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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.164.1992" ID-GBIF-Dataset="d7f2e460-57e5-490f-bfdc-5bca2ab00a39" ID-PMC="PMC3272622" ID-Pensoft-Pub="1313-2970-164-51" ID-PubMed="22328852" ModsDocAuthor="" ModsDocDate="2012" ModsDocID="1313-2970-164-51" ModsDocOrigin="ZooKeys 164" ModsDocTitle="Orthogonius species and diversity in Thailand (Coleoptera, Caraboidea, Orthogoniini), a result from the TIGER project" checkinTime="1451249383683" checkinUser="pensoft" docAuthor="Tian, Mingyi, Deuve, Thierry &amp; Felix, Ron" docDate="2012" docId="B5A9C15E6FE90AE2922CA619E890C8A8" docLanguage="en" docName="ZooKeys 164: 51-90" docOrigin="ZooKeys 164" docSource="http://dx.doi.org/10.3897/zookeys.164.1992" docTitle="Orthogonius pseudochaudoiri Tian, Deuve &amp; Felix, sp. n." docType="treatment" docVersion="3" lastPageNumber="63" masterDocId="FFF7FFDFFF9BFFF8FF99FFF2FFE0FFFD" masterDocTitle="Orthogonius species and diversity in Thailand (Coleoptera, Caraboidea, Orthogoniini), a result from the TIGER project" masterLastPageNumber="90" masterPageNumber="51" pageNumber="60" updateTime="1668153172466" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Orthogonius species and diversity in Thailand (Coleoptera, Caraboidea, Orthogoniini), a result from the TIGER project</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tian, Mingyi</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Deuve, Thierry</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Felix, Ron</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2012</mods:date>
<mods:detail type="volume">
<mods:number>164</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>51</mods:start>
<mods:end>90</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.164.1992</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.164.1992</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-164-51</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152033044" LSID="urn:lsid:plazi:treatment:B5A9C15E6FE90AE2922CA619E890C8A8" httpUri="http://treatment.plazi.org/id/B5A9C15E6FE90AE2922CA619E890C8A8" lastPageId="12" lastPageNumber="63" pageId="9" pageNumber="60">
<subSubSection pageId="9" pageNumber="60" type="nomenclature">
<paragraph pageId="9" pageNumber="60">
<taxonomicName LSID="urn:lsid:zoobank.org:act:597B2644-6DB2-48DC-81A7-24BC6E304379" authority="Tian, Deuve &amp; Felix" class="Insecta" family="Carabidae" genus="Orthogonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthogonius pseudochaudoiri" order="Coleoptera" pageId="9" pageNumber="60" phylum="Arthropoda" rank="species" species="pseudochaudoiri">Orthogonius pseudochaudoiri Tian, Deuve &amp; Felix</taxonomicName>
<taxonomicNameLabel pageId="9" pageNumber="60">sp. n.</taxonomicNameLabel>
Figures 7-921
</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="60" type="diagnosis">
<paragraph pageId="9" pageNumber="60">Diagnosis.</paragraph>
<paragraph pageId="9" pageNumber="60">
Small to medium sized, labrum straight at frontal margin; prosternal process well bordered at apex, abdominal ventrite VII slightly emarginate at apical margin; very similar to
<taxonomicName class="Insecta" family="Carabidae" genus="Orthogonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthogonius mouhoti" order="Coleoptera" pageId="9" pageNumber="60" phylum="Arthropoda" rank="species" species="mouhoti">Orthogonius mouhoti</taxonomicName>
Chaudoir, 1871, but apical lamella of the aedeagus much longer than that of the latter species.
</paragraph>
<paragraph pageId="9" pageNumber="60">Length: 12.5-16.0 mm; width: 5.5-7.0 mm. Habitus as in Figures 7-9.</paragraph>
</subSubSection>
<subSubSection lastPageId="10" lastPageNumber="61" pageId="9" pageNumber="60" type="description">
<paragraph pageId="9" pageNumber="60">Description.</paragraph>
<paragraph pageId="9" pageNumber="60">Dark brown to black, lateral expanded margin of pronotum, antennae, mouthparts palpi, legs and underside surface reddish brown.</paragraph>
<paragraph pageId="9" pageNumber="60">Wrinkles and punctures: surface smooth and impunctate; head and pronotum faintly striate, odd elytral intervals (3, 5, 7) with distinct fine punctures which are irregularly rowed; surface strongly shiny.</paragraph>
<paragraph pageId="9" pageNumber="60">Microsculptural meshes densely isodiametric, clear on elytra, but faint on pronotum and head.</paragraph>
<paragraph lastPageId="10" lastPageNumber="61" pageId="9" pageNumber="60">
Head stout, as long as wide; eyes very large, strongly prominent, frons and vertex moderately convex, frontal impressions small, short, fovea-like, clypeus bisetose, rather even, labrum sexsetose, nearly straight at apical margin; ligula very small and narrow, bisetose at apex; palpi slender, subcylindrical, normal; palpiger asetose, mentum without median tooth, asetose, mentum and submentum each with one pair of setae
<pageBreakToken pageId="10" pageNumber="61" start="start">.</pageBreakToken>
Antennae slender, extended beyond basal 1/3 of elytra, pubescent from apical 2/3 of antennomere 4; antennomeres 3, 4 and 5 subequal in length; antennomeres 1-3 glabrous; antennomeres 4-6 distinctly expanded laterally.
</paragraph>
<paragraph pageId="10" pageNumber="61">Pronotum strongly transverse, PW/PL=1.88-1.90, sides evenly rounded, widest at about middle, both basal and fore margins beaded, lateral expanded margins well defined, wide and even, flat and smooth; both fore and hind angles rounded; disc strongly convex, fore transverse impression faint, basal one moderate, basal foveae small.</paragraph>
<paragraph pageId="10" pageNumber="61">Elytra broadly ovate (EL/WL=1.55-1.57), strongly convex, basal border well complete; sides slightly expanded in middle portion, hardly parallel-sided, widest at middle; striae deep, punctate-striate, intervals distinctly convex; intervals 2-5 subequal in width, interval 6 much wider than 5; odd intervals with more distinct fine punctures, irregular in row; apex roundly truncate, inner angle finely toothed, and with a wider sutural angle; interval 3 with three well marked setiferous pores, near striae 3, 2 and 2, respectively; interval 7 simple, wide and not carinate, without seta throughout.</paragraph>
<paragraph pageId="10" pageNumber="61">Legs moderate, fore tibia with outer angle nearly rectangular, blunt, outer margin not serrate; middle and hind coxae smooth and glabrous; middle and hind tibia slender, apex slightly dilated, hind apical tibial spurs very long and sharp; tarsomere 3 much longer than 4, tarsomere 4 deeply emarginate at apex (about 1/3 deep as the joint); all tarsal claws strongly pectinate; hind femur with 2 posterior setae on ventral.</paragraph>
<paragraph pageId="10" pageNumber="61">Prosternal process well bordered at apex; apical margin of abdominal ventrite VII narrowly and shallowly emarginate in male.</paragraph>
<paragraph pageId="10" pageNumber="61">
Male genitalia (Figure 21): Very similar to that of
<taxonomicName class="Insecta" family="Carabidae" genus="Orthogonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthogonius chaudoiri" order="Coleoptera" pageId="10" pageNumber="61" phylum="Arthropoda" rank="species" species="chaudoiri">Orthogonius chaudoiri</taxonomicName>
, straight, and arrowhead-shaped at apex in dorsal view, but more distinctly so than in
<taxonomicName class="Insecta" family="Carabidae" genus="Orthogonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthogonius chaudoiri" order="Coleoptera" pageId="10" pageNumber="61" phylum="Arthropoda" rank="species" species="chaudoiri">Orthogonius chaudoiri</taxonomicName>
, upper margin less sinuate, and apical lamella in dorsal view much longer and more elongate.
</paragraph>
<paragraph pageId="10" pageNumber="61">Female. Unknown.</paragraph>
</subSubSection>
<subSubSection pageId="10" pageNumber="61" type="remarks">
<paragraph pageId="10" pageNumber="61">Remarks.</paragraph>
<paragraph pageId="10" pageNumber="61">
This species is very similar to
<taxonomicName class="Insecta" family="Carabidae" genus="Orthogonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthogonius chaudoiri" order="Coleoptera" pageId="10" pageNumber="61" phylum="Arthropoda" rank="species" species="chaudoiri">Orthogonius chaudoiri</taxonomicName>
, but the apex of its aedeagus is more distinctly arrowhead-shaped than that of
<taxonomicName class="Insecta" family="Carabidae" genus="Orthogonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthogonius chaudoiri" order="Coleoptera" pageId="10" pageNumber="61" phylum="Arthropoda" rank="species" species="chaudoiri">Orthogonius chaudoiri</taxonomicName>
, less sinuate, and apical lamella much longer; labrum slightly emarginate (straight in latter); body a little more slender; and ventrite VII of male with a small emargination at apical margin.
</paragraph>
</subSubSection>
<subSubSection pageId="10" pageNumber="61" type="variability">
<paragraph pageId="10" pageNumber="61">Variability.</paragraph>
<paragraph pageId="10" pageNumber="61">
Shape of the arrow-headed apex of the aedeagus is variable, wider in some specimens, but narrower in others; however, in all specimens of this species examined, the apical lamella is much longer than that of
<taxonomicName class="Insecta" family="Carabidae" genus="Orthogonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthogonius chaudoiri" order="Coleoptera" pageId="10" pageNumber="61" phylum="Arthropoda" rank="species" species="chaudoiri">Orthogonius chaudoiri</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="12" lastPageNumber="63" pageId="10" pageNumber="61" type="material examined">
<paragraph pageId="10" pageNumber="61">Material examined.</paragraph>
<paragraph pageId="10" pageNumber="61">
Holotype: male, &quot;Thailand: Phetchabum, Thung Salaeng Luang NP, Gang Wang Nam Yen, 16°
<geoCoordinate direction="north" orientation="latitude" precision="55" value="37.531">37.531N</geoCoordinate>
, 100°
<geoCoordinate direction="east" orientation="longitude" precision="55" value="53.745">53.745E</geoCoordinate>
, 721 m, Malaise trap, 17-24.v.2007, Pongpitak Pranee &amp; Sathit leg., T2082&quot;, in QSBG.
</paragraph>
<paragraph lastPageId="12" lastPageNumber="63" pageId="10" pageNumber="61">
Paratypes: 1 male, data as holotype; 6 males, &quot;Thailand: Phetchabun: Thung Salaeng Luang NP, pine forest,
<geoCoordinate direction="north" orientation="latitude" precision="1" value="16.596483">16°35.789'N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="100.879486">100°52.769'E</geoCoordinate>
, 732 m, Malaise trap,15-22.vi.2007, Pongpitak Pranee &amp; Sathit leg., T2059&quot;; 3 males, &quot;Thailand: Phetchabum, Thung Salaeng Luang NP, Gang Wang Nam Yen, 16°
<geoCoordinate direction="north" orientation="latitude" precision="55" value="36.284">36.284N</geoCoordinate>
, 100°
<geoCoordinate direction="east" orientation="longitude" precision="55" value="53.128">53.128E</geoCoordinate>
, 749 m, Malaise trap, 29.
<normalizedToken originalValue="vi">vi-</normalizedToken>
6.vii.2007, Pongpitak Pranee &amp; Sathit leg., T2087&quot;; 2 males, &quot;Thailand: Phetchabun: Thung Salaeng Luang NP, pine forest;G ang Wang Nam Yen,
<geoCoordinate direction="north" orientation="latitude" precision="1" value="16.604733">16°36.284'N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="100.88547">100°53.128'E</geoCoordinate>
, 749 m, 15.vi.2007-18.vi.2007, Litter sample, Pongpitak &amp; Sathit leg., T2050&quot;; 1 male,
<pageBreakToken pageId="11" pageNumber="62" start="start">&quot;</pageBreakToken>
Thailand: Phetchabun: Thung Salaeng Luang NP, pine forest; Gang Wang Nam Yen,
<geoCoordinate direction="north" orientation="latitude" precision="1" value="16.604733">16°36.284'N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="100.88547">100°53.128'E</geoCoordinate>
, 749 m, 30.vi.2007-3.vii.2007, Litter sample, Pongpitak &amp; Sathit leg., T2051&quot;; 6 males, &quot;Thailand: Phetchabun: Thung Salaeng Luang NP, pine forest; Gang Wang Nam Yen,
<geoCoordinate direction="north" orientation="latitude" precision="1" value="16.604733">16°36.284'N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="100.88547">100°53.128'E</geoCoordinate>
, 749 m, 16-17.vi.2007, Pan traps, Pongpitak &amp; Sathit leg., T2052&quot;; 1 male, &quot;Thailand: Phetchabun: Thung Salaeng Luang NP, pine forest; Gang Wang Nam Yen,
<geoCoordinate direction="north" orientation="latitude" precision="1" value="16.604733">16°36.284'N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="100.88547">100°53.128'E</geoCoordinate>
, 749 m, 18-19.vi.2007, Pan traps, Pongpitak &amp; Sathit leg., T2054&quot;; 5 males, &quot;Thailand: Phetchabun: Thung Salaeng Luang NP, pine forest; Gang Wang Nam Yen,
<geoCoordinate direction="north" orientation="latitude" precision="1" value="16.604733">16°36.284'N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="100.88547">100°53.128'E</geoCoordinate>
, 749 m, 20-21.vi.2007, Pan traps, Pongpitak &amp; Sathit leg., T2056&quot;; 1 male, &quot;Thailand: Phetchabun: Thung Salaeng Luang NP, pine forest; Gang Wang Nam Yen,
<geoCoordinate direction="north" orientation="latitude" precision="1" value="16.604733">16°36.284'N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="100.88547">100°53.128'E</geoCoordinate>
, 749 m, 21-22.vi.2007, Pan traps, Pongpitak &amp; Sathit leg., T2057&quot;; 7 males, &quot;Thailand: Phetchabun: Thung Salaeng Luang NP, pine forest; Gang Wang Nam Yen,
<geoCoordinate direction="north" orientation="latitude" precision="1" value="16.604733">16°36.284'N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="100.88547">100°53.128'E</geoCoordinate>
, 749 m, 22-23.vi.2007, Pan traps, Pongpitak &amp; Sathit leg., T2058&quot;; 1 male, &quot;Thailand: Phetchabun: Thung Salaeng Luang NP, pine forest; Gang Wang Nam Yen,
<geoCoordinate direction="north" orientation="latitude" precision="1" value="16.604733">16°36.284'N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="100.88547">100°53.128'E</geoCoordinate>
, 749 m, 22-29.vi.2007, Malaise trap, Pongpitak &amp; Sathit leg., T2063&quot;; 3 males, &quot;Thailand: Phetchabum, Thung Salaeng Luang NP, Gang Wang Nam Yen, pine forest, 16°
<geoCoordinate direction="north" orientation="latitude" precision="55" value="35.789">35.789N</geoCoordinate>
, 100°
<geoCoordinate direction="east" orientation="longitude" precision="55" value="52.769">52.769E</geoCoordinate>
, 723 m, Malaise trap, 6-13.vii.2007, Pongpitak Pranee &amp; Sathit leg. T 2068&quot;; 6 males, &quot;Thailand: Phetchabum, Thung Salaeng Luang NP, Gang Wang Nam Yen, 16°
<geoCoordinate direction="north" orientation="latitude" precision="55" value="36.284">36.284N</geoCoordinate>
, 100°
<geoCoordinate direction="east" orientation="longitude" precision="55" value="53.128">53.128E</geoCoordinate>
, 749 m, Malaise trap, 29.
<normalizedToken originalValue="vi">vi-</normalizedToken>
6.vii.2007, Pongpitak Pranee &amp; Sathit leg., T 2069&quot;; 2 males, &quot;Thailand: Phetchabum, Thung Salaeng Luang NP, Gang Wang Nam Yen, pine forest, 16°
<geoCoordinate direction="north" orientation="latitude" precision="55" value="35.805">35.805N</geoCoordinate>
, 100°
<geoCoordinate direction="east" orientation="longitude" precision="55" value="52.286">52.286E</geoCoordinate>
, 726 m, Malaise trap, 6-13.vii.2007, Pongpitak Pranee &amp; Sathit leg., T 2070&quot;; 3 males, &quot;Thailand: Phetchabun: Thung Salaeng Luang NP: Gang Wang Nam Yen, 750 m,
<geoCoordinate direction="north" orientation="latitude" precision="1" value="16.609783">16°36.587'N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="100.889915">100°53.395'E</geoCoordinate>
; 17-24.v.2007, Pongpitak Pranee &amp; Sathit leg., T2080&quot;; 7 males, &quot;Thailand: Phetchabum, Thung Salaeng Luang NP, Gang Wang Nam Yen, 16°
<geoCoordinate direction="north" orientation="latitude" precision="55" value="37.178">37.178N</geoCoordinate>
, 100°.
<geoCoordinate direction="east" orientation="longitude" precision="55" value="53.504">53.504E</geoCoordinate>
, 706 m, Malaise trap, 17-24.v.2007, Pongpitak Pranee &amp; Sathit leg., T2081&quot;; 1 males, &quot;Thailand: Phetchabun: Thung Salaeng Luang NP, Gang Wang Nam Yen,
<geoCoordinate direction="north" orientation="latitude" precision="1" value="16.609783">16°36.587'N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="100.889915">100°53.395'E</geoCoordinate>
, 753 m, 24-31.v.2007, Malaise trap, Pongpitak Pranee &amp; Sathit leg., T2083&quot;; 2 males, &quot;Thailand: Nakhon Nayok: Khao Yai NP, Lum Ta Kong View Point,
<geoCoordinate direction="north" orientation="latitude" precision="9" value="14.430333">14°25.82'N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="101.3959">101°23.754'E</geoCoordinate>
, 744 m, 26.iv.2007-2.v.2007, Malaise trap, Pong Sandao leg., T2132&quot;; 5 males, &quot;Thailand: Nakhon Ratchasima: Khao Yai NP, Cobra zone near fire protection office, 14°
<geoCoordinate direction="north" orientation="latitude" precision="55" value="27.511">27.511N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="101.37347">101°22.408'E</geoCoordinate>
, 760 m, 5-12.vi.2007, Malaise trap, Pong Sandao leg., T2223&quot;; 1 male, &quot;Thailand: Nakhon Ratchasima: Khao Yai NP, Cobra zone near fire protection office, 14°
<geoCoordinate direction="north" orientation="latitude" precision="55" value="28.285">28.285N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="9" value="101.37617">101°22.57'E</geoCoordinate>
, 751 m, 12-19.vi.2007, Malaise trap, Wirat Sukho leg., T2225&quot;; 1 male, &quot;Thailand: Nakhon Ratchasima: Khao Yai NP, Cobra zone near fire protection office, 14°
<geoCoordinate direction="north" orientation="latitude" precision="55" value="27.511">27.511N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="101.37347">101°22.408'E</geoCoordinate>
, 760 m, 12-19.vi.2007, Malaise trap, Wirat Sukho leg., T2226&quot;; 1 male, &quot;Thailand: Nakhon Nayok: Khao Yai NP, entrance of Hnong Pak Chee Trail,
<geoCoordinate direction="north" orientation="latitude" precision="1" value="14.451917">14°27.115'N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="101.36585">101°21.951'E</geoCoordinate>
, 733 m, 5-6.v.2007, Pan traps, Pong Sandao leg., T2256&quot;; 5 males, &quot;Thailand: Nakhon Nayok: Khao
<pageBreakToken pageId="12" pageNumber="63" start="start">Yai</pageBreakToken>
NP, entrance of Hnong Pak Chee Trail,
<geoCoordinate direction="north" orientation="latitude" precision="1" value="14.451917">14°27.115'N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="101.36585">101°21.951'E</geoCoordinate>
, 733 m, 5-12.v.2007, Malaise traps, Wirat Sukho leg., T2264&quot;; 4 males, &quot;Thailand: Nakhon Nayok: Khao Yai NP, entrance of Hnong Pak Chee Trail,
<geoCoordinate direction="north" orientation="latitude" precision="1" value="14.451917">14°27.115'N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="101.36585">101°21.951'E</geoCoordinate>
, 733 m, 12-19.v.2007, Malaise traps, Pong Sandao leg., T2267&quot;; 3 males, &quot;Thailand: Nakhon Nayok: Khao Yai NP, entrance of Hnong Pak Chee Trail,
<geoCoordinate direction="north" orientation="latitude" precision="1" value="14.451983">14°27.119'N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="101.35803">101°21.482'E</geoCoordinate>
, 699 m, 12-19.v.2007, Malaise traps, Wirat Sukho leg., T2268&quot;; 2 males, &quot;Thailand: Nakhon Nayok: Khao Yai NP, entrance of Hnong Pak Chee Trail,
<geoCoordinate direction="north" orientation="latitude" precision="1" value="14.452784">14°27.167'N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="9" value="101.36417">101°21.85'E</geoCoordinate>
, 758 m, 19-26.v.2007, Malaise traps, Wirat Sukho leg., T2269&quot;; 2 males, &quot;Thailand: Nakhon Nayok: Khao Yai NP, entrance of Hnong Pak Chee Trail,
<geoCoordinate direction="north" orientation="latitude" precision="1" value="14.451917">14°27.115'N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="101.36585">101°21.951'E</geoCoordinate>
, 733 m, 19-26.v.2007, Malaise traps, Pong Sandao leg., T2270&quot;; 2 males, &quot;Thailand: Nakhon Nayok: Khao Yai NP, entrance of Hnong Pak Chee Trail,
<geoCoordinate direction="north" orientation="latitude" precision="1" value="14.451983">14°27.119'N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="101.35803">101°21.482'E</geoCoordinate>
, 699 m, 19-26.v.2007, Malaise traps, Wirat Sukho leg., T2271&quot;; 1 male, &quot;Thailand: Nakhon Nayok: Khao Yai NP, entrance of Hnong Pak Chee Trail,
<geoCoordinate direction="north" orientation="latitude" precision="1" value="14.452784">14°27.167'N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="9" value="101.36417">101°21.85'E</geoCoordinate>
, 758 m, 26.v.2007-2.vi.2007, Malaise traps, Wirat Sukho leg., T2272&quot;; 2 males, &quot;Thailand: Nakhon Nayok: Khao Yai NP, entrance of Hnong Pak Chee Trail,
<geoCoordinate direction="north" orientation="latitude" precision="1" value="14.451917">14°27.115'N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="101.36585">101°21.951'E</geoCoordinate>
, 733 m, 26.v.2007-2.vi.2007, Malaise traps, Pong Sandao leg., T2273&quot;; 1 male, label lost; in QSBG, MNHN, SCAU and CRF, respectively.
</paragraph>
</subSubSection>
<subSubSection pageId="12" pageNumber="63" type="etymology">
<paragraph pageId="12" pageNumber="63">Etymology.</paragraph>
<paragraph pageId="12" pageNumber="63">
The name refers to the similarity of this new species to
<taxonomicName class="Insecta" family="Carabidae" genus="Orthogonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthogonius chaudoiri" order="Coleoptera" pageId="12" pageNumber="63" phylum="Arthropoda" rank="species" species="chaudoiri">Orthogonius chaudoiri</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="12" pageNumber="63" type="distribution">
<paragraph pageId="12" pageNumber="63">Distribution.</paragraph>
<paragraph pageId="12" pageNumber="63">Thailand. Known only from the type localities.</paragraph>
</subSubSection>
</treatment>
</document>