237 lines
36 KiB
XML
237 lines
36 KiB
XML
<document id="71937F6ED6C1795FAC9A78D0C1FFCBC4" ID-CLB-Dataset="27801" ID-DOI="10.11646/zootaxa.4571.4.7" ID-GBIF-Dataset="7080afbb-67f5-4742-be94-337881dc6935" ID-ISSN="1175-5326" ID-Zenodo-Dep="2614266" ID-ZooBank="F2BA103A-766F-4D1D-80E2-0303CF53AFF7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553782158970" checkinUser="plazi" docAuthor="Chaibi, Marwa, Antit, Mouna, Bouhedi, Marwa, Meca, Miguel A., Gillet, Patrick, Azzouna, Atf & Martin, Daniel" docDate="2019" docId="03C2621BFF874534FF7E268E755CFF2D" docLanguage="en" docName="zootaxa.4571.4.7.pdf" docOrigin="Zootaxa 4571 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Trophoniella radesiensis Chaibi & Antit & Bouhedi & Meca & Gillet & Azzouna & Martin 2019, n. sp." docType="treatment" docVersion="8" lastPageNumber="556" masterDocId="FFFB1A63FF844531FFE923497132FFD5" masterDocTitle="A new species of Flabelligeridae (Annelida), Trophoniella radesiensis n. sp., from Tunisia" masterLastPageNumber="561" masterPageNumber="551" pageNumber="554" updateTime="1698714785451" updateUser="ExternalLinkService">
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<mods:title id="39E1A7B2A96D60C861AEF50F39B4821C">A new species of Flabelligeridae (Annelida), Trophoniella radesiensis n. sp., from Tunisia</mods:title>
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<mods:namePart id="EBD5E030638E0C207A9B913D544AF58C">Chaibi, Marwa</mods:namePart>
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<mods:namePart id="8AD32BBDCD779D0543F2A2B0D4699DDF">Antit, Mouna</mods:namePart>
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<mods:roleTerm id="87F6CE4A952FCF18BEACCFEF33DB2631">Author</mods:roleTerm>
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<mods:namePart id="3E2CA803D2E2764F7CB9E96BEC8516F8">Bouhedi, Marwa</mods:namePart>
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<mods:roleTerm id="E4668F7B7C83519E4483AF057A70BDEC">Author</mods:roleTerm>
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<mods:namePart id="6825C9CFA2246E05BC8BACE2F07D4E81">Meca, Miguel A.</mods:namePart>
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<mods:roleTerm id="34FA0F90FD51AD4036A7E0B6B17ED2E7">Author</mods:roleTerm>
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<mods:namePart id="6CA0AF6BCBE56B8EBAE0927376F7933D">Gillet, Patrick</mods:namePart>
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<mods:namePart id="14CC4C07C7DF06E20359DCF22FB9B555">Azzouna, Atf</mods:namePart>
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<mods:namePart id="0FC9EC185E8FA8BC970B9524EF00B7B4">Martin, Daniel</mods:namePart>
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<mods:title id="105692B877F9ABCFFAC51B436DDF924D">Zootaxa</mods:title>
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<mods:part id="0A9409D5EA068D6B08A9ADBD198442D7">
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<mods:date id="731170E70F4A220490D61757FD498867">2019</mods:date>
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<mods:number id="0776B00CCF5BFEF4325DA1833F958578">2019-03-28</mods:number>
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<mods:identifier id="F866AA848076E22FEFE0CAD8A6A3987D" type="DOI">10.11646/zootaxa.4571.4.7</mods:identifier>
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<mods:identifier id="85C5068696FF6E32DCFAE75D77759FB4" type="GBIF-Dataset">7080afbb-67f5-4742-be94-337881dc6935</mods:identifier>
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<mods:identifier id="19F8493BF520DFC7379075C9CEE3BDB6" type="Zenodo-Dep">2614266</mods:identifier>
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<treatment id="03C2621BFF874534FF7E268E755CFF2D" ID-DOI="http://doi.org/10.5281/zenodo.5935765" ID-GBIF-Taxon="154520680" ID-Zenodo-Dep="5935765" LSID="urn:lsid:plazi:treatment:03C2621BFF874534FF7E268E755CFF2D" httpUri="http://treatment.plazi.org/id/03C2621BFF874534FF7E268E755CFF2D" lastPageId="5" lastPageNumber="556" pageId="3" pageNumber="554">
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<subSubSection id="C3718086FF874532FF7E268E73D8FA34" box="[151,746,1479,1505]" pageId="3" pageNumber="554" type="nomenclature">
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<paragraph id="8BD4D30DFF874532FF7E268E73D8FA34" blockId="3.[151,965,1479,1574]" box="[151,746,1479,1505]" pageId="3" pageNumber="554">
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<heading id="D09C6461FF874532FF7E268E73D8FA34" bold="true" box="[151,746,1479,1505]" fontSize="11" level="1" pageId="3" pageNumber="554" reason="1">
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<emphasis id="B91F0F1FFF874532FF7E268E73D8FA34" bold="true" box="[151,746,1479,1505]" pageId="3" pageNumber="554">
