221 lines
31 KiB
XML
221 lines
31 KiB
XML
<document id="DF605BFD480A82380D15B5DB92C5C2DA" ID-DOI="10.1649/0010-065X-72.mo4.1" ID-ISSN="1938-4394" ID-Zenodo-Dep="5102840" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1626277672172" checkinUser="carolina" docAuthor="Mullen, Logan J., Campbell, J. M. & Sikes, Derek S." docDate="2018" docId="03971078FFD1BC66FD39FD0DFEADCB7B" docLanguage="en" docName="TheColeopBull.mo.72.16.1-54.pdf" docOrigin="The Coleopterists Bulletin 72" docSource="http://dx.doi.org/10.1649/0010-065x-72.mo4.1" docStyle="DocumentStyle:A431C8F32F3D86E670C18C13DA9B5F2B.4:TheColeopBull.2000-2009.monograph" docStyleId="A431C8F32F3D86E670C18C13DA9B5F2B" docStyleName="TheColeopBull.2000-2009.monograph" docStyleVersion="4" docTitle="Phlaeopterus houkae Hatch 1957" docType="treatment" docVersion="3" lastPageNumber="35" masterDocId="FFAE6800FFF0BC44FFDAFFA6FFBDCE36" masterDocTitle="Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini)" masterLastPageNumber="54" masterPageNumber="1" pageNumber="34" updateTime="1699007718751" updateUser="plazi">
|
||
<mods:mods id="6461A51C5E5ECAF74DAE7E61A2D3DCC1" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="BE67B48F776C6ED8E7BD403C8B09D1EB">
|
||
<mods:title id="84DE13CA959F3ED15B16E8DA407BF942">Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="553C8FECE955A9F9EDFFC2AB076D286B" type="personal">
|
||
<mods:role id="621BE3265330BFD39F07C7E6ED9271D2">
|
||
<mods:roleTerm id="C397D512055DBA4EB1E1EFA613D91113">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="FB3EF75F083B85CEE42D310137B0CF2F">Mullen, Logan J.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="5D2392ABD7EF75DC4A6109FF30FD2446" type="personal">
|
||
<mods:role id="F6D24D7EA9C3D541F030728CC74CA0BB">
|
||
<mods:roleTerm id="1C3795B2296848AA9B2045D355E3B407">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="F9BEDABFA7800B1C793B569433B8BB94">Campbell, J. M.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="6A3D35BDC3F7E3CD18FD9566B9916071" type="personal">
|
||
<mods:role id="D663320B98165540C2F3E9FD0DACAEAF">
|
||
<mods:roleTerm id="12CC07ED4560EDA330A9A2DE91C8FDB7">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="08F0C09A9798D0CAA7E0A916ED70B4F8">Sikes, Derek S.</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="767E697E19888C61B428F7EA33D134A6">text</mods:typeOfResource>
|
||
<mods:relatedItem id="35F2B2B595640718144E328C9190AE78" type="host">
|
||
<mods:titleInfo id="526C12FC7ADFE3E0F44E9449445A86A5">
|
||
<mods:title id="A00ED1A00E53029937A1E1C65B29E978">The Coleopterists Bulletin</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="B17025229592DFA96B2B7433E3739E47">
|
||
<mods:date id="C9F4EBA5F0397465F119F19CB4C8C4AF">2018</mods:date>
|
||
<mods:detail id="DC0760522AA0E24F1C6DC7A801901550" type="pubDate">
|
||
<mods:number id="B9889745A3C0A38A3D12087F30E07373">2018-12-28</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="5BFB77BC4FB3330210B4DBA54AAD194F" type="volume">
|
||
<mods:number id="9138628C798F1F2EA7B04EFB6CCE6CE9">72</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="68B1B12028DA427190DDF63B547F97AC" unit="page">
|
||
<mods:start id="748EBF86F7CB6C6D0A04719F05794F40">1</mods:start>
|
||
<mods:end id="5133423CB9101FE7063EA687FCB7DA46">54</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location id="ECF03EDFC4E728F816A12F4C2A7F3FEB">
|
||
<mods:url id="B8EC1CD7269AB7B1A4D84996245CBD8F">http://dx.doi.org/10.1649/0010-065x-72.mo4.1</mods:url>
|
||
</mods:location>
|
||
<mods:classification id="4397E9A32CCF5B7C7AA954E49CFB11E0">journal article</mods:classification>
|
||
<mods:identifier id="BC1C1A731034B3C6447BC522353CE92E" type="DOI">10.1649/0010-065X-72.mo4.1</mods:identifier>
|
||
<mods:identifier id="85BBCB1E7D62EAD0984F476326DABADC" type="ISSN">1938-4394</mods:identifier>
|
||
<mods:identifier id="50F5B8A7D01C17B30DE1B93F1E6DB426" type="Zenodo-Dep">5102840</mods:identifier>
|
||
<mods:identifier id="27EEFBE9B73B74E31FC47D52D7C8E0AC" type="ZooBank">65F0E5A1-D396-4517-9E14-764B3073E0EF</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03971078FFD1BC66FD39FD0DFEADCB7B" LSID="urn:lsid:plazi:treatment:03971078FFD1BC66FD39FD0DFEADCB7B" httpUri="http://treatment.plazi.org/id/03971078FFD1BC66FD39FD0DFEADCB7B" lastPageId="34" lastPageNumber="35" pageId="33" pageNumber="34">
|
||
<subSubSection id="C324F2E5FFD1BC65FD39FD0DFBE3CCF6" box="[739,1118,683,704]" pageId="33" pageNumber="34" type="nomenclature">
|
||
<paragraph id="8B81A16EFFD1BC65FD39FD0DFBE3CCF6" blockId="33.[686,1172,683,758]" box="[739,1118,683,704]" pageId="33" pageNumber="34">
|
||
<emphasis id="B94A7D7CFFD1BC65FD39FD0DFBE3CCF6" bold="true" box="[739,1118,683,704]" pageId="33" pageNumber="34">
|
||
10.
|
||
<taxonomicName id="4C3EDAEDFFD1BC65FCD3FD0DFBE3CCF6" ID-CoL="76ZZZ" authority="Hatch, 1957" authorityName="Hatch" authorityYear="1957" box="[777,1118,683,704]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="houkae">
|
||
<emphasis id="B94A7D7CFFD1BC65FCD3FD0DFC66CCF6" bold="true" box="[777,987,683,704]" italics="true" pageId="33" pageNumber="34">Phlaeopterus houkae</emphasis>
|
||
<bibRefCitation id="EFAFDC9FFFD1BC65FC39FD0DFBE3CCF6" author="Hatch" box="[995,1118,683,704]" pageId="33" pageNumber="34" refId="ref28862" refString="Hatch, M. H. 1957. The Beetles of the Pacific Northwest. Part II: Staphyliniformia. University of Washington Publications in Biology, Seattle, WA." type="book" year="1957">Hatch, 1957</bibRefCitation>
|
||
</taxonomicName>
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C324F2E5FFD1BC65FD74FD60FBDACCC0" pageId="33" pageNumber="34" type="description">
|
||
<paragraph id="8B81A16EFFD1BC65FD74FD60FBDACCC0" blockId="33.[686,1172,683,758]" pageId="33" pageNumber="34">
|
||
<emphasis id="B94A7D7CFFD1BC65FD74FD60FD0BCCED" bold="true" box="[686,694,710,731]" pageId="33" pageNumber="34">(</emphasis>
|
||
<figureCitation id="1305BDEBFFD1BC65FD6FFD60FCB5CCED" box="[693,776,710,731]" captionStart="Fig" captionStartId="12.[197,228,1727,1746]" captionTargetBox="[177,1188,196,1710]" captionTargetId="figure-0@12.[175,1190,194,1712]" captionTargetPageId="12" captionText="Fig. 3. Phlaeopterus species, habitus. A) P. hatchi, male (UAMObs:Ento:232429), B) P. houkae, male (UAMObs:" figureDoi="http://doi.org/10.5281/zenodo.5102847" httpUri="https://zenodo.org/record/5102847/files/figure.png" pageId="33" pageNumber="34">Figs. 3B</figureCitation>
|
||
,
|
||
<figureCitation id="1305BDEBFFD1BC65FCC9FD60FC80CCED" box="[787,829,710,731]" captionStart="Fig" captionStartId="17.[709,740,1029,1048]" captionTargetBox="[154,1169,195,1012]" captionTargetPageId="17" captionText="Fig. 10. Distributions of Phlaeopterus species based on all georeferenced specimen records. A) P. hatchi, B) P. houkae. Grey dots represent localities where at least one Phlaeopterus species, but not the species of interest, was collected. Red dots represent localities where the species of interest was collected." figureDoi="http://doi.org/10.5281/zenodo.5102859" httpUri="https://zenodo.org/record/5102859/files/figure.png" pageId="33" pageNumber="34">10B</figureCitation>
|
||
,
|
||
<figureCitation id="1305BDEBFFD1BC65FC9DFD60FCD0CCED" box="[839,877,710,731]" captionStart="Fig" captionStartId="21.[175,206,1672,1691]" captionTargetBox="[157,1166,199,1650]" captionTargetId="figure-1@21.[154,1169,194,1656]" captionTargetPageId="21" captionText="Fig. 15. Pronota, dorsal view, of A) Lesteva pallipes LeConte, 1863, B) Lesteva longoelytrata (Goeze, 1777), C) Unamis sp. undescribed, D) Unamis sp., E) Phlaeopterus lagrandeuri, F) Phlaeopterus houkae, G) Phlaeopterus longipennis, H) Phlaeopterus obsoletus." figureDoi="http://doi.org/10.5281/zenodo.5102865" httpUri="https://zenodo.org/record/5102865/files/figure.png" pageId="33" pageNumber="34">15F</figureCitation>
|
||
,
|
||
