283 lines
39 KiB
XML
283 lines
39 KiB
XML
<document id="D9FD229A59122A49FBEC515EB29565AC" ID-DOI="10.11646/zootaxa.5353.4.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="8430604" ID-ZooBank="A3794DB2-0F35-412F-B88F-C5BA9600C10A" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1697019988276" checkinUser="plazi" docAuthor="Rocha, Isabela Cristina, Santos, Allan Paulo Moreira & Nessimian, Jorge Luiz" docDate="2023" docId="FB4B87B9C34CFFBFFF7FD0E9FD88FF26" docLanguage="en" docName="zootaxa.5353.4.1.pdf" docOrigin="Zootaxa 5353 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5353.4.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rhyacopsyche escamosa Rocha & Santos & Nessimian 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="326" masterDocId="0772FFC1C35AFFA6FFE8D267FFBFFFF3" masterDocTitle="Taxonomic diversity of Ochrotrichiinae (Trichoptera: Hydroptilidae) from Peru with the description of ten new species, a new distributional record, and an updated checklist" masterLastPageNumber="331" masterPageNumber="301" pageNumber="323" updateTime="1697020540242" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="7AD6F0C6D0F23CB3D48455518CA4980A">Taxonomic diversity of Ochrotrichiinae (Trichoptera: Hydroptilidae) from Peru with the description of ten new species, a new distributional record, and an updated checklist</mods:title>
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<mods:namePart id="9AA3019091B2D9348B1E5D72CE3BD6B7">Rocha, Isabela Cristina</mods:namePart>
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<mods:affiliation id="501CECC7C18417BFEAB93B7AFF271D7E">isaabio @ hotmail. com; https: // orcid. org / 0000 - 0001 - 6514 - 9509</mods:affiliation>
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<mods:nameIdentifier id="EE2A9107E9E95A2F56FA316BFBF55617" type="email">isaabio@hotmail.com</mods:nameIdentifier>
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<mods:namePart id="929BB61FEBC472BFE32DB6FFA5D0D4C9">Santos, Allan Paulo Moreira</mods:namePart>
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<mods:nameIdentifier id="CFC288EB405056470AAC0B6C19F6D6FD" type="ORCID">0000-0001-5567-0149</mods:nameIdentifier>
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<mods:affiliation id="8F67A15D00E3F49E8E363378518BE1FB">allanpms @ gmail. com; https: // orcid. org / 0000 - 0001 - 5567 - 0149</mods:affiliation>
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<mods:nameIdentifier id="E63478C000F9EE1234505AA8BFFEB01D" type="email">allanpms@gmail.com</mods:nameIdentifier>
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<mods:namePart id="A4092CAEBCAB9258FEA789D9DAF99EC9">Nessimian, Jorge Luiz</mods:namePart>
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<mods:affiliation id="7B5E658AE13BF42AA7D13AE545A5417E">jnessimian @ gmail. com; https: // orcid. org / 0000 - 0001 - 8795 - 2538</mods:affiliation>
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<mods:nameIdentifier id="61E6E35B8D62A34770CBEEB0EF160119" type="email">jnessimian@gmail.com</mods:nameIdentifier>
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<mods:title id="C2598E39F990792C012D1F19C703DC65">Zootaxa</mods:title>
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<mods:part id="0726331D8AC428C7F4BC68C540EDEF21">
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<mods:date id="A07B4BC9C75B028F2CDB0035276DC1F3">2023</mods:date>
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<mods:number id="09C4BA12D0AB19DCD2DBC830F8A45D5A">2023-10-11</mods:number>
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<mods:url id="CA5FF6EDF2D20D5952EDA7077A181186">http://dx.doi.org/10.11646/zootaxa.5353.4.1</mods:url>
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<mods:classification id="A522A187C7B2F4A8FC72B8308A368B96">journal article</mods:classification>
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<mods:identifier id="5E02133E875C16E63E24B213BB1C4072" type="DOI">10.11646/zootaxa.5353.4.1</mods:identifier>
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<mods:identifier id="50747E6097D250503ACFB6177A90A048" type="ISSN">1175-5326</mods:identifier>
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<treatment id="FB4B87B9C34CFFBFFF7FD0E9FD88FF26" ID-DOI="http://doi.org/10.5281/zenodo.8430658" ID-Zenodo-Dep="8430658" LSID="urn:lsid:plazi:treatment:FB4B87B9C34CFFBFFF7FD0E9FD88FF26" httpUri="http://treatment.plazi.org/id/FB4B87B9C34CFFBFFF7FD0E9FD88FF26" lastPageId="25" lastPageNumber="326" pageId="22" pageNumber="323">
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<subSubSection id="3BF86524C34CFFB0FF7FD0E9FD9BFD5A" box="[151,548,654,681]" pageId="22" pageNumber="323" type="nomenclature">
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<paragraph id="735D36AFC34CFFB0FF7FD0E9FD9BFD5A" blockId="22.[151,548,654,717]" box="[151,548,654,681]" pageId="22" pageNumber="323">
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<heading id="281581C3C34CFFB0FF7FD0E9FD9BFD5A" bold="true" box="[151,548,654,681]" fontSize="11" level="1" pageId="22" pageNumber="323" reason="1">
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<emphasis id="4196EABDC34CFFB0FF7FD0E9FD9BFD5A" bold="true" box="[151,548,654,681]" pageId="22" pageNumber="323">
