161 lines
22 KiB
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161 lines
22 KiB
XML
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<mods:title id="E8024EE54A7A0643A464D972CAE506AE">Replacement names for two species of Orthacanthus Agassiz, 1843 (Chondrichthyes, Xenacanthiformes), and discussion of Giebelodus Whitley, 1940, replacement name for Chilodus Giebel, 1848 (Chondrichthyes, Xenacanthiformes), preoccupied by Chilodus Mueller & Troschel, 1844 (Actinopterygii, Characiformes)</mods:title>
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<mods:namePart id="DE5B10CF5226F6239C32FDF9B765121B">Babcock, Loren E.</mods:namePart>
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<mods:affiliation id="F06330B8BC95A4902A22341825729248">School of Earth Sciences, Orton Geological Museum, The Ohio State University, Columbus, OH, 43210, USA</mods:affiliation>
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<mods:nameIdentifier id="C930BD83AA91D1A104AD3CA5B52BE32C" type="email">babcock.5@osu.edu</mods:nameIdentifier>
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<mods:date id="2F1D19978ADDA5CA3E32909E41F2C581">2024</mods:date>
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<mods:identifier id="7E1F7615BE699195F1A9F18D1D27AD8D" type="DOI">http://dx.doi.org/10.3897/zookeys.1188.108571</mods:identifier>
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<subSection id="0BDAC5EF2AABB128FEC79C315BAD3916" pageId="0" pageNumber="219" type="systematics">
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<treatment id="BDFD6863936D5C9A9E11D8AE14C1860C" LSID="urn:lsid:zoobank.org:act:3CAE1776-5270-4FAB-A6EB-C9808C448235" httpUri="http://treatment.plazi.org/id/BDFD6863936D5C9A9E11D8AE14C1860C" lastPageNumber="219" pageId="0" pageNumber="219">
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<subSubSection id="F8CFFB6ABB8081A2AF6C4F99BD80A91F" pageId="0" pageNumber="219" type="nomenclature">
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<paragraph id="913D5166FB41438FADF8D7CEB9750674" pageId="0" pageNumber="219">
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<taxonomicName id="DAAE51898AEDC146D02B2BF90D592596" LSID="BDFD6863-936D-5C9A-9E11-D8AE14C1860C" authorityName="Babcock" authorityYear="2024" class="Mammalia" family="Poaceae" genus="Orthacanthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthacanthus adamas" order="Poales" pageId="0" pageNumber="219" phylum="Tracheophyta" rank="species" species="adamas" status="nom. nov.">Orthacanthus adamas</taxonomicName>
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<taxonomicNameLabel id="5C3560CE0A30FF343E61465E559BC0DB" pageId="0" pageNumber="219">nom. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="8161787D59A227FC2D181144405A3FD4" pageId="0" pageNumber="219" type="description">
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<paragraph id="041814A886CA58072233A3550DABE49F" pageId="0" pageNumber="219">
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<figureCitation id="4339E501F5AF8ECF3936D268DC8F6D4D" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Original 19 th century figures of Carboniferous-age xenacanthiform shark fossils from Saxony-Anhalt, Germany, and Ohio, USA A Orthacanthus gracilis (Giebel, 1848), tooth, holotype (Geiseltalmuseum Halle, GTM 1095), two views; reproduced from Giebel (1849: pl. XXIX, fig. 2 a, b), 7.7 mm long. Wettin-Schichten, Wettin / Saalegebiet, Saxony-Anhalt, Germany B Orthacanthus lintonensis nom. nov., replacement name for Diplodus gracilis Newberry, 1857 a, two teeth, syntypes (repository unknown); reproduced from Newberry (1875 a: pl. LVIII, figs. 3, 3 a), ca 5 and 13 mm long. Upper Freeport Coal, Allegheny Group, Diamond Coal Mine, Linton, Ohio, USA C Orthacanthus adamas nom. nov., replacement name for Orthacanthus gracilis Newberry, 1875 a, dorsal spine, illustration is a composite based on syntypes (Orton Geological Museum, OSU 4467 A, 4467 B); reproduced from Newberry (1875 a, pl. LIX, fig. 7), ca 71 mm long. Upper Freeport Coal, Allegheny Group, Diamond Coal Mine, Linton, Ohio, USA D Orthacanthus carbonarius (Germar, 1844), two teeth, syntypes (repository unknown) of Chilodus tuberosus Giebel, 1848; reproduction of Giebel (1849: pl. XXIX, figs. 1, 1 a, 1 b as), length unknown. Wettin-Loebejun, Saxony-Anhalt, Germany E Orthacanthus carbonarius (Germar, 1844), tooth, syntype (repository unknown) of Lamna carbonaria Germar, 1844; reproduced from Germar (1844: pl. 1, fig. 1), ca 20 mm long. Presumably from Saxony-Anhalt, Germany." figureDoi="10.3897/zookeys.1188.108571.figure1" httpUri="https://binary.pensoft.net/fig/963874" pageId="0" pageNumber="219">Fig. 1C</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="17F58CDFE16B128CD85695A2E7EA2A90" pageId="0" pageNumber="219" type="reference_group">
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<paragraph id="2B0B3B2596CDEF2796E08A9FF9225D3E" pageId="0" pageNumber="219">
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<taxonomicName id="DF77EF587FAF422D4324E3169508014E" authorityName="Babcock" authorityYear="2024" class="Mammalia" family="Poaceae" genus="Orthacanthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthacanthus gracilis" order="Poales" pageId="0" pageNumber="219" phylum="Tracheophyta" rank="species" species="gracilis" status="nom. nov.">Orthacanthus gracilis</taxonomicName>
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Newberry, 1875a: 56-57, pl. LIX, fig. 7.
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</paragraph>
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<paragraph id="924AF1242D3AF94A53CCD413A589922B" pageId="0" pageNumber="219">
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<taxonomicName id="13A13AB30B3F11BFE0E05BA82DBF2343" authorityName="Babcock" authorityYear="2024" class="Mammalia" family="Poaceae" genus="Orthacanthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthacanthus gracilis" order="Poales" pageId="0" pageNumber="219" phylum="Tracheophyta" rank="species" species="gracilis" status="nom. nov.">Orthacanthus gracilis</taxonomicName>
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:
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<bibRefCitation id="8F2DEA0DA36C38114B4313081D2295AE" author="Newberry, JS" journalOrPublisher="Archiv fuer Naturgeschichte" pageId="0" pageNumber="219" refId="B28" refString="Newberry, JS, 1875b. Beschreibung fossiler Fische. Bericht ueber die Geologische Aufnahme von Ohio. II. Band. Geologie und Palaeontologie. II. Theil. Palaeontologie: 1-64." title="Beschreibung fossiler Fische. Bericht ueber die Geologische Aufnahme von Ohio. II. Band. Geologie und Palaeontologie. II. Theil. Palaeontologie: 1 - 64." year="1875 b">Newberry 1875b</bibRefCitation>
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: 56-57, pl. LIX, fig. 7.
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</paragraph>
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<paragraph id="1E46614E4DD3E22E3FAFAD09B2783117" pageId="0" pageNumber="219">
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<taxonomicName id="2747032094A8EEF2AD8D63F40AC03C5F" authorityName="Babcock" authorityYear="2024" class="Mammalia" family="Poaceae" genus="Orthacanthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthacanthus gracilis" order="Poales" pageId="0" pageNumber="219" phylum="Tracheophyta" rank="species" species="gracilis" status="nom. nov.">Orthacanthus gracilis</taxonomicName>
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:
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<bibRefCitation id="2F4953A554D0CAB32BA8884F8073C31B" author="Cope, ED" journalOrPublisher="American Naturalist" pageId="0" pageNumber="219" pagination="162 - 164" refId="B6" refString="Cope, ED, 1881. Catalogue of the Vertebrata of the Permian formation of the United States. American Naturalist 15 (2): 162 - 164" title="Catalogue of the Vertebrata of the Permian formation of the United States." volume="15" year="1881">Cope 1881</bibRefCitation>
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: 163.
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</paragraph>
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<paragraph id="60AC4AF21D14C7E0447F53EC36672D93" pageId="0" pageNumber="219">
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<taxonomicName id="7BBB2243B0D277A9DA7DF40D63173DDF" authorityName="Babcock" authorityYear="2024" class="Insecta" family="Poaceae" genus="Pleuracanthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pleuracanthus (Orthacanthus) gracilis" order="Poales" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="gracilis" status="nom. nov." subGenus="Orthacanthus">Pleuracanthus (Orthacanthus) gracilis</taxonomicName>
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:
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<bibRefCitation id="8086825BDD533BFE2BFDD974F11843E7" DOI="https://doi.org/10.1086/620866" author="Case, EC" journalOrPublisher="The Journal of Geology" pageId="0" pageNumber="219" pagination="698 - 729" refId="B5" refString="Case, EC, 1900. The vertebrates from the Permian bone bed of Vermilion County, Illinois. The Journal of Geology 8 (8): 698 - 729, DOI: https://doi.org/10.1086/620866" title="The vertebrates from the Permian bone bed of Vermilion County, Illinois." url="https://doi.org/10.1086/620866" volume="8" year="1900">Case 1900</bibRefCitation>
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: 701, pl. I, fig. 4.
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</paragraph>
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<paragraph id="02DE6903F1B07187A5900C490A110176" pageId="0" pageNumber="219">
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<taxonomicName id="A2E1AECBBB9153955744CA82DCD73153" authorityName="Babcock" authorityYear="2024" class="Mammalia" family="Poaceae" genus="Orthacanthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthacanthus gracilis" order="Poales" pageId="0" pageNumber="219" phylum="Tracheophyta" rank="species" species="gracilis" status="nom. nov.">Orthacanthus gracilis</taxonomicName>
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:
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<bibRefCitation id="D40DFFDBF7B971C80982C24353CD0DFA" author="Morningstar, H" journalOrPublisher="The Ohio Journal of Science" pageId="0" pageNumber="219" pagination="31 - 64" refId="B22" refString="Morningstar, H, 1924. Catalogue of type fossils in the Geological Museum of The Ohio State University. The Ohio Journal of Science 24: 31 - 64" title="Catalogue of type fossils in the Geological Museum of The Ohio State University." volume="24" year="1924">Morningstar 1924</bibRefCitation>
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: 53.
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</paragraph>
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<paragraph id="8C6CD3B4CCC3C696D18B55E0EB7B05E6" pageId="0" pageNumber="219">
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<taxonomicName id="0BADBBB25EAC3313825DFBA5C0DD20F1" authorityName="Babcock" authorityYear="2024" class="Chondrichthyes" family="Poaceae" genus="Xenacanthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xenacanthus gracilis" order="Poales" pageId="0" pageNumber="219" phylum="Tracheophyta" rank="species" species="gracilis" status="nom. nov.">Xenacanthus gracilis</taxonomicName>
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(Newberry, 1875a):
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<bibRefCitation id="E4CA5EBF4D6306484D08457D6A3A7FFA" DOI="https://doi.org/10.1086/625364" author="Olson, EC" journalOrPublisher="The Journal of Geology" pageId="0" pageNumber="219" pagination="281 - 305" refId="B29" refString="Olson, EC, 1946. Fresh- and brackish-water vertebrate-bearing deposits of the Pennsylvanian of Illinois. The Journal of Geology 54 (5): 281 - 305, DOI: https://doi.org/10.1086/625364" title="Fresh- and brackish-water vertebrate-bearing deposits of the Pennsylvanian of Illinois." url="https://doi.org/10.1086/625364" volume="54" year="1946">Olson 1946</bibRefCitation>
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: 287.
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</paragraph>
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<paragraph id="7AD1F885DA5945E7BC4C2503408C35D7" pageId="0" pageNumber="219">
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<taxonomicName id="6FAF1CB5FCD705F4CA7F77EB561E748A" authorityName="Babcock" authorityYear="2024" class="Chondrichthyes" family="Poaceae" genus="Xenacanthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xenacanthus gracilis" order="Poales" pageId="0" pageNumber="219" phylum="Tracheophyta" rank="species" species="gracilis" status="nom. nov.">Xenacanthus gracilis</taxonomicName>
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:
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<bibRefCitation id="78AE6477537A2BD06E1010F2E3DD7C1C" DOI="https://doi.org/10.1080/02724634.1986.10011609" author="Hook, RW" journalOrPublisher="Journal of Vertebrate Paleontology" pageId="0" pageNumber="219" pagination="174 - 190" refId="B16" refString="Hook, RW, Baird, D, 1986. The Diamond Coal Mine of Linton, Ohio, and its Pennsylvanian-age vertebrates. Journal of Vertebrate Paleontology 6 (2): 174 - 190, DOI: https://doi.org/10.1080/02724634.1986.10011609" title="The Diamond Coal Mine of Linton, Ohio, and its Pennsylvanian-age vertebrates." url="https://doi.org/10.1080/02724634.1986.10011609" volume="6" year="1986">Hook and Baird 1986</bibRefCitation>
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: 179, table 2.
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</paragraph>
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<paragraph id="31E4C2D1D828954CC11DA5D5C1B2751B" pageId="0" pageNumber="219">
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<taxonomicName id="28CCA15362C0FF65009E3C73726A4BD8" authorityName="Babcock" authorityYear="2024" class="Chondrichthyes" family="Poaceae" genus="Xenacanthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xenacanthus gracilis" order="Poales" pageId="0" pageNumber="219" phylum="Tracheophyta" rank="species" species="gracilis" status="nom. nov.">Xenacanthus gracilis</taxonomicName>
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: Hook and Baird 1988: table 1.
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</paragraph>
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<paragraph id="1BCE27C44A9AB20B4DDF95E7EAAB26B1" pageId="0" pageNumber="219">
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<taxonomicName id="9CC27B50A2DD0F90BBC6E38A5983C3E8" authorityName="Babcock" authorityYear="2024" class="Mammalia" family="Poaceae" genus="Orthacanthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthacanthus gracilis" order="Poales" pageId="0" pageNumber="219" phylum="Tracheophyta" rank="species" species="gracilis" status="nom. nov.">Orthacanthus gracilis</taxonomicName>
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: Hampe 2004: 209.
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</paragraph>
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</subSubSection>
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<subSubSection id="AF577E1BAAF25AA946D67618523541A7" pageId="0" pageNumber="219" type="syntypes">
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<paragraph id="1D0648FDE243FEE20ADF0E9D3878B316" pageId="0" pageNumber="219">Syntypes.</paragraph>
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<paragraph id="25FCDE9E22940DD22371EE0FE9577EC0" pageId="0" pageNumber="219">
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Two dorsal spines, Orton Geological Museum, The Ohio State University, Columbus, Ohio, USA (OSU) 4467A, 4467B, previously illustrated as a composite by
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<bibRefCitation id="7FE328A2EB45FF34F4DB2514B9B505D3" author="Newberry, JS" journalOrPublisher="Archiv fuer Naturgeschichte" pageId="0" pageNumber="219" refId="B27" refString="Newberry, JS, 1875a. Descriptions of fossil fishes. Report of the Geological Survey of Ohio. Volume II. Geology and Palaeontology. Part II. Palaeontology, 1-64." title="Descriptions of fossil fishes. Report of the Geological Survey of Ohio. Volume II. Geology and Palaeontology. Part II. Palaeontology, 1 - 64." year="1875 a">Newberry (1875a</bibRefCitation>
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: pl. LIX, fig. 7; 1875b: pl. LIX, fig. 7).
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</paragraph>
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</subSubSection>
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||
<subSubSection id="1637E29A8F156E9F212A5EB50C908FDD" pageId="0" pageNumber="219" type="type locality">
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<paragraph id="2F992928DFF9BAC6E862DB9A6C625DCB" pageId="0" pageNumber="219">Type locality.</paragraph>
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<paragraph id="0DC10DBC81366FF4C101A20E0885B93F" pageId="0" pageNumber="219">Upper Freeport Coal (Carboniferous), from the Diamond Coal Mine, Linton, Jefferson County, Ohio, USA.</paragraph>
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||
</subSubSection>
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||
<subSubSection id="23D74C82D663A5D7FE8FFFF124614110" pageId="0" pageNumber="219" type="etymology">
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<paragraph id="F5339B5D061994147677291F4A4142FD" pageId="0" pageNumber="219">Etymology.</paragraph>
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<paragraph id="2CC44A04CF61E4473581D1FBA01729D7" pageId="0" pageNumber="219">
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<taxonomicName id="9B8E1B8C3BE9C76F9D74489C1722C2DC" family="Nothobranchiidae" genus="Adamas" kingdom="Animalia" lsidName="Adamas" order="Cyprinodontiformes" pageId="0" pageNumber="219" phylum="Chordata" rank="genus">
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<emphasis id="83A314A20142B35D828C2329908CE6B0" italics="true" pageId="0" pageNumber="219">Adamas</emphasis>
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</taxonomicName>
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(Latin, diamond), in allusion to the Diamond Coal Mine, where the species was first collected.
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</paragraph>
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</subSubSection>
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<subSubSection id="619566819390BDF842C49BB92A9740FA" pageId="0" pageNumber="219" type="remarks">
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<paragraph id="5BC41AAB16ABA546382A4AE812C71BAB" pageId="0" pageNumber="219">Remarks.</paragraph>
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<paragraph id="973D5DA0D5511A9DD5CE80409FC88EF3" pageId="0" pageNumber="219">
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The new species-group name
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<taxonomicName id="9B65F8A60B9B56E351B5C34D8E834071" authorityName="Babcock" authorityYear="2024" class="Mammalia" family="Poaceae" genus="Orthacanthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthacanthus adamas" order="Poales" pageId="0" pageNumber="219" phylum="Tracheophyta" rank="species" species="adamas">
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<emphasis id="0858FE603A813A75BE17598FDD45A664" italics="true" pageId="0" pageNumber="219">Orthacanthus adamas</emphasis>
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</taxonomicName>
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nom. nov. replaces
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<taxonomicName id="9C56A0ECEFEB726D8F75F6A39A89757C" authorityName="Babcock" authorityYear="2024" class="Mammalia" family="Poaceae" genus="Orthacanthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthacanthus gracilis" order="Poales" pageId="0" pageNumber="219" phylum="Tracheophyta" rank="species" species="gracilis">
|
||
<emphasis id="589C616AACAA920A286C4A3C01C1BF9A" italics="true" pageId="0" pageNumber="219">Orthacanthus gracilis</emphasis>
|
||
</taxonomicName>
|
||
Newberry, 1875a, which is a junior homonym of
|
||
<taxonomicName id="F8DB4298C090AD4308694F1DAADB26EB" authorityName="Babcock" authorityYear="2024" class="Mammalia" family="Poaceae" genus="Orthacanthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthacanthus gracilis" order="Poales" pageId="0" pageNumber="219" phylum="Tracheophyta" rank="species" species="gracilis">
|
||
<emphasis id="FEB39B7E2665BEC6F783A275614AB72B" italics="true" pageId="0" pageNumber="219">Orthacanthus gracilis</emphasis>
|
||
</taxonomicName>
|
||
(Giebel, 1848).
|
||
<bibRefCitation id="628152E76ECD3427F70A64DDE6B2BB18" author="Newberry, JS" journalOrPublisher="Archiv fuer Naturgeschichte" pageId="0" pageNumber="219" refId="B27" refString="Newberry, JS, 1875a. Descriptions of fossil fishes. Report of the Geological Survey of Ohio. Volume II. Geology and Palaeontology. Part II. Palaeontology, 1-64." title="Descriptions of fossil fishes. Report of the Geological Survey of Ohio. Volume II. Geology and Palaeontology. Part II. Palaeontology, 1 - 64." year="1875 a">Newberry (1875a</bibRefCitation>
|
||
: pl. LIX, fig. 7; 1875b: pl. LIX, fig. 7) illustrated this species with a composite figure based on syntypic dorsal spines. This species should not be confused with the other xenacanthiform species from Linton bearing the species epithet
|
||
<taxonomicName id="685F1493D6E0D5BF8E38466B71C9FB5F" authorityName="Babcock" authorityYear="2024" class="Mammalia" family="Poaceae" genus="Orthacanthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthacanthus gracilis" order="Poales" pageId="0" pageNumber="219" phylum="Tracheophyta" rank="species" species="gracilis">
|
||
<emphasis id="DE33551544915F4514D61E838B9F6BA6" italics="true" pageId="0" pageNumber="219">Orthacanthus gracilis</emphasis>
|
||
</taxonomicName>
|
||
, based on teeth, and also referred to
|
||
<taxonomicName id="0145FA57444C182C40F102FFF0045A7E" class="Mammalia" family="Poaceae" genus="Orthacanthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthacanthus" order="Poales" pageId="0" pageNumber="219" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="41F1EA4B1A3FBA6EBE85C1057C620E39" italics="true" pageId="0" pageNumber="219">Orthacanthus</emphasis>
|
||
</taxonomicName>
|
||
, as discussed above. Replacement names for both taxa will reduce potential confusion.
|
||
<bibRefCitation id="EF0975B25E3927CC835D28A0A8C6C9F5" author="Cope, ED" journalOrPublisher="American Naturalist" pageId="0" pageNumber="219" pagination="162 - 164" refId="B6" refString="Cope, ED, 1881. Catalogue of the Vertebrata of the Permian formation of the United States. American Naturalist 15 (2): 162 - 164" title="Catalogue of the Vertebrata of the Permian formation of the United States." volume="15" year="1881">Cope (1881)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="FA7AE9FFD88AA6BE9F65F430D488FD5D" DOI="https://doi.org/10.1086/620866" author="Case, EC" journalOrPublisher="The Journal of Geology" pageId="0" pageNumber="219" pagination="698 - 729" refId="B5" refString="Case, EC, 1900. The vertebrates from the Permian bone bed of Vermilion County, Illinois. The Journal of Geology 8 (8): 698 - 729, DOI: https://doi.org/10.1086/620866" title="The vertebrates from the Permian bone bed of Vermilion County, Illinois." url="https://doi.org/10.1086/620866" volume="8" year="1900">Case (1900)</bibRefCitation>
|
||
extended the stratigraphic range of this species into the Permian.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</subSection>
|
||
</document> |