<figureCitation id="1305BDEBFFD1BC65FCADFD60FC20CCED" box="[887,925,710,731]" captionStart="Fig" captionStartId="23.[175,206,1547,1566]" captionTargetBox="[168,1155,197,1527]" captionTargetId="figure-55@23.[164,1159,194,1532]" captionTargetPageId="23" captionText="Fig. 17. Phlaeopterus species. Pronota, dorsal view, of A) P. fusconiger, B) P. frosti, C) P. filicornis, D) P. hatchi, E) P. elongatus. Abdominal segments 4–6, dorsal view, of F) P. houkae, G) P. obsoletus, H) P. elongatus." figureDoi="http://doi.org/10.5281/zenodo.5102867" httpUri="https://zenodo.org/record/5102867/files/figure.png" pageId="33" pageNumber="34">17F</figureCitation>
|
||
,
|
||
<figureCitation id="1305BDEBFFD1BC65FC7DFD60FC70CCED" box="[935,973,710,731]" captionStart="Fig" captionStartId="25.[175,206,1235,1254]" captionTargetBox="[154,634,195,1217]" captionTargetId="figure-213@25.[153,635,194,1219]" captionTargetPageId="25" captionText="Fig. 19. Mesosterna, ventral oblique view, of A) Lesteva pallipes, B) Lesteva longoelytrata, C) Unamis sp. undescribed, D) Unamis sp., E) Phlaeopterus lagrandeuri, F) Phlaeopterus houkae." figureDoi="http://doi.org/10.5281/zenodo.5102873" httpUri="https://zenodo.org/record/5102873/files/figure.png" pageId="33" pageNumber="34">19F</figureCitation>
|
||
, 21B, 28F, 31F, 33F, 34C–D, 36B, 37F–G, 39A–B, 40F, 42D)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C324F2E5FFD1BC65FD77FCADFB3ECD63" pageId="33" pageNumber="34" type="reference_group">
|
||
<paragraph id="8B81A16EFFD1BC65FD77FCADFB3ECD63" blockId="33.[685,1172,779,853]" pageId="33" pageNumber="34">
|
||
<treatmentCitationGroup id="AB2E8640FFD1BC65FD77FCADFB3ECD63" pageId="33" pageNumber="34">
|
||
<treatmentCitation id="0A9F877FFFD1BC65FD77FCADFB8ECD16" author="Hatch" box="[685,1075,779,800]" page="58" pageId="33" pageNumber="34" year="1957">
|
||
<taxonomicName id="4C3EDAEDFFD1BC65FD77FCADFB8ECD16" ID-CoL="76ZZZ" authority="Hatch 1957: 58" authorityName="Hatch" authorityPageNumber="58" authorityYear="1957" box="[685,1075,779,800]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="houkae">
|
||
<emphasis id="B94A7D7CFFD1BC65FD77FCADFC3DCD16" box="[685,896,779,800]" italics="true" pageId="33" pageNumber="34">Phlaeopterus houkae</emphasis>
|
||
<bibRefCitation id="EFAFDC9FFFD1BC65FC55FCADFB8ECD16" author="Hatch" box="[911,1075,779,800]" pageId="33" pageNumber="34" refId="ref28862" refString="Hatch, M. H. 1957. The Beetles of the Pacific Northwest. Part II: Staphyliniformia. University of Washington Publications in Biology, Seattle, WA." type="book" year="1957">Hatch 1957: 58</bibRefCitation>
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
[original description]. Campbell and Davies 1991: 5;
|
||
<treatmentCitation id="0A9F877FFFD1BC65FD12FCE6FC3CCD63" author="Herman" box="[712,897,832,853]" page="377" pageId="33" pageNumber="34" year="2001">
|
||
<bibRefCitation id="EFAFDC9FFFD1BC65FD12FCE6FC3CCD63" author="Herman" box="[712,897,832,853]" pageId="33" pageNumber="34" pagination="1 - 650" refId="ref28893" refString="Herman, L. H. 2001. Catalog of the Staphylinidae (Insecta, Coleoptera): 1758 to the end of the Second Millennium. I. Introduction, history, biogeographical sketches, and Omaliine group. Bulletin of the American Museum of Natural History 265: 1 - 650." type="journal article" year="2001">Herman 2001: 377</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0A9F877FFFD1BC65FC56FCE6FBC2CD63" author="Bousquet" box="[908,1151,832,853]" page="89" pageId="33" pageNumber="34" year="2013">
|
||
<bibRefCitation id="EFAFDC9FFFD1BC65FC56FCE6FBC2CD63" author="Bousquet" box="[908,1151,832,853]" pageId="33" pageNumber="34" refId="ref28340" refString="Bousquet, Y., P. Bouchard, A. E. Davies, and D. S. Sikes. 2013. Checklist of Beetles (Coleoptera) of Canada and Alaska. Second Edition. Pensoft, Sofia, Bulgaria." type="book" year="2013">
|
||
Bousquet
|
||
<emphasis id="B94A7D7CFFD1BC65FC37FCE6FBA2CD63" box="[1005,1055,832,853]" italics="true" pageId="33" pageNumber="34">et al.</emphasis>
|
||
2013: 89
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C324F2E5FFD1BC65FD1FFCD3FB3FCDBC" box="[709,1154,885,906]" pageId="33" pageNumber="34" type="materials_examined">
|
||
<paragraph id="8B81A16EFFD1BC65FD1FFCD3FB3FCDBC" blockId="33.[685,1173,885,1730]" box="[709,1154,885,906]" pageId="33" pageNumber="34">
|
||
<materialsCitation id="3B56AB33FFD1BC65FD1FFCD3FBC3CDBC" box="[709,1150,885,906]" country="United States of America" location="Mt. Baker" pageId="33" pageNumber="34" specimenCount="1" stateProvince="Washington" typeStatus="holotype">
|
||
<emphasis id="B94A7D7CFFD1BC65FD1FFCD3FCEBCDBC" bold="true" box="[709,854,885,906]" pageId="33" pageNumber="34">
|
||
<typeStatus id="54851FCCFFD1BC65FD1FFCD3FD4ACDBC" box="[709,759,885,906]" pageId="33" pageNumber="34">Type</typeStatus>
|
||
Locality.
|
||
</emphasis>
|
||
<location id="8EE1F7B5FFD1BC65FC84FCD3FC7CCDBC" LSID="urn:lsid:plazi:treatment:03971078FFD1BC66FD39FD0DFEADCB7B:8EE1F7B5FFD1BC65FC84FCD3FC7CCDBC" box="[862,961,885,906]" country="United States of America" name="Mt. Baker" pageId="33" pageNumber="34" stateProvince="Washington">Mt. Baker</location>
|
||
,
|
||
<collectingRegion id="49FA6F8CFFD1BC65FC17FCD3FBFCCDBC" box="[973,1089,885,906]" country="United States of America" name="Washington" pageId="33" pageNumber="34">Washington</collectingRegion>
|
||
,
|
||
<collectingCountry id="F329E1FEFFD1BC65FB91FCD3FBC3CDBC" box="[1099,1150,885,906]" name="United States of America" pageId="33" pageNumber="34">USA</collectingCountry>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C324F2E5FFD1BC66FD1FFC36FDDFCC53" lastPageId="34" lastPageNumber="35" pageId="33" pageNumber="34" type="description">
|
||
<paragraph id="8B81A16EFFD1BC66FD1FFC36FDDFCC53" blockId="33.[685,1173,885,1730]" lastBlockId="34.[173,660,193,1357]" lastPageId="34" lastPageNumber="35" pageId="33" pageNumber="34">
|
||
<emphasis id="B94A7D7CFFD1BC65FD1FFC36FC06CD93" bold="true" box="[709,955,912,933]" pageId="33" pageNumber="34">Redescription. Habitus:</emphasis>
|
||
Length 3.1–4.0 mm. Dark brown; elytra usually lighter; legs, antennae, and mouthparts yellowish brown; abdomen usually reddish brown (
|
||
<figureCitation id="1305BDEBFFD1BC65FC93FC46FC2ACDC3" box="[841,919,992,1013]" captionStart="Fig" captionStartId="12.[197,228,1727,1746]" captionTargetBox="[177,1188,196,1710]" captionTargetId="figure-0@12.[175,1190,194,1712]" captionTargetPageId="12" captionText="Fig. 3. Phlaeopterus species, habitus. A) P. hatchi, male (UAMObs:Ento:232429), B) P. houkae, male (UAMObs:" figureDoi="http://doi.org/10.5281/zenodo.5102847" httpUri="https://zenodo.org/record/5102847/files/figure.png" pageId="33" pageNumber="34">Fig. 3B</figureCitation>
|
||
).
