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<document id="3965FD7120D47E5CC2EC5DA77BE21965" ID-DOI="10.11646/phytotaxa.268.2.2" ID-ISSN="1179-3163" ID-Zenodo-Dep="13666769" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725392870972" checkinUser="felipe" docAuthor="Polyiam, Jutarat Kalb Wetchasart, Plata, Eimy Rivas, Bawingan, Paulina A., Kalb, Klaus &amp; Lücking, Robert" docDate="2016" docId="0397B65BFFBC5C3DFF75F974516D76E2" docLanguage="en" docName="phytotaxa.268.2.2.pdf" docOrigin="Phytotaxa 268 (2)" docSource="http://dx.doi.org/10.11646/phytotaxa.268.2.2" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Dyplolabia chumphonensis J. Kalb &amp; K. Kalb 2016, sp. nov." docType="treatment" docVersion="2" lastPageNumber="117" masterDocId="FFAECE23FFB95C3AFFFDFFF154697010" masterDocTitle=" Missing links alive? Novel taxa represent morphological transitions between distinctive phenotypes among extant Graphidaceae (lichenized Ascomycota: Ostropales)" masterLastPageNumber="122" masterPageNumber="110" pageNumber="115" updateTime="1725414794177" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="664EDB094AEFF150EB7ABC94D21A8B8B"> Missing links alive? Novel taxa represent morphological transitions between distinctive phenotypes among extant Graphidaceae (lichenized Ascomycota: Ostropales)</mods:title>
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<mods:namePart id="C18F8411CBF37BE52FA1CDCCAD2B530E">Polyiam, Jutarat Kalb Wetchasart</mods:namePart>
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<mods:namePart id="B1FD54CA96DD65492D7C1CDA7A5BDE8D">Plata, Eimy Rivas</mods:namePart>
<mods:affiliation id="21AB18DD6C4A36EF88F27002DF6B5E51">Science &amp; Education, Integrative Research &amp; Collections Center, The Field Museum, 1400 South Lake Shore Drive, Chicago, Illinois 60605 - 2496, U. S. A.</mods:affiliation>
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<mods:namePart id="26AB2E8B3215DCC1A110D176123C6B6D">Bawingan, Paulina A.</mods:namePart>
<mods:affiliation id="DF192417E335BF6B401DE065500A6517">School of Natural Sciences, Saint Louis University, Baguio City, Philippines.</mods:affiliation>
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<mods:namePart id="AADC719D16DB139C90810D8C1F0C6B92">Kalb, Klaus</mods:namePart>
<mods:affiliation id="E7412CD9AD91E28DDFDDAD65F3728AA1">Lichenologisches Institut Neumarkt, Im Tal 12, D- 92318 Neumarkt, Germany, and University of Regensburg, Institute of Plant Sciences, Universitätsstraβe 31, D- 93040 Regensburg, Germany.</mods:affiliation>
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<mods:namePart id="9582D75E6E05ACA3F42AA5398FBE3123">Lücking, Robert</mods:namePart>
<mods:affiliation id="E946B6A6EDF1FBA7512E5B7207354224">Botanical Garden and Botanical Museum, Königin-Luise-Strasse 6 - 8, 14195 Berlin, Germany.</mods:affiliation>
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<mods:title id="0F130101D74D555D40B3AF791A696C61">Phytotaxa</mods:title>
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<mods:date id="79D72966ED0660A7D7486A270629763B">2016</mods:date>
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<taxonomicName id="4C3E7CCEFFBC5C3FFF75F97456CE76B0" ID-CoL="386HZ" authority="J. Kalb &amp; K. Kalb" authorityName="J. Kalb &amp; K. Kalb" authorityYear="2016" box="[136,679,1669,1696]" class="Lecanoromycetes" family="Graphidaceae" genus="Dyplolabia" kingdom="Fungi" order="Ostropales" pageId="5" pageNumber="115" phylum="Ascomycota" rank="species" species="chumphonensis" status="sp. nov.">
<emphasis id="B94ADB5FFFBC5C3FFF75F97455AA76B0" bold="true" box="[136,451,1669,1696]" italics="true" pageId="5" pageNumber="115">Dyplolabia chumphonensis</emphasis>
J. Kalb &amp; K. Kalb
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,
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<taxonomicNameLabel id="A2796624FFBC5C3FFD4EF977576376B0" box="[691,778,1670,1696]" pageId="5" pageNumber="115" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</emphasis>
(
<figureCitation id="13051BC8FFBC5C3FFCE7F97657E576B0" box="[794,908,1671,1696]" captionStart="FIGURE 6" captionStartId="9.[136,229,1576,1597]" captionTargetBox="[189,1398,190,1551]" captionTargetId="figure-14@9.[189,1398,190,1551]" captionTargetPageId="9" captionText="FIGURE 6. A. Dyplolabia dalywaiana (holotype; thallus with ascomata). BD. D. oryzoides (B, Brazil, Kalb 31120; C, D, Sri Lanka, Weerakoon 116; thallus with ascomata and ascospores). EF. D. chumphonensis (holotype; thallus with ascomata and ascospores). Scale = 1 mm, in D, F = 10 μm." figureDoi="http://doi.org/10.5281/zenodo.13666795" httpUri="https://zenodo.org/record/13666795/files/figure.png" pageId="5" pageNumber="115">Fig. 6EF</figureCitation>
)
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<paragraph id="8B81074DFFBC5C3FFF75F95A55E276D4" blockId="5.[136,395,1707,1732]" box="[136,395,1707,1732]" pageId="5" pageNumber="115">MycoBank MB817436</paragraph>
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Differing from
<taxonomicName id="4C3E7CCEFFBC5C3FFEE1F9075583771B" box="[284,490,1782,1803]" class="Lecanoromycetes" family="Graphidaceae" genus="Dyplolabia" kingdom="Fungi" order="Ostropales" pageId="5" pageNumber="115" phylum="Ascomycota" rank="species" species="oryzoides">
<emphasis id="B94ADB5FFFBC5C3FFEE1F9075583771B" box="[284,490,1782,1803]" italics="true" pageId="5" pageNumber="115">Dyplolabia oryzoides</emphasis>
</taxonomicName>
by smaller ascospores.
</paragraph>
<paragraph id="8B81074DFFBC5C3FFF75F8EB511D7743" blockId="5.[136,1451,1782,1875]" pageId="5" pageNumber="115">
<materialsCitation id="3B560D10FFBC5C3FFF75F8EB570E773F" box="[136,871,1818,1839]" collectorName="Type" country="Thailand" location="Tambon Tha Sae" municipality="Tasae district" pageId="5" pageNumber="115" specimenCount="1" stateProvince="Chumphon">
<collectorName id="26CB629BFFBC5C3FFF75F8EB54D1773F" box="[136,184,1818,1839]" pageId="5" pageNumber="115">Type</collectorName>
:
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<collectingCountry id="F32947DDFFBC5C3FFF25F8EB553E773F" box="[216,343,1818,1839]" name="Thailand" pageId="5" pageNumber="115">THAILAND</collectingCountry>
.
<collectingRegion id="49FAC9AFFFBC5C3FFE9CF8EB5643773F" box="[353,554,1818,1839]" country="Thailand" name="Chumphon" pageId="5" pageNumber="115">Chumphon province</collectingRegion>
:
<collectingMunicipality id="6BE59D37FFBC5C3FFDC9F8EB56DF773F" box="[564,694,1818,1839]" pageId="5" pageNumber="115">Tasae district</collectingMunicipality>
,
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;
<materialsCitation id="3B560D10FFBC5C3FFC8FF8EB5175773F" box="[882,1308,1816,1839]" country="Thailand" location="Anniversary Queen Sirikit Park" pageId="5" pageNumber="115" specimenCount="1" stateProvince="Chumphon">
area of 72
<superScript id="7C4BAA05FFBC5C3FFC2EF8E957887734" attach="left" box="[979,993,1816,1828]" fontSize="5" pageId="5" pageNumber="115">nd</superScript>
<location id="8EE15196FFBC5C3FFC1AF8EB5175773F" LSID="urn:lsid:plazi:treatment:0397B65BFFBC5C3DFF75F974516D76E2:8EE15196FFBC5C3FFC1AF8EB5175773F" box="[999,1308,1818,1839]" country="Thailand" name="Anniversary Queen Sirikit Park" pageId="5" pageNumber="115" stateProvince="Chumphon">Anniversary Queen Sirikit Park</location>
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;
<geoCoordinate id="EE0A618AFFBC5C3FFADAF8EB51CE773F" box="[1319,1447,1818,1839]" degrees="10" direction="north" minutes="43" orientation="latitude" pageId="5" pageNumber="115" precision="15" seconds="24" value="10.723333">10°4324 N</geoCoordinate>
,
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,
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; on a ± free standing deciduous tree;
<materialsCitation id="3B560D10FFBC5C3FFD17F8CF51197743" box="[746,1392,1854,1875]" collectingDate="2011-03-19" collectionCode="RAMK" country="Thailand" pageId="5" pageNumber="115" specimenCode="RAMK 027820" specimenCount="1" stateProvince="Chumphon" typeStatus="holotype">
<date id="FF80218DFFBC5C3FFD17F8CF57177743" box="[746,894,1854,1875]" pageId="5" pageNumber="115" value="2011-03-19">
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,
<emphasis id="B94ADB5FFFBC5C3FFC75F8CF500E7743" box="[904,1127,1854,1875]" italics="true" pageId="5" pageNumber="115">J. Sutjaritturakan 3606</emphasis>
(
<typeStatus id="5485B9EFFFBC5C3FFB8EF8CF50A07743" box="[1139,1225,1854,1875]" pageId="5" pageNumber="115" type="holotype">holotype</typeStatus>
<specimenCode id="DB98AF36FFBC5C3FFB32F8CF51007743" box="[1231,1385,1854,1875]" collectionCode="RAMK" country="Thailand" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13907" lsid="urn:lsid:biocol.org:col:13907" name="Ramkhamhaeng University" pageId="5" pageNumber="115" type="Herbarium">RAMK 027820</specimenCode>
)
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.
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Thallus corticolous, epiperidermal, yellow-olive, smooth to uneven, with dense, prosoplectenchymatous cortex; photobiont layer without or with very rare scattered clusters of calcium oxalate crystals.Apothecia erumpent, elongate-lirellate,
<quantity id="4CC6AAA8FFBC5C3FFF0DF83A553877F4" box="[240,337,1995,2020]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="4.0" metricValueMin="1.0" pageId="5" pageNumber="115" unit="mm" value="2.5" valueMax="4.0" valueMin="1.0">14 mm</quantity>
long and
<quantity id="4CC6AAA8FFBC5C3FFE31F83A560C77F4" box="[460,613,1995,2020]" metricMagnitude="-4" metricUnit="m" metricValue="6.25" metricValueMax="7.5" metricValueMin="5.0" pageId="5" pageNumber="115" unit="mm" value="0.625" valueMax="0.75" valueMin="0.5">0.50.75 mm</quantity>
broad, straight or curved, very rarely branched; disc immersed, thickly white-pruinose, fully covered by margin; proper margin distinct, thick, erect, black but with a very thick, white cover; thalline margin absent. Excipulum apically and laterally carbonized, jet-black, without discernable structure. Periphysoids absent but hymenium laterally with a broad zone of widely spaced, anastomosing paraphyses embedded in a gelatinous matrix. Pseudocolumella absent. Hymenium 75110 μm high; paraphyses unbranched except for lateral areas as described above.Ascospores 46 (8)/ascus, uniseriate, submuriform with 35 transverse and 01 longitudinal septa per segment, 1619 × 89 μm, oval, with mostly thick septa (rarely thin septa) and diamond-shaped lumina (
<taxonomicName id="4C3E7CCEFFBF5C3CFF6CF8B45548774E" box="[145,289,1861,1886]" class="Dothideomycetes" family="Trypetheliaceae" genus="Astrothelium" kingdom="Fungi" order="Trypetheliales" pageId="6" pageNumber="116" phylum="Ascomycota" rank="genus">
<emphasis id="B94ADB5FFFBF5C3CFF6CF8B45548774E" box="[145,289,1861,1886]" italics="true" pageId="6" pageNumber="116">Astrothelium</emphasis>
</taxonomicName>
-
<typeStatus id="5485B9EFFFBF5C3CFED4F8B45537774E" box="[297,350,1861,1886]" pageId="6" pageNumber="116">type</typeStatus>
), colorless, I (non-amyloid).
</paragraph>
<caption id="DF4157C5FFBF5C3CFF75F9D957197695" ID-DOI="http://doi.org/10.5281/zenodo.13666781" ID-Zenodo-Dep="13666781" httpUri="https://zenodo.org/record/13666781/files/figure.png" pageId="6" pageNumber="116" startId="6.[136,229,1576,1597]" targetBox="[189,1398,190,1551]" targetPageId="6" targetType="figure">
<paragraph id="8B81074DFFBF5C3CFF75F9D957197695" blockId="6.[136,1452,1576,1669]" pageId="6" pageNumber="116">
<emphasis id="B94ADB5FFFBF5C3CFF75F9D95494762D" bold="true" box="[136,253,1576,1597]" pageId="6" pageNumber="116">FIGURE 3.</emphasis>
AD.
<taxonomicName id="4C3E7CCEFFBF5C3CFEC6F9D9564A762D" box="[315,547,1576,1597]" class="Lecanoromycetes" family="Graphidaceae" genus="Cruentotrema" kingdom="Fungi" order="Ostropales" pageId="6" pageNumber="116" phylum="Ascomycota" rank="species" species="puniceum">
<emphasis id="B94ADB5FFFBF5C3CFEC6F9D9564A762D" box="[315,547,1576,1597]" italics="true" pageId="6" pageNumber="116">Cruentotrema puniceum</emphasis>
</taxonomicName>
(A, D, Australia,
<emphasis id="B94ADB5FFFBF5C3CFD30F9D9574A762D" box="[717,803,1576,1597]" italics="true" pageId="6" pageNumber="116">Kalb s.n.</emphasis>
; B, holotype of
<taxonomicName id="4C3E7CCEFFBF5C3CFC40F9D950C4762D" box="[957,1197,1576,1597]" class="Lecanoromycetes" family="Graphidaceae" genus="Thelotrema" kingdom="Fungi" order="Ostropales" pageId="6" pageNumber="116" phylum="Ascomycota" rank="species" species="rhododiscum">
<emphasis id="B94ADB5FFFBF5C3CFC40F9D950C4762D" box="[957,1197,1576,1597]" italics="true" pageId="6" pageNumber="116">Thelotrema rhododiscum</emphasis>
</taxonomicName>
, C, Thailand,
<emphasis id="B94ADB5FFFBF5C3CFAC8F9D951CC762D" box="[1333,1445,1576,1597]" italics="true" pageId="6" pageNumber="116">Kalb 33412</emphasis>
; thallus with ascomata, section through ascoma showing excipulum, and ascospores). EF.
<taxonomicName id="4C3E7CCEFFBF5C3CFC05F9BD50EB7671" box="[1016,1154,1612,1633]" class="Lecanoromycetes" family="Graphidaceae" genus="Cruentotrema" kingdom="Fungi" order="Ostropales" pageId="6" pageNumber="116" phylum="Ascomycota" rank="species" species="cruentatum">
<emphasis id="B94ADB5FFFBF5C3CFC05F9BD50EB7671" box="[1016,1154,1612,1633]" italics="true" pageId="6" pageNumber="116">C. cruentatum</emphasis>
</taxonomicName>
(Brazil,
<emphasis id="B94ADB5FFFBF5C3CFB24F9BD51377671" box="[1241,1374,1612,1633]" italics="true" pageId="6" pageNumber="116">Cáceres 1070</emphasis>
; thallus with ascomata and ascospores). Scale = 1 mm, in C = 100 μm, D, F = 10 μm.
</paragraph>
</caption>
<paragraph id="8B81074DFFBF5C3CFF40F89850217792" blockId="6.[136,1452,1717,2030]" box="[189,1096,1897,1922]" pageId="6" pageNumber="116">
<emphasis id="B94ADB5FFFBF5C3CFF40F89855AB7792" bold="true" box="[189,450,1897,1922]" pageId="6" pageNumber="116">Secondary chemistry:</emphasis>
—White cover of ascoma margin C+ red (lecanoric acid).
</paragraph>
</subSubSection>
<subSubSection id="C32454C6FFBF5C3CFF40F87C50F477B6" box="[189,1181,1933,1958]" pageId="6" pageNumber="116" type="etymology">
<paragraph id="8B81074DFFBF5C3CFF40F87C50F477B6" blockId="6.[136,1452,1717,2030]" box="[189,1181,1933,1958]" pageId="6" pageNumber="116">
<emphasis id="B94ADB5FFFBF5C3CFF40F87C552177B6" bold="true" box="[189,328,1933,1958]" pageId="6" pageNumber="116">Etymology:</emphasis>
―This new species is named after the Thai province where it was collected.
</paragraph>
</subSubSection>
<subSubSection id="C32454C6FFBF5C3CFF40F840566F77FE" pageId="6" pageNumber="116" type="distribution">
<paragraph id="8B81074DFFBF5C3CFF40F840566F77FE" blockId="6.[136,1452,1717,2030]" pageId="6" pageNumber="116">
<emphasis id="B94ADB5FFFBF5C3CFF40F840558577DA" bold="true" box="[189,492,1969,1994]" pageId="6" pageNumber="116">Distribution and ecology:</emphasis>
<collectingCountry id="F32947DDFFBF5C3CFDF5F840560677DA" box="[520,623,1969,1994]" name="Thailand" pageId="6" pageNumber="116">Thailand</collectingCountry>
; thus far only known a single collection from the
<typeStatus id="5485B9EFFFBF5C3CFB6BF84050AF77DA" box="[1174,1222,1969,1994]" pageId="6" pageNumber="116">type</typeStatus>
locality, growing on bark of a deciduous tree in a park.
</paragraph>
</subSubSection>
<caption id="DF4157C5FFBE5C3DFF75F9D951267671" ID-DOI="http://doi.org/10.5281/zenodo.13666785" ID-Zenodo-Dep="13666785" httpUri="https://zenodo.org/record/13666785/files/figure.png" pageId="7" pageNumber="117" startId="7.[136,228,1576,1597]" targetBox="[189,1398,190,1551]" targetPageId="7" targetType="figure">
<paragraph id="8B81074DFFBE5C3DFF75F9D951267671" blockId="7.[136,1452,1576,1633]" pageId="7" pageNumber="117">
<emphasis id="B94ADB5FFFBE5C3DFF75F9D95495762D" bold="true" box="[136,252,1576,1597]" pageId="7" pageNumber="117">FIGURE 4.</emphasis>
AB.
<taxonomicName id="4C3E7CCEFFBE5C3DFEC5F9D9564F762D" box="[312,550,1576,1597]" class="Lecanoromycetes" family="Graphidaceae" genus="Cruentotrema" kingdom="Fungi" order="Ostropales" pageId="7" pageNumber="117" phylum="Ascomycota" rank="species" species="amazonum">
<emphasis id="B94ADB5FFFBE5C3DFEC5F9D9564F762D" box="[312,550,1576,1597]" italics="true" pageId="7" pageNumber="117">Cruentotrema amazonum</emphasis>
</taxonomicName>
(holotype; thallus with ascomata). CF.
<taxonomicName id="4C3E7CCEFFBE5C3DFC5FF9D95040762D" box="[930,1065,1576,1597]" class="Lecanoromycetes" family="Graphidaceae" genus="Cruentotrema" kingdom="Fungi" order="Ostropales" pageId="7" pageNumber="117" phylum="Ascomycota" rank="species" species="kurandense">
<emphasis id="B94ADB5FFFBE5C3DFC5FF9D95040762D" box="[930,1065,1576,1597]" italics="true" pageId="7" pageNumber="117">C. kurandense</emphasis>
</taxonomicName>
(C, holotype; D, India,
<emphasis id="B94ADB5FFFBE5C3DFAF4F9D951CC762D" box="[1289,1445,1576,1597]" italics="true" pageId="7" pageNumber="117">Lumbsch 19737l</emphasis>
; E, India,
<emphasis id="B94ADB5FFFBE5C3DFF23F9BD55167671" box="[222,383,1612,1633]" italics="true" pageId="7" pageNumber="117">Lumbsch 19736g</emphasis>
; F, Thailand,
<emphasis id="B94ADB5FFFBE5C3DFDFDF9BD56AB7671" box="[512,706,1612,1633]" italics="true" pageId="7" pageNumber="117">Jariangprasert 1352</emphasis>
; thallus with ascomata and ascospores). Scale = 1 mm, in F = 10 μm.
</paragraph>
</caption>
<subSubSection id="C32454C6FFBE5C3DFF40F960516D76E2" pageId="7" pageNumber="117" type="discussion">
<paragraph id="8B81074DFFBE5C3DFF40F960516D76E2" blockId="7.[136,1451,1681,1778]" pageId="7" pageNumber="117">
<emphasis id="B94ADB5FFFBE5C3DFF40F960555D76BA" bold="true" box="[189,308,1681,1706]" pageId="7" pageNumber="117">Remarks:</emphasis>
<taxonomicName id="4C3E7CCEFFBE5C3DFEADF96056EA76BA" box="[336,643,1681,1706]" class="Lecanoromycetes" family="Graphidaceae" genus="Dyplolabia" kingdom="Fungi" order="Ostropales" pageId="7" pageNumber="117" phylum="Ascomycota" rank="species" species="chumphonensis">
<emphasis id="B94ADB5FFFBE5C3DFEADF96056EA76BA" box="[336,643,1681,1706]" italics="true" pageId="7" pageNumber="117">Dyplolabia chumphonensis</emphasis>
</taxonomicName>
is very similar to
<taxonomicName id="4C3E7CCEFFBE5C3DFCA8F96057D476BA" box="[853,957,1681,1706]" class="Lecanoromycetes" family="Graphidaceae" genus="Dyplolabia" kingdom="Fungi" order="Ostropales" pageId="7" pageNumber="117" phylum="Ascomycota" rank="species" species="afzelii">
<emphasis id="B94ADB5FFFBE5C3DFCA8F96057D476BA" box="[853,957,1681,1706]" italics="true" pageId="7" pageNumber="117">D. afzelii</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3E7CCEFFBE5C3DFC0AF96050EF76BA" box="[1015,1158,1681,1706]" class="Lecanoromycetes" family="Graphidaceae" genus="Dyplolabia" kingdom="Fungi" order="Ostropales" pageId="7" pageNumber="117" phylum="Ascomycota" rank="species" species="oryzoides">
<emphasis id="B94ADB5FFFBE5C3DFC0AF96050EF76BA" box="[1015,1158,1681,1706]" italics="true" pageId="7" pageNumber="117">D. oryzoides</emphasis>
</taxonomicName>
. The first one very rarely has one longitudinal septum per spore (
<bibRefCitation id="EFAF7ABCFFBE5C3DFDB5F944572976DE" author="Kalb, K. &amp; Staiger, B." box="[584,832,1717,1742]" pageId="7" pageNumber="117" pagination="409 - 422" refId="ref6063" refString="Kalb, K. &amp; Staiger, B. (2000) Dyplolabia Massalongo. Monographie einer vergessenen Flechtengattung. Hoppea 61: 409 - 422." type="journal article" year="2000">Kalb &amp; Staiger, 2000</bibRefCitation>
), but is usually separated by only 4-locular ascospores without a longitudinal septum. The spores of
<taxonomicName id="4C3E7CCEFFBE5C3DFD79F928577876E2" box="[644,785,1753,1778]" class="Lecanoromycetes" family="Graphidaceae" genus="Dyplolabia" kingdom="Fungi" order="Ostropales" pageId="7" pageNumber="117" phylum="Ascomycota" rank="species" species="oryzoides">
<emphasis id="B94ADB5FFFBE5C3DFD79F928577876E2" box="[644,785,1753,1778]" italics="true" pageId="7" pageNumber="117">D. oryzoides</emphasis>
</taxonomicName>
are considerably larger, 1927 × 1014 μm.
</paragraph>
</subSubSection>
</treatment>
</document>