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<taxonomicName id="B4E24D2CC34CFFB0FF7FD0E9FE00FD5A" authority="Rocha & Santos & Nessimian, 2023" authorityName="Rocha & Santos & Nessimian" authorityYear="2023" box="[151,447,654,681]" class="Insecta" family="Hydroptilidae" genus="Rhyacopsyche" kingdom="Animalia" order="Trichoptera" pageId="22" pageNumber="323" phylum="Arthropoda" rank="species" species="escamosa" status="sp. nov.">
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<emphasis id="4196EABDC34CFFB0FF7FD0E9FE00FD5A" bold="true" box="[151,447,654,681]" italics="true" pageId="22" pageNumber="323">Rhyacopsyche escamosa</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="5AA557C6C34CFFB0FE2ED0E8FD9BFD5A" box="[454,548,655,681]" pageId="22" pageNumber="323" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="3BF86524C34CFFB0FF7FD0D4FE65F9DE" pageId="22" pageNumber="323" type="description">
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<paragraph id="735D36AFC34CFFB0FF7FD0D4FE87FD3E" blockId="22.[151,548,654,717]" box="[151,312,690,717]" pageId="22" pageNumber="323">
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<figureCitation id="EBD92A2AC34CFFB0FF7FD0D4FEAFFD3E" box="[151,272,691,717]" captionStart="FIGURE 14" captionStartId="23.[151,250,1809,1833]" captionTargetBox="[151,1432,190,1783]" captionTargetId="figure-20@23.[151,1436,181,1785]" captionTargetPageId="23" captionText="FIGURE 14. Rhyacopsyche escamosa sp. nov. 14A, head, dorsal. 14B, antenna. 14C, forewing, detail of costal overfold and scales. 14D, right forewing, dorsal. 14E, right hind wing, dorsal. Abbreviations: o.s.w = ocellar setal wart (paired); p.s.w = posterior setal wart (paired)." figureDoi="http://doi.org/10.5281/zenodo.8430646" httpUri="https://zenodo.org/record/8430646/files/figure.png" pageId="22" pageNumber="323">Figures 14</figureCitation>
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,
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<figureCitation id="EBD92A2AC34CFFB0FEF4D0D5FE87FD3E" box="[284,312,690,717]" captionStart="FIGURE 15" captionStartId="24.[151,250,1943,1967]" captionTargetBox="[186,1399,196,1917]" captionTargetId="figure-22@24.[175,1412,181,1919]" captionTargetPageId="24" captionText="FIGURE 15. Rhyacopsyche escamosa sp. nov., male genitalia. 15A, left lateral; 15B, dorsal; 15C, ventral; 15D, phallus, dorsal; 15E, phallus, left lateral. Abbreviations: IX = segment IX; X = tergum X." figureDoi="http://doi.org/10.5281/zenodo.8430650" httpUri="https://zenodo.org/record/8430650/files/figure.png" pageId="22" pageNumber="323">15</figureCitation>
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</paragraph>
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<paragraph id="735D36AFC34CFFB0FF7FD09DFA98FC72" blockId="22.[151,1437,762,1977]" pageId="22" pageNumber="323">
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<emphasis id="4196EABDC34CFFB0FF7FD09DFED0FCE6" bold="true" box="[151,367,762,789]" pageId="22" pageNumber="323">Description, male.</emphasis>
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<emphasis id="4196EABDC34CFFB0FE9ED09CFE0CFCE6" box="[374,435,763,789]" italics="true" pageId="22" pageNumber="323">Head</emphasis>
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. Wider than long, in dorsal view, frontal margin subtrapezoidal, ocellar setose warts large, subtriangular, posterior setose warts subtriangular with longitudinal length about 2/3 transverse width (
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<figureCitation id="EBD92A2AC34CFFB0FACED178FA2FFCCA" box="[1318,1424,799,825]" captionStart="FIGURE 14" captionStartId="23.[151,250,1809,1833]" captionTargetBox="[151,1432,190,1783]" captionTargetId="figure-20@23.[151,1436,181,1785]" captionTargetPageId="23" captionText="FIGURE 14. Rhyacopsyche escamosa sp. nov. 14A, head, dorsal. 14B, antenna. 14C, forewing, detail of costal overfold and scales. 14D, right forewing, dorsal. 14E, right hind wing, dorsal. Abbreviations: o.s.w = ocellar setal wart (paired); p.s.w = posterior setal wart (paired)." figureDoi="http://doi.org/10.5281/zenodo.8430646" httpUri="https://zenodo.org/record/8430646/files/figure.png" pageId="22" pageNumber="323">Fig. 14A</figureCitation>
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). Antennae each 40-articulated, scape about 3x as long as wide, median flagellomeres each about 1.5x as long as wide and with slender tapered setae scattered over entire flagellomere, apical flagellomere conical (
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<figureCitation id="EBD92A2AC34CFFB0FB5CD100FAA5FC72" box="[1204,1306,871,897]" captionStart="FIGURE 14" captionStartId="23.[151,250,1809,1833]" captionTargetBox="[151,1432,190,1783]" captionTargetId="figure-20@23.[151,1436,181,1785]" captionTargetPageId="23" captionText="FIGURE 14. Rhyacopsyche escamosa sp. nov. 14A, head, dorsal. 14B, antenna. 14C, forewing, detail of costal overfold and scales. 14D, right forewing, dorsal. 14E, right hind wing, dorsal. Abbreviations: o.s.w = ocellar setal wart (paired); p.s.w = posterior setal wart (paired)." figureDoi="http://doi.org/10.5281/zenodo.8430646" httpUri="https://zenodo.org/record/8430646/files/figure.png" pageId="22" pageNumber="323">Fig. 14B</figureCitation>
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).
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</paragraph>
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<paragraph id="735D36AFC34CFFB0FF2FD1ECFDD2FBE2" blockId="22.[151,1437,762,1977]" pageId="22" pageNumber="323">
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<emphasis id="4196EABDC34CFFB0FF2FD1ECFE9FFC56" box="[199,288,907,933]" italics="true" pageId="22" pageNumber="323">Thorax.</emphasis>
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<materialsCitation id="C38A3CF2C34CFFB0FECFD1ECFDD6FBE2" collectionCode="II, III" pageId="22" pageNumber="323" specimenCount="2" typeStatus="holotype">
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Metascutellum subpentagonal. Length of each forewing:
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<typeStatus id="AC59880DC34CFFB0FC42D1ECFBB3FC56" box="[938,1036,907,933]" pageId="22" pageNumber="323">holotype</typeStatus>
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3.2;
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<typeStatus id="AC59880DC34CFFB0FBAED1ECFB0EFC56" box="[1094,1201,907,933]" pageId="22" pageNumber="323">paratypes</typeStatus>
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3.0–
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<quantity id="B41A9B4AC34CFFB0FB02D1ECFAFFFC56" box="[1258,1344,907,933]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="22" pageNumber="323" unit="mm" value="3.2">3.2 mm</quantity>
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(n = 2). Wings each tapering to acute apex. Forewings elliptical, each with reflexed costal cell covered with dark brown, tear-drop-shape scales; forks I,
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<collectionCode id="15F3AE6AC34CFFB0FE16D1B4FDACFC1E" box="[510,531,979,1005]" pageId="22" pageNumber="323">II</collectionCode>
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, and
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<collectionCode id="15F3AE6AC34CFFB0FDB9D1B4FDD2FC1E" box="[593,621,979,1005]" pageId="22" pageNumber="323">III</collectionCode>
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present; jugal lobe present (
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<figureCitation id="EBD92A2AC34CFFB0FC5BD1B4FBDFFC1E" box="[947,1120,979,1005]" captionStart="FIGURE 14" captionStartId="23.[151,250,1809,1833]" captionTargetBox="[151,1432,190,1783]" captionTargetId="figure-20@23.[151,1436,181,1785]" captionTargetPageId="23" captionText="FIGURE 14. Rhyacopsyche escamosa sp. nov. 14A, head, dorsal. 14B, antenna. 14C, forewing, detail of costal overfold and scales. 14D, right forewing, dorsal. 14E, right hind wing, dorsal. Abbreviations: o.s.w = ocellar setal wart (paired); p.s.w = posterior setal wart (paired)." figureDoi="http://doi.org/10.5281/zenodo.8430646" httpUri="https://zenodo.org/record/8430646/files/figure.png" pageId="22" pageNumber="323">Figs 14C, 14D</figureCitation>
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). Hind wings tapering from basal 1/3, with fork
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<collectionCode id="15F3AE6AC34CFFB0FE90D190FE2BFBE2" box="[376,404,1015,1041]" pageId="22" pageNumber="323">III</collectionCode>
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present (
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<figureCitation id="EBD92A2AC34CFFB0FE13D190FDDEFBE2" box="[507,609,1015,1041]" captionStart="FIGURE 14" captionStartId="23.[151,250,1809,1833]" captionTargetBox="[151,1432,190,1783]" captionTargetId="figure-20@23.[151,1436,181,1785]" captionTargetPageId="23" captionText="FIGURE 14. Rhyacopsyche escamosa sp. nov. 14A, head, dorsal. 14B, antenna. 14C, forewing, detail of costal overfold and scales. 14D, right forewing, dorsal. 14E, right hind wing, dorsal. Abbreviations: o.s.w = ocellar setal wart (paired); p.s.w = posterior setal wart (paired)." figureDoi="http://doi.org/10.5281/zenodo.8430646" httpUri="https://zenodo.org/record/8430646/files/figure.png" pageId="22" pageNumber="323">Fig. 14E</figureCitation>
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)
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="735D36AFC34CFFB0FF2FD67CFEDAFBAA" blockId="22.[151,1437,762,1977]" pageId="22" pageNumber="323">
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<emphasis id="4196EABDC34CFFB0FF2FD67CFD90FBC7" box="[199,559,1051,1077]" italics="true" pageId="22" pageNumber="323">Pregenital abdominal segments</emphasis>
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. Segments V and VI with no modifications. Segment VII with ventromesal process triangular.
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</paragraph>
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<paragraph id="735D36AFC34CFFB0FF2FD604FE65F9DE" blockId="22.[151,1437,762,1977]" pageId="22" pageNumber="323">
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<emphasis id="4196EABDC34CFFB0FF2FD604FE8EFB8E" box="[199,305,1123,1149]" italics="true" pageId="22" pageNumber="323">Genitalia</emphasis>
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. Segment IX partially retracted within segment VIII, open dorsally between pair of dorsolateral lobes; in lateral view, anterior and posterior margins round (