|
||
<emphasis id="B94A7D7CFFD1BC65FC71FC46FC54CDC3" bold="true" box="[939,1001,992,1013]" pageId="33" pageNumber="34">Head:</emphasis>
|
||
Width moderate, width across eyes to head length slightly wider than long (
|
||
<figureCitation id="1305BDEBFFD1BC65FCFDFBB3FC1ACA1C" box="[807,935,1044,1066]" captionStart="Fig" captionStartId="41.[175,206,1236,1255]" captionTargetBox="[154,634,195,1220]" captionTargetId="figure-493@41.[153,635,194,1221]" captionTargetPageId="41" captionText="Fig. 34. Phlaeopterus species, head, dorsal and dorsal oblique views. A–B) P. lagrandeuri, C–D) P. houkae, E–F) P. obsoletus." figureDoi="http://doi.org/10.5281/zenodo.5102897" httpUri="https://zenodo.org/record/5102897/files/figure.png" pageId="33" pageNumber="34">Fig. 34C–D</figureCitation>
|
||
). Interantennal groove vague or absent at least across midline of head. Anteocellar foveae vague, shallowly impressed. Eyes moderately pubescent on ventral half, with more than 10 setae (
|
||
<figureCitation id="1305BDEBFFD1BC65FC56FBD9FC50CAA2" box="[908,1005,1151,1172]" captionStart="Fig" captionStartId="42.[731,762,1272,1291]" captionTargetBox="[711,1189,196,1255]" captionTargetId="figure-251@42.[709,1191,194,1257]" captionTargetPageId="42" captionText="Fig. 36. Phlaeopterus species, eyes, dorsal view. A) P. lagrandeuri, B) P. houkae, C) P. longipennis, D) P. frosti, E) P. bakerensis, F) P. olympicus." figureDoi="http://doi.org/10.5281/zenodo.5102901" httpUri="https://zenodo.org/record/5102901/files/figure.png" pageId="33" pageNumber="34">Fig. 36B</figureCitation>
|
||
). Antennomeres 3–10 each 1.6–2.0 times longer than wide; antennomeres 4–11 each with many sensory pits with papilliform projections (
|
||
<figureCitation id="1305BDEBFFD1BC65FC0AFB69FBFBCAD2" box="[976,1094,1230,1252]" captionStart="Fig" captionStartId="43.[175,206,1294,1313]" captionTargetBox="[156,1167,196,1276]" captionTargetId="figure-165@43.[154,1169,194,1278]" captionTargetPageId="43" captionText="Fig. 37. Sensory structures on antenna of A) Lesteva pallipes, B–C) Lesteva pubescens, D–E) Unamis sp. undescribed, F–G) Phlaeopterus houkae, H–I) Phlaeopterus obsoletus, J–K) Phlaeopterus elongatus. In B, D, F, H, and J, white boxes indicate inset area that is magnified in the image to the right." figureDoi="http://doi.org/10.5281/zenodo.5102903" httpUri="https://zenodo.org/record/5102903/files/figure.png" pageId="33" pageNumber="34">Fig. 37F–G</figureCitation>
|
||
). Ocelli present. Nuchal constriction absent. Epipharynx as in Fig.
|
||
<specimenCount id="9D386AE7FFD1BC65FCFEFAA2FCEDCB2F" box="[804,848,1284,1305]" pageId="33" pageNumber="34" type="generic">33F.</specimenCount>
|
||
Hypopharynx as in
|
||
<figureCitation id="1305BDEBFFD1BC65FBEEFAA2FB2CCB2F" box="[1076,1169,1284,1305]" captionStart="Fig" captionStartId="45.[175,206,1177,1196]" captionText="Fig. 40. Hypopharynges, dorsal view, of A) Lesteva pallipes, B) Lesteva longoelytrata, C) Unamis sp. undescribed, D) Unamis sp., E) Phlaeopterus lagrandeuri, F) Phlaeopterus houkae." pageId="33" pageNumber="34">
|
||
Fig.
|
||
<specimenCount id="9D386AE7FFD1BC65FBB3FAA2FB2CCB2F" box="[1129,1169,1284,1305]" pageId="33" pageNumber="34" type="generic">40F</specimenCount>
|
||
</figureCitation>
|
||
. Mandibular molar area with L-shaped row of setae (
|
||
<figureCitation id="1305BDEBFFD1BC65FD6FFA9FFCA9CB78" box="[693,788,1337,1358]" captionStart="Fig" captionStartId="36.[197,228,1167,1186]" captionTargetBox="[176,656,195,1150]" captionTargetId="figure-220@36.[175,657,194,1151]" captionTargetPageId="36" captionText="Fig. 28. Ventral view of A) Left mandible of Lesteva longoelytrata, B) Right mandible of Lesteva pallipes, C) Right mandible of Unamis sp. undescribed, D) Right mandible of Unamis sp., E) Left mandible of Phlaeopterus lagrandeuri, F) Left mandible of Phlaeopterus houkae." figureDoi="http://doi.org/10.5281/zenodo.5102889" httpUri="https://zenodo.org/record/5102889/files/figure.png" pageId="33" pageNumber="34">Fig. 28F</figureCitation>
|
||
). Labrum with sensory pores along anterior margin only (
|
||
<figureCitation id="1305BDEBFFD1BC65FC43FAF2FC4ACB5F" box="[921,1015,1364,1385]" captionStart="Fig" captionStartId="38.[197,228,1598,1617]" captionTargetBox="[177,1188,196,1581]" captionTargetId="figure-0@38.[175,1190,194,1583]" captionTargetPageId="38" captionText="Fig. 31. Labra of A) Lesteva pallipes, B) Lesteva pubescens, C) Unamis sp. undescribed, D) Unamis sp., E) Phlaeopterus lagrandeuri, F) Phlaeopterus houkae, G) Phlaeopterus obsoletus, H) Phlaeopterus filicornis. White arrows indicate labrum." figureDoi="http://doi.org/10.5281/zenodo.5102893" httpUri="https://zenodo.org/record/5102893/files/figure.png" pageId="33" pageNumber="34">Fig. 31F</figureCitation>
|
||
). Maxilla with lacinia and galea as in
|
||
<figureCitation id="1305BDEBFFD1BC65FC4DFAC8FBADCBB5" box="[919,1040,1390,1412]" captionStart="Fig" captionStartId="44.[731,762,1213,1232]" captionTargetBox="[711,1189,196,1195]" captionTargetId="figure-182@44.[709,1191,194,1197]" captionTargetPageId="44" captionText="Fig. 39. Phlaeopterus species, maxillae, dorsal and ventral views. A–B) P. houkae, C–D) P. obsoletus, E–F) P. elongatus." figureDoi="http://doi.org/10.5281/zenodo.5102905" httpUri="https://zenodo.org/record/5102905/files/figure.png" pageId="33" pageNumber="34">Fig. 39A–B</figureCitation>
|
||
. Labial palpi with 3
|
||
<superScript id="7C4B0C26FFD1BC65FD36FA25FD46CBA7" attach="left" box="[748,763,1411,1425]" fontSize="6" pageId="33" pageNumber="34">rd</superScript>
|
||
palpomere 1.2–1.5 times longer than 2
|
||
<superScript id="7C4B0C26FFD1BC65FB5BFA25FB2ECBA7" attach="none" box="[1153,1171,1411,1425]" fontSize="6" pageId="33" pageNumber="34">nd</superScript>
|
||
palpomere. Gula as in
|
||
<figureCitation id="1305BDEBFFD1BC65FC46FA05FC41CB8E" box="[924,1020,1443,1464]" captionStart="Fig" captionStartId="46.[197,228,1197,1216]" captionTargetBox="[176,656,195,1180]" captionTargetId="figure-522@46.[175,657,194,1181]" captionTargetPageId="46" captionText="Fig. 42. Heads, ventral view, of A) Unamis sp. undescribed, B) Unamis sp., C) Phlaeopterus lagrandeuri, D) Phlaeopterus houkae, E) Phlaeopterus longipennis, F) Phlaeopterus elongatus." figureDoi="http://doi.org/10.5281/zenodo.5102911" httpUri="https://zenodo.org/record/5102911/files/figure.png" pageId="33" pageNumber="34">Fig. 42D</figureCitation>
|
||
.
|
||
<emphasis id="B94A7D7CFFD1BC65FBD3FA05FBE2CB8E" bold="true" box="[1033,1119,1443,1464]" pageId="33" pageNumber="34">Thorax:</emphasis>
|
||
Pronotum narrow (
|
||
<figureCitation id="1305BDEBFFD1BC65FC9CFA18FC23CBE5" box="[838,926,1470,1491]" captionStart="Fig" captionStartId="21.[175,206,1672,1691]" captionTargetBox="[157,1166,199,1650]" captionTargetId="figure-1@21.[154,1169,194,1656]" captionTargetPageId="21" captionText="Fig. 15. Pronota, dorsal view, of A) Lesteva pallipes LeConte, 1863, B) Lesteva longoelytrata (Goeze, 1777), C) Unamis sp. undescribed, D) Unamis sp., E) Phlaeopterus lagrandeuri, F) Phlaeopterus houkae, G) Phlaeopterus longipennis, H) Phlaeopterus obsoletus." figureDoi="http://doi.org/10.5281/zenodo.5102865" httpUri="https://zenodo.org/record/5102865/files/figure.png" pageId="33" pageNumber="34">Fig. 15F</figureCitation>
|
||
), length to width ratio = 0.7–0.8; ratio of pronotal width to head width = 1.3–1.4; maximum width subequal to width of elytra at bases; punctures separated by average distance equal to slightly less than diameter of a puncture; lateral margins narrowly explanate posterad lateral fovea, not explanate anterad lateral foveae; lateral foveae moderately impressed, contiguous with lateral margins. Elytra with humeral angles convex; epipleural carina not projecting; 1.9–2.1 times longer than pronotum; apical margins convex. Wings brachypterous. Mesosternum with projecting tooth; longitudinal carina along midline of mesosternum present but vague, especially near midline and posterior margin (
|
||
<figureCitation id="1305BDEBFFD2BC66FEDEFEE0FEDDCF6D" box="[260,352,326,347]" captionStart="Fig" captionStartId="25.[175,206,1235,1254]" captionTargetBox="[154,634,195,1217]" captionTargetId="figure-213@25.[153,635,194,1219]" captionTargetPageId="25" captionText="Fig. 19. Mesosterna, ventral oblique view, of A) Lesteva pallipes, B) Lesteva longoelytrata, C) Unamis sp. undescribed, D) Unamis sp., E) Phlaeopterus lagrandeuri, F) Phlaeopterus houkae." figureDoi="http://doi.org/10.5281/zenodo.5102873" httpUri="https://zenodo.org/record/5102873/files/figure.png" pageId="34" pageNumber="35">Fig. 19F</figureCitation>
|
||
).
|
||
<emphasis id="B94A7D7CFFD2BC66FEADFEE0FE12CF6D" bold="true" box="[375,431,326,347]" pageId="34" pageNumber="35">Legs:</emphasis>
|
||
All tibiae evenly pubescent to apex. Metatrochanter without tooth on apical margin.
|
||
<emphasis id="B94A7D7CFFD2BC66FEEBFEDAFE24CFA7" bold="true" box="[305,409,380,401]" pageId="34" pageNumber="35">Abdomen:</emphasis>
|
||
Tergites IVand Veach with wing-folding spicules very broad, nearly contiguous or contiguous in shape of single transverse band (
|
||
<figureCitation id="1305BDEBFFD2BC66FDB4FE17FF6ECFD6" captionStart="Fig" captionStartId="23.[175,206,1547,1566]" captionTargetBox="[168,1155,197,1527]" captionTargetId="figure-55@23.[164,1159,194,1532]" captionTargetPageId="23" captionText="Fig. 17. Phlaeopterus species. Pronota, dorsal view, of A) P. fusconiger, B) P. frosti, C) P. filicornis, D) P. hatchi, E) P. elongatus. Abdominal segments 4–6, dorsal view, of F) P. houkae, G) P. obsoletus, H) P. elongatus." figureDoi="http://doi.org/10.5281/zenodo.5102867" httpUri="https://zenodo.org/record/5102867/files/figure.png" pageId="34" pageNumber="35">Fig. 17F</figureCitation>
|
||
). Apical palisade fringe on tergite VII absent.
|
||
<emphasis id="B94A7D7CFFD2BC66FF77FE40FEAFCFCD" bold="true" box="[173,274,486,507]" pageId="34" pageNumber="35">Aedeagus:</emphasis>
|
||
Length
|
||
<quantity id="4CC60C8BFFD2BC66FE85FE40FE55CFCD" box="[351,488,485,507]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" metricValueMax="7.5" metricValueMin="6.5" pageId="34" pageNumber="35" unit="mm" value="0.7" valueMax="0.75" valueMin="0.65">0.65–0.75 mm</quantity>
|
||
. Median lobe with lateral margins subparallel from base to past midline, then abruptly narrowed, apex acute (Fig. 21B). Parameres broad, parallel and nearly straight. Internal sac with 2 dense patches of spicules lateroapically.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C324F2E5FFD2BC66FF1FFDCDFD82CD0C" pageId="34" pageNumber="35" type="materials_examined">
|
||
<paragraph id="8B81A16EFFD2BC66FF1FFDCDFD82CD0C" blockId="34.[173,660,193,1357]" pageId="34" pageNumber="35">
|
||
<emphasis id="B94A7D7CFFD2BC66FF1FFDCDFED7CCB6" bold="true" box="[197,362,619,640]" pageId="34" pageNumber="35">Type Specimen.</emphasis>
|
||
<typeStatus id="54851FCCFFD2BC66FEACFDCDFE73CCB6" box="[374,462,619,640]" pageId="34" pageNumber="35" type="holotype">Holotype</typeStatus>
|
||
female (UAMObs: Ento:235216) labeled as follows: Mt. Baker Lodge,
|
||
<date id="FF8087AEFFD2BC66FF77FD06FE84CC83" box="[173,313,672,693]" pageId="34" pageNumber="35" value="1935-08-26">Aug. 26, 1935</date>
|
||
, WASH., M.H. Hatch/Det. 1954, H. Houk/not
|
||
<taxonomicName id="4C3EDAEDFFD2BC66FED4FD1CFE47CCF9" authority="Csy." authorityName="Casey" authorityYear="1885" box="[270,506,698,719]" class="Insecta" family="Staphylinidae" genus="Vellica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="longipennis">Vellica longipennis Csy.</taxonomicName>
|
||
, compared with type, M.H. Hatch 1958/TYPE
|
||
<specimenCount id="9D386AE7FFD2BC66FE21FD71FDB5CCDF" box="[507,520,727,745]" pageId="34" pageNumber="35" type="female">♀</specimenCount>
|
||
<taxonomicName id="4C3EDAEDFFD2BC66FDC3FD73FE92CD33" authorityName="Hatch" authorityYear="1957" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="houkae">Phlaeopterus houkae 1951</taxonomicName>
|
||
– M.H. Hatch/
|
||
<taxonomicName id="4C3EDAEDFFD2BC66FE0EFD56FF64CD29" authority="Csy." authorityName="Casey" authorityYear="1885" class="Insecta" family="Staphylinidae" genus="Vellica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="longipennis">Vellica longipennis Csy.</taxonomicName>
|
||
, compared with Fall coll., M.H. Hatch 1952. The specimen is deposited in the USNM.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C324F2E5FFD2BC66FF1FFC99FEACCA68" pageId="34" pageNumber="35" type="distribution">
|
||
<paragraph id="8B81A16EFFD2BC66FF1FFC99FE24CDC2" blockId="34.[173,660,193,1357]" pageId="34" pageNumber="35">
|
||
<emphasis id="B94A7D7CFFD2BC66FF1FFC99FEFBCD62" bold="true" box="[197,326,831,852]" pageId="34" pageNumber="35">Distribution.</emphasis>
|
||
<taxonomicName id="4C3EDAEDFFD2BC66FE8EFC99FD9BCD62" authorityName="Hatch" authorityYear="1957" box="[340,550,831,852]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="houkae">
|
||
<emphasis id="B94A7D7CFFD2BC66FE8EFC99FD9BCD62" box="[340,550,831,852]" italics="true" pageId="34" pageNumber="35">Phlaeopterus houkae</emphasis>
|
||
</taxonomicName>
|
||
is known broadly from the Pacific Coast and Cascade Ranges, from Siskiyou County, California and Oregon north to the Alaska Range in Alaska (
|
||
<figureCitation id="1305BDEBFFD2BC66FE01FC29FD8FCD92" box="[475,562,911,932]" captionStart="Fig" captionStartId="17.[709,740,1029,1048]" captionTargetBox="[154,1169,195,1012]" captionTargetPageId="17" captionText="Fig. 10. Distributions of Phlaeopterus species based on all georeferenced specimen records. A) P. hatchi, B) P. houkae. Grey dots represent localities where at least one Phlaeopterus species, but not the species of interest, was collected. Red dots represent localities where the species of interest was collected." figureDoi="http://doi.org/10.5281/zenodo.5102859" httpUri="https://zenodo.org/record/5102859/files/figure.png" pageId="34" pageNumber="35">Fig. 10B</figureCitation>
|
||
). It is also known coastally from Baranof Island, Prince of
|
||
<collectingCountry id="F329E1FEFFD2BC66FF77FC62FE97CDEF" box="[173,298,964,985]" name="Wales Island" pageId="34" pageNumber="35">Wales Island</collectingCountry>
|
||
, Haida Gwaii, Vancouver Island, and the Olympic Mountains.
|
||
</paragraph>
|
||
<paragraph id="8B81A16EFFD2BC66FF1FFC5FFEACCA68" blockId="34.[173,660,193,1357]" pageId="34" pageNumber="35">
|
||
<emphasis id="B94A7D7CFFD2BC66FF1FFC5FFE89CA38" bold="true" box="[197,308,1017,1038]" pageId="34" pageNumber="35">Bionomics.</emphasis>
|
||
Adults have been collected at
|
||
<quantity id="4CC60C8BFFD2BC66FF77FBB2FE9CCA1F" box="[173,289,1043,1065]" metricMagnitude="3" metricUnit="m" metricValue="1.2449999999999999" metricValueMax="2.4" metricValueMin="0.09" pageId="34" pageNumber="35" unit="m" value="1245.0" valueMax="2400.0" valueMin="90.0">90–2,400 m</quantity>
|
||
elevation during June–August in wet moss at the edge of cold water, usually small streams and pools.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C324F2E5FFD2BC66FF1FFBC5FEADCB7B" pageId="34" pageNumber="35" type="discussion">
|
||
<paragraph id="8B81A16EFFD2BC66FF1FFBC5FEADCB7B" blockId="34.[173,660,193,1357]" pageId="34" pageNumber="35">
|
||
<emphasis id="B94A7D7CFFD2BC66FF1FFBC5FE9ACA4E" bold="true" box="[197,295,1123,1144]" pageId="34" pageNumber="35">Remarks.</emphasis>
|
||
<taxonomicName id="4C3EDAEDFFD2BC66FEEBFBC5FDBDCA4E" authorityName="Hatch" authorityYear="1957" box="[305,512,1123,1144]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="houkae">
|
||
<emphasis id="B94A7D7CFFD2BC66FEEBFBC5FDBDCA4E" box="[305,512,1123,1144]" italics="true" pageId="34" pageNumber="35">Phlaeopterus houkae</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from all other
|
||
<taxonomicName id="4C3EDAEDFFD2BC66FE40FBD8FDCFCAA5" box="[410,626,1150,1171]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B94A7D7CFFD2BC66FE40FBD8FD9DCAA5" box="[410,544,1150,1171]" italics="true" pageId="34" pageNumber="35">Phlaeopterus</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
by its small size, presence of ocelli, vague anteocellar foveae, slight nuchal constriction, shape and punctation of the pronotum, shape of the wing-folding spicules on tergites IV and V, lack of wing-folding spicules on tergite VI, and the unique structure of the internal sac of the aedeagus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |