185 lines
20 KiB
XML
185 lines
20 KiB
XML
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<mods:title id="1E7859FD8E8A73571140BA39029F9A38">A mountain of millipedes V: three new genera of Odontopygidae from the Udzungwa mountains, Tanzania (Diplopoda, Spirostreptida, Odontopygidae)</mods:title>
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<mods:namePart id="FA7475AC1274F118B0B3616C36FC221F">Enghoff, Henrik</mods:namePart>
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<mods:title id="279F02D4CB11E36591BFB20DA6B48D32">European Journal of Taxonomy</mods:title>
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<mods:part id="C15DF4EA04E8EA78FDA4E9C89891C798">
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<mods:date id="4E9EEE27B5835C0E1667E03E969A2FB8">2016</mods:date>
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<mods:number id="4351C86EE2CAE64E037731E585FAFBE0">2016-08-12</mods:number>
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<treatment id="8D3887BCFFE2FF9FDDFF4C428A542F2A" ID-DOI="http://doi.org/10.5281/zenodo.3853134" ID-GBIF-Taxon="164252619" ID-Zenodo-Dep="3853134" LSID="urn:lsid:zoobank.org:act:5071EA1A-F2FB-4A8C-AE0C-4F18EA27C973" httpUri="http://treatment.plazi.org/id/8D3887BCFFE2FF9FDDFF4C428A542F2A" lastPageNumber="4" pageId="3" pageNumber="4">
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<subSubSection id="4D8B6521FFE2FF9FDDFF4C428AD52A5B" pageId="3" pageNumber="4" type="nomenclature">
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<paragraph id="052E36AAFFE2FF9FDDFF4C428DBC2A3C" blockId="3.[359,1228,527,589]" box="[655,933,527,554]" pageId="3" pageNumber="4">
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<heading id="5E6681C6FFE2FF9FDDFF4C428DBC2A3C" box="[655,933,527,554]" centered="true" fontSize="11" level="2" pageId="3" pageNumber="4" reason="2">
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<taxonomicName id="C2914D29FFE2FF9FDDFF4C428D2B2A3F" ID-CoL="7NWFK" authority="Enghoff, 2016" authorityName="Enghoff" authorityYear="2016" box="[655,818,527,553]" class="Diplopoda" family="Odontopygidae" genus="Casuariverpa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="37E5EAB8FFE2FF9FDDFF4C428D2B2A3F" bold="true" box="[655,818,527,553]" italics="true" pageId="3" pageNumber="4">Casuariverpa</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="2CD657C3FFE2FF9FDC4A4C5D8DBC2A3C" box="[826,933,528,554]" pageId="3" pageNumber="4" rank="genus">gen. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection id="4D8B6521FFE2FF9FDFCD4C398F1D2B3B" pageId="3" pageNumber="4" type="type_taxon">
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<paragraph id="052E36AAFFE2FF9FDFCD4C398F412A9B" blockId="3.[189,344,627,654]" box="[189,344,627,654]" pageId="3" pageNumber="4">
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<heading id="5E6681C6FFE2FF9FDFCD4C398F412A9B" bold="true" box="[189,344,627,654]" fontSize="11" level="3" pageId="3" pageNumber="4" reason="3">
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<emphasis id="37E5EAB8FFE2FF9FDFCD4C398F412A9B" bold="true" box="[189,344,627,654]" pageId="3" pageNumber="4">
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<typeStatus id="DA2A8808FFE2FF9FDFCD4C398EE32A98" box="[189,250,628,654]" pageId="3" pageNumber="4">Type</typeStatus>
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species
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="052E36AAFFE2FF9FDFCD4CEE8C742AAB" blockId="3.[189,621,675,701]" box="[189,621,675,701]" pageId="3" pageNumber="4">
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<taxonomicName id="C2914D29FFE2FF9FDFCD4CEE8FAF2AAA" ID-CoL="RQZ6" authority="Enghoff, 2016" authorityName="Enghoff" authorityYear="2016" box="[189,438,675,701]" class="Diplopoda" family="Odontopygidae" genus="Casuariverpa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="scarpa" status="gen. et sp. nov.">
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<emphasis id="37E5EAB8FFE2FF9FDFCD4CEE8FAF2AAA" box="[189,438,675,701]" italics="true" pageId="3" pageNumber="4">Casuariverpa scarpa</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="2CD657C3FFE2FF9FDECD4CEE8C742AAB" box="[445,621,675,701]" pageId="3" pageNumber="4" rank="species">gen. et sp. nov.</taxonomicNameLabel>
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</paragraph>
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<paragraph id="052E36AAFFE2FF9FDFCD4CAE8FC12AEB" blockId="3.[189,472,739,765]" box="[189,472,739,765]" pageId="3" pageNumber="4">
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<heading id="5E6681C6FFE2FF9FDFCD4CAE8FC12AEB" bold="true" box="[189,472,739,765]" fontSize="11" level="3" pageId="3" pageNumber="4" reason="3">
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<emphasis id="37E5EAB8FFE2FF9FDFCD4CAE8FC12AEB" bold="true" box="[189,472,739,765]" pageId="3" pageNumber="4">Other included species</emphasis>
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</heading>
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</paragraph>
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<paragraph id="052E36AAFFE2FF9FDFCD4D5E8F1D2B3B" blockId="3.[189,260,787,813]" box="[189,260,787,813]" pageId="3" pageNumber="4">None.</paragraph>
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</subSubSection>
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<subSubSection id="4D8B6521FFE2FF9FDFCD4D1E8CCC2CCF" pageId="3" pageNumber="4" type="diagnosis">
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<paragraph id="052E36AAFFE2FF9FDFCD4D1E8F2F2B7B" blockId="3.[189,310,851,877]" box="[189,310,851,877]" pageId="3" pageNumber="4">
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<heading id="5E6681C6FFE2FF9FDFCD4D1E8F2F2B7B" bold="true" box="[189,310,851,877]" fontSize="11" level="3" pageId="3" pageNumber="4" reason="3">
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<emphasis id="37E5EAB8FFE2FF9FDFCD4D1E8F2F2B7B" bold="true" box="[189,310,851,877]" pageId="3" pageNumber="4">Diagnosis</emphasis>
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</heading>
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||
</paragraph>
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<paragraph id="052E36AAFFE2FF9FDFCD4DCF8CCC2CCF" blockId="3.[189,1399,898,1241]" pageId="3" pageNumber="4">
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A genus of Odontopygidae-Prionopetalini characterized by: anal valves with a well-developed dorsal spine, a similarly well-developed ventral spine and strongly protruding setiferous tubercles – male body ring seven with large ventral lobes – limbus very broad, with a simply denticulate margin – ventral postfemoral and tibial pads on some male legs - gonopod coxa simple, metaplica (
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<emphasis id="37E5EAB8FFE2FF9FDBF34DA08ABE2C10" box="[1155,1191,1005,1030]" italics="true" pageId="3" pageNumber="4">mp</emphasis>
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) with a subapical basad spine (
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<emphasis id="37E5EAB8FFE2FF9FDE2A4A5D8F902C3F" box="[346,393,1040,1065]" italics="true" pageId="3" pageNumber="4">msp</emphasis>
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) – gonopod telopodite with a compact torsotope (
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<emphasis id="37E5EAB8FFE2FF9FDC964A5D8DEF2C3F" box="[998,1014,1040,1065]" italics="true" pageId="3" pageNumber="4">tt</emphasis>
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) – no pretorsal or torsal spines/ processes – a pronounced posttorsal narrowing (
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<emphasis id="37E5EAB8FFE2FF9FDC704A7E8D042C5A" box="[768,797,1075,1100]" italics="true" pageId="3" pageNumber="4">pn</emphasis>
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) without spines – a division of the telopodite into solenomere (
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<emphasis id="37E5EAB8FFE2FF9FDE254A188F672C79" box="[341,382,1109,1135]" italics="true" pageId="3" pageNumber="4">slm</emphasis>
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) and telomere (
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<emphasis id="37E5EAB8FFE2FF9FDD474A188C452C79" box="[567,604,1109,1135]" italics="true" pageId="3" pageNumber="4">tlm</emphasis>
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) immediately distal to posttorsal narrowing – solenomere whiplike, with a very long accessory spine (
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<emphasis id="37E5EAB8FFE2FF9FDD124A348C652C84" box="[610,636,1145,1170]" italics="true" pageId="3" pageNumber="4">ps</emphasis>
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) branching off near its base and with a small subdistal lobe – telomere simple, with a longitudinal ridge on concave side enveloping solenomere, distally expanded, with micro-spinose fields basally and distally.
