treatments-xml/data/03/A0/87/03A087A90F762D04FF70F926FE503C97.xml

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<mods:title id="5FE176DF007CFDFDCF2A4267550067CE">Heissiella donguri sp. nov. (Hemiptera: Heteroptera: Tingidae) feeding on Quercus gilva (Fagaceae): First representative of the genus from Japan</mods:title>
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<mods:date id="79A6F62F2D10A98AB6F27BBC4D36A9FA">2020</mods:date>
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<taxonomicName id="4C094D3C0F762D03FF70F926FEEE3DB7" authority="Souma &amp; Ishikawa, 2020" authorityName="Souma &amp; Ishikawa" authorityYear="2020" box="[151,373,1746,1773]" class="Insecta" family="Tingidae" genus="Heissiella" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="378" phylum="Arthropoda" rank="species" species="donguri" status="sp. nov.">
<emphasis id="B97DEAAD0F762D03FF70F926FEEE3DB7" bold="true" box="[151,373,1746,1773]" italics="true" pageId="1" pageNumber="378">Heissiella donguri</emphasis>
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(
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A–D, 2A–G, 3A–E, 4A)
</paragraph>
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<treatmentCitationGroup id="AB1911910F762D03FF70F8CFFBA73C09" box="[151,1084,1850,1875]" pageId="1" pageNumber="378">
<taxonomicName id="4C094D3C0F762D03FF70F8CFFEE23C09" ID-CoL="8GPV" authority="" box="[151,377,1850,1875]" class="Insecta" family="Tingidae" genus="Tingis" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="378" phylum="Arthropoda" rank="subGenus" subGenus="Tropidocheila">
<emphasis id="B97DEAAD0F762D03FF70F8CFFF4E3C08" box="[151,213,1851,1874]" italics="true" pageId="1" pageNumber="378">Tingis</emphasis>
(
<emphasis id="B97DEAAD0F762D03FF03F8CEFEEA3C08" box="[228,369,1850,1874]" italics="true" pageId="1" pageNumber="378">Tropidocheila</emphasis>
)
</taxonomicName>
sp.: 
<treatmentCitation id="0AA810AE0F762D03FE4DF8CFFD183C09" author="Komatsu, T." box="[426,643,1850,1875]" page="101" pageId="1" pageNumber="378" year="2016">
<bibRefCitation id="EF984B4E0F762D03FE4DF8CFFD183C09" author="Komatsu, T." box="[426,643,1850,1875]" pageId="1" pageNumber="378" refId="ref5230" refString="Komatsu, T. (2016) [Terrestrial heteropterans from Miyazaki-ken, Kyushu, Japan]. Kuroshio Bunko, Miyazaki, 139 pp. [in Japanese]" type="book" year="2016">Komatsu (2016: 101)</bibRefCitation>
</treatmentCitation>
(distribution); 
<treatmentCitation id="0AA810AE0F762D03FCFBF8CFFC323C09" author="Ito, R." box="[796,937,1851,1875]" page="77" pageId="1" pageNumber="378" year="2019">
<bibRefCitation id="EF984B4E0F762D03FCFBF8CFFC323C09" author="Ito, R." box="[796,937,1851,1875]" pageId="1" pageNumber="378" pagination="77 - 78" refId="ref5096" refString="Ito, R. (2019) [First record of Dictyla evidens (Drake, 1927) from Oita-ken, Kyushu, Japan, with a correction of distributional records of Tingidae from Oita-ken]. Niho-no-Mushi, 57, 77 - 78. [in Japanese]" type="journal article" year="2019">Ito (2019: 77)</bibRefCitation>
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(distribution).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BB636BF0F762D03FF70F8AEFE463CCA" blockId="1.[151,1436,1850,1936]" pageId="1" pageNumber="378">
<treatmentCitationGroup id="AB1911910F762D03FF70F8AEFE463CCA" pageId="1" pageNumber="378">
<taxonomicName id="4C094D3C0F762D03FF70F8AEFE693C2B" ID-CoL="572ZQ" authorityName="Drake &amp; Maa" authorityYear="1953" box="[151,498,1881,1905]" class="Insecta" family="Tingidae" genus="Tingis" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="378" phylum="Arthropoda" rank="species" species="shaowuana" subGenus="Tropidocheila">
<emphasis id="B97DEAAD0F762D03FF70F8AEFF4E3C2B" box="[151,213,1882,1905]" italics="true" pageId="1" pageNumber="378">Tingis</emphasis>
(
<emphasis id="B97DEAAD0F762D03FF03F8ADFEEA3C2B" box="[228,369,1881,1905]" italics="true" pageId="1" pageNumber="378">Tropidocheila</emphasis>
) 
<emphasis id="B97DEAAD0F762D03FE98F8ADFE693C2B" box="[383,498,1881,1905]" italics="true" pageId="1" pageNumber="378">shaowuana</emphasis>
</taxonomicName>
(non Drake &amp; Maa, 1953): 
<treatmentCitation id="0AA810AE0F762D03FCEAF8ACFC513C2B" author="Miyake, T." box="[781,970,1880,1905]" page="22" pageId="1" pageNumber="378" year="2017">
<bibRefCitation id="EF984B4E0F762D03FCEAF8ACFC513C2B" author="Miyake, T." box="[781,970,1880,1905]" pageId="1" pageNumber="378" pagination="22" refId="ref5292" refString="Miyake, T. (2017) [Some distributional records of insects from Ume district, Saeki-shi, Oita-ken, Kyushu, Japan]. Niho-no- Mushi, 55, 22. [in Japanese]" type="journal article" year="2017">Miyake (2017: 22)</bibRefCitation>
</treatmentCitation>
(distribution); 
<treatmentCitation id="0AA810AE0F762D03FB84F8AEFACA3C2B" author="Ito, R. &amp; Sasaki, S." box="[1123,1361,1880,1905]" page="21" pageId="1" pageNumber="378" year="2018">
<bibRefCitation id="EF984B4E0F762D03FB84F8AEFACA3C2B" author="Ito, R. &amp; Sasaki, S." box="[1123,1361,1880,1905]" pageId="1" pageNumber="378" pagination="17 - 24" refId="ref5145" refString="Ito, R. &amp; Sasaki, S. (2018) [Tingidae from Oita-ken, Kyushu, Japan (Heteroptera)]. Niho-no-Mushi, 56, 17 - 24. [in Japanese]" type="journal article" year="2018">Ito &amp; Sasaki (2018: 21)</bibRefCitation>
</treatmentCitation>
(distribution). Misidentifications.
</treatmentCitationGroup>
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<emphasis id="B97DEAAD0F762D03FF70F84AFEBF3C82" bold="true" box="[151,292,1981,2008]" pageId="1" pageNumber="378">Type series.</emphasis>
<materialsCitation id="3B613CE20F762D03FECCF84AFD283CA6" ID-GBIF-Occurrence="2577306107" collectingDate="2019-10-01" collectionCode="TUA" collectorName="J. Souma" country="Japan" county="Takabatake-cho" latitude="34.67553" location="Honshu" longLatPrecision="1" longitude="135.852" municipality="Nara-shi" pageId="1" pageNumber="378" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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<subScript id="178D34FA0F762D03FECCF84AFE3A3C8C" attach="left" box="[299,417,1982,2008]" fontSize="7" pageId="1" pageNumber="378">HOLOTYPE</subScript>
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(macropterous 
<specimenCount id="9D0FFD360F762D03FDB5F84BFDF23C83" box="[594,617,1983,2009]" pageId="1" pageNumber="378" type="male">♂</specimenCount>
), 
<emphasis id="B97DEAAD0F762D03FD98F849FCCA3C82" bold="true" box="[639,849,1981,2008]" pageId="1" pageNumber="378">
<collectingCountry id="F31E762F0F762D03FD98F849FD463C82" box="[639,733,1981,2008]" name="Japan" pageId="1" pageNumber="378">JAPAN</collectingCountry>
: 
<location id="8ED660640F762D03FD0DF84AFCD73C82" LSID="urn:lsid:plazi:treatment:03A087A90F762D04FF70F926FE503C97:8ED660640F762D03FD0DF84AFCD73C82" box="[746,844,1982,2008]" country="Japan" county="Takabatake-cho" latitude="34.67553" longLatPrecision="1" longitude="135.852" municipality="Nara-shi" name="Honshu" pageId="1" pageNumber="378">Honshu</location>
:
</emphasis>
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, 
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, 
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, 
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, 
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<collectingDate id="EFF3E9970F762D03FEB7F816FE2F3CA6" box="[336,436,2018,2044]" pageId="1" pageNumber="378" value="2019-10-01">1.x.2019</collectingDate>
</date>
, leg. 
<collectorName id="26FC53690F762D03FE14F817FDC03CA6" box="[499,603,2018,2045]" pageId="1" pageNumber="378">J. Souma</collectorName>
(
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)
</materialsCitation>
. 
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</typeStatus>
(macropterous 
<specimenCount id="9D0FFD360F762D03FC0BF816FBA03CA7" box="[1004,1083,2018,2045]" pageId="1" pageNumber="378" type="male">41 ♂♂</specimenCount>
<specimenCount id="9D0FFD360F762D03FBA4F816FB0A3CA7" box="[1091,1169,2018,2045]" pageId="1" pageNumber="378" type="female">27 ♀♀</specimenCount>
), 
<emphasis id="B97DEAAD0F762D03FB4DF815FAE73CA6" bold="true" box="[1194,1404,2017,2044]" pageId="1" pageNumber="378">
<collectingCountry id="F31E762F0F762D03FB4DF815FA933CA6" box="[1194,1288,2017,2044]" name="Japan" pageId="1" pageNumber="378">JAPAN</collectingCountry>
: 
<collectingRegion id="49CDF85D0F762D03FAF1F816FAE33CA6" box="[1302,1400,2018,2044]" pageId="1" pageNumber="378">Honshu</collectingRegion>
:
</emphasis>
as
</materialsCitation>
</paragraph>
<paragraph id="8BB636BF0F752D00FF70FF6DFBDC3A45" blockId="2.[151,1437,152,1007]" pageId="2" pageNumber="379">
<materialsCitation id="3B613CE20F752D00FF70FF6DFC9A3BE9" ID-GBIF-Occurrence="2577306108" box="[151,769,153,179]" collectionCode="TUA, HNHM" pageId="2" pageNumber="379" specimenCount="20" specimenCount-female="9" specimenCount-male="11" typeStatus="holotype">
<typeStatus id="54B2881D0F752D00FF70FF6DFF623BE9" box="[151,249,153,179]" pageId="2" pageNumber="379" type="holotype">holotype</typeStatus>
(
<specimenCount id="9D0FFD360F752D00FEEEFF6DFED23BE9" box="[265,329,153,179]" pageId="2" pageNumber="379" type="male">9 ♂♂</specimenCount>
<specimenCount id="9D0FFD360F752D00FEB6FF6DFE0A3BE9" box="[337,401,153,179]" pageId="2" pageNumber="379" type="female">7 ♀♀</specimenCount>
, 
<collectionCode id="ED18AE7A0F752D00FE79FF6DFE403BE9" box="[414,475,153,179]" pageId="2" pageNumber="379">TUA</collectionCode>
; 
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<specimenCount id="9D0FFD360F752D00FDC9FF6DFDF43BE9" box="[558,623,153,179]" pageId="2" pageNumber="379" type="female">2 ♀♀</specimenCount>
, 
<collectionCode id="ED18AE7A0F752D00FD9BFF6DFD4C3BE9" box="[636,727,153,179]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="2" pageNumber="379">HNHM</collectionCode>
); as
</materialsCitation>
<materialsCitation id="3B613CE20F752D00FCEEFF6DFBF83BE9" ID-GBIF-Occurrence="2577306109" box="[777,1123,153,179]" collectingDate="2017-10-14" collectionCode="R" pageId="2" pageNumber="379" specimenCount="1" typeStatus="holotype">
<typeStatus id="54B2881D0F752D00FCEEFF6DFCF03BE9" box="[777,875,153,179]" pageId="2" pageNumber="379" type="holotype">holotype</typeStatus>
but 
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<collectingDate id="EFF3E9970F752D00FC79FF6DFB8B3BE9" box="[926,1040,153,179]" pageId="2" pageNumber="379" value="2017-10-14">14.x.2017</collectingDate>
</date>
, leg. 
<collectionCode id="ED18AE7A0F752D00FBAAFF6DFBF83BE9" box="[1101,1123,153,179]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="2" pageNumber="379">R</collectionCode>
</materialsCitation>
. 
<materialsCitation id="3B613CE20F752D00FB89FF6DFA073BE9" ID-GBIF-Occurrence="2577306104" box="[1134,1436,152,179]" collectionCode="TUA" location="Ito" pageId="2" pageNumber="379" specimenCount="12" specimenCount-female="5" specimenCount-male="7" typeStatus="holotype">
<location id="8ED660640F752D00FB89FF6DFB163BE9" LSID="urn:lsid:plazi:treatment:03A087A90F762D04FF70F926FE503C97:8ED660640F752D00FB89FF6DFB163BE9" box="[1134,1165,153,179]" name="Ito" pageId="2" pageNumber="379">Ito</location>
(
<specimenCount id="9D0FFD360F752D00FB79FF6DFB453BE9" box="[1182,1246,153,179]" pageId="2" pageNumber="379" type="male">7 ♂♂</specimenCount>
<specimenCount id="9D0FFD360F752D00FB02FF6CFABD3BE9" box="[1253,1318,152,179]" pageId="2" pageNumber="379" type="female">5 ♀♀</specimenCount>
, 
<collectionCode id="ED18AE7A0F752D00FAD4FF6DFAEA3BE9" box="[1331,1393,153,179]" pageId="2" pageNumber="379">TUA</collectionCode>
); as
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<materialsCitation id="3B613CE20F752D00FF70FF49FC1E3B8D" ID-GBIF-Occurrence="2577306113" box="[151,901,189,215]" collectingDate="2017-06-10" collectingDateMax="2017-06-11" collectingDateMin="2017-06-10" collectionCode="TUA" location="Akita" pageId="2" pageNumber="379" specimenCount="27" specimenCount-female="8" specimenCount-male="19" stateProvince="Akita" typeStatus="holotype">
<typeStatus id="54B2881D0F752D00FF70FF49FF623B8D" box="[151,249,189,215]" pageId="2" pageNumber="379" type="holotype">holotype</typeStatus>
but 
<date id="FFB7107F0F752D00FECBFF49FE543B8D" box="[300,463,189,215]" pageId="2" pageNumber="379" value="2017-06-10" valueMax="2017-06-11" valueMin="2017-06-10">
<collectingDate id="EFF3E9970F752D00FECBFF49FE543B8D" box="[300,463,189,215]" pageId="2" pageNumber="379" value="2017-06-10" valueMax="2017-06-11" valueMin="2017-06-10">10–11.vi.2017</collectingDate>
</date>
, leg. K. 
<collectingRegion id="49CDF85D0F752D00FDCBFF49FDF13B8D" box="[556,618,189,215]" country="Japan" name="Akita" pageId="2" pageNumber="379">Akita</collectingRegion>
(
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, 
<collectionCode id="ED18AE7A0F752D00FCFBFF49FCC13B8D" box="[796,858,189,215]" pageId="2" pageNumber="379">TUA</collectionCode>
); as
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<materialsCitation id="3B613CE20F752D00FC6AFF49FE8E3BA1" ID-GBIF-Occurrence="2577306111" collectingDate="2017-11-23" collectionCode="TUA" location="Akita" pageId="2" pageNumber="379" specimenCount="6" specimenCount-female="3" specimenCount-male="3" stateProvince="Akita" typeStatus="holotype">
<typeStatus id="54B2881D0F752D00FC6AFF49FC743B8D" box="[909,1007,189,215]" pageId="2" pageNumber="379" type="holotype">holotype</typeStatus>
but 
<date id="FFB7107F0F752D00FBC6FF49FB003B8D" box="[1057,1179,189,215]" pageId="2" pageNumber="379" value="2017-11-23">
<collectingDate id="EFF3E9970F752D00FBC6FF49FB003B8D" box="[1057,1179,189,215]" pageId="2" pageNumber="379" value="2017-11-23">23.xi.2017</collectingDate>
</date>
, leg. K. 
<collectingRegion id="49CDF85D0F752D00FB1EFF49FAAC3B8D" box="[1273,1335,189,215]" country="Japan" name="Akita" pageId="2" pageNumber="379">Akita</collectingRegion>
(
<specimenCount id="9D0FFD360F752D00FAAFFF49FA1C3B8D" box="[1352,1415,189,215]" pageId="2" pageNumber="379" type="male">3 ♂♂</specimenCount>
<specimenCount id="9D0FFD360F752D00FA69FF49FF593BA1" pageId="2" pageNumber="379" type="female">3 ♀♀</specimenCount>
, 
<collectionCode id="ED18AE7A0F752D00FF29FF15FE963BA1" box="[206,269,225,251]" pageId="2" pageNumber="379">TUA</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="3B613CE20F752D00FEF8FF15FBA13BA1" ID-GBIF-Occurrence="2577306110" box="[287,1082,225,251]" collectingDate="2011-12-28" collectionCode="T" collectorName="Kyushu &amp; Oita-ken &amp; Saeki-shi &amp; Ume &amp; Kitakawa Dam" pageId="2" pageNumber="379" specimenCount="1" stateProvince="Akita" typeStatus="holotype">
<emphasis id="B97DEAAD0F752D00FEF8FF15FE1D3BA1" bold="true" box="[287,390,225,251]" pageId="2" pageNumber="379">
<collectorName id="26FC53690F752D00FEF8FF15FE1A3BA1" box="[287,385,225,251]" pageId="2" pageNumber="379">Kyushu</collectorName>
:
</emphasis>
<collectorName id="26FC53690F752D00FE6BFF15FE6A3BA1" box="[396,497,225,251]" pageId="2" pageNumber="379">Oita-ken</collectorName>
, 
<collectorName id="26FC53690F752D00FE1BFF15FDFC3BA1" box="[508,615,225,251]" pageId="2" pageNumber="379">Saeki-shi</collectorName>
, 
<collectorName id="26FC53690F752D00FD95FF15FD303BA1" box="[626,683,225,251]" pageId="2" pageNumber="379">Ume</collectorName>
, 
<collectorName id="26FC53690F752D00FD51FF15FCFA3BA1" box="[694,865,225,251]" pageId="2" pageNumber="379">Kitakawa Dam</collectorName>
, 
<date id="FFB7107F0F752D00FC8BFF15FC763BA1" box="[876,1005,225,251]" pageId="2" pageNumber="379" value="2011-12-28">
<collectingDate id="EFF3E9970F752D00FC8BFF15FC763BA1" box="[876,1005,225,251]" pageId="2" pageNumber="379" value="2011-12-28">28.xii.2011</collectingDate>
</date>
, leg. 
<collectionCode id="ED18AE7A0F752D00FBCFFF15FBA13BA1" box="[1064,1082,225,251]" name="Tavera, Department of Geology and Geophysics" pageId="2" pageNumber="379">T</collectionCode>
</materialsCitation>
. 
<materialsCitation id="3B613CE20F752D00FBA2FF15FCC13A45" ID-GBIF-Occurrence="2577306112" collectingDate="2018-09-24" collectionCode="TUA, R" collectorName="Aya-cho &amp; Kitamata &amp; Kawanaka Shrine" location="Miyake" pageId="2" pageNumber="379" specimenCount="1" specimenCount-female="1" stateProvince="Miyazaki" typeStatus="holotype">
<location id="8ED660640F752D00FBA2FF15FB003BA1" LSID="urn:lsid:plazi:treatment:03A087A90F762D04FF70F926FE503C97:8ED660640F752D00FBA2FF15FB003BA1" box="[1093,1179,225,251]" name="Miyake" pageId="2" pageNumber="379" stateProvince="Miyazaki">Miyake</location>
(
<specimenCount id="9D0FFD360F752D00FB4BFF15FB4F3BA1" box="[1196,1236,225,251]" pageId="2" pageNumber="379" type="female">1 ♀</specimenCount>
, 
<collectionCode id="ED18AE7A0F752D00FB06FF15FA843BA1" box="[1249,1311,225,251]" pageId="2" pageNumber="379">TUA</collectionCode>
); 
<collectingRegion id="49CDF85D0F752D00FAD5FF15FF4B3A45" country="Japan" name="Miyazaki" pageId="2" pageNumber="379">Miyazaki Pref.</collectingRegion>
, 
<collectorName id="26FC53690F752D00FF3EFEF1FEA13A45" box="[217,314,261,287]" pageId="2" pageNumber="379">Aya-chô</collectorName>
, 
<collectorName id="26FC53690F752D00FEA2FEF1FE343A45" box="[325,431,261,287]" pageId="2" pageNumber="379">Kitamata</collectorName>
, 
<collectorName id="26FC53690F752D00FE5DFEF1FD183A45" box="[442,643,261,287]" pageId="2" pageNumber="379">Kawanaka Shrine</collectorName>
, 
<date id="FFB7107F0F752D00FD69FEF1FC933A45" box="[654,776,261,287]" pageId="2" pageNumber="379" value="2018-09-24">
<collectingDate id="EFF3E9970F752D00FD69FEF1FC933A45" box="[654,776,261,287]" pageId="2" pageNumber="379" value="2018-09-24">24.ix.2018</collectingDate>
</date>
, leg. 
<collectionCode id="ED18AE7A0F752D00FCA3FEF1FCC13A45" box="[836,858,261,287]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="2" pageNumber="379">R</collectionCode>
</materialsCitation>
. 
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<location id="8ED660640F752D00FC82FEF1FC1F3A45" LSID="urn:lsid:plazi:treatment:03A087A90F762D04FF70F926FE503C97:8ED660640F752D00FC82FEF1FC1F3A45" box="[869,900,261,287]" name="Ito" pageId="2" pageNumber="379" stateProvince="Miyazaki">Ito</location>
(
<specimenCount id="9D0FFD360F752D00FC71FEF1FC253A45" box="[918,958,261,287]" pageId="2" pageNumber="379" type="male">1 ♂</specimenCount>
<specimenCount id="9D0FFD360F752D00FC22FEF1FC743A45" box="[965,1007,261,287]" pageId="2" pageNumber="379" type="female">1 ♀</specimenCount>
, 
<collectionCode id="ED18AE7A0F752D00FC1AFEF1FBA73A45" box="[1021,1084,261,287]" pageId="2" pageNumber="379">TUA</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31365340F752D00FF20FEDCFC083AAD" pageId="2" pageNumber="379" type="diagnosis">
<paragraph id="8BB636BF0F752D00FF20FEDCFC083AAD" blockId="2.[151,1437,152,1007]" pageId="2" pageNumber="379">
<emphasis id="B97DEAAD0F752D00FF20FEDCFED93A19" bold="true" box="[199,322,296,323]" pageId="2" pageNumber="379">Diagnosis.</emphasis>
Recognized among other species of 
<taxonomicName id="4C094D3C0F752D00FD3AFEDDFCD63A19" authorityName="Pericart" authorityYear="1984" box="[733,845,297,323]" class="Insecta" family="Tingidae" genus="Heissiella" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="379" phylum="Arthropoda" rank="genus">
<emphasis id="B97DEAAD0F752D00FD3AFEDDFCD63A19" box="[733,845,297,323]" italics="true" pageId="2" pageNumber="379">Heissiella</emphasis>
</taxonomicName>
by a combination of the following characters: dorsal outline of hood nearly straight in lateral view (
<figureCitation id="13322A3A0F752D00FD45FEB9FD613A3D" box="[674,762,333,359]" captionStart="FIGURE 2" captionStartId="3.[151,250,1942,1967]" captionTargetBox="[201,1385,993,1917]" captionTargetId="figure@3.[201,1385,993,1917]" captionTargetPageId="3" captionText="FIGURE 2. Detail morphological images of Heissiella donguri sp. nov. A–C: cephalic and thoracic structures, dorsal, lateral and ventral views; D, E, hemelytron, dorsal and lateral views; F, apical part of abdomen of male, ventral view; G, same of female, ventral view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3713028" httpUri="https://zenodo.org/record/3713028/files/figure.png" pageId="2" pageNumber="379">Fig. 2B</figureCitation>
); outer margin of paranotum resting on pronotal disc except anterior end (
<figureCitation id="13322A3A0F752D00FECCFE85FE1E3AD1" box="[299,389,369,395]" captionStart="FIGURE 2" captionStartId="3.[151,250,1942,1967]" captionTargetBox="[201,1385,993,1917]" captionTargetId="figure@3.[201,1385,993,1917]" captionTargetPageId="3" captionText="FIGURE 2. Detail morphological images of Heissiella donguri sp. nov. A–C: cephalic and thoracic structures, dorsal, lateral and ventral views; D, E, hemelytron, dorsal and lateral views; F, apical part of abdomen of male, ventral view; G, same of female, ventral view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3713028" httpUri="https://zenodo.org/record/3713028/files/figure.png" pageId="2" pageNumber="379">Fig. 2A</figureCitation>
); costal area of hemelytron with 2–3 rows of areolae throughout its length (
<figureCitation id="13322A3A0F752D00FB2FFE85FAB93AD1" box="[1224,1314,369,395]" captionStart="FIGURE 2" captionStartId="3.[151,250,1942,1967]" captionTargetBox="[201,1385,993,1917]" captionTargetId="figure@3.[201,1385,993,1917]" captionTargetPageId="3" captionText="FIGURE 2. Detail morphological images of Heissiella donguri sp. nov. A–C: cephalic and thoracic structures, dorsal, lateral and ventral views; D, E, hemelytron, dorsal and lateral views; F, apical part of abdomen of male, ventral view; G, same of female, ventral view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3713028" httpUri="https://zenodo.org/record/3713028/files/figure.png" pageId="2" pageNumber="379">Fig. 2D</figureCitation>
); hypocostal lamina with 2 rows of areolae in basal half and a single row in apical half (
<figureCitation id="13322A3A0F752D00FBEBFE61FBFF3AF5" box="[1036,1124,405,431]" captionStart="FIGURE 2" captionStartId="3.[151,250,1942,1967]" captionTargetBox="[201,1385,993,1917]" captionTargetId="figure@3.[201,1385,993,1917]" captionTargetPageId="3" captionText="FIGURE 2. Detail morphological images of Heissiella donguri sp. nov. A–C: cephalic and thoracic structures, dorsal, lateral and ventral views; D, E, hemelytron, dorsal and lateral views; F, apical part of abdomen of male, ventral view; G, same of female, ventral view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3713028" httpUri="https://zenodo.org/record/3713028/files/figure.png" pageId="2" pageNumber="379">Fig. 2E</figureCitation>
); endophallus lacking inner sclerite basally (
<figureCitation id="13322A3A0F752D00FEAAFE4DFE5C3A89" box="[333,455,441,467]" captionStart="FIGURE 3" captionStartId="4.[151,250,1494,1519]" captionTargetBox="[190,1402,459,1464]" captionTargetId="figure@4.[177,1410,453,1469]" captionTargetPageId="4" captionText="FIGURE 3. Male and female terminalia of Heissiella donguri sp. nov. A, pygophore, dorsal view; B, female terminalia, ventral view; C, D, phallus, dorsal and ventral views; E, paramere, dorsal view. Abbreviations: st 7, abdominal sternite VII; ov, ovivalvula; gcp 1, gonocoxopodite I; pt 8, paratergite VIII; pt 9, paratergite IX; ol, outer lobe; il, inner lobe; bp, basal plate; cp, capitate process; ph, phallotheca; en, endophallus. Scale bars: A–D, 0.2 mm; E, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3713030" httpUri="https://zenodo.org/record/3713030/files/figure.png" pageId="2" pageNumber="379">Fig. 3C, D</figureCitation>
); outer lobe of paratergite IX in female protruding laterad, shorter than inner lobe (
<figureCitation id="13322A3A0F752D00FA83FE4DFF273AAD" captionStart="FIGURE 2" captionStartId="3.[151,250,1942,1967]" captionTargetBox="[201,1385,993,1917]" captionTargetId="figure@3.[201,1385,993,1917]" captionTargetPageId="3" captionText="FIGURE 2. Detail morphological images of Heissiella donguri sp. nov. A–C: cephalic and thoracic structures, dorsal, lateral and ventral views; D, E, hemelytron, dorsal and lateral views; F, apical part of abdomen of male, ventral view; G, same of female, ventral view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3713028" httpUri="https://zenodo.org/record/3713028/files/figure.png" pageId="2" pageNumber="379">Figs. 2G</figureCitation>
, 
<figureCitation id="13322A3A0F752D00FF20FE29FF703AAD" box="[199,235,477,503]" captionStart="FIGURE 3" captionStartId="4.[151,250,1494,1519]" captionTargetBox="[190,1402,459,1464]" captionTargetId="figure@4.[177,1410,453,1469]" captionTargetPageId="4" captionText="FIGURE 3. Male and female terminalia of Heissiella donguri sp. nov. A, pygophore, dorsal view; B, female terminalia, ventral view; C, D, phallus, dorsal and ventral views; E, paramere, dorsal view. Abbreviations: st 7, abdominal sternite VII; ov, ovivalvula; gcp 1, gonocoxopodite I; pt 8, paratergite VIII; pt 9, paratergite IX; ol, outer lobe; il, inner lobe; bp, basal plate; cp, capitate process; ph, phallotheca; en, endophallus. Scale bars: A–D, 0.2 mm; E, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3713030" httpUri="https://zenodo.org/record/3713030/files/figure.png" pageId="2" pageNumber="379">3B</figureCitation>
); and inner lobe of paratergite IX in female rounded at apex.
</paragraph>
</subSubSection>
<subSubSection id="C31365340F752D06FF20FDF4FD503AF5" lastPageId="4" lastPageNumber="381" pageId="2" pageNumber="379" type="description">
<paragraph id="8BB636BF0F752D00FF20FDF4FF723939" blockId="2.[151,1437,152,1007]" pageId="2" pageNumber="379">
<emphasis id="B97DEAAD0F752D00FF20FDF4FEC23941" bold="true" box="[199,345,512,539]" pageId="2" pageNumber="379">Description.</emphasis>
Macropterous male. General color brown; compound eyes dark red; calli, antennal segment IV except basal part, thorax in ventral surface, coxa and apex of tarsus black; pubescence on body yellowish (
<figureCitation id="13322A3A0F752D00FAD3FDD1FF373939" captionStart="FIGURE 1" captionStartId="2.[151,250,1794,1819]" captionTargetBox="[156,1431,1029,1761]" captionTargetId="figure@2.[151,1436,1026,1766]" captionTargetPageId="2" captionText="FIGURE 1. Habitus images of Heissiella donguri sp. nov. from Japan, dorsal and lateral views. A, B: male from Nara, Honshu; C, D: female from Oita, Kyushu. Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3713026" httpUri="https://zenodo.org/record/3713026/files/figure.png" pageId="2" pageNumber="379">Figs. 1A, B</figureCitation>
, 
<figureCitation id="13322A3A0F752D00FF5FFDBDFF473939" box="[184,220,585,611]" captionStart="FIGURE 2" captionStartId="3.[151,250,1942,1967]" captionTargetBox="[201,1385,993,1917]" captionTargetId="figure@3.[201,1385,993,1917]" captionTargetPageId="3" captionText="FIGURE 2. Detail morphological images of Heissiella donguri sp. nov. A–C: cephalic and thoracic structures, dorsal, lateral and ventral views; D, E, hemelytron, dorsal and lateral views; F, apical part of abdomen of male, ventral view; G, same of female, ventral view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3713028" httpUri="https://zenodo.org/record/3713028/files/figure.png" pageId="2" pageNumber="379">2C</figureCitation>
).
</paragraph>
<paragraph id="8BB636BF0F752D00FF20FD99FDA538B5" blockId="2.[151,1437,152,1007]" pageId="2" pageNumber="379">
Body covered with minute pubescence, 2.3 times as long as maximum width across hemelytra (
<figureCitation id="13322A3A0F752D00FB14FD99FAD739DD" box="[1267,1356,621,647]" captionStart="FIGURE 1" captionStartId="2.[151,250,1794,1819]" captionTargetBox="[156,1431,1029,1761]" captionTargetId="figure@2.[151,1436,1026,1766]" captionTargetPageId="2" captionText="FIGURE 1. Habitus images of Heissiella donguri sp. nov. from Japan, dorsal and lateral views. A, B: male from Nara, Honshu; C, D: female from Oita, Kyushu. Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3713026" httpUri="https://zenodo.org/record/3713026/files/figure.png" pageId="2" pageNumber="379">Fig. 1A</figureCitation>
). Head (
<figureCitation id="13322A3A0F752D00FF47FD65FE8039F1" box="[160,283,657,683]" captionStart="FIGURE 2" captionStartId="3.[151,250,1942,1967]" captionTargetBox="[201,1385,993,1917]" captionTargetId="figure@3.[201,1385,993,1917]" captionTargetPageId="3" captionText="FIGURE 2. Detail morphological images of Heissiella donguri sp. nov. A–C: cephalic and thoracic structures, dorsal, lateral and ventral views; D, E, hemelytron, dorsal and lateral views; F, apical part of abdomen of male, ventral view; G, same of female, ventral view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3713028" httpUri="https://zenodo.org/record/3713028/files/figure.png" pageId="2" pageNumber="379">Fig. 2A, B</figureCitation>
) smooth; a pair of frontal spines touching each other at apices, reaching beyond tip of tylus; median spine shorter than frontal spines, extending beyond bases of frontal spines; a pair of occipital spines slightly reflexed upward, shorter than median spine, reaching middle part of compound eyes; antenniferous tubercles obtuse, slightly curved inward. Compound eyes prominent laterally, round in dorsal view. Antennae (
<figureCitation id="13322A3A0F752D00FB86FD09FB23384D" box="[1121,1208,765,791]" captionStart="FIGURE 1" captionStartId="2.[151,250,1794,1819]" captionTargetBox="[156,1431,1029,1761]" captionTargetId="figure@2.[151,1436,1026,1766]" captionTargetPageId="2" captionText="FIGURE 1. Habitus images of Heissiella donguri sp. nov. from Japan, dorsal and lateral views. A, B: male from Nara, Honshu; C, D: female from Oita, Kyushu. Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3713026" httpUri="https://zenodo.org/record/3713026/files/figure.png" pageId="2" pageNumber="379">Fig. 1A</figureCitation>
) smooth; segment I approximately 1.6 times as long as its width; segment II cone-shaped, 1.4 times as long as its maximum width; segment III longest among antennal segments, 1.8 times as long as maximum width of head across compound eyes; segment IV fusiform, widest a little beyond middle, irregularly covered with long pubescence; ratios of lengths from segments I to IV as 1.1: 1.0: 6.9: 2.9. Bucculae long, approximately 2.6 times as long as their maximum height, with 2 rows of areolae throughout its length. Rostrum (
<figureCitation id="13322A3A0F752D00FD1DFC45FCD43891" box="[762,847,945,971]" captionStart="FIGURE 2" captionStartId="3.[151,250,1942,1967]" captionTargetBox="[201,1385,993,1917]" captionTargetId="figure@3.[201,1385,993,1917]" captionTargetPageId="3" captionText="FIGURE 2. Detail morphological images of Heissiella donguri sp. nov. A–C: cephalic and thoracic structures, dorsal, lateral and ventral views; D, E, hemelytron, dorsal and lateral views; F, apical part of abdomen of male, ventral view; G, same of female, ventral view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3713028" httpUri="https://zenodo.org/record/3713028/files/figure.png" pageId="2" pageNumber="379">Fig. 2C</figureCitation>
) approximately 0.6 times as long as antennae, reaching posterior margin of mesosternum.
</paragraph>
<caption id="DF7666370F752D00FF70F8F6FD303C64" ID-DOI="http://doi.org/10.5281/zenodo.3713026" ID-Zenodo-Dep="3713026" httpUri="https://zenodo.org/record/3713026/files/figure.png" pageId="2" pageNumber="379" startId="2.[151,250,1794,1819]" targetBox="[156,1431,1029,1761]" targetPageId="2">
<paragraph id="8BB636BF0F752D00FF70F8F6FD303C64" blockId="2.[151,1436,1794,1855]" pageId="2" pageNumber="379">
<emphasis id="B97DEAAD0F752D00FF70F8F6FE893C41" bold="true" box="[151,274,1794,1819]" pageId="2" pageNumber="379">FIGURE 1.</emphasis>
Habitus images of 
<taxonomicName id="4C094D3C0F752D00FE33F8F7FD143C41" authority="Souma &amp; Ishikawa, 2020" authorityName="Souma &amp; Ishikawa" authorityYear="2020" box="[468,655,1795,1819]" class="Insecta" family="Tingidae" genus="Heissiella" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="379" phylum="Arthropoda" rank="species" species="donguri" status="sp. nov.">
<emphasis id="B97DEAAD0F752D00FE33F8F7FD143C41" box="[468,655,1795,1819]" italics="true" pageId="2" pageNumber="379">Heissiella donguri</emphasis>
</taxonomicName>
<emphasis id="B97DEAAD0F752D00FD72F8F7FD7E3C41" bold="true" box="[661,741,1795,1819]" pageId="2" pageNumber="379">
<taxonomicNameLabel id="A24E57D60F752D00FD72F8F7FD7E3C41" box="[661,741,1795,1819]" pageId="2" pageNumber="379" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
from Japan, dorsal and lateral views. A, B: male from Nara, Honshu; C, D: female from Oita, Kyushu. Scale bar: 1.0 mm.
</paragraph>
</caption>
<paragraph id="8BB636BF0F752D01FF20F89AFBFE3A45" blockId="2.[151,1437,1901,2036]" lastBlockId="3.[151,1437,153,971]" lastPageId="3" lastPageNumber="380" pageId="2" pageNumber="379">
Pronotum (
<figureCitation id="13322A3A0F752D00FEA1F899FE133CDD" box="[326,392,1901,1927]" captionStart="FIGURE 2" captionStartId="3.[151,250,1942,1967]" captionTargetBox="[201,1385,993,1917]" captionTargetId="figure@3.[201,1385,993,1917]" captionTargetPageId="3" captionText="FIGURE 2. Detail morphological images of Heissiella donguri sp. nov. A–C: cephalic and thoracic structures, dorsal, lateral and ventral views; D, E, hemelytron, dorsal and lateral views; F, apical part of abdomen of male, ventral view; G, same of female, ventral view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3713028" httpUri="https://zenodo.org/record/3713028/files/figure.png" pageId="2" pageNumber="379">Fig. 2</figureCitation>
A–C) 1.4 times as long as maximum width across paranota. Pronotal disc coarsely punctate. Hood obtusely protruded forward, concealing basal fourth of vertex; dorsal outline nearly straight in lateral view. Pronotal carinae with a single row of minute areolae throughout their length; median carina straight, extending to apex of posterior process, higher than hood at maximum height; lateral carinae nearly parallel to each other except anterior end, not concealed by paranota throughout their length, as high as median carina. Calli coarsely punctate. Paranotum widened posteriorly, with 3 rows of areolae at widest part, approximately 4 times as long as its maximum width, slightly covering pronotal disc in posterior part, not touching lateral carinae; outer margin gently curved inward throughout its length. Posterior process triangular, as long as its maximum width.
</paragraph>
<paragraph id="8BB636BF0F742D01FF20FEDDFD433965" blockId="3.[151,1437,153,971]" pageId="3" pageNumber="380">
Hemelytron (
<figureCitation id="13322A3A0F742D01FEBBFEDDFE493A19" box="[348,466,297,323]" captionStart="FIGURE 2" captionStartId="3.[151,250,1942,1967]" captionTargetBox="[201,1385,993,1917]" captionTargetId="figure@3.[201,1385,993,1917]" captionTargetPageId="3" captionText="FIGURE 2. Detail morphological images of Heissiella donguri sp. nov. A–C: cephalic and thoracic structures, dorsal, lateral and ventral views; D, E, hemelytron, dorsal and lateral views; F, apical part of abdomen of male, ventral view; G, same of female, ventral view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3713028" httpUri="https://zenodo.org/record/3713028/files/figure.png" pageId="3" pageNumber="380">Fig. 2D, E</figureCitation>
) narrow, 2.9 times as long as its maximum width, considerably extending beyond apex of abdomen; maximum width across hemelytra 1.2 times as much as maximum width across paranota; clavus with 4 rows of areolae a widest part; costal area wider than subcostal area at widest part, slightly reflexed upward in basal part, with 2–3 rows of areolae throughout its length; subcostal area 0.2 times as wide as discoidal area at middle of hemelytron, with 2 rows of areolae throughout its length; discoidal area distinctly expanding beyond middle of hemelytron, with 10–11 rows of areolae at widest part; sutural area well-developed, with 11–12 rows of areolae at widest part; hypocostal lamina 0.8 times as high as width of costal area at middle of hemelytron, with 2 rows of areolae in basal half and a single row in apical half.
</paragraph>
<paragraph id="8BB636BF0F742D01FF20FDBDFD173995" blockId="3.[151,1437,153,971]" pageId="3" pageNumber="380">
Thoracic pleura coarsely and evenly punctate (
<figureCitation id="13322A3A0F742D01FD30FDBDFCAB3939" box="[727,816,585,611]" captionStart="FIGURE 2" captionStartId="3.[151,250,1942,1967]" captionTargetBox="[201,1385,993,1917]" captionTargetId="figure@3.[201,1385,993,1917]" captionTargetPageId="3" captionText="FIGURE 2. Detail morphological images of Heissiella donguri sp. nov. A–C: cephalic and thoracic structures, dorsal, lateral and ventral views; D, E, hemelytron, dorsal and lateral views; F, apical part of abdomen of male, ventral view; G, same of female, ventral view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3713028" httpUri="https://zenodo.org/record/3713028/files/figure.png" pageId="3" pageNumber="380">Fig. 2B</figureCitation>
). Sternal laminae (
<figureCitation id="13322A3A0F742D01FBE1FDBDFBC73939" box="[1030,1116,585,611]" captionStart="FIGURE 2" captionStartId="3.[151,250,1942,1967]" captionTargetBox="[201,1385,993,1917]" captionTargetId="figure@3.[201,1385,993,1917]" captionTargetPageId="3" captionText="FIGURE 2. Detail morphological images of Heissiella donguri sp. nov. A–C: cephalic and thoracic structures, dorsal, lateral and ventral views; D, E, hemelytron, dorsal and lateral views; F, apical part of abdomen of male, ventral view; G, same of female, ventral view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3713028" httpUri="https://zenodo.org/record/3713028/files/figure.png" pageId="3" pageNumber="380">Fig. 2C</figureCitation>
) apparently lower than bucculae; pro- and mesosternal laminae nearly parallel to each other, open in both anterior and posterior ends; metasternal laminae as high as mesosternal laminae, open at anterior end, angularly curved inward at posterior end. Legs smooth; femora thickest at middle (
<figureCitation id="13322A3A0F742D01FDC2FD41FDE53995" box="[549,638,693,719]" captionStart="FIGURE 1" captionStartId="2.[151,250,1794,1819]" captionTargetBox="[156,1431,1029,1761]" captionTargetId="figure@2.[151,1436,1026,1766]" captionTargetPageId="2" captionText="FIGURE 1. Habitus images of Heissiella donguri sp. nov. from Japan, dorsal and lateral views. A, B: male from Nara, Honshu; C, D: female from Oita, Kyushu. Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3713026" httpUri="https://zenodo.org/record/3713026/files/figure.png" pageId="3" pageNumber="380">Fig. 1A</figureCitation>
).
</paragraph>
<paragraph id="8BB636BF0F742D01FF20FD2DFC713891" blockId="3.[151,1437,153,971]" pageId="3" pageNumber="380">
Abdomen ellipsoidal, 1.5 times as long as its maximum width. Pygophore (
<figureCitation id="13322A3A0F742D01FBC0FD2DFB1D39A9" box="[1063,1158,729,755]" captionStart="FIGURE 2" captionStartId="3.[151,250,1942,1967]" captionTargetBox="[201,1385,993,1917]" captionTargetId="figure@3.[201,1385,993,1917]" captionTargetPageId="3" captionText="FIGURE 2. Detail morphological images of Heissiella donguri sp. nov. A–C: cephalic and thoracic structures, dorsal, lateral and ventral views; D, E, hemelytron, dorsal and lateral views; F, apical part of abdomen of male, ventral view; G, same of female, ventral view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3713028" httpUri="https://zenodo.org/record/3713028/files/figure.png" pageId="3" pageNumber="380">Figs. 2F</figureCitation>
, 
<figureCitation id="13322A3A0F742D01FB74FD2DFB2C39A9" box="[1171,1207,729,755]" captionStart="FIGURE 3" captionStartId="4.[151,250,1494,1519]" captionTargetBox="[190,1402,459,1464]" captionTargetId="figure@4.[177,1410,453,1469]" captionTargetPageId="4" captionText="FIGURE 3. Male and female terminalia of Heissiella donguri sp. nov. A, pygophore, dorsal view; B, female terminalia, ventral view; C, D, phallus, dorsal and ventral views; E, paramere, dorsal view. Abbreviations: st 7, abdominal sternite VII; ov, ovivalvula; gcp 1, gonocoxopodite I; pt 8, paratergite VIII; pt 9, paratergite IX; ol, outer lobe; il, inner lobe; bp, basal plate; cp, capitate process; ph, phallotheca; en, endophallus. Scale bars: A–D, 0.2 mm; E, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3713030" httpUri="https://zenodo.org/record/3713030/files/figure.png" pageId="3" pageNumber="380">3A</figureCitation>
) compressed dorsoventrally, semicircular in ventral view, weakly concave at anterior margin of dorsum, slightly elevated at center of venter, smooth, irregularly punctate in middle part of dorsum. Parameres (
<figureCitation id="13322A3A0F742D01FBE3FCD5FBE63861" box="[1028,1149,801,827]" captionStart="FIGURE 3" captionStartId="4.[151,250,1494,1519]" captionTargetBox="[190,1402,459,1464]" captionTargetId="figure@4.[177,1410,453,1469]" captionTargetPageId="4" captionText="FIGURE 3. Male and female terminalia of Heissiella donguri sp. nov. A, pygophore, dorsal view; B, female terminalia, ventral view; C, D, phallus, dorsal and ventral views; E, paramere, dorsal view. Abbreviations: st 7, abdominal sternite VII; ov, ovivalvula; gcp 1, gonocoxopodite I; pt 8, paratergite VIII; pt 9, paratergite IX; ol, outer lobe; il, inner lobe; bp, basal plate; cp, capitate process; ph, phallotheca; en, endophallus. Scale bars: A–D, 0.2 mm; E, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3713030" httpUri="https://zenodo.org/record/3713030/files/figure.png" pageId="3" pageNumber="380">Fig. 3A, E</figureCitation>
) thick and long, strongly expanded in middle part, strongly curved inward in apical part; outer and inner margins covered with pubescence in middle part; suspensory arms of parameres completely visible in dorsal view. Phallus (
<figureCitation id="13322A3A0F742D01FB99FC9DFB6D38D9" box="[1150,1270,873,899]" captionStart="FIGURE 3" captionStartId="4.[151,250,1494,1519]" captionTargetBox="[190,1402,459,1464]" captionTargetId="figure@4.[177,1410,453,1469]" captionTargetPageId="4" captionText="FIGURE 3. Male and female terminalia of Heissiella donguri sp. nov. A, pygophore, dorsal view; B, female terminalia, ventral view; C, D, phallus, dorsal and ventral views; E, paramere, dorsal view. Abbreviations: st 7, abdominal sternite VII; ov, ovivalvula; gcp 1, gonocoxopodite I; pt 8, paratergite VIII; pt 9, paratergite IX; ol, outer lobe; il, inner lobe; bp, basal plate; cp, capitate process; ph, phallotheca; en, endophallus. Scale bars: A–D, 0.2 mm; E, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3713030" httpUri="https://zenodo.org/record/3713030/files/figure.png" pageId="3" pageNumber="380">Fig. 3C, D</figureCitation>
) oblong; basal plate and capitate process distinctly sclerotized; phallotheca greatly membranous, with a dorsal sclerite; endophallus membranous except sclerites in its apical part, lacking inner sclerite basally.
</paragraph>
<caption id="DF7666370F742D01FF70F862FDBC3CAC" ID-DOI="http://doi.org/10.5281/zenodo.3713028" ID-Zenodo-Dep="3713028" httpUri="https://zenodo.org/record/3713028/files/figure.png" pageId="3" pageNumber="380" startId="3.[151,250,1942,1967]" targetBox="[201,1385,993,1917]" targetPageId="3">
<paragraph id="8BB636BF0F742D01FF70F862FDBC3CAC" blockId="3.[151,1436,1942,2039]" pageId="3" pageNumber="380">
<emphasis id="B97DEAAD0F742D01FF70F862FE8F3CF5" bold="true" box="[151,276,1942,1967]" pageId="3" pageNumber="380">FIGURE 2.</emphasis>
Detail morphological images of 
<taxonomicName id="4C094D3C0F742D01FD80F863FCB83CF5" authority="Souma &amp; Ishikawa, 2020" authorityName="Souma &amp; Ishikawa" authorityYear="2020" box="[615,803,1943,1967]" class="Insecta" family="Tingidae" genus="Heissiella" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="380" phylum="Arthropoda" rank="species" species="donguri" status="sp. nov.">
<emphasis id="B97DEAAD0F742D01FD80F863FCB83CF5" box="[615,803,1943,1967]" italics="true" pageId="3" pageNumber="380">Heissiella donguri</emphasis>
</taxonomicName>
<emphasis id="B97DEAAD0F742D01FCCCF863FCE73CF5" bold="true" box="[811,892,1943,1967]" pageId="3" pageNumber="380">
<taxonomicNameLabel id="A24E57D60F742D01FCCCF863FCE73CF5" box="[811,892,1943,1967]" pageId="3" pageNumber="380" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A–C: cephalic and thoracic structures, dorsal, lateral and ventral views; D, E, hemelytron, dorsal and lateral views; F, apical part of abdomen of male, ventral view; G, same of female, ventral view. Scale bars: 0.2 mm.
</paragraph>
</caption>
<paragraph id="8BB636BF0F732D06FF20FF6DFDCF3B8D" blockId="4.[151,1437,153,431]" pageId="4" pageNumber="381">
Measurements (
<typeStatus id="54B2881D0F732D06FE9BFF6DFE7E3BEE" box="[380,485,153,180]" pageId="4" pageNumber="381" type="holotype">holotype</typeStatus>
). Body length with hemelytra 
<quantity id="4CF19B5A0F732D06FCA2FF6DFC3B3BE9" box="[837,928,153,179]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="4" pageNumber="381" unit="mm" value="3.6">3.6 mm</quantity>
; maximum width across hemelytra 
<quantity id="4CF19B5A0F732D06FADBFF6DFA033BE9" box="[1340,1432,153,179]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="4" pageNumber="381" unit="mm" value="1.6">1.6 mm</quantity>
; pronotal width across paranota 
<quantity id="4CF19B5A0F732D06FE10FF49FDCB3B8D" box="[503,592,189,215]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="4" pageNumber="381" unit="mm" value="1.3">1.3 mm</quantity>
.
</paragraph>
<paragraph id="8BB636BF0F732D06FF20FF15FE743A19" blockId="4.[151,1437,153,431]" pageId="4" pageNumber="381">
Macropterous female. General appearance very similar to that of male (
<figureCitation id="13322A3A0F732D06FC1CFF15FBEC3BA1" box="[1019,1143,225,251]" captionStart="FIGURE 1" captionStartId="2.[151,250,1794,1819]" captionTargetBox="[156,1431,1029,1761]" captionTargetId="figure@2.[151,1436,1026,1766]" captionTargetPageId="2" captionText="FIGURE 1. Habitus images of Heissiella donguri sp. nov. from Japan, dorsal and lateral views. A, B: male from Nara, Honshu; C, D: female from Oita, Kyushu. Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3713026" httpUri="https://zenodo.org/record/3713026/files/figure.png" pageId="4" pageNumber="381">Fig. 1C, D</figureCitation>
) except for the following characters: outer lobe of paratergite IX protruding laterad, shorter than inner lobe; inner lobe of paratergite IX rounded at apex (
<figureCitation id="13322A3A0F732D06FEBEFEDDFE293A19" box="[345,434,297,323]" captionStart="FIGURE 2" captionStartId="3.[151,250,1942,1967]" captionTargetBox="[201,1385,993,1917]" captionTargetId="figure@3.[201,1385,993,1917]" captionTargetPageId="3" captionText="FIGURE 2. Detail morphological images of Heissiella donguri sp. nov. A–C: cephalic and thoracic structures, dorsal, lateral and ventral views; D, E, hemelytron, dorsal and lateral views; F, apical part of abdomen of male, ventral view; G, same of female, ventral view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3713028" httpUri="https://zenodo.org/record/3713028/files/figure.png" pageId="4" pageNumber="381">Fig. 2G</figureCitation>
, 
<figureCitation id="13322A3A0F732D06FE59FEDDFE7A3A19" box="[446,481,297,323]" captionStart="FIGURE 3" captionStartId="4.[151,250,1494,1519]" captionTargetBox="[190,1402,459,1464]" captionTargetId="figure@4.[177,1410,453,1469]" captionTargetPageId="4" captionText="FIGURE 3. Male and female terminalia of Heissiella donguri sp. nov. A, pygophore, dorsal view; B, female terminalia, ventral view; C, D, phallus, dorsal and ventral views; E, paramere, dorsal view. Abbreviations: st 7, abdominal sternite VII; ov, ovivalvula; gcp 1, gonocoxopodite I; pt 8, paratergite VIII; pt 9, paratergite IX; ol, outer lobe; il, inner lobe; bp, basal plate; cp, capitate process; ph, phallotheca; en, endophallus. Scale bars: A–D, 0.2 mm; E, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3713030" httpUri="https://zenodo.org/record/3713030/files/figure.png" pageId="4" pageNumber="381">3B</figureCitation>
).
</paragraph>
<paragraph id="8BB636BF0F732D06FF20FEB9FB063AD1" blockId="4.[151,1437,153,431]" pageId="4" pageNumber="381">
Infraspecific variation among both sexes (
<typeStatus id="54B2881D0F732D06FD7FFEB9FD673A3D" box="[664,764,333,359]" pageId="4" pageNumber="381" type="holotype">holotype</typeStatus>
and 
<specimenCount id="9D0FFD360F732D06FCD7FEB9FC583A3D" box="[816,963,333,359]" pageId="4" pageNumber="381" type="generic" typeStatus="paratypes">68 paratypes</specimenCount>
). Body length with hemelytra 
<quantity id="4CF19B5A0F732D06FAE8FEB9FA033A3D" box="[1295,1432,333,359]" metricMagnitude="-3" metricUnit="m" metricValue="3.45" metricValueMax="3.6" metricValueMin="3.3" pageId="4" pageNumber="381" unit="mm" value="3.45" valueMax="3.6" valueMin="3.3">3.3–3.6 mm</quantity>
; maximum width across hemelytra 
<quantity id="4CF19B5A0F732D06FDFDFE85FD3F3AD1" box="[538,676,369,395]" metricMagnitude="-3" metricUnit="m" metricValue="1.5499999999999998" metricValueMax="1.7" metricValueMin="1.4" pageId="4" pageNumber="381" unit="mm" value="1.5499999999999998" valueMax="1.7" valueMin="1.4">1.4–1.7 mm</quantity>
; pronotal width across paranota 
<quantity id="4CF19B5A0F732D06FBE8FE85FB023AD1" box="[1039,1177,369,395]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.4" metricValueMin="1.1" pageId="4" pageNumber="381" unit="mm" value="1.25" valueMax="1.4" valueMin="1.1">1.1–1.4 mm</quantity>
.
</paragraph>
<paragraph id="8BB636BF0F732D06FF20FE61FD503AF5" blockId="4.[151,1437,153,431]" box="[199,715,405,431]" pageId="4" pageNumber="381">Brachypterous morph unknown in both sexes.</paragraph>
</subSubSection>
<caption id="DF7666370F732D06FF70FA22FC5A3D00" ID-DOI="http://doi.org/10.5281/zenodo.3713030" ID-Zenodo-Dep="3713030" httpUri="https://zenodo.org/record/3713030/files/figure.png" pageId="4" pageNumber="381" startId="4.[151,250,1494,1519]" targetBox="[190,1402,459,1464]" targetPageId="4">
<paragraph id="8BB636BF0F732D06FF70FA22FC5A3D00" blockId="4.[151,1436,1494,1627]" pageId="4" pageNumber="381">
<emphasis id="B97DEAAD0F732D06FF70FA22FE893EB5" bold="true" box="[151,274,1494,1519]" pageId="4" pageNumber="381">FIGURE 3.</emphasis>
Male and female terminalia of 
<taxonomicName id="4C094D3C0F732D06FDA9FA22FC923EB4" authority="Souma &amp; Ishikawa, 2020" authorityName="Souma &amp; Ishikawa" authorityYear="2020" box="[590,777,1494,1518]" class="Insecta" family="Tingidae" genus="Heissiella" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="381" phylum="Arthropoda" rank="species" species="donguri" status="sp. nov.">
<emphasis id="B97DEAAD0F732D06FDA9FA22FC923EB4" box="[590,777,1494,1518]" italics="true" pageId="4" pageNumber="381">Heissiella donguri</emphasis>
</taxonomicName>
<emphasis id="B97DEAAD0F732D06FCE8FA23FCC43EB5" bold="true" box="[783,863,1495,1519]" pageId="4" pageNumber="381">
<taxonomicNameLabel id="A24E57D60F732D06FCE8FA23FCC43EB5" box="[783,863,1495,1519]" pageId="4" pageNumber="381" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A, pygophore, dorsal view; B, female terminalia, ventral view; C, D, phallus, dorsal and ventral views; E, paramere, dorsal view. Abbreviations: st 7, abdominal sternite VII; ov, ovivalvula; gcp 1, gonocoxopodite I; pt 8, paratergite VIII; pt 9, paratergite IX; ol, outer lobe; il, inner lobe; bp, basal plate; cp, capitate process; ph, phallotheca; en, endophallus. Scale bars: A–D, 0.2 mm; E, 0.1 mm.
</paragraph>
</caption>
<subSubSection id="C31365340F732D07FF20F97DFA0639F1" lastPageId="5" lastPageNumber="382" pageId="4" pageNumber="381" type="discussion">
<paragraph id="8BB636BF0F732D06FF20F97DFD923C55" blockId="4.[151,1436,1673,2023]" pageId="4" pageNumber="381">
<emphasis id="B97DEAAD0F732D06FF20F97DFEA03DF9" bold="true" box="[199,315,1673,1699]" pageId="4" pageNumber="381">Remarks.</emphasis>
<taxonomicName id="4C094D3C0F732D06FEA6F97DFE2A3DF9" authorityName="Pericart" authorityYear="1984" box="[321,433,1673,1699]" class="Insecta" family="Tingidae" genus="Heissiella" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="381" phylum="Arthropoda" rank="genus">
<emphasis id="B97DEAAD0F732D06FEA6F97DFE2A3DF9" box="[321,433,1673,1699]" italics="true" pageId="4" pageNumber="381">Heissiella</emphasis>
</taxonomicName>
species were diagnosed by previous authors based on the differences of different morphological characters, such as shape of the pronotum, arrangement of areolae of the hemelytron, structure of the phallus, and shape of the female paratergite IX (
<bibRefCitation id="EF984B4E0F732D06FDB8F925FD673DB1" author="Pericart, J." box="[607,764,1745,1771]" pageId="4" pageNumber="381" pagination="23 - 47" refId="ref5423" refString="Pericart, J. (1984) Tingidae nouveaux ou interessants du Nord du sous-continent indien (Hemiptera). I. Entomologica Basiliensia, 9, 23 - 47. [in French, English summary]" type="journal article" year="1984">Péricart 1984</bibRefCitation>
; 
<bibRefCitation id="EF984B4E0F732D06FCEEF925FC4F3DB1" author="Dang, K. &amp; Bu, W." box="[777,980,1745,1771]" pageId="4" pageNumber="381" pagination="4" refId="ref4837" refString="Dang, K. &amp; Bu, W. (2012) The discovery of the genus Heissiella Pericart, 1984 (Hemiptera: Heteroptera: Tingidae) from China, with description of a new species. Entomotaxonomia, 34 (4), 654 - 628." type="journal article" year="2012">Dang &amp; Bu 2012</bibRefCitation>
). These characters are also useful in the identification of the new species.
</paragraph>
<paragraph id="8BB636BF0F732D06FF20F8EDFA8C3C99" blockId="4.[151,1436,1673,2023]" pageId="4" pageNumber="381">
The structure of the hypocostal lamina of hemelytron being formed by 2 rows of areolae in basal half and a single row in apical half is a feature in which the new species differs from the diagnosis of 
<taxonomicName id="4C094D3C0F732D06FB75F8C9FA993C0D" authorityName="Pericart" authorityYear="1984" box="[1170,1282,1853,1879]" class="Insecta" family="Tingidae" genus="Heissiella" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="381" phylum="Arthropoda" rank="genus">
<emphasis id="B97DEAAD0F732D06FB75F8C9FA993C0D" box="[1170,1282,1853,1879]" italics="true" pageId="4" pageNumber="381">Heissiella</emphasis>
</taxonomicName>
and all members of the closely related genus 
<taxonomicName id="4C094D3C0F732D06FDE7F895FD0C3C21" authorityName="Horvath" authorityYear="1926" box="[512,663,1889,1915]" class="Insecta" family="Tingidae" genus="Trachypeplus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="381" phylum="Arthropoda" rank="genus">
<emphasis id="B97DEAAD0F732D06FDE7F895FD0C3C21" box="[512,663,1889,1915]" italics="true" pageId="4" pageNumber="381">Trachypeplus</emphasis>
</taxonomicName>
(
<bibRefCitation id="EF984B4E0F732D06FD42F895FCFC3C21" author="Dang, K. &amp; Bu, W." box="[677,871,1889,1915]" pageId="4" pageNumber="381" pagination="4" refId="ref4837" refString="Dang, K. &amp; Bu, W. (2012) The discovery of the genus Heissiella Pericart, 1984 (Hemiptera: Heteroptera: Tingidae) from China, with description of a new species. Entomotaxonomia, 34 (4), 654 - 628." type="journal article" year="2012">Dang &amp; Bu 2012</bibRefCitation>
; 
<bibRefCitation id="EF984B4E0F732D06FC96F895FBB63C21" author="Dang, K. &amp; Guilbert, E. &amp; Bu, W." box="[881,1069,1889,1915]" pageId="4" pageNumber="381" pagination="531 - 550" refId="ref4886" refString="Dang, K., Guilbert, E. &amp; Bu, W. (2013) New species and records of the genus Trachypeplus Horvath (Hemiptera: Tingidae) from China. Zootaxa, 3669 (4), 531 - 550. https: // doi. org / 10.11646 / zootaxa. 3669.4.7" type="journal article" year="2013">
Dang 
<emphasis id="B97DEAAD0F732D06FC53F896FC7F3C21" box="[948,996,1889,1915]" italics="true" pageId="4" pageNumber="381">et al</emphasis>
. 2013
</bibRefCitation>
), allowing an easy recognition of 
<taxonomicName id="4C094D3C0F732D06FF70F872FE883CC5" authority="Souma &amp; Ishikawa, 2020" authorityName="Souma &amp; Ishikawa" authorityYear="2020" box="[151,275,1925,1951]" class="Insecta" family="Tingidae" genus="Heissiella" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="381" phylum="Arthropoda" rank="species" species="donguri" status="sp. nov.">
<emphasis id="B97DEAAD0F732D06FF70F872FF303CC5" box="[151,171,1926,1951]" italics="true" pageId="4" pageNumber="381">H</emphasis>
. 
<emphasis id="B97DEAAD0F732D06FF5DF871FE883CC5" box="[186,275,1925,1951]" italics="true" pageId="4" pageNumber="381">donguri</emphasis>
</taxonomicName>
<emphasis id="B97DEAAD0F732D06FEFDF871FEEF3CC5" bold="true" box="[282,372,1925,1951]" pageId="4" pageNumber="381">
<taxonomicNameLabel id="A24E57D60F732D06FEFDF871FEEF3CC5" box="[282,372,1925,1951]" pageId="4" pageNumber="381" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
However, the new species can be provisionally placed into 
<taxonomicName id="4C094D3C0F732D06FBFFF871FB133CC5" authorityName="Pericart" authorityYear="1984" box="[1048,1160,1925,1951]" class="Insecta" family="Tingidae" genus="Heissiella" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="381" phylum="Arthropoda" rank="genus">
<emphasis id="B97DEAAD0F732D06FBFFF871FB133CC5" box="[1048,1160,1925,1951]" italics="true" pageId="4" pageNumber="381">Heissiella</emphasis>
</taxonomicName>
based on the paranotum with a longitudinal ridge dividing it into dorsal and ventral parts and the bilobed female paratergite IX.
</paragraph>
<paragraph id="8BB636BF0F732D07FF20F839FF743A19" blockId="4.[151,1436,1673,2023]" lastBlockId="5.[151,1437,153,1151]" lastPageId="5" lastPageNumber="382" pageId="4" pageNumber="381">
In the key to all known species of 
<taxonomicName id="4C094D3C0F732D06FDDAF839FC1F3CBD" authority="(Dang &amp; Bu 2012)" baseAuthorityName="Dang &amp; Bu" baseAuthorityYear="2012" box="[573,900,1997,2023]" class="Insecta" family="Tingidae" genus="Heissiella" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="381" phylum="Arthropoda" rank="genus">
<emphasis id="B97DEAAD0F732D06FDDAF839FD363CBD" box="[573,685,1997,2023]" italics="true" pageId="4" pageNumber="381">Heissiella</emphasis>
(
<bibRefCitation id="EF984B4E0F732D06FD5DF839FCE73CBD" author="Dang, K. &amp; Bu, W." box="[698,892,1997,2023]" pageId="4" pageNumber="381" pagination="4" refId="ref4837" refString="Dang, K. &amp; Bu, W. (2012) The discovery of the genus Heissiella Pericart, 1984 (Hemiptera: Heteroptera: Tingidae) from China, with description of a new species. Entomotaxonomia, 34 (4), 654 - 628." type="journal article" year="2012">Dang &amp; Bu 2012</bibRefCitation>
)
</taxonomicName>
, this new species does not run to any species due to having a combination of the following characters: dorsal outline of hood nearly straight in lateral view (
<figureCitation id="13322A3A0F722D07FAD0FF6DFA153BE9" box="[1335,1422,153,179]" captionStart="FIGURE 2" captionStartId="3.[151,250,1942,1967]" captionTargetBox="[201,1385,993,1917]" captionTargetId="figure@3.[201,1385,993,1917]" captionTargetPageId="3" captionText="FIGURE 2. Detail morphological images of Heissiella donguri sp. nov. A–C: cephalic and thoracic structures, dorsal, lateral and ventral views; D, E, hemelytron, dorsal and lateral views; F, apical part of abdomen of male, ventral view; G, same of female, ventral view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3713028" httpUri="https://zenodo.org/record/3713028/files/figure.png" pageId="5" pageNumber="382">Fig. 2B</figureCitation>
); outer margin of paranotum resting on pronotal disc except anterior end (
<figureCitation id="13322A3A0F722D07FC56FF49FB913B8D" box="[945,1034,189,215]" captionStart="FIGURE 2" captionStartId="3.[151,250,1942,1967]" captionTargetBox="[201,1385,993,1917]" captionTargetId="figure@3.[201,1385,993,1917]" captionTargetPageId="3" captionText="FIGURE 2. Detail morphological images of Heissiella donguri sp. nov. A–C: cephalic and thoracic structures, dorsal, lateral and ventral views; D, E, hemelytron, dorsal and lateral views; F, apical part of abdomen of male, ventral view; G, same of female, ventral view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3713028" httpUri="https://zenodo.org/record/3713028/files/figure.png" pageId="5" pageNumber="382">Fig. 2A</figureCitation>
); costal area of hemelytron with 2–3 rows of areolae throughout its length (
<figureCitation id="13322A3A0F722D07FDDAFF15FD0D3BA1" box="[573,662,225,251]" captionStart="FIGURE 2" captionStartId="3.[151,250,1942,1967]" captionTargetBox="[201,1385,993,1917]" captionTargetId="figure@3.[201,1385,993,1917]" captionTargetPageId="3" captionText="FIGURE 2. Detail morphological images of Heissiella donguri sp. nov. A–C: cephalic and thoracic structures, dorsal, lateral and ventral views; D, E, hemelytron, dorsal and lateral views; F, apical part of abdomen of male, ventral view; G, same of female, ventral view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3713028" httpUri="https://zenodo.org/record/3713028/files/figure.png" pageId="5" pageNumber="382">Fig. 2D</figureCitation>
); endophallus lacking inner sclerite basally (
<figureCitation id="13322A3A0F722D07FB64FF15FB603BA1" box="[1155,1275,225,251]" captionStart="FIGURE 3" captionStartId="4.[151,250,1494,1519]" captionTargetBox="[190,1402,459,1464]" captionTargetId="figure@4.[177,1410,453,1469]" captionTargetPageId="4" captionText="FIGURE 3. Male and female terminalia of Heissiella donguri sp. nov. A, pygophore, dorsal view; B, female terminalia, ventral view; C, D, phallus, dorsal and ventral views; E, paramere, dorsal view. Abbreviations: st 7, abdominal sternite VII; ov, ovivalvula; gcp 1, gonocoxopodite I; pt 8, paratergite VIII; pt 9, paratergite IX; ol, outer lobe; il, inner lobe; bp, basal plate; cp, capitate process; ph, phallotheca; en, endophallus. Scale bars: A–D, 0.2 mm; E, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3713030" httpUri="https://zenodo.org/record/3713030/files/figure.png" pageId="5" pageNumber="382">Fig. 3C, D</figureCitation>
); outer lobe of paratergite IX in female shorter than inner lobe (
<figureCitation id="13322A3A0F722D07FD5DFEF1FC853A45" box="[698,798,261,287]" captionStart="FIGURE 2" captionStartId="3.[151,250,1942,1967]" captionTargetBox="[201,1385,993,1917]" captionTargetId="figure@3.[201,1385,993,1917]" captionTargetPageId="3" captionText="FIGURE 2. Detail morphological images of Heissiella donguri sp. nov. A–C: cephalic and thoracic structures, dorsal, lateral and ventral views; D, E, hemelytron, dorsal and lateral views; F, apical part of abdomen of male, ventral view; G, same of female, ventral view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3713028" httpUri="https://zenodo.org/record/3713028/files/figure.png" pageId="5" pageNumber="382">Figs. 2G</figureCitation>
, 
<figureCitation id="13322A3A0F722D07FCCCFEF1FCD53A45" box="[811,846,261,287]" captionStart="FIGURE 3" captionStartId="4.[151,250,1494,1519]" captionTargetBox="[190,1402,459,1464]" captionTargetId="figure@4.[177,1410,453,1469]" captionTargetPageId="4" captionText="FIGURE 3. Male and female terminalia of Heissiella donguri sp. nov. A, pygophore, dorsal view; B, female terminalia, ventral view; C, D, phallus, dorsal and ventral views; E, paramere, dorsal view. Abbreviations: st 7, abdominal sternite VII; ov, ovivalvula; gcp 1, gonocoxopodite I; pt 8, paratergite VIII; pt 9, paratergite IX; ol, outer lobe; il, inner lobe; bp, basal plate; cp, capitate process; ph, phallotheca; en, endophallus. Scale bars: A–D, 0.2 mm; E, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3713030" httpUri="https://zenodo.org/record/3713030/files/figure.png" pageId="5" pageNumber="382">3B</figureCitation>
); and inner lobe of paratergite IX in female rounded at apex.
</paragraph>
<paragraph id="8BB636BF0F722D07FF20FEB9FD0E3939" blockId="5.[151,1437,153,1151]" pageId="5" pageNumber="382">
<taxonomicName id="4C094D3C0F722D07FF20FEB9FE0C3A3D" authority="Souma &amp; Ishikawa, 2020" authorityName="Souma &amp; Ishikawa" authorityYear="2020" box="[199,407,333,359]" class="Insecta" family="Tingidae" genus="Heissiella" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="382" phylum="Arthropoda" rank="species" species="donguri" status="sp. nov.">
<emphasis id="B97DEAAD0F722D07FF20FEB9FE0C3A3D" box="[199,407,333,359]" italics="true" pageId="5" pageNumber="382">Heissiella donguri</emphasis>
</taxonomicName>
<emphasis id="B97DEAAD0F722D07FE79FEB9FE633A3D" bold="true" box="[414,504,333,359]" pageId="5" pageNumber="382">
<taxonomicNameLabel id="A24E57D60F722D07FE79FEB9FE633A3D" box="[414,504,333,359]" pageId="5" pageNumber="382" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
resembles 
<taxonomicName id="4C094D3C0F722D07FD91FEBAFD763A3D" baseAuthorityName="Drake &amp; Poor" baseAuthorityYear="1936" box="[630,749,333,359]" class="Insecta" family="Tingidae" genus="Physatocheila" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="382" phylum="Arthropoda" rank="species" species="dryadis">
<emphasis id="B97DEAAD0F722D07FD91FEBAFD113A3D" box="[630,650,334,359]" italics="true" pageId="5" pageNumber="382">H</emphasis>
. 
<emphasis id="B97DEAAD0F722D07FD7EFEB9FD763A3D" box="[665,749,333,359]" italics="true" pageId="5" pageNumber="382">dryadis</emphasis>
</taxonomicName>
in the shape of hood and the structure of phallus, but the former is easily distinguished from the latter by the following character states: outer margin of paranotum resting on pronotal disc except anterior end (
<figureCitation id="13322A3A0F722D07FDF6FE61FDF13AF5" box="[529,618,405,431]" captionStart="FIGURE 2" captionStartId="3.[151,250,1942,1967]" captionTargetBox="[201,1385,993,1917]" captionTargetId="figure@3.[201,1385,993,1917]" captionTargetPageId="3" captionText="FIGURE 2. Detail morphological images of Heissiella donguri sp. nov. A–C: cephalic and thoracic structures, dorsal, lateral and ventral views; D, E, hemelytron, dorsal and lateral views; F, apical part of abdomen of male, ventral view; G, same of female, ventral view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3713028" httpUri="https://zenodo.org/record/3713028/files/figure.png" pageId="5" pageNumber="382">Fig. 2A</figureCitation>
); costal area of hemelytron with 2–3 rows of areolae throughout its length (
<figureCitation id="13322A3A0F722D07FF47FE4DFF623A89" box="[160,249,441,467]" captionStart="FIGURE 2" captionStartId="3.[151,250,1942,1967]" captionTargetBox="[201,1385,993,1917]" captionTargetId="figure@3.[201,1385,993,1917]" captionTargetPageId="3" captionText="FIGURE 2. Detail morphological images of Heissiella donguri sp. nov. A–C: cephalic and thoracic structures, dorsal, lateral and ventral views; D, E, hemelytron, dorsal and lateral views; F, apical part of abdomen of male, ventral view; G, same of female, ventral view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3713028" httpUri="https://zenodo.org/record/3713028/files/figure.png" pageId="5" pageNumber="382">Fig. 2D</figureCitation>
); hypocostal lamina with 2 rows of areolae in basal half and a single row in apical half (
<figureCitation id="13322A3A0F722D07FB2EFE4DFABB3A89" box="[1225,1312,441,467]" captionStart="FIGURE 2" captionStartId="3.[151,250,1942,1967]" captionTargetBox="[201,1385,993,1917]" captionTargetId="figure@3.[201,1385,993,1917]" captionTargetPageId="3" captionText="FIGURE 2. Detail morphological images of Heissiella donguri sp. nov. A–C: cephalic and thoracic structures, dorsal, lateral and ventral views; D, E, hemelytron, dorsal and lateral views; F, apical part of abdomen of male, ventral view; G, same of female, ventral view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3713028" httpUri="https://zenodo.org/record/3713028/files/figure.png" pageId="5" pageNumber="382">Fig. 2E</figureCitation>
); and outer lobe of paratergite IX in female protruding laterad (
<figureCitation id="13322A3A0F722D07FD36FE29FCAF3AAD" box="[721,820,477,503]" captionStart="FIGURE 2" captionStartId="3.[151,250,1942,1967]" captionTargetBox="[201,1385,993,1917]" captionTargetId="figure@3.[201,1385,993,1917]" captionTargetPageId="3" captionText="FIGURE 2. Detail morphological images of Heissiella donguri sp. nov. A–C: cephalic and thoracic structures, dorsal, lateral and ventral views; D, E, hemelytron, dorsal and lateral views; F, apical part of abdomen of male, ventral view; G, same of female, ventral view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3713028" httpUri="https://zenodo.org/record/3713028/files/figure.png" pageId="5" pageNumber="382">Figs. 2G</figureCitation>
, 
<figureCitation id="13322A3A0F722D07FCD8FE29FCF93AAD" box="[831,866,477,503]" captionStart="FIGURE 3" captionStartId="4.[151,250,1494,1519]" captionTargetBox="[190,1402,459,1464]" captionTargetId="figure@4.[177,1410,453,1469]" captionTargetPageId="4" captionText="FIGURE 3. Male and female terminalia of Heissiella donguri sp. nov. A, pygophore, dorsal view; B, female terminalia, ventral view; C, D, phallus, dorsal and ventral views; E, paramere, dorsal view. Abbreviations: st 7, abdominal sternite VII; ov, ovivalvula; gcp 1, gonocoxopodite I; pt 8, paratergite VIII; pt 9, paratergite IX; ol, outer lobe; il, inner lobe; bp, basal plate; cp, capitate process; ph, phallotheca; en, endophallus. Scale bars: A–D, 0.2 mm; E, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3713030" httpUri="https://zenodo.org/record/3713030/files/figure.png" pageId="5" pageNumber="382">3B</figureCitation>
). In contrast, 
<taxonomicName id="4C094D3C0F722D07FC1EFE2AFBF43AAD" baseAuthorityName="Drake &amp; Poor" baseAuthorityYear="1936" box="[1017,1135,477,503]" class="Insecta" family="Tingidae" genus="Physatocheila" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="382" phylum="Arthropoda" rank="species" species="dryadis">
<emphasis id="B97DEAAD0F722D07FC1EFE2AFB963AAD" box="[1017,1037,478,503]" italics="true" pageId="5" pageNumber="382">H</emphasis>
. 
<emphasis id="B97DEAAD0F722D07FBFCFE29FBF43AAD" box="[1051,1135,477,503]" italics="true" pageId="5" pageNumber="382">dryadis</emphasis>
</taxonomicName>
has the following features: outer margin of paranotum resting on pronotal disc at posterior end; costal area of hemelytron with 2 rows of areolae throughout its length; hypocostal lamina with a single row of areolae throughout its length; and outer lobe of paratergite IX in female protruding posteriad.
</paragraph>
<paragraph id="8BB636BF0F722D07FF20FD99FA0639F1" blockId="5.[151,1437,153,1151]" pageId="5" pageNumber="382">
The tingid species recorded as ‘ 
<taxonomicName id="4C094D3C0F722D07FDC2FD9AFC8539DD" authority="" box="[549,798,621,647]" class="Insecta" family="Tingidae" genus="Tingis" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="382" phylum="Arthropoda" rank="subGenus" subGenus="Tropidocheila">
<emphasis id="B97DEAAD0F722D07FDC2FD9AFDF239DD" box="[549,617,622,647]" italics="true" pageId="5" pageNumber="382">Tingis</emphasis>
(
<emphasis id="B97DEAAD0F722D07FD9EFD99FC8E39DD" box="[633,789,621,647]" italics="true" pageId="5" pageNumber="382">Tropidocheila</emphasis>
)
</taxonomicName>
sp.’ or ‘ 
<taxonomicName id="4C094D3C0F722D07FC9FFD9AFF4939F1" authority="Drake &amp; Maa, 1953" authorityName="Drake &amp; Maa" authorityYear="1953" class="Insecta" family="Tingidae" genus="Tingis" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="382" phylum="Arthropoda" rank="species" species="shaowuana" subGenus="Tropidocheila">
<emphasis id="B97DEAAD0F722D07FC9FFD9AFC2739DD" box="[888,956,622,647]" italics="true" pageId="5" pageNumber="382">Tingis</emphasis>
(
<emphasis id="B97DEAAD0F722D07FC2BFD99FBF339DD" box="[972,1128,621,647]" italics="true" pageId="5" pageNumber="382">Tropidocheila</emphasis>
) 
<emphasis id="B97DEAAD0F722D07FB9FFD99FB6339DD" box="[1144,1272,621,647]" italics="true" pageId="5" pageNumber="382">shaowuana</emphasis>
Drake &amp; Maa, 1953
</taxonomicName>
’ by previous authors (
<bibRefCitation id="EF984B4E0F722D07FE2EFD65FDE839F1" author="Komatsu, T." box="[457,627,657,683]" pageId="5" pageNumber="382" refId="ref5230" refString="Komatsu, T. (2016) [Terrestrial heteropterans from Miyazaki-ken, Kyushu, Japan]. Kuroshio Bunko, Miyazaki, 139 pp. [in Japanese]" type="book" year="2016">Komatsu 2016</bibRefCitation>
; 
<bibRefCitation id="EF984B4E0F722D07FD99FD65FC8E39F1" author="Miyake, T." box="[638,789,657,683]" pageId="5" pageNumber="382" pagination="22" refId="ref5292" refString="Miyake, T. (2017) [Some distributional records of insects from Ume district, Saeki-shi, Oita-ken, Kyushu, Japan]. Niho-no- Mushi, 55, 22. [in Japanese]" type="journal article" year="2017">Miyake 2017</bibRefCitation>
; 
<bibRefCitation id="EF984B4E0F722D07FCC6FD65FC7439F1" author="Ito, R. &amp; Sasaki, S." box="[801,1007,657,683]" pageId="5" pageNumber="382" pagination="17 - 24" refId="ref5145" refString="Ito, R. &amp; Sasaki, S. (2018) [Tingidae from Oita-ken, Kyushu, Japan (Heteroptera)]. Niho-no-Mushi, 56, 17 - 24. [in Japanese]" type="journal article" year="2018">Ito &amp; Sasaki 2018</bibRefCitation>
; 
<bibRefCitation id="EF984B4E0F722D07FC1CFD65FBC039F1" author="Ito, R." box="[1019,1115,657,683]" pageId="5" pageNumber="382" pagination="77 - 78" refId="ref5096" refString="Ito, R. (2019) [First record of Dictyla evidens (Drake, 1927) from Oita-ken, Kyushu, Japan, with a correction of distributional records of Tingidae from Oita-ken]. Niho-no-Mushi, 57, 77 - 78. [in Japanese]" type="journal article" year="2019">Ito 2019</bibRefCitation>
) pertains to the new species.
</paragraph>
</subSubSection>
<subSubSection id="C31365340F722D07FF20FD40FC113861" pageId="5" pageNumber="382" type="distribution">
<paragraph id="8BB636BF0F722D07FF20FD40FD443995" blockId="5.[151,1437,153,1151]" box="[199,735,692,720]" pageId="5" pageNumber="382">
<emphasis id="B97DEAAD0F722D07FF20FD40FEFA3995" bold="true" box="[199,353,692,719]" pageId="5" pageNumber="382">Distribution.</emphasis>
<collectingCountry id="F31E762F0F722D07FE80FD41FE3C3995" box="[359,423,693,719]" name="Japan" pageId="5" pageNumber="382">Japan</collectingCountry>
(Honshu, Kyushu) (Fig. 5).
</paragraph>
<paragraph id="8BB636BF0F722D07FF20FD2DFC113861" blockId="5.[151,1437,153,1151]" pageId="5" pageNumber="382">
This new species inhabits the laurilignosa ecosystem of 
<collectingCountry id="F31E762F0F722D07FCA0FD2DFC1C39A9" box="[839,903,729,755]" name="Japan" pageId="5" pageNumber="382">Japan</collectingCountry>
proper in a warm-temperate climate and is the first representative of 
<taxonomicName id="4C094D3C0F722D07FE6BFD09FE67384D" baseAuthorityName="Dang &amp; Bu" baseAuthorityYear="2012" box="[396,508,765,791]" class="Insecta" family="Tingidae" genus="Heissiella" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="382" phylum="Arthropoda" rank="genus">
<emphasis id="B97DEAAD0F722D07FE6BFD09FE67384D" box="[396,508,765,791]" italics="true" pageId="5" pageNumber="382">Heissiella</emphasis>
</taxonomicName>
from 
<collectingCountry id="F31E762F0F722D07FDD8FD09FDE4384D" box="[575,639,765,791]" name="Japan" pageId="5" pageNumber="382">Japan</collectingCountry>
as well as the Palaearctic Region. The discovery of 
<taxonomicName id="4C094D3C0F722D07FB25FD0AFAA6384D" authority="Souma &amp; Ishikawa, 2020" authorityName="Souma &amp; Ishikawa" authorityYear="2020" box="[1218,1341,765,791]" class="Insecta" family="Tingidae" genus="Heissiella" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="382" phylum="Arthropoda" rank="species" species="donguri" status="sp. nov.">
<emphasis id="B97DEAAD0F722D07FB25FD0AFB4D384D" box="[1218,1238,766,791]" italics="true" pageId="5" pageNumber="382">H</emphasis>
. 
<emphasis id="B97DEAAD0F722D07FB03FD09FAA6384D" box="[1252,1341,765,791]" italics="true" pageId="5" pageNumber="382">donguri</emphasis>
</taxonomicName>
<emphasis id="B97DEAAD0F722D07FAA4FD09FA07384D" bold="true" box="[1347,1436,765,791]" pageId="5" pageNumber="382">
<taxonomicNameLabel id="A24E57D60F722D07FAA4FD09FA07384D" box="[1347,1436,765,791]" pageId="5" pageNumber="382" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
represents the easternmost distribution record of 
<taxonomicName id="4C094D3C0F722D07FD5CFCD5FCB03861" baseAuthorityName="Dang &amp; Bu" baseAuthorityYear="2012" box="[699,811,801,827]" class="Insecta" family="Tingidae" genus="Heissiella" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="382" phylum="Arthropoda" rank="genus">
<emphasis id="B97DEAAD0F722D07FD5CFCD5FCB03861" box="[699,811,801,827]" italics="true" pageId="5" pageNumber="382">Heissiella</emphasis>
</taxonomicName>
species.
</paragraph>
</subSubSection>
<subSubSection id="C31365340F722D07FF20FCB1FC5D38D9" pageId="5" pageNumber="382" type="etymology">
<paragraph id="8BB636BF0F722D07FF20FCB1FC5D38D9" blockId="5.[151,1437,153,1151]" pageId="5" pageNumber="382">
<emphasis id="B97DEAAD0F722D07FF20FCB1FED53805" bold="true" box="[199,334,837,863]" pageId="5" pageNumber="382">Etymology.</emphasis>
The specific epithet is named after a Japanese common name “donguri” (= acorns of 
<taxonomicName id="4C094D3C0F722D07FAEEFCB1FAFC3805" box="[1289,1383,837,863]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="5" pageNumber="382" phylum="Tracheophyta" rank="genus">
<emphasis id="B97DEAAD0F722D07FAEEFCB1FAFC3805" box="[1289,1383,837,863]" italics="true" pageId="5" pageNumber="382">Quercus</emphasis>
</taxonomicName>
species), referring to the host plant of the new species; a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C31365340F722D07FF20FC79FABC3F25" pageId="5" pageNumber="382" type="biology_ecology">
<paragraph id="8BB636BF0F722D07FF20FC79FC343891" blockId="5.[151,1437,153,1151]" pageId="5" pageNumber="382">
<emphasis id="B97DEAAD0F722D07FF20FC79FED738FD" bold="true" box="[199,332,909,935]" pageId="5" pageNumber="382">Host plant.</emphasis>
<taxonomicName id="4C094D3C0F722D07FEB4FC79FD5F38FD" authority="Blume" authorityName="Blume" authorityYear="1851" box="[339,708,909,935]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="5" pageNumber="382" phylum="Tracheophyta" rank="species" species="gilva">
<emphasis id="B97DEAAD0F722D07FEB4FC79FE6B38FD" box="[339,496,909,935]" italics="true" pageId="5" pageNumber="382">Quercus gilva</emphasis>
Blume (Fagaceae)
</taxonomicName>
(
<figureCitation id="13322A3A0F722D07FD33FC79FCB138FD" box="[724,810,909,935]" captionStart="FIGURE 4" captionStartId="5.[151,250,1680,1705]" captionTargetBox="[151,1437,1173,1655]" captionTargetId="figure@5.[151,1437,1173,1655]" captionTargetPageId="5" captionText="FIGURE 4. Heissiella donguri sp. nov. A, Habitus image of a living adult (photograph taken indoor); B, host plant (Quercus gilva Blume) from Nara, Honshu, Japan." figureDoi="http://doi.org/10.5281/zenodo.3713032" httpUri="https://zenodo.org/record/3713032/files/figure.png" pageId="5" pageNumber="382">Fig. 4B</figureCitation>
) has been confirmed as a host plant for the new species by several researchers (cf. 
<bibRefCitation id="EF984B4E0F722D07FE25FC45FDC13891" author="Miyake, T." box="[450,602,945,971]" pageId="5" pageNumber="382" pagination="22" refId="ref5292" refString="Miyake, T. (2017) [Some distributional records of insects from Ume district, Saeki-shi, Oita-ken, Kyushu, Japan]. Niho-no- Mushi, 55, 22. [in Japanese]" type="journal article" year="2017">Miyake 2017</bibRefCitation>
; 
<bibRefCitation id="EF984B4E0F722D07FD81FC45FCAF3891" author="Ito, R. &amp; Sasaki, S." box="[614,820,945,971]" pageId="5" pageNumber="382" pagination="17 - 24" refId="ref5145" refString="Ito, R. &amp; Sasaki, S. (2018) [Tingidae from Oita-ken, Kyushu, Japan (Heteroptera)]. Niho-no-Mushi, 56, 17 - 24. [in Japanese]" type="journal article" year="2018">Ito &amp; Sasaki 2018</bibRefCitation>
; 
<bibRefCitation id="EF984B4E0F722D07FCA7FC45FC383891" author="Ito, R." box="[832,931,945,971]" pageId="5" pageNumber="382" pagination="77 - 78" refId="ref5096" refString="Ito, R. (2019) [First record of Dictyla evidens (Drake, 1927) from Oita-ken, Kyushu, Japan, with a correction of distributional records of Tingidae from Oita-ken]. Niho-no-Mushi, 57, 77 - 78. [in Japanese]" type="journal article" year="2019">Ito 2019</bibRefCitation>
).
</paragraph>
<paragraph id="8BB636BF0F722D07FF20FC20FDAF3F6D" blockId="5.[151,1437,153,1151]" pageId="5" pageNumber="382">
<emphasis id="B97DEAAD0F722D07FF20FC20FEBC38B5" bold="true" box="[199,295,980,1007]" pageId="5" pageNumber="382">Biology.</emphasis>
All the 
<typeStatus id="54B2881D0F722D07FE67FC21FE2B38B5" box="[384,432,981,1007]" pageId="5" pageNumber="382">type</typeStatus>
materials of 
<taxonomicName id="4C094D3C0F722D07FDA3FC21FC8F38B5" authority="Souma &amp; Ishikawa, 2020" authorityName="Souma &amp; Ishikawa" authorityYear="2020" box="[580,788,981,1007]" class="Insecta" family="Tingidae" genus="Heissiella" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="382" phylum="Arthropoda" rank="species" species="donguri" status="sp. nov.">
<emphasis id="B97DEAAD0F722D07FDA3FC21FC8F38B5" box="[580,788,981,1007]" italics="true" pageId="5" pageNumber="382">Heissiella donguri</emphasis>
</taxonomicName>
<emphasis id="B97DEAAD0F722D07FCFDFC21FCEF38B5" bold="true" box="[794,884,981,1007]" pageId="5" pageNumber="382">
<taxonomicNameLabel id="A24E57D60F722D07FCFDFC21FCEF38B5" box="[794,884,981,1007]" pageId="5" pageNumber="382" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
were only collected from the underside of leaves of the host plant 
<taxonomicName id="4C094D3C0F722D07FEBEFC0DFE6C3F49" authorityName="Blume" authorityYear="1851" box="[345,503,1017,1043]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="5" pageNumber="382" phylum="Tracheophyta" rank="species" species="gilva">
<emphasis id="B97DEAAD0F722D07FEBEFC0DFE6C3F49" box="[345,503,1017,1043]" italics="true" pageId="5" pageNumber="382">Quercus gilva</emphasis>
</taxonomicName>
, suggesting that this new species appears to feed on the underside of leaves as do many tingids (
<bibRefCitation id="EF984B4E0F722D07FED0FBE9FDBC3F6D" author="Schuh, R. T. &amp; Slater, J. A." box="[311,551,1052,1079]" pageId="5" pageNumber="382" refId="ref5526" refString="Schuh, R. T. &amp; Slater, J. A. (1995) True bugs of the world (Hemiptera: Heteroptera). Classification and natural history. Cornell University Press, Ithaca, New York, xii + 336 pp." type="book" year="1995">Schuh &amp; Slater 1995</bibRefCitation>
).
</paragraph>
<paragraph id="8BB636BF0F722D07FF20FBB5FABC3F25" blockId="5.[151,1437,153,1151]" pageId="5" pageNumber="382">
Adults of the new species were collected in December. Thus, the overwintering form of 
<taxonomicName id="4C094D3C0F722D07FB5CFBB6FAA33F01" authority="Souma &amp; Ishikawa, 2020" authorityName="Souma &amp; Ishikawa" authorityYear="2020" box="[1211,1336,1089,1115]" class="Insecta" family="Tingidae" genus="Heissiella" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="382" phylum="Arthropoda" rank="species" species="donguri" status="sp. nov.">
<emphasis id="B97DEAAD0F722D07FB5CFBB6FB543F01" box="[1211,1231,1090,1115]" italics="true" pageId="5" pageNumber="382">H</emphasis>
. 
<emphasis id="B97DEAAD0F722D07FB38FBB5FAA33F01" box="[1247,1336,1089,1115]" italics="true" pageId="5" pageNumber="382">donguri</emphasis>
</taxonomicName>
<emphasis id="B97DEAAD0F722D07FAA6FBB5FA073F01" bold="true" box="[1345,1436,1089,1115]" pageId="5" pageNumber="382">
<taxonomicNameLabel id="A24E57D60F722D07FAA6FBB5FA073F01" box="[1345,1436,1089,1115]" pageId="5" pageNumber="382" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
appears to be adult, in accordance with many other tingids in the temperate zone (
<bibRefCitation id="EF984B4E0F722D07FBCDFB91FA813F25" author="Schuh, R. T. &amp; Slater, J. A." box="[1066,1306,1124,1151]" pageId="5" pageNumber="382" refId="ref5526" refString="Schuh, R. T. &amp; Slater, J. A. (1995) True bugs of the world (Hemiptera: Heteroptera). Classification and natural history. Cornell University Press, Ithaca, New York, xii + 336 pp." type="book" year="1995">Schuh &amp; Slater 1995</bibRefCitation>
).
</paragraph>
</subSubSection>
<caption id="DF7666370F722D07FF70F964FDAF3D96" ID-DOI="http://doi.org/10.5281/zenodo.3713032" ID-Zenodo-Dep="3713032" httpUri="https://zenodo.org/record/3713032/files/figure.png" pageId="5" pageNumber="382" startId="5.[151,250,1680,1705]" targetBox="[151,1437,1173,1655]" targetPageId="5">
<paragraph id="8BB636BF0F722D07FF70F964FDAF3D96" blockId="5.[151,1436,1680,1849]" pageId="5" pageNumber="382">
<emphasis id="B97DEAAD0F722D07FF70F964FE8F3DF3" bold="true" box="[151,276,1680,1705]" pageId="5" pageNumber="382">FIGURE 4.</emphasis>
<taxonomicName id="4C094D3C0F722D07FEFCF965FE4C3DF3" authority="Souma &amp; Ishikawa, 2020" authorityName="Souma &amp; Ishikawa" authorityYear="2020" box="[283,471,1681,1705]" class="Insecta" family="Tingidae" genus="Heissiella" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="382" phylum="Arthropoda" rank="species" species="donguri" status="sp. nov.">
<emphasis id="B97DEAAD0F722D07FEFCF965FE4C3DF3" box="[283,471,1681,1705]" italics="true" pageId="5" pageNumber="382">Heissiella donguri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24E57D60F722D07FE38F965FDAB3DF3" box="[479,560,1681,1705]" pageId="5" pageNumber="382" rank="species">
<emphasis id="B97DEAAD0F722D07FE38F965FDAB3DF3" bold="true" box="[479,560,1681,1705]" pageId="5" pageNumber="382">sp. nov.</emphasis>
</taxonomicNameLabel>
A, Habitus image of a living adult (photograph taken indoor); B, host plant (
<taxonomicName id="4C094D3C0F722D07FAA0F965FE8E3D97" authority="Blume" authorityName="Blume" authorityYear="1851" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="5" pageNumber="382" phylum="Tracheophyta" rank="species" species="gilva">
<emphasis id="B97DEAAD0F722D07FAA0F965FF513D97" italics="true" pageId="5" pageNumber="382">Quercus gilva</emphasis>
Blume
</taxonomicName>
) from Nara, Honshu, Japan.
</paragraph>
</caption>
<subSubSection id="C31365340F722D04FF70F8EBFE803DD3" lastPageId="6" lastPageNumber="383" pageId="5" pageNumber="382" type="key">
<paragraph id="8BB636BF0F722D07FF70F8EBFD983C63" blockId="5.[151,1436,1680,1849]" box="[151,515,1822,1849]" pageId="5" pageNumber="382">
<emphasis id="B97DEAAD0F722D07FF70F8EBFD983C63" bold="true" box="[151,515,1822,1849]" pageId="5" pageNumber="382">
Key to the species of 
<taxonomicName id="4C094D3C0F722D07FE76F8EAFD983C63" ID-CoL="62TGP" baseAuthorityName="Dang &amp; Bu" baseAuthorityYear="2012" box="[401,515,1822,1849]" class="Insecta" family="Tingidae" genus="Heissiella" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="382" phylum="Arthropoda" rank="genus">
<emphasis id="B97DEAAD0F722D07FE76F8EAFD983C63" bold="true" box="[401,515,1822,1849]" italics="true" pageId="5" pageNumber="382">Heissiella</emphasis>
</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="8BB636BF0F722D07FF70F893FEBF3CB7" blockId="5.[151,1436,1895,2030]" pageId="5" pageNumber="382">
The key below is based on a comparison of the 
<typeStatus id="54B2881D0F722D07FD50F89CFD7C3CD8" box="[695,743,1896,1922]" pageId="5" pageNumber="382">type</typeStatus>
material of the new species with the photographs of the 
<typeStatus id="54B2881D0F722D07FA8BF89CFA073CD8" box="[1388,1436,1896,1922]" pageId="5" pageNumber="382">type</typeStatus>
material of 
<taxonomicName id="4C094D3C0F722D07FEFBF878FE0F3CFF" ID-CoL="4HGGB" baseAuthorityName="Drake &amp; Poor" baseAuthorityYear="1936" box="[284,404,1931,1958]" class="Insecta" family="Tingidae" genus="Physatocheila" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="382" phylum="Arthropoda" rank="species" species="dryadis">
<emphasis id="B97DEAAD0F722D07FEFBF878FEAB3CFF" box="[284,304,1932,1957]" italics="true" pageId="5" pageNumber="382">H</emphasis>
. 
<emphasis id="B97DEAAD0F722D07FEA7F87FFE0F3CFF" box="[320,404,1931,1957]" italics="true" pageId="5" pageNumber="382">dryadis</emphasis>
</taxonomicName>
(http://n 
<quantity id="4CF19B5A0F722D07FE11F87EFD973CFF" box="[502,524,1930,1957]" metricMagnitude="3" metricUnit="kg" metricValue="2.0" pageId="5" pageNumber="382" unit="t" value="2.0">2t</quantity>
.net/ark:/65665/3e0c902f4- 
<date id="FFB7107F0F722D07FCA7F87EFC063CFF" box="[832,925,1930,1957]" pageId="5" pageNumber="382" value="1126" valueMax="1147">1126-47</date>
d9-8345-340fe4a365bc), the descriptions and illustrations of other congeners (
<bibRefCitation id="EF984B4E0F722D07FE19F85BFD783C93" author="Drake, C. J. &amp; Poor, M. E." box="[510,739,1967,1994]" pageId="5" pageNumber="382" pagination="141 - 149" refId="ref4943" refString="Drake, C. J. &amp; Poor, M. E. (1936) New Indian Tingitidae (Hemiptera). Indian forest records (New Series). Entomology, 2 (5), 141 - 149." type="journal article" year="1936">Drake &amp; Poor, 1936</bibRefCitation>
; 
<bibRefCitation id="EF984B4E0F722D07FD09F85BFC1D3C93" author="Pericart, J." box="[750,902,1967,1993]" pageId="5" pageNumber="382" pagination="23 - 47" refId="ref5423" refString="Pericart, J. (1984) Tingidae nouveaux ou interessants du Nord du sous-continent indien (Hemiptera). I. Entomologica Basiliensia, 9, 23 - 47. [in French, English summary]" type="journal article" year="1984">Péricart 1984</bibRefCitation>
; 
<bibRefCitation id="EF984B4E0F722D07FC76F844FBC83C93" author="Dang, K. &amp; Bu, W." box="[913,1107,1967,1994]" pageId="5" pageNumber="382" pagination="4" refId="ref4837" refString="Dang, K. &amp; Bu, W. (2012) The discovery of the genus Heissiella Pericart, 1984 (Hemiptera: Heteroptera: Tingidae) from China, with description of a new species. Entomotaxonomia, 34 (4), 654 - 628." type="journal article" year="2012">Dang &amp; Bu 2012</bibRefCitation>
), and the available key (
<bibRefCitation id="EF984B4E0F722D07FAB9F85BFE823CB7" author="Dang, K. &amp; Bu, W." pageId="5" pageNumber="382" pagination="4" refId="ref4837" refString="Dang, K. &amp; Bu, W. (2012) The discovery of the genus Heissiella Pericart, 1984 (Hemiptera: Heteroptera: Tingidae) from China, with description of a new species. Entomotaxonomia, 34 (4), 654 - 628." type="journal article" year="2012">Dang &amp; Bu 2012</bibRefCitation>
).
</paragraph>
<key id="FF98618E0F712D04FF70FF4EFA073948" pageId="6" pageNumber="383">
<keyStep id="30FD2E1A0F712D04FF70FF4EFA073A7D" pageId="6" pageNumber="383">
<paragraph id="8BB636BF0F712D04FF70FF4EFA073BB6" blockId="6.[151,1437,185,530]" pageId="6" pageNumber="383">
<keyLead id="30F8958A0F712D04FF70FF4EFA073BB6" pageId="6" pageNumber="383">
1 Hypocostal lamina of hemelytron with 2 rows of areolae in basal half and a single row in apical half; outer lobe of paratergite IX in female protruding laterad........................................................... 
<taxonomicName id="4C094D3C0F712D04FB07FF2CFAD13BB6" authority="Souma &amp; Ishikawa, 2020" authorityName="Souma &amp; Ishikawa" authorityYear="2020" box="[1248,1354,215,237]" class="Insecta" family="Tingidae" genus="Heissiella" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="383" phylum="Arthropoda" rank="species" species="donguri" status="sp. nov.">
<emphasis id="B97DEAAD0F712D04FB07FF2CFB6A3BB7" box="[1248,1265,216,237]" italics="true" pageId="6" pageNumber="383">H</emphasis>
. 
<emphasis id="B97DEAAD0F712D04FB19FF23FAD13BB6" box="[1278,1354,215,236]" italics="true" pageId="6" pageNumber="383">donguri</emphasis>
</taxonomicName>
<emphasis id="B97DEAAD0F712D04FAB7FF23FA073BB6" bold="true" box="[1360,1436,215,236]" pageId="6" pageNumber="383">
<taxonomicNameLabel id="A24E57D60F712D04FAB7FF23FA073BB6" box="[1360,1436,215,236]" pageId="6" pageNumber="383" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BB636BF0F712D04FF70FF01FA073A7D" blockId="6.[151,1437,185,530]" pageId="6" pageNumber="383">
<keyLead id="30F8958A0F712D04FF70FF01FA073A7D" pageId="6" pageNumber="383">- Hypocostal lamina of hemelytron with a single row of areolae throughout its length; outer lobe of paratergite IX in female protruding posteriad................................................................................... 2</keyLead>
</paragraph>
</keyStep>
<keyStep id="30FD2E1A0F712D04FF70FEDBFA073AC6" pageId="6" pageNumber="383">
<paragraph id="8BB636BF0F712D04FF70FEDBFA073A3B" blockId="6.[151,1437,185,530]" pageId="6" pageNumber="383">
<keyLead id="30F8958A0F712D04FF70FEDBFA073A3B" pageId="6" pageNumber="383">
2 Outer margin of paranotum resting on pronotal disc except anterior end; outer lobe of paratergite IX in female longer than inner lobe; inner lobe of paratergite IX in female angular at apex............................... 
<taxonomicName id="4C094D3C0F712D04FB75FEB9FA073A3B" ID-CoL="3K3VZ" authority="Pericart, 1984" authorityName="Pericart" authorityYear="1984" box="[1170,1436,332,354]" class="Insecta" family="Tingidae" genus="Heissiella" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="383" phylum="Arthropoda" rank="species" species="bicaudata">
<emphasis id="B97DEAAD0F712D04FB75FEB9FB383A38" box="[1170,1187,333,354]" italics="true" pageId="6" pageNumber="383">H</emphasis>
. 
<emphasis id="B97DEAAD0F712D04FB57FEB8FA8B3A3B" box="[1200,1296,332,353]" italics="true" pageId="6" pageNumber="383">bicaudata</emphasis>
<bibRefCitation id="EF984B4E0F712D04FAF1FEB8FA073A3B" author="Pericart, J." box="[1302,1436,332,354]" pageId="6" pageNumber="383" pagination="23 - 47" refId="ref5423" refString="Pericart, J. (1984) Tingidae nouveaux ou interessants du Nord du sous-continent indien (Hemiptera). I. Entomologica Basiliensia, 9, 23 - 47. [in French, English summary]" type="journal article" year="1984">Péricart, 1984</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BB636BF0F712D04FF70FE9EFA073AC6" blockId="6.[151,1437,185,530]" pageId="6" pageNumber="383">
<keyLead id="30F8958A0F712D04FF70FE9EFA073AC6" pageId="6" pageNumber="383">- Outer margin of paranotum resting on pronotal disc at posterior end; outer lobe of paratergite IX in female shorter than inner lobe; inner lobe of paratergite IX in female rounded at apex................................................... 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="30FD2E1A0F712D04FF70FE50FA073948" pageId="6" pageNumber="383">
<paragraph id="8BB636BF0F712D04FF70FE50FA073A8D" blockId="6.[151,1437,185,530]" pageId="6" pageNumber="383">
<keyLead id="30F8958A0F712D04FF70FE50FA073A8D" pageId="6" pageNumber="383">
3 Dorsal outline of hood arched in lateral view; costal area of hemelytron with 2–3 rows of areolae throughout its length; endophallus with a pair of inner sclerites basally........................................... 
<taxonomicName id="4C094D3C0F712D04FB72FE36FA073A8D" authority="Dang &amp; Bu, 2012" authorityName="Dang &amp; Bu" authorityYear="2012" box="[1173,1436,450,471]" class="Insecta" family="Tingidae" genus="Heissiella" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="383" phylum="Arthropoda" rank="species" species="sinica">
<emphasis id="B97DEAAD0F712D04FB72FE36FB3D3A8D" box="[1173,1190,450,471]" italics="true" pageId="6" pageNumber="383">H</emphasis>
. 
<emphasis id="B97DEAAD0F712D04FB55FE36FB703A8D" box="[1202,1259,450,471]" italics="true" pageId="6" pageNumber="383">sinica</emphasis>
<bibRefCitation id="EF984B4E0F712D04FB15FE36FA073A8D" author="Dang, K. &amp; Bu, W." box="[1266,1436,450,471]" pageId="6" pageNumber="383" pagination="4" refId="ref4837" refString="Dang, K. &amp; Bu, W. (2012) The discovery of the genus Heissiella Pericart, 1984 (Hemiptera: Heteroptera: Tingidae) from China, with description of a new species. Entomotaxonomia, 34 (4), 654 - 628." type="journal article" year="2012">Dang &amp; Bu, 2012</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BB636BF0F712D04FF70FE2BFA073948" blockId="6.[151,1437,185,530]" pageId="6" pageNumber="383">
<keyLead id="30F8958A0F712D04FF70FE2BFA073948" pageId="6" pageNumber="383">
- Dorsal outline of hood nearly straight in lateral view; costal area of hemelytron with 2 rows of areolae throughout its length; endophallus lacking inner sclerite basally........................................ 
<taxonomicName id="4C094D3C0F712D04FBB8FE09FA073948" ID-CoL="4HGGB" authority="(Drake &amp; Poor, 1936)" baseAuthorityName="Drake &amp; Poor" baseAuthorityYear="1936" box="[1119,1436,508,530]" class="Insecta" family="Tingidae" genus="Physatocheila" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="383" phylum="Arthropoda" rank="species" species="dryadis">
<emphasis id="B97DEAAD0F712D04FBB8FE09FBEB3948" box="[1119,1136,509,530]" italics="true" pageId="6" pageNumber="383">H</emphasis>
. 
<emphasis id="B97DEAAD0F712D04FB9AFE08FB5E394B" box="[1149,1221,508,529]" italics="true" pageId="6" pageNumber="383">dryadis</emphasis>
(
<bibRefCitation id="EF984B4E0F712D04FB36FE08FA0E3948" author="Drake, C. J. &amp; Poor, M. E." box="[1233,1429,508,530]" pageId="6" pageNumber="383" pagination="141 - 149" refId="ref4943" refString="Drake, C. J. &amp; Poor, M. E. (1936) New Indian Tingitidae (Hemiptera). Indian forest records (New Series). Entomology, 2 (5), 141 - 149." type="journal article" year="1936">Drake &amp; Poor, 1936</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
<paragraph id="8BB636BF0F712D04FF70F9F0FE803DD3" blockId="6.[151,1431,1540,1673]" pageId="6" pageNumber="383">
<emphasis id="B97DEAAD0F712D04FF70F9F0FE883D47" bold="true" box="[151,275,1540,1565]" pageId="6" pageNumber="383">FIGURE 5.</emphasis>
Distribution of 
<taxonomicName id="4C094D3C0F712D04FE51F9F1FDEA3D47" authority="Souma &amp; Ishikawa, 2020" authorityName="Souma &amp; Ishikawa" authorityYear="2020" box="[438,625,1541,1565]" class="Insecta" family="Tingidae" genus="Heissiella" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="383" phylum="Arthropoda" rank="species" species="donguri" status="sp. nov.">
<emphasis id="B97DEAAD0F712D04FE51F9F1FDEA3D47" box="[438,625,1541,1565]" italics="true" pageId="6" pageNumber="383">Heissiella donguri</emphasis>
</taxonomicName>
<emphasis id="B97DEAAD0F712D04FD9FF9F1FD533D47" bold="true" box="[632,712,1541,1565]" pageId="6" pageNumber="383">
<taxonomicNameLabel id="A24E57D60F712D04FD9FF9F1FD533D47" box="[632,712,1541,1565]" pageId="6" pageNumber="383" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in 
<collectingCountry id="F31E762F0F712D04FD0EF9F1FCB83D46" box="[745,803,1541,1564]" name="Japan" pageId="6" pageNumber="383">Japan</collectingCountry>
proper. 
<typeStatus id="54B2881D0F712D04FC90F9F0FC4E3D47" box="[887,981,1540,1565]" pageId="6" pageNumber="383" type="holotype">Holotype</typeStatus>
locality=circle; 
<typeStatus id="54B2881D0F712D04FB9CF9F1FB493D47" box="[1147,1234,1541,1565]" pageId="6" pageNumber="383" type="paratype">paratype</typeStatus>
localities=triangle. 
<emphasis id="B97DEAAD0F712D04FF70F99BFE803DD3" bold="true" box="[151,283,1647,1673]" pageId="6" pageNumber="383">Discussion</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C31365340F712D04FF70F943FE503C97" pageId="6" pageNumber="383" type="description">
<paragraph id="8BB636BF0F712D04FF70F943FAC33C3B" blockId="6.[151,1437,1719,1997]" pageId="6" pageNumber="383">
The Japanese species of 
<taxonomicName id="4C094D3C0F712D04FE4FF943FD9D3D8B" box="[424,518,1719,1745]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="6" pageNumber="383" phylum="Tracheophyta" rank="genus">
<emphasis id="B97DEAAD0F712D04FE4FF943FD9D3D8B" box="[424,518,1719,1745]" italics="true" pageId="6" pageNumber="383">Quercus</emphasis>
</taxonomicName>
are confirmed as host plants for a number of the phytophagous insects (e.g. 
<bibRefCitation id="EF984B4E0F712D04FAB1F943FF6F3DAF" author="Shirozu, T." pageId="6" pageNumber="383" refId="ref5572" refString="Shirozu, T. (2006) The butterflies of Japan in color. Gakken, Tokyo, 336 pp. [in Japanese]" type="book" year="2006">Shirozu 2006</bibRefCitation>
; 
<bibRefCitation id="EF984B4E0F712D04FF18F92FFDB63DAF" author="Ohbayashi, N. &amp; Niisato, T." box="[255,557,1755,1781]" pageId="6" pageNumber="383" refId="ref5386" refString="Ohbayashi, N. &amp; Niisato, T. (2007) Longicorn beetles of Japan. Tokai University Press, Hadano, xii + 818 pp., 22 figs., 130 color pls." type="book" year="2007">Ohbayashi &amp; Niisato 2007</bibRefCitation>
; 
<bibRefCitation id="EF984B4E0F712D04FDDFF92FFD7C3DAF" author="Takizawa, H." box="[568,743,1755,1781]" pageId="6" pageNumber="383" pagination="1 - 11" refId="ref5839" refString="Takizawa, H. (2009) [Remarks on biology of Chrysomelidae from Japan. IV]. Kanagawa-Chuho, 168, 1 - 11. [in Japanese]" type="journal article" year="2009">Takizawa 2009</bibRefCitation>
). However, only two species belonging to the family 
<taxonomicName id="4C094D3C0F712D04FADDF92FFA063DAF" authorityName="Laporte" authorityYear="1832" box="[1338,1437,1755,1781]" class="Insecta" family="Tingidae" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="383" phylum="Arthropoda" rank="family">Tingidae</taxonomicName>
have been found on 
<taxonomicName id="4C094D3C0F712D04FE9CF90BFE423C43" box="[379,473,1791,1817]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="6" pageNumber="383" phylum="Tracheophyta" rank="genus">
<emphasis id="B97DEAAD0F712D04FE9CF90BFE423C43" box="[379,473,1791,1817]" italics="true" pageId="6" pageNumber="383">Quercus</emphasis>
</taxonomicName>
species in 
<collectingCountry id="F31E762F0F712D04FDB1F90BFD0D3C43" box="[598,662,1791,1817]" name="Japan" pageId="6" pageNumber="383">Japan</collectingCountry>
(
<bibRefCitation id="EF984B4E0F712D04FD43F90BFCF93C43" author="Yano, S. &amp; Takechi, L. &amp; Kikuhara, Y." box="[676,866,1791,1817]" pageId="6" pageNumber="383" pagination="23 - 52" refId="ref6017" refString="Yano, S., Takechi, L. &amp; Kikuhara, Y. (2013) Heteroptera of Ehime Prefecture, Shikoku, Japan (Insecta, Hemiptera) I. Bulletin of Omogo mountain museum, 5, 23 - 52. [in Japanese]" type="journal article" year="2013">
Yano 
<emphasis id="B97DEAAD0F712D04FD00F8F4FC823C43" box="[743,793,1791,1817]" italics="true" pageId="6" pageNumber="383">et al</emphasis>
. 2013
</bibRefCitation>
; 
<bibRefCitation id="EF984B4E0F712D04FC89F90BFB8F3C43" author="Maehara, S." box="[878,1044,1791,1817]" pageId="6" pageNumber="383" pagination="57 - 64" refId="ref5260" refString="Maehara, S. (2014) [Tingidae from Tochigi-ken, Honshu, Japan]. Insekuto, 65 (1), 57 - 64. [in Japanese]" type="journal article" year="2014">Maehara 2014</bibRefCitation>
). Therefore, the discovery of 
<taxonomicName id="4C094D3C0F712D04FA87F8F4FEA83C67" authority="Souma &amp; Ishikawa, 2020" authorityName="Souma &amp; Ishikawa" authorityYear="2020" class="Insecta" family="Tingidae" genus="Heissiella" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="383" phylum="Arthropoda" rank="species" species="donguri" status="sp. nov.">
<emphasis id="B97DEAAD0F712D04FA87F8F4FEA83C67" italics="true" pageId="6" pageNumber="383">Heissiella donguri</emphasis>
</taxonomicName>
<emphasis id="B97DEAAD0F712D04FEDFF8D7FE0B3C67" bold="true" box="[312,400,1827,1853]" pageId="6" pageNumber="383">
<taxonomicNameLabel id="A24E57D60F712D04FEDFF8D7FE0B3C67" box="[312,400,1827,1853]" pageId="6" pageNumber="383" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
represents the third record of the Japanese lace bugs feeding on 
<taxonomicName id="4C094D3C0F712D04FBB2F8D7FB283C67" box="[1109,1203,1827,1853]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="6" pageNumber="383" phylum="Tracheophyta" rank="genus">
<emphasis id="B97DEAAD0F712D04FBB2F8D7FB283C67" box="[1109,1203,1827,1853]" italics="true" pageId="6" pageNumber="383">Quercus</emphasis>
</taxonomicName>
species and suggests that far more unknown tingids may be newly found on the Japanese oaks with more extensive field research.
</paragraph>
<paragraph id="8BB636BF0F712D04FF20F89FFE503C97" blockId="6.[151,1437,1719,1997]" pageId="6" pageNumber="383">
This new species feeds on 
<taxonomicName id="4C094D3C0F712D04FE13F89FFDD53CDF" authorityName="Blume" authorityYear="1851" box="[500,590,1899,1925]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="6" pageNumber="383" phylum="Tracheophyta" rank="species" species="gilva">
<emphasis id="B97DEAAD0F712D04FE13F89FFD933CDF" box="[500,520,1899,1925]" italics="true" pageId="6" pageNumber="383">Q</emphasis>
. 
<emphasis id="B97DEAAD0F712D04FDF1F89FFDD53CDF" box="[534,590,1899,1925]" italics="true" pageId="6" pageNumber="383">gilva</emphasis>
</taxonomicName>
, and a number of individuals of 
<taxonomicName id="4C094D3C0F712D04FC5AF898FBAF3CDF" baseAuthorityName="Drake &amp; Poor" baseAuthorityYear="1936" box="[957,1076,1899,1925]" class="Insecta" family="Tingidae" genus="Physatocheila" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="383" phylum="Arthropoda" rank="species" species="dryadis">
<emphasis id="B97DEAAD0F712D04FC5AF898FC4A3CDF" box="[957,977,1900,1925]" italics="true" pageId="6" pageNumber="383">H</emphasis>
. 
<emphasis id="B97DEAAD0F712D04FC07F89FFBAF3CDF" box="[992,1076,1899,1925]" italics="true" pageId="6" pageNumber="383">dryadis</emphasis>
</taxonomicName>
were collected from 
<taxonomicName id="4C094D3C0F712D04FAC4F89FFE613CF3" authority="Lindl. ex Royle (Pericart 1984)" authorityName="Lindl. ex Royle (Pericart" authorityYear="1984" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="6" pageNumber="383" phylum="Tracheophyta" rank="species" species="dilatata">
<emphasis id="B97DEAAD0F712D04FAC4F89FFAAC3CDF" box="[1315,1335,1899,1925]" italics="true" pageId="6" pageNumber="383">Q</emphasis>
. 
<emphasis id="B97DEAAD0F712D04FAA2F89FFA073CDF" box="[1349,1436,1899,1925]" italics="true" pageId="6" pageNumber="383">dilatata</emphasis>
Lindl. ex Royle (
<bibRefCitation id="EF984B4E0F712D04FEB2F87BFE6A3CF3" author="Pericart, J." box="[341,497,1935,1961]" pageId="6" pageNumber="383" pagination="23 - 47" refId="ref5423" refString="Pericart, J. (1984) Tingidae nouveaux ou interessants du Nord du sous-continent indien (Hemiptera). I. Entomologica Basiliensia, 9, 23 - 47. [in French, English summary]" type="journal article" year="1984">Péricart 1984</bibRefCitation>
)
</taxonomicName>
. Therefore, unrecorded host plants for the remaining two described congeners may be 
<taxonomicName id="4C094D3C0F712D04FF5EF847FE8C3C97" box="[185,279,1971,1997]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="6" pageNumber="383" phylum="Tracheophyta" rank="genus">
<emphasis id="B97DEAAD0F712D04FF5EF847FE8C3C97" box="[185,279,1971,1997]" italics="true" pageId="6" pageNumber="383">Quercus</emphasis>
</taxonomicName>
species as well.
</paragraph>
</subSubSection>
</treatment>
</document>