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<taxonomicName id="4C6BA88EFF874532FF7E268E70F2FA34" ID-CoL="30ae9de1-e872-4bb2-ae62-3b654c560048" authority="Chaibi & Antit & Bouhedi & Meca & Gillet & Azzouna & Martin, 2019" authorityName="Chaibi & Antit & Bouhedi & Meca & Gillet & Azzouna & Martin" authorityYear="2019" box="[151,448,1479,1505]" class="Polychaeta" family="Flabelligeridae" genus="Trophoniella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="554" phylum="Annelida" rank="species" species="radesiensis" status="n. sp .">
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<emphasis id="B91F0F1FFF874532FF7E268E70F2FA34" bold="true" box="[151,448,1479,1505]" italics="true" pageId="3" pageNumber="554">Trophoniella radesiensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22CB264FF874532FE21268E7323FA34" box="[456,529,1479,1505]" pageId="3" pageNumber="554">n. sp.</taxonomicNameLabel>
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Chaibi and Gillet
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3718086FF874532FF7E26A372F7F9F3" pageId="3" pageNumber="554" type="description">
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<paragraph id="8BD4D30DFF874532FF7E26A3702EF9D6" blockId="3.[151,965,1479,1574]" box="[151,284,1514,1539]" pageId="3" pageNumber="554">
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<figureCitation id="1350CF88FF874532FF7E26A3702EF9D6" box="[151,284,1514,1539]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStart-2="FIGURE 5" captionStart-3="FIGURE 6" captionStartId-0="4.[151,250,1939,1961]" captionStartId-1="5.[151,250,1207,1229]" captionStartId-2="6.[151,250,1448,1470]" captionStartId-3="7.[151,250,1574,1596]" captionTargetBox-0="[340,1246,975,1898]" captionTargetBox-1="[343,1246,297,1196]" captionTargetBox-2="[343,1247,194,1426]" captionTargetBox-3="[341,1247,517,1550]" captionTargetId-0="figure@4.[340,1247,974,1898]" captionTargetId-1="figure@5.[340,1247,296,1196]" captionTargetId-2="figure@6.[340,1247,193,1426]" captionTargetId-3="figure@7.[340,1247,517,1553]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionText-0="FIGURE 3. Trophoniella radesiensis n. sp. A. Lateral view. B. Mid body region showing the attached sand grains, lateral view. C. Anterior region, lateral view. D. Anterior region, dorsal view. E. Anterior region, ventral view. A, C–D: Holotype MNCN 16.01/18453; B: Paratype MNCN 16.01/18454." captionText-1="FIGURE 4. Trophoniella radesiensis n. sp., stained with methyl blue. A. Detail of the anterior end, ventral view, prostomium non-everted, showing the tips of the branchial plates, the position of the first four chaetigers (1 to 4) and the crenulated cephalic veil (white arrow). B. Palps of a dissected specimen, dorsal view, surrounded by branchiae. C. Whole ventral view of the anterior region of a dissected specimen. D. Detail of C indicate the relevant characters. BP: branchial plates; CA: caruncle; E: eyes; LL: lateral lips; PS: palp scars; Pr: prostomium. A: Holotype MNCN16.01/18453; B–E: Paratype MNCN16.01/18454." captionText-2="FIGURE 5. Trophoniella radesiensis n. sp. A: Notopodium of chaetiger 16 (with two broken notochaetae). B–D. Notochaetae. Tip. C. Median region. D. Basal region. A–D: Paratype MNCN16.01/18454." captionText-3="FIGURE 6. Trophoniella radesiensis n. sp. A. Neuropodium of chaetiger 15. A1, A2, A5. Detail of tips from three short neurochaetal hooks of chaetiger 15. B. Neuropodium of chaetiger 30. B1, B2, B3. Detail of tips from three anchylosed neurochaetal hooks of chaetiger 30. A–B: Paratype MNCN16.01/18454." httpUri-0="https://zenodo.org/record/2614270/files/figure.png" httpUri-1="https://zenodo.org/record/2614272/files/figure.png" httpUri-2="https://zenodo.org/record/2614274/files/figure.png" httpUri-3="https://zenodo.org/record/2614276/files/figure.png" pageId="3" pageNumber="554">Figures 3–6</figureCitation>
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</paragraph>
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<paragraph id="8BD4D30DFF874532FF7E254472F7F9F3" blockId="3.[151,965,1479,1574]" box="[151,965,1549,1574]" pageId="3" pageNumber="554">
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urn:lsid:zoobank.org:act:
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<uuid id="FFCDE9D8FF874532FE47254472F7F9F3" box="[430,965,1549,1574]" pageId="3" pageNumber="554">D7C85C70-87FF-4AF4-8599-A8462B3FB8B1</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3718086FF874532FF7E251C7238F963" pageId="3" pageNumber="554" type="materials_examined">
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<paragraph id="8BD4D30DFF874532FF7E251C7238F963" blockId="3.[151,1437,1621,2006]" pageId="3" pageNumber="554">
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<emphasis id="B91F0F1FFF874532FF7E251C70B6F9BB" bold="true" box="[151,388,1621,1646]" pageId="3" pageNumber="554">Material examined.</emphasis>
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<materialsCitation id="3B03D950FF874532FE66251C736FF946" ID-GBIF-Occurrence="2107540862" collectingDate="2015-11-25" collectionCode="MNCN" country="Tunisia" latitude="15.916667" location="Rades Station" longLatPrecision="1284" longitude="-97.683334" municipality="Gulf" pageId="3" pageNumber="554" specimenCode="MNCN16.01" specimenCount="1" typeStatus="holotype">
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<typeStatus id="54D06DAFFF874532FE66251C70CEF9BB" box="[399,508,1621,1646]" pageId="3" pageNumber="554" type="holotype">Holotype</typeStatus>
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:
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<specimenCode id="DBCD7B76FF874532FDE0251C73ADF9BB" box="[521,671,1621,1646]" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="3" pageNumber="554" type="Museum">MNCN16.01</specimenCode>
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/18453, collected on
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<date id="FFD5F5CDFF874532FC7A251C7510F9BB" box="[915,1058,1621,1646]" pageId="3" pageNumber="554" value="2015-11-25">
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<collectingDate id="EF910C25FF874532FC7A251C7510F9BB" box="[915,1058,1621,1646]" pageId="3" pageNumber="554" value="2015-11-25">
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25 Nov
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<quantity id="4C937EE8FF874532FC03251C7510F9BB" box="[1002,1058,1621,1646]" metricMagnitude="1" metricUnit="m" metricValue="5.1181" pageId="3" pageNumber="554" unit="in" value="2015.0">2015</quantity>
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</collectingDate>
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</date>
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in
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<location id="8EB485D6FF874532FBA4251C75DCF9BB" LSID="urn:lsid:plazi:treatment:03C2621BFF874534FF7E268E755CFF2D:8EB485D6FF874532FBA4251C75DCF9BB" box="[1101,1262,1621,1646]" country="Tunisia" latitude="15.916667" longLatPrecision="1284" longitude="-97.683334" municipality="Gulf" name="Rades Station" pageId="3" pageNumber="554">Radés Station</location>
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,
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<collectingMunicipality id="6BB04977FF874532FB14251C7403F9BB" box="[1277,1329,1621,1646]" pageId="3" pageNumber="554">Gulf</collectingMunicipality>
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of
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<collectingCountry id="F37C939DFF874532FAB4251C74AEF9BB" box="[1373,1436,1621,1646]" name="Tunisia" pageId="3" pageNumber="554">Tunis</collectingCountry>
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(
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<geoCoordinate id="EE5FB5CAFF874532FF762533703AF947" box="[159,264,1658,1683]" direction="north" orientation="latitude" pageId="3" pageNumber="554" precision="925" value="15.916667">15°55′ N</geoCoordinate>
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,
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<geoCoordinate id="EE5FB5CAFF874532FEFD253370B3F947" box="[276,385,1658,1683]" direction="west" orientation="longitude" pageId="3" pageNumber="554" precision="925" value="-97.683334">97°41′ W</geoCoordinate>
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) by the first author
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</materialsCitation>
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.
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<materialsCitation id="3B03D950FF874532FD8E253274AAF946" ID-GBIF-Occurrence="2107545720" box="[615,1432,1658,1683]" collectingDate="2015-11-25" collectionCode="MNCN" country="Tunisia" latitude="15.916667" location="Rades Station" longLatPrecision="1284" longitude="-97.683334" municipality="Gulf" pageId="3" pageNumber="554" specimenCode="MNCN 16.01" specimenCount="6" typeStatus="paratype">
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<typeStatus id="54D06DAFFF874532FD8E253273EBF946" box="[615,729,1659,1683]" pageId="3" pageNumber="554" type="paratype">Paratypes</typeStatus>
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:
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<specimenCode id="DBCD7B76FF874532FD0D2533724CF946" box="[740,894,1658,1683]" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="3" pageNumber="554" type="Museum">MNCN 16.01</specimenCode>
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/18454.
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<specimenCount id="9D6D1884FF874532FC322533755AF946" box="[987,1128,1658,1683]" pageId="3" pageNumber="554" type="generic">6 specimens</specimenCount>
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, same data as for holotype
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</materialsCitation>
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;
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<materialsCitation id="3B03D950FF874532FF7E25D77234F963" ID-GBIF-Occurrence="2107544658" box="[151,774,1693,1718]" collectingDate="2015-11-25" collectionCode="MNCN" country="Tunisia" latitude="15.916667" location="Rades Station" longLatPrecision="1284" longitude="-97.683334" municipality="Gulf" pageId="3" pageNumber="554" specimenCount="1" typeStatus="paratype">UCO T FLA 025.1specimen, same data as for holotype</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3718086FF874532FF2E258B7295F893" pageId="3" pageNumber="554" type="diagnosis">
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<paragraph id="8BD4D30DFF874532FF2E258B7295F893" blockId="3.[151,1437,1621,2006]" pageId="3" pageNumber="554">
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<emphasis id="B91F0F1FFF874532FF2E258B7073F90E" bold="true" box="[199,321,1730,1755]" pageId="3" pageNumber="554">Diagnosis.</emphasis>
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Body covered by sediments of different grain size (50–1000 µm of long axe) embedded on tunic, completely concealed except in posterior region. Tunic pale, dorsally smooth, ventrally rough, with large papillae, carrying sediment grains and particles. Body papillae arranged in two dorsal and four ventral rows. Branchial plate tongue-shaped. Unidentate anchylosed neurohooks from chaetiger 20.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3718086FF874535FF2E241B7463FC55" lastPageId="4" lastPageNumber="555" pageId="3" pageNumber="554" type="description">
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<paragraph id="8BD4D30DFF874535FF2E241B7463FC55" blockId="3.[151,1437,1621,2006]" lastBlockId="4.[151,1437,151,896]" lastPageId="4" lastPageNumber="555" pageId="3" pageNumber="554">
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<emphasis id="B91F0F1FFF874532FF2E241B706BF8BE" bold="true" box="[199,345,1874,1899]" pageId="3" pageNumber="554">Description.</emphasis>
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<typeStatus id="54D06DAFFF874532FE88241B70FBF8BE" box="[353,457,1874,1899]" pageId="3" pageNumber="554" type="holotype">Holotype</typeStatus>
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with some chaetae broken, non-reproductive adult, unknown sex.
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<quantity id="4C937EE8FF874532FB29241B7423F8BF" box="[1216,1297,1874,1899]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="3" pageNumber="554" unit="mm" value="35.0">35 mm</quantity>
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long,
|
||
<quantity id="4C937EE8FF874532FAB3241B74AFF8BF" box="[1370,1437,1874,1898]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="3" pageNumber="554" unit="mm" value="4.0">4 mm</quantity>
|
||
wide, with 60 chaetigers;
|
||
<typeStatus id="54D06DAFFF874532FE5C243E7313F85A" box="[437,545,1911,1935]" pageId="3" pageNumber="554" type="paratype">paratypes</typeStatus>
|
||
varying from
|
||
<quantity id="4C937EE8FF874532FD29243F720AF85B" box="[704,824,1910,1934]" metricMagnitude="-2" metricUnit="m" metricValue="3.65" metricValueMax="4.3" metricValueMin="3.0" pageId="3" pageNumber="554" unit="mm" value="36.5" valueMax="43.0" valueMin="30.0">30–43 mm</quantity>
|
||
long and
|
||
<quantity id="4C937EE8FF874532FC4C243F7530F85B" box="[933,1026,1910,1934]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="4.0" metricValueMin="1.0" pageId="3" pageNumber="554" unit="mm" value="2.5" valueMax="4.0" valueMin="1.0">1–4 mm</quantity>
|
||
wide for 43–74 chaetigers (
|
||
<figureCitation id="1350CF88FF874532FAD1243C744AF85B" box="[1336,1400,1909,1934]" captionStart="FIGURE 3" captionStartId="4.[151,250,1939,1961]" captionTargetBox="[340,1246,975,1898]" captionTargetId="figure@4.[340,1247,974,1898]" captionTargetPageId="4" captionText="FIGURE 3. Trophoniella radesiensis n. sp. A. Lateral view. B. Mid body region showing the attached sand grains, lateral view. C. Anterior region, lateral view. D. Anterior region, dorsal view. E. Anterior region, ventral view. A, C–D: Holotype MNCN 16.01/18453; B: Paratype MNCN 16.01/18454." httpUri="https://zenodo.org/record/2614270/files/figure.png" pageId="3" pageNumber="554">Fig. 3</figureCitation>
|
||
A– E). Anterior body sub-cylindrical in cross-section, tapering towards pygidium. Tunic transparent, dorsally smooth, ventrally rough with large papillae, carrying sediment grains and particles with long axes ranging from 50 to 1000 µm, totally embedded in tunic, completely concealing it dorsally and ventrally, absent in posterior region. Pale brown, slightly reddish anteriorly. Cephalic cage
|
||
<quantity id="4C937EE8FF804535FD2F23F57224FF01" box="[710,790,188,212]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="4" pageNumber="555" unit="mm" value="12.0">12 mm</quantity>
|
||
long, with chaetae ca. 1.5 times longer than body width, formed by chaetigers 1–5; chaetiger 2 dorsolateral, chaetiger 3–4 lateral. Chaetal transition from cephalic cage to body chaetae gradual (
|
||
<figureCitation id="1350CF88FF804535FE7C224D7335FEC9" box="[405,519,260,285]" captionStart="FIGURE 3" captionStartId="4.[151,250,1939,1961]" captionTargetBox="[340,1246,975,1898]" captionTargetId="figure@4.[340,1247,974,1898]" captionTargetPageId="4" captionText="FIGURE 3. Trophoniella radesiensis n. sp. A. Lateral view. B. Mid body region showing the attached sand grains, lateral view. C. Anterior region, lateral view. D. Anterior region, dorsal view. E. Anterior region, ventral view. A, C–D: Holotype MNCN 16.01/18453; B: Paratype MNCN 16.01/18454." httpUri="https://zenodo.org/record/2614270/files/figure.png" pageId="4" pageNumber="555">Fig. 3A, 3</figureCitation>
|
||
C–E). Body papillae similar in colour to body wall, mostly eroded, arising in two dorsal and four ventral longitudinal rows from first chaetiger to posterior end, better preserved anterior-most body region (
|
||
<figureCitation id="1350CF88FF804535FF1922057055FEB1" box="[240,359,332,357]" captionStart="FIGURE 3" captionStartId="4.[151,250,1939,1961]" captionTargetBox="[340,1246,975,1898]" captionTargetId="figure@4.[340,1247,974,1898]" captionTargetPageId="4" captionText="FIGURE 3. Trophoniella radesiensis n. sp. A. Lateral view. B. Mid body region showing the attached sand grains, lateral view. C. Anterior region, lateral view. D. Anterior region, dorsal view. E. Anterior region, ventral view. A, C–D: Holotype MNCN 16.01/18453; B: Paratype MNCN 16.01/18454." httpUri="https://zenodo.org/record/2614270/files/figure.png" pageId="4" pageNumber="555">Fig. 3A, 3</figureCitation>
|
||
C–E). Parapodia well developed. Noto and neuropodia have four prechaetal papillae and five postchaetal papillae (
|
||
<figureCitation id="1350CF88FF804535FE60222670D1FE5D" box="[393,483,367,392]" captionStart="FIGURE 3" captionStartId="4.[151,250,1939,1961]" captionTargetBox="[340,1246,975,1898]" captionTargetId="figure@4.[340,1247,974,1898]" captionTargetPageId="4" captionText="FIGURE 3. Trophoniella radesiensis n. sp. A. Lateral view. B. Mid body region showing the attached sand grains, lateral view. C. Anterior region, lateral view. D. Anterior region, dorsal view. E. Anterior region, ventral view. A, C–D: Holotype MNCN 16.01/18453; B: Paratype MNCN 16.01/18454." httpUri="https://zenodo.org/record/2614270/files/figure.png" pageId="4" pageNumber="555">Fig. 3C</figureCitation>
|
||
). Especially long papillae absent from anterior chaetigers. Gonopodial papillae not seen. Chaetiger 1 with about six notochaetae and eight neurochaetae; anterior dorsal margin with dorsal papillae, arising as a multifid cephalic veil (
|
||
<figureCitation id="1350CF88FF804535FDA822FE7390FE05" box="[577,674,439,464]" captionStart="FIGURE 4" captionStartId="5.[151,250,1207,1229]" captionTargetBox="[343,1246,297,1196]" captionTargetId="figure@5.[340,1247,296,1196]" captionTargetPageId="5" captionText="FIGURE 4. Trophoniella radesiensis n. sp., stained with methyl blue. A. Detail of the anterior end, ventral view, prostomium non-everted, showing the tips of the branchial plates, the position of the first four chaetigers (1 to 4) and the crenulated cephalic veil (white arrow). B. Palps of a dissected specimen, dorsal view, surrounded by branchiae. C. Whole ventral view of the anterior region of a dissected specimen. D. Detail of C indicate the relevant characters. BP: branchial plates; CA: caruncle; E: eyes; LL: lateral lips; PS: palp scars; Pr: prostomium. A: Holotype MNCN16.01/18453; B–E: Paratype MNCN16.01/18454." httpUri="https://zenodo.org/record/2614272/files/figure.png" pageId="4" pageNumber="555">Fig. 4A</figureCitation>
|
||
). Chaetiger 5 widening posteriorly. Cephalic hood not exposed. Caruncle short and triangular (
|
||
<figureCitation id="1350CF88FF804535FDE8229573A6FE21" box="[513,660,476,501]" captionStart="FIGURE 4" captionStartId="5.[151,250,1207,1229]" captionTargetBox="[343,1246,297,1196]" captionTargetId="figure@5.[340,1247,296,1196]" captionTargetPageId="5" captionText="FIGURE 4. Trophoniella radesiensis n. sp., stained with methyl blue. A. Detail of the anterior end, ventral view, prostomium non-everted, showing the tips of the branchial plates, the position of the first four chaetigers (1 to 4) and the crenulated cephalic veil (white arrow). B. Palps of a dissected specimen, dorsal view, surrounded by branchiae. C. Whole ventral view of the anterior region of a dissected specimen. D. Detail of C indicate the relevant characters. BP: branchial plates; CA: caruncle; E: eyes; LL: lateral lips; PS: palp scars; Pr: prostomium. A: Holotype MNCN16.01/18453; B–E: Paratype MNCN16.01/18454." httpUri="https://zenodo.org/record/2614272/files/figure.png" pageId="4" pageNumber="555">Fig. 4C, 4D</figureCitation>
|
||
). Branchiae cirriform, arising from tongue-shaped branchial plate, arranged in two lateral lobes (
|
||
<figureCitation id="1350CF88FF804535FE0522B6734BFDCD" box="[492,633,511,536]" captionStart="FIGURE 4" captionStartId="5.[151,250,1207,1229]" captionTargetBox="[343,1246,297,1196]" captionTargetId="figure@5.[340,1247,296,1196]" captionTargetPageId="5" captionText="FIGURE 4. Trophoniella radesiensis n. sp., stained with methyl blue. A. Detail of the anterior end, ventral view, prostomium non-everted, showing the tips of the branchial plates, the position of the first four chaetigers (1 to 4) and the crenulated cephalic veil (white arrow). B. Palps of a dissected specimen, dorsal view, surrounded by branchiae. C. Whole ventral view of the anterior region of a dissected specimen. D. Detail of C indicate the relevant characters. BP: branchial plates; CA: caruncle; E: eyes; LL: lateral lips; PS: palp scars; Pr: prostomium. A: Holotype MNCN16.01/18453; B–E: Paratype MNCN16.01/18454." httpUri="https://zenodo.org/record/2614272/files/figure.png" pageId="4" pageNumber="555">Fig. 4A, 4D</figureCitation>
|
||
), thin, long (
|
||
<quantity id="4C937EE8FF804535FCE522B6724EFDCD" box="[780,892,511,536]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="3.0" metricValueMin="0.5" pageId="4" pageNumber="555" unit="mm" value="1.75" valueMax="3.0" valueMin="0.5">0.5–3mm</quantity>
|
||
), whitish once preserved in ethanol, with ca. 60 filaments. Palp, long, corrugated, pale, as long as largest branchiae,
|
||
<quantity id="4C937EE8FF804535FC4F216D72DBFDE9" box="[934,1001,548,573]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="4" pageNumber="555" unit="mm" value="6.0">6 mm</quantity>
|
||
long (
|
||
<figureCitation id="1350CF88FF804535FBDC216D75F3FDE9" box="[1077,1217,548,573]" captionStart="FIGURE 4" captionStartId="5.[151,250,1207,1229]" captionTargetBox="[343,1246,297,1196]" captionTargetId="figure@5.[340,1247,296,1196]" captionTargetPageId="5" captionText="FIGURE 4. Trophoniella radesiensis n. sp., stained with methyl blue. A. Detail of the anterior end, ventral view, prostomium non-everted, showing the tips of the branchial plates, the position of the first four chaetigers (1 to 4) and the crenulated cephalic veil (white arrow). B. Palps of a dissected specimen, dorsal view, surrounded by branchiae. C. Whole ventral view of the anterior region of a dissected specimen. D. Detail of C indicate the relevant characters. BP: branchial plates; CA: caruncle; E: eyes; LL: lateral lips; PS: palp scars; Pr: prostomium. A: Holotype MNCN16.01/18453; B–E: Paratype MNCN16.01/18454." httpUri="https://zenodo.org/record/2614272/files/figure.png" pageId="4" pageNumber="555">Fig. 4B, 4C</figureCitation>
|
||
). Prostomium lowcone, with two large and two small black eyes (
|
||
<figureCitation id="1350CF88FF804535FD47210E7204FDB5" box="[686,822,583,608]" captionStart="FIGURE 3" captionStartId="4.[151,250,1939,1961]" captionTargetBox="[340,1246,975,1898]" captionTargetId="figure@4.[340,1247,974,1898]" captionTargetPageId="4" captionText="FIGURE 3. Trophoniella radesiensis n. sp. A. Lateral view. B. Mid body region showing the attached sand grains, lateral view. C. Anterior region, lateral view. D. Anterior region, dorsal view. E. Anterior region, ventral view. A, C–D: Holotype MNCN 16.01/18453; B: Paratype MNCN 16.01/18454." httpUri="https://zenodo.org/record/2614270/files/figure.png" pageId="4" pageNumber="555">Fig. 3D, 3E</figureCitation>
|
||
). Lateral lip expanded; dorsal and ventral lips not well developed (
|
||
<figureCitation id="1350CF88FF804535FEF321257091FD51" box="[282,419,620,645]" captionStart="FIGURE 3" captionStartId="4.[151,250,1939,1961]" captionTargetBox="[340,1246,975,1898]" captionTargetId="figure@4.[340,1247,974,1898]" captionTargetPageId="4" captionText="FIGURE 3. Trophoniella radesiensis n. sp. A. Lateral view. B. Mid body region showing the attached sand grains, lateral view. C. Anterior region, lateral view. D. Anterior region, dorsal view. E. Anterior region, ventral view. A, C–D: Holotype MNCN 16.01/18453; B: Paratype MNCN 16.01/18454." httpUri="https://zenodo.org/record/2614270/files/figure.png" pageId="4" pageNumber="555">Fig. 3D, 3E</figureCitation>
|
||
). Notochaetae all multiarticulated capillaries; articles progressively longer towards falcate tips; medial ones in short longitudinal series, 4–7 per bundle; some yellowish some dark brown; unidentate tips (
|
||
<figureCitation id="1350CF88FF804535FF7621FD71D3FD18" box="[159,225,692,717]" captionStart="FIGURE 5" captionStartId="6.[151,250,1448,1470]" captionTargetBox="[343,1247,194,1426]" captionTargetId="figure@6.[340,1247,193,1426]" captionTargetPageId="6" captionText="FIGURE 5. Trophoniella radesiensis n. sp. A: Notopodium of chaetiger 16 (with two broken notochaetae). B–D. Notochaetae. Tip. C. Median region. D. Basal region. A–D: Paratype MNCN16.01/18454." httpUri="https://zenodo.org/record/2614274/files/figure.png" pageId="4" pageNumber="555">Fig. 5</figureCitation>
|
||
A–5D). Multiarticulated capillary neurochaeta from chaetiger 2 to 5, then short unidentate neurohooks from chaetiger 6 to 19 (
|
||
<figureCitation id="1350CF88FF804535FE8F219E708DFD25" box="[358,447,727,752]" captionStart="FIGURE 6" captionStartId="7.[151,250,1574,1596]" captionTargetBox="[341,1247,517,1550]" captionTargetId="figure@7.[340,1247,517,1553]" captionTargetPageId="7" captionText="FIGURE 6. Trophoniella radesiensis n. sp. A. Neuropodium of chaetiger 15. A1, A2, A5. Detail of tips from three short neurochaetal hooks of chaetiger 15. B. Neuropodium of chaetiger 30. B1, B2, B3. Detail of tips from three anchylosed neurochaetal hooks of chaetiger 30. A–B: Paratype MNCN16.01/18454." httpUri="https://zenodo.org/record/2614276/files/figure.png" pageId="4" pageNumber="555">Fig. 6A</figureCitation>
|
||
) and anchylosed hooks from chaetiger 20 to body end (
|
||
<figureCitation id="1350CF88FF804535FBD1219E75A2FD25" box="[1080,1168,727,752]" captionStart="FIGURE 6" captionStartId="7.[151,250,1574,1596]" captionTargetBox="[341,1247,517,1550]" captionTargetId="figure@7.[340,1247,517,1553]" captionTargetPageId="7" captionText="FIGURE 6. Trophoniella radesiensis n. sp. A. Neuropodium of chaetiger 15. A1, A2, A5. Detail of tips from three short neurochaetal hooks of chaetiger 15. B. Neuropodium of chaetiger 30. B1, B2, B3. Detail of tips from three anchylosed neurochaetal hooks of chaetiger 30. A–B: Paratype MNCN16.01/18454." httpUri="https://zenodo.org/record/2614276/files/figure.png" pageId="4" pageNumber="555">Fig. 6B</figureCitation>
|
||
), darker than preceding ones, arranged in transverse series of 4–6 units per bundle, similarly wide along their length, subdistally not or slightly expanded, with short rings continued up to a subdistal, non-annulated hyaline region, hooked, tapering to roughly pointed, unidentate tips (
|
||
<figureCitation id="1350CF88FF804535FDCE200D73BAFC88" box="[551,648,836,861]" captionStart="FIGURE 5" captionStartId="6.[151,250,1448,1470]" captionTargetBox="[343,1247,194,1426]" captionTargetId="figure@6.[340,1247,193,1426]" captionTargetPageId="6" captionText="FIGURE 5. Trophoniella radesiensis n. sp. A: Notopodium of chaetiger 16 (with two broken notochaetae). B–D. Notochaetae. Tip. C. Median region. D. Basal region. A–D: Paratype MNCN16.01/18454." httpUri="https://zenodo.org/record/2614274/files/figure.png" pageId="4" pageNumber="555">Fig. 5A</figureCitation>
|
||
,
|
||
<figureCitation id="1350CF88FF804535FD73200D738CFC88" box="[666,702,836,861]" captionStart="FIGURE 6" captionStartId="7.[151,250,1574,1596]" captionTargetBox="[341,1247,517,1550]" captionTargetId="figure@7.[340,1247,517,1553]" captionTargetPageId="7" captionText="FIGURE 6. Trophoniella radesiensis n. sp. A. Neuropodium of chaetiger 15. A1, A2, A5. Detail of tips from three short neurochaetal hooks of chaetiger 15. B. Neuropodium of chaetiger 30. B1, B2, B3. Detail of tips from three anchylosed neurochaetal hooks of chaetiger 30. A–B: Paratype MNCN16.01/18454." httpUri="https://zenodo.org/record/2614276/files/figure.png" pageId="4" pageNumber="555">6B</figureCitation>
|
||
). Pygidium simple, with single anal cirrus. No intra-specific variability in morphological characters was observed, except for lacking sediment gains in some specimens.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF148385FF804535FF7E24DA734BF833" httpUri="https://zenodo.org/record/2614270/files/figure.png" pageId="4" pageNumber="555" startId="4.[151,250,1939,1961]" targetBox="[340,1246,975,1898]" targetPageId="4">
|
||
<paragraph id="8BD4D30DFF804535FF7E24DA734BF833" blockId="4.[151,1437,1939,2022]" pageId="4" pageNumber="555">
|
||
<emphasis id="B91F0F1FFF804535FF7E24DA7020F87D" bold="true" box="[151,274,1939,1961]" pageId="4" pageNumber="555">FIGURE 3.</emphasis>
|
||
<taxonomicName id="4C6BA88EFF804535FEF124DA7323F87C" authority="Chaibi & Antit & Bouhedi & Meca & Gillet & Azzouna & Martin, 2019" authorityName="Chaibi & Antit & Bouhedi & Meca & Gillet & Azzouna & Martin" authorityYear="2019" box="[280,529,1939,1961]" class="Polychaeta" family="Flabelligeridae" genus="Trophoniella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebellida" pageId="4" pageNumber="555" phylum="Annelida" rank="species" species="radesiensis" status="sp. nov.">
|
||
<emphasis id="B91F0F1FFF804535FEF124DA7323F87C" box="[280,529,1939,1961]" italics="true" pageId="4" pageNumber="555">Trophoniella radesiensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B91F0F1FFF804535FDFE24DA7362F87D" bold="true" box="[535,592,1939,1960]" pageId="4" pageNumber="555">
|
||
<taxonomicNameLabel id="A22CB264FF804535FDFE24DA7362F87D" box="[535,592,1939,1960]" pageId="4" pageNumber="555" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
A. Lateral view. B. Mid body region showing the attached sand grains, lateral view. C. Anterior region, lateral view. D. Anterior region, dorsal view. E. Anterior region, ventral view. A, C–D: Holotype MNCN 16.01/18453; B: Paratype MNCN 16.01/18454.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C3718086FF814534FF2E23DE755CFF2D" pageId="5" pageNumber="556" type="distribution">
|
||
<paragraph id="8BD4D30DFF814534FF2E23DE755CFF2D" blockId="5.[151,1436,151,248]" pageId="5" pageNumber="556">
|
||
<emphasis id="B91F0F1FFF814534FF2E23DE7052FF65" bold="true" box="[199,352,151,176]" pageId="5" pageNumber="556">Distribution.</emphasis>
|
||
Known only from the
|
||
<typeStatus id="54D06DAFFF814534FD6D23D17386FF65" box="[644,692,152,176]" pageId="5" pageNumber="556">type</typeStatus>
|
||
locality, Radés Station (Gulf of
|
||
<collectingCountry id="F37C939DFF814534FBA323DE7592FF65" box="[1098,1184,151,176]" name="Tunisia" pageId="5" pageNumber="556">Tunisia</collectingCountry>
|
||
, Mediterranean Sea). Collected from both soft and rocky bottoms, from
|
||
<quantity id="4C937EE8FF814534FD2523F57221FF01" box="[716,787,188,212]" metricMagnitude="0" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="5" pageNumber="556" unit="m" value="3.5" valueMax="4.0" valueMin="3.0">3–4 m</quantity>
|
||
to
|
||
<quantity id="4C937EE8FF814534FCDE23F57242FF01" box="[823,880,188,212]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="556" unit="m" value="10.0">10 m</quantity>
|
||
depth.
|
||
<emphasis id="B91F0F1FFF814534FF2E2396707CFF2D" bold="true" box="[199,334,223,248]" pageId="5" pageNumber="556">Etymology.</emphasis>
|
||
The species name
|
||
<taxonomicName id="4C6BA88EFF814534FDCB23967392FF2D" authorityName="Chaibi & Antit & Bouhedi & Meca & Gillet & Azzouna & Martin" authorityYear="2019" box="[546,672,223,248]" class="Polychaeta" family="Flabelligeridae" genus="Trophoniella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebellida" pageId="5" pageNumber="556" phylum="Annelida" rank="species" species="radesiensis">
|
||
<emphasis id="B91F0F1FFF814534FDCB23967392FF2D" box="[546,672,223,248]" italics="true" pageId="5" pageNumber="556">radesiensis</emphasis>
|
||
</taxonomicName>
|
||
refers to the
|
||
<typeStatus id="54D06DAFFF814534FCDD23A97256FF2D" box="[820,868,224,248]" pageId="5" pageNumber="556">type</typeStatus>
|
||
locality, Radés Station.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |