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<mods:title id="22AFA5B7CE29BD1ABD7C32632073D6A8">Contribution to the genus Anomala Samouelle, 1819 (Coleoptera: Scarabaeidae Rutelinae) of China and adjacent regions. Part I: descriptions of two new species and remarks on four species</mods:title>
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<taxonomicName id="85F54D3AF02E5F0A919FFF6294FCFF72" authority="Arrow, 1917" authorityName=", Arrow" authorityYear="1917" box="[151,488,223,250]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="587" phylum="Arthropoda" rank="species" species="bella">
<emphasis id="7081EAABF02E5F0A919FFF6294FCFF72" bold="true" box="[151,488,223,250]" pageId="9" pageNumber="587">
<emphasis id="7081EAABF02E5F0A919FFF62945CFF71" bold="true" box="[151,328,223,250]" italics="true" pageId="9" pageNumber="587">Anomala bella</emphasis>
<bibRefCitation id="26644B48F02E5F0A9046FF6294FCFF72" author="Arrow, G. J." box="[334,488,223,249]" pageId="9" pageNumber="587" refId="ref5916" refString="Arrow, G. J. (1917) The fauna of British India, including Ceylon and Burma. Coleoptera Lamellicornia part II (Rutelinae, Des - monycinae, and Euchirinae). Taylor &amp; Francis, London, 387 pp." type="book" year="1917">Arrow, 1917</bibRefCitation>
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(
<figureCitation id="DACE2A3CF02E5F0A9197FEBE9432FE95" box="[159,294,259,286]" captionStart="FIGURES 4953" captionStartId="9.[151,264,1696,1720]" captionTargetBox="[172,1416,916,1666]" captionTargetId="figure-220@9.[164,1424,910,1672]" captionTargetPageId="9" captionText="FIGURES 4953. Habitus and male genitalia of Anomala bella Arrow, 1917 from Yunnan. 49, habitus of male; 50, habitus of female; 51, male genitalia in right lateral view; 52, male genitalia in left lateral view; 53, male genitalia in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4431475" httpUri="https://zenodo.org/record/4431475/files/figure.png" pageId="9" pageNumber="587">Figs. 4953</figureCitation>
)
</paragraph>
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<subSubSection id="0AEF6532F02E5F0A919FFEF595E0FE29" pageId="9" pageNumber="587" type="reference_group">
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<treatmentCitation id="C35410A8F02E5F0A919FFEF594F3FEEB" author="Arrow, G. J." box="[151,487,328,352]" page="208" pageId="9" pageNumber="587" year="1917">
<taxonomicName id="85F54D3AF02E5F0A919FFEF594F3FEEB" authority="Arrow, 1917: 208" authorityName=", Arrow" authorityPageNumber="208" authorityYear="1917" box="[151,487,328,352]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="587" phylum="Arthropoda" rank="species" species="bella">
<emphasis id="7081EAABF02E5F0A919FFEF59439FEEB" box="[151,301,328,352]" italics="true" pageId="9" pageNumber="587">Anomala bella</emphasis>
<bibRefCitation id="26644B48F02E5F0A903AFEF594F3FEEB" author="Arrow, G. J." box="[306,487,328,352]" pageId="9" pageNumber="587" refId="ref5916" refString="Arrow, G. J. (1917) The fauna of British India, including Ceylon and Burma. Coleoptera Lamellicornia part II (Rutelinae, Des - monycinae, and Euchirinae). Taylor &amp; Francis, London, 387 pp." type="book" year="1917">Arrow, 1917: 208</bibRefCitation>
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[
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designated but
<typeStatus id="9D4E881BF02E5F0A93CCFEF597E4FEEB" box="[708,752,328,352]" pageId="9" pageNumber="587">type</typeStatus>
locality unspecified], Plate I, fig. 9;
<treatmentCitation id="C35410A8F02E5F0A9561FEF59074FEEB" author="Machatschke, J. W." box="[1129,1376,328,352]" page="124" pageId="9" pageNumber="587" year="1972">
<bibRefCitation id="26644B48F02E5F0A9561FEF59074FEEB" author="Machatschke, J. W." box="[1129,1376,328,352]" pageId="9" pageNumber="587" refId="ref6393" refString="Machatschke, J. W. (1972) Coleopterorum catalogus supplementa. Pars 66. Fascicle 1. Superfamilie Scarabaeoidea, Familie Melolonthidae, Subfamilie Rutelinae. Editio Secunda. Dr. W. Junk, Gravenhage, 361 pp." type="book" year="1972">Machatschke 1972: 124</bibRefCitation>
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;
<treatmentCitation id="C35410A8F02E5F0A9463FEF59424FEF5" author="Zorn, C." page="257" pageId="9" pageNumber="587" year="2006">
<bibRefCitation id="26644B48F02E5F0A9463FEF59424FEF5" author="Zorn, C." pageId="9" pageNumber="587" pagination="276 - 277" refId="ref7080" refString="Zorn, C. (2006) Subfamily Rutelinae, tribe Anomalini C. E. Blanchard, 1851. In: Lobl, I. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera, Volume 3, Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. Apollo Books, Stenstrup, pp. 276 - 277." type="book chapter" year="2006">Zorn 2006: 257</bibRefCitation>
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;
<treatmentCitation id="C35410A8F02E5F0A9032FEDB971FFEF5" author="Prokofiev, A. M." box="[314,523,358,382]" page="39" pageId="9" pageNumber="587" year="2015">
<bibRefCitation id="26644B48F02E5F0A9032FEDB971FFEF5" author="Prokofiev, A. M." box="[314,523,358,382]" pageId="9" pageNumber="587" pagination="37 - 59" refId="ref6681" refString="Prokofiev, A. M. (2015 a) New Anomala Samouelle, 1819 from South-East Asia. Russian Entomological Journal, 24, 37 - 59. https: // doi. org / 10.15298 / rusentj. 24.1.04" type="journal article" year="2015">Prokofiev 2015a: 39</bibRefCitation>
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, figs. 1013 [recorded from North
<collectingCountry id="3AE27629F02E5F0A9266FEDB96DFFEF5" box="[878,971,358,382]" name="Thailand" pageId="9" pageNumber="587">Thailand</collectingCountry>
, without detailed data];
<bibRefCitation id="26644B48F02E5F0A95B7FEDA9089FEF5" author="Zorn, C. &amp; Bezdek, A." box="[1215,1437,358,382]" pageId="9" pageNumber="587" refId="ref7190" refString="Zorn, C. &amp; Bezdek, A. (2016) Subfamily Rutelinae. In: Lobl, I. &amp; Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 3. Scarabaeoidea, Scirtoidea, Dascilloidea, Buprestoidea, Byrrhoidea. Revised and updated edition. Brill, Leiden, pp. 317 - 358." type="book" year="2016">Zorn &amp; Bezděk 2016:</bibRefCitation>
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</paragraph>
<paragraph id="424A36B9F02E5F0A91CFFE3795E0FE29" blockId="9.[151,1437,328,418]" box="[199,244,394,418]" pageId="9" pageNumber="587">330.</paragraph>
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<paragraph id="424A36B9F02E5F0A919FFE6C97F1FDB8" blockId="9.[151,1437,464,887]" pageId="9" pageNumber="587">
<emphasis id="7081EAABF02E5F0A919FFE6C94A9FE61" bold="true" box="[151,445,464,491]" pageId="9" pageNumber="587">Type material examined.</emphasis>
<materialsCitation id="F29D3CE4F02E5F0A90CCFE6D97F5FDB8" ID-GBIF-Occurrence="3014871322" collectionCode="BMNH, T" country="Germany" county="It" location="Carsten Zorn" municipality="Gnoien" pageId="9" pageNumber="587" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
The type material was not examined by the author. Labels of the male
<typeStatus id="9D4E881BF02E5F0A95C5FE6D903BFE61" box="[1229,1327,464,490]" pageId="9" pageNumber="587" type="holotype">holotype</typeStatus>
(
<collectionCode id="24E4AE7CF02E5F0A9435FE6C9080FE60" box="[1341,1428,465,491]" pageId="9" pageNumber="587">BMNH</collectionCode>
) were provided by
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(
<collectingMunicipality id="A22EACC3F02E5F0A930DFE499748FD85" box="[517,604,500,526]" pageId="9" pageNumber="587">Gnoien</collectingMunicipality>
,
<collectingCountry id="3AE27629F02E5F0A936FFE4897C5FD84" box="[615,721,501,527]" name="Germany" pageId="9" pageNumber="587">Germany</collectingCountry>
) and transcribed here.
<collectingCounty id="AB2B4E35F02E5F0A92D9FE4896F6FD84" box="[977,994,501,527]" pageId="9" pageNumber="587">It</collectingCounty>
is labeled “
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// Type H.
<collectionCode id="24E4AE7CF02E5F0A95F3FE489019FD84" box="[1275,1293,501,527]" name="Tavera, Department of Geology and Geophysics" pageId="9" pageNumber="587">T</collectionCode>
. Mungphu // Atkinson Coll. 92-3. //
<taxonomicName id="85F54D3AF02E5F0A9091FDA59787FDB8" authority=", Arrow" authorityName=", Arrow" authorityYear="1917" box="[409,659,536,563]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="587" phylum="Arthropoda" rank="species" species="bella">Anomala bella, Arrow</taxonomicName>
Type”
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.
</paragraph>
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<emphasis id="7081EAABF02E5F0A91CFFD819721FDDD" bold="true" box="[199,565,572,598]" pageId="9" pageNumber="587">Additional material examined.</emphasis>
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<collectingCountry id="3AE27629F02E5F0A9334FD81978AFDDD" box="[572,670,572,598]" name="China" pageId="9" pageNumber="587">CHINA</collectingCountry>
:
</emphasis>
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,
<specimenCount id="54F3FD30F02E5F0A93F8FD809607FDDC" box="[752,787,573,599]" pageId="9" pageNumber="587" type="female">1♀</specimenCount>
(
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<collectionCode id="24E4AE7CF02E5F0A926BFD8096B9FDDC" box="[867,941,573,599]" pageId="9" pageNumber="587">CCPC</collectionCode>
,
<specimenCount id="54F3FD30F02E5F0A92B1FD8096CAFDDC" box="[953,990,573,599]" pageId="9" pageNumber="587" type="male">1♂</specimenCount>
,
<specimenCount id="54F3FD30F02E5F0A92E3FD809119FDDC" box="[1003,1037,573,599]" pageId="9" pageNumber="587" type="female">1♀</specimenCount>
<collectionCode id="24E4AE7CF02E5F0A951DFD809172FDDC" box="[1045,1126,573,599]" pageId="9" pageNumber="587">ZMPC</collectionCode>
), CHINA:
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, Dehong Pref.,
<collectingCounty id="AB2B4E35F02E5F0A91D2FDDD94B0FDF0" box="[218,420,608,635]" pageId="9" pageNumber="587">Yingjiang County</collectingCounty>
,
<collectingMunicipality id="A22EACC3F02E5F0A90A7FDDD975BFDF0" box="[431,591,608,635]" pageId="9" pageNumber="587">Nabang Town</collectingMunicipality>
,
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,
<date id="364B1079F02E5F0A9261FDDC96DEFDF0" box="[873,970,609,635]" pageId="9" pageNumber="587" value="2013-05-09">
<collectingDate id="260FE991F02E5F0A9261FDDC96DEFDF0" box="[873,970,609,635]" pageId="9" pageNumber="587" value="2013-05-09">2013.v.9</collectingDate>
</date>
<collectorName id="EF00536FF02E5F0A92D9FDDD919BFDF0" box="[977,1167,608,635]" pageId="9" pageNumber="587">Xiao-Dong Yang</collectorName>
leg.
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;
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<specimenCount id="54F3FD30F02E5F0A95C7FDDC91E6FDF0" box="[1231,1266,609,635]" pageId="9" pageNumber="587" type="male">1♂</specimenCount>
(
<collectionCode id="24E4AE7CF02E5F0A9409FDDC905AFDF0" box="[1281,1358,609,635]" pageId="9" pageNumber="587">CCPC</collectionCode>
), ditto, but
<date id="364B1079F02E5F0A91CAFD389423FD14" box="[194,311,645,671]" pageId="9" pageNumber="587" value="2013-05-10">
<collectingDate id="260FE991F02E5F0A91CAFD389423FD14" box="[194,311,645,671]" pageId="9" pageNumber="587" value="2013-05-10">2013.v.10.</collectingDate>
</date>
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</paragraph>
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<subSubSection id="0AEF6532F02E5F0A91CFFD159673FCD8" pageId="9" pageNumber="587" type="discussion">
<paragraph id="424A36B9F02E5F0A91CFFD15960DFC80" blockId="9.[151,1437,464,887]" pageId="9" pageNumber="587">
<emphasis id="7081EAABF02E5F0A91CFFD15947BFD48" bold="true" box="[199,367,680,707]" pageId="9" pageNumber="587">Comments on</emphasis>
<materialsCitation id="F29D3CE4F02E5F0A9070FD14966CFD49" ID-GBIF-Occurrence="3014871311" box="[376,888,680,707]" location="Even" pageId="9" pageNumber="587" specimenCount="1">
<emphasis id="7081EAABF02E5F0A9070FD149705FD49" bold="true" box="[376,529,680,707]" pageId="9" pageNumber="587">
<typeStatus id="9D4E881BF02E5F0A9070FD1494BFFD48" box="[376,427,681,707]" pageId="9" pageNumber="587">type</typeStatus>
locality.
</emphasis>
<location id="472A6062F02E5F0A9312FD149740FD48" LSID="urn:lsid:plazi:treatment:CA5C87AFF02E5F0A919FFF62915BFCFD:472A6062F02E5F0A9312FD149740FD48" box="[538,596,681,707]" name="Even" pageId="9" pageNumber="587">Even</location>
<bibRefCitation id="26644B48F02E5F0A9353FD1497E3FD48" author="Arrow, G. J." box="[603,759,681,707]" pageId="9" pageNumber="587" refId="ref5916" refString="Arrow, G. J. (1917) The fauna of British India, including Ceylon and Burma. Coleoptera Lamellicornia part II (Rutelinae, Des - monycinae, and Euchirinae). Taylor &amp; Francis, London, 387 pp." type="book" year="1917">Arrow (1917)</bibRefCitation>
designated
</materialsCitation>
<typeStatus id="9D4E881BF02E5F0A9289FD1496A5FD48" box="[897,945,681,707]" pageId="9" pageNumber="587">type</typeStatus>
and cotypes for this species originally, the
<materialsCitation id="F29D3CE4F02E5F0A919FFD7094FFFC80" ID-GBIF-Occurrence="3014871307" country="India" county="Mungpoo" location="According" municipality="Mungphu" pageId="9" pageNumber="587" specimenCount="1">
<typeStatus id="9D4E881BF02E5F0A919FFD7095D3FD6C" box="[151,199,717,743]" pageId="9" pageNumber="587">type</typeStatus>
locality was not specified by him.
<location id="472A6062F02E5F0A934EFD7197A8FD6D" LSID="urn:lsid:plazi:treatment:CA5C87AFF02E5F0A919FFF62915BFCFD:472A6062F02E5F0A934EFD7197A8FD6D" box="[582,700,716,742]" country="India" county="Mungpoo" municipality="Mungphu" name="According" pageId="9" pageNumber="587">According</location>
to the above labels,
<collectingMunicipality id="A22EACC3F02E5F0A9294FD71911DFD6D" box="[924,1033,716,742]" pageId="9" pageNumber="587">Mungphu</collectingMunicipality>
(currently
<collectingCounty id="AB2B4E35F02E5F0A958BFD7091E4FD6C" box="[1155,1264,717,743]" pageId="9" pageNumber="587">Mungpoo</collectingCounty>
in northeastern
<collectingCountry id="3AE27629F02E5F0A919FFD4D95C6FC80" box="[151,210,752,779]" name="India" pageId="9" pageNumber="587">India</collectingCountry>
) should be considered as
</materialsCitation>
<typeStatus id="9D4E881BF02E5F0A90FBFD4C9737FC80" box="[499,547,753,779]" pageId="9" pageNumber="587">type</typeStatus>
locality of this taxon.
</paragraph>
<paragraph id="424A36B9F02E5F0A91CFFCA89673FCD8" blockId="9.[151,1437,464,887]" pageId="9" pageNumber="587">
<emphasis id="7081EAABF02E5F0A91CFFCA8942FFCA4" bold="true" box="[199,315,789,815]" pageId="9" pageNumber="587">Remarks.</emphasis>
This species is widely distributed from northern
<collectingCountry id="3AE27629F02E5F0A9266FCA996BCFCA5" box="[878,936,788,814]" name="India" pageId="9" pageNumber="587">India</collectingCountry>
to northern
<collectingCountry id="3AE27629F02E5F0A953EFCA99189FCA5" box="[1078,1181,788,814]" name="Thailand" pageId="9" pageNumber="587">Thailand</collectingCountry>
. The male genitalia of Chinese specimens well fits those figured by
<bibRefCitation id="26644B48F02E5F0A939BFC859677FCD8" author="Prokofiev, A. M." box="[659,867,824,851]" pageId="9" pageNumber="587" pagination="37 - 59" refId="ref6681" refString="Prokofiev, A. M. (2015 a) New Anomala Samouelle, 1819 from South-East Asia. Russian Entomological Journal, 24, 37 - 59. https: // doi. org / 10.15298 / rusentj. 24.1.04" type="journal article" year="2015">Prokofiev (2015a)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="0AEF6532F02E5F0A91CFFCE1915BFCFD" box="[199,1103,860,887]" pageId="9" pageNumber="587" type="distribution">
<paragraph id="424A36B9F02E5F0A91CFFCE1915BFCFD" blockId="9.[151,1437,464,887]" box="[199,1103,860,887]" pageId="9" pageNumber="587">
<emphasis id="7081EAABF02E5F0A91CFFCE19475FCFD" bold="true" box="[199,353,860,886]" pageId="9" pageNumber="587">Distribution.</emphasis>
<collectingCountry id="3AE27629F02E5F0A906FFCE194BAFCFD" box="[359,430,860,886]" name="China" pageId="9" pageNumber="587">China</collectingCountry>
:
<collectingRegion id="8031F85BF02E5F0A90B0FCE0971AFCFC" box="[440,526,861,887]" country="China" name="Yunnan" pageId="9" pageNumber="587">Yunnan</collectingRegion>
<emphasis id="7081EAABF02E5F0A931DFCE19606FCFD" bold="true" box="[533,786,860,887]" pageId="9" pageNumber="587">(new country record)</emphasis>
;
<collectingCountry id="3AE27629F02E5F0A9228FCE19649FCFD" box="[800,861,860,886]" name="India" pageId="9" pageNumber="587">India</collectingCountry>
;
<collectingCountry id="3AE27629F02E5F0A9261FCE096CEFCFC" box="[873,986,861,887]" name="Myanmar" pageId="9" pageNumber="587">Myanmar</collectingCountry>
;
<collectingCountry id="3AE27629F02E5F0A92ECFCE1915FFCFD" box="[996,1099,860,886]" name="Thailand" pageId="9" pageNumber="587">Thailand</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>