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<mods:title id="F3AB9F4254CC3F92B55CD2153D8E5C40">Revision of the carnivorous snail genus Indoartemon Forcart, 1946 and a new genus Carinartemis from Thailand (Pulmonata: Streptaxidae)</mods:title>
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Genus
<taxonomicName id="37B34D20C44D9C3EFEA23899FD8B99B6" ID-CoL="7P5QV" authority="Forcart, 1946" authorityName="Forcart" authorityYear="1946" box="[320,626,1757,1781]" class="Gastropoda" family="Streptaxidae" genus="Indoartemon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="162" phylum="Mollusca" rank="genus">
<emphasis id="C2C7EAB1C44D9C3EFEA23899FE2999B6" bold="true" box="[320,464,1757,1781]" italics="true" pageId="1" pageNumber="162">Indoartemon</emphasis>
<bibRefCitation id="94224B52C44D9C3EFE35389AFD8B99B6" author="Forcart P" box="[471,626,1758,1781]" pageId="1" pageNumber="162" pagination="215" refId="ref8591" refString="Forcart P (1946) Indoartemon subgen. nov. for Odonartemon Kobelt, 1905 (non Pfeiffer, 1856); Streptaxidae. Journal of Conchology, 22: 215." type="journal article" year="1946">Forcart, 1946</bibRefCitation>
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<taxonomicName id="37B34D20C44D9C3EFF933958FE5E9872" authority="(Odontartemon)" box="[113,423,1820,1841]" class="Gastropoda" family="Streptaxidae" genus="Odontartemon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="162" phylum="Mollusca" rank="subGenus" subGenus="Odontartemon">
<emphasis id="C2C7EAB1C44D9C3EFF933958FE5E9872" box="[113,423,1820,1841]" italics="true" pageId="1" pageNumber="162">Odontartemon (Odontartemon)</emphasis>
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:
<treatmentCitation id="711210B2C44D9C3EFE553958FD989872" author="Kobelt W &amp; Die Raublungenschnecken" box="[439,609,1820,1841]" page="91" pageId="1" pageNumber="162" year="1905">
<bibRefCitation id="94224B52C44D9C3EFE553958FD989872" author="Kobelt W &amp; Die Raublungenschnecken" box="[439,609,1820,1841]" pageId="1" pageNumber="162" refId="ref8839" refString="Kobelt W (1905 - 1906) Die Raublungenschnecken (Agnatha). Zweite Abtheilung: Streptaxidae und Daudebardiidae. Systematisches Conchylien-Cabinet von Martini und Chemnitz, 1 (12 b) (2): 1 - 96, pls 42 - 59 [1905]; 97 - 211, pls 60 - 71 [1906]." type="book" year="1905">Kobelt, 1905: 91</bibRefCitation>
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(non
<bibRefCitation id="94224B52C44D9C3EFD433958FF32980D" author="Pfeiffer L" pageId="1" pageNumber="162" pagination="1 - 192" refId="ref9049" refString="Pfeiffer L (1856) Uebersicht der Gattung Cylindrella. Malakozoologische Blatter. 2 (1855): 1 - 192." type="journal article" year="1856">Pfeiffer, 1856</bibRefCitation>
).
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<bibRefCitation id="94224B52C44D9C3EFF3E397DFE72980D" author="Thiele J &amp; Handbuch der Systematischen Weichtierkunde &amp; Gustav Fischer, Jena" box="[220,395,1849,1870]" pageId="1" pageNumber="162" pagination="1023 - 1134" refId="ref9468" refString="Thiele J (1929 - 1935) Handbuch der Systematischen Weichtierkunde. Gustav Fischer, Jena. 1134 pp. [p. 1 - 376 (1929); p. 377 - 778 (1931); p. 779 - 1022 (1934); p. 1023 - 1134 (1935)]." type="book chapter" year="1931">Thiele, 1931: 730</bibRefCitation>
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<treatmentCitationGroup id="D0A3118DC44D9C3EFF933912FF0D98CB" pageId="1" pageNumber="162">
<treatmentCitation id="711210B2C44D9C3EFF933912FDCC9828" author="Forcart P" box="[113,565,1878,1899]" page="215" pageId="1" pageNumber="162" year="1946">
<taxonomicName id="37B34D20C44D9C3EFF933912FDCC9828" authority="Forcart, 1946: 215" authorityName="Forcart" authorityPageNumber="215" authorityYear="1946" box="[113,565,1878,1899]" class="Gastropoda" family="Streptaxidae" genus="Oophana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="162" phylum="Mollusca" rank="subGenus" subGenus="Indoartemon">
<emphasis id="C2C7EAB1C44D9C3EFF933912FE919828" box="[113,360,1878,1899]" italics="true" pageId="1" pageNumber="162">Oophana (Indoartemon)</emphasis>
<bibRefCitation id="94224B52C44D9C3EFE903912FDCC9828" author="Forcart P" box="[370,565,1878,1899]" pageId="1" pageNumber="162" pagination="215" refId="ref8591" refString="Forcart P (1946) Indoartemon subgen. nov. for Odonartemon Kobelt, 1905 (non Pfeiffer, 1856); Streptaxidae. Journal of Conchology, 22: 215." type="journal article" year="1946">Forcart, 1946: 215</bibRefCitation>
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.
<treatmentCitation id="711210B2C44D9C3EFDA03912FF0998CB" author="van Benthem Jutting WSS" page="95" pageId="1" pageNumber="162" year="1954">
<bibRefCitation id="94224B52C44D9C3EFDA03912FF0998CB" author="van Benthem Jutting WSS" pageId="1" pageNumber="162" pagination="71 - 106" refId="ref8247" refString="van Benthem Jutting WSS (1954) The Malayan Streptaxidae of the genera Discartemon and Oophana. Bulletin of the Raffles Museum, 25: 71 - 106." type="journal article" year="1954">Benthem Jutting, 1954: 95</bibRefCitation>
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.
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<treatmentCitationGroup id="D0A3118DC44D9C3EFF9339CBFE679882" pageId="1" pageNumber="162">
<treatmentCitation id="711210B2C44D9C3EFF9339CBFE4398E7" author="Zilch A" box="[113,442,1935,1956]" page="562" pageId="1" pageNumber="162" year="1960">
<taxonomicName id="37B34D20C44D9C3EFF9339CBFE4398E7" ID-CoL="7P5QV" authority=": Zilch, 1960: 562" authorityName=": Zilch" authorityPageNumber="562" authorityYear="1960" box="[113,442,1935,1956]" class="Gastropoda" family="Streptaxidae" genus="Indoartemon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="162" phylum="Mollusca" rank="genus">
<emphasis id="C2C7EAB1C44D9C3EFF9339CBFF0D98E7" box="[113,244,1935,1956]" italics="true" pageId="1" pageNumber="162">Indoartemon</emphasis>
:
<bibRefCitation id="94224B52C44D9C3EFEE439CBFE4398E7" author="Zilch A" box="[262,442,1935,1956]" pageId="1" pageNumber="162" pagination="401 - 834" refId="ref9662" refString="Zilch A (1960) Gastropoda, Euthyneura. In: Schinderwolf OH (ed.) Handbuch der Palaozoologie. Gebruder Borntraeger, Berlin, 6: 401 - 834." type="journal article" year="1960">Zilch, 1960: 562</bibRefCitation>
</taxonomicName>
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.
<treatmentCitation id="711210B2C44D9C3EFE2A39CBFD1598E7" author="Richardson L" box="[456,748,1935,1956]" page="223" pageId="1" pageNumber="162" year="1988">
<bibRefCitation id="94224B52C44D9C3EFE2A39CBFD1598E7" author="Richardson L" box="[456,748,1935,1956]" pageId="1" pageNumber="162" pagination="1 - 326" refId="ref9221" refString="Richardson L (1988) Streptaxacea: Catalog of species, Part I, Streptaxidae. Tryonia, 16: 1 - 326." type="journal article" year="1988">Richardson, 1988: 223225</bibRefCitation>
</treatmentCitation>
.
<treatmentCitation id="711210B2C44D9C3EFF7539E8FE919882" author="Schileyko AA" box="[151,360,1964,1985]" page="776" pageId="1" pageNumber="162" year="2000">
<bibRefCitation id="94224B52C44D9C3EFF7539E8FE919882" author="Schileyko AA" box="[151,360,1964,1985]" pageId="1" pageNumber="162" pagination="731 - 880" refId="ref9288" refString="Schileyko AA (2000) Treatise on recent terrestrial pulmonate molluscs: Rhytididae; Chlamydephoridae; Systrophiidae; Haplotrematidae; Streptaxidae; Spiraxidae; Oleacinidae; Testacellidae. Ruthenica, Supplement 2, Part 6: 731 - 880." type="journal article" year="2000">Schileyko, 2000: 776</bibRefCitation>
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, 777.
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<emphasis id="C2C7EAB1C44D9C3EFF9339AEFEFF9742" bold="true" box="[113,262,2026,2049]" pageId="1" pageNumber="162">
<typeStatus id="2F088801C44D9C3EFF9339AEFF509742" box="[113,169,2026,2049]" pageId="1" pageNumber="162">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="37B34D20C44D9C3EFEEE39AEFD889742" ID-CoL="52XK6" authority="Pfeiffer, 1861" authorityName="Pfeiffer" authorityYear="1861" box="[268,625,2026,2049]" class="Gastropoda" family="Streptaxidae" genus="Streptaxis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="162" phylum="Mollusca" rank="species" species="eburnea">
<emphasis id="C2C7EAB1C44D9C3EFEEE39AEFE2D9742" box="[268,468,2026,2049]" italics="true" pageId="1" pageNumber="162">Streptaxis eburnea</emphasis>
<bibRefCitation id="94224B52C44D9C3EFE3939AEFD889742" author="Pfeiffer L" box="[475,625,2026,2049]" pageId="1" pageNumber="162" pagination="20 - 29" refId="ref9071" refString="Pfeiffer L (1861) Description of forty-seven new species of landshells, from the collection of H. Cuming, Esq. Proceedings of the Zoological Society of London, 1861: 20 - 29." type="journal article" year="1861">Pfeiffer, 1861</bibRefCitation>
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, by original designation.
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<paragraph id="F00C36A3C44D9C3EFCCF3EFAFA559D56" blockId="1.[813,1452,190,533]" pageId="1" pageNumber="162">
<emphasis id="C2C7EAB1C44D9C3EFCCF3EFAFC699F96" bold="true" box="[813,912,190,213]" pageId="1" pageNumber="162">Remark.</emphasis>
Shells of
<taxonomicName id="37B34D20C44D9C3EFC143EFAFBA19F96" authorityName="Ancey" authorityYear="1884" box="[1014,1112,190,213]" class="Gastropoda" family="Streptaxidae" genus="Oophana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="162" phylum="Mollusca" rank="genus">
<emphasis id="C2C7EAB1C44D9C3EFC143EFAFBA19F96" box="[1014,1112,190,213]" italics="true" pageId="1" pageNumber="162">Oophana</emphasis>
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and
<taxonomicName id="37B34D20C44D9C3EFB6A3EFAFAE99F96" authorityName=": Zilch" authorityYear="1960" box="[1160,1296,190,213]" class="Gastropoda" family="Streptaxidae" genus="Indoartemon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="162" phylum="Mollusca" rank="genus">
<emphasis id="C2C7EAB1C44D9C3EFB6A3EFAFAE99F96" box="[1160,1296,190,213]" italics="true" pageId="1" pageNumber="162">Indoartemon</emphasis>
</taxonomicName>
are similar but there are differences in the apertural dentition.
<taxonomicName id="37B34D20C44D9C3EFAC03E9AFA559FB6" authorityName=": Zilch" authorityYear="1960" box="[1314,1452,222,245]" class="Gastropoda" family="Streptaxidae" genus="Indoartemon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="162" phylum="Mollusca" rank="genus">
<emphasis id="C2C7EAB1C44D9C3EFAC03E9AFA559FB6" box="[1314,1452,222,245]" italics="true" pageId="1" pageNumber="162">Indoartemon</emphasis>
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possesses one parietal lamella and one palatal lamella; in addition basal and columellar lamellae are present in
<taxonomicName id="37B34D20C44D9C3EFA793F5AFB909E16" authority="(Benson, 1853)" baseAuthorityName="Benson" baseAuthorityYear="1853" class="Gastropoda" family="Streptaxidae" genus="Indoartemon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="162" phylum="Mollusca" rank="species" species="layardianus">
<emphasis id="C2C7EAB1C44D9C3EFA793F5AFC499E16" italics="true" pageId="1" pageNumber="162">I. layardianus</emphasis>
(
<bibRefCitation id="94224B52C44D9C3EFC203F7AFB999E16" author="Benson WH" box="[962,1120,318,341]" pageId="1" pageNumber="162" pagination="90 - 97" refId="ref8167" refString="Benson WH (1853) Characters of new Land Shells in Ceylon. The Annals and Magazine of Natural History, Series 2, 12: 90 - 97." type="journal article" year="1853">Benson, 1853</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="37B34D20C44D9C3EFB443F7AFC899E36" authority="(Benson, 1853)" baseAuthorityName="Benson" baseAuthorityYear="1853" class="Gastropoda" family="Streptaxidae" genus="Indoartemon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="162" phylum="Mollusca" rank="species" species="cingalensis">
<emphasis id="C2C7EAB1C44D9C3EFB443F7AFAC79E16" box="[1190,1342,318,341]" italics="true" pageId="1" pageNumber="162">I. cingalensis</emphasis>
(
<bibRefCitation id="94224B52C44D9C3EFAB23F7AFC919E36" author="Benson WH" pageId="1" pageNumber="162" pagination="90 - 97" refId="ref8167" refString="Benson WH (1853) Characters of new Land Shells in Ceylon. The Annals and Magazine of Natural History, Series 2, 12: 90 - 97." type="journal article" year="1853">Benson, 1853</bibRefCitation>
)
</taxonomicName>
. Apertural dentition of
<taxonomicName id="37B34D20C44D9C3EFB603F1AFB119E36" authorityName="Ancey" authorityYear="1884" box="[1154,1256,350,373]" class="Gastropoda" family="Streptaxidae" genus="Oophana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="162" phylum="Mollusca" rank="genus">
<emphasis id="C2C7EAB1C44D9C3EFB603F1AFB119E36" box="[1154,1256,350,373]" italics="true" pageId="1" pageNumber="162">Oophana</emphasis>
</taxonomicName>
usually includes a single parietal lamella, palatal lamella, basal lamella, and columellar lamella. An upper palatal lamella or supracolumellar lamella are present in some species such as
<taxonomicName id="37B34D20C44D9C3EFCA93F9AFAE79EB6" authority="(Mollendorff, 1894)" baseAuthorityName="Mollendorff" baseAuthorityYear="1894" box="[843,1310,478,501]" class="Gastropoda" family="Streptaxidae" genus="Oophana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="162" phylum="Mollusca" rank="species" species="strangulatus">
<emphasis id="C2C7EAB1C44D9C3EFCA93F9AFBC29EB6" box="[843,1083,478,501]" italics="true" pageId="1" pageNumber="162">Oophana strangulatus</emphasis>
(
<bibRefCitation id="94224B52C44D9C3EFBA93F9AFAEF9EB6" author="von Mollendorff O" box="[1099,1302,478,501]" pageId="1" pageNumber="162" pagination="146 - 156" refId="ref8963" refString="von Mollendorff O (1894) On a Collection of Land-Shells from the Samui Islands, Gulf of Siam. Proceedings of the Zoological Society of London, 1894: 146 - 156, pl. 16." type="journal article" year="1894">Möllendorff, 1894</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="37B34D20C44D9C3EFAC83F9AFC069D56" authority="(Mollendorff, 1900)" baseAuthorityName="Mollendorff" baseAuthorityYear="1900" class="Gastropoda" family="Streptaxidae" genus="Oophana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="162" phylum="Mollusca" rank="species" species="diplodon">
<emphasis id="C2C7EAB1C44D9C3EFAC83F9AFA559EB6" box="[1322,1452,478,501]" italics="true" pageId="1" pageNumber="162">O. diplodon</emphasis>
(
<bibRefCitation id="94224B52C44D9C3EFCD13FBAFC0E9D56" author="von Mollendorff O" box="[819,1015,510,533]" pageId="1" pageNumber="162" pagination="117 - 121" refId="ref9001" refString="von Mollendorff O (1900) Zur Binnenmollusken-Fauna Annams III. Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft, 32: 117 - 121." type="journal article" year="1900">Möllendorff, 1900</bibRefCitation>
)
</taxonomicName>
, and
<taxonomicName id="37B34D20C44D9C3EFBD03FBAFA519D56" authority="(Mollendorff, 1900)" baseAuthorityName="Mollendorff" baseAuthorityYear="1900" box="[1074,1448,510,533]" class="Gastropoda" family="Streptaxidae" genus="Oophana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="162" phylum="Mollusca" rank="species" species="pachyglottis">
<emphasis id="C2C7EAB1C44D9C3EFBD03FBAFB299D56" box="[1074,1232,510,533]" italics="true" pageId="1" pageNumber="162">O. pachyglottis</emphasis>
(
<bibRefCitation id="94224B52C44D9C3EFB3E3FBAFA599D56" author="von Mollendorff O" box="[1244,1440,510,533]" pageId="1" pageNumber="162" pagination="117 - 121" refId="ref9001" refString="von Mollendorff O (1900) Zur Binnenmollusken-Fauna Annams III. Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft, 32: 117 - 121." type="journal article" year="1900">Möllendorff, 1900</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="F00C36A3C44D9C3EFCCF3C7AFBBB9C36" blockId="1.[813,1452,574,885]" pageId="1" pageNumber="162">
The first genital morphological study of
<taxonomicName id="37B34D20C44D9C3EFAFA3C7AFA529D16" authorityName=": Zilch" authorityYear="1960" box="[1304,1451,574,597]" class="Gastropoda" family="Streptaxidae" genus="Indoartemon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="162" phylum="Mollusca" rank="genus">
<emphasis id="C2C7EAB1C44D9C3EFAFA3C7AFA529D16" box="[1304,1451,574,597]" italics="true" pageId="1" pageNumber="162">Indoartemon</emphasis>
</taxonomicName>
presented here indicates that the following characters may be diagnostic for the genus; a long penis with a thin penial sheath extending about half of the penis length; penial hooks present. The vas deferens does not pass through the penial sheath. The seminal vesicle is about one and a half times longer than the length from the talon to the branching point of the seminal vesicle. In
<taxonomicName id="37B34D20C44D9C3EFBAA3D5AFB529C76" authorityName="Ancey" authorityYear="1884" box="[1096,1195,798,821]" class="Gastropoda" family="Streptaxidae" genus="Oophana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="162" phylum="Mollusca" rank="genus">
<emphasis id="C2C7EAB1C44D9C3EFBAA3D5AFB529C76" box="[1096,1195,798,821]" italics="true" pageId="1" pageNumber="162">Oophana</emphasis>
</taxonomicName>
the vas deferens enters the penial sheath distally, penial hooks and a seminal vesicle are present (
<bibRefCitation id="94224B52C44D9C3EFC533D1AFBCF9C36" author="Berry AJ" box="[945,1078,862,885]" pageId="1" pageNumber="162" pagination="139 - 150" refId="ref8276" refString="Berry AJ (1963) The anatomy of two Malayan limestone hill Streptaxidae, Sinoennea kanchingensis Tomlin and Oophana diaphanopepla van Benthem Jutting with special reference to the genital system. Proceedings of the Malacological Society of London, 35: 139 - 150." type="journal article" year="1963">Berry, 1963</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>