp. nov." subGenus="Volutocorbis">
|
||
<emphasis id="B97DEAE40E5EFFA5FBBBF929FB2DF974" box="[1107,1271,1734,1759]" italics="true" pageId="23" pageNumber="124">A. (V.) lasharii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24E579F0E5EFFA5FB16F928FA8DF974" box="[1278,1367,1734,1759]" pageId="23" pageNumber="124" rank="species">
|
||
<emphasis id="B97DEAE40E5EFFA5FB16F928FA8DF974" bold="true" box="[1278,1367,1734,1759]" pageId="23" pageNumber="124">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
by its early whorls having two more abapical spiral rows of crenulations. In
|
||
<taxonomicName id="4C094D750E5EFFA5FC21F905FB5AF8A8" box="[969,1152,1770,1795]" class="Gastropoda" family="Volutidae" genus="Athleta" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="124" phylum="Mollusca" rank="species" species="bicorona" subGenus="Volutocorbis">
|
||
<emphasis id="B97DEAE40E5EFFA5FC21F905FB5AF8A8" box="[969,1152,1770,1795]" italics="true" pageId="23" pageNumber="124">A. (V.) bicorona</emphasis>
|
||
</taxonomicName>
|
||
, a residual color pattern observed under UV light has been reported (
|
||
<bibRefCitation id="EF984B070E5EFFA5FD67F8E0FC83F88C" author="Merle" box="[655,857,1806,1831]" pageId="23" pageNumber="124" refString="Merle, D., Pacaud, J. - M., Kriloff, A. & Loubry, P. (2008) Les motifs colores residuels des coquilles lutetiennes du bassin de Paris. In: Merle, D. (Ed.), Stratotype Lutetien. Collection Patrimoine geologique, Biotope, Meze, MNHN, Paris, BRGM Orleans, pp. 182 - 227." type="book chapter" year="2008">
|
||
Merle
|
||
<emphasis id="B97DEAE40E5EFFA5FD34F8E1FCCFF88C" box="[732,789,1806,1831]" italics="true" pageId="23" pageNumber="124">et al.</emphasis>
|
||
2008
|
||
</bibRefCitation>
|
||
), but the recrystallized shells of
|
||
<taxonomicName id="4C094D750E5EFFA5FB21F8E1FAAAF88C" box="[1225,1392,1806,1831]" class="Gastropoda" family="Volutidae" genus="Athleta" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="124" phylum="Mollusca" rank="species" species="lasharii" subGenus="Volutocorbis">
|
||
<emphasis id="B97DEAE40E5EFFA5FB21F8E1FAAAF88C" box="[1225,1392,1806,1831]" italics="true" pageId="23" pageNumber="124">A. (V.) lasharii</emphasis>
|
||
</taxonomicName>
|
||
did not reveal any traces of a residual color pattern.
|
||
</paragraph>
|
||
<paragraph id="8BB636F60E5EFFA5FF2FF8B8FCE1F8C5" blockId="23.[151,1437,654,1903]" box="[199,827,1878,1903]" pageId="23" pageNumber="124">
|
||
<emphasis id="B97DEAE40E5EFFA5FF2FF8B8FE62F8C4" bold="true" box="[199,440,1878,1903]" pageId="23" pageNumber="124">Stratigraphic range.</emphasis>
|
||
Lakhra Formation: Lakhra Dome.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |