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<document id="FB68704ED87FDAE01E3FF128A388C95A" ID-DOI="10.11646/zootaxa.4756.1.1" ID-GBIF-Dataset="4d48d52e-6505-4f62-909f-5da5c2cfb489" ID-ISSN="1175-5334" ID-Zenodo-Dep="3736895" ID-ZooBank="urn:lsid:zoobank.org:pub:9FF62616-A7FA-4331-AC51-0F534400631D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1585767638433" checkinUser="carolina" docAuthor="Kluge, Nikita J." docDate="2020" docId="039787A6FF1884E38CFBFB10DA6DFD9D" docLanguage="en" docName="zootaxa.4756.1.1.pdf.imf" docOrigin="Zootaxa 4756 (1)" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Thraulodes Ulmer 1920" docType="treatment" docVersion="7" lastPageNumber="136" masterDocId="FFAEFFDEFF9D84648C6CFFE6DB3EFFA2" masterDocTitle="Systematic position of Thraulodes Ulmer 1920 (Ephemeroptera: Leptophlebiidae) and descriptions of new and little-known species" masterLastPageNumber="142" masterPageNumber="1" pageNumber="134" updateTime="1698734461607" updateUser="plazi">
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<mods:title id="E84BC02B47CEF0064F6E6CD662C3E6AD">Systematic position of Thraulodes Ulmer 1920 (Ephemeroptera: Leptophlebiidae) and descriptions of new and little-known species</mods:title>
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<mods:namePart id="41A18A42E49CDAFD41657AE7287BD958">Kluge, Nikita J.</mods:namePart>
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<mods:date id="B6FAB029D04347A0F6FE9409BFDD578F">2020</mods:date>
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<mods:number id="EBECF4D73C10F5E2ED1139EC455B996F">4756</mods:number>
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<mods:identifier id="21B7395CF048535F30BA44BD5D99D7FD" type="DOI">10.11646/zootaxa.4756.1.1</mods:identifier>
<mods:identifier id="BFB9112FF7D67BC558B393E8A3865784" type="GBIF-Dataset">4d48d52e-6505-4f62-909f-5da5c2cfb489</mods:identifier>
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<treatment id="039787A6FF1884E38CFBFB10DA6DFD9D" ID-DOI="http://doi.org/10.5281/zenodo.3811749" ID-GBIF-Taxon="163420767" ID-Zenodo-Dep="3811749" LSID="urn:lsid:plazi:treatment:039787A6FF1884E38CFBFB10DA6DFD9D" httpUri="http://treatment.plazi.org/id/039787A6FF1884E38CFBFB10DA6DFD9D" lastPageId="135" lastPageNumber="136" pageId="133" pageNumber="134">
<subSubSection id="C324653BFF1884E18CFBFB10DAE0FAB2" box="[151,478,1270,1297]" pageId="133" pageNumber="134" type="nomenclature">
<paragraph id="8B8136B0FF1884E18CFBFB10DAE0FAB2" blockId="133.[151,478,1270,1332]" box="[151,478,1270,1297]" pageId="133" pageNumber="134">
<heading id="D0C981DCFF1884E18CFBFB10DAE0FAB2" bold="true" box="[151,478,1270,1297]" fontSize="10" level="2" pageId="133" pageNumber="134" reason="2">
<emphasis id="B94AEAA2FF1884E18CFBFB10DAE0FAB2" bold="true" box="[151,478,1270,1297]" pageId="133" pageNumber="134">
20.
<taxonomicName id="4C3E4D33FF1884E18CAFFB10DA70FAB3" ID-CoL="648C7" ID-ENA="309571" authorityName="Ulmer" authorityYear="1920" box="[195,334,1270,1297]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="133" pageNumber="134" phylum="Arthropoda" rank="genus">
<emphasis id="B94AEAA2FF1884E18CAFFB10DA70FAB3" bold="true" box="[195,334,1270,1297]" italics="true" pageId="133" pageNumber="134">Thraulodes</emphasis>
</taxonomicName>
sp. «Itaya»
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C324653BFF1884E18CFBFAFCDA7DFA96" box="[151,323,1305,1332]" pageId="133" pageNumber="134" type="description">
<paragraph id="8B8136B0FF1884E18CFBFAFCDA7DFA96" blockId="133.[151,478,1270,1332]" box="[151,323,1305,1332]" pageId="133" pageNumber="134">
(
<figureCitation id="13052A35FF1884E18CF3FAFCDA05FA96" box="[159,315,1305,1332]" captionStart-0="FIGURES 637–649" captionStart-1="FIGURES 650–653" captionStartId-0="134.[151,264,1864,1889]" captionStartId-1="135.[151,264,1222,1247]" captionTargetBox-0="[151,1436,196,1765]" captionTargetBox-1="[151,1435,589,1202]" captionTargetId-0="figure@134.[151,1436,196,1765]" captionTargetId-1="figure@135.[151,1435,589,1202]" captionTargetPageId-0="134" captionTargetPageId-1="135" captionText-0="FIGURES 637–649. Thraulodes sp. «Itaya», larva. 637, tergalius III; 638–645, larval exuviae at the same magnification: 638, part of head with labrum; 639–640, left and right mandibles; 641, right half of pronotum and mesonotum; 642–644, fore, middle and hind legs; 645, abdominal terga and sterna; 646, anterior side of hind femur; 647, anterior side of fore tibia; 648, hind tibia with focus on outer-posterior row of stout setae; 649, the same with focus on outer-anterior and inner-anterior rows of stout setae." captionText-1="FIGURES 650–653. Thraulodes sp. «Itaya», labrum. 650, general view; 651, focus on ventral side and six sensilla (shown by arrows); 652, with focus on anterior transverse setal row (transformed to field of irregularly situated setae); 653, with focus on posterior transverse setal row." figureDoi-0="http://doi.org/10.5281/zenodo.3737062" figureDoi-1="http://doi.org/10.5281/zenodo.3737064" httpUri-0="https://zenodo.org/record/3737062/files/figure.png" httpUri-1="https://zenodo.org/record/3737064/files/figure.png" pageId="133" pageNumber="134">Figs 637–653</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C324653BFF1884E18CFBFA87DE7FFA03" pageId="133" pageNumber="134" type="materials_examined">
<paragraph id="8B8136B0FF1884E18CFBFA87DE7FFA03" blockId="133.[151,1437,1377,2016]" pageId="133" pageNumber="134">
<emphasis id="B94AEAA2FF1884E18CFBFA87DABDFADE" bold="true" box="[151,387,1377,1404]" pageId="133" pageNumber="134">Material examined.</emphasis>
<materialsCitation id="3B563CEDFF1884E18DE1FA84DE7FFA03" ID-GBIF-Occurrence="2594509893" collectingDate="2006-11-15" collectorName="N. Kluge" country="Peru" location="Puente Itaya" municipality="Rio Itaya" pageId="133" pageNumber="134" specimenCount="3" specimenCount-female="2" specimenCount-male="1">
<collectingCountry id="F3297620FF1884E18DE1FA84DAE9FADE" box="[397,471,1378,1404]" name="Peru" pageId="133" pageNumber="134">PERU</collectingCountry>
,
<collectingMunicipality id="6BE5ACCAFF1884E18D89FA84D96CFADE" box="[485,594,1378,1404]" pageId="133" pageNumber="134">Rio Itaya</collectingMunicipality>
, upstream
<location id="8EE1606BFF1884E18EBCFA84D860FADE" LSID="urn:lsid:plazi:treatment:039787A6FF1884E38CFBFB10DA6DFD9D:8EE1606BFF1884E18EBCFA84D860FADE" box="[720,862,1378,1404]" country="Peru" municipality="Rio Itaya" name="Puente Itaya" pageId="133" pageNumber="134">Puente Itaya</location>
(
<quantity id="4CC69B55FF1884E18F03FA87D884FADE" box="[879,954,1377,1404]" metricMagnitude="4" metricUnit="m" metricValue="5.7" pageId="133" pageNumber="134" unit="km" value="57.0">
<locationDeviation id="9A332F92FF1884E18F03FA87D884FADE" box="[879,954,1377,1404]" metricMagnitude="4" metricUnit="m" metricValue="5.7" pageId="133" pageNumber="134" unit="km" value="57.0">57 km</locationDeviation>
</quantity>
by road from
<location id="8EE1606BFF1884E18808FA84DF88FADF" LSID="urn:lsid:plazi:treatment:039787A6FF1884E38CFBFB10DA6DFD9D:8EE1606BFF1884E18808FA84DF88FADF" box="[1124,1206,1378,1405]" country="Peru" municipality="Rio Itaya" name="Iquitos" pageId="133" pageNumber="134">Iquitos</location>
), near
<location id="8EE1606BFF1884E1896AFA84DE4DFADE" LSID="urn:lsid:plazi:treatment:039787A6FF1884E38CFBFB10DA6DFD9D:8EE1606BFF1884E1896AFA84DE4DFADE" box="[1286,1395,1378,1404]" country="Peru" municipality="Rio Itaya" name="Omaguas" pageId="133" pageNumber="134">Omaguas</location>
, 1–
<date id="FF801070FF1884E18CFBFA63DA2AFA02" box="[151,276,1413,1440]" pageId="133" pageNumber="134" value="2006-11-15">
<collectingDate id="EFC4E998FF1884E18CFBFA63DA2AFA02" box="[151,276,1413,1440]" pageId="133" pageNumber="134" value="2006-11-15">15.11.2006</collectingDate>
</date>
(
<collectorName id="26CB5366FF1884E18D48FA60DAAFFA03" box="[292,401,1414,1441]" pageId="133" pageNumber="134">N. Kluge</collectorName>
): 3 incomplete larval exuviae (last instar
<specimenCount id="9D38FD39FF1884E18F31FA60D8A8FA02" box="[861,918,1414,1440]" pageId="133" pageNumber="134" type="male">male</specimenCount>
, last instar
<specimenCount id="9D38FD39FF1884E18878FA60DF5EFA02" box="[1044,1120,1414,1440]" pageId="133" pageNumber="134" type="female">female</specimenCount>
and young
<specimenCount id="9D38FD39FF1884E18888FA60DE0BFA03" box="[1252,1333,1414,1441]" pageId="133" pageNumber="134" type="female">female</specimenCount>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C324653BFF1884E38CABFA4FD969FE71" lastPageId="135" lastPageNumber="136" pageId="133" pageNumber="134" type="description">
<paragraph id="8B8136B0FF1884E18CABFA4FDA5AFA66" blockId="133.[151,1437,1377,2016]" box="[199,356,1449,1476]" pageId="133" pageNumber="134">
<emphasis id="B94AEAA2FF1884E18CABFA4FDA5AFA66" bold="true" box="[199,356,1449,1476]" pageId="133" pageNumber="134">Descriptions.</emphasis>
</paragraph>
<paragraph id="8B8136B0FF1884E18CABFA28DA7FF9DA" blockId="133.[151,1437,1377,2016]" pageId="133" pageNumber="134">
<emphasis id="B94AEAA2FF1884E18CABFA28DA2CFA4A" bold="true" box="[199,274,1486,1512]" italics="true" pageId="133" pageNumber="134">Larva.</emphasis>
CUTICULAR COLORATION (
<figureCitation id="13052A35FF1884E18EE9FA28D818FA4A" box="[645,806,1485,1512]" captionStart="FIGURES 637–649" captionStartId="134.[151,264,1864,1889]" captionTargetBox="[151,1436,196,1765]" captionTargetId="figure@134.[151,1436,196,1765]" captionTargetPageId="134" captionText="FIGURES 637–649. Thraulodes sp. «Itaya», larva. 637, tergalius III; 638–645, larval exuviae at the same magnification: 638, part of head with labrum; 639–640, left and right mandibles; 641, right half of pronotum and mesonotum; 642–644, fore, middle and hind legs; 645, abdominal terga and sterna; 646, anterior side of hind femur; 647, anterior side of fore tibia; 648, hind tibia with focus on outer-posterior row of stout setae; 649, the same with focus on outer-anterior and inner-anterior rows of stout setae." figureDoi="http://doi.org/10.5281/zenodo.3737062" httpUri="https://zenodo.org/record/3737062/files/figure.png" pageId="133" pageNumber="134">Figs 638–645</figureCitation>
). Dorsal side of head brown. Pronotum and mesonotum brown with diffusive blanks; fore protopteron with veins darker than background. Femur light brown with diffusive blanks; tibia with 2 wide brown bands (proximal and middle); tarsus colorless.Abdominal terga without contrasting markings, posterior terga gradually darker than anterior ones. Abdominal sterna lighter, posterior sterna with brown arched macula.
</paragraph>
<paragraph id="8B8136B0FF1884E18CABF964DFD6F93F" blockId="133.[151,1437,1377,2016]" box="[199,1256,1666,1693]" pageId="133" pageNumber="134">
HYPODERMAL COLORATION. Coloration of body unknown. Tergalii dark gray (
<figureCitation id="13052A35FF1884E18815F964DFE2F93F" box="[1145,1244,1666,1693]" captionStart="FIGURES 637–649" captionStartId="134.[151,264,1864,1889]" captionTargetBox="[151,1436,196,1765]" captionTargetId="figure@134.[151,1436,196,1765]" captionTargetPageId="134" captionText="FIGURES 637–649. Thraulodes sp. «Itaya», larva. 637, tergalius III; 638–645, larval exuviae at the same magnification: 638, part of head with labrum; 639–640, left and right mandibles; 641, right half of pronotum and mesonotum; 642–644, fore, middle and hind legs; 645, abdominal terga and sterna; 646, anterior side of hind femur; 647, anterior side of fore tibia; 648, hind tibia with focus on outer-posterior row of stout setae; 649, the same with focus on outer-anterior and inner-anterior rows of stout setae." figureDoi="http://doi.org/10.5281/zenodo.3737062" httpUri="https://zenodo.org/record/3737062/files/figure.png" pageId="133" pageNumber="134">Fig. 637</figureCitation>
).
</paragraph>
<paragraph id="8B8136B0FF1884E18CABF940D989F88E" blockId="133.[151,1437,1377,2016]" pageId="133" pageNumber="134">
SHAPE AND SETATION. Clypeus slightly widened anteriorly; labrum 1.2 times wider than clypeus (
<figureCitation id="13052A35FF1884E18943F940DEAEF962" box="[1327,1424,1701,1728]" captionStart="FIGURES 650–653" captionStartId="135.[151,264,1222,1247]" captionTargetBox="[151,1435,589,1202]" captionTargetId="figure@135.[151,1435,589,1202]" captionTargetPageId="135" captionText="FIGURES 650–653. Thraulodes sp. «Itaya», labrum. 650, general view; 651, focus on ventral side and six sensilla (shown by arrows); 652, with focus on anterior transverse setal row (transformed to field of irregularly situated setae); 653, with focus on posterior transverse setal row." figureDoi="http://doi.org/10.5281/zenodo.3737064" httpUri="https://zenodo.org/record/3737064/files/figure.png" pageId="133" pageNumber="134">Fig. 650</figureCitation>
). Labrum widest at midlength, with narrow median emargination exposed ventrally; anterior transverse setal row wider than all 5 denticles, in its median part represents field of irregularly situated setae (
<figureCitation id="13052A35FF1884E188FCF908DE0FF8AA" box="[1168,1329,1773,1800]" captionStart="FIGURES 650–653" captionStartId="135.[151,264,1222,1247]" captionTargetBox="[151,1435,589,1202]" captionTargetId="figure@135.[151,1435,589,1202]" captionTargetPageId="135" captionText="FIGURES 650–653. Thraulodes sp. «Itaya», labrum. 650, general view; 651, focus on ventral side and six sensilla (shown by arrows); 652, with focus on anterior transverse setal row (transformed to field of irregularly situated setae); 653, with focus on posterior transverse setal row." figureDoi="http://doi.org/10.5281/zenodo.3737064" httpUri="https://zenodo.org/record/3737064/files/figure.png" pageId="133" pageNumber="134">Figs 651–653</figureCitation>
). Maxilla with 12–14 pectinate setae in apical-ventral row.
</paragraph>
<paragraph id="8B8136B0FF1884E18CABF8D0D805F8D6" blockId="133.[151,1437,1377,2016]" pageId="133" pageNumber="134">
Femora: Stout setae on anterior surface narrowed apically and truncate (
<figureCitation id="13052A35FF1884E18F84F8D0DF77F8F3" box="[1000,1097,1846,1873]" captionStart="FIGURES 637–649" captionStartId="134.[151,264,1864,1889]" captionTargetBox="[151,1436,196,1765]" captionTargetId="figure@134.[151,1436,196,1765]" captionTargetPageId="134" captionText="FIGURES 637–649. Thraulodes sp. «Itaya», larva. 637, tergalius III; 638–645, larval exuviae at the same magnification: 638, part of head with labrum; 639–640, left and right mandibles; 641, right half of pronotum and mesonotum; 642–644, fore, middle and hind legs; 645, abdominal terga and sterna; 646, anterior side of hind femur; 647, anterior side of fore tibia; 648, hind tibia with focus on outer-posterior row of stout setae; 649, the same with focus on outer-anterior and inner-anterior rows of stout setae." figureDoi="http://doi.org/10.5281/zenodo.3737062" httpUri="https://zenodo.org/record/3737062/files/figure.png" pageId="133" pageNumber="134">Fig. 646</figureCitation>
). Row of hairs near inner margin absent on fore and middle femora, sparse on hind femur.
</paragraph>
<paragraph id="8B8136B0FF1884E18CABF898DF90F842" blockId="133.[151,1437,1377,2016]" pageId="133" pageNumber="134">
Fore tibia (
<figureCitation id="13052A35FF1884E18D2AF898DA95F83B" box="[326,427,1918,1945]" captionStart="FIGURES 637–649" captionStartId="134.[151,264,1864,1889]" captionTargetBox="[151,1436,196,1765]" captionTargetId="figure@134.[151,1436,196,1765]" captionTargetPageId="134" captionText="FIGURES 637–649. Thraulodes sp. «Itaya», larva. 637, tergalius III; 638–645, larval exuviae at the same magnification: 638, part of head with labrum; 639–640, left and right mandibles; 641, right half of pronotum and mesonotum; 642–644, fore, middle and hind legs; 645, abdominal terga and sterna; 646, anterior side of hind femur; 647, anterior side of fore tibia; 648, hind tibia with focus on outer-posterior row of stout setae; 649, the same with focus on outer-anterior and inner-anterior rows of stout setae." figureDoi="http://doi.org/10.5281/zenodo.3737062" httpUri="https://zenodo.org/record/3737062/files/figure.png" pageId="133" pageNumber="134">Fig. 647</figureCitation>
): outer hairs form two irregular rows; inner-anterior row of recurved hairs absent; inneranterior row of stout setae represented by few (2–3) stout blunt setae in proximal part of tibia; inner field of stout pointed setae consists of pointed smooth setae, situated irregularly, 1–2 setae in cross section.
</paragraph>
<caption id="DF416638FF1B84E28CFBF8AEDBEFF852" ID-DOI="http://doi.org/10.5281/zenodo.3737062" ID-Zenodo-Dep="3737062" httpUri="https://zenodo.org/record/3737062/files/figure.png" pageId="134" pageNumber="135" startId="134.[151,264,1864,1889]" targetBox="[151,1436,196,1765]" targetPageId="134">
<paragraph id="8B8136B0FF1B84E28CFBF8AEDBEFF852" blockId="134.[151,1436,1864,2032]" pageId="134" pageNumber="135">
<emphasis id="B94AEAA2FF1B84E28CFBF8AEDA50F8C3" bold="true" box="[151,366,1864,1889]" pageId="134" pageNumber="135">FIGURES 637–649.</emphasis>
<taxonomicName id="4C3E4D33FF1B84E28D18F8AEDAD9F8C2" authorityName="Ulmer" authorityYear="1920" box="[372,487,1864,1888]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="134" pageNumber="135" phylum="Arthropoda" rank="genus">
<emphasis id="B94AEAA2FF1B84E28D18F8AEDAD9F8C2" box="[372,487,1864,1888]" italics="true" pageId="134" pageNumber="135">Thraulodes</emphasis>
</taxonomicName>
sp. «Itaya», larva. 637, tergalius III; 638–645, larval exuviae at the same magnification: 638, part of head with labrum; 639–640, left and right mandibles; 641, right half of pronotum and mesonotum; 642–644, fore, middle and hind legs; 645, abdominal terga and sterna; 646, anterior side of hind femur; 647, anterior side of fore tibia; 648, hind tibia with focus on outer-posterior row of stout setae; 649, the same with focus on outer-anterior and inner-anterior rows of stout setae.
</paragraph>
</caption>
<paragraph id="8B8136B0FF1A84E38CABFF7FD915FEBD" blockId="135.[151,1436,153,575]" pageId="135" pageNumber="136">
Hind tibia (
<figureCitation id="13052A35FF1A84E38D20FF7FDAD0FF16" box="[332,494,153,180]" captionStart="FIGURES 637–649" captionStartId="134.[151,264,1864,1889]" captionTargetBox="[151,1436,196,1765]" captionTargetId="figure@134.[151,1436,196,1765]" captionTargetPageId="134" captionText="FIGURES 637–649. Thraulodes sp. «Itaya», larva. 637, tergalius III; 638–645, larval exuviae at the same magnification: 638, part of head with labrum; 639–640, left and right mandibles; 641, right half of pronotum and mesonotum; 642–644, fore, middle and hind legs; 645, abdominal terga and sterna; 646, anterior side of hind femur; 647, anterior side of fore tibia; 648, hind tibia with focus on outer-posterior row of stout setae; 649, the same with focus on outer-anterior and inner-anterior rows of stout setae." figureDoi="http://doi.org/10.5281/zenodo.3737062" httpUri="https://zenodo.org/record/3737062/files/figure.png" pageId="135" pageNumber="136">Figs 648–649</figureCitation>
): outer-anterior and outer-posterior rows of stout setae consist of blunt stout setae of various lengths and long spoon-like stout setae; posterior hairs located between these rows form more than one row (besides row of hairs posteriad of outer-posterior row of stout setae); stout setae of inner-anterior row elongate, arched, narrowed distally and blunt.
</paragraph>
<paragraph id="8B8136B0FF1A84E38CABFECFDF8CFEE1" blockId="135.[151,1436,153,575]" box="[199,1202,296,323]" pageId="135" pageNumber="136">Claws with 4–5 denticles on rigid portion, often without denticles on articulatory portion.</paragraph>
<paragraph id="8B8136B0FF1A84E38CABFEABD8E2FE0D" blockId="135.[151,1436,153,575]" pageId="135" pageNumber="136">
Tergalii (
<figureCitation id="13052A35FF1A84E38D41FEABDAAEFECA" box="[301,400,333,360]" captionStart="FIGURES 637–649" captionStartId="134.[151,264,1864,1889]" captionTargetBox="[151,1436,196,1765]" captionTargetId="figure@134.[151,1436,196,1765]" captionTargetPageId="134" captionText="FIGURES 637–649. Thraulodes sp. «Itaya», larva. 637, tergalius III; 638–645, larval exuviae at the same magnification: 638, part of head with labrum; 639–640, left and right mandibles; 641, right half of pronotum and mesonotum; 642–644, fore, middle and hind legs; 645, abdominal terga and sterna; 646, anterior side of hind femur; 647, anterior side of fore tibia; 648, hind tibia with focus on outer-posterior row of stout setae; 649, the same with focus on outer-anterior and inner-anterior rows of stout setae." figureDoi="http://doi.org/10.5281/zenodo.3737062" httpUri="https://zenodo.org/record/3737062/files/figure.png" pageId="135" pageNumber="136">Fig. 637</figureCitation>
): wide, but with few side trachea; dorsal lamella oval, widest in distal part, apically abruptly turned to slender apical filament; ventral lamella widest near base, gradually narrowed toward apex, with slender apical filament, with main trachea arched repeating outline of anal margin.
</paragraph>
<paragraph id="8B8136B0FF1A84E38CABFE5ED969FE71" blockId="135.[151,1436,153,575]" box="[199,599,440,467]" pageId="135" pageNumber="136">
<emphasis id="B94AEAA2FF1A84E38CABFE5EDAE4FE71" bold="true" box="[199,474,440,467]" italics="true" pageId="135" pageNumber="136">Subimago, imago, eggs.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C324653BFF1A84E38CABFE3ADA6DFD9D" pageId="135" pageNumber="136" type="discussion">
<paragraph id="8B8136B0FF1A84E38CABFE3ADA6DFD9D" blockId="135.[151,1436,153,575]" pageId="135" pageNumber="136">
<emphasis id="B94AEAA2FF1A84E38CABFE3ADA72FE55" bold="true" box="[199,332,476,503]" pageId="135" pageNumber="136">Discussion.</emphasis>
By cuticular coloration of larval femora and structure of tergalii this species resembles
<taxonomicName id="4C3E4D33FF1A84E38958FE3BDBECFDB9" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="135" pageNumber="136" phylum="Arthropoda" rank="species" species="alboniger" status="sp. nov.">
<emphasis id="B94AEAA2FF1A84E38958FE3BDBECFDB9" italics="true" pageId="135" pageNumber="136">Th. alboniger</emphasis>
</taxonomicName>
<emphasis id="B94AEAA2FF1A84E38CB0FDE7DA20FDB9" bold="true" box="[220,286,513,539]" pageId="135" pageNumber="136">
<taxonomicNameLabel id="A27957D9FF1A84E38CB0FDE7DA20FDB9" box="[220,286,513,539]" pageId="135" pageNumber="136" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, but clearly differs from
<taxonomicName id="4C3E4D33FF1A84E38E2FFDE7D9E0FDB9" box="[579,734,513,539]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="135" pageNumber="136" phylum="Arthropoda" rank="species" species="alboniger">
<emphasis id="B94AEAA2FF1A84E38E2FFDE7D9E0FDB9" box="[579,734,513,539]" italics="true" pageId="135" pageNumber="136">Th. alboniger</emphasis>
</taxonomicName>
by structure of labrum and absence of contrasting blanks on abdominal terga.
</paragraph>
</subSubSection>
</treatment>
</document>
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