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<mods:title id="29FC782B9AC4B24F6D1171F3FDAEFCF9">Provisional checklist of freshwater fish diversity and distribution in Perak, Malaysia, and some latest taxonomic concerns</mods:title>
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<mods:namePart id="0E3F0768F69DF4C021ACE631A7964E56">Ng, Casey Keat Chuan</mods:namePart>
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<mods:namePart id="EEA7C8D75D6B1316E9C5716B8E201FF7">Lim, Teow Yeong</mods:namePart>
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<mods:namePart id="7EE03E8A6C4C05AA904EAEBF849962E2">Ahmad, Amirrudin</mods:namePart>
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<treatment id="039787C5FFB74D48FF28F9DEFAB1F809" ID-DOI="http://doi.org/10.5281/zenodo.5450171" ID-GBIF-Taxon="154435371" ID-Zenodo-Dep="5450171" LSID="urn:lsid:plazi:treatment:039787C5FFB74D48FF28F9DEFAB1F809" httpUri="http://treatment.plazi.org/id/039787C5FFB74D48FF28F9DEFAB1F809" lastPageNumber="525" pageId="10" pageNumber="525">
<subSubSection id="C3246558FFB74D48FF28F9DEFD8BF961" box="[151,523,1641,1666]" pageId="10" pageNumber="525" type="nomenclature">
<paragraph id="8B8136D3FFB74D48FF28F9DEFD8BF961" blockId="10.[151,1437,1641,2026]" box="[151,523,1641,1666]" pageId="10" pageNumber="525">
<heading id="D0C981BFFFB74D48FF28F9DEFD8BF961" bold="true" box="[151,523,1641,1666]" fontSize="10" level="2" pageId="10" pageNumber="525" reason="2">
<taxonomicName id="4C3E4D50FFB74D48FF28F9DEFD8BF961" ID-CoL="7CHBY" authority="(Valenciennes 1842)" baseAuthorityName="Valenciennes" baseAuthorityYear="1842" box="[151,523,1641,1666]" class="Actinopterygii" family="Cyprinidae" genus="Tor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="525" phylum="Chordata" rank="species" species="tambra">
<emphasis id="B94AEAC1FFB74D48FF28F9DEFD8BF961" bold="true" box="[151,523,1641,1666]" pageId="10" pageNumber="525">
<emphasis id="B94AEAC1FFB74D48FF28F9DEFE97F961" bold="true" box="[151,279,1641,1666]" italics="true" pageId="10" pageNumber="525">Tor tambra</emphasis>
(Valenciennes 1842)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3246558FFB74D48FF28F93AFCB3F90D" pageId="10" pageNumber="525" type="reference_group">
<paragraph id="8B8136D3FFB74D48FF28F93AFBFBF945" blockId="10.[151,1437,1641,2026]" box="[151,1147,1677,1702]" pageId="10" pageNumber="525">
<treatmentCitationGroup id="AB2E11FDFFB74D48FF28F93AFBFBF945" box="[151,1147,1677,1702]" pageId="10" pageNumber="525">
<taxonomicName id="4C3E4D50FFB74D48FF28F93AFDEBF945" ID-CoL="57FBV" authority=": Herre &amp; Myers, 1937: 60" authorityName=": Herre &amp; Myers" authorityPageNumber="60" authorityYear="1937" box="[151,619,1677,1702]" class="Actinopterygii" family="Cyprinidae" genus="Tor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="525" phylum="Chordata" rank="species" species="tambroides">
<emphasis id="B94AEAC1FFB74D48FF28F93AFEBFF945" box="[151,319,1677,1702]" italics="true" pageId="10" pageNumber="525">Tor tambroides</emphasis>
:
<treatmentCitation id="0A9F10C2FFB74D48FEF0F93AFDEBF945" author="Herre, A. W. C. T. &amp; Myers, G. S." box="[335,619,1677,1702]" page="60" pageId="10" pageNumber="525" year="1937">
<bibRefCitation id="EFAF4B22FFB74D48FEF0F93AFDC4F945" author="Herre, A. W. C. T. &amp; Myers, G. S." box="[335,580,1678,1702]" pageId="10" pageNumber="525" pagination="5 - 75" refId="ref15145" refString="Herre, A. W. C. T. &amp; Myers, G. S. (1937) A contribution to the ichthyology of the Malay Peninsula. Bulletin of the Raffles Museum, 13, 5 - 75." type="journal article" year="1937">Herre &amp; Myers, 1937</bibRefCitation>
: 60
</treatmentCitation>
</taxonomicName>
(Chandra Dam; River Plus near Kuala Legap).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B8136D3FFB74D48FF28F906FCB3F90D" blockId="10.[151,1437,1641,2026]" pageId="10" pageNumber="525">
<treatmentCitationGroup id="AB2E11FDFFB74D48FF28F906FCB3F90D" pageId="10" pageNumber="525">
<taxonomicName id="4C3E4D50FFB74D48FF28F906FDD1F929" ID-CoL="7CHBY" authority=": Herre &amp; Myers, 1937: 60" authorityName=": Herre &amp; Myers" authorityPageNumber="60" authorityYear="1937" box="[151,593,1713,1738]" class="Actinopterygii" family="Cyprinidae" genus="Tor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="525" phylum="Chordata" rank="species" species="tambra">
<emphasis id="B94AEAC1FFB74D48FF28F906FE98F929" box="[151,280,1713,1738]" italics="true" pageId="10" pageNumber="525">Tor tambra</emphasis>
:
<treatmentCitation id="0A9F10C2FFB74D48FE96F906FDD1F929" author="Herre, A. W. C. T. &amp; Myers, G. S." box="[297,593,1713,1738]" page="60" pageId="10" pageNumber="525" year="1937">
<bibRefCitation id="EFAF4B22FFB74D48FE96F906FDA7F929" author="Herre, A. W. C. T. &amp; Myers, G. S." box="[297,551,1714,1738]" pageId="10" pageNumber="525" pagination="5 - 75" refId="ref15145" refString="Herre, A. W. C. T. &amp; Myers, G. S. (1937) A contribution to the ichthyology of the Malay Peninsula. Bulletin of the Raffles Museum, 13, 5 - 75." type="journal article" year="1937">Herre &amp; Myers, 1937</bibRefCitation>
: 60
</treatmentCitation>
</taxonomicName>
(River Plus);
<treatmentCitation id="0A9F10C2FFB74D48FD47F905FC4DF929" author="Tweedie, M. W. F." box="[760,973,1713,1738]" page="74" pageId="10" pageNumber="525" year="1940">
<bibRefCitation id="EFAF4B22FFB74D48FD47F905FC23F929" author="Tweedie, M. W. F." box="[760,931,1713,1738]" pageId="10" pageNumber="525" pagination="68 - 82" refId="ref18809" refString="Tweedie, M. W. F. (1940) Additions to the collection of fishes in the Raffles Museum. Bulletin of the Raffles Museum, 16, 68 - 82." type="journal article" year="1940">Tweedie, 1940</bibRefCitation>
: 74
</treatmentCitation>
(River Plus);
<treatmentCitation id="0A9F10C2FFB74D48FBCBF905FE87F90D" author="Ahmad, A. &amp; Zakaria-Ismail, M." page="413" pageId="10" pageNumber="525" year="2014">
<bibRefCitation id="EFAF4B22FFB74D48FBCBF905FF54F90D" author="Ahmad, A. &amp; Zakaria-Ismail, M." pageId="10" pageNumber="525" pagination="407 - 419" refId="ref13660" refString="Ahmad, A. &amp; Zakaria-Ismail, M. (2014) Additional native fish fauna of rivers &amp; streams flowing into Temengor reservoir, Perak, Malaysia. Malayan Nature Journal, 664, 407 - 419." type="journal article" year="2014">Ahmad &amp; Zakaria-Ismail, 2014</bibRefCitation>
: 413
</treatmentCitation>
(Sg. Ketir; Sg. Tiang); Authors data (Sg. Kuak).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3246558FFB74D48FF28F94DFAB1F809" pageId="10" pageNumber="525" type="discussion">
<paragraph id="8B8136D3FFB74D48FF28F94DFAB1F809" blockId="10.[151,1437,1641,2026]" pageId="10" pageNumber="525">
<emphasis id="B94AEAC1FFB74D48FF28F94DFE8BF8F1" bold="true" box="[151,267,1785,1810]" pageId="10" pageNumber="525">Remarks.</emphasis>
To be conservative, we regard all specimens found by studies in
<collectingRegion id="49FAF831FFB74D48FC5BF94DFBA4F8F1" box="[996,1060,1785,1810]" country="Malaysia" name="Perak" pageId="10" pageNumber="525">Perak</collectingRegion>
as
<taxonomicName id="4C3E4D50FFB74D48FBF6F94EFB46F8F1" authorityName=": Herre &amp; Myers" authorityYear="1937" box="[1097,1222,1785,1810]" class="Actinopterygii" family="Cyprinidae" genus="Tor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="525" phylum="Chordata" rank="species" species="tambra">
<emphasis id="B94AEAC1FFB74D48FBF6F94EFB46F8F1" box="[1097,1222,1785,1810]" italics="true" pageId="10" pageNumber="525">Tor tambra</emphasis>
</taxonomicName>
for the time being. Past literature has treated
<emphasis id="B94AEAC1FFB74D48FE05F8AAFC01F8D5" box="[442,897,1821,1846]" italics="true" pageId="10" pageNumber="525">
<taxonomicName id="4C3E4D50FFB74D48FE05F8AAFDA9F8D5" authorityName=": Herre &amp; Myers" authorityYear="1937" box="[442,553,1821,1846]" class="Actinopterygii" family="Cyprinidae" genus="Tor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="525" phylum="Chordata" rank="species" species="tambra">T. tambra</taxonomicName>
,
<taxonomicName id="4C3E4D50FFB74D48FD8AF8AAFD51F8D5" authorityName=": Herre &amp; Myers" authorityYear="1937" box="[565,721,1821,1846]" class="Actinopterygii" family="Cyprinidae" genus="Tor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="525" phylum="Chordata" rank="species" species="tambroides">T. tambroides</taxonomicName>
,
<taxonomicName id="4C3E4D50FFB74D48FD62F8AAFC01F8D5" baseAuthorityName="Valenciennes" baseAuthorityYear="1842" box="[733,897,1821,1846]" class="Actinopterygii" family="Cyprinidae" genus="Tor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="525" phylum="Chordata" rank="species" species="douronensis">T. douronensis</taxonomicName>
</emphasis>
and
<taxonomicName id="4C3E4D50FFB74D48FC07F8AAFB83F8D6" baseAuthorityName="non Valenciennes" baseAuthorityYear="1842" box="[952,1027,1822,1845]" class="Actinopterygii" family="Cyprinidae" genus="Tor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="525" phylum="Chordata" rank="species" species="soro">
<emphasis id="B94AEAC1FFB74D48FC07F8AAFB83F8D6" box="[952,1027,1822,1845]" italics="true" pageId="10" pageNumber="525">T. soro</emphasis>
</taxonomicName>
as valid species (
<bibRefCitation id="EFAF4B22FFB74D48FB75F8AAFAE7F8D5" author="Nguyen, T. T. T. &amp; Na-Nakorn, U. &amp; Sukmanomon, S. &amp; ZiMing, C." box="[1226,1383,1822,1846]" pageId="10" pageNumber="525" pagination="1223 - 1231" refId="ref17677" refString="Nguyen, T. T. T., Na-Nakorn, U., Sukmanomon, S. &amp; ZiMing, C. (2008) A study on phylogeny and biogeography of mahseer species (Pisces: Cyprinidae) using sequences of three mitochondrial DNA gene regions. Molecular Phylogenetics and Evolution, 48, 1223 - 1231. https: // doi. org / 10.1016 / j. ympev. 2008.01.006" type="journal article" year="2008">Nguyen 2008</bibRefCitation>
;
<bibRefCitation id="EFAF4B22FFB74D48FACCF8AAFE9DF8B9" author="Esa, Y. B. &amp; Rahim, K. A. A." pageId="10" pageNumber="525" pagination="1 - 14" refId="ref14982" refString="Esa, Y. B. &amp; Rahim, K. A. A. (2013) Genetic structure and preliminary findings of cryptic diversity of the Malaysian Mahseer (Tor tambroides Valenciennes: Cyprinidae) inferred from mitochondrial DNA and microsatellite analyses. BioMed Research International, 1 - 14. https: // doi. org / 10.1155 / 2013 / 170980" type="book chapter" year="2013">
Esa
<emphasis id="B94AEAC1FFB74D48FF28F8F6FF59F8B9" box="[151,217,1857,1882]" italics="true" pageId="10" pageNumber="525">et al.,</emphasis>
2013
</bibRefCitation>
;
<bibRefCitation id="EFAF4B22FFB74D48FE93F8F5FDBFF8B9" author="Norfatimah, M. Y. &amp; Teh, L. K. &amp; Salleh, M. Z. &amp; Mat Isa, M. N. &amp; SitiAzizah, M. N." box="[300,575,1857,1882]" pageId="10" pageNumber="525" pagination="263 - 269" refId="ref17775" refString="Norfatimah, M. Y., Teh, L. K., Salleh, M. Z., Mat Isa, M. N. &amp; SitiAzizah, M. N. (2014) Complete mitochondrial genome of Malaysian Mahseer (Tor tambroides). Gene, 548, 263 - 269. https: // doi. org / 10.1016 / j. gene. 2014.07.044" type="journal article" year="2014">
Norfatimah
<emphasis id="B94AEAC1FFB74D48FE09F8F6FE78F8B9" box="[438,504,1857,1882]" italics="true" pageId="10" pageNumber="525">et al.,</emphasis>
2014
</bibRefCitation>
).
<taxonomicName id="4C3E4D50FFB74D48FDEBF8F6FD21F8BA" baseAuthorityName="non Valenciennes" baseAuthorityYear="1842" box="[596,673,1858,1881]" class="Actinopterygii" family="Cyprinidae" genus="Tor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="525" phylum="Chordata" rank="species" species="soro">
<emphasis id="B94AEAC1FFB74D48FDEBF8F6FD21F8BA" box="[596,673,1858,1881]" italics="true" pageId="10" pageNumber="525">T. soro</emphasis>
</taxonomicName>
is now valid as
<taxonomicName id="4C3E4D50FFB74D48FCDAF8F5FAB8F8B9" authority="(Fricke et al., 2018)" baseAuthorityName="Fricke et al." baseAuthorityYear="2018" box="[869,1336,1857,1882]" class="Actinopterygii" family="Cyprinidae" genus="Neolissochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="525" phylum="Chordata" rank="species" species="soro">
<emphasis id="B94AEAC1FFB74D48FCDAF8F5FBC4F8BA" box="[869,1092,1857,1882]" italics="true" pageId="10" pageNumber="525">Neolissochilus soro</emphasis>
(
<bibRefCitation id="EFAF4B22FFB74D48FBEBF8F5FAB0F8B9" author="Fricke, R. &amp; Eschmeyer, W. N. &amp; van der Laan, R." box="[1108,1328,1857,1882]" pageId="10" pageNumber="525" refId="ref15047" refString="Fricke, R., Eschmeyer, W. N. &amp; van der Laan, R. (eds) (2018) Catalogue of Fishes: Genera, Species, References. California Academy of Sciences. Available from http: // researcharchive. calacademy. org / research / ichthyology / catalog / fishcatmain. asp (Accessed 20 October 2018)" type="url" year="2018">
Fricke
<emphasis id="B94AEAC1FFB74D48FB18F8F6FB62F8B9" box="[1191,1250,1857,1882]" italics="true" pageId="10" pageNumber="525">et al.</emphasis>
, 2018
</bibRefCitation>
)
</taxonomicName>
.
<bibRefCitation id="EFAF4B22FFB74D48FAFAF8F5FF61F89D" author="Roberts, T. R." pageId="10" pageNumber="525" pagination="225 - 236" refId="ref18096" refString="Roberts, T. R. (1999) Fishes of the cyprinid genus Tor in the Nam Theun watershed (Mekong Basin) of Laos, with description of a new species. The Raffles Bulletin of Zoology, 47 (1), 225 - 236." type="journal article" year="1999">Roberts (1999)</bibRefCitation>
and
<bibRefCitation id="EFAF4B22FFB74D48FEA5F8D1FE46F89D" author="Kottelat, M." box="[282,454,1893,1918]" pageId="10" pageNumber="525" pagination="1 - 663" refId="ref15672" refString="Kottelat, M. (2013) The fishes of the inland waters of Southeast Asia: A catalogue and core bibliography of the fishes known to occur in freshwaters, mangroves and estuaries. The Raffles Bulletin of Zoology, 27, 1 - 663." type="journal article" year="2013">Kottelat (2013)</bibRefCitation>
were unable to conclusively determine the unique morphological markers between the mentioned species. They have also treated
<taxonomicName id="4C3E4D50FFB74D48FD30F83EFCB7F841" baseAuthorityName="Valenciennes" baseAuthorityYear="1842" box="[655,823,1929,1954]" class="Actinopterygii" family="Cyprinidae" genus="Tor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="525" phylum="Chordata" rank="species" species="douronensis">
<emphasis id="B94AEAC1FFB74D48FD30F83EFCB7F841" box="[655,823,1929,1954]" italics="true" pageId="10" pageNumber="525">T. douronensis</emphasis>
</taxonomicName>
as junior synonym of
<taxonomicName id="4C3E4D50FFB74D48FBF9F83EFB35F841" authorityName=": Herre &amp; Myers" authorityYear="1937" box="[1094,1205,1929,1954]" class="Actinopterygii" family="Cyprinidae" genus="Tor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="525" phylum="Chordata" rank="species" species="tambra">
<emphasis id="B94AEAC1FFB74D48FBF9F83EFB35F841" box="[1094,1205,1929,1954]" italics="true" pageId="10" pageNumber="525">T. tambra</emphasis>
</taxonomicName>
. A recent study by
<bibRefCitation id="EFAF4B22FFB74D48FF28F819FEF9F825" author="Walton, S. E. &amp; Gan, H. M. &amp; Raghavan, R. &amp; Pinder, A. &amp; Ahmad, A." box="[151,377,1965,1990]" pageId="10" pageNumber="525" pagination="171 - 183" refId="ref18901" refString="Walton, S. E., Gan, H. M., Raghavan, R., Pinder, A. &amp; Ahmad, A. (2017) Disentangling the Taxonomy of the Mahseers (Tor spp.) of Malaysia: An Integrated Approach Using Morphology, Genetics and Historical Records. Reviews in Fisheries Science and Aquaculture, 171 - 183." type="book chapter" year="2017">
Walton
<emphasis id="B94AEAC1FFB74D48FF51F81AFEA7F825" box="[238,295,1965,1990]" italics="true" pageId="10" pageNumber="525">et al.</emphasis>
(2017)
</bibRefCitation>
also does not support multiple species designation for
<taxonomicName id="4C3E4D50FFB74D48FC5BF81AFB87F826" box="[996,1031,1966,1989]" class="Actinopterygii" family="Cyprinidae" genus="Tor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="525" phylum="Chordata" rank="genus">
<emphasis id="B94AEAC1FFB74D48FC5BF81AFB87F826" box="[996,1031,1966,1989]" italics="true" pageId="10" pageNumber="525">Tor</emphasis>
</taxonomicName>
that occurs in Peninsular
<collectingCountry id="F3297643FFB74D48FA91F819FA18F825" box="[1326,1432,1965,1990]" name="Malaysia" pageId="10" pageNumber="525">Malaysia</collectingCountry>
. Thus, following
<bibRefCitation id="EFAF4B22FFB74D48FEEEF865FE7FF809" author="Kottelat, M." box="[337,511,2001,2026]" pageId="10" pageNumber="525" pagination="1 - 663" refId="ref15672" refString="Kottelat, M. (2013) The fishes of the inland waters of Southeast Asia: A catalogue and core bibliography of the fishes known to occur in freshwaters, mangroves and estuaries. The Raffles Bulletin of Zoology, 27, 1 - 663." type="journal article" year="2013">Kottelat (2013)</bibRefCitation>
, our checklist adopted the name
<taxonomicName id="4C3E4D50FFB74D48FCD1F866FC5AF809" authorityName=": Herre &amp; Myers" authorityYear="1937" box="[878,986,2001,2026]" class="Actinopterygii" family="Cyprinidae" genus="Tor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="525" phylum="Chordata" rank="species" species="tambra">
<emphasis id="B94AEAC1FFB74D48FCD1F866FC5AF809" box="[878,986,2001,2026]" italics="true" pageId="10" pageNumber="525">T. tambra</emphasis>
</taxonomicName>
for specimens found in
<collectingRegion id="49FAF831FFB74D48FB55F865FAADF809" box="[1258,1325,2001,2026]" country="Malaysia" name="Perak" pageId="10" pageNumber="525">Perak</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>