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<mods:title id="F9D663A13C9837DDB0264F542AFD7CA9">Phylogenetics and comparative morphology of crab spiders (Araneae: Dionycha, Thomisidae)</mods:title>
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<treatment id="03978793FFB5E13341E6FCA8FCC90B36" ID-DOI="http://doi.org/10.5281/zenodo.6186489" ID-GBIF-Taxon="137306782" ID-Zenodo-Dep="6186489" LSID="urn:lsid:plazi:treatment:03978793FFB5E13341E6FCA8FCC90B36" httpUri="http://treatment.plazi.org/id/03978793FFB5E13341E6FCA8FCC90B36" lastPageNumber="11" pageId="10" pageNumber="11">
<subSubSection id="C324650EFFB5E13341E6FCA8FDCA0D66" box="[151,555,825,850]" pageId="10" pageNumber="11" type="nomenclature">
<paragraph id="8B813685FFB5E13341E6FCA8FDCA0D66" blockId="10.[151,555,825,850]" box="[151,555,825,850]" pageId="10" pageNumber="11">
<emphasis id="B94AEA97FFB5E13341E6FCA8FF020D65" bold="true" box="[151,227,825,850]" pageId="10" pageNumber="11">Genus</emphasis>
<taxonomicName id="4C3E4D06FFB5E133419BFCA8FDCA0D66" ID-CoL="62GXC" ID-ENA="635227" authority="Simon, 1884" authorityName="Simon" authorityYear="1884" box="[234,555,825,850]" class="Arachnida" family="Thomisidae" genus="Borboropactus" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B94AEA97FFB5E133419BFCA8FE740D65" bold="true" box="[234,405,825,850]" italics="true" pageId="10" pageNumber="11">Borboropactus</emphasis>
<bibRefCitation id="EFAF4B74FFB5E13340EDFCA8FDCA0D66" author="Simon" box="[412,555,825,849]" pageId="10" pageNumber="11" refString="Simon, E. (1884) Description d'une nouvelle famille de l'ordre des Araneae (Bradystichidae). Annales de la Societe entomologique de Belgique. Comptes Rendus [Les comptes Rendus sont reunis en un bulletin, place apres les Annales], 28, 297 - 301." type="journal article" year="1884">Simon, 1884</bibRefCitation>
</taxonomicName>
</paragraph>
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<subSubSection id="C324650EFFB5E13341E6FC13FF130D8A" pageId="10" pageNumber="11" type="materials_examined">
<paragraph id="8B813685FFB5E13341E6FC13FF130D8A" blockId="10.[151,747,897,957]" pageId="10" pageNumber="11">
<emphasis id="B94AEA97FFB5E13341E6FC13FED50DAD" bold="true" box="[151,308,897,922]" pageId="10" pageNumber="11">
<typeStatus id="54858827FFB5E13341E6FC13FF330DAD" box="[151,210,898,922]" pageId="10" pageNumber="11">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="4C3E4D06FFB5E133404AFC10FD0A0DAE" authority="Simon, 1884" authorityName="Simon" authorityYear="1884" box="[315,747,897,921]" class="Arachnida" family="Thomisidae" genus="Borboropactus" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="squalidus">
<emphasis id="B94AEA97FFB5E133404AFC10FDB40DAE" box="[315,597,897,921]" italics="true" pageId="10" pageNumber="11">Borboropactus squalidus</emphasis>
<bibRefCitation id="EFAF4B74FFB5E133432DFC10FD0A0DAE" author="Simon" box="[604,747,897,921]" pageId="10" pageNumber="11" refString="Simon, E. (1884) Description d'une nouvelle famille de l'ordre des Araneae (Bradystichidae). Annales de la Societe entomologique de Belgique. Comptes Rendus [Les comptes Rendus sont reunis en un bulletin, place apres les Annales], 28, 297 - 301." type="journal article" year="1884">Simon, 1884</bibRefCitation>
</taxonomicName>
<figureCitation id="13052A00FFB5E13341E6FC34FF3E0D8A" box="[151,223,933,957]" captionStart="FIGURE 20. A, B, D" captionStartId="52.[151,250,1954,1977]" captionTargetBox="[157,1425,179,1925]" captionTargetId="figure@52.[151,1436,173,1933]" captionTargetPageId="52" captionText="FIGURE 20. A, B, D Borboropactus bituberculatus. C Borboropactus squalidus (MNHN 5045 / 1572). A B male palp from Irian Jaya, Waigeo Island (A prolateral, B retrolateral); C, D epigynum, ventral view (MNHN 5460 / 1572). Scale bars = 0.2 mm." httpUri="https://zenodo.org/record/1049480/files/figure.png" pageId="10" pageNumber="11">Fig 20</figureCitation>
C
</paragraph>
</subSubSection>
<subSubSection id="C324650EFFB5E13341E6FC7CFE830AEA" pageId="10" pageNumber="11" type="diagnosis">
<paragraph id="8B813685FFB5E13341E6FC7CFC9D0AA2" blockId="10.[151,1436,1005,1282]" pageId="10" pageNumber="11">
<emphasis id="B94AEA97FFB5E13341E6FC7CFEF00A31" bold="true" box="[151,273,1005,1030]" pageId="10" pageNumber="11">Diagnosis.</emphasis>
<taxonomicName id="4C3E4D06FFB5E1334069FC7CFE5E0A32" box="[280,447,1005,1029]" class="Arachnida" family="Thomisidae" genus="Borboropactus" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B94AEA97FFB5E1334069FC7CFE5E0A32" box="[280,447,1005,1029]" italics="true" pageId="10" pageNumber="11">Borboropactus</emphasis>
</taxonomicName>
can be diagnosed by the presence of an epigynal plate or folds (
<figureCitation id="13052A00FFB5E13345FEFC7CFB050A32" box="[1167,1252,1005,1029]" captionStart="FIGURE 18" captionStartId="50.[151,250,1949,1972]" captionTargetBox="[154,1433,197,1908]" captionTargetId="figure@50.[151,1436,193,1928]" captionTargetPageId="50" captionText="FIGURE 18. Borboropactus cinerascens (MHNG). A. male palp (A ventral, B prolateral, C retrolateral); D epigynum, ventral view; E vulva, ventral view. Scale bars = 0.2 mm (D, E), 0.5 mm (A C)." httpUri="https://zenodo.org/record/1049476/files/figure.png" pageId="10" pageNumber="11">Figs 18</figureCitation>
D, 19C, 20C, D, 21A, B, 23A, B), the unique sensory patch on tarsi (
<figureCitation id="13052A00FFB5E13343AAFB80FCD10A1E" box="[731,816,1041,1065]" captionStart="FIGURE 24" captionStartId="56.[151,250,1569,1592]" captionTargetBox="[161,1429,204,1539]" captionTargetId="figure@56.[151,1436,193,1548]" captionTargetPageId="56" captionText="FIGURE 24. Scanning electron micrographs of Borboropactus nyerere sp. nov. (USNM), female. A, B Chelicera; C F left leg 1 (C, D dorsal, E, F lateral). Scale bars = 10 µm (D,) 100 µm (A C, E, F)." httpUri="https://zenodo.org/record/1049488/files/figure.png" pageId="10" pageNumber="11">Figs 24</figureCitation>
CE), canoe-shaped tapetum and straight PER. Further,
<taxonomicName id="4C3E4D06FFB5E13341E6FBA4FEDF0A7A" box="[151,318,1077,1101]" class="Arachnida" family="Thomisidae" genus="Borboropactus" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B94AEA97FFB5E13341E6FBA4FEDF0A7A" box="[151,318,1077,1101]" italics="true" pageId="10" pageNumber="11">Borboropactus</emphasis>
</taxonomicName>
can be separated from all other thomisids, except for
<taxonomicName id="4C3E4D06FFB5E13342E6FBA7FC1F0A7A" box="[919,1022,1078,1101]" class="Arachnida" family="Thomisidae" genus="Geraesta" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B94AEA97FFB5E13342E6FBA7FC1F0A7A" box="[919,1022,1078,1101]" italics="true" pageId="10" pageNumber="11">Geraesta</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3E4D06FFB5E1334545FBA4FAD70A7A" authority="Thorell, 1881" authorityName="Thorell" authorityYear="1881" box="[1076,1334,1077,1101]" class="Arachnida" family="Thomisidae" genus="Angaeus" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B94AEA97FFB5E1334545FBA4FB740A7A" box="[1076,1173,1077,1101]" italics="true" pageId="10" pageNumber="11">Angaeus</emphasis>
Thorell, 1881
</taxonomicName>
, by presence of a rudimentary RTA in males and ET in females. Both
<taxonomicName id="4C3E4D06FFB5E133423CFBCBFC550A46" box="[845,948,1114,1137]" class="Arachnida" family="Thomisidae" genus="Geraesta" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B94AEA97FFB5E133423CFBCBFC550A46" box="[845,948,1114,1137]" italics="true" pageId="10" pageNumber="11">Geraesta</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3E4D06FFB5E133429AFBC8FBAD0A46" box="[1003,1100,1113,1137]" class="Arachnida" family="Thomisidae" genus="Angaeus" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B94AEA97FFB5E133429AFBC8FBAD0A46" box="[1003,1100,1113,1137]" italics="true" pageId="10" pageNumber="11">Angaeus</emphasis>
</taxonomicName>
posses a well developed RTA and sometimes DTA/VTA (
<figureCitation id="13052A00FFB5E13340BBFBECFDC00AA2" box="[458,545,1149,1173]" captionStart="FIGURE 39" captionStartId="71.[151,250,1891,1914]" captionTargetBox="[169,1416,199,1858]" captionTargetId="figure@71.[151,1436,193,1870]" captionTargetPageId="71" captionText="FIGURE 39. Geraesta hirta (USNM). A C male palp (A, C retrolateral, B ventral); D epigynum, ventral view; E vulva, ventral view. Scale bars = 0.1 mm (D, E), 0.2 mm (A C)." httpUri="https://zenodo.org/record/1049518/files/figure.png" pageId="10" pageNumber="11">Figs 39</figureCitation>
B, 39C, 40E;
<bibRefCitation id="EFAF4B74FFB5E13343C9FBEFFC910AA2" author="Tang" box="[696,880,1149,1174]" pageId="10" pageNumber="11" refString="Tang, G. &amp; Li, S. (2009) Paraborboropactus gen. nov., with description of three new species of crab spiders from Xishuangbanna, Yunnan, China (Araneae, Thomisidae). Acta Zootaxonomica Sinica, 34, 712 - 721." type="journal article" year="2009">Tang &amp; Li 2009</bibRefCitation>
).
</paragraph>
<paragraph id="8B813685FFB5E13341B6FB30FE830AEA" blockId="10.[151,1436,1005,1282]" pageId="10" pageNumber="11">
<emphasis id="B94AEA97FFB5E13341B6FB30FE860A8D" bold="true" box="[199,359,1185,1210]" pageId="10" pageNumber="11">Composition.</emphasis>
15 species, plus the two new species described below; see
<bibRefCitation id="EFAF4B74FFB5E1334569FB30FB260A8E" author="Platnick" box="[1048,1223,1185,1209]" pageId="10" pageNumber="11" refString="Platnick, N. I. (2011) The World Spiders Catalog, Version 11.5, American Museum of Natural History, New York. Available from: http: // research. amnh. org / entomology / spiders / catalog 81 - 87 / index. html (accessed 16. III. 2011)." type="journal article" year="2011">Platnick (2011)</bibRefCitation>
for a listing of all described species.
</paragraph>
</subSubSection>
<subSubSection id="C324650EFFB5E13341B6FB78FCC90B36" box="[199,808,1257,1282]" pageId="10" pageNumber="11" type="distribution">
<paragraph id="8B813685FFB5E13341B6FB78FCC90B36" blockId="10.[151,1436,1005,1282]" box="[199,808,1257,1282]" pageId="10" pageNumber="11">
<emphasis id="B94AEA97FFB5E13341B6FB78FE810B35" bold="true" box="[199,352,1257,1282]" pageId="10" pageNumber="11">Distribution.</emphasis>
Old World, from Africa to New
<collectingCountry id="F3297615FFB5E13343A1FB78FCC50B36" box="[720,804,1257,1281]" name="Guinea-Bissau" pageId="10" pageNumber="11">Guinea</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
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