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<figureCitation id="EBD92A2AC34CFFB0FD19D6E0FCE5FB52" box="[753,858,1158,1185]" captionStart="FIGURE 15" captionStartId="24.[151,250,1943,1967]" captionTargetBox="[186,1399,196,1917]" captionTargetId="figure-22@24.[175,1412,181,1919]" captionTargetPageId="24" captionText="FIGURE 15. Rhyacopsyche escamosa sp. nov., male genitalia. 15A, left lateral; 15B, dorsal; 15C, ventral; 15D, phallus, dorsal; 15E, phallus, left lateral. Abbreviations: IX = segment IX; X = tergum X." figureDoi="http://doi.org/10.5281/zenodo.8430650" httpUri="https://zenodo.org/record/8430650/files/figure.png" pageId="22" pageNumber="323">Fig. 15A</figureCitation>
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); in ventral view, anterior margin widely excavated mesally and V-shaped, posterior margin produced into round process (
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<figureCitation id="EBD92A2AC34CFFB0FC44D6CCFBACFB36" box="[940,1043,1194,1221]" captionStart="FIGURE 15" captionStartId="24.[151,250,1943,1967]" captionTargetBox="[186,1399,196,1917]" captionTargetId="figure-22@24.[175,1412,181,1919]" captionTargetPageId="24" captionText="FIGURE 15. Rhyacopsyche escamosa sp. nov., male genitalia. 15A, left lateral; 15B, dorsal; 15C, ventral; 15D, phallus, dorsal; 15E, phallus, left lateral. Abbreviations: IX = segment IX; X = tergum X." figureDoi="http://doi.org/10.5281/zenodo.8430650" httpUri="https://zenodo.org/record/8430650/files/figure.png" pageId="22" pageNumber="323">Fig. 15C</figureCitation>
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). Dorsolateral lobes of segment IX elongate, 2x as long as segment, bearing spiniform setae (
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<figureCitation id="EBD92A2AC34CFFB0FCF8D6A8FCC8FB1A" box="[784,887,1230,1257]" captionStart="FIGURE 15" captionStartId="24.[151,250,1943,1967]" captionTargetBox="[186,1399,196,1917]" captionTargetId="figure-22@24.[175,1412,181,1919]" captionTargetPageId="24" captionText="FIGURE 15. Rhyacopsyche escamosa sp. nov., male genitalia. 15A, left lateral; 15B, dorsal; 15C, ventral; 15D, phallus, dorsal; 15E, phallus, left lateral. Abbreviations: IX = segment IX; X = tergum X." figureDoi="http://doi.org/10.5281/zenodo.8430650" httpUri="https://zenodo.org/record/8430650/files/figure.png" pageId="22" pageNumber="323">Fig. 15A</figureCitation>
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); in dorsal view, deeply divided basally, gradually convergent and almost meeting along median line, apically subtruncate bearing fine setae subapicoventrally (
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<figureCitation id="EBD92A2AC34CFFB0FA87D694FF76FAC2" captionStart="FIGURE 15" captionStartId="24.[151,250,1943,1967]" captionTargetBox="[186,1399,196,1917]" captionTargetId="figure-22@24.[175,1412,181,1919]" captionTargetPageId="24" captionText="FIGURE 15. Rhyacopsyche escamosa sp. nov., male genitalia. 15A, left lateral; 15B, dorsal; 15C, ventral; 15D, phallus, dorsal; 15E, phallus, left lateral. Abbreviations: IX = segment IX; X = tergum X." figureDoi="http://doi.org/10.5281/zenodo.8430650" httpUri="https://zenodo.org/record/8430650/files/figure.png" pageId="22" pageNumber="323">Fig. 15B</figureCitation>
|
||
). Inferior appendages elongate, almost as long as dorsolateral lobes of segment IX and extending beyond them, almost meeting basally, with dorsal spiniform setae apically; in lateral view, apex tapered and upturned, ventral lobe broad basally and produced into acute lobe (
|
||
<figureCitation id="EBD92A2AC34CFFB0FD71D738FCBAFA8A" box="[665,773,1374,1401]" captionStart="FIGURE 15" captionStartId="24.[151,250,1943,1967]" captionTargetBox="[186,1399,196,1917]" captionTargetId="figure-22@24.[175,1412,181,1919]" captionTargetPageId="24" captionText="FIGURE 15. Rhyacopsyche escamosa sp. nov., male genitalia. 15A, left lateral; 15B, dorsal; 15C, ventral; 15D, phallus, dorsal; 15E, phallus, left lateral. Abbreviations: IX = segment IX; X = tergum X." figureDoi="http://doi.org/10.5281/zenodo.8430650" httpUri="https://zenodo.org/record/8430650/files/figure.png" pageId="22" pageNumber="323">Fig. 15A</figureCitation>
|
||
). Preanal appendages absent. Pair of subgenital processes slender, ventrally fused into transverse plate (
|
||
<figureCitation id="EBD92A2AC34CFFB0FD7DD7E4FD43FA6E" box="[661,764,1410,1437]" captionStart="FIGURE 15" captionStartId="24.[151,250,1943,1967]" captionTargetBox="[186,1399,196,1917]" captionTargetId="figure-22@24.[175,1412,181,1919]" captionTargetPageId="24" captionText="FIGURE 15. Rhyacopsyche escamosa sp. nov., male genitalia. 15A, left lateral; 15B, dorsal; 15C, ventral; 15D, phallus, dorsal; 15E, phallus, left lateral. Abbreviations: IX = segment IX; X = tergum X." figureDoi="http://doi.org/10.5281/zenodo.8430650" httpUri="https://zenodo.org/record/8430650/files/figure.png" pageId="22" pageNumber="323">Fig. 15C</figureCitation>
|
||
); in dorsal view, enclosed by segment X, processes meeting apically along median line (
|
||
<figureCitation id="EBD92A2AC34CFFB0FE30D7C0FDFFFA32" box="[472,576,1446,1473]" captionStart="FIGURE 15" captionStartId="24.[151,250,1943,1967]" captionTargetBox="[186,1399,196,1917]" captionTargetId="figure-22@24.[175,1412,181,1919]" captionTargetPageId="24" captionText="FIGURE 15. Rhyacopsyche escamosa sp. nov., male genitalia. 15A, left lateral; 15B, dorsal; 15C, ventral; 15D, phallus, dorsal; 15E, phallus, left lateral. Abbreviations: IX = segment IX; X = tergum X." figureDoi="http://doi.org/10.5281/zenodo.8430650" httpUri="https://zenodo.org/record/8430650/files/figure.png" pageId="22" pageNumber="323">Fig. 15B</figureCitation>
|
||
). Tergum X membranous, apically round (
|
||
<figureCitation id="EBD92A2AC34CFFB0FBC0D7C0FB6AFA32" box="[1064,1237,1446,1473]" captionStart="FIGURE 15" captionStartId="24.[151,250,1943,1967]" captionTargetBox="[186,1399,196,1917]" captionTargetId="figure-22@24.[175,1412,181,1919]" captionTargetPageId="24" captionText="FIGURE 15. Rhyacopsyche escamosa sp. nov., male genitalia. 15A, left lateral; 15B, dorsal; 15C, ventral; 15D, phallus, dorsal; 15E, phallus, left lateral. Abbreviations: IX = segment IX; X = tergum X." figureDoi="http://doi.org/10.5281/zenodo.8430650" httpUri="https://zenodo.org/record/8430650/files/figure.png" pageId="22" pageNumber="323">Figs 15A, 15B</figureCitation>
|
||
). Phallus tubular, elongate, slightly broader basally, thin membranous sheath surrounding phallus subapically, ejaculatory duct short, slightly sclerotized, not protruding apically, apical membrane with two small sclerites, two lanceolate processes subapically (
|
||
<figureCitation id="EBD92A2AC34CFFB0FECED474FE72F9DE" box="[294,461,1554,1581]" captionStart="FIGURE 15" captionStartId="24.[151,250,1943,1967]" captionTargetBox="[186,1399,196,1917]" captionTargetId="figure-22@24.[175,1412,181,1919]" captionTargetPageId="24" captionText="FIGURE 15. Rhyacopsyche escamosa sp. nov., male genitalia. 15A, left lateral; 15B, dorsal; 15C, ventral; 15D, phallus, dorsal; 15E, phallus, left lateral. Abbreviations: IX = segment IX; X = tergum X." figureDoi="http://doi.org/10.5281/zenodo.8430650" httpUri="https://zenodo.org/record/8430650/files/figure.png" pageId="22" pageNumber="323">Figs 15D, 15E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3BF86524C34CFFB0FF2FD450FB63F8BE" pageId="22" pageNumber="323" type="discussion">
|
||
<paragraph id="735D36AFC34CFFB0FF2FD450FB63F8BE" blockId="22.[151,1437,762,1977]" pageId="22" pageNumber="323">
|
||
<emphasis id="4196EABDC34CFFB0FF2FD450FE85F9A2" bold="true" box="[199,314,1591,1617]" pageId="22" pageNumber="323">Remarks.</emphasis>
|
||
<taxonomicName id="B4E24D2CC34CFFB0FED6D450FDF2F9A3" authority="Rocha & Santos & Nessimian, 2023" authorityName="Rocha & Santos & Nessimian" authorityYear="2023" box="[318,589,1591,1617]" class="Insecta" family="Hydroptilidae" genus="Rhyacopsyche" kingdom="Animalia" order="Trichoptera" pageId="22" pageNumber="323" phylum="Arthropoda" rank="species" species="escamosa" status="sp. nov.">
|
||
<emphasis id="4196EABDC34CFFB0FED6D450FDF2F9A3" box="[318,589,1591,1617]" italics="true" pageId="22" pageNumber="323">Rhyacopsyche escamosa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="4196EABDC34CFFB0FDB8D450FD19F9A2" bold="true" box="[592,678,1591,1617]" pageId="22" pageNumber="323">
|
||
<taxonomicNameLabel id="5AA557C6C34CFFB0FDB8D450FD19F9A2" box="[592,678,1591,1617]" pageId="22" pageNumber="323" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
shares with
|
||
<taxonomicName id="B4E24D2CC34CFFB0FCC1D450FABCF9A2" authority="Wasmund & Holzenthal 2007" authorityName="Wasmund & Holzenthal" authorityYear="2007" box="[809,1283,1591,1617]" class="Insecta" family="Hydroptilidae" genus="Rhyacopsyche" kingdom="Animalia" order="Trichoptera" pageId="22" pageNumber="323" phylum="Arthropoda" rank="species" species="tanylobosa">
|
||
<emphasis id="4196EABDC34CFFB0FCC1D450FC00F9A2" box="[809,959,1591,1617]" italics="true" pageId="22" pageNumber="323">R. tanylobosa</emphasis>
|
||
<bibRefCitation id="17734B5EC34CFFB0FC2AD450FABCF9A2" author="Wasmund, A. M. & Holzenthal, R. W." box="[962,1283,1591,1617]" pageId="22" pageNumber="323" pagination="1 - 59" refId="ref16166" refString="Wasmund, A. M. & Holzenthal, R. W. (2007) A revision of the caddisfly genus Rhyacopsyche, with the description of 13 new species (Trichoptera: Hydroptilidae). Zootaxa, 1634 (1), 1 - 59. https: // doi. org / 10.11646 / zootaxa. 1634.1.1" type="journal article" year="2007">Wasmund &Holzenthal 2007</bibRefCitation>
|
||
</taxonomicName>
|
||
the extremely long dorsolateral lobes of segment IX and inferior appendages. Additionally, the inferior appendages of both species are broad basally. However, in
|
||
<taxonomicName id="B4E24D2CC34CFFB0FE18D418FD36F96A" authorityName="Wasmund & Holzenthal" authorityYear="2007" box="[496,649,1663,1689]" class="Insecta" family="Hydroptilidae" genus="Rhyacopsyche" kingdom="Animalia" order="Trichoptera" pageId="22" pageNumber="323" phylum="Arthropoda" rank="species" species="tanylobosa">
|
||
<emphasis id="4196EABDC34CFFB0FE18D418FD36F96A" box="[496,649,1663,1689]" italics="true" pageId="22" pageNumber="323">R. tanylobosa</emphasis>
|
||
</taxonomicName>
|
||
the ventral lobe of each inferior appendage is more acute and elongate than in
|
||
<taxonomicName id="B4E24D2CC34CFFB0FF04D4C4FEC8F94F" authority="Rocha & Santos & Nessimian, 2023" authorityName="Rocha & Santos & Nessimian" authorityYear="2023" box="[236,375,1699,1724]" class="Insecta" family="Hydroptilidae" genus="Rhyacopsyche" kingdom="Animalia" order="Trichoptera" pageId="22" pageNumber="323" phylum="Arthropoda" rank="species" species="escamosa" status="sp. nov.">
|
||
<emphasis id="4196EABDC34CFFB0FF04D4C4FEC8F94F" box="[236,375,1699,1724]" italics="true" pageId="22" pageNumber="323">R. escamosa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="4196EABDC34CFFB0FE96D4C4FE67F94E" bold="true" box="[382,472,1699,1725]" pageId="22" pageNumber="323">
|
||
<taxonomicNameLabel id="5AA557C6C34CFFB0FE96D4C4FE67F94E" box="[382,472,1699,1725]" pageId="22" pageNumber="323" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
These species are also distinguished from each other by elements of the phallus: In
|
||
<taxonomicName id="B4E24D2CC34CFFB0FA6CD4C4FEAEF912" authorityName="Wasmund & Holzenthal" authorityYear="2007" class="Insecta" family="Hydroptilidae" genus="Rhyacopsyche" kingdom="Animalia" order="Trichoptera" pageId="22" pageNumber="323" phylum="Arthropoda" rank="species" species="tanylobosa">
|
||
<emphasis id="4196EABDC34CFFB0FA6CD4C4FEAEF912" italics="true" pageId="22" pageNumber="323">R. tanylobosa</emphasis>
|
||
</taxonomicName>
|
||
, a long paramere is present, while in
|
||
<taxonomicName id="B4E24D2CC34CFFB0FD42D4A0FC8CF913" authority="Rocha & Santos & Nessimian, 2023" authorityName="Rocha & Santos & Nessimian" authorityYear="2023" box="[682,819,1735,1760]" class="Insecta" family="Hydroptilidae" genus="Rhyacopsyche" kingdom="Animalia" order="Trichoptera" pageId="22" pageNumber="323" phylum="Arthropoda" rank="species" species="escamosa" status="sp. nov.">
|
||
<emphasis id="4196EABDC34CFFB0FD42D4A0FC8CF913" box="[682,819,1735,1760]" italics="true" pageId="22" pageNumber="323">R. escamosa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="4196EABDC34CFFB0FCD1D4A0FC2EF912" bold="true" box="[825,913,1735,1761]" pageId="22" pageNumber="323">
|
||
<taxonomicNameLabel id="5AA557C6C34CFFB0FCD1D4A0FC2EF912" box="[825,913,1735,1761]" pageId="22" pageNumber="323" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, a pair of small sclerites is situated in the apical membrane and two lanceolate processes are present subapically. The forewings each have a reflexed costal cell which is shared with
|
||
<taxonomicName id="B4E24D2CC34CFFB0FEDFD568FD75F8DA" authority="Olah & Johanson 2011" authorityName="Olah & Johanson" authorityYear="2011" box="[311,714,1807,1833]" class="Insecta" family="Hydroptilidae" genus="Rhyacopsyche" kingdom="Animalia" order="Trichoptera" pageId="22" pageNumber="323" phylum="Arthropoda" rank="species" species="bunkotala">
|
||
<emphasis id="4196EABDC34CFFB0FEDFD568FE7BF8DA" box="[311,452,1807,1833]" italics="true" pageId="22" pageNumber="323">R. bunkotala</emphasis>
|
||
<bibRefCitation id="17734B5EC34CFFB0FE23D568FD75F8DA" author="Olah, J. & Johanson, K. A." box="[459,714,1807,1833]" pageId="22" pageNumber="323" pagination="1 - 143" refId="ref15371" refString="Olah, J. & Johanson, K. A. (2011) New Neotropical Hydroptilidae (Trichoptera). Annales Historico-Naturales Musei Nationalis Hungarici, 103, 1 - 143." type="journal article" year="2011">Oláh & Johanson 2011</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="B4E24D2CC34CFFB0FD3DD568FBF2F8DA" authority="Olah & Johanson 2011" authorityName="Olah & Johanson" authorityYear="2011" box="[725,1101,1807,1833]" class="Insecta" family="Hydroptilidae" genus="Rhyacopsyche" kingdom="Animalia" order="Trichoptera" pageId="22" pageNumber="323" phylum="Arthropoda" rank="species" species="hajtoka">
|
||
<emphasis id="4196EABDC34CFFB0FD3DD568FCF8F8DA" box="[725,839,1807,1833]" italics="true" pageId="22" pageNumber="323">R. hajtoka</emphasis>
|
||
<bibRefCitation id="17734B5EC34CFFB0FCA5D568FBF2F8DA" author="Olah, J. & Johanson, K. A." box="[845,1101,1807,1833]" pageId="22" pageNumber="323" pagination="1 - 143" refId="ref15371" refString="Olah, J. & Johanson, K. A. (2011) New Neotropical Hydroptilidae (Trichoptera). Annales Historico-Naturales Musei Nationalis Hungarici, 103, 1 - 143." type="journal article" year="2011">Oláh & Johanson 2011</bibRefCitation>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="B4E24D2CC34CFFB0FB6ED568FEC7F8BE" authority="(Wasmund & Holzenthal 2007)" baseAuthorityName="Wasmund & Holzenthal" baseAuthorityYear="2007" class="Insecta" family="Hydroptilidae" genus="Rhyacopsyche" kingdom="Animalia" order="Trichoptera" pageId="22" pageNumber="323" phylum="Arthropoda" rank="species" species="tanylobosa">
|
||
<emphasis id="4196EABDC34CFFB0FB6ED568FAA0F8DA" box="[1158,1311,1807,1833]" italics="true" pageId="22" pageNumber="323">R. tanylobosa</emphasis>
|
||
(
|
||
<bibRefCitation id="17734B5EC34CFFB0FAC4D568FED0F8BE" author="Wasmund, A. M. & Holzenthal, R. W." pageId="22" pageNumber="323" pagination="1 - 59" refId="ref16166" refString="Wasmund, A. M. & Holzenthal, R. W. (2007) A revision of the caddisfly genus Rhyacopsyche, with the description of 13 new species (Trichoptera: Hydroptilidae). Zootaxa, 1634 (1), 1 - 59. https: // doi. org / 10.11646 / zootaxa. 1634.1.1" type="journal article" year="2007">Wasmund & Holzenthal 2007</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, but in none of these species were the presence of scales in this cell described.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3BF86524C34CFFB0FF2FD531FE7DF84A" pageId="22" pageNumber="323" type="materials_examined">
|
||
<paragraph id="735D36AFC34CFFB0FF2FD531FE7DF84A" blockId="22.[151,1437,762,1977]" pageId="22" pageNumber="323">
|
||
<emphasis id="4196EABDC34CFFB0FF2FD531FE0CF883" bold="true" box="[199,435,1878,1904]" pageId="22" pageNumber="323">Material examined.</emphasis>
|
||
<materialsCitation id="C38A3CF2C34CFFB0FE54D530FBC6F866" collectingDate="2012-08-20" collectionCode="DM, MUSM" collectorName="Malaise & RR Cavichioli & JA Rafael & Santos & Takiya" country="Peru" elevation="633" latitude="-13.2175" location="Quincemil" longLatPrecision="21" longitude="-70.727776" pageId="22" pageNumber="323" specimenCount="1" stateProvince="Cusco" typeStatus="holotype">
|
||
<typeStatus id="AC59880DC34CFFB0FE54D530FD97F882" box="[444,552,1879,1905]" pageId="22" pageNumber="323">Holotype</typeStatus>
|
||
.
|
||
<collectingCountry id="0BF5763FC34CFFB0FDDDD530FDD2F882" box="[565,621,1879,1905]" name="Peru" pageId="22" pageNumber="323">Peru</collectingCountry>
|
||
:
|
||
<collectingRegion id="B126F84DC34CFFB0FD91D530FD7CF882" box="[633,707,1879,1905]" country="Peru" name="Cusco" pageId="22" pageNumber="323">Cusco</collectingRegion>
|
||
:
|
||
<location id="763D6074C34CFFB0FD38D530FCF6F882" LSID="urn:lsid:plazi:treatment:FB4B87B9C34CFFBFFF7FD0E9FD88FF26:763D6074C34CFFB0FD38D530FCF6F882" box="[720,841,1879,1905]" country="Peru" latitude="-13.2175" longLatPrecision="21" longitude="-70.727776" name="Quincemil" pageId="22" pageNumber="323" stateProvince="Cusco">Quincemil</location>
|
||
,
|
||
<quantity id="B41A9B4AC34CFFB0FCBFD530FC2DF882" box="[855,914,1879,1905]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="22" pageNumber="323" unit="km" value="3.0">3 km</quantity>
|
||
E,
|
||
<geoCoordinate id="16D65068C34CFFB0FC55D530FB80F882" box="[957,1087,1879,1905]" degrees="13" direction="south" minutes="13" orientation="latitude" pageId="22" pageNumber="323" precision="15" seconds="03" value="-13.2175">13º13’03”S</geoCoordinate>
|
||
<geoCoordinate id="16D65068C34CFFB0FBA0D530FB69F882" box="[1096,1238,1879,1905]" degrees="70" direction="west" minutes="43" orientation="longitude" pageId="22" pageNumber="323" precision="15" seconds="40" value="-70.727776">70º43’40”W</geoCoordinate>
|
||
,
|
||
<quantity id="B41A9B4AC34CFFB0FB0BD530FA91F882" box="[1251,1326,1879,1905]" metricMagnitude="2" metricUnit="m" metricValue="6.33" pageId="22" pageNumber="323" unit="m" value="633.0">
|
||
<elevation id="F8CFD19CC34CFFB0FB0BD530FA91F882" box="[1251,1326,1879,1905]" metricMagnitude="2" metricUnit="m" metricValue="6.33" pageId="22" pageNumber="323" unit="m" value="633.0">633 m</elevation>
|
||
</quantity>
|
||
,
|
||
<collectorName id="DE175379C34CFFB0FAD5D530FA27F882" box="[1341,1432,1879,1905]" pageId="22" pageNumber="323">Malaise</collectorName>
|
||
,
|
||
<date id="075C106FC34CFFB0FF7FD51CFE9EF866" box="[151,289,1915,1941]" pageId="22" pageNumber="323" value="2012-08-20">
|
||
<collectingDate id="1718E987C34CFFB0FF7FD51CFE9EF866" box="[151,289,1915,1941]" pageId="22" pageNumber="323" value="2012-08-20">20.viii.2012</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="DE175379C34CFFB0FEC7D51CFE68F866" box="[303,471,1915,1941]" pageId="22" pageNumber="323">RR Cavichioli</collectorName>
|
||
,
|
||
<collectorName id="DE175379C34CFFB0FE0CD51CFDE7F866" box="[484,600,1915,1941]" pageId="22" pageNumber="323">JA Rafael</collectorName>
|
||
, APM
|
||
<collectorName id="DE175379C34CFFB0FD43D51CFD47F866" box="[683,760,1915,1941]" pageId="22" pageNumber="323">Santos</collectorName>
|
||
,
|
||
<collectionCode id="15F3AE6AC34CFFB0FCEFD51CFC8BF866" box="[775,820,1915,1941]" country="New Zealand" name="Dominion Museum" pageId="22" pageNumber="323">DM</collectionCode>
|
||
<collectorName id="DE175379C34CFFB0FCD5D51CFC36F866" box="[829,905,1915,1941]" pageId="22" pageNumber="323">Takiya</collectorName>
|
||
leg., male (
|
||
<collectionCode id="15F3AE6AC34CFFB0FBFDD51CFBCEF866" box="[1045,1137,1915,1941]" pageId="22" pageNumber="323">MUSM</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="C38A3CF2C34CFFB0FB6ED51CFE01F84A" collectingDate="2012-08-20" collectionCode="DZRJ" collectorName="Malaise & RR Cavichioli & JA Rafael & Santos & Takiya" country="Peru" elevation="633" latitude="-13.2175" location="Quincemil" longLatPrecision="21" longitude="-70.727776" pageId="22" pageNumber="323" specimenCount="2" specimenCount-male="2" stateProvince="Cusco" typeStatus="paratype">
|
||
<typeStatus id="AC59880DC34CFFB0FB6ED51CFB48F866" box="[1158,1271,1915,1941]" pageId="22" pageNumber="323">Paratypes</typeStatus>
|
||
. Same data as holotype,
|
||
<specimenCount id="65E4FD26C34CFFB0FEEFD5F8FEE2F84A" box="[263,349,1951,1977]" pageId="22" pageNumber="323" type="male">2 males</specimenCount>
|
||
(
|
||
<collectionCode id="15F3AE6AC34CFFB0FE84D5F8FE0AF84A" box="[364,437,1951,1977]" pageId="22" pageNumber="323">DZRJ</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="279D6627C34DFFB1FF7FD576FE08F882" ID-DOI="http://doi.org/10.5281/zenodo.8430646" ID-Zenodo-Dep="8430646" httpUri="https://zenodo.org/record/8430646/files/figure.png" pageId="23" pageNumber="324" startId="23.[151,250,1809,1833]" targetBox="[151,1432,190,1783]" targetPageId="23" targetType="figure">
|
||
<paragraph id="735D36AFC34DFFB1FF7FD576FE08F882" blockId="23.[151,1437,1809,1905]" pageId="23" pageNumber="324">
|
||
<emphasis id="4196EABDC34DFFB1FF7FD576FE9EF8DA" bold="true" box="[151,289,1809,1833]" pageId="23" pageNumber="324">FIGURE 14.</emphasis>
|
||
<taxonomicName id="B4E24D2CC34DFFB1FEC0D576FD9DF8DA" authority="Rocha & Santos & Nessimian, 2023" authorityName="Rocha & Santos & Nessimian" authorityYear="2023" box="[296,546,1809,1833]" class="Insecta" family="Hydroptilidae" genus="Rhyacopsyche" kingdom="Animalia" order="Trichoptera" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="escamosa" status="sp. nov.">
|
||
<emphasis id="4196EABDC34DFFB1FEC0D576FD9DF8DA" box="[296,546,1809,1833]" italics="true" pageId="23" pageNumber="324">Rhyacopsyche escamosa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="4196EABDC34DFFB1FDC2D576FDC3F8DA" bold="true" box="[554,636,1809,1833]" pageId="23" pageNumber="324">
|
||
<taxonomicNameLabel id="5AA557C6C34DFFB1FDC2D576FDC3F8DA" box="[554,636,1809,1833]" pageId="23" pageNumber="324" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
14A, head, dorsal. 14B, antenna. 14C, forewing, detail of costal overfold and scales. 14D, right forewing, dorsal. 14E, right hind wing, dorsal. Abbreviations: o.s.w = ocellar setal wart (paired); p.s.w = posterior setal wart (paired).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="279D6627C342FFBEFF7FD5F0FCC0F820" ID-DOI="http://doi.org/10.5281/zenodo.8430650" ID-Zenodo-Dep="8430650" httpUri="https://zenodo.org/record/8430650/files/figure.png" pageId="24" pageNumber="325" startId="24.[151,250,1943,1967]" targetBox="[186,1399,196,1917]" targetPageId="24" targetType="figure">
|
||
<paragraph id="735D36AFC342FFBEFF7FD5F0FCC0F820" blockId="24.[151,1436,1943,2004]" pageId="24" pageNumber="325">
|
||
<emphasis id="4196EABDC342FFBEFF7FD5F0FEA0F85C" bold="true" box="[151,287,1943,1967]" pageId="24" pageNumber="325">FIGURE 15.</emphasis>
|
||
<taxonomicName id="B4E24D2CC342FFBEFECCD5F0FDA3F85C" authority="Rocha & Santos & Nessimian, 2023" authorityName="Rocha & Santos & Nessimian" authorityYear="2023" box="[292,540,1943,1967]" class="Insecta" family="Hydroptilidae" genus="Rhyacopsyche" kingdom="Animalia" order="Trichoptera" pageId="24" pageNumber="325" phylum="Arthropoda" rank="species" species="escamosa" status="sp. nov.">
|
||
<emphasis id="4196EABDC342FFBEFECCD5F0FDA3F85C" box="[292,540,1943,1967]" italics="true" pageId="24" pageNumber="325">Rhyacopsyche escamosa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="4196EABDC342FFBEFDCAD5F0FDCDF85C" bold="true" box="[546,626,1943,1967]" pageId="24" pageNumber="325">
|
||
<taxonomicNameLabel id="5AA557C6C342FFBEFDCAD5F0FDCDF85C" box="[546,626,1943,1967]" pageId="24" pageNumber="325" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male genitalia. 15A, left lateral; 15B, dorsal; 15C, ventral; 15D, phallus, dorsal; 15E, phallus, left lateral. Abbreviations: IX = segment IX; X = tergum X.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="3BF86524C343FFBFFF2FD2F0FD88FF26" pageId="25" pageNumber="326" type="etymology">
|
||
<paragraph id="735D36AFC343FFBFFF2FD2F0FD88FF26" blockId="25.[151,1436,151,213]" pageId="25" pageNumber="326">
|
||
<emphasis id="4196EABDC343FFBFFF2FD2F0FEF1FF42" bold="true" box="[199,334,151,177]" pageId="25" pageNumber="326">Etymology.</emphasis>
|
||
<emphasis id="4196EABDC343FFBFFEBFD2F0FE76FF43" box="[343,457,151,176]" italics="true" pageId="25" pageNumber="326">Escamosa</emphasis>
|
||
is the Spanish adjective for
|
||
<emphasis id="4196EABDC343FFBFFCFBD2F0FCF2FF42" box="[787,845,151,177]" italics="true" pageId="25" pageNumber="326">scaly</emphasis>
|
||
, in reference to the dark brown scales covering the reflexed costal cell of each forewing.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |