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<document id="D464C82F81AB04B82A952324CDE31D9E" ID-CLB-Dataset="31881" ID-DOI="10.5852/ejt.2016.255" ID-GBIF-Dataset="2719d968-2e75-4336-8e59-4c9563abdda2" ID-Zenodo-Dep="831094" ID-ZooBank="http://zoobank.org/urn:lsid:zoobank.org:pub:0E8DA4DC-FF4C-436E-94FB-CB89F6416C6E" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1500286020512" checkinUser="jeremy" docAuthor="Nadine Duperre &amp; Elicio Tapia" docDate="2016" docId="F319EF09070AFB10FD267E7C888FC6D5" docLanguage="en" docName="Dupérré, N., _ Tapia, E. (2016). Overview of the Anyphaenids (Araneae, Anyphaeninae, Anyphaenidae) (Done).pdf.imf" docOrigin="European Jornal of Taxonomy 255" docSource="http://sciencepress.mnhn.fr/en/periodiques/european-journal-taxonomy/2016/overview-anyphaenids-araneae-anyphaeninae-anyphaenidae-spider-fauna-choco-forest-ecuador-description-thirteen-new-species" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Shuyushka Duperre &amp; Tapia, 2016, gen. nov." docType="treatment" docUuid="22C28CEB-BAE6-4902-8657-9A922D775514" docUuidSource="ZooBank" docVersion="13" lastPageNumber="22" masterDocId="0F209771071EFB05FFBB7F778B3DC26A" masterDocTitle="Overview of the Anyphaenids (Araneae, Anyphaeninae, Anyphaenidae) spider fauna from the Chocó forest of Ecuador, with the description of thirteen new species" masterLastPageNumber="50" masterPageNumber="1" pageNumber="21" updateTime="1698465609934" updateUser="ExternalLinkService">
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<mods:title id="3BAF2BAC27917D635F35D14FF922ACBD">Overview of the Anyphaenids (Araneae, Anyphaeninae, Anyphaenidae) spider fauna from the Chocó forest of Ecuador, with the description of thirteen new species</mods:title>
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<mods:namePart id="41A929579C339EF65F332964151EA61E">Nadine Duperre</mods:namePart>
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<mods:namePart id="ABD83679C0C7F5E1FD35FDAB193832A2">Elicio Tapia</mods:namePart>
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<mods:title id="4BA3D6C1763A23FAA8199A59D8897A85">European Jornal of Taxonomy</mods:title>
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<paragraph id="7B0F5E1F070AFB11FD267E7C88AAC34C" blockId="20.[370,1217,267,329]" box="[669,919,267,294]" pageId="20" pageNumber="21">
<heading id="2047E973070AFB11FD267E7C88AAC34C" box="[669,919,267,294]" centered="true" fontSize="11" level="5" pageId="20" pageNumber="21" reason="4">
<taxonomicName id="BCB0259C070AFB11FD267E7C8819C34C" ID-CoL="7GGX" box="[669,804,267,294]" class="Arachnida" family="Anyphaenidae" genus="Shuyushka" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="49C4820D070AFB11FD267E7C8819C34C" bold="true" box="[669,804,267,294]" italics="true" pageId="20" pageNumber="21">Shuyushka</emphasis>
</taxonomicName>
<taxonomicNameLabel id="52F73F76070AFB11FC977E7B88AAC34C" box="[812,919,268,294]" pageId="20" pageNumber="21" rank="genus">gen. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="7B0F5E1F070AFB11FEC97E598FFCC323" blockId="20.[370,1217,267,329]" box="[370,1217,302,329]" pageId="20" pageNumber="21">urn:lsid:zoobank.org:act:22C28CEB-BAE6-4902-8657-9A922D775514</paragraph>
<paragraph id="7B0F5E1F070AFB11FF067E078A65C3E0" blockId="20.[189,575,368,441]" box="[189,344,368,394]" pageId="20" pageNumber="21">
<heading id="2047E973070AFB11FF067E078A65C3E0" bold="true" box="[189,344,368,394]" fontSize="11" level="3" pageId="20" pageNumber="21" reason="6">
<emphasis id="49C4820D070AFB11FF067E078A65C3E0" bold="true" box="[189,344,368,394]" pageId="20" pageNumber="21">
<typeStatus id="A40BE0BD070AFB11FF067E078BC7C3E0" box="[189,250,368,394]" pageId="20" pageNumber="21">Type</typeStatus>
species
</emphasis>
</heading>
</paragraph>
<paragraph id="7B0F5E1F070AFB11FF067EE98902C3D3" blockId="20.[189,575,368,441]" box="[189,575,414,441]" pageId="20" pageNumber="21">
<taxonomicName id="BCB0259C070AFB11FF067EE98AB5C3D2" box="[189,392,414,440]" class="Arachnida" family="Anyphaenidae" genus="Shuyushka" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="wachi">
<emphasis id="49C4820D070AFB11FF067EE98AB5C3D2" box="[189,392,414,440]" italics="true" pageId="20" pageNumber="21">Shuyushka wachi</emphasis>
</taxonomicName>
gen. et sp. nov.
</paragraph>
<subSubSection id="33AA0D94070AFB11FF067E9788CEC096" pageId="20" pageNumber="21" type="diagnosis">
<paragraph id="7B0F5E1F070AFB11FF067E978A0BC390" blockId="20.[189,1399,480,764]" box="[189,310,480,506]" pageId="20" pageNumber="21">
<heading id="2047E973070AFB11FF067E978A0BC390" bold="true" box="[189,310,480,506]" fontSize="11" level="3" pageId="20" pageNumber="21" reason="6">
<emphasis id="49C4820D070AFB11FF067E978A0BC390" bold="true" box="[189,310,480,506]" pageId="20" pageNumber="21">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="7B0F5E1F070AFB11FF067D7988CEC096" blockId="20.[189,1399,480,764]" pageId="20" pageNumber="21">
<taxonomicName id="BCB0259C070AFB11FF067D798A07C042" box="[189,314,526,552]" class="Arachnida" family="Anyphaenidae" genus="Shuyushka" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="49C4820D070AFB11FF067D798A07C042" box="[189,314,526,552]" italics="true" pageId="20" pageNumber="21">Shuyushka</emphasis>
</taxonomicName>
<taxonomicNameLabel id="52F73F76070AFB11FEFD7D788A8BC043" box="[326,438,527,553]" pageId="20" pageNumber="21" rank="genus">gen. nov.</taxonomicNameLabel>
can be distinguished from other Anyphaeninae genera by the presence of the following characters: tracheal spiracle in the middle of abdomen; lateral margin of endites concave; posterior eye row procurved; prolateral apex of tibia I with patch of short setae (
<figureCitation id="E38B429A070AFB11FBF27D228F9DC005" box="[1097,1184,597,623]" captionStart="Figs 37 – 40" captionStartId="28.[189,241,1964,1990]" captionTargetBox="[222,1369,267,1916]" captionTargetId="figure@28.[208,1381,257,1924]" captionTargetPageId="28" captionText="Figs 37 – 40. Shuyushka achachay gen. et sp. nov. 37 .  Ƌ,   abdomen,  dorsal  view.   38 .  Ƌ,   palp,  ventral  view.   39. Ƌ,   palp,  retrolateral  view. 40 .  Ƌ,  tibia  I,  prolateral  view.  Scale  bars:   38 – 39  =  0.05   mm;   37,   40  =  1   mm.  " httpUri="https://zenodo.org/record/1038252/files/figure.png" pageId="20" pageNumber="21">Fig. 40</figureCitation>
). From
<emphasis id="49C4820D070AFB11FB4D7D228E4AC005" box="[1270,1399,597,623]" italics="true" pageId="20" pageNumber="21">WulFlopsis</emphasis>
Soares &amp; Camargo, 1955 by the presence of ventral patellar apophysis (
<figureCitation id="E38B429A070AFB11FBAC7D0F8F49C0F8" box="[1047,1140,632,658]" captionStart="Figs 27 – 31" captionStartId="22.[189,241,1930,1956]" captionTargetBox="[229,1361,257,1892]" captionTargetId="figure@22.[229,1361,257,1892]" captionTargetPageId="22" captionText="Figs 27 – 31. Shuyushka wachi gen. et sp. nov. 27 .  Ƌ,   habitus,  dorsal  view.   28 .  Ƌ,   palp,  ventral  view.   29 .  Ƌ,   palp,  retrolateral  view. 30 .  ♀,   epigynum,  ventral  view.   31 .  ♀,  internal  genitalia,  dorsal  view.  Scale  bars: 28 – 31 = 0.1 mm; 27 = 1 mm." httpUri="https://zenodo.org/record/1038248/files/figure.png" pageId="20" pageNumber="21">Figs 29</figureCitation>
,
<figureCitation id="E38B429A070AFB11FB397D0F8F9FC0F8" box="[1154,1186,632,658]" captionStart="Figs 32 – 36" captionStartId="25.[189,241,1928,1954]" captionTargetBox="[254,1331,261,1866]" captionTargetId="figure@25.[239,1344,251,1880]" captionTargetPageId="25" captionText="Figs 32 – 36. Shuyushka moscai gen. et sp. nov. 32 .  Ƌ,   abdomen,  dorsal  view.   33 .  Ƌ,   palp,  ventral  view.   34. Ƌ,   palp,  retrolateral  view. 35 .  ♀,   epigynum,  ventral  view.   36 .  ♀,  internal  genitalia,  dorsal  view.  Scale  bars: 33 – 36 = 0.1 mm; 32 = 1 mm." httpUri="https://zenodo.org/record/1038250/files/figure.png" pageId="20" pageNumber="21">34</figureCitation>
,
<figureCitation id="E38B429A070AFB11FB147D0F8FF3C0F8" box="[1199,1230,632,658]" captionStart="Figs 37 – 40" captionStartId="28.[189,241,1964,1990]" captionTargetBox="[222,1369,267,1916]" captionTargetId="figure@28.[208,1381,257,1924]" captionTargetPageId="28" captionText="Figs 37 – 40. Shuyushka achachay gen. et sp. nov. 37 .  Ƌ,   abdomen,  dorsal  view.   38 .  Ƌ,   palp,  ventral  view.   39. Ƌ,   palp,  retrolateral  view. 40 .  Ƌ,  tibia  I,  prolateral  view.  Scale  bars:   38 – 39  =  0.05   mm;   37,   40  =  1   mm.  " httpUri="https://zenodo.org/record/1038252/files/figure.png" pageId="20" pageNumber="21">39</figureCitation>
) absent in the latter genera; from
<taxonomicName id="BCB0259C070AFB11FE187DEC8984C0DF" authority="Brescovit, 1997" authorityName="Brescovit" authorityYear="1997" box="[419,697,667,693]" class="Arachnida" family="Anyphaenidae" genus="Thaloe" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis id="49C4820D070AFB11FE187DEC8AC9C0DF" box="[419,500,667,693]" italics="true" pageId="20" pageNumber="21">Thaloe</emphasis>
Brescovit, 1997
</taxonomicName>
by the absence of medial projections on the lateral margin of the endites, present in the latter (Brescovit 1997: Fg. 41). Females of
<taxonomicName id="BCB0259C070AFB11FBA47DC98FA1C0B2" box="[1055,1180,702,728]" class="Arachnida" family="Anyphaenidae" genus="Shuyushka" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="49C4820D070AFB11FBA47DC98FA1C0B2" box="[1055,1180,702,728]" italics="true" pageId="20" pageNumber="21">Shuyushka</emphasis>
</taxonomicName>
<taxonomicNameLabel id="52F73F76070AFB11FB1D7DC88E2FC0B3" box="[1190,1298,703,729]" pageId="20" pageNumber="21" rank="genus">gen. nov.</taxonomicNameLabel>
are also distinguished by the absence of lateral epigynal grooves (
<figureCitation id="E38B429A070AFB11FCE57D958884C096" box="[862,953,738,764]" captionStart="Figs 27 – 31" captionStartId="22.[189,241,1930,1956]" captionTargetBox="[229,1361,257,1892]" captionTargetId="figure@22.[229,1361,257,1892]" captionTargetPageId="22" captionText="Figs 27 – 31. Shuyushka wachi gen. et sp. nov. 27 .  Ƌ,   habitus,  dorsal  view.   28 .  Ƌ,   palp,  ventral  view.   29 .  Ƌ,   palp,  retrolateral  view. 30 .  ♀,   epigynum,  ventral  view.   31 .  ♀,  internal  genitalia,  dorsal  view.  Scale  bars: 28 – 31 = 0.1 mm; 27 = 1 mm." httpUri="https://zenodo.org/record/1038248/files/figure.png" pageId="20" pageNumber="21">Figs 30</figureCitation>
,
<figureCitation id="E38B429A070AFB11FC7E7D9688DEC096" box="[965,995,737,764]" captionStart="Figs 32 – 36" captionStartId="25.[189,241,1928,1954]" captionTargetBox="[254,1331,261,1866]" captionTargetId="figure@25.[239,1344,251,1880]" captionTargetPageId="25" captionText="Figs 32 – 36. Shuyushka moscai gen. et sp. nov. 32 .  Ƌ,   abdomen,  dorsal  view.   33 .  Ƌ,   palp,  ventral  view.   34. Ƌ,   palp,  retrolateral  view. 35 .  ♀,   epigynum,  ventral  view.   36 .  ♀,  internal  genitalia,  dorsal  view.  Scale  bars: 33 – 36 = 0.1 mm; 32 = 1 mm." httpUri="https://zenodo.org/record/1038250/files/figure.png" pageId="20" pageNumber="21">35</figureCitation>
).
</paragraph>
</subSubSection>
<paragraph id="7B0F5E1F070AFB11FF067C548A79C157" blockId="20.[189,324,803,829]" box="[189,324,803,829]" pageId="20" pageNumber="21">
<heading id="2047E973070AFB11FF067C548A79C157" bold="true" box="[189,324,803,829]" fontSize="11" level="3" pageId="20" pageNumber="21" reason="2">
<emphasis id="49C4820D070AFB11FF067C548A79C157" bold="true" box="[189,324,803,829]" pageId="20" pageNumber="21">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="7B0F5E1F070AFB11FF067C258973C1E5" blockId="20.[189,1399,850,911]" pageId="20" pageNumber="21">The generic name is taken from the Kichwa language meaning ‘spotted’ for the dark marks on the abdomen. The gender is feminine.</paragraph>
<subSubSection id="33AA0D94070AFB11FF067CC18FCEC195" box="[189,1267,950,1023]" pageId="20" pageNumber="21" type="description">
<paragraph id="7B0F5E1F070AFB11FF067CC18A60C1BA" blockId="20.[189,349,950,976]" box="[189,349,950,976]" pageId="20" pageNumber="21">
<heading id="2047E973070AFB11FF067CC18A60C1BA" bold="true" box="[189,349,950,976]" fontSize="11" level="3" pageId="20" pageNumber="21" reason="2">
<emphasis id="49C4820D070AFB11FF067CC18A60C1BA" bold="true" box="[189,349,950,976]" pageId="20" pageNumber="21">Composition</emphasis>
</heading>
</paragraph>
<paragraph id="7B0F5E1F070AFB11FF067C928FCEC195" blockId="20.[189,1267,997,1023]" box="[189,1267,997,1023]" pageId="20" pageNumber="21">
<taxonomicName id="BCB0259C070AFB11FF067C928A8FC195" box="[189,434,997,1023]" class="Arachnida" family="Anyphaenidae" genus="Shuyushka" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="achachay">
<emphasis id="49C4820D070AFB11FF067C928A8FC195" box="[189,434,997,1023]" italics="true" pageId="20" pageNumber="21">Shuyushka achachay</emphasis>
</taxonomicName>
gen. et sp. nov.,
<taxonomicName id="BCB0259C070AFB11FDC37C9289D5C195" box="[632,744,997,1023]" class="Arachnida" family="Anyphaenidae" genus="Shuyushka" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="moscai">
<emphasis id="49C4820D070AFB11FDC37C9289D5C195" box="[632,744,997,1023]" italics="true" pageId="20" pageNumber="21">S. moscai</emphasis>
</taxonomicName>
gen. et sp. nov. and
<taxonomicName id="BCB0259C070AFB11FC637C928F01C195" box="[984,1084,997,1023]" class="Arachnida" family="Anyphaenidae" genus="Shuyushka" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="wachi">
<emphasis id="49C4820D070AFB11FC637C928F01C195" box="[984,1084,997,1023]" italics="true" pageId="20" pageNumber="21">S. wachi</emphasis>
</taxonomicName>
gen. et sp. nov.
</paragraph>
</subSubSection>
<subSubSection id="33AA0D94070AFB11FF067B518A18C605" box="[189,342,1062,1135]" pageId="20" pageNumber="21" type="distribution">
<paragraph id="7B0F5E1F070AFB11FF067B518A6BC62A" blockId="20.[189,342,1062,1088]" box="[189,342,1062,1088]" pageId="20" pageNumber="21">
<heading id="2047E973070AFB11FF067B518A6BC62A" bold="true" box="[189,342,1062,1088]" fontSize="11" level="3" pageId="20" pageNumber="21" reason="2">
<emphasis id="49C4820D070AFB11FF067B518A6BC62A" bold="true" box="[189,342,1062,1088]" pageId="20" pageNumber="21">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="7B0F5E1F070AFB11FF067B228A18C605" blockId="20.[189,293,1109,1135]" box="[189,293,1109,1135]" pageId="20" pageNumber="21">Ecuador.</paragraph>
</subSubSection>
<subSubSection id="33AA0D94070AFB10FF067BE1888FC6D5" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21" type="description">
<paragraph id="7B0F5E1F070AFB11FF067BE18A73C6DA" blockId="20.[189,334,1174,1200]" box="[189,334,1174,1200]" pageId="20" pageNumber="21">
<heading id="2047E973070AFB11FF067BE18A73C6DA" bold="true" box="[189,334,1174,1200]" fontSize="11" level="3" pageId="20" pageNumber="21" reason="2">
<emphasis id="49C4820D070AFB11FF067BE18A73C6DA" bold="true" box="[189,334,1174,1200]" pageId="20" pageNumber="21">Description</emphasis>
</heading>
</paragraph>
<paragraph id="7B0F5E1F070AFB11FF517BB28A17C6B5" blockId="20.[189,798,1221,1283]" box="[234,298,1221,1247]" pageId="20" pageNumber="21">
<heading id="2047E973070AFB11FF517BB28A17C6B5" bold="true" box="[234,298,1221,1247]" fontSize="11" level="3" pageId="20" pageNumber="21" reason="6">
<emphasis id="49C4820D070AFB11FF517BB28A17C6B5" bold="true" box="[234,298,1221,1247]" pageId="20" pageNumber="21">Male</emphasis>
</heading>
</paragraph>
<paragraph id="7B0F5E1F070AFB11FF067B9E8823C768" blockId="20.[189,798,1221,1283]" box="[189,798,1256,1283]" pageId="20" pageNumber="21">MEASUREMENTS. Medium (5.9–7.8 mm) size spiders.</paragraph>
<paragraph id="7B0F5E1F070AFB11FF067A5D8E27C7C4" blockId="20.[189,1399,1321,1454]" pageId="20" pageNumber="21">
CEPHALOTHORAX. Sub-rectangular, narrower anteriorly, slightly longer than wide, pars cephalica not elevated; longitudinal fovea short and shallow (
<figureCitation id="E38B429A070AFB11FD547A3A8875C70D" box="[751,840,1357,1383]" captionStart="Figs 27 – 31" captionStartId="22.[189,241,1930,1956]" captionTargetBox="[229,1361,257,1892]" captionTargetId="figure@22.[229,1361,257,1892]" captionTargetPageId="22" captionText="Figs 27 – 31. Shuyushka wachi gen. et sp. nov. 27 .  Ƌ,   habitus,  dorsal  view.   28 .  Ƌ,   palp,  ventral  view.   29 .  Ƌ,   palp,  retrolateral  view. 30 .  ♀,   epigynum,  ventral  view.   31 .  ♀,  internal  genitalia,  dorsal  view.  Scale  bars: 28 – 31 = 0.1 mm; 27 = 1 mm." httpUri="https://zenodo.org/record/1038248/files/figure.png" pageId="20" pageNumber="21">Fig. 27</figureCitation>
). Chilum present, trapezoidal. Clypeus low (
<date id="0F0E78DF070AFB11FAE27A3A8E4BC70D" box="[1369,1398,1357,1383]" pageId="20" pageNumber="21">1x</date>
AME). Endites longer than wide, lateral margin concave medially. Labium longer than wide, constricted submedially, apex excavated. Sternum oval, Fat, longer than wide. Precoxal triangular present.
</paragraph>
<paragraph id="7B0F5E1F070AFB11FF067AA28FF9C784" blockId="20.[189,1220,1492,1519]" box="[189,1220,1492,1519]" pageId="20" pageNumber="21">CHELICERAE. Oblique and robuste, geniculate, promargin and retromargin with 4-5 teeth.</paragraph>
<paragraph id="7B0F5E1F070AFB11FF0679618E10C41C" blockId="20.[189,1399,1558,1655]" pageId="20" pageNumber="21">
EYES. Eight eyes; AME smallest, less than half the diameter of the others; LE-PME about the same size; AME touching; AME-ALE almost touching, LE contiguous, LE-PME separated by half their width; PME separated by half their width; posterior eye row slightly procurved in dorsal view (
<figureCitation id="E38B429A070AFB11FB73792B8E1DC41C" box="[1224,1312,1628,1654]" captionStart="Figs 27 – 31" captionStartId="22.[189,241,1930,1956]" captionTargetBox="[229,1361,257,1892]" captionTargetId="figure@22.[229,1361,257,1892]" captionTargetPageId="22" captionText="Figs 27 – 31. Shuyushka wachi gen. et sp. nov. 27 .  Ƌ,   habitus,  dorsal  view.   28 .  Ƌ,   palp,  ventral  view.   29 .  Ƌ,   palp,  retrolateral  view. 30 .  ♀,   epigynum,  ventral  view.   31 .  ♀,  internal  genitalia,  dorsal  view.  Scale  bars: 28 – 31 = 0.1 mm; 27 = 1 mm." httpUri="https://zenodo.org/record/1038248/files/figure.png" pageId="20" pageNumber="21">Fig. 27</figureCitation>
).
</paragraph>
<paragraph id="7B0F5E1F070AFB11FF0679E988D1C494" blockId="20.[189,1398,1694,1791]" pageId="20" pageNumber="21">
LEGS. Trochanters notched; scopula sparse, entire on metatarsi and tarsi I–II; absent on metatarsi III–IV, entire on tarsi III–IV; apex of metatarsi III–IV with apical brush; prolateral apex of tibia I with patch of short setae (
<figureCitation id="E38B429A070AFB11FEF179938A9EC494" box="[330,419,1764,1790]" captionStart="Figs 37 – 40" captionStartId="28.[189,241,1964,1990]" captionTargetBox="[222,1369,267,1916]" captionTargetId="figure@28.[208,1381,257,1924]" captionTargetPageId="28" captionText="Figs 37 – 40. Shuyushka achachay gen. et sp. nov. 37 .  Ƌ,   abdomen,  dorsal  view.   38 .  Ƌ,   palp,  ventral  view.   39. Ƌ,   palp,  retrolateral  view. 40 .  Ƌ,  tibia  I,  prolateral  view.  Scale  bars:   38 – 39  =  0.05   mm;   37,   40  =  1   mm.  " httpUri="https://zenodo.org/record/1038252/files/figure.png" pageId="20" pageNumber="21">Fig. 40</figureCitation>
). Tarsal claws with 5–6 teeth. Legs formula 1423.
</paragraph>
<paragraph id="7B0F5E1F070AFB11FF067851880FC509" blockId="20.[189,1398,1829,1891]" pageId="20" pageNumber="21">
LEGS SPINATION RECURRENCE. Femora I–IV d1-1-1, r0-1-1; patellae III–IV r1; tibiae III–IV d1-0-0,
<date id="0F0E78DF070AFB11FA9B78518E64C52A" box="[1312,1369,1830,1856]" pageId="20" pageNumber="21">v2-2</date>
-2; metatarsi I–II
<date id="0F0E78DF070AFB11FED8783E8AA4C509" box="[355,409,1865,1891]" pageId="20" pageNumber="21">v2-1</date>
-0, r0-1-0, metatarsi III–IV
<date id="0F0E78DF070AFB11FD66783E882BC509" box="[733,790,1865,1891]" pageId="20" pageNumber="21">v1-1</date>
-1.
</paragraph>
<paragraph id="7B0F5E1F070AFB11FF0678FD8A2BC581" blockId="20.[189,1398,1930,2027]" pageId="20" pageNumber="21">ABDOMEN. Elongated cylindrical, covered with a mix of pale and dark short, none-erected setae and a few dark erected setae; tracheal spiracle recurved, situated in the middle between spinnerets and epigastric groove.</paragraph>
<paragraph id="7B0F5E1F070BFB10FF067E7C8F3AC393" blockId="21.[189,1399,267,505]" pageId="21" pageNumber="22">
GENITALIA. Palpal femur without apophysis. Palpal patella with two ventral apophyses, and one dorsal apophysis (
<figureCitation id="E38B429A070BFB10FEFA7E598AA6C323" box="[321,411,302,329]" captionStart="Figs 27 – 31" captionStartId="22.[189,241,1930,1956]" captionTargetBox="[229,1361,257,1892]" captionTargetId="figure@22.[229,1361,257,1892]" captionTargetPageId="22" captionText="Figs 27 – 31. Shuyushka wachi gen. et sp. nov. 27 .  Ƌ,   habitus,  dorsal  view.   28 .  Ƌ,   palp,  ventral  view.   29 .  Ƌ,   palp,  retrolateral  view. 30 .  ♀,   epigynum,  ventral  view.   31 .  ♀,  internal  genitalia,  dorsal  view.  Scale  bars: 28 – 31 = 0.1 mm; 27 = 1 mm." httpUri="https://zenodo.org/record/1038248/files/figure.png" pageId="21" pageNumber="22">Figs 29</figureCitation>
,
<figureCitation id="E38B429A070BFB10FE1D7E588AFBC323" box="[422,454,303,329]" captionStart="Figs 32 – 36" captionStartId="25.[189,241,1928,1954]" captionTargetBox="[254,1331,261,1866]" captionTargetId="figure@25.[239,1344,251,1880]" captionTargetPageId="25" captionText="Figs 32 – 36. Shuyushka moscai gen. et sp. nov. 32 .  Ƌ,   abdomen,  dorsal  view.   33 .  Ƌ,   palp,  ventral  view.   34. Ƌ,   palp,  retrolateral  view. 35 .  ♀,   epigynum,  ventral  view.   36 .  ♀,  internal  genitalia,  dorsal  view.  Scale  bars: 33 – 36 = 0.1 mm; 32 = 1 mm." httpUri="https://zenodo.org/record/1038250/files/figure.png" pageId="21" pageNumber="22">34</figureCitation>
,
<figureCitation id="E38B429A070BFB10FE6A7E588ACFC323" box="[465,498,303,329]" captionStart="Figs 37 – 40" captionStartId="28.[189,241,1964,1990]" captionTargetBox="[222,1369,267,1916]" captionTargetId="figure@28.[208,1381,257,1924]" captionTargetPageId="28" captionText="Figs 37 – 40. Shuyushka achachay gen. et sp. nov. 37 .  Ƌ,   abdomen,  dorsal  view.   38 .  Ƌ,   palp,  ventral  view.   39. Ƌ,   palp,  retrolateral  view. 40 .  Ƌ,  tibia  I,  prolateral  view.  Scale  bars:   38 – 39  =  0.05   mm;   37,   40  =  1   mm.  " httpUri="https://zenodo.org/record/1038252/files/figure.png" pageId="21" pageNumber="22">39</figureCitation>
). Palpal tibia short, shorter than cymbium; with (
<figureCitation id="E38B429A070BFB10FB927E598FBEC323" box="[1065,1155,302,329]" captionStart="Figs 27 – 31" captionStartId="22.[189,241,1930,1956]" captionTargetBox="[229,1361,257,1892]" captionTargetId="figure@22.[229,1361,257,1892]" captionTargetPageId="22" captionText="Figs 27 – 31. Shuyushka wachi gen. et sp. nov. 27 .  Ƌ,   habitus,  dorsal  view.   28 .  Ƌ,   palp,  ventral  view.   29 .  Ƌ,   palp,  retrolateral  view. 30 .  ♀,   epigynum,  ventral  view.   31 .  ♀,  internal  genitalia,  dorsal  view.  Scale  bars: 28 – 31 = 0.1 mm; 27 = 1 mm." httpUri="https://zenodo.org/record/1038248/files/figure.png" pageId="21" pageNumber="22">Figs 29</figureCitation>
,
<figureCitation id="E38B429A070BFB10FB357E588F90C323" box="[1166,1197,303,329]" captionStart="Figs 32 – 36" captionStartId="25.[189,241,1928,1954]" captionTargetBox="[254,1331,261,1866]" captionTargetId="figure@25.[239,1344,251,1880]" captionTargetPageId="25" captionText="Figs 32 – 36. Shuyushka moscai gen. et sp. nov. 32 .  Ƌ,   abdomen,  dorsal  view.   33 .  Ƌ,   palp,  ventral  view.   34. Ƌ,   palp,  retrolateral  view. 35 .  ♀,   epigynum,  ventral  view.   36 .  ♀,  internal  genitalia,  dorsal  view.  Scale  bars: 33 – 36 = 0.1 mm; 32 = 1 mm." httpUri="https://zenodo.org/record/1038250/files/figure.png" pageId="21" pageNumber="22">34</figureCitation>
) or without blunt ventral apophysis (
<figureCitation id="E38B429A070BFB10FE247E258AC5C306" box="[415,504,338,364]" captionStart="Figs 37 – 40" captionStartId="28.[189,241,1964,1990]" captionTargetBox="[222,1369,267,1916]" captionTargetId="figure@28.[208,1381,257,1924]" captionTargetPageId="28" captionText="Figs 37 – 40. Shuyushka achachay gen. et sp. nov. 37 .  Ƌ,   abdomen,  dorsal  view.   38 .  Ƌ,   palp,  ventral  view.   39. Ƌ,   palp,  retrolateral  view. 40 .  Ƌ,  tibia  I,  prolateral  view.  Scale  bars:   38 – 39  =  0.05   mm;   37,   40  =  1   mm.  " httpUri="https://zenodo.org/record/1038252/files/figure.png" pageId="21" pageNumber="22">Fig. 39</figureCitation>
); retrolateral tibial apophysis bipartite, anterior part globular (
<figureCitation id="E38B429A070BFB10FB667E258E07C306" box="[1245,1338,338,364]" captionStart="Figs 27 – 31" captionStartId="22.[189,241,1930,1956]" captionTargetBox="[229,1361,257,1892]" captionTargetId="figure@22.[229,1361,257,1892]" captionTargetPageId="22" captionText="Figs 27 – 31. Shuyushka wachi gen. et sp. nov. 27 .  Ƌ,   habitus,  dorsal  view.   28 .  Ƌ,   palp,  ventral  view.   29 .  Ƌ,   palp,  retrolateral  view. 30 .  ♀,   epigynum,  ventral  view.   31 .  ♀,  internal  genitalia,  dorsal  view.  Scale  bars: 28 – 31 = 0.1 mm; 27 = 1 mm." httpUri="https://zenodo.org/record/1038248/files/figure.png" pageId="21" pageNumber="22">Figs 29</figureCitation>
,
<figureCitation id="E38B429A070BFB10FAF37E258E55C306" box="[1352,1384,338,364]" captionStart="Figs 37 – 40" captionStartId="28.[189,241,1964,1990]" captionTargetBox="[222,1369,267,1916]" captionTargetId="figure@28.[208,1381,257,1924]" captionTargetPageId="28" captionText="Figs 37 – 40. Shuyushka achachay gen. et sp. nov. 37 .  Ƌ,   abdomen,  dorsal  view.   38 .  Ƌ,   palp,  ventral  view.   39. Ƌ,   palp,  retrolateral  view. 40 .  Ƌ,  tibia  I,  prolateral  view.  Scale  bars:   38 – 39  =  0.05   mm;   37,   40  =  1   mm.  " httpUri="https://zenodo.org/record/1038252/files/figure.png" pageId="21" pageNumber="22">39</figureCitation>
), triangular (
<figureCitation id="E38B429A070BFB10FEFB7E028AABC3E5" box="[320,406,373,399]" captionStart="Figs 32 – 36" captionStartId="25.[189,241,1928,1954]" captionTargetBox="[254,1331,261,1866]" captionTargetId="figure@25.[239,1344,251,1880]" captionTargetPageId="25" captionText="Figs 32 – 36. Shuyushka moscai gen. et sp. nov. 32 .  Ƌ,   abdomen,  dorsal  view.   33 .  Ƌ,   palp,  ventral  view.   34. Ƌ,   palp,  retrolateral  view. 35 .  ♀,   epigynum,  ventral  view.   36 .  ♀,  internal  genitalia,  dorsal  view.  Scale  bars: 33 – 36 = 0.1 mm; 32 = 1 mm." httpUri="https://zenodo.org/record/1038250/files/figure.png" pageId="21" pageNumber="22">Fig. 34</figureCitation>
) and less sclerotized, posterior part well sclerotized, elongated (
<figureCitation id="E38B429A070BFB10FB307E028FDAC3E5" box="[1163,1255,373,399]" captionStart="Figs 27 – 31" captionStartId="22.[189,241,1930,1956]" captionTargetBox="[229,1361,257,1892]" captionTargetId="figure@22.[229,1361,257,1892]" captionTargetPageId="22" captionText="Figs 27 – 31. Shuyushka wachi gen. et sp. nov. 27 .  Ƌ,   habitus,  dorsal  view.   28 .  Ƌ,   palp,  ventral  view.   29 .  Ƌ,   palp,  retrolateral  view. 30 .  ♀,   epigynum,  ventral  view.   31 .  ♀,  internal  genitalia,  dorsal  view.  Scale  bars: 28 – 31 = 0.1 mm; 27 = 1 mm." httpUri="https://zenodo.org/record/1038248/files/figure.png" pageId="21" pageNumber="22">Figs 29</figureCitation>
,
<figureCitation id="E38B429A070BFB10FB487E028E2FC3E5" box="[1267,1298,373,399]" captionStart="Figs 32 – 36" captionStartId="25.[189,241,1928,1954]" captionTargetBox="[254,1331,261,1866]" captionTargetId="figure@25.[239,1344,251,1880]" captionTargetPageId="25" captionText="Figs 32 – 36. Shuyushka moscai gen. et sp. nov. 32 .  Ƌ,   abdomen,  dorsal  view.   33 .  Ƌ,   palp,  ventral  view.   34. Ƌ,   palp,  retrolateral  view. 35 .  ♀,   epigynum,  ventral  view.   36 .  ♀,  internal  genitalia,  dorsal  view.  Scale  bars: 33 – 36 = 0.1 mm; 32 = 1 mm." httpUri="https://zenodo.org/record/1038250/files/figure.png" pageId="21" pageNumber="22">34</figureCitation>
) or with short spine (
<figureCitation id="E38B429A070BFB10FEF57EEF8A9BC3D8" box="[334,422,408,434]" captionStart="Figs 37 – 40" captionStartId="28.[189,241,1964,1990]" captionTargetBox="[222,1369,267,1916]" captionTargetId="figure@28.[208,1381,257,1924]" captionTargetPageId="28" captionText="Figs 37 – 40. Shuyushka achachay gen. et sp. nov. 37 .  Ƌ,   abdomen,  dorsal  view.   38 .  Ƌ,   palp,  ventral  view.   39. Ƌ,   palp,  retrolateral  view. 40 .  Ƌ,  tibia  I,  prolateral  view.  Scale  bars:   38 – 39  =  0.05   mm;   37,   40  =  1   mm.  " httpUri="https://zenodo.org/record/1038252/files/figure.png" pageId="21" pageNumber="22">Fig. 39</figureCitation>
). Subtegulum rounded apically; tegulum rounded basally; median apophysis large, hook-shaped; embolus black, heavily sclerotized, prolaterally positioned, wide basally, distally with small triangular extensions and pale terminal segment (
<figureCitation id="E38B429A070BFB10FCFE7EA9889DC393" box="[837,928,478,505]" captionStart="Figs 27 – 31" captionStartId="22.[189,241,1930,1956]" captionTargetBox="[229,1361,257,1892]" captionTargetId="figure@22.[229,1361,257,1892]" captionTargetPageId="22" captionText="Figs 27 – 31. Shuyushka wachi gen. et sp. nov. 27 .  Ƌ,   habitus,  dorsal  view.   28 .  Ƌ,   palp,  ventral  view.   29 .  Ƌ,   palp,  retrolateral  view. 30 .  ♀,   epigynum,  ventral  view.   31 .  ♀,  internal  genitalia,  dorsal  view.  Scale  bars: 28 – 31 = 0.1 mm; 27 = 1 mm." httpUri="https://zenodo.org/record/1038248/files/figure.png" pageId="21" pageNumber="22">Figs 28</figureCitation>
,
<figureCitation id="E38B429A070BFB10FC177EA888F1C393" box="[940,972,479,505]" captionStart="Figs 32 – 36" captionStartId="25.[189,241,1928,1954]" captionTargetBox="[254,1331,261,1866]" captionTargetId="figure@25.[239,1344,251,1880]" captionTargetPageId="25" captionText="Figs 32 – 36. Shuyushka moscai gen. et sp. nov. 32 .  Ƌ,   abdomen,  dorsal  view.   33 .  Ƌ,   palp,  ventral  view.   34. Ƌ,   palp,  retrolateral  view. 35 .  ♀,   epigynum,  ventral  view.   36 .  ♀,  internal  genitalia,  dorsal  view.  Scale  bars: 33 – 36 = 0.1 mm; 32 = 1 mm." httpUri="https://zenodo.org/record/1038250/files/figure.png" pageId="21" pageNumber="22">33</figureCitation>
,
<figureCitation id="E38B429A070BFB10FC637EA888C4C393" box="[984,1017,479,505]" captionStart="Figs 37 – 40" captionStartId="28.[189,241,1964,1990]" captionTargetBox="[222,1369,267,1916]" captionTargetId="figure@28.[208,1381,257,1924]" captionTargetPageId="28" captionText="Figs 37 – 40. Shuyushka achachay gen. et sp. nov. 37 .  Ƌ,   abdomen,  dorsal  view.   38 .  Ƌ,   palp,  ventral  view.   39. Ƌ,   palp,  retrolateral  view. 40 .  Ƌ,  tibia  I,  prolateral  view.  Scale  bars:   38 – 39  =  0.05   mm;   37,   40  =  1   mm.  " httpUri="https://zenodo.org/record/1038252/files/figure.png" pageId="21" pageNumber="22">38</figureCitation>
).
</paragraph>
<paragraph id="7B0F5E1F070BFB10FF517D578A78C050" blockId="21.[189,798,544,606]" box="[234,325,544,570]" pageId="21" pageNumber="22">
<heading id="2047E973070BFB10FF517D578A78C050" bold="true" box="[234,325,544,570]" fontSize="11" level="3" pageId="21" pageNumber="22" reason="6">
<emphasis id="49C4820D070BFB10FF517D578A78C050" bold="true" box="[234,325,544,570]" pageId="21" pageNumber="22">Female</emphasis>
</heading>
</paragraph>
<paragraph id="7B0F5E1F070BFB10FF067D338823C037" blockId="21.[189,798,544,606]" box="[189,798,579,606]" pageId="21" pageNumber="22">MEASUREMENTS. Medium (5.6–5.7 mm) size spiders.</paragraph>
<paragraph id="7B0F5E1F070BFB10FF067DF28944C0F5" blockId="21.[189,633,645,671]" box="[189,633,645,671]" pageId="21" pageNumber="22">CEPHALOTHORAX AND EYES. As in male.</paragraph>
<paragraph id="7B0F5E1F070BFB10FF067DB08A72C120" blockId="21.[189,1399,710,842]" pageId="21" pageNumber="22">LEGS. As in male; somewhat shorter in relation to length of the body. Tarsi and metatarsi I–II with dense, entire scopula, metatarsi III–IV without scopula, tarsi III–IV with dense, entire scopula; metatarsi III–IV with apical brush; prolateral apex of tibia I with patch of short setae. Legs formula 4123. Palpal claws with 4 teeth.</paragraph>
<paragraph id="7B0F5E1F070BFB10FF067C058F42C1C4" blockId="21.[189,1399,881,943]" pageId="21" pageNumber="22">
LEGS SPINATION RECURRENCE. Femora I–IV d1-1-1; patellae III–IV r1; tibiae I–II
<date id="0F0E78DF070BFB10FBE37C068FACC1E1" box="[1112,1169,881,907]" pageId="21" pageNumber="22">v2-2</date>
-0; tibiae III–IV d1- 0-0, p1-1-1; metatarsi I–II
<date id="0F0E78DF070BFB10FE4C7CE3892AC1C4" box="[503,535,916,942]" pageId="21" pageNumber="22">v2</date>
-0-0, metatarsi III–IV d0-1-0,
<date id="0F0E78DF070BFB10FCCE7CE38890C1C4" box="[885,941,916,942]" pageId="21" pageNumber="22">v2-2</date>
-2, r1-1-1, p1-1-1.
</paragraph>
<paragraph id="7B0F5E1F070BFB10FF067CA18E09C679" blockId="21.[189,1399,982,1043]" pageId="21" pageNumber="22">ABDOMEN. Oval; covered with a mix of pale and dark short, none-erected setae and a few dark erected setae; tracheal spiracle recurved, situated in the middle between spinnerets and epigastric groove.</paragraph>
<paragraph id="7B0F5E1F070BFB10FF067B4D888FC6D5" blockId="21.[189,1406,1082,1215]" pageId="21" pageNumber="22">
GENITALIA. Epigynum well sclerotized with (
<figureCitation id="E38B429A070BFB10FD607B4D8809C63F" box="[731,820,1082,1109]" captionStart="Figs 32 – 36" captionStartId="25.[189,241,1928,1954]" captionTargetBox="[254,1331,261,1866]" captionTargetId="figure@25.[239,1344,251,1880]" captionTargetPageId="25" captionText="Figs 32 – 36. Shuyushka moscai gen. et sp. nov. 32 .  Ƌ,   abdomen,  dorsal  view.   33 .  Ƌ,   palp,  ventral  view.   34. Ƌ,   palp,  retrolateral  view. 35 .  ♀,   epigynum,  ventral  view.   36 .  ♀,  internal  genitalia,  dorsal  view.  Scale  bars: 33 – 36 = 0.1 mm; 32 = 1 mm." httpUri="https://zenodo.org/record/1038250/files/figure.png" pageId="21" pageNumber="22">Fig. 35</figureCitation>
) or without epigynal Fap (
<figureCitation id="E38B429A070BFB10FB387B4D8FE2C63E" box="[1155,1247,1082,1109]" captionStart="Figs 27 – 31" captionStartId="22.[189,241,1930,1956]" captionTargetBox="[229,1361,257,1892]" captionTargetId="figure@22.[229,1361,257,1892]" captionTargetPageId="22" captionText="Figs 27 – 31. Shuyushka wachi gen. et sp. nov. 27 .  Ƌ,   habitus,  dorsal  view.   28 .  Ƌ,   palp,  ventral  view.   29 .  Ƌ,   palp,  retrolateral  view. 30 .  ♀,   epigynum,  ventral  view.   31 .  ♀,  internal  genitalia,  dorsal  view.  Scale  bars: 28 – 31 = 0.1 mm; 27 = 1 mm." httpUri="https://zenodo.org/record/1038248/files/figure.png" pageId="21" pageNumber="22">Fig. 30</figureCitation>
); copulatory openings situated lateromedially (
<figureCitation id="E38B429A070BFB10FDD07B2A89ECC612" box="[619,721,1117,1144]" captionStart="Figs 27 – 31" captionStartId="22.[189,241,1930,1956]" captionTargetBox="[229,1361,257,1892]" captionTargetId="figure@22.[229,1361,257,1892]" captionTargetPageId="22" captionText="Figs 27 – 31. Shuyushka wachi gen. et sp. nov. 27 .  Ƌ,   habitus,  dorsal  view.   28 .  Ƌ,   palp,  ventral  view.   29 .  Ƌ,   palp,  retrolateral  view. 30 .  ♀,   epigynum,  ventral  view.   31 .  ♀,  internal  genitalia,  dorsal  view.  Scale  bars: 28 – 31 = 0.1 mm; 27 = 1 mm." httpUri="https://zenodo.org/record/1038248/files/figure.png" pageId="21" pageNumber="22">Figs 30</figureCitation>
,
<figureCitation id="E38B429A070BFB10FD537B2A8834C612" box="[744,777,1117,1144]" captionStart="Figs 32 – 36" captionStartId="25.[189,241,1928,1954]" captionTargetBox="[254,1331,261,1866]" captionTargetId="figure@25.[239,1344,251,1880]" captionTargetPageId="25" captionText="Figs 32 – 36. Shuyushka moscai gen. et sp. nov. 32 .  Ƌ,   abdomen,  dorsal  view.   33 .  Ƌ,   palp,  ventral  view.   34. Ƌ,   palp,  retrolateral  view. 35 .  ♀,   epigynum,  ventral  view.   36 .  ♀,  internal  genitalia,  dorsal  view.  Scale  bars: 33 – 36 = 0.1 mm; 32 = 1 mm." httpUri="https://zenodo.org/record/1038250/files/figure.png" pageId="21" pageNumber="22">35</figureCitation>
). Internal genitalia with large copulatory ducts; spermathecae not well deFned, seminal receptacles positioned at the middle of copulatory ducts course; fertilization ducts long and curved (
<figureCitation id="E38B429A070BFB10FDE47BD38989C6D4" box="[607,692,1188,1214]" captionStart="Figs 27 – 31" captionStartId="22.[189,241,1930,1956]" captionTargetBox="[229,1361,257,1892]" captionTargetId="figure@22.[229,1361,257,1892]" captionTargetPageId="22" captionText="Figs 27 – 31. Shuyushka wachi gen. et sp. nov. 27 .  Ƌ,   habitus,  dorsal  view.   28 .  Ƌ,   palp,  ventral  view.   29 .  Ƌ,   palp,  retrolateral  view. 30 .  ♀,   epigynum,  ventral  view.   31 .  ♀,  internal  genitalia,  dorsal  view.  Scale  bars: 28 – 31 = 0.1 mm; 27 = 1 mm." httpUri="https://zenodo.org/record/1038248/files/figure.png" pageId="21" pageNumber="22">Fig. 31</figureCitation>
) or straight (
<figureCitation id="E38B429A070BFB10FCF67BD38899C6D5" box="[845,932,1188,1215]" captionStart="Figs 32 – 36" captionStartId="25.[189,241,1928,1954]" captionTargetBox="[254,1331,261,1866]" captionTargetId="figure@25.[239,1344,251,1880]" captionTargetPageId="25" captionText="Figs 32 – 36. Shuyushka moscai gen. et sp. nov. 32 .  Ƌ,   abdomen,  dorsal  view.   33 .  Ƌ,   palp,  ventral  view.   34. Ƌ,   palp,  retrolateral  view. 35 .  ♀,   epigynum,  ventral  view.   36 .  ♀,  internal  genitalia,  dorsal  view.  Scale  bars: 33 – 36 = 0.1 mm; 32 = 1 mm." httpUri="https://zenodo.org/record/1038250/files/figure.png" pageId="21" pageNumber="22">Fig. 36</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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