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</paragraph>
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</subSubSection>
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<subSubSection id="4D8B6521FFE2FF9FDFCD4B4C8FF12D99" pageId="3" pageNumber="4" type="etymology">
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<paragraph id="052E36AAFFE2FF9FDFCD4B4C8F5D2D0D" blockId="3.[189,324,1281,1307]" box="[189,324,1281,1307]" pageId="3" pageNumber="4">
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<heading id="5E6681C6FFE2FF9FDFCD4B4C8F5D2D0D" bold="true" box="[189,324,1281,1307]" fontSize="11" level="3" pageId="3" pageNumber="4" reason="3">
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<emphasis id="37E5EAB8FFE2FF9FDFCD4B4C8F5D2D0D" bold="true" box="[189,324,1281,1307]" pageId="3" pageNumber="4">Etymology</emphasis>
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</heading>
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</paragraph>
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<paragraph id="052E36AAFFE2FF9FDFCD4B7D8FF12D99" blockId="3.[189,1399,1327,1424]" pageId="3" pageNumber="4">
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From Latin
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<taxonomicName id="C2914D29FFE2FF9FDE384B628FAC2D5F" authorityName="Walker" authorityYear="1866" box="[328,437,1327,1353]" class="Insecta" family="Pyralidae" genus="Casuaria" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
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<emphasis id="37E5EAB8FFE2FF9FDE384B628FAC2D5F" box="[328,437,1327,1353]" italics="true" pageId="3" pageNumber="4">Casuaria</emphasis>
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</taxonomicName>
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, “cassowary”, and “
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<emphasis id="37E5EAB8FFE2FF9FDDD34B7D8CFF2D5F" box="[675,742,1328,1353]" italics="true" pageId="3" pageNumber="4">verpa</emphasis>
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”, penis. Refers to the shape of the solenomere tip which is somewhat reminiscent of the head profile of the Australian cassowary (
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<taxonomicName id="C2914D29FFE2FF9FDB744B1E8F162D86" authority="casuarius (Linnaeus, 1758))" authorityName="casuarius (Linnaeus" authorityYear="1758" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Aves" family="Casuariidae" genus="Casuarius" kingdom="Animalia" order="Struthioniformes" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="casuarius">
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<emphasis id="37E5EAB8FFE2FF9FDB744B1E8AED2D7A" box="[1028,1268,1363,1389]" italics="true" pageId="3" pageNumber="4">Casuarius casuarius</emphasis>
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(Linnaeus, 1758))
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</taxonomicName>
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. Gender feminine.
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</paragraph>
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</subSubSection>
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<subSubSection id="4D8B6521FFE2FF9FDFCD4BFA8A542F2A" pageId="3" pageNumber="4" type="discussion">
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<paragraph id="052E36AAFFE2FF9FDFCD4BFA8F362DC7" blockId="3.[189,303,1463,1489]" box="[189,303,1463,1489]" pageId="3" pageNumber="4">
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<heading id="5E6681C6FFE2FF9FDFCD4BFA8F362DC7" bold="true" box="[189,303,1463,1489]" fontSize="11" level="3" pageId="3" pageNumber="4" reason="3">
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<emphasis id="37E5EAB8FFE2FF9FDFCD4BFA8F362DC7" bold="true" box="[189,303,1463,1489]" pageId="3" pageNumber="4">Remarks</emphasis>
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</heading>
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</paragraph>
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<paragraph id="052E36AAFFE2FF9FDFCD4BAB8A542F2A" blockId="3.[189,1399,1509,1853]" pageId="3" pageNumber="4">
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In the key of
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<bibRefCitation id="61004B5BFFE2FF9FDE3D4BAB8FF02E16" author="Kraus O." box="[333,489,1509,1536]" pageId="3" pageNumber="4" refId="ref6370" refString="Kraus O. 1966. Phylogenie, Chorologie und Systematik der Odontopygoideen (Diplopoda, Spirostreptomorpha). Abhandlungen der senckenbergischen naturforschenden Gesellschaft 512, Naturmuseum Senckenberg, Frankfurt am Main." type="book" year="1966">Kraus (1966)</bibRefCitation>
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,
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<taxonomicName id="C2914D29FFE2FF9FDE824BAB8C782DE9" authority="Enghoff, 2016" authorityName="Enghoff" authorityYear="2016" box="[498,609,1510,1536]" class="Diplopoda" family="Odontopygidae" genus="Casuariverpa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="scarpa" status="gen. et sp. nov.">
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<emphasis id="37E5EAB8FFE2FF9FDE824BAB8C782DE9" box="[498,609,1510,1536]" italics="true" pageId="3" pageNumber="4">C. scarpa</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="2CD657C3FFE2FF9FDD154BAB8D152E16" box="[613,780,1510,1536]" pageId="3" pageNumber="4" rank="species">gen. et sp. nov.</taxonomicNameLabel>
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runs to
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<taxonomicName id="C2914D29FFE2FF9FDC124BA88AF92E16" authority="Attems, 1928" authorityName="Attems" authorityYear="1928" box="[866,1248,1509,1536]" class="Diplopoda" family="Odontopygidae" genus="Trichochaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
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<emphasis id="37E5EAB8FFE2FF9FDC124BA88A582DE9" box="[866,1089,1509,1535]" italics="true" pageId="3" pageNumber="4">Trichochaleponcus</emphasis>
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Attems, 1928
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</taxonomicName>
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(couplet 21). They key character for this genus is the very broad limbus (which is, however, paralleled in certain other odontopygids, e.g.,
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<taxonomicName id="C2914D29FFE2FF9FDE8248608DB72E50" authority="Kraus, 1960" authorityName="Kraus" authorityYear="1960" box="[498,942,1580,1606]" class="Diplopoda" family="Odontopygidae" genus="Kompsoprium" kingdom="Animalia" order="Spirostreptida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="firmosum">
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<emphasis id="37E5EAB8FFE2FF9FDE8248608D092E50" box="[498,784,1580,1606]" italics="true" pageId="3" pageNumber="4">Kompsoprium firmosum</emphasis>
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||
<bibRefCitation id="61004B5BFFE2FF9FDC6B48618DB72E50" author="Kraus O." box="[795,942,1580,1606]" pageId="3" pageNumber="4" refId="ref6335" refString="Kraus O. 1960. Athiopische Diplopoden I. Monographie der Odontopygidae-Odontopyginae (Diplopoda, Spirostreptoidea). Annalen van het Koninklijk Museum van Belgisch-Congo 82, Koninklijk Museum voor Midden Afrika, Tervuren." type="book" year="1960">Kraus, 1960</bibRefCitation>
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</taxonomicName>
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(
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<bibRefCitation id="61004B5BFFE2FF9FDCB148618A4B2E50" author="Kraus O." box="[961,1106,1580,1606]" pageId="3" pageNumber="4" refId="ref6335" refString="Kraus O. 1960. Athiopische Diplopoden I. Monographie der Odontopygidae-Odontopyginae (Diplopoda, Spirostreptoidea). Annalen van het Koninklijk Museum van Belgisch-Congo 82, Koninklijk Museum voor Midden Afrika, Tervuren." type="book" year="1960">
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Kraus
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<date id="712F106AFFE2FF9FDB6348618A4B2E50" box="[1043,1106,1580,1606]" pageId="3" pageNumber="4" value="1960">1960</date>
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</bibRefCitation>
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: fig. 69)).
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<taxonomicName id="C2914D29FFE2FF9FDBA548618F142E7F" authority="Enghoff, 2016" authorityName="Enghoff" authorityYear="2016" class="Diplopoda" family="Odontopygidae" genus="Casuariverpa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="scarpa" status="gen. et sp. nov.">
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<emphasis id="37E5EAB8FFE2FF9FDBA548618F142E7F" italics="true" pageId="3" pageNumber="4">Casuariverpa scarpa</emphasis>
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||
</taxonomicName>
|
||
<taxonomicNameLabel id="2CD657C3FFE2FF9FDE65481D8FD02E7C" box="[277,457,1616,1642]" pageId="3" pageNumber="4" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||
also agrees with
|
||
<taxonomicName id="C2914D29FFE2FF9FDDE848028D6E2E7F" authorityName="Attems" authorityYear="1928" box="[664,887,1615,1641]" class="Diplopoda" family="Odontopygidae" genus="Trichochaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="37E5EAB8FFE2FF9FDDE848028D6E2E7F" box="[664,887,1615,1641]" italics="true" pageId="3" pageNumber="4">Trichochaleponcus</emphasis>
|
||
</taxonomicName>
|
||
species in some other characters, but there are differences enough, at least vis-a-vis the
|
||
<typeStatus id="DA2A8808FFE2FF9FDDAC483E8D162E9B" box="[732,783,1651,1677]" pageId="3" pageNumber="4">type</typeStatus>
|
||
species of
|
||
<emphasis id="37E5EAB8FFE2FF9FDCEB483F8A982E9A" box="[923,1153,1650,1676]" italics="true" pageId="3" pageNumber="4">
|
||
<taxonomicName id="C2914D29FFE2FF9FDCEB483F8A642E9A" authorityName="Attems" authorityYear="1928" box="[923,1149,1650,1676]" class="Diplopoda" family="Odontopygidae" genus="Trichochaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">Trichochaleponcus</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
that a new genus is warranted, cf.
|
||
<bibRefCitation id="61004B5BFFE2FF9FDE0548D88C5A2EA6" author="Enghoff H." box="[373,579,1685,1712]" pageId="3" pageNumber="4" pagination="1 - 19" refId="ref6008" refString="Enghoff H. 2016 a. A mountain of millipedes III: A new genus for three new species from the Udzungwa mountains and surroundings, Tanzania, as well as several ' orphaned' species previously assigned to Odontopyge Brandt, 1841 (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 177: 1 - 19. http: // dx. doi. org / 10.5852 / ejt. 2016.177" type="journal article" year="2016">
|
||
Enghoff (
|
||
<date id="712F106AFFE2FF9FDE9C48D88C352EA6" box="[492,556,1685,1712]" pageId="3" pageNumber="4" value="2016">2016</date>
|
||
a)
|
||
</bibRefCitation>
|
||
. Thus,
|
||
<taxonomicName id="C2914D29FFE2FF9FDDD548DB8D5F2EA6" authority="Enghoff, 2016" authorityName="Enghoff" authorityYear="2016" box="[677,838,1686,1712]" class="Diplopoda" family="Odontopygidae" genus="Casuariverpa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="37E5EAB8FFE2FF9FDDD548DB8D5F2EA6" box="[677,838,1686,1712]" italics="true" pageId="3" pageNumber="4">Casuariverpa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="2CD657C3FFE2FF9FDC2548DB8DDE2EA6" box="[853,967,1686,1712]" pageId="3" pageNumber="4" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
has strongly protruding setiferous tubercles and an exceptionally well-developed ventral spine on the anal valves, its gonopod coxa has no lateral process or spine at mid-length, its gonopod telopodite has no spine between torsotope and posttorsal narrowing, the solenomere has no second basal spine in addition to the very long one, and the telomere is expanded distally instead of being strongly attenuate. See
|
||
<tableCitation id="48130311FFE2FF9FDC9F496F8A5E2F2A" box="[1007,1095,1826,1852]" captionStart="Table 1" captionStartId="4.[189,258,268,294]" captionTargetPageId="4" captionText="Table 1. (continued on next page) Comparison of Casuariverpa scarpa gen. et sp. nov. and Yia geminispina gen. et sp. nov. with Trichochaleponcus Attems, 1928 and its constituent species. Information on Trichochaleponcus taken from the cited papers." httpUri="http://table.plazi.org/id/51EE6622FFE5FF98DFCD4F418CF2297A" pageId="3" pageNumber="4" tableUuid="51EE6622FFE5FF98DFCD4F418CF2297A">Table 1</tableCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |