treatments-xml/data/03/F8/87/03F887B9C5000E4AFF2DFE295BA5FD82.xml
2024-06-21 12:22:17 +02:00

1780 lines
214 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="28E988D39591563CFC996D490A652D2E" ID-CLB-Dataset="27515" ID-DOI="10.11646/zootaxa.4576.2.1" ID-GBIF-Dataset="3e329e8f-2f23-4832-b7e1-748e6c1f2925" ID-ISSN="1175-5326" ID-Zenodo-Dep="2624785" ID-ZooBank="C21B66E0-E7BF-4E82-9EBE-24F7CDEFC8A5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1554273053661" checkinUser="plazi" docAuthor="Dessouassi, Comlan Eugène, Lalèyè, Philippe A. &amp; Udekem, Cédric" docDate="2019" docId="03F887B9C5000E4AFF2DFE295BA5FD82" docLanguage="en" docName="zootaxa.4576.2.1.pdf" docOrigin="Zootaxa 4576 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Charybdis hellerii" docType="treatment" docVersion="9" lastPageNumber="224" masterDocId="FFC1FFC1C5030E5DFFBAFFA05F2AFFE2" masterDocTitle="First record of the globally invasive crab, Charybdis hellerii (A. Milne-Edwards, 1867), in Benin, with notes on its taxonomy (Crustacea, Decapoda, Brachyura, Portunidae)" masterLastPageNumber="238" masterPageNumber="201" pageNumber="204" updateTime="1698716647082" updateUser="ExternalLinkService">
<mods:mods id="22E8847030461242305375B3369C3A6E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="C6DDC5F7363BAA56D1B154A517AE7419">
<mods:title id="5142E431CDD1C8589FB773F2E164E04A">First record of the globally invasive crab, Charybdis hellerii (A. Milne-Edwards, 1867), in Benin, with notes on its taxonomy (Crustacea, Decapoda, Brachyura, Portunidae)</mods:title>
</mods:titleInfo>
<mods:name id="4787346B9AE3FA4B14747830DBF00400" type="personal">
<mods:role id="C555A3EC16BC790F7A241EA100B453DF">
<mods:roleTerm id="A12119892C4853092E809433C467F25B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="77B2C6C9292D35C63492810F3109379B">Dessouassi, Comlan Eugène</mods:namePart>
</mods:name>
<mods:name id="39889A32B1DB8627349B327AA33AF7B2" type="personal">
<mods:role id="884DD6A36BBF32B8B0C8031397C5BBF7">
<mods:roleTerm id="0DBBADE17E14B02784D25FA4F7862C89">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F8999EB1DF0EDC1BBDBD8D1138FFB767">Lalèyè, Philippe A.</mods:namePart>
</mods:name>
<mods:name id="F7CD4127A05E16BE7657A22180331A8E" type="personal">
<mods:role id="5E921E0F1341BFD0AC2CFE289EA6D70D">
<mods:roleTerm id="98512885A997681DF00AE3D0769EBB0E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B5C07330EEBD748C6C56C583CDF5A12E"> Udekem, Cédric</mods:namePart>
</mods:name>
<mods:typeOfResource id="A82FC4B0D024A445FABB3F400E213BB1">text</mods:typeOfResource>
<mods:relatedItem id="B32EC130B2DEE78DEA01AFCBC843426C" type="host">
<mods:titleInfo id="B7F87933809A395AA41AE1961561A271">
<mods:title id="414F5008EBCBB9CB9BA239C1141C5EC8">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="A1A8567BA7F71BF5C7687219D67CA9DF">
<mods:date id="08E168AF385B95B679C4F8F80BB85F61">2019</mods:date>
<mods:detail id="31DC98414474E48790C9B05D93016EA7" type="pubDate">
<mods:number id="2BB8AE13170EA673D398EBB5EDC0F5FC">2019-04-02</mods:number>
</mods:detail>
<mods:detail id="CD7FD96009E83AB4D05D3A1738C6C757" type="volume">
<mods:number id="9689F400251B999AB5C7C0F8ED69CF4F">4576</mods:number>
</mods:detail>
<mods:detail id="18E91E0226C017B771FFA9942C885F6B" type="issue">
<mods:number id="8EA0D4E77C94D47FD869699CBEE4AD0E">2</mods:number>
</mods:detail>
<mods:extent id="E5711542768CA8CCAEBE7DD04766EEA1" unit="page">
<mods:start id="608E87851E9BC5FC493D83AA5D1F60C0">201</mods:start>
<mods:end id="EA773323AF1DDA98CD0C7D37642E2589">238</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="54CBB57696A87DBB470B70C505DC6052">journal article</mods:classification>
<mods:identifier id="0EDC83F45537D69D849D840619802862" type="CLB-Dataset">27515</mods:identifier>
<mods:identifier id="4F4A53AA43ECF20A034228C1A91DED77" type="DOI">10.11646/zootaxa.4576.2.1</mods:identifier>
<mods:identifier id="47F45A1485DFFBA75AF9EF4F7DBCB134" type="GBIF-Dataset">3e329e8f-2f23-4832-b7e1-748e6c1f2925</mods:identifier>
<mods:identifier id="BF03BDB6F5C09D333EC3195848E408C1" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="52E73719E1321A4B8DC715A711D23230" type="Zenodo-Dep">2624785</mods:identifier>
<mods:identifier id="C2729511C82D5B4FF6E6A4080641A985" type="ZooBank">C21B66E0-E7BF-4E82-9EBE-24F7CDEFC8A5</mods:identifier>
</mods:mods>
<treatment id="03F887B9C5000E4AFF2DFE295BA5FD82" ID-DOI="http://doi.org/10.5281/zenodo.5469552" ID-GBIF-Taxon="154533294" ID-Zenodo-Dep="5469552" LSID="urn:lsid:plazi:treatment:03F887B9C5000E4AFF2DFE295BA5FD82" httpUri="http://treatment.plazi.org/id/03F887B9C5000E4AFF2DFE295BA5FD82" lastPageId="23" lastPageNumber="224" pageId="3" pageNumber="204">
<subSubSection id="C34B6524C5000E5EFF2DFE295D91FE41" box="[151,699,393,419]" pageId="3" pageNumber="204" type="nomenclature">
<paragraph id="8BEE36AFC5000E5EFF2DFE295D91FE41" blockId="3.[151,699,393,453]" box="[151,699,393,419]" pageId="3" pageNumber="204">
<heading id="D0A681C3C5000E5EFF2DFE295D91FE41" bold="true" box="[151,699,393,419]" fontSize="11" level="1" pageId="3" pageNumber="204" reason="1">
<taxonomicName id="4C514D2CC5000E5EFF2DFE295D91FE41" ID-CoL="TPTB" authority="(A. Milne-Edwards, 1867)" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1867" box="[151,699,393,419]" class="Malacostraca" family="Portunidae" genus="Charybdis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="hellerii">
<emphasis id="B925EABDC5000E5EFF2DFE295D91FE41" bold="true" box="[151,699,393,419]" pageId="3" pageNumber="204">
<emphasis id="B925EABDC5000E5EFF2DFE295E5AFE41" bold="true" box="[151,368,393,419]" italics="true" pageId="3" pageNumber="204">Charybdis hellerii</emphasis>
(
<bibRefCitation id="EFC04B5EC5000E5EFE3AFE295D99FE41" author="Milne-Edwards, A." box="[384,691,393,419]" pageId="3" pageNumber="204" pagination="263 - 288" refId="ref16786" refString="Milne-Edwards, A. (1867) Descriptions de quelques especes nouvelles de Crustaces Brachyures. Annales de la Societe entomologique de France, Series 4, 7, 263 - 288." type="journal article" year="1867">A. Milne-Edwards, 1867</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34B6524C5000E5EFF2DFE0C5E33FE26" box="[151,281,428,453]" pageId="3" pageNumber="204" type="description">
<paragraph id="8BEE36AFC5000E5EFF2DFE0C5E33FE26" blockId="3.[151,699,393,453]" box="[151,281,428,453]" pageId="3" pageNumber="204">
(
<figureCitation id="136A2A2AC5000E5EFF25FE0C5E3AFE26" box="[159,272,428,453]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-10="FIGURE 11" captionStart-11="FIGURE 12" captionStart-12="FIGURE 13" captionStart-13="FIGURE 14" captionStart-14="FIGURE 15" captionStart-15="FIGURE 16" captionStart-16="FIGURE 17" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStart-6="FIGURE 7" captionStart-7="FIGURE 8" captionStart-8="FIGURE 9" captionStart-9="FIGURE 10" captionStartId-0="5.[151,250,1927,1949]" captionStartId-1="6.[151,250,1919,1941]" captionStartId-10="15.[151,250,1928,1950]" captionStartId-11="16.[151,250,1928,1950]" captionStartId-12="17.[151,250,1928,1950]" captionStartId-13="18.[151,250,1928,1950]" captionStartId-14="19.[151,250,1928,1950]" captionStartId-15="20.[151,250,1928,1950]" captionStartId-16="21.[151,250,1928,1950]" captionStartId-2="7.[151,250,1871,1893]" captionStartId-3="8.[151,250,1942,1964]" captionStartId-4="9.[151,250,1881,1903]" captionStartId-5="10.[151,250,1928,1950]" captionStartId-6="11.[151,250,1928,1950]" captionStartId-7="12.[151,250,1928,1950]" captionStartId-8="13.[151,250,1928,1950]" captionStartId-9="14.[151,250,1928,1950]" captionTargetBox-0="[195,1377,193,1901]" captionTargetBox-1="[179,1408,193,1898]" captionTargetBox-10="[150,1436,193,1907]" captionTargetBox-11="[150,1436,193,1907]" captionTargetBox-12="[150,1436,193,1907]" captionTargetBox-13="[150,1436,193,1907]" captionTargetBox-14="[150,1436,193,1907]" captionTargetBox-15="[150,1436,193,1907]" captionTargetBox-16="[150,1436,193,1907]" captionTargetBox-2="[151,1437,193,1850]" captionTargetBox-3="[169,1454,193,1907]" captionTargetBox-4="[151,1435,193,1860]" captionTargetBox-5="[150,1436,193,1907]" captionTargetBox-6="[150,1436,193,1907]" captionTargetBox-7="[150,1436,193,1907]" captionTargetBox-8="[150,1436,193,1907]" captionTargetBox-9="[150,1436,193,1907]" captionTargetId-0="figure@5.[195,1377,193,1901]" captionTargetId-1="figure@6.[179,1408,193,1898]" captionTargetId-10="figure@15.[150,1436,193,1907]" captionTargetId-11="figure@16.[150,1436,193,1907]" captionTargetId-12="figure@17.[150,1436,193,1907]" captionTargetId-13="figure@18.[150,1436,193,1907]" captionTargetId-14="figure@19.[150,1436,193,1907]" captionTargetId-15="figure@20.[150,1436,193,1907]" captionTargetId-16="figure@21.[150,1436,193,1907]" captionTargetId-2="figure@7.[151,1437,193,1850]" captionTargetId-3="figure@8.[169,1454,193,1907]" captionTargetId-4="figure@9.[151,1435,193,1860]" captionTargetId-5="figure@10.[150,1436,193,1907]" captionTargetId-6="figure@11.[150,1436,193,1907]" captionTargetId-7="figure@12.[150,1436,193,1907]" captionTargetId-8="figure@13.[150,1436,193,1907]" captionTargetId-9="figure@14.[150,1436,193,1907]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-10="15" captionTargetPageId-11="16" captionTargetPageId-12="17" captionTargetPageId-13="18" captionTargetPageId-14="19" captionTargetPageId-15="20" captionTargetPageId-16="21" captionTargetPageId-2="7" captionTargetPageId-3="8" captionTargetPageId-4="9" captionTargetPageId-5="10" captionTargetPageId-6="11" captionTargetPageId-7="12" captionTargetPageId-8="13" captionTargetPageId-9="14" captionText-0="FIGURE 1. Charybdis hellerii (A. Milne-Edwards, 1867), New Caledonia. A. Lectotype, male, 60.0 x 85.0 mm, MNHN-IU- 2000-732 (= MNHN-B732). B, C. Paralectotype, male 49.0 x 72.0 mm, MNHN-IU-2000-761 (= MNHN-B761). A, B. Dorsal habitus. C. Frontal region." captionText-1="FIGURE 2. Charybdis hellerii (A. Milne-Edwards, 1867), New Caledonia. A. Lectotype, male, 60 x 85 mm, MNHN-IU-2000- 732 (= MNHN-B732). B, C. Paralectotype, male 49.0 x 72.0 mm, MNHN-IU-2000-761 (= MNHN-B761). A, B. Ventral habitus. C. Carpus and extremity of adjacent articles of left pereiopod 5 (ventral view)." captionText-10="FIGURE 11. Charybdis hellerii (A. Milne-Edwards, 1867), male, 47.0 x 68.0 mm, New Caledonia, RBINS, INV. 129463. A. Lateral view of chelipeds. B. Ventral habitus. C. Telson. D. Carpus and extremity of adjacent articles of left P5, ventral view). The colour have been enhanced to increase the contrasts." captionText-11="FIGURE 12. Charybdis hellerii (A. Milne-Edwards, 1867), male, 47.0 x 68.0 mm, New Caledonia, RBINS, INV. 129463. A. Right orbital region (ventral view). B. Right basal antennal article." captionText-12="FIGURE 13. Charybdis hellerii (A. Milne-Edwards, 1867), male, 47.0 x 68.0 mm, New Caledonia, RBINS, INV. 129463. A. Right gonopod 1. B. Tip of right gonopod 1. The colour have been enhanced to increase the contrasts." captionText-13="FIGURE 14. Charybdis hellerii (A. Milne-Edwards, 1867), male, 37.0 x 56.0 mm, Queensland, RBINS, INV. 129466. A. Dorsal habitus. B. Frontal region." captionText-14="FIGURE 15. Charybdis hellerii (A. Milne-Edwards, 1867), male, 37.0 x 56.0 mm, Queensland, RBINS, INV. 129466. A. Lateral view of the chelipeds. B. Ventral habitus. C. Telson. D. Carpus and extremity of adjacent articles of right pereiopod 5 (ventral view). The colour have been enhanced to increase the contrasts." captionText-15="FIGURE 16. Charybdis hellerii (A. Milne-Edwards, 1867), male, 37.0 x 56.0 mm, Queensland, RBINS, INV. 129466. A. Right orbital region (ventral view). B. Right basal antennal article. The colour have been enhanced to increase the contrasts." captionText-16="FIGURE 17. Charybdis hellerii (A. Milne-Edwards, 1867), male, 37.0 x 56.0 mm, Queensland, RBINS, INV. 129466. A. Left gonopod 1 B. Tip of gonopod 1." captionText-2="FIGURE 3. Charybdis hellerii (A. Milne-Edwards, 1867), males, New Caledonia, left gonopod1s (rehydrated). A, B. Lectotype, male 60.0 x 85.0 mm, MNHN-IU-2000-732 (= MNHN-B732). C, D, E. Paralectotype, male 49.0 x 72.0 mm, MNHN-IU-2000-761 (=MNHN-B761). A, C, medial view. B, D, external view. E. Tip, external view." captionText-3="FIGURE 4. Charybdis hellerii (A. Milne-Edwards, 1867), male holotype of Goniosoma spiniferum Miers, 1884, about 13 x 17 mm, Queensland, Port Molle, NNM 1881.31. A. Dorsal habitus. B. Ventral habitus. C. Right pereiopod 5 (dorsal view). D. Right pereiopod 5 (ventral view)." captionText-4="FIGURE 5. Charybdis hellerii (A. Milne-Edwards, 1867), males, dorsal habitus, Mergui Archipelago, syntypes of Goniosoma merguiensis De Man, 1887, RMNH.CRUS.D.13101886. A. 25.0 x 40.0 mm. B. 21.0 x 32.0 mm." captionText-5="FIGURE 6. Charybdis hellerii (A. Milne-Edwards, 1867), males, Benin, Channel of Cotonou. A, B. 37 x 56 mm, RBINS, INV. 150111. C. 46.6 x 68.7 mm, RBINS, INV. 150110. A. Habitus dorsal. B, C. Frontal region." captionText-6="FIGURE 7. Charybdis hellerii (A. Milne-Edwards, 1867), male, 37.0 x 56.0 mm, Benin, Channel of Cotonou, RBINS, INV. 150111. A. Lateral view of the chelipeds. B. Ventral habitus. C. Telson. D. Carpus and extremity of adjacent articles of left pereiopod 5 (ventral view)." captionText-7="FIGURE 8. Charybdis hellerii (A. Milne-Edwards, 1867), male, 37.0 x 56.0 mm, Benin, Channel of Cotonou, RBINS, INV. 150111. A. Right orbital region (ventral view). B. Right basal antennal article." captionText-8="FIGURE 9. Charybdis hellerii (A. Milne-Edwards, 1867), male, 37.0 x 56.0 mm, Benin, Channel of Cotonou, RBINS, INV. 150111. A. Gonopod 1. B. Tip of gonopod 1. The colour have been enhanced to increase the contrasts." captionText-9="FIGURE 10. Charybdis hellerii (A. Milne-Edwards, 1867), male, 47.0 x 68.0 mm, New Caledonia, RBINS, INV. 129463. A. Dorsal habitus. B. Frontal region." httpUri-0="https://zenodo.org/record/2624787/files/figure.png" httpUri-1="https://zenodo.org/record/2624789/files/figure.png" httpUri-10="https://zenodo.org/record/2624807/files/figure.png" httpUri-11="https://zenodo.org/record/2624809/files/figure.png" httpUri-12="https://zenodo.org/record/2624811/files/figure.png" httpUri-13="https://zenodo.org/record/2624813/files/figure.png" httpUri-14="https://zenodo.org/record/2624817/files/figure.png" httpUri-15="https://zenodo.org/record/2624819/files/figure.png" httpUri-16="https://zenodo.org/record/2624821/files/figure.png" httpUri-2="https://zenodo.org/record/2624791/files/figure.png" httpUri-3="https://zenodo.org/record/2624793/files/figure.png" httpUri-4="https://zenodo.org/record/2624795/files/figure.png" httpUri-5="https://zenodo.org/record/2624797/files/figure.png" httpUri-6="https://zenodo.org/record/2624799/files/figure.png" httpUri-7="https://zenodo.org/record/2624801/files/figure.png" httpUri-8="https://zenodo.org/record/2624803/files/figure.png" httpUri-9="https://zenodo.org/record/2624805/files/figure.png" pageId="3" pageNumber="204">Figs 117</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34B6524C5000E5EFF2DFE545D59F936" pageId="3" pageNumber="204" type="reference_group">
<paragraph id="8BEE36AFC5000E5EFF2DFE545C33FDE8" blockId="3.[151,1437,500,1933]" box="[151,793,500,522]" pageId="3" pageNumber="204">
<treatmentCitationGroup id="AB411181C5000E5EFF2DFE545C33FDE8" box="[151,793,500,522]" pageId="3" pageNumber="204">
<taxonomicName id="4C514D2CC5000E5EFF2DFE545D6FFDE8" authority="Ruppell, 1830: 4" authorityName="Ruppell" authorityPageNumber="4" authorityYear="1830" box="[151,581,500,522]" class="Malacostraca" family="Portunidae" genus="Talamita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="sexdentata">
<emphasis id="B925EABDC5000E5EFF2DFE545FDAFDE8" box="[151,240,500,522]" italics="true" pageId="3" pageNumber="204">Talamita</emphasis>
[sic]
<emphasis id="B925EABDC5000E5EFE90FE545EBCFDE8" box="[298,406,500,522]" italics="true" pageId="3" pageNumber="204">sexdentata</emphasis>
<treatmentCitation id="0AF010BEC5000E5EFE26FE545D6FFDE8" author="Ruppell, E." box="[412,581,500,522]" page="4" pageId="3" pageNumber="204" year="1830">
<bibRefCitation id="EFC04B5EC5000E5EFE26FE545D05FDE8" author="Ruppell, E." box="[412,559,500,522]" pageId="3" pageNumber="204" pagination="1 - 6" refId="ref18354" refString="Ruppell, E. (1830) Beschreibung und Abbildung von 24 Arten kurzschwanzigen Krabben, also Beitrag zur Naturgeschichte des Rothen Meeres. H. L. Bronner, Frankfurt am Main, 28 pp., pls 1 - 6." type="book chapter" year="1830">Rüppell, 1830</bibRefCitation>
: 4
</treatmentCitation>
</taxonomicName>
(in part), pl. 1 fig. 1.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BEE36AFC5000E5EFF2DFDB35B91FDCA" blockId="3.[151,1437,500,1933]" box="[151,1211,530,552]" pageId="3" pageNumber="204">
<treatmentCitationGroup id="AB411181C5000E5EFF2DFDB35B91FDCA" box="[151,1211,530,552]" pageId="3" pageNumber="204">
<taxonomicName id="4C514D2CC5000E5EFF2DFDB35EBAFDCA" ID-CoL="3GXMV" authorityName="De Man" authorityYear="1879" box="[151,400,530,552]" class="Malacostraca" family="Portunidae" genus="Goniosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="sexdentatum">
<emphasis id="B925EABDC5000E5EFF2DFDB35EBAFDCA" box="[151,400,530,552]" italics="true" pageId="3" pageNumber="204">Goniosoma sexdentatum</emphasis>
</taxonomicName>
.—
<treatmentCitation id="0AF010BEC5000E5EFE0BFDB35DF3FDCA" author="Milne-Edwards, A." box="[433,729,530,552]" page="372" pageId="3" pageNumber="204" year="1861">
<bibRefCitation id="EFC04B5EC5000E5EFE0BFDB35D80FDCA" author="Milne-Edwards, A." box="[433,682,530,552]" pageId="3" pageNumber="204" pagination="309 - 430" refId="ref16732" refString="Milne-Edwards, A. (1861) Etudes zoologiques sur les Crustaces recents de la famille des Portuniens. Archives du Museum national d'Histoire naturelle, Paris, 10, 309 - 430, pls. 28 - 38. https: // doi. org / 10.5962 / bhl. title. 10629" type="journal article" year="1861">A. Milne-Edwards 1861</bibRefCitation>
: 372
</treatmentCitation>
(fide
<bibRefCitation id="EFC04B5EC5000E5EFCAFFDB35CA4FDCA" author="Leene, J. E." box="[789,910,530,552]" pageId="3" pageNumber="204" pagination="1 - 156" refId="ref15785" refString="Leene, J. E. (1938) The Decapoda Brachyura of the Siboga-Expedition. VII. Brachygnatha: Portunidae. Siboga Expeditie Monografie, 39 c 3, 1 - 156." type="journal article" year="1938">Leene 1938</bibRefCitation>
, who examined the material).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BEE36AFC5000E5EFF2DFD925CDDFDA5" blockId="3.[151,1437,500,1933]" box="[151,1015,561,583]" pageId="3" pageNumber="204">
<treatmentCitationGroup id="AB411181C5000E5EFF2DFD925CDDFDA5" box="[151,1015,561,583]" pageId="3" pageNumber="204">
<treatmentCitation id="0AF010BEC5000E5EFF2DFD925DB3FDA5" author="Milne-Edwards, A." box="[151,665,561,583]" page="282" pageId="3" pageNumber="204" year="1867">
<taxonomicName id="4C514D2CC5000E5EFF2DFD925D4CFDA5" ID-CoL="3GXMJ" authority="A. Milne-Edwards, 1867" authorityName="A. Milne-Edwards" authorityYear="1867" box="[151,614,561,583]" class="Malacostraca" family="Portunidae" genus="Goniosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="225" phylum="Arthropoda" rank="species" species="hellerii">
<emphasis id="B925EABDC5000E5EFF2DFD925E4BFDA5" box="[151,353,561,583]" italics="true" pageId="3" pageNumber="204">Goniosoma Hellerii</emphasis>
<bibRefCitation id="EFC04B5EC5000E5EFEDDFD925D4CFDA5" author="Milne-Edwards, A." box="[359,614,561,583]" pageId="3" pageNumber="204" pagination="263 - 288" refId="ref16786" refString="Milne-Edwards, A. (1867) Descriptions de quelques especes nouvelles de Crustaces Brachyures. Annales de la Societe entomologique de France, Series 4, 7, 263 - 288." type="journal article" year="1867">A. Milne-Edwards, 1867</bibRefCitation>
</taxonomicName>
: 282
</treatmentCitation>
; 1873: 167;
<treatmentCitation id="0AF010BEC5000E5EFCACFD915CD9FDA5" author="Henderson, J. R." box="[790,1011,561,583]" page="375" pageId="3" pageNumber="204" year="1893">
<bibRefCitation id="EFC04B5EC5000E5EFCACFD915CEAFDA5" author="Henderson, J. R." box="[790,960,561,583]" pageId="3" pageNumber="204" pagination="325 - 458" refId="ref15042" refString="Henderson, J. R. (1893) A contribution to Indian carcinology. Transactions of the Linnaean Society of London Zoology, Series 2, 5, 325 - 458, pls. 36 - 40. https: // doi. org / 10.1111 / j. 1096 - 3642.1893. tb 00653. x" type="journal article" year="1893">Henderson 1893</bibRefCitation>
: 375
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BEE36AFC5000E5EFF2DFDF15C35FD84" blockId="3.[151,1437,500,1933]" box="[151,799,592,614]" pageId="3" pageNumber="204">
<treatmentCitationGroup id="AB411181C5000E5EFF2DFDF15C35FD84" box="[151,799,592,614]" pageId="3" pageNumber="204">
<taxonomicName id="4C514D2CC5000E5EFF2DFDF15E54FD84" box="[151,382,592,614]" class="Malacostraca" family="Portunidae" genus="Goniosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="annulatum">
<emphasis id="B925EABDC5000E5EFF2DFDF15E54FD84" box="[151,382,592,614]" italics="true" pageId="3" pageNumber="204">Goniosoma annulatum</emphasis>
</taxonomicName>
.—
<treatmentCitation id="0AF010BEC5000E5EFE26FDF15D76FD84" author="De Man, J. G." box="[412,604,592,614]" page="151" pageId="3" pageNumber="204" year="1883">
<bibRefCitation id="EFC04B5EC5000E5EFE26FDF15D06FD84" author="De Man, J. G." box="[412,556,592,614]" pageId="3" pageNumber="204" pagination="150 - 169" refId="ref13013" refString="De Man, J. G. (1883) Note XV. Carcinological studies in the Leyden Museum. No 3. Notes from the Leyden Museum, 5, 150 - 169." type="journal article" year="1883">De Man 1883</bibRefCitation>
: 151
</treatmentCitation>
(fide
<bibRefCitation id="EFC04B5EC5000E5EFD23FDF15C3EFD84" author="Leene, J. E." box="[665,788,592,614]" pageId="3" pageNumber="204" pagination="165 - 176" refId="ref15738" refString="Leene, J. E. (1937) Notes on Charybdis demani nov. spec., Charybdis variegata var. brevispinosa nov. var. and other Charybdis - species. Zoologische Mededelingen, Leiden, 19 (13), 165 - 176." type="journal article" year="1937">Leene 1937</bibRefCitation>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BEE36AFC5000E5EFF2DFDCF5B2CFD66" blockId="3.[151,1437,500,1933]" box="[151,1030,622,644]" pageId="3" pageNumber="204">
<treatmentCitationGroup id="AB411181C5000E5EFF2DFDCF5B2CFD66" box="[151,1030,622,644]" pageId="3" pageNumber="204">
<taxonomicName id="4C514D2CC5000E5EFF2DFDCF5D1EFD66" ID-CoL="3GXMW" authority="Miers, 1884: 233" authorityName="Miers" authorityPageNumber="233" authorityYear="1884" box="[151,564,622,644]" class="Malacostraca" family="Portunidae" genus="Goniosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="spiniferum">
<emphasis id="B925EABDC5000E5EFF2DFDCF5E54FD66" box="[151,382,622,644]" italics="true" pageId="3" pageNumber="204">Goniosoma spiniferum</emphasis>
<treatmentCitation id="0AF010BEC5000E5EFE3EFDCE5D1EFD66" author="Miers, E. J." box="[388,564,622,644]" page="233" pageId="3" pageNumber="204" year="1884">
<bibRefCitation id="EFC04B5EC5000E5EFE3EFDCE5D28FD66" author="Miers, E. J." box="[388,514,622,644]" pageId="3" pageNumber="204" pagination="513 - 575" refId="ref16627" refString="Miers, E. J. (1884) Crustacea. In: Report on the zoological collections in the Indopacific Ocean during the voyage of H. M. S. &quot; Alert &quot; 1881 - 82. Part II. Collections from the Western Indian Ocean. Printed by order of the trustees, London, pp. 513 - 575, pls. 46 - 51." type="book chapter" year="1884">Miers, 1884</bibRefCitation>
: 233
</treatmentCitation>
</taxonomicName>
, pl. 23 fig. C.; De Man 1887: 85 (discussion).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BEE36AFC5000E5EFF2DFD2E5CA4FD20" blockId="3.[151,1437,500,1933]" pageId="3" pageNumber="204">
<treatmentCitationGroup id="AB411181C5000E5EFF2DFD2E5CA4FD20" bridged="true" pageId="3" pageNumber="204">
<taxonomicName id="4C514D2CC5000E5EFF2DFD2E5D03FD41" ID-CoL="9CBDG" authority="De Man, 1887" authorityName="De Man" authorityYear="1887" box="[151,553,653,675]" class="Malacostraca" family="Portunidae" genus="Goniosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="merguiense">
<emphasis id="B925EABDC5000E5EFF2DFD2E5EA3FD41" box="[151,393,654,675]" italics="true" pageId="3" pageNumber="204">Goniosoma merguiense</emphasis>
De Man, 1887
</taxonomicName>
[
<date id="FFEF106FC5000E5EFD82FD2D5D9BFD41" box="[568,689,653,675]" pageId="3" pageNumber="204" value="1887" valueMax="1888">18871888</date>
a]: 82, pl. 5 figs 34; 1888 [
<date id="FFEF106FC5000E5EFC63FD2D5B78FD41" box="[985,1106,653,675]" pageId="3" pageNumber="204" value="1887" valueMax="1888">18871888</date>
b]: 335 (discussion); 1895: 560; 1897: pl. 12 fig. 9;
<treatmentCitation id="0AF010BEC5000E5EFE30FD0D5D06FD20" author="Lenz, H." box="[394,556,684,706]" page="360" pageId="3" pageNumber="204" year="1905">
<bibRefCitation id="EFC04B5EC5000E5EFE30FD0D5ED3FD20" author="Lenz, H." box="[394,505,684,706]" pageId="3" pageNumber="204" pagination="341 - 392" refId="ref15918" refString="Lenz, H. (1905) Ostafrikanische Dekapoden und Stomatopoden gesammelt von Herrn Prof. Dr. A. Voeltzkow. Abhandlungen der Senckenbergischen naturforschenden Gesellschaft, 27 (4), 341 - 392, pls. 47 - 48." type="journal article" year="1905">Lenz 1905</bibRefCitation>
: 360
</treatmentCitation>
; 1910: 556;
<treatmentCitation id="0AF010BEC5000E5EFD13FD0C5CA0FD20" author="Klunzinger, C. B." box="[681,906,684,706]" page="367" pageId="3" pageNumber="204" year="1913">
<bibRefCitation id="EFC04B5EC5000E5EFD13FD0C5C7DFD20" author="Klunzinger, C. B." box="[681,855,684,706]" pageId="3" pageNumber="204" pagination="97 - 402" refId="ref15555" refString="Klunzinger, C. B. (1913) Die Rundkrabben (Cyclometopa) des Roten Meeres. Abhandlungen der kaiserlich Leop. - Carol. Deutschen Akademie der Naturforscher Halle, 99 (2), 97 - 402, pls. 5 - 11." type="journal article" year="1913">Klunzinger 1913</bibRefCitation>
: 367
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BEE36AFC5000E5EFF2DFD6A5A1FFD1D" blockId="3.[151,1437,500,1933]" pageId="3" pageNumber="204">
<treatmentCitationGroup id="AB411181C5000E5EFF2DFD6A5A1FFD1D" pageId="3" pageNumber="204">
<taxonomicName id="4C514D2CC5000E5EFF2DFD6A5D24FD02" ID-CoL="TPV8" baseAuthorityName="De Man" baseAuthorityYear="1887" box="[151,526,714,736]" class="Malacostraca" family="Portunidae" genus="Charybdis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="merguiensis" subGenus="Goniosoma">
<emphasis id="B925EABDC5000E5EFF2DFD6A5E2BFD02" box="[151,257,714,736]" italics="true" pageId="3" pageNumber="204">Charybdis</emphasis>
(
<emphasis id="B925EABDC5000E5EFEAAFD6B5EAFFD02" box="[272,389,715,736]" italics="true" pageId="3" pageNumber="204">Goniosoma</emphasis>
)
<emphasis id="B925EABDC5000E5EFE2FFD6B5D24FD02" box="[405,526,715,736]" italics="true" pageId="3" pageNumber="204">merguiensis</emphasis>
</taxonomicName>
.—
<treatmentCitation id="0AF010BEC5000E5EFD91FD6A5DF1FD02" author="Alcock, A." box="[555,731,714,736]" page="49" pageId="3" pageNumber="204" year="1899">
<bibRefCitation id="EFC04B5EC5000E5EFD91FD6A5D9FFD02" author="Alcock, A." box="[555,693,714,736]" pageId="3" pageNumber="204" pagination="1 - 104" refId="ref10886" refString="Alcock, A. (1899) Materials for a Carcinological Fauna of India. No. 4. The Brachyura Cyclometopa. Part II. A Revision, of the Cyclometopa, with an Account of the Families Portunidae, Cancridae and Corystidae. Journal of the Asiatic Society of Bengal, part 2 (natural science), 68 (1), 1 - 104. https: // doi. org / 10.5962 / bhl. title. 15343" type="journal article" year="1899">Alcock 1899</bibRefCitation>
: 49
</treatmentCitation>
, 55;
<treatmentCitation id="0AF010BEC5000E5EFCB4FD6A5CEAFD02" author="Nobili, G." box="[782,960,714,736]" page="254" pageId="3" pageNumber="204" year="1899">
<bibRefCitation id="EFC04B5EC5000E5EFCB4FD6A5CA6FD02" author="Nobili, G." box="[782,908,714,736]" pageId="3" pageNumber="204" pagination="230 - 282" refId="ref17540" refString="Nobili, G. (1899) Contribuzioni alla conoscenza della fauna carcinologica della Papuasia, delle Molluche e dell' Australia. Annali del Museo Civico di Storia Naturale di Genova, Series 2, 40, 230 - 282." type="journal article" year="1899">Nobili 1899</bibRefCitation>
: 254
</treatmentCitation>
; 1903: 31; 1906: 194, 196;
<treatmentCitation id="0AF010BEC5000E5EFB64FD6A5AB2FD02" author="Monod, T." box="[1246,1432,714,736]" page="140" pageId="3" pageNumber="204" year="1930">
<bibRefCitation id="EFC04B5EC5000E5EFB64FD6A5A4EFD02" author="Monod, T." box="[1246,1380,714,736]" pageId="3" pageNumber="204" pagination="135 - 141" refId="ref16912" refString="Monod, T. (1930) Uber einige indo-pazifische Dekapoden der Meeresfauna Syriens. Zoologischer Anzeiger, 92 (5 / 6), 135 - 141." type="journal article" year="1930">Monod 1930</bibRefCitation>
: 140
</treatmentCitation>
, fig. 7;
<treatmentCitation id="0AF010BEC5000E5EFEB0FD495EE9FD1D" author="Chopra, B." box="[266,451,745,767]" page="484" pageId="3" pageNumber="204" year="1935">
<bibRefCitation id="EFC04B5EC5000E5EFEB0FD495EBBFD1D" author="Chopra, B." box="[266,401,745,767]" pageId="3" pageNumber="204" pagination="463 - 514" refId="ref12083" refString="Chopra, B. (1935) Further notes on the Crustacea Decapoda in the Indian Museum. VIII. On the Decapod Crustacea collected by the Bengal Pilot Service off the mouth of the River Hooghly. Brachygnatha (Oxyrhyncha and Brachyrhyncha). Records of the Indian Museum, 37, 463 - 514." type="journal article" year="1935">Chopra 1935</bibRefCitation>
: 484
</treatmentCitation>
, fig. 8;
<treatmentCitation id="0AF010BEC5000E5EFDAAFD495D84FD1D" author="Shen, C. - J." box="[528,686,745,767]" page="116" pageId="3" pageNumber="204" year="1937">
<bibRefCitation id="EFC04B5EC5000E5EFDAAFD495DAAFD1D" author="Shen, C. - J." box="[528,640,745,767]" pageId="3" pageNumber="204" pagination="96 - 139" refId="ref18781" refString="Shen, C. - J. (1937) Notes on a collection of swimming crabs (Portunidae) from Singapore. Bulletin of the Raffles Museum, 13, 96 - 139." type="journal article" year="1937">Shen 1937</bibRefCitation>
: 116
</treatmentCitation>
(key), 121, fig. 12;
<treatmentCitation id="0AF010BEC5000E5EFCC2FD4A5B0FFD1D" author="Leene, J. E." box="[888,1061,745,767]" page="165" pageId="3" pageNumber="204" year="1937">
<bibRefCitation id="EFC04B5EC5000E5EFCC2FD4A5CD8FD1D" author="Leene, J. E." box="[888,1010,745,767]" pageId="3" pageNumber="204" pagination="165 - 176" refId="ref15738" refString="Leene, J. E. (1937) Notes on Charybdis demani nov. spec., Charybdis variegata var. brevispinosa nov. var. and other Charybdis - species. Zoologische Mededelingen, Leiden, 19 (13), 165 - 176." type="journal article" year="1937">Leene 1937</bibRefCitation>
: 165
</treatmentCitation>
;
<treatmentCitation id="0AF010BEC5000E5EFB8AFD495A1BFD1D" author="Chopra, B. &amp; Das, K. N." box="[1072,1329,745,767]" page="394" pageId="3" pageNumber="204" year="1938">
<bibRefCitation id="EFC04B5EC5000E5EFB8AFD495BD4FD1D" author="Chopra, B. &amp; Das, K. N." box="[1072,1278,745,767]" pageId="3" pageNumber="204" pagination="377 - 434" refId="ref12140" refString="Chopra, B. &amp; Das, K. N. (1938) Further notes on the Crustacea Decapoda in the Indian Museum. IX. On three collections of crabs from Tavoy and Mergui Archipelago. Records of the Indian Museum, 39, 377 - 434." type="journal article" year="1938">Chopra &amp; Das 1938</bibRefCitation>
: 394
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BEE36AFC5000E5EFF2DFCA85C6FFCFC" blockId="3.[151,1437,500,1933]" box="[151,837,776,798]" pageId="3" pageNumber="204">
<treatmentCitationGroup id="AB411181C5000E5EFF2DFCA85C6FFCFC" box="[151,837,776,798]" pageId="3" pageNumber="204">
<taxonomicName id="4C514D2CC5000E5EFF2DFCA85E41FCFC" box="[151,363,776,798]" class="Malacostraca" family="Portunidae" genus="Charybdis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="spiniferus">
<emphasis id="B925EABDC5000E5EFF2DFCA85E41FCFC" box="[151,363,776,798]" italics="true" pageId="3" pageNumber="204">Charybdis spiniferus</emphasis>
</taxonomicName>
.—
<treatmentCitation id="0AF010BEC5000E5EFE32FCA85D9FFCFC" author="Grant, F. E. &amp; McCulloch, A. R." box="[392,693,776,798]" page="18" pageId="3" pageNumber="204" year="1906">
<bibRefCitation id="EFC04B5EC5000E5EFE32FCA85DB9FCFC" author="Grant, F. E. &amp; McCulloch, A. R." box="[392,659,776,798]" pageId="3" pageNumber="204" pagination="2 - 53" refId="ref14915" refString="Grant, F. E. &amp; McCulloch, A. R. (1906) On a collection of Crustacea from the Port Curtis district, Queensland. Proceedings of the Linnean Society of New South Wales, 31 (1), No. 121, 2 - 53, pls. 1 - 4." type="journal article" year="1906">Grant &amp; McCulloch 1906</bibRefCitation>
: 18
</treatmentCitation>
(new record).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BEE36AFC5000E5EFF2DFC865A49FC34" blockId="3.[151,1437,500,1933]" pageId="3" pageNumber="204">
<treatmentCitationGroup id="AB411181C5000E5EFF2DFC865A49FC34" pageId="3" pageNumber="204">
<taxonomicName id="4C514D2CC5000E5EFF2DFC865EF3FCDE" ID-CoL="TPTB" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1867" box="[151,473,806,828]" class="Malacostraca" family="Portunidae" genus="Charybdis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="hellerii" subGenus="Charybdis">
<emphasis id="B925EABDC5000E5EFF2DFC865E2BFCDE" box="[151,257,806,828]" italics="true" pageId="3" pageNumber="204">Charybdis</emphasis>
(
<emphasis id="B925EABDC5000E5EFEA9FC865E57FCDE" box="[275,381,806,828]" italics="true" pageId="3" pageNumber="204">Charybdis</emphasis>
)
<emphasis id="B925EABDC5000E5EFE2AFC865EF3FCDE" box="[400,473,806,828]" italics="true" pageId="3" pageNumber="204">hellerii</emphasis>
</taxonomicName>
.—
<treatmentCitation id="0AF010BEC5000E5EFE4CFC875DB5FCDE" author="Leene, J. E." box="[502,671,806,828]" page="44" pageId="3" pageNumber="204" year="1938">
<bibRefCitation id="EFC04B5EC5000E5EFE4CFC875D52FCDE" author="Leene, J. E." box="[502,632,806,828]" pageId="3" pageNumber="204" pagination="1 - 156" refId="ref15785" refString="Leene, J. E. (1938) The Decapoda Brachyura of the Siboga-Expedition. VII. Brachygnatha: Portunidae. Siboga Expeditie Monografie, 39 c 3, 1 - 156." type="journal article" year="1938">Leene 1938</bibRefCitation>
: 44
</treatmentCitation>
49, figs 1517; 1940: 182 (examination of the
<typeStatus id="54EA880DC5000E5EFB1EFC865BD7FCDE" box="[1188,1277,806,828]" pageId="3" pageNumber="204" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C514D2CC5000E5EFA9DFC875E19FCB9" ID-CoL="3GXMW" authorityName="Miers" authorityYear="1884" class="Malacostraca" family="Portunidae" genus="Goniosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="spiniferum">
<emphasis id="B925EABDC5000E5EFA9DFC875E19FCB9" italics="true" pageId="3" pageNumber="204">Goniosoma spiniferum</emphasis>
</taxonomicName>
);
<treatmentCitation id="0AF010BEC5000E5EFEF0FCE55D01FCB9" author="Stephenson, W." box="[330,555,837,859]" page="11" pageId="3" pageNumber="204" year="1972">
<bibRefCitation id="EFC04B5EC5000E5EFEF0FCE55D22FCB9" author="Stephenson, W." box="[330,520,837,859]" pageId="3" pageNumber="204" pagination="1 - 64" refId="ref18960" refString="Stephenson, W. (1972 b) An annotated check list and key to the Indo-West Pacific swimming crabs (Crustacea: Decapoda: Portunidae). Bulletin of the Royal Society of New Zealand, 10, 1 - 64." type="journal article" year="1972">Stephenson 1972b</bibRefCitation>
: 11
</treatmentCitation>
(key), 32; 1976: 14;
<treatmentCitation id="0AF010BEC5000E5EFCBCFCE55BB0FCB9" author="Kathirvel, M. &amp; Gopalakrishnan, K. N." box="[774,1178,837,859]" page="286" pageId="3" pageNumber="204" year="1974">
<bibRefCitation id="EFC04B5EC5000E5EFCBCFCE55B4CFCB9" author="Kathirvel, M. &amp; Gopalakrishnan, K. N." box="[774,1126,837,859]" pageId="3" pageNumber="204" pagination="286 - 287" refId="ref15472" refString="Kathirvel, M. &amp; Gopalakrishnan, K. N. (1974) On the occurence of Charybdis (Charybdis) hellerii (A. Milne Edwards) (Decapoda: Portunidae) along the West Coast of India. Journal of the Marine Biological Association of India, 16, 286 - 287." type="journal article" year="1974">Kathirvel &amp; Gopalakrishnan 1974</bibRefCitation>
: 286
</treatmentCitation>
;
<treatmentCitation id="0AF010BEC5000E5EFB1FFCE55AA5FCB9" author="Dai, A. &amp; Yang, S." box="[1189,1423,837,859]" page="233" pageId="3" pageNumber="204" year="1991">
<bibRefCitation id="EFC04B5EC5000E5EFB1FFCE55A74FCB9" author="Dai, A. &amp; Yang, S." box="[1189,1374,837,859]" pageId="3" pageNumber="204" refId="ref12533" refString="Dai, A. &amp; Yang, S. (1991) Crabs of the China Seas. China Ocean Press, Beijing and Springer-Verlag, Berlin, Heidelberg, New York, Tokyo, 608 pp. [translation from Chinese original 1986]. https: // doi. org / 10.1163 / 1937240 X 91 X 00518" type="book" year="1991">Dai &amp; Yang 1991</bibRefCitation>
: 233
</treatmentCitation>
234, fig. 126 (1), pl. 28 (6);
<treatmentCitation id="0AF010BEC5000E5EFE5FFCC55D99FC98" author="Wee, D. P. C. &amp; Ng, P. K. L." box="[485,691,868,890]" page="32" pageId="3" pageNumber="204" year="1995">
<bibRefCitation id="EFC04B5EC5000E5EFE5FFCC55DA4FC98" author="Wee, D. P. C. &amp; Ng, P. K. L." box="[485,654,868,890]" pageId="3" pageNumber="204" pagination="1 - 128" refId="ref19791" refString="Wee, D. P. C. &amp; Ng, P. K. L. (1995) Swimming crabs of the genera Charybdis de Haan, 1833, and Thalamita Latreille, 1829 (Crustacea: Decapoda: Brachyura: Portunidae) from Peninsular Malaysia and Singapore. Raffles Bulletin of Zoology, 1 (Supplement), 1 - 128." type="journal article" year="1995">Wee &amp; Ng 1995</bibRefCitation>
: 32
</treatmentCitation>
34, fig. 14ag;
<treatmentCitation id="0AF010BEC5000E5EFCE3FCC45BA3FC98" author="Apel, M. &amp; Spiridonov, V. A." box="[857,1161,868,890]" page="194" pageId="3" pageNumber="204" year="1998">
<bibRefCitation id="EFC04B5EC5000E5EFCE3FCC45B72FC98" author="Apel, M. &amp; Spiridonov, V. A." box="[857,1112,868,890]" pageId="3" pageNumber="204" pagination="159 - 331" refId="ref11122" refString="Apel, M. &amp; Spiridonov, V. A. (1998) Taxonomy and zoogeography of the portunid crabs (Crustacea: Decapoda: Brachyura: Portunidae) of the Arabian Gulf and adjacent waters. Fauna of Arabia, 17, 159 - 331." type="journal article" year="1998">Apel &amp; Spiridonov 1998</bibRefCitation>
: 194
</treatmentCitation>
197 (synonymy), figs 13 15, 17;
<treatmentCitation id="0AF010BEC5000E5EFEAEFC225E97FC7A" author="Davie, P. J. F." box="[276,445,898,920]" page="473" pageId="3" pageNumber="204" year="2002">
<bibRefCitation id="EFC04B5EC5000E5EFEAEFC225EA4FC7A" author="Davie, P. J. F." box="[276,398,898,920]" pageId="3" pageNumber="204" pagination="1 - 641" refId="ref12763" refString="Davie, P. J. F. (2002) Crustacea: Malacostraca Eucarida (Part 2). Decapoda-Anomura, Brachyura. In: Wells, A. &amp; Houston, W. W. K. (Eds.), Zoological Catalogue of Australia. Vol. 19.3 B. CSIRO Publishing, Melbourne, pp. i-xiv + 1 - 641." type="book chapter" year="2002">Davie 2002</bibRefCitation>
: 473
</treatmentCitation>
(synonymy and list of types);
<treatmentCitation id="0AF010BEC5000E5EFD4EFC225CE4FC7A" author="Galil, B. &amp; Froglia, C. &amp; Noel, P." box="[756,974,898,920]" page="110" pageId="3" pageNumber="204" year="2002">
<bibRefCitation id="EFC04B5EC5000E5EFD4EFC225CB4FC7A" author="Galil, B. &amp; Froglia, C. &amp; Noel, P." box="[756,926,898,920]" pageId="3" pageNumber="204" refId="ref14584" refString="Galil, B., Froglia, C. &amp; Noel, P. (2002) CIESM Atlas of Exotic Species in the Mediterranean Vol. 2. Crustaceans: Decapods and Stomatopods. CIESM Publishers, Monte Carlo, 192 pp." type="book" year="2002">
Galil
<emphasis id="B925EABDC5000E5EFC97FC235C70FC7A" box="[813,858,898,920]" italics="true" pageId="3" pageNumber="204">et al</emphasis>
. 2002
</bibRefCitation>
: 110
</treatmentCitation>
, unnumbered photograph;
<treatmentCitation id="0AF010BEC5000E5EFB65FC225FC7FC55" author="Padate, V. P. &amp; Rivonker, C. U. &amp; Anil, A. C. &amp; Krishnamurthy, V." page="587" pageId="3" pageNumber="204" year="2006">
<bibRefCitation id="EFC04B5EC5000E5EFB65FC225AB2FC7A" author="Padate, V. P. &amp; Rivonker, C. U. &amp; Anil, A. C. &amp; Krishnamurthy, V." box="[1247,1432,898,920]" pageId="3" pageNumber="204" pagination="579 - 590" refId="ref18028" refString="Padate, V. P., Rivonker, C. U., Anil, A. C. &amp; Krishnamurthy, V. (2006) A new species of portunid crab of the genus Charybdis (De Haan, 1833) (Crustacea: Decapoda: Brachyura) from Goa, India. Marine Biology Research, 6, 579 - 590. https: // doi. org / 10.1080 / 17451000903505608" type="journal article" year="2006">
Padate
<emphasis id="B925EABDC5000E5EFA93FC235A76FC7A" box="[1321,1372,898,920]" italics="true" pageId="3" pageNumber="204">et al.</emphasis>
2006
</bibRefCitation>
: 587
</treatmentCitation>
(key);
<treatmentCitation id="0AF010BEC5000E5EFE8DFC015D11FC55" ID-DOI="http://doi.org/10.5281/zenodo.5469518" ID-Zenodo-Dep="5469518" author="Yokes, B. &amp; Galil, B. S." box="[311,571,929,951]" httpUri="http://treatment.plazi.org/id/03BF7F58FFF4FF8861940BE40CB8CD79" page="754" pageId="3" pageNumber="204" year="2006">
<bibRefCitation id="EFC04B5EC5000E5EFE8DFC015D2DFC55" author="Yokes, B. &amp; Galil, B. S." box="[311,519,929,951]" pageId="3" pageNumber="204" pagination="747 - 755" refId="ref19856" refString="Yokes, B. &amp; Galil, B. S. (2006) New records of alien decapods (Crustacea) from the Mediterranean coast of Turkey, with a description of a new palaemonid species. Zoosystema, 28 (3), 747 - 755." type="journal article" year="2006">Yokes &amp; Galil 2006</bibRefCitation>
: 754
</treatmentCitation>
, fig. 2F;
<treatmentCitation id="0AF010BEC5000E5EFD22FC015CAAFC55" author="Yokes, M. B. &amp; Karhan, S. U. &amp; Okus, E. &amp; Yusek, A. &amp; Aslan-Yilmaz, A. &amp; Yilmaz, I. N. &amp; Demirel, N. &amp; Demir, V. &amp; Galil, B. S." box="[664,896,929,951]" page="165" pageId="3" pageNumber="204" year="2007">
<bibRefCitation id="EFC04B5EC5000E5EFD22FC015C64FC55" author="Yokes, M. B. &amp; Karhan, S. U. &amp; Okus, E. &amp; Yusek, A. &amp; Aslan-Yilmaz, A. &amp; Yilmaz, I. N. &amp; Demirel, N. &amp; Demir, V. &amp; Galil, B. S." box="[664,846,929,951]" pageId="3" pageNumber="204" pagination="162 - 168" refId="ref19905" refString="Yokes, M. B., Karhan, S. U., Okus, E., Yusek, A., Aslan-Yilmaz, A., Yilmaz, I. N., Demirel, N., Demir, V. &amp; Galil, B. S. (2007) Alien Crustacean Decapods from the Aegean Coast of Turkey. Aquatic Invasions, 2 (3), 162 - 168. https: // doi. org / 10.3391 / ai. 2007.2.3.2" type="journal article" year="2007">
Yokes
<emphasis id="B925EABDC5000E5EFD67FC025C3BFC55" box="[733,785,929,951]" italics="true" pageId="3" pageNumber="204">et al.</emphasis>
2007
</bibRefCitation>
: 165
</treatmentCitation>
, fig. 2d;
<treatmentCitation id="0AF010BEC5000E5EFC67FC025B8EFC55" author="Ng, P. K. L. &amp; Guinot, D. &amp; Davie, P. J. F." box="[989,1188,929,951]" page="153" pageId="3" pageNumber="204" year="2008">
<bibRefCitation id="EFC04B5EC5000E5EFC67FC025B59FC55" author="Ng, P. K. L. &amp; Guinot, D. &amp; Davie, P. J. F." box="[989,1139,929,951]" pageId="3" pageNumber="204" pagination="1 - 286" refId="ref17354" refString="Ng, P. K. L., Guinot, D. &amp; Davie, P. J. F. (2008) Systema Brachyurorum: part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, 17 (Supplement), 1 - 286." type="journal article" year="2008">
Ng
<emphasis id="B925EABDC5000E5EFBB9FC025B1AFC55" box="[1027,1072,929,951]" italics="true" pageId="3" pageNumber="204">et al</emphasis>
. 2008
</bibRefCitation>
: 153
</treatmentCitation>
(list);
<collectingRegion id="4995F84DC5000E5EFB53FC025A3AFC55" box="[1257,1296,930,951]" country="Turkey" name="Van" pageId="3" pageNumber="204">Van</collectingRegion>
Chung 2012: 168 (key), 174, fig. 12;
<treatmentCitation id="0AF010BEC5000E5EFE0DFC605DAEFC34" author="Naderloo, R." box="[439,644,960,982]" page="174" pageId="3" pageNumber="204" year="2017">
<bibRefCitation id="EFC04B5EC5000E5EFE0DFC605D78FC34" author="Naderloo, R." box="[439,594,960,982]" pageId="3" pageNumber="204" refId="ref17022" refString="Naderloo, R. (2017) Atlas of the crabs of the Persian Gulf. Springer International Publishing AG, Basel, 444 pp. https: // doi. org / 10.1007 / 978 - 3 - 319 - 49374 - 9" type="book" year="2017">Naderloo 2017</bibRefCitation>
: 174
</treatmentCitation>
, figs 20.3d, 20.5, 20.7;
<treatmentCitation id="0AF010BEC5000E5EFCC9FC605B7FFC34" author="Trivedi, J. N. &amp; Trivedi, D. J. &amp; Vachhrajani, K. D. &amp; Ng, P. K. L." box="[883,1109,960,982]" page="64" pageId="3" pageNumber="204" year="2018">
<bibRefCitation id="EFC04B5EC5000E5EFCC9FC605B18FC34" author="Trivedi, J. N. &amp; Trivedi, D. J. &amp; Vachhrajani, K. D. &amp; Ng, P. K. L." box="[883,1074,960,982]" pageId="3" pageNumber="204" pagination="1 - 83" refId="ref19538" refString="Trivedi, J. N., Trivedi, D. J., Vachhrajani, K. D. &amp; Ng, P. K. L. (2018) An annotated checklist of the marine brachyuran crabs (Crustacea: Decapoda: Brachyura) of India. Zootaxa, 4502 (1), 1 - 83. https: // doi. org / 10.11646 / zootaxa. 4502.1.1" type="journal article" year="2018">
Trivedi
<emphasis id="B925EABDC5000E5EFC79FC615CDCFC34" box="[963,1014,960,982]" italics="true" pageId="3" pageNumber="204">et al.</emphasis>
2018
</bibRefCitation>
: 64
</treatmentCitation>
(list of Indian references).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BEE36AFC5000E5EFF2DFC7E5C4EFBF1" blockId="3.[151,1437,500,1933]" pageId="3" pageNumber="204">
<treatmentCitationGroup id="AB411181C5000E5EFF2DFC7E5C4EFBF1" pageId="3" pageNumber="204">
<taxonomicName id="4C514D2CC5000E5EFF2DFC7E5EABFC17" ID-CoL="TPV8" baseAuthorityName="De Man" baseAuthorityYear="1887" box="[151,385,990,1013]" class="Malacostraca" family="Portunidae" genus="Charybdis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="merguiensis">
<emphasis id="B925EABDC5000E5EFF2DFC7E5EABFC17" box="[151,385,990,1013]" italics="true" pageId="3" pageNumber="204">Charybdis merguiensis</emphasis>
</taxonomicName>
.—
<treatmentCitation id="0AF010BEC5000E5EFE25FC7E5D49FC16" author="Bouvier, E. - L." box="[415,611,990,1012]" page="252" pageId="3" pageNumber="204" year="1940">
<bibRefCitation id="EFC04B5EC5000E5EFE25FC7E5D1AFC16" author="Bouvier, E. - L." box="[415,560,990,1012]" pageId="3" pageNumber="204" pagination="1 - 404" refId="ref11553" refString="Bouvier, E. - L. (1940) Decapodes marcheurs. In: Faune de France. Vol. 37. Paul Lechevalier, Paris, pp. 1 - 404, pls. 1 - 14." type="book chapter" year="1940">Bouvier 1940</bibRefCitation>
: 252
</treatmentCitation>
, fig. 162;
<treatmentCitation id="0AF010BEC5000E5EFD76FC7E5C70FC16" author="Ward, M." box="[716,858,990,1012]" page="5" pageId="3" pageNumber="204" year="1941">
<bibRefCitation id="EFC04B5EC5000E5EFD76FC7E5C6AFC16" author="Ward, M." box="[716,832,990,1012]" pageId="3" pageNumber="204" pagination="1 - 15" refId="ref19686" refString="Ward, M. (1941) New Brachyura from the Gulf of Davao, Mindanao, Philippine Islands. American Museum Novitates, 1104, 1 - 15." type="journal article" year="1941">Ward 1941</bibRefCitation>
: 5
</treatmentCitation>
;
<treatmentCitation id="0AF010BEC5000E5EFCDCFC7E5B02FC16" author="Barnard, K. H." box="[870,1064,990,1012]" page="168" pageId="3" pageNumber="204" year="1950">
<bibRefCitation id="EFC04B5EC5000E5EFCDCFC7E5CDFFC16" author="Barnard, K. H." box="[870,1013,990,1012]" pageId="3" pageNumber="204" pagination="1 - 837" refId="ref11170" refString="Barnard, K. H. (1950) Descriptive catalogue of South African decapod Crustacea (crabs and shrimps). Annals of the South African Museum, 38, 1 - 837." type="journal article" year="1950">Barnard 1950</bibRefCitation>
: 168
</treatmentCitation>
, figs 27d, 32b;
<treatmentCitation id="0AF010BEC5000E5EFB73FC7E5ABDFC16" author="Fourmanoir, P." box="[1225,1431,990,1012]" page="8" pageId="3" pageNumber="204" year="1954">
<bibRefCitation id="EFC04B5EC5000E5EFB73FC7E5A54FC16" author="Fourmanoir, P." box="[1225,1406,990,1012]" pageId="3" pageNumber="204" pagination="1 - 16" refId="ref14212" refString="Fourmanoir, P. (1954) Crabes de la cote ouest de Madagascar. Le Naturaliste Malgache, 6 (1 / 2), 1 - 16." type="journal article" year="1954">Fourmanoir 1954</bibRefCitation>
: 8
</treatmentCitation>
, fig. 8; Badhra 1995: 257 (key), 258;
<treatmentCitation id="0AF010BEC5000E5EFD80FC5D5DDAFBF1" author="Bhadra, S." box="[570,752,1021,1043]" page="410" pageId="3" pageNumber="204" year="1999">
<bibRefCitation id="EFC04B5EC5000E5EFD80FC5D5DEAFBF1" author="Bhadra, S." box="[570,704,1021,1043]" pageId="3" pageNumber="204" pagination="405 - 415" refId="ref11342" refString="Bhadra, S. (1999) Crustacea: Decapoda: Portunidae. Fauna of West Bengal, State Fauna Series, 3 (10), 405 - 415." type="journal article" year="1999">Bhadra 1999</bibRefCitation>
: 410
</treatmentCitation>
(key), 411.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BEE36AFC5000E5EFF2DFBBC5B97FBD0" blockId="3.[151,1437,500,1933]" box="[151,1213,1052,1074]" pageId="3" pageNumber="204">
<treatmentCitationGroup id="AB411181C5000E5EFF2DFBBC5B97FBD0" box="[151,1213,1052,1074]" pageId="3" pageNumber="204">
<taxonomicName id="4C514D2CC5000E5EFF2DFBBC5D2AFBD0" ID-CoL="84GRH" authority="Ward, 1941: 4" authorityName="Ward" authorityPageNumber="4" authorityYear="1941" box="[151,512,1052,1074]" class="Malacostraca" family="Portunidae" genus="Charybdis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="vannamei">
<emphasis id="B925EABDC5000E5EFF2DFBBC5E43FBD0" box="[151,361,1052,1074]" italics="true" pageId="3" pageNumber="204">Charybdis vannamei</emphasis>
<treatmentCitation id="0AF010BEC5000E5EFED5FBBC5D2AFBD0" author="Ward, M." box="[367,512,1052,1074]" page="4" pageId="3" pageNumber="204" year="1941">
<bibRefCitation id="EFC04B5EC5000E5EFED5FBBC5EC2FBD0" author="Ward, M." box="[367,488,1052,1074]" pageId="3" pageNumber="204" pagination="1 - 15" refId="ref19686" refString="Ward, M. (1941) New Brachyura from the Gulf of Davao, Mindanao, Philippine Islands. American Museum Novitates, 1104, 1 - 15." type="journal article" year="1941">Ward, 1941</bibRefCitation>
: 4
</treatmentCitation>
</taxonomicName>
, figs 5, 6;
<treatmentCitation id="0AF010BEC5000E5EFDD1FBBC5C61FBD0" author="Stephenson, W." box="[619,843,1052,1074]" page="32" pageId="3" pageNumber="204" year="1972">
<bibRefCitation id="EFC04B5EC5000E5EFDD1FBBC5C02FBD0" author="Stephenson, W." box="[619,808,1052,1074]" pageId="3" pageNumber="204" pagination="1 - 64" refId="ref18960" refString="Stephenson, W. (1972 b) An annotated check list and key to the Indo-West Pacific swimming crabs (Crustacea: Decapoda: Portunidae). Bulletin of the Royal Society of New Zealand, 10, 1 - 64." type="journal article" year="1972">Stephenson 1972b</bibRefCitation>
: 32
</treatmentCitation>
(discussion);
<treatmentCitation id="0AF010BEC5000E5EFC60FBBC5B93FBD0" author="Santhanam, R." box="[986,1209,1052,1074]" page="160" pageId="3" pageNumber="204" year="2018">
<bibRefCitation id="EFC04B5EC5000E5EFC60FBBC5BADFBD0" author="Santhanam, R." box="[986,1159,1052,1074]" pageId="3" pageNumber="204" refId="ref18734" refString="Santhanam, R. (2018) Biology and Culture of Portunid Crabs of World Seas. Series: Biology and Ecology of Marine Life. Apple Academic Press, Waretown, NJ, 404 pp. https: // doi. org / 10.1201 / 9781315205816" type="book" year="2018">Santhanam 2018</bibRefCitation>
: 160
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BEE36AFC5000E5EFF2DFB9A5C43FBB2" blockId="3.[151,1437,500,1933]" box="[151,873,1082,1104]" pageId="3" pageNumber="204">
<treatmentCitationGroup id="AB411181C5000E5EFF2DFB9A5C43FBB2" box="[151,873,1082,1104]" pageId="3" pageNumber="204">
<taxonomicName id="4C514D2CC5000E5EFF2DFB9A5E59FBB2" ID-CoL="3GXMW" box="[151,371,1082,1104]" class="Malacostraca" family="Portunidae" genus="Charybdis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="spiniferum">
<emphasis id="B925EABDC5000E5EFF2DFB9A5E59FBB2" box="[151,371,1082,1104]" italics="true" pageId="3" pageNumber="204">Charybdis spiniferum</emphasis>
</taxonomicName>
.—
<treatmentCitation id="0AF010BEC5000E5EFE2AFB9A5D34FBB2" author="Ward, M." box="[400,542,1082,1104]" page="5" pageId="3" pageNumber="204" year="1941">
<bibRefCitation id="EFC04B5EC5000E5EFE2AFB9A5D2FFBB2" author="Ward, M." box="[400,517,1082,1104]" pageId="3" pageNumber="204" pagination="1 - 15" refId="ref19686" refString="Ward, M. (1941) New Brachyura from the Gulf of Davao, Mindanao, Philippine Islands. American Museum Novitates, 1104, 1 - 15." type="journal article" year="1941">Ward 1941</bibRefCitation>
: 5
</treatmentCitation>
;
<treatmentCitation id="0AF010BEC5000E5EFD93FB9B5DCAFBB2" author="Ng, P. K. L. &amp; Wang, C. - H. &amp; Ho, P. - H. &amp; Shih, H. - T." box="[553,736,1082,1104]" page="20" pageId="3" pageNumber="204" year="2001">
<bibRefCitation id="EFC04B5EC5000E5EFD93FB9B5D97FBB2" author="Ng, P. K. L. &amp; Wang, C. - H. &amp; Ho, P. - H. &amp; Shih, H. - T." box="[553,701,1082,1104]" pageId="3" pageNumber="204" pagination="1 - 86" refId="ref17478" refString="Ng, P. K. L., Wang, C. - H., Ho, P. - H. &amp; Shih, H. - T. (2001) An annotated checklist of brachyuran crabs from Taiwan (Crustacea: Decapoda). National Taiwan Museum Special Publication Series, 11, 1 - 86." type="journal article" year="2001">
Ng
<emphasis id="B925EABDC5000E5EFDF4FB9B5DABFBB2" box="[590,641,1082,1104]" italics="true" pageId="3" pageNumber="204">et al.</emphasis>
2001
</bibRefCitation>
: 20
</treatmentCitation>
(discussion).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BEE36AFC5000E5EFF2DFBF95BB6FB6C" blockId="3.[151,1437,500,1933]" pageId="3" pageNumber="204">
<treatmentCitationGroup id="AB411181C5000E5EFF2DFBF95BB6FB6C" pageId="3" pageNumber="204">
<taxonomicName id="4C514D2CC5000E5EFF2DFBF95EE1FB8D" box="[151,459,1113,1135]" class="Malacostraca" family="Portunidae" genus="Charybdis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="helleri" subGenus="Charybdis">
<emphasis id="B925EABDC5000E5EFF2DFBF95E2BFB8D" box="[151,257,1113,1135]" italics="true" pageId="3" pageNumber="204">Charybdis</emphasis>
(
<emphasis id="B925EABDC5000E5EFEAAFBF95E50FB8D" box="[272,378,1113,1135]" italics="true" pageId="3" pageNumber="204">Charybdis</emphasis>
)
<emphasis id="B925EABDC5000E5EFE33FBF95EE1FB8D" box="[393,459,1113,1135]" italics="true" pageId="3" pageNumber="204">helleri</emphasis>
</taxonomicName>
.—
<treatmentCitation id="0AF010BEC5000E5EFE52FBF95DECFB8D" author="Buitendijk, A. M." box="[488,710,1113,1135]" page="281" pageId="3" pageNumber="204" year="1947">
<bibRefCitation id="EFC04B5EC5000E5EFE52FBF95DB9FB8D" author="Buitendijk, A. M." box="[488,659,1113,1135]" pageId="3" pageNumber="204" pagination="280 - 284" refId="ref11799" refString="Buitendijk, A. M. (1947) Zoological notes from Port Dickson, III. Crustacea Anomura and Brachyura. Zoologische Mededelingen, Leiden, 28, 280 - 284." type="journal article" year="1947">Buitendijk 1947</bibRefCitation>
: 281
</treatmentCitation>
;
<treatmentCitation id="0AF010BEC5000E5EFD68FBF95CC4FB8D" author="Stephenson, W. &amp; Hudson, J. J. &amp; Campbell, B." box="[722,1006,1113,1135]" page="492" pageId="3" pageNumber="204" year="1957">
<bibRefCitation id="EFC04B5EC5000E5EFD68FBF95C94FB8D" author="Stephenson, W. &amp; Hudson, J. J. &amp; Campbell, B." box="[722,958,1113,1135]" pageId="3" pageNumber="204" pagination="491 - 507" refId="ref19047" refString="Stephenson, W., Hudson, J. J. &amp; Campbell, B. (1957) The Australian Portunids (Crustacea; Portunidae). II. The genus Charybdis. Australian Journal of Marine and Freshwater Research, 8 (4), 491 - 507. https: // doi. org / 10.1071 / MF 9570491" type="journal article" year="1957">
Stephenson
<emphasis id="B925EABDC5000E5EFCF7FBFA5CABFB8D" box="[845,897,1113,1135]" italics="true" pageId="3" pageNumber="204">et al.</emphasis>
1957
</bibRefCitation>
: 492
</treatmentCitation>
(key), 493 (key), 497498, figs 1a,
<date id="FFEF106FC5000E5EFADBFBF95A5DFB8D" box="[1377,1399,1113,1135]" pageId="3" pageNumber="204">2i</date>
, 3j, pl. 1, fig. 4, pls 4c, 5b;
<treatmentCitation id="0AF010BEC5000E5EFE08FBD85D42FB6C" author="Crosnier, A." box="[434,616,1144,1166]" page="75" pageId="3" pageNumber="204" year="1962">
<bibRefCitation id="EFC04B5EC5000E5EFE08FBD85D6FFB6C" author="Crosnier, A." box="[434,581,1144,1166]" pageId="3" pageNumber="204" pagination="1 - 154" refId="ref12372" refString="Crosnier, A. (1962) Crustaces Decapodes Portunidae. In: Faune de Madagascar. Vol. 16. ORSTOM, CNRS, Paris, pp. 1 - 154." type="book chapter" year="1962">Crosnier 1962</bibRefCitation>
: 75
</treatmentCitation>
(key), 7778, figs 133135, pl. 5 fig. 1;
<treatmentCitation id="0AF010BEC5000E5EFBBEFBD85BB2FB6C" author="Apel, M." box="[1028,1176,1144,1166]" page="70" pageId="3" pageNumber="204" year="2001">
<bibRefCitation id="EFC04B5EC5000E5EFBBEFBD85B58FB6C" author="Apel, M." box="[1028,1138,1144,1166]" pageId="3" pageNumber="204" refId="ref11073" refString="Apel, M. (2001) Taxonomie und Zoogeographie der Brachyura, Paguridea und Porcellanidae (Crustacea: Decapoda) des Persisch-Arabischen Golfes. Dissertation zur Erlangung des Doktorgrades der Naturwissenschaften, vorgelegt beim Fachbereich Biologie der Johann Wolfgang Goethe-Universitat in Frankfurt am Main, 255 pp." type="book" year="2001">Apel 2001</bibRefCitation>
: 70
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BEE36AFC5000E5EFF2DFB365CB2F99A" blockId="3.[151,1437,500,1933]" pageId="3" pageNumber="204">
<treatmentCitationGroup id="AB411181C5000E5EFF2DFB365CB2F99A" pageId="3" pageNumber="204">
<taxonomicName id="4C514D2CC5000E5EFF2DFB365E7AFB4E" ID-CoL="TPTB" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1867" box="[151,336,1174,1196]" class="Malacostraca" family="Portunidae" genus="Charybdis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="hellerii">
<emphasis id="B925EABDC5000E5EFF2DFB365E7AFB4E" box="[151,336,1174,1196]" italics="true" pageId="3" pageNumber="204">Charybdis hellerii</emphasis>
</taxonomicName>
.—
<treatmentCitation id="0AF010BEC5000E5EFED4FB365D72FB4E" author="Edmondson, C. H." box="[366,600,1174,1196]" page="247" pageId="3" pageNumber="204" year="1954">
<bibRefCitation id="EFC04B5EC5000E5EFED4FB365D0CFB4E" author="Edmondson, C. H." box="[366,550,1174,1196]" pageId="3" pageNumber="204" pagination="217 - 274" refId="ref13723" refString="Edmondson, C. H. (1954) Hawaiian Portunidae. Occasional Papers of Bernice P. Bishop Museum, 21 (12), 217 - 274." type="journal article" year="1954">Edmondson 1954</bibRefCitation>
: 247
</treatmentCitation>
248, fig. 32af (Hawai'i);
<treatmentCitation id="0AF010BEC5000E5EFCD7FB365BB2FB4E" author="Campos, N. H. &amp; Turkay, M." box="[877,1176,1174,1196]" page="119" pageId="3" pageNumber="204" year="1989">
<bibRefCitation id="EFC04B5EC5000E5EFCD7FB365B42FB4E" author="Campos, N. H. &amp; Turkay, M." box="[877,1128,1174,1196]" pageId="3" pageNumber="204" pagination="119 - 123" refId="ref11832" refString="Campos, N. H. &amp; Turkay, M. (1989) On a record of Charybdis hellerii from the Caribbean Coast of Colombia (Crustacea: Decapoda: Portunidae) Senckenbergiana maritima, 20 (3 / 4), 119 - 123." type="journal article" year="1989">Campos &amp; Türkay 1989</bibRefCitation>
: 119
</treatmentCitation>
(first published record in the Western Atlantic);
<treatmentCitation id="0AF010BEC5000E5EFE10FB155D5AFB29" author="Lemaitre, R." box="[426,624,1205,1227]" page="643" pageId="3" pageNumber="204" year="1995">
<bibRefCitation id="EFC04B5EC5000E5EFE10FB155D68FB29" author="Lemaitre, R." box="[426,578,1205,1227]" pageId="3" pageNumber="204" pagination="643 - 648" refId="ref15864" refString="Lemaitre, R. (1995) Charybdis hellerii (Milne Edwards, 1867), a nonindigenous portunid crab (Crustacea: Decapoda: Brachyura) discovered in the Indian River lagoon system of Florida. Proceedings of the Biological Society of Washington, 108 (4), 643 - 648." type="journal article" year="1995">Lemaitre 1995</bibRefCitation>
: 643
</treatmentCitation>
647, fig. 2;
<treatmentCitation id="0AF010BEC5000E5EFD4BFB155CD7FB29" author="Tirmizi, N. M. &amp; Kazmi, Q. B." box="[753,1021,1205,1227]" page="29" pageId="3" pageNumber="204" year="1996">
<bibRefCitation id="EFC04B5EC5000E5EFD4BFB155CF1FB29" author="Tirmizi, N. M. &amp; Kazmi, Q. B." box="[753,987,1205,1227]" pageId="3" pageNumber="204" refId="ref19406" refString="Tirmizi, N. M. &amp; Kazmi, Q. B. (1996) Marine Fauna of Pakistan. 6. Crustacea: Brachyura, Brachyrhyncha. Part 2. Portunidae. Marine Reference Collection Center, University of Karachi, Karachi, 72 pp., 2 pls." type="book" year="1996">Tirmizi &amp; Kazmi 1996</bibRefCitation>
: 29
</treatmentCitation>
(key), 4547 (possibly in part), pl. 1, fig. B, possibly not fig. 23ai; Tavares &amp; Braga de
<treatmentCitation id="0AF010BEC5000E5EFD1AFB745C53FB08" author="Tavares, M. &amp; Braga de Mendonca, J. Jr." box="[672,889,1236,1258]" page="151" pageId="3" pageNumber="204" year="1996">
<bibRefCitation id="EFC04B5EC5000E5EFD1AFB745C6DFB08" author="Tavares, M. &amp; Braga de Mendonca, J. Jr." box="[672,839,1236,1258]" pageId="3" pageNumber="204" pagination="151 - 157" refId="ref19268" refString="Tavares, M. &amp; Braga de Mendonca, J. Jr. (1996) Charybdis hellerii (A. Milne Edwards, 1867) (Brachyura: Portunidae) eighth nonindigenous marine decapod recorded from Brazil. Crustacean Research, 25, 151 - 157. https: // doi. org / 10.18353 / crustacea. 25.0 _ 151" type="journal article" year="1996">Mendonça 1996</bibRefCitation>
: 151
</treatmentCitation>
, 154 (key);
<treatmentCitation id="0AF010BEC5000E5EFC48FB755BCCFB08" author="Fransen, C. H. J. M. &amp; Holthuis, L. B. &amp; Adema, J. P. H. M." box="[1010,1254,1236,1258]" page="179" pageId="3" pageNumber="204" year="1997">
<bibRefCitation id="EFC04B5EC5000E5EFC48FB755B9CFB08" author="Fransen, C. H. J. M. &amp; Holthuis, L. B. &amp; Adema, J. P. H. M." box="[1010,1206,1236,1258]" pageId="3" pageNumber="204" pagination="1 - 344" refId="ref14385" refString="Fransen, C. H. J. M., Holthuis, L. B. &amp; Adema, J. P. H. M. (1997) Type-catalogue of the decapod Crustacea in the collections of the Nationaal Natuurhistorisch Museum, with appendices of pre- 1900 collectors and material. Zoologische Verhandelingen, Leiden, 311, i-xvi + 1 - 344." type="journal article" year="1997">
Fransen
<emphasis id="B925EABDC5000E5EFBF2FB755B51FB08" box="[1096,1147,1236,1258]" italics="true" pageId="3" pageNumber="204">et al.</emphasis>
1997
</bibRefCitation>
: 179
</treatmentCitation>
(original material of
<taxonomicName id="4C514D2CC5000E5EFF58FB525D6EFAEA" authority="Ruppell, 1830" authorityName="Ruppell" authorityYear="1830" box="[226,580,1266,1288]" class="Malacostraca" family="Portunidae" genus="Talamita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="sexdentata">
<emphasis id="B925EABDC5000E5EFF58FB525E87FAEA" box="[226,429,1266,1288]" italics="true" pageId="3" pageNumber="204">Talamita sexdentata</emphasis>
<bibRefCitation id="EFC04B5EC5000E5EFE0EFB525D6EFAEA" author="Ruppell, E." box="[436,580,1266,1288]" pageId="3" pageNumber="204" pagination="1 - 6" refId="ref18354" refString="Ruppell, E. (1830) Beschreibung und Abbildung von 24 Arten kurzschwanzigen Krabben, also Beitrag zur Naturgeschichte des Rothen Meeres. H. L. Bronner, Frankfurt am Main, 28 pp., pls 1 - 6." type="book chapter" year="1830">Rüppell, 1830</bibRefCitation>
</taxonomicName>
listed);
<treatmentCitation id="0AF010BEC5000E5EFD2CFB525CF8FAEA" author="Vannini, M. &amp; Innocenti, G." box="[662,978,1266,1288]" page="269" pageId="3" pageNumber="204" year="2000">
<bibRefCitation id="EFC04B5EC5000E5EFD2CFB525C8BFAEA" author="Vannini, M. &amp; Innocenti, G." box="[662,929,1266,1288]" pageId="3" pageNumber="204" pagination="251 - 298" refId="ref19642" refString="Vannini, M. &amp; Innocenti, G. (2000) Research on the coast of Somalia. Portunidae (Crustacea Brachyura). Tropical Zoology, 13, 251 - 298. https: // doi. org / 10.1080 / 03946975.2000.10531136" type="journal article" year="2000">Vannini &amp; Innocenti 2000</bibRefCitation>
: 269
</treatmentCitation>
, figs 30, 35, 93;
<treatmentCitation id="0AF010BEC5000E5EFBC1FB525A41FAEA" author="Dineen, J. F. &amp; Clark, P. F. &amp; Hines, A. H. &amp; Reed, S. A. &amp; Walton, H. P." box="[1147,1387,1266,1288]" page="778" pageId="3" pageNumber="204" year="2001">
<bibRefCitation id="EFC04B5EC5000E5EFBC1FB525A13FAEA" author="Dineen, J. F. &amp; Clark, P. F. &amp; Hines, A. H. &amp; Reed, S. A. &amp; Walton, H. P." box="[1147,1337,1266,1288]" pageId="3" pageNumber="204" pagination="774 - 805" refId="ref13561" refString="Dineen, J. F., Clark, P. F., Hines, A. H., Reed, S. A. &amp; Walton, H. P. (2001) Life history, larval description, and natural history of Charybdis hellerii (Decapoda, Brachyura, Portunidae), an invasive crab in the Western Atlantic. Journal of Crustacean Biology, 21 (3), 774 - 805. https: // doi. org / 10.1163 / 20021975 - 99990173" type="journal article" year="2001">
Dineen
<emphasis id="B925EABDC5000E5EFB71FB535BD4FAEA" box="[1227,1278,1266,1288]" italics="true" pageId="3" pageNumber="204">et al.</emphasis>
2001
</bibRefCitation>
: 778
</treatmentCitation>
, figs 324 (larval and juvenile stages);
<treatmentCitation id="0AF010BEC5000E5EFDA5FAB25C41FAC5" author="Tavares, M. &amp; Amouroux, J. M." box="[543,875,1297,1319]" page="625" pageId="3" pageNumber="204" year="2003">
<bibRefCitation id="EFC04B5EC5000E5EFDA5FAB25C13FAC5" author="Tavares, M. &amp; Amouroux, J. M." box="[543,825,1297,1319]" pageId="3" pageNumber="204" pagination="625 - 630" refId="ref19210" refString="Tavares, M. &amp; Amouroux, J. M. (2003) First record of the non-indigenous crab Charybdis hellerii (A. Milne-Edwards, 1867) from French Guyana (Decapoda, Brachyura, Portunidae). Crustaceana, 76, 625 - 630. https: // doi. org / 10.1163 / 156854003322316254" type="journal article" year="2003">Tavares &amp; Amouroux 2003</bibRefCitation>
: 625
</treatmentCitation>
, fig. 1; Alves Coelho &amp; do
<treatmentCitation id="0AF010BEC5000E5EFB3EFAB15FC5FAA4" author="Alves Coelho, P. &amp; do Carmo Ferrao Santos, M." page="167" pageId="3" pageNumber="204" year="2003">
<bibRefCitation id="EFC04B5EC5000E5EFB3EFAB15AB2FAC5" author="Alves Coelho, P. &amp; do Carmo Ferrao Santos, M." box="[1156,1432,1297,1319]" pageId="3" pageNumber="204" pagination="167 - 173" refId="ref10969" refString="Alves Coelho, P. &amp; do Carmo Ferrao Santos, M. (2003) Ocorrencia de Charybdis hellerii (Milne Edwards, 1867) (Crustacea, Decapoda, Portunidae) no litoral de Pernambuco. Boletim Tecnico-Cientifico do CEPENE, 11 (1), 167 - 173." type="journal article" year="2003">Carmo Ferrão Santos 2003</bibRefCitation>
: 167
</treatmentCitation>
, fig. 1; 2004: 192, fig. 5;
<treatmentCitation id="0AF010BEC5000E5EFDB9FA915DCFFAA4" author="Braga, A. A. &amp; Fransozo, A. &amp; Bertini, G. &amp; Fumis, P. B." box="[515,741,1328,1350]" page="30" pageId="3" pageNumber="204" year="2005">
<bibRefCitation id="EFC04B5EC5000E5EFDB9FA915D96FAA4" author="Braga, A. A. &amp; Fransozo, A. &amp; Bertini, G. &amp; Fumis, P. B." box="[515,700,1328,1350]" pageId="3" pageNumber="204" pagination="1 - 34" refId="ref11593" refString="Braga, A. A., Fransozo, A., Bertini, G. &amp; Fumis, P. B. (2005). Composition and abundance of the crabs (Decapoda, Brachyura) off Ubatuba and Caraguatatuba, northern coast of Sao Paulo, Brazil. Biota Neotropica, 5 (2), 1 - 34." type="journal article" year="2005">
Braga
<emphasis id="B925EABDC5000E5EFDF2FA915D54FAA4" box="[584,638,1328,1350]" italics="true" pageId="3" pageNumber="204">et al.</emphasis>
2005
</bibRefCitation>
: 30
</treatmentCitation>
, fig. 30;
<treatmentCitation id="0AF010BEC5000E5EFCFCFA905B77FAA4" author="Sampaio, C. &amp; LS &amp; Rosa, I. L." box="[838,1117,1328,1350]" page="58" pageId="3" pageNumber="204" year="2005">
<bibRefCitation id="EFC04B5EC5000E5EFCFCFA905B1FFAA4" author="Sampaio, C. &amp; LS &amp; Rosa, I. L." box="[838,1077,1328,1350]" pageId="3" pageNumber="204" pagination="58" refId="ref18397" refString="Sampaio, C. LS. &amp; Rosa, I. L. (2005) Predation of an alien species of crab (Charybdis hellerii Milne Edwards) by a native Octopus species on NE Brazilian reefs. Coral Reefs, 25 (1), 58. https: // doi. org / 10.1007 / s 00338 - 005 - 0050 - 0" type="journal article" year="2005">Sampaio &amp; Rosa 2005</bibRefCitation>
: 58
</treatmentCitation>
, fig. 1 (predation by octopus);
<treatmentCitation id="0AF010BEC5000E5EFF7DFAEE5EC0FA86" author="Moran, R. &amp; Atencio, M." box="[199,490,1358,1380]" page="205" pageId="3" pageNumber="204" year="2006">
<bibRefCitation id="EFC04B5EC5000E5EFF7DFAEE5E9DFA86" author="Moran, R. &amp; Atencio, M." box="[199,439,1358,1380]" pageId="3" pageNumber="204" pagination="202 - 209" refId="ref16976" refString="Moran, R. &amp; Atencio, M. (2006) Charybdis hellerii (Crustacea: Decapoda: Portunidae), especie invasora en la Peninsula de Paraguanua, estado Falcon, Venezuela. Multiciencias, 6 (2), 202 - 209." type="journal article" year="2006">Morán &amp; Atencio 2006</bibRefCitation>
: 205
</treatmentCitation>
; Oliveira de
<treatmentCitation id="0AF010BEC5000E5EFDD6FAEE5C74FA86" author="Oliveira de Almeida, A. &amp; Alves Coelho, P. &amp; Almeida dos Santos, J. R. &amp; Ribeiro Ferraz, N." box="[620,862,1358,1381]" page="13" pageId="3" pageNumber="204" year="2006">
<bibRefCitation id="EFC04B5EC5000E5EFDD6FAEE5C10FA86" author="Oliveira de Almeida, A. &amp; Alves Coelho, P. &amp; Almeida dos Santos, J. R. &amp; Ribeiro Ferraz, N." box="[620,826,1358,1381]" pageId="3" pageNumber="204" pagination="1 - 24" refId="ref17868" refString="Oliveira de Almeida, A., Alves Coelho, P., Almeida dos Santos, J. R. &amp; Ribeiro Ferraz, N. 2006. Crustaceos decapodos estuarinos de Ilheus, Bahia, Brasil. Biota Neotropica, 6 (2), 1 - 24." type="journal article" year="2006">
Almeida
<emphasis id="B925EABDC5000E5EFD70FAF05DDCFA86" box="[714,758,1358,1381]" italics="true" pageId="3" pageNumber="204">et al</emphasis>
. 2006
</bibRefCitation>
: 13
</treatmentCitation>
, fig. 8;
<treatmentCitation id="0AF010BEC5000E5EFC11FAEE5AB2FA86" author="Figueiredo Frigotto, S. &amp; Serafim-Junior, M." box="[939,1432,1358,1380]" page="228" pageId="3" pageNumber="204" year="2007">
<bibRefCitation id="EFC04B5EC5000E5EFC11FAEE5A4CFA86" author="Figueiredo Frigotto, S. &amp; Serafim-Junior, M." box="[939,1382,1358,1380]" pageId="3" pageNumber="204" pagination="227 - 230" refId="ref14150" refString="Figueiredo Frigotto, S. &amp; Serafim-Junior, M. (2007) Primeiro registro de Charybdis hellerii (Milne Edwards, 1867) (Crustacea) no litoral do Estado do Parana. Estudos de Biologia, 29 (7), 227 - 230. https: // doi. org / 10.7213 / reb. v 29 i 67.22771" type="journal article" year="2007">Figueiredo Frigotto &amp; Serafim-Junior 2007</bibRefCitation>
: 228
</treatmentCitation>
, fig. 1;
<treatmentCitation id="0AF010BEC5000E5EFEB5FACD5D75FA61" author="Brockerhoff, A. &amp; McLay, C. L." box="[271,607,1389,1411]" page="71" pageId="3" pageNumber="204" year="2008">
<bibRefCitation id="EFC04B5EC5000E5EFEB5FACD5D10FA61" author="Brockerhoff, A. &amp; McLay, C. L." box="[271,570,1389,1411]" pageId="3" pageNumber="204" pagination="1 - 107" refId="ref11656" refString="Brockerhoff, A. &amp; McLay, C. L. (2008) No frontiers in the sea for marine invaders and their parasites? Biosecurity New Zealand Technical Paper, 2008 / 10, 1 - 107." type="journal article" year="2008">Brockerhoff &amp; McLay 2008</bibRefCitation>
: 71
</treatmentCitation>
(extensive data compilation);
<treatmentCitation id="0AF010BEC5000E5EFC25FACD5BFBFA61" author="McMillen-Jackson, A. L." box="[927,1233,1389,1411]" page="889" pageId="3" pageNumber="204" year="2008">
<bibRefCitation id="EFC04B5EC5000E5EFC25FACD5BB6FA61" author="McMillen-Jackson, A. L." box="[927,1180,1389,1411]" pageId="3" pageNumber="204" pagination="889 - 894" refId="ref16496" refString="McMillen-Jackson, A. L. (2008) First record of the Indo-pacific swimming crab, Charybdis hellerii (A. Milne-Edwards, 1867) in the Gulf of Mexico. Crustaceana, 81 (7), 889 - 894. https: // doi. org / 10.1163 / 156854008784771720" type="journal article" year="2008">McMillen-Jackson 2008</bibRefCitation>
: 889
</treatmentCitation>
, fig. 2;
<treatmentCitation id="0AF010BEC5000E5EFA92FACD5E08FA40" author="Lima, T. B. Jr. &amp; da Costa Aragao, M. I. &amp; Pereira da Silva, J. &amp; Melo, G. A. S. &amp; Leite, J. R. S. A." page="37" pageId="3" pageNumber="204" year="2008">
<bibRefCitation id="EFC04B5EC5000E5EFA92FACD5FD7FA40" author="Lima, T. B. Jr. &amp; da Costa Aragao, M. I. &amp; Pereira da Silva, J. &amp; Melo, G. A. S. &amp; Leite, J. R. S. A." pageId="3" pageNumber="204" pagination="25 - 40" refId="ref16038" refString="Lima, T. B. Jr., da Costa Aragao, M. I., Pereira da Silva, J., Melo, G. A. S. &amp; Leite, J. R. S. A. (2008) occurence of two Indo-pacific species of Brachyura on the coast of Piaui, Brazil. Boletim do Laboratorio de Hidrobiologia, 21, 25 - 40." type="journal article" year="2008">
Lima
<emphasis id="B925EABDC5000E5EFADCFACE5AB7FA61" box="[1382,1437,1389,1411]" italics="true" pageId="3" pageNumber="204">et al.</emphasis>
2008
</bibRefCitation>
: 37
</treatmentCitation>
, fig. 3;
<treatmentCitation id="0AF010BEC5000E5EFED5FA2C5D7DFA40" author="Felder, D. L. &amp; Dworschak, P. C. &amp; Robles, R. &amp; Bracken, H. D. &amp; Windsor, A. M. &amp; Felder, J. M. &amp; Lemaitre, R." box="[367,599,1420,1442]" page="183" pageId="3" pageNumber="204" year="2009">
<bibRefCitation id="EFC04B5EC5000E5EFED5FA2C5D0CFA40" author="Felder, D. L. &amp; Dworschak, P. C. &amp; Robles, R. &amp; Bracken, H. D. &amp; Windsor, A. M. &amp; Felder, J. M. &amp; Lemaitre, R." box="[367,550,1420,1442]" pageId="3" pageNumber="204" pagination="181 - 188" refId="ref13997" refString="Felder, D. L., Dworschak, P. C., Robles, R., Bracken, H. D., Windsor, A. M., Felder, J. M., Lemaitre, R. (2009) Obvious invaders and overlooked infauna: unexpected constituents of the decapod crustacean fauna at Twin Cays, Belize. Smithsonian Contributions to Marine Sciences, 38, 181 - 188." type="journal article" year="2009">
Felder
<emphasis id="B925EABDC5000E5EFE0CFA2D5EC3FA40" box="[438,489,1420,1442]" italics="true" pageId="3" pageNumber="204">et al.</emphasis>
2009
</bibRefCitation>
: 183
</treatmentCitation>
, fig. 1b;
<treatmentCitation id="0AF010BEC5000E5EFD0BFA2C5CFEFA40" author="Mizzan, L. &amp; Vianello, C." box="[689,980,1420,1442]" page="29" pageId="3" pageNumber="204" year="2009">
<bibRefCitation id="EFC04B5EC5000E5EFD0BFA2C5C85FA40" author="Mizzan, L. &amp; Vianello, C." box="[689,943,1420,1442]" pageId="3" pageNumber="204" pagination="27 - 30" refId="ref16859" refString="Mizzan, L. &amp; Vianello, C. (2009) First record of Charybdis (Charybdis) lucifera (Fabricius, 1798) (Crustacea, Decapoda, Portunidae) in the Mediterranean Sea. Bolletino del Museo civico di Storia naturale di Venezia, 59, 27 - 30." type="journal article" year="2009">Mizzan &amp; Vianello 2009</bibRefCitation>
: 29
</treatmentCitation>
, fig. 6;
<treatmentCitation id="0AF010BEC5000E5EFB9BFA2D5BCBFA40" author="Boos, H. Jr. &amp; Mattar de Oliveira, M. &amp; Delfim, R." box="[1057,1249,1420,1442]" page="1" pageId="3" pageNumber="204" year="2010">
<bibRefCitation id="EFC04B5EC5000E5EFB9BFA2D5BE1FA40" author="Boos, H. Jr. &amp; Mattar de Oliveira, M. &amp; Delfim, R." box="[1057,1227,1420,1442]" pageId="3" pageNumber="204" pagination="1 - 7" refId="ref11455" refString="Boos, H. Jr., Mattar de Oliveira, M. &amp; Delfim, R. (2010) Novos registros do siri exotico Charybdis hellerii (A. Milne-Edwards, 1867) (Crustacea, Portunidae), no litoral do Estado de Santa Catarina, Brasil. Revista CEPSUL - Biodiversidade e Conservacao Marinha, 1 (1), 1 - 7." type="journal article" year="2010">
Boos
<emphasis id="B925EABDC5000E5EFBE1FA2D5BADFA40" box="[1115,1159,1420,1442]" italics="true" pageId="3" pageNumber="204">et al</emphasis>
. 2010
</bibRefCitation>
: 1
</treatmentCitation>
, fig. 1;
<treatmentCitation id="0AF010BEC5000E5EFA8AFA2C5E5FFA22" author="Loebmann, D. &amp; Mai, A. C. G. &amp; Lee, J. T." page="918" pageId="3" pageNumber="204" year="2010">
<bibRefCitation id="EFC04B5EC5000E5EFA8AFA2C5E6AFA22" author="Loebmann, D. &amp; Mai, A. C. G. &amp; Lee, J. T." pageId="3" pageNumber="204" pagination="909 - 923" refId="ref16112" refString="Loebmann, D., Mai, A. C. G. &amp; Lee, J. T. (2010) The invasion of five alien species in the Delta do Parnaiba Environmental Protection Area, Northeastern Brazil. Revista de Biologia Tropical, 58 (3), 909 - 923. https: // doi. org / 10.15517 / rbt. v 58 i 2.5254" type="journal article" year="2010">
Loebmann
<emphasis id="B925EABDC5000E5EFF7DFA0B5FD5FA22" box="[199,255,1450,1472]" italics="true" pageId="3" pageNumber="204">et al.</emphasis>
2010
</bibRefCitation>
: 918
</treatmentCitation>
, fig. 5; Medellín
<emphasis id="B925EABDC5000E5EFD8DFA0B5D42FA22" box="[567,616,1450,1472]" italics="true" pageId="3" pageNumber="204">et al</emphasis>
. 2001: 73, unnumbered fig.;
<treatmentCitation id="0AF010BEC5000E5EFC1CFA0B5B40FA22" author="Tavares, M." box="[934,1130,1450,1472]" page="255" pageId="3" pageNumber="204" year="2011">
<bibRefCitation id="EFC04B5EC5000E5EFC1CFA0B5B1EFA22" author="Tavares, M." box="[934,1076,1450,1472]" pageId="3" pageNumber="204" pagination="251 - 268" refId="ref19109" refString="Tavares, M. (2011) Alien decapod crustaceans in the Southwestern Atlantic Ocean. In: Galil, B. S., Clark, P. F. &amp; Carlton, J. T. (Eds.) In the wrong place-Alien marine crustaceans: distribution, biology and impacts. Invading Nature Springer Series in Invasion Ecology 6. Springer, Dordrecht, Heidelberg, London and New York, pp. 251 - 268. https: // doi. org / 10.1007 / 978 - 94 - 007 - 0591 - 3 _ 7" type="book chapter" year="2011">Tavares 2011</bibRefCitation>
: 255
</treatmentCitation>
, fig. 1b;
<treatmentCitation id="0AF010BEC5000E5EFB68FA0A5AB2FA22" author="Khvorov, S." box="[1234,1432,1450,1472]" page="39" pageId="3" pageNumber="204" year="2012">
<bibRefCitation id="EFC04B5EC5000E5EFB68FA0A5A44FA22" author="Khvorov, S." box="[1234,1390,1450,1472]" pageId="3" pageNumber="204" refId="ref15528" refString="Khvorov, S. (2012) Crabs of Sultanate of Oman. Field guide. Marine Science and Fisheries Centre, Muscat, 60 pp." type="book" year="2012">Khvorov 2012</bibRefCitation>
: 39
</treatmentCitation>
, unnumbered photographs;
<treatmentCitation id="0AF010BEC5000E5EFE63FA6A5DE3FA3D" author="Bentes, A. B. &amp; Gomes Lima, W. &amp; Fernandes, S. &amp; Duarte Paula, J. &amp; Silva, K. &amp; Abrunhosa, F. &amp; Bentes, B." box="[473,713,1481,1503]" page="181" pageId="3" pageNumber="204" year="2013">
<bibRefCitation id="EFC04B5EC5000E5EFE63FA6A5DBCFA3D" author="Bentes, A. B. &amp; Gomes Lima, W. &amp; Fernandes, S. &amp; Duarte Paula, J. &amp; Silva, K. &amp; Abrunhosa, F. &amp; Bentes, B." box="[473,662,1481,1503]" pageId="3" pageNumber="204" pagination="181 - 184" refId="ref11205" refString="Bentes, A. B., Gomes Lima, W., Fernandes, S., Duarte Paula, J., Silva, K., Abrunhosa, F. &amp; Bentes, B. (2013) Occurrence of Charybdis hellerii (Milne Edwards, 1867) (Crustacea, Decapoda, Portunidae) in an Amazonian Estuary. Biota Amazonica, 3 (3), 181 - 184. https: // doi. org / 10.18561 / 2179 - 5746 / biotaamazonia. v 3 n 3 p 181 - 184" type="journal article" year="2013">
Bentes
<emphasis id="B925EABDC5000E5EFD9FFA6A5D73FA3D" box="[549,601,1481,1503]" italics="true" pageId="3" pageNumber="204">et al.</emphasis>
2013
</bibRefCitation>
: 181
</treatmentCitation>
, fig. 1; Cruz da
<treatmentCitation id="0AF010BEC5000E5EFCC8FA6A5B3CFA3D" author="Cruz da Rosa, L." box="[882,1046,1481,1503]" page="445" pageId="3" pageNumber="204" year="2014">
<bibRefCitation id="EFC04B5EC5000E5EFCC8FA6A5CC9FA3D" author="Cruz da Rosa, L." box="[882,995,1481,1503]" pageId="3" pageNumber="204" pagination="445 - 447" refId="ref12405" refString="Cruz da Rosa, L. (2014) New records of portunid crabs (Decapoda: Brachyura: Portunidae) from Sergipe, NE Brazil. Check List, 10 (2), 445 - 447. https: // doi. org / 10.15560 / 10.2.445" type="journal article" year="2014">Rosa 2014</bibRefCitation>
: 445
</treatmentCitation>
, figs 1gh;
<treatmentCitation id="0AF010BEC5000E5EFB2AFA695ABDFA3D" author="Elumalai, V. &amp; Viswanathan, C. &amp; Prakinkumar, M. &amp; Raffi, S. M." box="[1168,1431,1481,1503]" page="338" pageId="3" pageNumber="204" year="2014">
<bibRefCitation id="EFC04B5EC5000E5EFB2AFA695A4EFA3D" author="Elumalai, V. &amp; Viswanathan, C. &amp; Prakinkumar, M. &amp; Raffi, S. M." box="[1168,1380,1481,1503]" pageId="3" pageNumber="204" pagination="337 - 339" refId="ref13753" refString="Elumalai, V., Viswanathan, C., Prakinkumar, M. &amp; Raffi, S. M. (2014) Infestation of parasitic barnacle Sacculina spp. in commercial marine crabs. Journal of Parasitic Diseases, 38 (3), 337 - 339. https: // doi. org / 10.1007 / s 12639 - 013 - 0247 - z" type="journal article" year="2014">
Elumalai
<emphasis id="B925EABDC5000E5EFB49FA6A5A02FA3D" box="[1267,1320,1481,1503]" italics="true" pageId="3" pageNumber="204">et al.</emphasis>
2014
</bibRefCitation>
: 338
</treatmentCitation>
, fig. 1 (right, third row,
<taxonomicName id="4C514D2CC5000E5EFE0EFA485D32FA1C" ID-CoL="7BHH" box="[436,536,1512,1534]" class="Maxillopoda" family="Sacculinidae" genus="Sacculina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Kentrogonida" pageId="3" pageNumber="204" phylum="Arthropoda" rank="genus">
<emphasis id="B925EABDC5000E5EFE0EFA485D32FA1C" box="[436,536,1512,1534]" italics="true" pageId="3" pageNumber="204">Sacculina</emphasis>
</taxonomicName>
infestation);
<treatmentCitation id="0AF010BEC5000E5EFD1BFA485C84FA1C" author="Watanabe, T. &amp; Zara, F. J. &amp; Hattori, G. Y. &amp; Turra, A. &amp; Sant'Anna, B. S." box="[673,942,1512,1534]" page="219" pageId="3" pageNumber="204" year="2015">
<bibRefCitation id="EFC04B5EC5000E5EFD1BFA485C51FA1C" author="Watanabe, T. &amp; Zara, F. J. &amp; Hattori, G. Y. &amp; Turra, A. &amp; Sant'Anna, B. S." box="[673,891,1512,1534]" pageId="3" pageNumber="204" pagination="219 - 232" refId="ref19716" refString="Watanabe, T., Zara, F. J., Hattori, G. Y., Turra, A. &amp; Sant'Anna, B. S. (2015) Biological associations of color variation in the Indo- Pacific swimming crab Charybdis hellerii. Anais da Academia Brasileira de Ciencias, 87 (1), 219 - 232. https: // doi. org / 10.1590 / 0001 - 3765201520130388" type="journal article" year="2015">
Watanabe
<emphasis id="B925EABDC5000E5EFCB1FA495C15FA1C" box="[779,831,1512,1534]" italics="true" pageId="3" pageNumber="204">et al.</emphasis>
2015
</bibRefCitation>
: 219
</treatmentCitation>
, fig. 3;
<treatmentCitation id="0AF010BEC5000E5EFC44FA495B9DFA1C" author="McLay, C. L." box="[1022,1207,1512,1534]" page="830" pageId="3" pageNumber="204" year="2015">
<bibRefCitation id="EFC04B5EC5000E5EFC44FA495BAFFA1C" author="McLay, C. L." box="[1022,1157,1512,1534]" pageId="3" pageNumber="204" pagination="811 - 846" refId="ref16400" refString="McLay, C. L. (2015) Invasive Brachyura. In: Castro, P., Davie, P. J. F., Guinot, D., Schram, F. R. &amp; Vaupel Klein, J. C. von (Eds.), Treatise on Zoology - Anatomy, taxonomy, biology. Crustacea. Vol. 9. Part C-II. Brill, Leiden and Boston, pp. 811 - 846. https: // doi. org / 10.1163 / 9789004190832 _ 017" type="book chapter" year="2015">McLay 2015</bibRefCitation>
: 830
</treatmentCitation>
, fig. 71b;
<treatmentCitation id="0AF010BEC5000E5EFA9BFA485E05F9FE" author="Abbas, E. M. &amp; Abdelsalam, K. M. &amp; Mohammed-Geba, K. &amp; Ahmed, H. O. &amp; Kato, M." page="321" pageId="3" pageNumber="204" year="2016">
<bibRefCitation id="EFC04B5EC5000E5EFA9BFA485FD7F9FE" author="Abbas, E. M. &amp; Abdelsalam, K. M. &amp; Mohammed-Geba, K. &amp; Ahmed, H. O. &amp; Kato, M." pageId="3" pageNumber="204" pagination="319 - 329" refId="ref10772" refString="Abbas, E. M., Abdelsalam, K. M., Mohammed-Geba, K., Ahmed, H. O. &amp; Kato, M. (2016) Genetic and morphological identification of some crabs from the Gulf of Suez, Northern Red Sea, Egypt. Egyptian Journal of Aquatic Research, 42, 319 - 329. https: // doi. org / 10.1016 / j. ejar. 2016.08.003" type="journal article" year="2016">
Abbas
<emphasis id="B925EABDC5000E5EFAD2FA495AB7FA1C" box="[1384,1437,1512,1534]" italics="true" pageId="3" pageNumber="204">et al.</emphasis>
2016
</bibRefCitation>
: 321
</treatmentCitation>
, fig. 3;
<treatmentCitation id="0AF010BEC5000E5EFEC4F9A65D46F9FE" author="Cuesta, J. A. &amp; Almon, B. &amp; Perez-Dieste, J. &amp; Trigo, J. E. &amp; Banon, R." box="[382,620,1542,1565]" page="622" pageId="3" pageNumber="204" year="2016">
<bibRefCitation id="EFC04B5EC5000E5EFEC4F9A65D11F9FE" author="Cuesta, J. A. &amp; Almon, B. &amp; Perez-Dieste, J. &amp; Trigo, J. E. &amp; Banon, R." box="[382,571,1542,1565]" pageId="3" pageNumber="204" pagination="619 - 630" refId="ref12454" refString="Cuesta, J. A., Almon, B., Perez-Dieste, J., Trigo, J. E. &amp; Banon, R. (2016) Role of ships' hull fouling and tropicalization process on European carcinofauna: new records in Galician waters (NW Spain). Biological Invasions, 19, 619 - 630. https: // doi. org / 10.1007 / s 10530 - 015 - 1034 - 9" type="journal article" year="2016">
Cuesta
<emphasis id="B925EABDC5000E5EFE70F9A85ED4F9FE" box="[458,510,1542,1565]" italics="true" pageId="3" pageNumber="204">et al.</emphasis>
2016
</bibRefCitation>
: 622
</treatmentCitation>
, 624, fig. 2B (
<collectingCountry id="F346763FC5000E5EFCBAF9A65C15F9FE" box="[768,831,1542,1564]" name="Spain" pageId="3" pageNumber="204">Spain</collectingCountry>
); da
<treatmentCitation id="0AF010BEC5000E5EFCD5F9A65B4CF9FE" author="da Silva Reis, N. &amp; da Conceicao Silva, D. &amp; Pinho Rabelo, A. C. &amp; dos Santos Nascimento, M. &amp; Eocha Pereira, D." box="[879,1126,1542,1565]" page="1" pageId="3" pageNumber="204" year="2016">
<bibRefCitation id="EFC04B5EC5000E5EFCD5F9A65B64F9FE" author="da Silva Reis, N. &amp; da Conceicao Silva, D. &amp; Pinho Rabelo, A. C. &amp; dos Santos Nascimento, M. &amp; Eocha Pereira, D." box="[879,1102,1542,1565]" pageId="3" pageNumber="204" refId="ref12695" refString="da Silva Reis, N., da Conceicao Silva, D., Pinho Rabelo, A. C., dos Santos Nascimento, M. &amp; Eocha Pereira, D. (2016) Agua de Lastro: bioinvasao por siri Charybdis hellerii. In: 10 ° Simposio Internacional de Qualidade Ambiental, Predio 41, PUCRS, Porto Alegre, 13 pp." type="book" year="2016">
Silva Reis
<emphasis id="B925EABDC5000E5EFC64F9A85B38F9FE" box="[990,1042,1542,1565]" italics="true" pageId="3" pageNumber="204">et al.</emphasis>
2016
</bibRefCitation>
: 1
</treatmentCitation>
, figs 13;
<treatmentCitation id="0AF010BEC5000E5EFB69F9A65FFEF9D9" author="Negri Pereira, M." page="1" pageId="3" pageNumber="204" year="2016">
<bibRefCitation id="EFC04B5EC5000E5EFB69F9A65AB3F9FE" author="Negri Pereira, M." box="[1235,1433,1542,1564]" pageId="3" pageNumber="204" refId="ref17215" refString="Negri Pereira, M. (2016) Registro de Charybdis hellerii (Milne Edwards, 1867) em aguas do litoral brasileiro (Decapoda: Portunidae). Tese apresentada a Faculdade de Filosofia, Ciencias e Letras de Ribeirao Preto da USP, como parte das exigencias para a obtencao do titulo de Doutor em Ciencias Area: Biologia Comparada, viii + 164 pp." type="book" year="2016">Negri Pereira 2016</bibRefCitation>
: 1
</treatmentCitation>
, figs 1, 17, 18, 19, 20b;
<treatmentCitation id="0AF010BEC5000E5EFE6FF9865DB9F9D9" author="Ng, P. K. L. &amp; Shih, H. - T. &amp; Ho, P. - H. &amp; Wang, C. - H." box="[469,659,1573,1595]" page="72" pageId="3" pageNumber="204" year="2017">
<bibRefCitation id="EFC04B5EC5000E5EFE6FF9865D44F9D9" author="Ng, P. K. L. &amp; Shih, H. - T. &amp; Ho, P. - H. &amp; Wang, C. - H." box="[469,622,1573,1595]" pageId="3" pageNumber="204" pagination="1 - 185" refId="ref17410" refString="Ng, P. K. L., Shih, H. - T., Ho, P. - H. &amp; Wang, C. - H. (2017) An updated annotated checklist of brachyuran crabs from Taiwan (Crustacea: Decapoda). Journal of the National Taiwan Museum, 70 (3 &amp; 4), 1 - 185." type="journal article" year="2017">
Ng
<emphasis id="B925EABDC5000E5EFE46F9865D00F9D9" box="[508,554,1573,1595]" italics="true" pageId="3" pageNumber="204">et al</emphasis>
. 2017
</bibRefCitation>
: 72
</treatmentCitation>
(list of
<collectingCountry id="F346763FC5000E5EFD5DF9855C1BF9D9" box="[743,817,1573,1595]" name="Taiwan" pageId="3" pageNumber="204">Taiwan</collectingCountry>
records);
<treatmentCitation id="0AF010BEC5000E5EFC21F9865BAAF9D9" author="Ferry, R. &amp; Buske, Y. &amp; Poupin, J. &amp; Smith-Ravin, J." box="[923,1152,1573,1595]" page="240" pageId="3" pageNumber="204" year="2017">
<bibRefCitation id="EFC04B5EC5000E5EFC21F9865B66F9D9" author="Ferry, R. &amp; Buske, Y. &amp; Poupin, J. &amp; Smith-Ravin, J." box="[923,1100,1573,1595]" pageId="3" pageNumber="204" pagination="239 - 247" refId="ref14075" refString="Ferry, R., Buske, Y., Poupin, J. &amp; Smith-Ravin, J. (2017) First record of the invasive swimming crab Charybdis hellerii (A. Milne Edwards, 1867) (Crustacea, Portunidae) off Martinique, French Lesser Antilles. BioInvasions Records, 6 (3), 239 - 247. https: // doi. org / 10.3391 / bir. 20176.3.09" type="journal article" year="2017">
Ferry
<emphasis id="B925EABDC5000E5EFC60F9865B22F9D9" box="[986,1032,1573,1595]" italics="true" pageId="3" pageNumber="204">et al</emphasis>
. 2017
</bibRefCitation>
: 240
</treatmentCitation>
, fig. 1;
<treatmentCitation id="0AF010BEC5000E5EFB6BF9855E1BF9B8" author="Negri, M. &amp; Mantelatto, F. L." page="282" pageId="3" pageNumber="204" year="2017">
<bibRefCitation id="EFC04B5EC5000E5EFB6BF9855FD7F9B8" author="Negri, M. &amp; Mantelatto, F. L." pageId="3" pageNumber="204" pagination="278 - 284" refId="ref17066" refString="Negri, M. &amp; Mantelatto, F. L. (2017) Integrative taxonomy reveals that Charybdis variegata (Fabricius, 1798) (Brachyura: Portunidae) has not been introduced in the South Atlantic Ocean. Journal of Crustacean Biology, 37 (3), 278 - 284. https: // doi. org / 10.1093 / jcbiol / rux 023" type="journal article" year="2017">Negri &amp; Mantelatto 2017</bibRefCitation>
: 282
</treatmentCitation>
, figs 2a, 3a;
<treatmentCitation id="0AF010BEC5000E5EFE03F9E45D77F9B8" author="Questel, K." box="[441,605,1604,1626]" page="1" pageId="3" pageNumber="204" year="2017">
<bibRefCitation id="EFC04B5EC5000E5EFE03F9E45D6FF9B8" author="Questel, K." box="[441,581,1604,1626]" pageId="3" pageNumber="204" pagination="1" refId="ref18182" refString="Questel, K. (2017) Le crabe Charybdis hellerii. Une nouvelle espece exotique marine pour Saint-Barthelemy. Le Bulletin de l'ATE [Agence Territoriale de l'environnement St Barthelemy], 1, 1." type="journal article" year="2017">Questel 2017</bibRefCitation>
: 1
</treatmentCitation>
, unnumbered photographs;
<treatmentCitation id="0AF010BEC5000E5EFCC4F9E45B4BF9B8" author="Santhanam, R." box="[894,1121,1604,1626]" page="136" pageId="3" pageNumber="204" year="2018">
<bibRefCitation id="EFC04B5EC5000E5EFCC4F9E45B07F9B8" author="Santhanam, R." box="[894,1069,1604,1626]" pageId="3" pageNumber="204" refId="ref18734" refString="Santhanam, R. (2018) Biology and Culture of Portunid Crabs of World Seas. Series: Biology and Ecology of Marine Life. Apple Academic Press, Waretown, NJ, 404 pp. https: // doi. org / 10.1201 / 9781315205816" type="book" year="2018">Santhanam 2018</bibRefCitation>
: 136
</treatmentCitation>
, unnumbered fig.;
<treatmentCitation id="0AF010BEC5000E5EFA9FF9E45E16F99A" author="Negri, M. &amp; Schubart, C. D. &amp; Mantelatto, F. L." page="1771" pageId="3" pageNumber="204" year="2018">
<bibRefCitation id="EFC04B5EC5000E5EFA9FF9E45FD7F99A" author="Negri, M. &amp; Schubart, C. D. &amp; Mantelatto, F. L." pageId="3" pageNumber="204" pagination="1771 - 1798" refId="ref17133" refString="Negri, M., Schubart, C. D. &amp; Mantelatto, F. L. (2018) Tracing the introduction history of the invasive swimming crab Charybdis hellerii (A. Milne-Edwards, 1867) in the Western Atlantic: evidences of high genetic diversity and multiple introductions. Biological Invasions, 20 (7), 1771 - 1798. https: // doi. org / 10.1007 / s 10530 - 018 - 1660 - 0" type="journal article" year="2018">
Negri
<emphasis id="B925EABDC5000E5EFADDF9E55AB7F9B8" box="[1383,1437,1604,1626]" italics="true" pageId="3" pageNumber="204">et al.</emphasis>
2018
</bibRefCitation>
: 1771
</treatmentCitation>
, figs 45; Evans
<emphasis id="B925EABDC5000E5EFE50F9C35D37F99A" box="[490,541,1634,1656]" italics="true" pageId="3" pageNumber="204">et al.</emphasis>
2018: 641, figs 1, 2A, 4, 5 (Hawai'i).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BEE36AFC5000E5EFF2DF9215D85F954" blockId="3.[151,1437,500,1933]" pageId="3" pageNumber="204">
<treatmentCitationGroup id="AB411181C5000E5EFF2DF9215D85F954" pageId="3" pageNumber="204">
<taxonomicName id="4C514D2CC5000E5EFF2DF9215E63F975" box="[151,329,1665,1687]" class="Malacostraca" family="Portunidae" genus="Charybdis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="helleri">
<emphasis id="B925EABDC5000E5EFF2DF9215E63F975" box="[151,329,1665,1687]" italics="true" pageId="3" pageNumber="204">Charybdis helleri</emphasis>
</taxonomicName>
.—
<treatmentCitation id="0AF010BEC5000E5EFEDCF9215D24F975" author="Riedl, R." box="[358,526,1665,1687]" page="501" pageId="3" pageNumber="204" year="1983">
<bibRefCitation id="EFC04B5EC5000E5EFEDCF9215EF6F975" author="Riedl, R." box="[358,476,1665,1687]" pageId="3" pageNumber="204" refId="ref18280" refString="Riedl, R. (1983) Fauna und Flora des Mittelmeeres. Paul Parey, Hamburg &amp; Berlin, 836 pp., 15 pls." type="book" year="1983">Riedl 1983</bibRefCitation>
: 501
</treatmentCitation>
, pl. 501, unnumbered fig.;
<treatmentCitation id="0AF010BEC5000E5EFC9BF9215CCDF975" author="Holthuis, L. B." box="[801,999,1665,1687]" page="350" pageId="3" pageNumber="204" year="1987">
<bibRefCitation id="EFC04B5EC5000E5EFC9BF9215C9EF975" author="Holthuis, L. B." box="[801,948,1665,1687]" pageId="3" pageNumber="204" pagination="321 - 367" refId="ref15211" refString="Holthuis, L. B. (1987) Vrais Crabes. In: Fischer, W., Schneider, M. &amp; Bauchot, M. - L. (Eds.), Fiches FAO d'identification des especes pour les besoins de la peche. Mediterranee et Mer Noire. Zone de peche 37. Revision 1. Vol. 1. Vegetaux et invertebres. Food and Agriculture Organization of the United Nations, Rome, pp. 321 - 367." type="book chapter" year="1987">Holthuis 1987</bibRefCitation>
: 350
</treatmentCitation>
, fig. 10;
<treatmentCitation id="0AF010BEC5000E5EFBFBF9215FC9F954" author="Gomez, C. &amp; Martinez-Iglesias, J. C." page="70" pageId="3" pageNumber="204" year="1990">
<bibRefCitation id="EFC04B5EC5000E5EFBFBF9215AB2F975" author="Gomez, C. &amp; Martinez-Iglesias, J. C." box="[1089,1432,1665,1687]" pageId="3" pageNumber="204" pagination="70 - 72" refId="ref14802" refString="Gomez, C. &amp; Martinez-Iglesias, J. C. (1990) Reciente hallazgo de la especie indopacifica Charybdis helleri (A. Milne Edwards, 1867) (Crustacea: Decapoda: Portunidae:) en aguas cubanas. Caribbean Journal of Science, 26 (l- 2), 70 - 72." type="journal article" year="1990">Gómez &amp; Martínez-Iglesias 1990</bibRefCitation>
: 70
</treatmentCitation>
, fig. 1;
<treatmentCitation id="0AF010BEC5000E5EFE8AF9015ED6F954" author="Fransen, C. H. J. M." box="[304,508,1696,1718]" page="316" pageId="3" pageNumber="204" year="2014">
<bibRefCitation id="EFC04B5EC5000E5EFE8AF9015EE3F954" author="Fransen, C. H. J. M." box="[304,457,1696,1718]" pageId="3" pageNumber="204" pagination="249 - 358" refId="ref14314" refString="Fransen, C. H. J. M. (2014 b) True crabs. In: Carpenter, K. E. &amp; De Angelis, N. (Eds.), The living marine resources of the Eastern Central Atlantic. FAO species identification guide for fishery purposes. Foods and Agriculture Organization of the United Nations, Rome, pp. 249 - 358." type="book chapter" year="2014">Fransen 2014b</bibRefCitation>
: 316
</treatmentCitation>
, fig. 31 [
<collectingCountry id="F346763FC5000E5EFDEEF9005D8FF954" box="[596,677,1696,1718]" name="Guinea-Bissau" pageId="3" pageNumber="204">Guinea</collectingCountry>
].
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BEE36AFC5000E5EFF2DF91E5D59F936" blockId="3.[151,1437,500,1933]" box="[151,627,1726,1749]" pageId="3" pageNumber="204">
<treatmentCitationGroup id="AB411181C5000E5EFF2DF91E5D59F936" box="[151,627,1726,1749]" pageId="3" pageNumber="204">
<taxonomicName id="4C514D2CC5000E5EFF2DF91E5E51F937" ID-CoL="9CBDG" box="[151,379,1726,1749]" class="Malacostraca" family="Portunidae" genus="Charybdis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="merguiense">
<emphasis id="B925EABDC5000E5EFF2DF91E5E51F937" box="[151,379,1726,1749]" italics="true" pageId="3" pageNumber="204">Charybdis merguiense</emphasis>
</taxonomicName>
.—
<treatmentCitation id="0AF010BEC5000E5EFE22F91E5D1BF936" author="Deb, M." box="[408,561,1726,1748]" page="144" pageId="3" pageNumber="204" year="1998">
<bibRefCitation id="EFC04B5EC5000E5EFE22F91E5D2BF936" author="Deb, M." box="[408,513,1726,1748]" pageId="3" pageNumber="204" pagination="129 - 159" refId="ref12830" refString="Deb, M. (1998) Crustacea. Zoological Survey of India. Estuarine Ecosystem Series, 3, Mahanadi Estuary, pp. 129 - 159." type="book chapter" year="1998">Deb 1998</bibRefCitation>
: 144
</treatmentCitation>
(key).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C34B6524C5000E5EFF2DF97D5D94F911" box="[151,702,1757,1779]" pageId="3" pageNumber="204" type="description">
<paragraph id="8BEE36AFC5000E5EFF2DF97D5D94F911" blockId="3.[151,1437,500,1933]" box="[151,702,1757,1779]" pageId="3" pageNumber="204">
<treatmentCitationGroup id="AB411181C5000E5EFF2DF97D5D94F911" box="[151,702,1757,1779]" pageId="3" pageNumber="204">
<taxonomicName id="4C514D2CC5000E5EFF2DF97D5D2AF911" ID-CoL="TPV8" baseAuthorityName="De Man" baseAuthorityYear="1887" box="[151,512,1757,1779]" class="Malacostraca" family="Portunidae" genus="Charybdis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="merguiensis" subGenus="Charybdis">
<emphasis id="B925EABDC5000E5EFF2DF97D5E2BF911" box="[151,257,1757,1779]" italics="true" pageId="3" pageNumber="204">Charybdis</emphasis>
(
<emphasis id="B925EABDC5000E5EFEB5F97D5E52F911" box="[271,376,1757,1779]" italics="true" pageId="3" pageNumber="204">Charybdis</emphasis>
)
<emphasis id="B925EABDC5000E5EFE3DF97E5D2AF911" box="[391,512,1758,1779]" italics="true" pageId="3" pageNumber="204">merguiensis</emphasis>
</taxonomicName>
.—
<treatmentCitation id="0AF010BEC5000E5EFDA5F97D5D90F911" author="Deb, M." box="[543,698,1757,1779]" page="145" pageId="3" pageNumber="204" year="1998">
<bibRefCitation id="EFC04B5EC5000E5EFDA5F97D5DA2F911" author="Deb, M." box="[543,648,1757,1779]" pageId="3" pageNumber="204" pagination="129 - 159" refId="ref12830" refString="Deb, M. (1998) Crustacea. Zoological Survey of India. Estuarine Ecosystem Series, 3, Mahanadi Estuary, pp. 129 - 159." type="book chapter" year="1998">Deb 1998</bibRefCitation>
: 145
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C34B6524C5000E5EFF2DF95C5C28F8AD" pageId="3" pageNumber="204" type="reference_group">
<paragraph id="8BEE36AFC5000E5EFF2DF95C5D41F8D2" blockId="3.[151,1437,500,1933]" pageId="3" pageNumber="204">
<treatmentCitationGroup id="AB411181C5000E5EFF2DF95C5D41F8D2" pageId="3" pageNumber="204">
<taxonomicName id="4C514D2CC5000E5EFF2DF95C5EE6F8F0" box="[151,460,1788,1810]" class="Malacostraca" family="Portunidae" genus="Charybdis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="helleri" subGenus="Charybdis">
<emphasis id="B925EABDC5000E5EFF2DF95C5E2BF8F0" box="[151,257,1788,1810]" italics="true" pageId="3" pageNumber="204">Charybdis</emphasis>
(
<emphasis id="B925EABDC5000E5EFEAAF95C5E53F8F0" box="[272,377,1788,1810]" italics="true" pageId="3" pageNumber="204">Charybdis</emphasis>
)
<emphasis id="B925EABDC5000E5EFE30F95C5EE6F8F0" box="[394,460,1788,1810]" italics="true" pageId="3" pageNumber="204">helleri</emphasis>
</taxonomicName>
.—
<treatmentCitation id="0AF010BEC5000E5EFE53F95C5C39F8F0" author="Jeyabaskaran, R. &amp; Ajmal Khan, S. &amp; Ramaiyan, V." box="[489,787,1788,1810]" page="52" pageId="3" pageNumber="204" year="2000">
<bibRefCitation id="EFC04B5EC5000E5EFE53F95C5DC7F8F0" author="Jeyabaskaran, R. &amp; Ajmal Khan, S. &amp; Ramaiyan, V." box="[489,749,1788,1810]" pageId="3" pageNumber="204" refId="ref15427" refString="Jeyabaskaran, R., Ajmal Khan, S. &amp; Ramaiyan, V. (2000) Brachyuran crabs of Gulf of Mannar. Annamalai University, Centre of Advanced Study in Marine Biology: Parangipettai, 99 pp., 78 pls." type="book" year="2000">
Jeyabaskaran
<emphasis id="B925EABDC5000E5EFDC0F95D5D85F8F0" box="[634,687,1788,1810]" italics="true" pageId="3" pageNumber="204">et al.</emphasis>
2000
</bibRefCitation>
: 52
</treatmentCitation>
, pl. 38A;
<treatmentCitation id="0AF010BEC5000E5EFCC0F95D5BAFF8F0" author="Dev Roy, M. K. &amp; Das, A. K." box="[890,1157,1788,1810]" page="36" pageId="3" pageNumber="204" year="2000">
<bibRefCitation id="EFC04B5EC5000E5EFCC0F95D5B75F8F0" author="Dev Roy, M. K. &amp; Das, A. K." box="[890,1119,1788,1810]" pageId="3" pageNumber="204" pagination="1 - 211" refId="ref13498" refString="Dev Roy, M. K. &amp; Das, A. K. (2000) Taxonomy, ecobiology and distribution pattern of the Brachyuran Crabs of mangrove ecosystem in Andaman Islands. In: Records of the zoological Survey of India Occasional Paper. Vol. 185. Published- Director, ZSI, Calcutta, pp. 1 - 211." type="book chapter" year="2000">Dev Roy &amp; Das 2000</bibRefCitation>
: 36
</treatmentCitation>
, pl. 2 fig. 1, pl. 9 figs 78;
<treatmentCitation id="0AF010BEC5000E5EFF7DF8BB5EDCF8D2" author="Dev Roy, M. K. &amp; Bhadra, S." box="[199,502,1818,1840]" page="150" pageId="3" pageNumber="204" year="2011">
<bibRefCitation id="EFC04B5EC5000E5EFF7DF8BB5EECF8D2" author="Dev Roy, M. K. &amp; Bhadra, S." box="[199,454,1818,1840]" pageId="3" pageNumber="204" pagination="109 - 269" refId="ref13450" refString="Dev Roy, M. K. &amp; Bhadra, S. (2011) Brachyuran crabs (Crustacea: Decapoda: Brachyura). Zoological Survey of India. Fauna of Tamil Nadu, State Fauna Series, 17 (2), 109 - 269." type="journal article" year="2011">Dev Roy &amp; Bhadra 2011</bibRefCitation>
: 150
</treatmentCitation>
(key), 151.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BEE36AFC5000E5EFF2DF8995C28F8AD" blockId="3.[151,1437,500,1933]" box="[151,770,1849,1871]" pageId="3" pageNumber="204">
<treatmentCitationGroup id="AB411181C5000E5EFF2DF8995C28F8AD" box="[151,770,1849,1871]" pageId="3" pageNumber="204">
<taxonomicName id="4C514D2CC5000E5EFF2DF8995ECBF8AD" ID-CoL="5XQ2Z" baseAuthorityName="Miers" baseAuthorityYear="1884" box="[151,481,1849,1871]" class="Malacostraca" family="Portunidae" genus="Charybdis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="spinifera" subGenus="Charybdis">
<emphasis id="B925EABDC5000E5EFF2DF8995E2BF8AD" box="[151,257,1849,1871]" italics="true" pageId="3" pageNumber="204">Charybdis</emphasis>
(
<emphasis id="B925EABDC5000E5EFEB5F8995E52F8AD" box="[271,376,1849,1871]" italics="true" pageId="3" pageNumber="204">Charybdis</emphasis>
)
<emphasis id="B925EABDC5000E5EFE3DF8995ECBF8AD" box="[391,481,1849,1871]" italics="true" pageId="3" pageNumber="204">spinifera</emphasis>
</taxonomicName>
.—
<treatmentCitation id="0AF010BEC5000E5EFDBAF89A5DEFF8AD" author="Ng, P. K. L. &amp; Guinot, D. &amp; Davie, P. J. F." box="[512,709,1849,1871]" page="153" pageId="3" pageNumber="204" year="2008">
<bibRefCitation id="EFC04B5EC5000E5EFDBAF89A5DBFF8AD" author="Ng, P. K. L. &amp; Guinot, D. &amp; Davie, P. J. F." box="[512,661,1849,1871]" pageId="3" pageNumber="204" pagination="1 - 286" refId="ref17354" refString="Ng, P. K. L., Guinot, D. &amp; Davie, P. J. F. (2008) Systema Brachyurorum: part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, 17 (Supplement), 1 - 286." type="journal article" year="2008">
Ng
<emphasis id="B925EABDC5000E5EFD9CF89A5D79F8AD" box="[550,595,1849,1871]" italics="true" pageId="3" pageNumber="204">et al</emphasis>
. 2008
</bibRefCitation>
: 153
</treatmentCitation>
(list).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C34B6524C5000E5EFF2DF8F85DFFF86E" pageId="3" pageNumber="204" type="description">
<paragraph id="8BEE36AFC5000E5EFF2DF8F85C21F88C" blockId="3.[151,1437,500,1933]" box="[151,779,1880,1902]" pageId="3" pageNumber="204">
<treatmentCitationGroup id="AB411181C5000E5EFF2DF8F85C21F88C" box="[151,779,1880,1902]" pageId="3" pageNumber="204">
<taxonomicName id="4C514D2CC5000E5EFF2DF8F85EC3F88C" ID-CoL="84GRH" authorityName="Ward" authorityYear="1941" box="[151,489,1880,1902]" class="Malacostraca" family="Portunidae" genus="Charybdis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="vannamei" subGenus="Charybdis">
<emphasis id="B925EABDC5000E5EFF2DF8F85E2BF88C" box="[151,257,1880,1902]" italics="true" pageId="3" pageNumber="204">Charybdis</emphasis>
(
<emphasis id="B925EABDC5000E5EFEB5F8F85E52F88C" box="[271,376,1880,1902]" italics="true" pageId="3" pageNumber="204">Charybdis</emphasis>
)
<emphasis id="B925EABDC5000E5EFE3DF8F95EC3F88C" box="[391,489,1881,1902]" italics="true" pageId="3" pageNumber="204">vannamei</emphasis>
</taxonomicName>
.—
<treatmentCitation id="0AF010BEC5000E5EFDB2F8F95DE7F88C" author="Ng, P. K. L. &amp; Guinot, D. &amp; Davie, P. J. F." box="[520,717,1880,1902]" page="153" pageId="3" pageNumber="204" year="2008">
<bibRefCitation id="EFC04B5EC5000E5EFDB2F8F95DB4F88C" author="Ng, P. K. L. &amp; Guinot, D. &amp; Davie, P. J. F." box="[520,670,1880,1902]" pageId="3" pageNumber="204" pagination="1 - 286" refId="ref17354" refString="Ng, P. K. L., Guinot, D. &amp; Davie, P. J. F. (2008) Systema Brachyurorum: part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, 17 (Supplement), 1 - 286." type="journal article" year="2008">
Ng
<emphasis id="B925EABDC5000E5EFD94F8F95D71F88C" box="[558,603,1880,1902]" italics="true" pageId="3" pageNumber="204">et al</emphasis>
. 2008
</bibRefCitation>
: 153
</treatmentCitation>
(list).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BEE36AFC5000E5EFF2DF8D65DFFF86E" blockId="3.[151,1437,500,1933]" box="[151,725,1910,1933]" pageId="3" pageNumber="204">
<taxonomicName id="4C514D2CC5000E5EFF2DF8D65E42F86F" box="[151,360,1910,1933]" class="Malacostraca" family="Portunidae" genus="Charybdis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="204" phylum="Arthropoda" rank="genus">
<emphasis id="B925EABDC5000E5EFF2DF8D65E42F86F" box="[151,360,1910,1933]" italics="true" pageId="3" pageNumber="204">Charybdis variegata</emphasis>
</taxonomicName>
.—Sant'Anna
<emphasis id="B925EABDC5000E5EFE4CF8D85D03F86E" box="[502,553,1910,1933]" italics="true" pageId="3" pageNumber="204">et al.</emphasis>
2012a: 12, fig. 2
</paragraph>
</subSubSection>
<subSubSection id="C34B6524C5000E59FF2DF8155BC1FEE8" lastPageId="4" lastPageNumber="205" pageId="3" pageNumber="204" type="reference_group">
<paragraph id="8BEE36AFC5000E5EFF2DF8155AB6F828" blockId="3.[151,1436,1972,2024]" box="[151,1436,1972,1994]" pageId="3" pageNumber="204">
non
<taxonomicName id="4C514D2CC5000E5EFF7DF8145D75F828" authority="Herbst, 1783: 153" authorityName="Herbst" authorityPageNumber="153" authorityYear="1783" box="[199,607,1972,1994]" class="Malacostraca" family="Cancridae" genus="Cancer" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="sexdentatus">
<emphasis id="B925EABDC5000E5EFF7DF8145EB8F828" box="[199,402,1972,1994]" italics="true" pageId="3" pageNumber="204">Cancer sexdentatus</emphasis>
Herbst, 1783: 153
</taxonomicName>
, pl. 7 fig. 52 [=
<taxonomicName id="4C514D2CC5000E5EFCA7F8145BBAF828" ID-CoL="TPT6" authority="(Linnaeus, 1758)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[797,1168,1972,1994]" class="Malacostraca" family="Portunidae" genus="Charybdis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="feriata">
<emphasis id="B925EABDC5000E5EFCA7F8145CFFF828" box="[797,981,1972,1994]" italics="true" pageId="3" pageNumber="204">Charybdis feriata</emphasis>
(Linnaeus, 1758)
</taxonomicName>
by
<typeStatus id="54EA880DC5000E5EFB7AF8145A37F828" box="[1216,1309,1972,1994]" pageId="3" pageNumber="204" type="lectotype">lectotype</typeStatus>
designation
</paragraph>
<paragraph id="8BEE36AFC5000E5EFF7DF8725E5DF80A" blockId="3.[151,1436,1972,2024]" box="[199,375,2002,2024]" pageId="3" pageNumber="204">
(
<bibRefCitation id="EFC04B5EC5000E5EFF74F8725E4FF80A" author="Holthuis, L. B." box="[206,357,2002,2024]" pageId="3" pageNumber="204" pagination="232 - 253" refId="ref15163" refString="Holthuis, L. B. (1962) Forty-seven genera of Decapoda (Crustacea); proposed addition to the official list. Z. N. (S.) 1499. Bulletin of Zoological Nomenclature, 19 (4), 232 - 253." type="journal article" year="1962">Holthuis 1962</bibRefCitation>
)].
</paragraph>
<paragraph id="8BEE36AFC5070E59FF2DFF395D42FF4C" blockId="4.[151,1390,152,266]" box="[151,616,152,174]" pageId="4" pageNumber="205">
<treatmentCitationGroup id="AB411181C5070E59FF2DFF395D42FF4C" box="[151,616,152,174]" pageId="4" pageNumber="205">
non
<taxonomicName id="4C514D2CC5070E59FF79FF395D4EFF4C" ID-CoL="4M2MN" authority="Fabricius, 1798: 364" authorityName="Fabricius" authorityPageNumber="364" authorityYear="1798" box="[195,612,152,174]" class="Malacostraca" family="Portunidae" genus="Portunus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="annulatus">
<emphasis id="B925EABDC5070E59FF79FF395EA0FF4C" box="[195,394,152,174]" italics="true" pageId="4" pageNumber="205">Portunus annulatus</emphasis>
<treatmentCitation id="0AF010BEC5070E59FE2AFF385D4EFF4C" author="Fabricius, J. C." box="[400,612,152,174]" page="364" pageId="4" pageNumber="205" year="1798">
<bibRefCitation id="EFC04B5EC5070E59FE2AFF385D1BFF4C" author="Fabricius, J. C." box="[400,561,152,174]" pageId="4" pageNumber="205" refId="ref13952" refString="Fabricius, J. C. (1798) Entomologia Systematica emendata et aucta, secundum classes, ordines, genera, species adjectis synonimis locis observationibus descriptionibus. Supplementum Entomologiae Systematicae. Impensis Christ. Gottl. Proft, Hafniae, 572 pp." type="book" year="1798">Fabricius, 1798</bibRefCitation>
: 364
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BEE36AFC5070E59FF2DFF175D45FF2E" blockId="4.[151,1390,152,266]" box="[151,623,182,205]" pageId="4" pageNumber="205">
<treatmentCitationGroup id="AB411181C5070E59FF2DFF175D45FF2E" box="[151,623,182,205]" pageId="4" pageNumber="205">
non
<taxonomicName id="4C514D2CC5070E59FF79FF185D41FF2E" ID-CoL="4M2Q4" authority="Fabricius, 1798: 364" authorityName="Fabricius" authorityPageNumber="364" authorityYear="1798" box="[195,619,182,205]" class="Malacostraca" family="Portunidae" genus="Portunus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="variegatus">
<emphasis id="B925EABDC5070E59FF79FF185EBBFF2F" box="[195,401,184,205]" italics="true" pageId="4" pageNumber="205">Portunus variegatus</emphasis>
<treatmentCitation id="0AF010BEC5070E59FE2DFF165D41FF2E" author="Fabricius, J. C." box="[407,619,182,204]" page="364" pageId="4" pageNumber="205" year="1798">
<bibRefCitation id="EFC04B5EC5070E59FE2DFF165D12FF2E" author="Fabricius, J. C." box="[407,568,182,204]" pageId="4" pageNumber="205" refId="ref13952" refString="Fabricius, J. C. (1798) Entomologia Systematica emendata et aucta, secundum classes, ordines, genera, species adjectis synonimis locis observationibus descriptionibus. Supplementum Entomologiae Systematicae. Impensis Christ. Gottl. Proft, Hafniae, 572 pp." type="book" year="1798">Fabricius, 1798</bibRefCitation>
: 364
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BEE36AFC5070E59FF2DFF765A44FF09" blockId="4.[151,1390,152,266]" box="[151,1390,213,235]" pageId="4" pageNumber="205">
<treatmentCitationGroup id="AB411181C5070E59FF2DFF765A44FF09" box="[151,1390,213,235]" pageId="4" pageNumber="205">
non
<taxonomicName id="4C514D2CC5070E59FF79FF755D3BFF09" authority="Hellerii." authorityName="Hellerii." box="[195,529,213,235]" class="Malacostraca" family="Portunidae" genus="Charybdis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="205" phylum="Arthropoda" rank="subGenus" subGenus="Goniosoma">
<emphasis id="B925EABDC5070E59FF79FF755E06FF09" box="[195,300,213,235]" italics="true" pageId="4" pageNumber="205">Charybdis</emphasis>
(
<emphasis id="B925EABDC5070E59FE81FF765E9AFF09" box="[315,432,214,235]" italics="true" pageId="4" pageNumber="205">Goniosoma</emphasis>
)
<emphasis id="B925EABDC5070E59FE05FF755D24FF09" box="[447,526,213,235]" italics="true" pageId="4" pageNumber="205">Hellerii</emphasis>
.
</taxonomicName>
<treatmentCitation id="0AF010BEC5070E59FD91FF755DF6FF09" author="Nobili, G." box="[555,732,213,235]" page="194" pageId="4" pageNumber="205" year="1906">
<bibRefCitation id="EFC04B5EC5070E59FD91FF755D80FF09" author="Nobili, G." box="[555,682,213,235]" pageId="4" pageNumber="205" pagination="1 - 347" refId="ref17619" refString="Nobili, G. (1906) Faune carcinologique de la Mer Rouge. Decapodes et stomatopodes. Annales des Sciences Naturelles, Serie 9, 4, 1 - 347. https: // doi. org / 10.5962 / bhl. title. 10635" type="journal article" year="1906">Nobili 1906</bibRefCitation>
: 194
</treatmentCitation>
, 195 [=
<taxonomicName id="4C514D2CC5070E59FC8AFF755B54FF09" ID-CoL="TPTR" authority="Dana, 1852" authorityName="Dana" authorityYear="1852" box="[816,1150,213,235]" class="Malacostraca" family="Portunidae" genus="Charybdis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis id="B925EABDC5070E59FC8AFF755B2BFF09" box="[816,1025,213,235]" italics="true" pageId="4" pageNumber="205">Charybdis orientalis</emphasis>
<bibRefCitation id="EFC04B5EC5070E59FBBDFF765B54FF09" author="Dana, J. D." box="[1031,1150,213,235]" pageId="4" pageNumber="205" pagination="73 - 86" refId="ref12644" refString="Dana, J. D. (1852) Conspectus of the Crustacea of the Exploring Expedition under Capt. Wilkes, U. S. N., including the Crustacea Cancroidea Corystoidea. Proceedings of the Academy of Natural Sciences of Philadelphia, 6, 73 - 86." type="journal article" year="1852">Dana, 1852</bibRefCitation>
</taxonomicName>
] (fide
<treatmentCitation id="0AF010BEC5070E59FB7BFF765A48FF09" author="Leene, J. E." box="[1217,1378,213,235]" page="70" pageId="4" pageNumber="205" year="1938">
<bibRefCitation id="EFC04B5EC5070E59FB7BFF765A11FF09" author="Leene, J. E." box="[1217,1339,213,235]" pageId="4" pageNumber="205" pagination="1 - 156" refId="ref15785" refString="Leene, J. E. (1938) The Decapoda Brachyura of the Siboga-Expedition. VII. Brachygnatha: Portunidae. Siboga Expeditie Monografie, 39 c 3, 1 - 156." type="journal article" year="1938">Leene 1938</bibRefCitation>
: 70
</treatmentCitation>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BEE36AFC5070E59FF2DFF555BC1FEE8" blockId="4.[151,1390,152,266]" box="[151,1259,244,266]" pageId="4" pageNumber="205">
<treatmentCitationGroup id="AB411181C5070E59FF2DFF555BC1FEE8" box="[151,1259,244,266]" pageId="4" pageNumber="205">
non
<taxonomicName id="4C514D2CC5070E59FF79FF545EA7FEE8" ID-CoL="3GXMJ" authorityName="A. Milne-Edwards" authorityYear="1867" box="[195,397,244,266]" class="Malacostraca" family="Portunidae" genus="Goniosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="225" phylum="Arthropoda" rank="species" species="hellerii">
<emphasis id="B925EABDC5070E59FF79FF545EA7FEE8" box="[195,397,244,266]" italics="true" pageId="4" pageNumber="205">Goniosoma Hellerii</emphasis>
</taxonomicName>
.—
<treatmentCitation id="0AF010BEC5070E59FE10FF545DA1FEE8" author="Klunzinger, C. B." box="[426,651,244,266]" page="367" pageId="4" pageNumber="205" year="1913">
<bibRefCitation id="EFC04B5EC5070E59FE10FF545D71FEE8" author="Klunzinger, C. B." box="[426,603,244,266]" pageId="4" pageNumber="205" pagination="97 - 402" refId="ref15555" refString="Klunzinger, C. B. (1913) Die Rundkrabben (Cyclometopa) des Roten Meeres. Abhandlungen der kaiserlich Leop. - Carol. Deutschen Akademie der Naturforscher Halle, 99 (2), 97 - 402, pls. 5 - 11." type="journal article" year="1913">Klunzinger 1913</bibRefCitation>
: 367
</treatmentCitation>
[=
<taxonomicName id="4C514D2CC5070E59FD14FF545CD6FEE8" ID-CoL="TPTR" authority="Dana, 1852" authorityName="Dana" authorityYear="1852" box="[686,1020,244,266]" class="Malacostraca" family="Portunidae" genus="Charybdis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis id="B925EABDC5070E59FD14FF545C55FEE8" box="[686,895,244,266]" italics="true" pageId="4" pageNumber="205">Charybdis orientalis</emphasis>
<bibRefCitation id="EFC04B5EC5070E59FC3FFF555CD6FEE8" author="Dana, J. D." box="[901,1020,244,266]" pageId="4" pageNumber="205" pagination="73 - 86" refId="ref12644" refString="Dana, J. D. (1852) Conspectus of the Crustacea of the Exploring Expedition under Capt. Wilkes, U. S. N., including the Crustacea Cancroidea Corystoidea. Proceedings of the Academy of Natural Sciences of Philadelphia, 6, 73 - 86." type="journal article" year="1852">Dana, 1852</bibRefCitation>
</taxonomicName>
] (fide
<treatmentCitation id="0AF010BEC5070E59FB85FF555BCAFEE8" author="Leene, J. E." box="[1087,1248,244,266]" page="70" pageId="4" pageNumber="205" year="1938">
<bibRefCitation id="EFC04B5EC5070E59FB85FF555B93FEE8" author="Leene, J. E." box="[1087,1209,244,266]" pageId="4" pageNumber="205" pagination="1 - 156" refId="ref15785" refString="Leene, J. E. (1938) The Decapoda Brachyura of the Siboga-Expedition. VII. Brachygnatha: Portunidae. Siboga Expeditie Monografie, 39 c 3, 1 - 156." type="journal article" year="1938">Leene 1938</bibRefCitation>
: 70
</treatmentCitation>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C34B6524C5070E59FF2DFE965BC3FAF9" pageId="4" pageNumber="205" type="materials_examined">
<paragraph id="8BEE36AFC5070E59FF2DFE965D97FE3D" blockId="4.[151,1437,310,2027]" pageId="4" pageNumber="205">
<emphasis id="B925EABDC5070E59FF2DFE965C49FEAD" bold="true" box="[151,867,310,335]" pageId="4" pageNumber="205">Material examined (directly or after new photographs).</emphasis>
Type material of
<taxonomicName id="4C514D2CC5070E59FBFEFE975A34FEAD" authorityName="A. Milne-Edwards" authorityYear="1867" box="[1092,1310,310,335]" class="Malacostraca" family="Portunidae" genus="Goniosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="hellerii">
<emphasis id="B925EABDC5070E59FBFEFE975A34FEAD" box="[1092,1310,310,335]" italics="true" pageId="4" pageNumber="205">Goniosoma hellerii</emphasis>
</taxonomicName>
A. Milne-
<bibRefCitation id="EFC04B5EC5070E59FF2DFEFA5E6FFE91" author="Milne-Edwards, A." box="[151,325,346,371]" pageId="4" pageNumber="205" pagination="263 - 288" refId="ref16786" refString="Milne-Edwards, A. (1867) Descriptions de quelques especes nouvelles de Crustaces Brachyures. Annales de la Societe entomologique de France, Series 4, 7, 263 - 288." type="journal article" year="1867">Edwards, 1867</bibRefCitation>
.
<collectingCountry id="F346763FC5070E59FEE0FEFB5D3BFE91" box="[346,529,346,371]" name="New Caledonia" pageId="4" pageNumber="205">New Caledonia</collectingCountry>
, no exact locality, in dry condition:
<typeStatus id="54EA880DC5070E59FC6AFEFA5B1DFE91" box="[976,1079,346,371]" pageId="4" pageNumber="205" type="lectotype">lectotype</typeStatus>
male, 60.0 x 85.0 mm,
<collectingCountry id="F346763FC5070E59FADDFEFB5E26FE75" name="New Caledonia" pageId="4" pageNumber="205">New Caledonia</collectingCountry>
, M. Aubry-Lecomte, MNHN-IU-2000-732 (= MNHN-B732);
<typeStatus id="54EA880DC5070E59FC76FEDE5B49FE75" box="[972,1123,382,407]" pageId="4" pageNumber="205" type="paralectotype">paralectotype</typeStatus>
male 49.0 x 72.0 mm,
<collectingCountry id="F346763FC5070E59FADDFEDF5E26FE59" name="New Caledonia" pageId="4" pageNumber="205">New Caledonia</collectingCountry>
, coll. M. Jouan, MNHN-IU-2000-761 (= MNHN-B761) [photographs provided by Danièle Guinot; specimens not directly examined by the authors].
</paragraph>
<paragraph id="8BEE36AFC5070E59FF7DFE4B5BF4FDA9" blockId="4.[151,1437,310,2027]" pageId="4" pageNumber="205">
Type material of
<taxonomicName id="4C514D2CC5070E59FE24FE4B5C46FDE1" authority="De Man, 1887" authorityName="De Man" authorityYear="1887" box="[414,876,491,515]" class="Malacostraca" family="Portunidae" genus="Goniosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="merguiense">
<emphasis id="B925EABDC5070E59FE24FE4B5D86FDE1" box="[414,684,491,515]" italics="true" pageId="4" pageNumber="205">Goniosoma merguiense</emphasis>
De Man, 1887
</taxonomicName>
. Republic of the Union of
<collectingCountry id="F346763FC5070E59FB70FE4B5A12FDE1" box="[1226,1336,491,515]" name="Myanmar" pageId="4" pageNumber="205">Myanmar</collectingCountry>
, Mergui Archipelago, 1886:
<specimenCount id="9D57FD26C5070E59FEC9FDAF5E94FDC5" box="[371,446,526,551]" pageId="4" pageNumber="205" type="male">2 male</specimenCount>
<typeStatus id="54EA880DC5070E59FE7FFDAF5D0CFDC5" box="[453,550,527,551]" pageId="4" pageNumber="205" type="syntype">syntypes</typeStatus>
(25.0 x 40.0 mm and 21.0 x 32.0 mm), leg. J. Anderson, RMNH.CRUS.D.1310 [photographs communicated by Charles Fransen; specimens not directly examined by the authors].
</paragraph>
<paragraph id="8BEE36AFC5070E59FF7DFDF75BF8FD55" blockId="4.[151,1437,310,2027]" pageId="4" pageNumber="205">
Type material of
<taxonomicName id="4C514D2CC5070E59FE36FDF75C08FD8D" authority="Miers, 1884" authorityName="Miers" authorityYear="1884" box="[396,802,598,623]" class="Malacostraca" family="Portunidae" genus="Goniosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="spiniferum">
<emphasis id="B925EABDC5070E59FE36FDF75DA7FD8D" box="[396,653,599,623]" italics="true" pageId="4" pageNumber="205">Goniosoma spiniferum</emphasis>
<bibRefCitation id="EFC04B5EC5070E59FD2CFDF65C08FD8D" author="Miers, E. J." box="[662,802,598,623]" pageId="4" pageNumber="205" pagination="513 - 575" refId="ref16627" refString="Miers, E. J. (1884) Crustacea. In: Report on the zoological collections in the Indopacific Ocean during the voyage of H. M. S. &quot; Alert &quot; 1881 - 82. Part II. Collections from the Western Indian Ocean. Printed by order of the trustees, London, pp. 513 - 575, pls. 46 - 51." type="book chapter" year="1884">Miers, 1884</bibRefCitation>
</taxonomicName>
.
<specimenCount id="9D57FD26C5070E59FC95FDF75C56FD8D" box="[815,892,598,623]" pageId="4" pageNumber="205" type="male">1 male</specimenCount>
<typeStatus id="54EA880DC5070E59FC3FFDF65CC0FD8D" box="[901,1002,598,623]" pageId="4" pageNumber="205" type="holotype">holotype</typeStatus>
, about
<date id="FFEF106FC5070E59FB84FDF75BBFFD8D" box="[1086,1173,599,623]" pageId="4" pageNumber="205" value="2017-10-13">
13 x
<quantity id="4CA99B4AC5070E59FBC3FDF75BBFFD8D" box="[1145,1173,599,623]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="4" pageNumber="205" unit="mm" value="17.0">17</quantity>
</date>
mm,
<collectingRegion id="4995F84DC5070E59FB63FDF65A4AFD8D" box="[1241,1376,598,623]" country="Australia" name="Queensland" pageId="4" pageNumber="205">Queensland</collectingRegion>
, Port Molle, between tide marks, Natural History Museum, NNM 1881.31 [photographs taken by Harry Taylor, NHM photo Unit and communicated by
<collectingRegion id="4995F84DC5070E59FDAFFD3E5D6DFD55" box="[533,583,670,695]" country="Cape Verde" name="Paul" pageId="4" pageNumber="205">Paul</collectingRegion>
F. Clark; specimen not directly examined by the authors].
</paragraph>
<paragraph id="8BEE36AFC5070E59FF7DFD625BC3FAF9" blockId="4.[151,1437,310,2027]" pageId="4" pageNumber="205">
<emphasis id="B925EABDC5070E59FF7DFD625E45FD39" box="[199,367,706,731]" italics="true" pageId="4" pageNumber="205">Other material</emphasis>
.
<materialsCitation id="3B393CF2C5070E59FEC4FD625B70FD1C" ID-GBIF-Occurrence="2234218885" collectingDate="2017-05-24" collectionCode="RBINS, INV" collectorName="C. E. Dessouassi" country="Benin" elevation="4" location="Ancien Pont" municipality="Channel of Cotonou" pageId="4" pageNumber="205" specimenCount="1" specimenCount-male="1">
<collectingCountry id="F346763FC5070E59FEC4FD625EEEFD39" box="[382,452,706,731]" name="Benin" pageId="4" pageNumber="205">Benin</collectingCountry>
,
<collectingMunicipality id="6B8AACD5C5070E59FE6AFD625D9FFD39" box="[464,693,706,731]" pageId="4" pageNumber="205">Channel of Cotonou</collectingMunicipality>
(
<location id="8E8E6074C5070E59FD7FFD625C7CFD39" LSID="urn:lsid:plazi:treatment:03F887B9C5000E4AFF2DFE295BA5FD82:8E8E6074C5070E59FD7FFD625C7CFD39" box="[709,854,706,731]" country="Benin" municipality="Channel of Cotonou" name="Ancien Pont" pageId="4" pageNumber="205">Ancien Pont</location>
),
<elevation id="007CD19CC5070E59FCD1FD635CB3FD38" box="[875,921,707,731]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="4" pageNumber="205" unit="m" value="4.0">
<quantity id="4CA99B4AC5070E59FCD1FD635CB3FD38" box="[875,921,707,731]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="4" pageNumber="205" unit="m" value="4.0">4 m</quantity>
</elevation>
,
<collectingMethod id="52104EB8C5070E59FC1CFD635CD4FD38" box="[934,1022,707,730]" pageId="4" pageNumber="205">cast net</collectingMethod>
,
<collectingDate id="EFABE987C5070E59FBB1FD635B57FD39" box="[1035,1149,707,731]" pageId="4" pageNumber="205" value="2017-05-24">
<date id="FFEF106FC5070E59FBB1FD635B57FD39" box="[1035,1149,707,731]" pageId="4" pageNumber="205" value="2017-05-24">24.v.2017</date>
</collectingDate>
:
<specimenCount id="9D57FD26C5070E59FB33FD635BFFFD39" box="[1161,1237,706,731]" pageId="4" pageNumber="205" type="male">1 male</specimenCount>
(pubescent, front with teratological anomalies), 43.5 x
<quantity id="4CA99B4AC5070E59FDFEFD465D86FD1C" box="[580,684,742,767]" metricMagnitude="-2" metricUnit="m" metricValue="6.67" pageId="4" pageNumber="205" unit="mm" value="66.7">66.7 mm</quantity>
, coll.
<collectorName id="26A45379C5070E59FD4EFD475C9DFD1D" box="[756,951,742,767]" pageId="4" pageNumber="205">C. E. Dessouassi</collectorName>
,
<collectionCode id="ED40AE6AC5070E59FC7FFD475B30FD1D" box="[965,1050,743,767]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="4" pageNumber="205" type="Museum">RBINS</collectionCode>
,
<collectionCode id="ED40AE6AC5070E59FB92FD475B70FD1C" box="[1064,1114,743,766]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13356" name="Inverness Museum and Art Gallery" pageId="4" pageNumber="205" type="Herbarium">INV</collectionCode>
</materialsCitation>
.
<date id="FFEF106FC5070E59FBDCFD465B94FD1D" box="[1126,1214,742,767]" pageId="4" pageNumber="205" value="09-01-15">150109</date>
;
<materialsCitation id="3B393CF2C5070E59FB71FD465ED5FCA4" ID-GBIF-Occurrence="2234218852" collectingDate="2017-05-24" collectionCode="RBINS, INV" collectorName="C. E. Dessouassi" country="Benin" elevation="4" location="Ancien Pont" municipality="Channel of Cotonou" pageId="4" pageNumber="205" specimenCount="2" specimenCount-male="2">
<collectingCountry id="F346763FC5070E59FB71FD465A3BFD1D" box="[1227,1297,742,767]" name="Benin" pageId="4" pageNumber="205">Benin</collectingCountry>
,
<collectingMunicipality id="6B8AACD5C5070E59FAA4FD465FD2FCC1" pageId="4" pageNumber="205">Channel of Cotonou</collectingMunicipality>
(
<location id="8E8E6074C5070E59FEBFFCAA5EBFFCC1" LSID="urn:lsid:plazi:treatment:03F887B9C5000E4AFF2DFE295BA5FD82:8E8E6074C5070E59FEBFFCAA5EBFFCC1" box="[261,405,778,803]" country="Benin" municipality="Channel of Cotonou" name="Ancien Pont" pageId="4" pageNumber="205">Ancien Pont</location>
),
<elevation id="007CD19CC5070E59FE13FCAB5EFFFCC0" box="[425,469,779,803]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="4" pageNumber="205" unit="m" value="4.0">
<quantity id="4CA99B4AC5070E59FE13FCAB5EFFFCC0" box="[425,469,779,803]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="4" pageNumber="205" unit="m" value="4.0">4 m</quantity>
</elevation>
,
<collectingMethod id="52104EB8C5070E59FE5AFCAA5D06FCC0" box="[480,556,778,803]" pageId="4" pageNumber="205">lift net</collectingMethod>
,
<collectingDate id="EFABE987C5070E59FD8DFCAB5D80FCC1" box="[567,682,779,803]" pageId="4" pageNumber="205" value="2017-05-24">
<date id="FFEF106FC5070E59FD8DFCAB5D80FCC1" box="[567,682,779,803]" pageId="4" pageNumber="205" value="2017-05-24">24.v.2017</date>
</collectingDate>
:
<specimenCount id="9D57FD26C5070E59FD0EFCAB5C23FCC1" box="[692,777,778,803]" pageId="4" pageNumber="205" type="male">2 males</specimenCount>
(not pubescent), 46.4 x
<quantity id="4CA99B4AC5070E59FBACFCAA5B50FCC0" box="[1046,1146,778,803]" metricMagnitude="-2" metricUnit="m" metricValue="6.85" pageId="4" pageNumber="205" unit="mm" value="68.5">68.5 mm</quantity>
and 46.6 x
<quantity id="4CA99B4AC5070E59FB41FCAA5A4AFCC0" box="[1275,1376,778,803]" metricMagnitude="-2" metricUnit="m" metricValue="6.87" pageId="4" pageNumber="205" unit="mm" value="68.7">68.7 mm</quantity>
, coll.
<collectorName id="26A45379C5070E59FF2DFC8F5E71FCA5" box="[151,347,814,839]" pageId="4" pageNumber="205">C. E. Dessouassi</collectorName>
,
<collectionCode id="ED40AE6AC5070E59FED0FC8F5E95FCA5" box="[362,447,815,839]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="4" pageNumber="205" type="Museum">RBINS</collectionCode>
,
<collectionCode id="ED40AE6AC5070E59FE77FC8F5ED5FCA4" box="[461,511,815,838]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13356" name="Inverness Museum and Art Gallery" pageId="4" pageNumber="205" type="Herbarium">INV</collectionCode>
</materialsCitation>
.
<date id="FFEF106FC5070E59FDB6FC8E5D49FCA5" box="[524,611,814,839]" pageId="4" pageNumber="205" value="10-01-15">150110</date>
;
<materialsCitation id="3B393CF2C5070E59FDCAFC8E5BCCFC88" ID-GBIF-Occurrence="2234218832" collectingDate="2017-06-19" collectionCode="RBINS, INV" collectorName="C. E. Dessouassi" country="Benin" elevation="4" location="Ancien Pont" municipality="Channel of Cotonou" pageId="4" pageNumber="205" specimenCount="2" specimenCount-male="2">
<collectingCountry id="F346763FC5070E59FDCAFC8E5D9CFCA5" box="[624,694,814,839]" name="Benin" pageId="4" pageNumber="205">Benin</collectingCountry>
,
<collectingMunicipality id="6B8AACD5C5070E59FD7EFC8E5C84FCA5" box="[708,942,814,839]" pageId="4" pageNumber="205">Channel of Cotonou</collectingMunicipality>
(
<location id="8E8E6074C5070E59FC04FC8E5B78FCA5" LSID="urn:lsid:plazi:treatment:03F887B9C5000E4AFF2DFE295BA5FD82:8E8E6074C5070E59FC04FC8E5B78FCA5" box="[958,1106,814,839]" country="Benin" municipality="Channel of Cotonou" name="Ancien Pont" pageId="4" pageNumber="205">Ancien Pont</location>
),
<elevation id="007CD19CC5070E59FBD3FC8F5BB3FCA4" box="[1129,1177,815,839]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="4" pageNumber="205" unit="m" value="4.0">
<quantity id="4CA99B4AC5070E59FBD3FC8F5BB3FCA4" box="[1129,1177,815,839]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="4" pageNumber="205" unit="m" value="4.0">4 m</quantity>
</elevation>
,
<collectingMethod id="52104EB8C5070E59FB1DFC8E5BDCFCA4" box="[1191,1270,814,839]" pageId="4" pageNumber="205">lift net</collectingMethod>
,
<collectingDate id="EFABE987C5070E59FABFFC8E5AABFCA5" box="[1285,1409,814,839]" pageId="4" pageNumber="205" value="2017-06-19">
<date id="FFEF106FC5070E59FABFFC8E5AABFCA5" box="[1285,1409,814,839]" pageId="4" pageNumber="205" value="2017-06-19">19.vi.2017</date>
</collectingDate>
:
<specimenCount id="9D57FD26C5070E59FA34FC8F5FF2FC89" pageId="4" pageNumber="205" type="male">2 males</specimenCount>
(not pubescent), 49.2 x 76.0 mm and 37.0 x 56.0 mm, coll.
<collectorName id="26A45379C5070E59FC3FFCF35B6CFC89" box="[901,1094,850,875]" pageId="4" pageNumber="205">C. E. Dessouassi</collectorName>
,
<collectionCode id="ED40AE6AC5070E59FBE8FCF35B8DFC89" box="[1106,1191,851,875]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="4" pageNumber="205" type="Museum">RBINS</collectionCode>
,
<collectionCode id="ED40AE6AC5070E59FB0EFCF35BCCFC88" box="[1204,1254,851,874]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13356" name="Inverness Museum and Art Gallery" pageId="4" pageNumber="205" type="Herbarium">INV</collectionCode>
</materialsCitation>
.
<date id="FFEF106FC5070E59FB4BFCF25A6CFC89" box="[1265,1350,850,875]" pageId="4" pageNumber="205" value="11-01-15">150111</date>
;
<materialsCitation id="3B393CF2C5070E59FAE8FCF25D8BFC50" ID-GBIF-Occurrence="2234218859" collectingDate="2017-05-24" collectionCode="RBINS, INV" collectorName="C. E. Dessouassi" country="Benin" elevation="4" location="Ancien Pont" municipality="Channel of Cotonou" pageId="4" pageNumber="205" specimenCount="2" specimenCount-male="2">
<collectingCountry id="F346763FC5070E59FAE8FCF25AB2FC89" box="[1362,1432,850,875]" name="Benin" pageId="4" pageNumber="205">Benin</collectingCountry>
,
<collectingMunicipality id="6B8AACD5C5070E59FF2DFCD65E57FC6D" box="[151,381,886,911]" pageId="4" pageNumber="205">Channel of Cotonou</collectingMunicipality>
(
<location id="8E8E6074C5070E59FE37FCD65D35FC6D" LSID="urn:lsid:plazi:treatment:03F887B9C5000E4AFF2DFE295BA5FD82:8E8E6074C5070E59FE37FCD65D35FC6D" box="[397,543,886,911]" country="Benin" municipality="Channel of Cotonou" name="Ancien Pont" pageId="4" pageNumber="205">Ancien Pont</location>
),
<elevation id="007CD19CC5070E59FD8EFCD75D48FC6C" box="[564,610,887,911]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="4" pageNumber="205" unit="m" value="4.0">
<quantity id="4CA99B4AC5070E59FD8EFCD75D48FC6C" box="[564,610,887,911]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="4" pageNumber="205" unit="m" value="4.0">4 m</quantity>
</elevation>
,
<collectingMethod id="52104EB8C5070E59FDD5FCD65D97FC6C" box="[623,701,886,911]" pageId="4" pageNumber="205">lift net</collectingMethod>
,
<collectingDate id="EFABE987C5070E59FD70FCD75C17FC6D" box="[714,829,887,911]" pageId="4" pageNumber="205" value="2017-05-24">
<date id="FFEF106FC5070E59FD70FCD75C17FC6D" box="[714,829,887,911]" pageId="4" pageNumber="205" value="2017-05-24">24.v.2017</date>
</collectingDate>
:
<specimenCount id="9D57FD26C5070E59FCF2FCD75C8AFC6D" box="[840,928,886,911]" pageId="4" pageNumber="205" type="male">2 males</specimenCount>
(not pubescent), 38.0 x 58.0 mm and 34.5 x
<quantity id="4CA99B4AC5070E59FF2DFC3A5FD7FC50" box="[151,253,922,947]" metricMagnitude="-2" metricUnit="m" metricValue="5.24" pageId="4" pageNumber="205" unit="mm" value="52.4">52.4 mm</quantity>
, coll.
<collectorName id="26A45379C5070E59FEF8FC3B5D28FC51" box="[322,514,922,947]" pageId="4" pageNumber="205">C. E. Dessouassi</collectorName>
,
<collectionCode id="ED40AE6AC5070E59FDB4FC3B5D49FC51" box="[526,611,923,947]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="4" pageNumber="205" type="Museum">RBINS</collectionCode>
,
<collectionCode id="ED40AE6AC5070E59FDD5FC3B5D8BFC50" box="[623,673,923,946]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13356" name="Inverness Museum and Art Gallery" pageId="4" pageNumber="205" type="Herbarium">INV</collectionCode>
</materialsCitation>
.
<date id="FFEF106FC5070E59FD16FC3A5C28FC51" box="[684,770,922,947]" pageId="4" pageNumber="205" value="12-01-15">150112</date>
; 1 ovigerous female (not pubescent), 31.3 x 48.0 mm, coll.
<collectorName id="26A45379C5070E59FF2DFC1F5E76FC35" box="[151,348,958,983]" pageId="4" pageNumber="205">C. E. Dessouassi</collectorName>
, RBINS, INV.
<date id="FFEF106FC5070E59FDB4FC1E5D4FFC35" box="[526,613,958,983]" pageId="4" pageNumber="205" value="13-01-15">150113</date>
;
<materialsCitation id="3B393CF2C5070E59FDC9FC1E5CA5FC18" ID-GBIF-Occurrence="2234218895" collectingDate="2017-05-24" collectionCode="RBINS, INV" collectorName="C. E. Dessouassi" country="Benin" elevation="5" location="Agbato" municipality="Nokoue Lake" pageId="4" pageNumber="205" specimenCount="1" specimenCount-female="1">
<collectingCountry id="F346763FC5070E59FDC9FC1E5D93FC35" box="[627,697,958,983]" name="Benin" pageId="4" pageNumber="205">Benin</collectingCountry>
,
<collectingMunicipality id="6B8AACD5C5070E59FD7DFC1E5C48FC35" box="[711,866,958,983]" pageId="4" pageNumber="205">Nokoué Lake</collectingMunicipality>
(
<location id="8E8E6074C5070E59FCCEFC1E5CE6FC35" LSID="urn:lsid:plazi:treatment:03F887B9C5000E4AFF2DFE295BA5FD82:8E8E6074C5070E59FCCEFC1E5CE6FC35" box="[884,972,958,983]" country="Benin" municipality="Nokoue Lake" name="Agbato" pageId="4" pageNumber="205">Agbato</location>
),
<elevation id="007CD19CC5070E59FC58FC1E5B39FC34" box="[994,1043,958,983]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="205" unit="m" value="5.0">
<quantity id="4CA99B4AC5070E59FC58FC1E5B39FC34" box="[994,1043,958,983]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="205" unit="m" value="5.0">5 m</quantity>
</elevation>
,
<collectingMethod id="52104EB8C5070E59FB98FC1E5B58FC34" box="[1058,1138,958,983]" pageId="4" pageNumber="205">lift net</collectingMethod>
,
<collectingDate id="EFABE987C5070E59FB3AFC1F5BD9FC35" box="[1152,1267,959,983]" pageId="4" pageNumber="205" value="2017-05-24">
<date id="FFEF106FC5070E59FB3AFC1F5BD9FC35" box="[1152,1267,959,983]" pageId="4" pageNumber="205" value="2017-05-24">24.v.2017</date>
</collectingDate>
:
<specimenCount id="9D57FD26C5070E59FABAFC1F5A4FFC35" box="[1280,1381,958,983]" pageId="4" pageNumber="205" type="female">1 female</specimenCount>
(not pubescent), 34.2 x 53.0 mm, coll.
<collectorName id="26A45379C5070E59FD9CFC435DC1FC19" box="[550,747,994,1019]" pageId="4" pageNumber="205">C. E. Dessouassi</collectorName>
,
<collectionCode id="ED40AE6AC5070E59FD40FC435C65FC19" box="[762,847,995,1019]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="4" pageNumber="205" type="Museum">RBINS</collectionCode>
,
<collectionCode id="ED40AE6AC5070E59FCE7FC435CA5FC18" box="[861,911,995,1018]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13356" name="Inverness Museum and Art Gallery" pageId="4" pageNumber="205" type="Herbarium">INV</collectionCode>
</materialsCitation>
.
<date id="FFEF106FC5070E59FC26FC425CD9FC19" box="[924,1011,994,1019]" pageId="4" pageNumber="205" value="14-01-15">150114</date>
;
<materialsCitation id="3B393CF2C5070E59FBBBFC435B02FBFC" ID-GBIF-Occurrence="2234218826" collectingDate="1966-05-20" collectionCode="RBINS, INV" collectorName="Fierlant" country="New Caledonia" location="Noumea" pageId="4" pageNumber="205" specimenCount="1" specimenCount-male="1">
<collectingCountry id="F346763FC5070E59FBBBFC435B9FFC19" box="[1025,1205,994,1019]" name="New Caledonia" pageId="4" pageNumber="205">New Caledonia</collectingCountry>
,
<location id="8E8E6074C5070E59FB78FC435A0BFC18" LSID="urn:lsid:plazi:treatment:03F887B9C5000E4AFF2DFE295BA5FD82:8E8E6074C5070E59FB78FC435A0BFC18" box="[1218,1313,995,1018]" country="New Caledonia" name="Noumea" pageId="4" pageNumber="205">Noumea</location>
aquarium,
<collectingDate id="EFABE987C5070E59FF2DFBA65E23FBFD" box="[151,265,1030,1055]" pageId="4" pageNumber="205" value="1966-05-20">
<date id="FFEF106FC5070E59FF2DFBA65E23FBFD" box="[151,265,1030,1055]" pageId="4" pageNumber="205" value="1966-05-20">20.v.1966</date>
</collectingDate>
:
<specimenCount id="9D57FD26C5070E59FEACFBA75E4EFBFD" box="[278,356,1030,1055]" pageId="4" pageNumber="205" type="male">1 male</specimenCount>
(not pubescent), 47.0 x 68.0 mm, leg.
<collectorName id="26A45379C5070E59FC91FBA65CAFFBFD" box="[811,901,1030,1055]" pageId="4" pageNumber="205">Fierlant</collectorName>
,
<collectionCode id="ED40AE6AC5070E59FC28FBA75CCDFBFD" box="[914,999,1031,1055]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="4" pageNumber="205" type="Museum">RBINS</collectionCode>
,
<collectionCode id="ED40AE6AC5070E59FC4CFBA75B02FBFC" box="[1014,1064,1031,1054]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13356" name="Inverness Museum and Art Gallery" pageId="4" pageNumber="205" type="Herbarium">INV</collectionCode>
</materialsCitation>
. 129463;
<materialsCitation id="3B393CF2C5070E59FB20FBA65C0EFB84" ID-GBIF-Occurrence="2234218802" collectingDate="1983-04-14" collectionCode="RBINS, INV" collectorName="R. De Blauwe" country="Australia" location="Shoal Point" pageId="4" pageNumber="205" specimenCount="3" specimenCount-male="3" stateProvince="Queensland">
<collectingCountry id="F346763FC5070E59FB20FBA65A29FBFD" box="[1178,1283,1030,1055]" name="Australia" pageId="4" pageNumber="205">Australia</collectingCountry>
,
<collectingRegion id="4995F84DC5070E59FAABFBA65AB2FBFD" box="[1297,1432,1030,1055]" country="Australia" name="Queensland" pageId="4" pageNumber="205">Queensland</collectingRegion>
,
<location id="8E8E6074C5070E59FF2DFB8A5E31FBA1" LSID="urn:lsid:plazi:treatment:03F887B9C5000E4AFF2DFE295BA5FD82:8E8E6074C5070E59FF2DFB8A5E31FBA1" box="[151,283,1066,1091]" country="Australia" name="Shoal Point" pageId="4" pageNumber="205" stateProvince="Queensland">Shoal Point</location>
, under stones, at low tide,
<collectingDate id="EFABE987C5070E59FDFBFB8A5D91FBA1" box="[577,699,1066,1091]" pageId="4" pageNumber="205" value="1983-04-14">
<date id="FFEF106FC5070E59FDFBFB8A5D91FBA1" box="[577,699,1066,1091]" pageId="4" pageNumber="205" value="1983-04-14">14.iv.1983</date>
</collectingDate>
:
<specimenCount id="9D57FD26C5070E59FD7FFB8B5C30FBA1" box="[709,794,1066,1091]" pageId="4" pageNumber="205" type="male">3 males</specimenCount>
(two strongly pubescent; smallest one weakly pubescent) (largest 37.0 x 56.0 mm), leg.
<collectorName id="26A45379C5070E59FE5DFBEF5DADFB85" box="[487,647,1102,1127]" pageId="4" pageNumber="205">R. De Blauwe</collectorName>
,
<collectionCode id="ED40AE6AC5070E59FD28FBEF5DCDFB85" box="[658,743,1103,1127]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="4" pageNumber="205" type="Museum">RBINS</collectionCode>
,
<collectionCode id="ED40AE6AC5070E59FD48FBEF5C0EFB84" box="[754,804,1103,1126]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13356" name="Inverness Museum and Art Gallery" pageId="4" pageNumber="205" type="Herbarium">INV</collectionCode>
</materialsCitation>
. 129466;
<materialsCitation id="3B393CF2C5070E59FC2AFBEE5B30FB4C" ID-GBIF-Occurrence="2234218806" collectingDate="1983-05-26" collectionCode="RBINS, INV" collectorName="R. De Blauwe" country="Australia" location="Shoal Point" municipality="Mackay" pageId="4" pageNumber="205" specimenCount="3" specimenCount-male="3" stateProvince="Queensland">
<collectingCountry id="F346763FC5070E59FC2AFBEE5CD0FB85" box="[912,1018,1102,1127]" name="Australia" pageId="4" pageNumber="205">Australia</collectingCountry>
,
<collectingRegion id="4995F84DC5070E59FBBFFBEE5BA6FB85" box="[1029,1164,1102,1127]" country="Australia" name="Queensland" pageId="4" pageNumber="205">Queensland</collectingRegion>
, north of
<collectingMunicipality id="6B8AACD5C5070E59FB4CFBEE5A7BFB85" box="[1270,1361,1102,1127]" pageId="4" pageNumber="205">Mackay</collectingMunicipality>
,
<location id="8E8E6074C5070E59FAE6FBEE5FFFFB69" LSID="urn:lsid:plazi:treatment:03F887B9C5000E4AFF2DFE295BA5FD82:8E8E6074C5070E59FAE6FBEE5FFFFB69" country="Australia" municipality="Mackay" name="Shoal Point" pageId="4" pageNumber="205" stateProvince="Queensland">Shoal Point</location>
, under stones, at low tide,
<collectingDate id="EFABE987C5070E59FDB9FBD25D5CFB69" box="[515,630,1138,1163]" pageId="4" pageNumber="205" value="1983-05-26">
<date id="FFEF106FC5070E59FDB9FBD25D5CFB69" box="[515,630,1138,1163]" pageId="4" pageNumber="205" value="1983-05-26">26.v.1983</date>
</collectingDate>
:
<specimenCount id="9D57FD26C5070E59FD3BFBD35DF2FB69" box="[641,728,1138,1163]" pageId="4" pageNumber="205" type="male">3 males</specimenCount>
(largest: 41.0 x 63.0 mm) and 1 immature female (30.0 x 47.0 mm) [all specimens strongly pubescent], leg.
<collectorName id="26A45379C5070E59FD05FB375C44FB4D" box="[703,878,1174,1199]" pageId="4" pageNumber="205">R. De Blauwe</collectorName>
,
<collectionCode id="ED40AE6AC5070E59FC3AFB375CFFFB4D" box="[896,981,1175,1199]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="4" pageNumber="205" type="Museum">RBINS</collectionCode>
,
<collectionCode id="ED40AE6AC5070E59FC52FB375B30FB4C" box="[1000,1050,1175,1198]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13356" name="Inverness Museum and Art Gallery" pageId="4" pageNumber="205" type="Herbarium">INV</collectionCode>
</materialsCitation>
. 129467;
<materialsCitation id="3B393CF2C5070E59FB2FFB365FE3FB14" ID-GBIF-Occurrence="2234218812" collectionCode="RBINS, INV" collectorName="B. Kaspiew" country="Australia" location="Brampton Island" pageId="4" pageNumber="205" specimenCount="1" specimenCount-male="1" stateProvince="Queensland">
<collectingCountry id="F346763FC5070E59FB2FFB365BD4FB4D" box="[1173,1278,1174,1199]" name="Australia" pageId="4" pageNumber="205">Australia</collectingCountry>
,
<collectingRegion id="4995F84DC5070E59FAABFB365AB2FB4D" box="[1297,1432,1174,1199]" country="Australia" name="Queensland" pageId="4" pageNumber="205">Queensland</collectingRegion>
,
<location id="8E8E6074C5070E59FF2DFB1B5E7CFB31" LSID="urn:lsid:plazi:treatment:03F887B9C5000E4AFF2DFE295BA5FD82:8E8E6074C5070E59FF2DFB1B5E7CFB31" box="[151,342,1210,1235]" country="Australia" name="Brampton Island" pageId="4" pageNumber="205" stateProvince="Queensland">Brampton Island</location>
, mangrove shore, no date:
<specimenCount id="9D57FD26C5070E59FD31FB1B5DF2FB31" box="[651,728,1210,1235]" pageId="4" pageNumber="205" type="male">1 male</specimenCount>
(pubescent, 27.0 x 43.0 mm), leg.
<collectorName id="26A45379C5070E59FBD3FB1B5BC4FB31" box="[1129,1262,1210,1235]" pageId="4" pageNumber="205">B. Kaspiew</collectorName>
, 1953,
<collectionCode id="ED40AE6AC5070E59FAF9FB1B5AB2FB31" box="[1347,1432,1211,1235]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="4" pageNumber="205" type="Museum">RBINS</collectionCode>
,
<collectionCode id="ED40AE6AC5070E59FF2DFB7F5FE3FB14" box="[151,201,1247,1270]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13356" name="Inverness Museum and Art Gallery" pageId="4" pageNumber="205" type="Herbarium">INV</collectionCode>
</materialsCitation>
. 129465;
<materialsCitation id="3B393CF2C5070E59FE85FB7E5BA9FAF8" ID-GBIF-Occurrence="2234218853" collectingDate="1976-03-04" collectionCode="RBINS, INV" collectorName="J. Van Goethem" country="Philippines" location="Langsheen National Park" pageId="4" pageNumber="205" specimenCount="5" specimenCount-male="5" stateProvince="Manila">
<collectingCountry id="F346763FC5070E59FE85FB7E5EEAFB15" box="[319,448,1246,1271]" name="Philippines" pageId="4" pageNumber="205">Philippines</collectingCountry>
,
<collectingRegion id="4995F84DC5070E59FE75FB7E5D03FB15" box="[463,553,1246,1271]" country="Philippines" name="Manila" pageId="4" pageNumber="205">Manilla</collectingRegion>
,
<location id="8E8E6074C5070E59FD83FB7E5C4AFB15" LSID="urn:lsid:plazi:treatment:03F887B9C5000E4AFF2DFE295BA5FD82:8E8E6074C5070E59FD83FB7E5C4AFB15" box="[569,864,1246,1271]" country="Philippines" name="Langsheen National Park" pageId="4" pageNumber="205" stateProvince="Manila">Langsheen National Park</location>
, rocks at the coast,
<collectingDate id="EFABE987C5070E59FBE8FB7E5BEBFB15" box="[1106,1217,1246,1271]" pageId="4" pageNumber="205" value="1976-03-04">
<date id="FFEF106FC5070E59FBE8FB7E5BEBFB15" box="[1106,1217,1246,1271]" pageId="4" pageNumber="205" value="1976-03-04">4.iii.1976</date>
</collectingDate>
:
<specimenCount id="9D57FD26C5070E59FB6BFB7E5A01FB15" box="[1233,1323,1246,1271]" pageId="4" pageNumber="205" type="male">5 males</specimenCount>
(scarcely pubescent and not pubescent; largest 30.0 x 46.0 mm), leg.
<collectorName id="26A45379C5070E59FC8AFAA35CCFFAF9" box="[816,997,1282,1307]" pageId="4" pageNumber="205">J. Van Goethem</collectorName>
,
<collectionCode id="ED40AE6AC5070E59FC4AFAA35B6FFAF9" box="[1008,1093,1283,1307]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="4" pageNumber="205" type="Museum">RBINS</collectionCode>
,
<collectionCode id="ED40AE6AC5070E59FBEBFAA35BA9FAF8" box="[1105,1155,1283,1306]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13356" name="Inverness Museum and Art Gallery" pageId="4" pageNumber="205" type="Herbarium">INV</collectionCode>
</materialsCitation>
. 129464.
</paragraph>
</subSubSection>
<subSubSection id="C34B6524C5070E4BFF7DFA865D43F806" lastPageId="22" lastPageNumber="223" pageId="4" pageNumber="205" type="description">
<paragraph id="8BEE36AFC5070E59FF7DFA865BB0F9D9" blockId="4.[151,1437,310,2027]" pageId="4" pageNumber="205">
<emphasis id="B925EABDC5070E59FF7DFA865E73FADD" bold="true" box="[199,345,1318,1343]" pageId="4" pageNumber="205">Description.</emphasis>
<emphasis id="B925EABDC5070E59FEDAFA875EE4FADC" box="[352,462,1319,1342]" italics="true" pageId="4" pageNumber="205">Carapace</emphasis>
strongly pubescent, sparsely pubescent or glabrous, with a pair of short frontal ridges (sometimes hard to see), with a pair of nearly straight anterior protogastric ridges, with a continuous nearly straight mesogastric ridge, with shorter straight metagastric ridge (continuous or with short median gap), laterally followed (after a gap) by pair of broadly curved epibranchial ridges; no ridges on posterior half of carapace. Frontal region with 8 well developed teeth separated by deep incisions (3 pairs of true frontal teeth + 1 pair of teeth forming the inner border of orbit), subequal in length (very blunt in small specimens, blunt or sharp in large specimens). Anterolateral margin with 6 sharp teeth (the 5 first ones are subequal, the sixth one is subequal to the others or a bit longer). Junction between posterior margin and posterolateral margins arched (not angular).
</paragraph>
<paragraph id="8BEE36AFC5070E59FF7DF9E65D8FF9BD" blockId="4.[151,1437,310,2027]" box="[199,677,1606,1631]" pageId="4" pageNumber="205">
<emphasis id="B925EABDC5070E59FF7DF9E65E95F9BD" box="[199,447,1606,1631]" italics="true" pageId="4" pageNumber="205">Basal antennal article</emphasis>
with granular ridge.
</paragraph>
<paragraph id="8BEE36AFC5070E59FF7DF9CA5A78F89D" blockId="4.[151,1437,310,2027]" pageId="4" pageNumber="205">
<emphasis id="B925EABDC5070E59FF7DF9CA5E1DF961" box="[199,311,1642,1667]" italics="true" pageId="4" pageNumber="205">Chelipeds</emphasis>
: merus with 3 strong anterior spines, posterior border toothless; carpus with strong spine on distomedial corner (this spine is posteriorly followed by a ridge running all along the carpus), with 3 lateral spines, of which only the two uppermost are visible in dorsal view, and of which the uppermost and the lowermost are posteriorly followed by a long ridge; propodus with 5 spines on its upper half: one spine in proximolateral position, 2 spines on the middle of propodus (1 dorsolateral and 1 dorsomedial which is posteriorly followed by carina), 2 distal spines (1 dorsolateral that is sometimes reduced to an angular protrusion and 1 dorsomedial which is posteriorly followed by carina), with 3 lateral carinae, with 1 medial carina terminated by a granule; tip of the various cheliped spines brown (except sometimes dorsolateral distal spine of propodus, when it is obsolete).
</paragraph>
<paragraph id="8BEE36AFC5070E59FF7DF82A5CADF809" blockId="4.[151,1437,310,2027]" pageId="4" pageNumber="205">
<emphasis id="B925EABDC5070E59FF7DF82A5E65F840" box="[199,335,1930,1955]" italics="true" pageId="4" pageNumber="205">Pereiopod 5</emphasis>
: posterior border of merus with subdistal spine; posterior border of carpus with distal spine (better seen in ventral view) [this character is very unusual in the genus
<taxonomicName id="4C514D2CC5070E59FCCAF80E5CCFF825" authorityName="Zanclea Cocco" authorityYear="1832" box="[880,997,1966,1991]" class="Malacostraca" family="Portunidae" genus="Charybdis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="205" phylum="Arthropoda" rank="genus">
<emphasis id="B925EABDC5070E59FCCAF80E5CCFF825" box="[880,997,1966,1991]" italics="true" pageId="4" pageNumber="205">Charybdis</emphasis>
</taxonomicName>
]; posterior border of propodus with 5 13 denticles (sometimes up to 18 according to
<bibRefCitation id="EFC04B5EC5070E59FD1BF8725C56F809" author="Negri Pereira, M." box="[673,892,2002,2027]" pageId="4" pageNumber="205" refId="ref17215" refString="Negri Pereira, M. (2016) Registro de Charybdis hellerii (Milne Edwards, 1867) em aguas do litoral brasileiro (Decapoda: Portunidae). Tese apresentada a Faculdade de Filosofia, Ciencias e Letras de Ribeirao Preto da USP, como parte das exigencias para a obtencao do titulo de Doutor em Ciencias Area: Biologia Comparada, viii + 164 pp." type="book" year="2016">Negri Pereira 2016</bibRefCitation>
).
</paragraph>
<caption id="DF2E6627C5060E58FF2DF8275E88F838" httpUri="https://zenodo.org/record/2624787/files/figure.png" pageId="5" pageNumber="206" startId="5.[151,250,1927,1949]" targetBox="[195,1377,193,1901]" targetPageId="5">
<paragraph id="8BEE36AFC5060E58FF2DF8275E88F838" blockId="5.[151,1436,1927,2010]" pageId="5" pageNumber="206">
<emphasis id="B925EABDC5060E58FF2DF8275E3EF87E" bold="true" box="[151,276,1927,1949]" pageId="5" pageNumber="206">FIGURE 1.</emphasis>
<taxonomicName id="4C514D2CC5060E58FEA1F8275DC1F87F" authority="(A. Milne-Edwards, 1867)" authorityName="A. Milne-Edwards" authorityYear="1867" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1867" box="[283,747,1927,1949]" class="Malacostraca" family="Portunidae" genus="Charybdis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="206" phylum="Arthropoda" rank="species" species="hellerii">
<emphasis id="B925EABDC5060E58FEA1F8275EFEF87F" box="[283,468,1927,1949]" italics="true" pageId="5" pageNumber="206">Charybdis hellerii</emphasis>
(A. Milne-Edwards, 1867)
</taxonomicName>
, New Caledonia. A. Lectotype, male, 60.0 x 85.0 mm, MNHN-IU- 2000-732 (= MNHN-B732). B, C. Paralectotype, male 49.0 x 72.0 mm, MNHN-IU-2000-761 (= MNHN-B761). A, B. Dorsal habitus. C. Frontal region.
</paragraph>
</caption>
<caption id="DF2E6627C5050E5BFF2DF8DF5B22F830" httpUri="https://zenodo.org/record/2624789/files/figure.png" pageId="6" pageNumber="207" startId="6.[151,250,1919,1941]" targetBox="[179,1408,193,1898]" targetPageId="6">
<paragraph id="8BEE36AFC5050E5BFF2DF8DF5B22F830" blockId="6.[151,1436,1919,2002]" pageId="6" pageNumber="207">
<emphasis id="B925EABDC5050E5BFF2DF8DF5E38F876" bold="true" box="[151,274,1919,1941]" pageId="6" pageNumber="207">FIGURE 2.</emphasis>
<taxonomicName id="4C514D2CC5050E5BFEA2F8DF5DC9F877" authority="(A. Milne-Edwards, 1867)" authorityName="A. Milne-Edwards" authorityYear="1867" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1867" box="[280,739,1919,1941]" class="Malacostraca" family="Portunidae" genus="Charybdis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="207" phylum="Arthropoda" rank="species" species="hellerii">
<emphasis id="B925EABDC5050E5BFEA2F8DF5EFAF877" box="[280,464,1919,1941]" italics="true" pageId="6" pageNumber="207">Charybdis hellerii</emphasis>
(A. Milne-Edwards, 1867)
</taxonomicName>
, New Caledonia. A. Lectotype, male, 60 x 85 mm, MNHN-IU-2000- 732 (= MNHN-B732). B, C. Paralectotype, male 49.0 x 72.0 mm, MNHN-IU-2000-761 (= MNHN-B761). A, B. Ventral habitus. C. Carpus and extremity of adjacent articles of left pereiopod 5 (ventral view).
</paragraph>
</caption>
<caption id="DF2E6627C5040E5AFF2DF8EF5BB7F840" httpUri="https://zenodo.org/record/2624791/files/figure.png" pageId="7" pageNumber="208" startId="7.[151,250,1871,1893]" targetBox="[151,1437,193,1850]" targetPageId="7">
<paragraph id="8BEE36AFC5040E5AFF2DF8EF5BB7F840" blockId="7.[151,1436,1871,1954]" pageId="7" pageNumber="208">
<emphasis id="B925EABDC5040E5AFF2DF8EF5E33F886" bold="true" box="[151,281,1871,1893]" pageId="7" pageNumber="208">FIGURE 3.</emphasis>
<taxonomicName id="4C514D2CC5040E5AFE9CF8EF5C25F887" authority="(A. Milne-Edwards, 1867)" authorityName="A. Milne-Edwards" authorityYear="1867" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1867" box="[294,783,1871,1893]" class="Malacostraca" family="Portunidae" genus="Charybdis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="hellerii">
<emphasis id="B925EABDC5040E5AFE9CF8EF5ECFF887" box="[294,485,1871,1893]" italics="true" pageId="7" pageNumber="208">Charybdis hellerii</emphasis>
(A. Milne-Edwards, 1867)
</taxonomicName>
, males, New Caledonia, left gonopod1s (rehydrated). A, B. Lectotype, male 60.0 x 85.0 mm, MNHN-IU-2000-732 (= MNHN-B732). C, D, E. Paralectotype, male 49.0 x 72.0 mm, MNHN-IU-2000-761 (=MNHN-B761). A, C, medial view. B, D, external view. E. Tip, external view.
</paragraph>
</caption>
<caption id="DF2E6627C50B0E55FF2DF8365EC0F808" httpUri="https://zenodo.org/record/2624793/files/figure.png" pageId="8" pageNumber="209" startId="8.[151,250,1942,1964]" targetBox="[169,1454,193,1907]" targetPageId="8">
<paragraph id="8BEE36AFC50B0E55FF2DF8365EC0F808" blockId="8.[151,1436,1942,2026]" pageId="8" pageNumber="209">
<emphasis id="B925EABDC50B0E55FF2DF8365E38F849" bold="true" box="[151,274,1942,1964]" pageId="8" pageNumber="209">FIGURE 4.</emphasis>
<taxonomicName id="4C514D2CC50B0E55FEA2F8365DCEF84E" authority="(A. Milne-Edwards, 1867)" authorityName="A. Milne-Edwards" authorityYear="1867" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1867" box="[280,740,1942,1964]" class="Malacostraca" family="Portunidae" genus="Charybdis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="209" phylum="Arthropoda" rank="species" species="hellerii">
<emphasis id="B925EABDC50B0E55FEA2F8365EFBF84E" box="[280,465,1942,1964]" italics="true" pageId="8" pageNumber="209">Charybdis hellerii</emphasis>
(A. Milne-Edwards, 1867)
</taxonomicName>
, male holotype of
<taxonomicName id="4C514D2CC50B0E55FC25F8375A22F84E" authority="Miers, 1884" authorityName="Miers" authorityYear="1884" box="[927,1288,1942,1964]" class="Malacostraca" family="Portunidae" genus="Goniosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="209" phylum="Arthropoda" rank="species" species="spiniferum">
<emphasis id="B925EABDC50B0E55FC25F8375BADF84E" box="[927,1159,1942,1964]" italics="true" pageId="8" pageNumber="209">Goniosoma spiniferum</emphasis>
Miers, 1884
</taxonomicName>
, about 13 x 17 mm, Queensland, Port Molle, NNM 1881.31. A. Dorsal habitus. B. Ventral habitus. C. Right pereiopod 5 (dorsal view). D. Right pereiopod 5 (ventral view).
</paragraph>
</caption>
<caption id="DF2E6627C50A0E54FF2DF8F95B43F86C" httpUri="https://zenodo.org/record/2624795/files/figure.png" pageId="9" pageNumber="210" startId="9.[151,250,1881,1903]" targetBox="[151,1435,193,1860]" targetPageId="9">
<paragraph id="8BEE36AFC50A0E54FF2DF8F95B43F86C" blockId="9.[151,1436,1881,1934]" pageId="9" pageNumber="210">
<emphasis id="B925EABDC50A0E54FF2DF8F95E38F88D" bold="true" box="[151,274,1881,1903]" pageId="9" pageNumber="210">FIGURE 5.</emphasis>
<taxonomicName id="4C514D2CC50A0E54FEA3F8F95DCCF88D" authority="(A. Milne-Edwards, 1867)" authorityName="A. Milne-Edwards" authorityYear="1867" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1867" box="[281,742,1881,1903]" class="Malacostraca" family="Portunidae" genus="Charybdis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="hellerii">
<emphasis id="B925EABDC50A0E54FEA3F8F95EFBF88D" box="[281,465,1881,1903]" italics="true" pageId="9" pageNumber="210">Charybdis hellerii</emphasis>
(A. Milne-Edwards, 1867)
</taxonomicName>
, males, dorsal habitus, Mergui Archipelago, syntypes of
<taxonomicName id="4C514D2CC50A0E54FA9DF8FA5E80F86C" authority="De Man, 1887" authorityName="De Man" authorityYear="1887" class="Malacostraca" family="Portunidae" genus="Goniosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="merguiensis">
<emphasis id="B925EABDC50A0E54FA9DF8FA5E3BF86C" italics="true" pageId="9" pageNumber="210">Goniosoma merguiensis</emphasis>
De Man, 1887
</taxonomicName>
, RMNH.CRUS.D.13101886. A. 25.0 x 40.0 mm. B. 21.0 x 32.0 mm.
</paragraph>
</caption>
<caption id="DF2E6627C5090E57FF2DF8285B07F85F" httpUri="https://zenodo.org/record/2624797/files/figure.png" pageId="10" pageNumber="211" startId="10.[151,250,1928,1950]" targetBox="[150,1436,193,1907]" targetPageId="10">
<paragraph id="8BEE36AFC5090E57FF2DF8285B07F85F" blockId="10.[151,1437,1928,1981]" pageId="10" pageNumber="211">
<emphasis id="B925EABDC5090E57FF2DF8285E38F87F" bold="true" box="[151,274,1928,1950]" pageId="10" pageNumber="211">FIGURE 6.</emphasis>
<taxonomicName id="4C514D2CC5090E57FEA2F8285DC9F87C" authority="(A. Milne-Edwards, 1867)" authorityName="A. Milne-Edwards" authorityYear="1867" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1867" box="[280,739,1928,1950]" class="Malacostraca" family="Portunidae" genus="Charybdis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="211" phylum="Arthropoda" rank="species" species="hellerii">
<emphasis id="B925EABDC5090E57FEA2F8285EFAF87C" box="[280,464,1928,1950]" italics="true" pageId="10" pageNumber="211">Charybdis hellerii</emphasis>
(A. Milne-Edwards, 1867)
</taxonomicName>
, males, Benin, Channel of Cotonou. A, B. 37 x 56 mm, RBINS, INV. 150111. C. 46.6 x 68.7 mm, RBINS, INV. 150110. A. Habitus dorsal. B, C. Frontal region.
</paragraph>
</caption>
<caption id="DF2E6627C5080E56FF2DF8285E81F83E" httpUri="https://zenodo.org/record/2624799/files/figure.png" pageId="11" pageNumber="212" startId="11.[151,250,1928,1950]" targetBox="[150,1436,193,1907]" targetPageId="11">
<paragraph id="8BEE36AFC5080E56FF2DF8285E81F83E" blockId="11.[151,1437,1928,2012]" pageId="11" pageNumber="212">
<emphasis id="B925EABDC5080E56FF2DF8285E3EF87C" bold="true" box="[151,276,1928,1950]" pageId="11" pageNumber="212">FIGURE 7.</emphasis>
<taxonomicName id="4C514D2CC5080E56FEA6F8285DC5F87C" authority="(A. Milne-Edwards, 1867)" authorityName="A. Milne-Edwards" authorityYear="1867" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1867" box="[284,751,1928,1950]" class="Malacostraca" family="Portunidae" genus="Charybdis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="hellerii">
<emphasis id="B925EABDC5080E56FEA6F8285EFFF87C" box="[284,469,1928,1950]" italics="true" pageId="11" pageNumber="212">Charybdis hellerii</emphasis>
(A. Milne-Edwards, 1867)
</taxonomicName>
, male, 37.0 x 56.0 mm, Benin, Channel of Cotonou, RBINS, INV. 150111. A. Lateral view of the chelipeds. B. Ventral habitus. C.
<taxonomicName id="4C514D2CC5080E56FC8DF8075C54F85F" authorityName="Pearse" authorityYear="1952" box="[823,894,1959,1981]" class="Maxillopoda" family="Telsidae" genus="Telson" higherTaxonomySource="GBIF" kingdom="Animalia" order="Poecilostomatoida" pageId="11" pageNumber="223" phylum="Arthropoda" rank="genus">Telson</taxonomicName>
. D. Carpus and extremity of adjacent articles of left pereiopod 5 (ventral view).
</paragraph>
</caption>
<caption id="DF2E6627C50F0E51FF2DF8285C85F85F" httpUri="https://zenodo.org/record/2624801/files/figure.png" pageId="12" pageNumber="213" startId="12.[151,250,1928,1950]" targetBox="[150,1436,193,1907]" targetPageId="12">
<paragraph id="8BEE36AFC50F0E51FF2DF8285C85F85F" blockId="12.[151,1437,1928,1981]" pageId="12" pageNumber="213">
<emphasis id="B925EABDC50F0E51FF2DF8285E3EF87F" bold="true" box="[151,276,1928,1950]" pageId="12" pageNumber="213">FIGURE 8.</emphasis>
<taxonomicName id="4C514D2CC50F0E51FEA6F8285DC5F87C" authority="(A. Milne-Edwards, 1867)" authorityName="A. Milne-Edwards" authorityYear="1867" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1867" box="[284,751,1928,1950]" class="Malacostraca" family="Portunidae" genus="Charybdis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="hellerii">
<emphasis id="B925EABDC50F0E51FEA6F8285EFFF87C" box="[284,469,1928,1950]" italics="true" pageId="12" pageNumber="213">Charybdis hellerii</emphasis>
(A. Milne-Edwards, 1867)
</taxonomicName>
, male, 37.0 x 56.0 mm, Benin, Channel of Cotonou, RBINS, INV. 150111. A. Right orbital region (ventral view). B. Right basal antennal article.
</paragraph>
</caption>
<caption id="DF2E6627C50E0E50FF2DF8285B8FF85F" httpUri="https://zenodo.org/record/2624803/files/figure.png" pageId="13" pageNumber="214" startId="13.[151,250,1928,1950]" targetBox="[150,1436,193,1907]" targetPageId="13">
<paragraph id="8BEE36AFC50E0E50FF2DF8285B8FF85F" blockId="13.[151,1437,1928,1981]" pageId="13" pageNumber="214">
<emphasis id="B925EABDC50E0E50FF2DF8285E3EF87F" bold="true" box="[151,276,1928,1950]" pageId="13" pageNumber="214">FIGURE 9.</emphasis>
<taxonomicName id="4C514D2CC50E0E50FEA6F8285DC5F87C" authority="(A. Milne-Edwards, 1867)" authorityName="A. Milne-Edwards" authorityYear="1867" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1867" box="[284,751,1928,1950]" class="Malacostraca" family="Portunidae" genus="Charybdis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="214" phylum="Arthropoda" rank="species" species="hellerii">
<emphasis id="B925EABDC50E0E50FEA6F8285EFFF87C" box="[284,469,1928,1950]" italics="true" pageId="13" pageNumber="214">Charybdis hellerii</emphasis>
(A. Milne-Edwards, 1867)
</taxonomicName>
, male, 37.0 x 56.0 mm, Benin, Channel of Cotonou, RBINS, INV. 150111. A. Gonopod 1. B. Tip of gonopod 1. The colour have been enhanced to increase the contrasts.
</paragraph>
</caption>
<caption id="DF2E6627C50D0E53FF2DF8285EC1F85F" httpUri="https://zenodo.org/record/2624805/files/figure.png" pageId="14" pageNumber="215" startId="14.[151,250,1928,1950]" targetBox="[150,1436,193,1907]" targetPageId="14">
<paragraph id="8BEE36AFC50D0E53FF2DF8285EC1F85F" blockId="14.[151,1436,1928,1981]" pageId="14" pageNumber="215">
<emphasis id="B925EABDC50D0E53FF2DF8285E0AF87F" bold="true" box="[151,288,1928,1950]" pageId="14" pageNumber="215">FIGURE 10.</emphasis>
<taxonomicName id="4C514D2CC50D0E53FE9DF8285DD2F87C" authority="(A. Milne-Edwards, 1867)" authorityName="A. Milne-Edwards" authorityYear="1867" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1867" box="[295,760,1928,1950]" class="Malacostraca" family="Portunidae" genus="Charybdis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="215" phylum="Arthropoda" rank="species" species="hellerii">
<emphasis id="B925EABDC50D0E53FE9DF8285ECAF87C" box="[295,480,1928,1950]" italics="true" pageId="14" pageNumber="215">Charybdis hellerii</emphasis>
(A. Milne-Edwards, 1867)
</taxonomicName>
, male, 47.0 x 68.0 mm, New Caledonia, RBINS, INV. 129463. A. Dorsal habitus. B. Frontal region.
</paragraph>
</caption>
<caption id="DF2E6627C50C0E52FF2DF8285DFDF83E" httpUri="https://zenodo.org/record/2624807/files/figure.png" pageId="15" pageNumber="216" startId="15.[151,250,1928,1950]" targetBox="[150,1436,193,1907]" targetPageId="15">
<paragraph id="8BEE36AFC50C0E52FF2DF8285DFDF83E" blockId="15.[151,1436,1928,2012]" pageId="15" pageNumber="216">
<emphasis id="B925EABDC50C0E52FF2DF8285E35F87F" bold="true" box="[151,287,1928,1950]" pageId="15" pageNumber="216">FIGURE 11.</emphasis>
<taxonomicName id="4C514D2CC50C0E52FE9CF8285DDCF87C" authority="(A. Milne-Edwards, 1867)" authorityName="A. Milne-Edwards" authorityYear="1867" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1867" box="[294,758,1928,1950]" class="Malacostraca" family="Portunidae" genus="Charybdis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="216" phylum="Arthropoda" rank="species" species="hellerii">
<emphasis id="B925EABDC50C0E52FE9CF8285EF5F87C" box="[294,479,1928,1950]" italics="true" pageId="15" pageNumber="216">Charybdis hellerii</emphasis>
(A. Milne-Edwards, 1867)
</taxonomicName>
, male, 47.0 x 68.0 mm, New Caledonia, RBINS, INV. 129463. A. Lateral view of chelipeds. B. Ventral habitus. C.
<taxonomicName id="4C514D2CC50C0E52FD33F8075DFAF85F" authorityName="Pearse" authorityYear="1952" box="[649,720,1959,1981]" class="Maxillopoda" family="Telsidae" genus="Telson" higherTaxonomySource="GBIF" kingdom="Animalia" order="Poecilostomatoida" pageId="15" pageNumber="223" phylum="Arthropoda" rank="genus">Telson</taxonomicName>
. D. Carpus and extremity of adjacent articles of left P5, ventral view). The colour have been enhanced to increase the contrasts.
</paragraph>
</caption>
<caption id="DF2E6627C5130E4DFF2DF8285C13F85F" httpUri="https://zenodo.org/record/2624809/files/figure.png" pageId="16" pageNumber="217" startId="16.[151,250,1928,1950]" targetBox="[150,1436,193,1907]" targetPageId="16">
<paragraph id="8BEE36AFC5130E4DFF2DF8285C13F85F" blockId="16.[151,1436,1928,1981]" pageId="16" pageNumber="217">
<emphasis id="B925EABDC5130E4DFF2DF8285E0AF87F" bold="true" box="[151,288,1928,1950]" pageId="16" pageNumber="217">FIGURE 12.</emphasis>
<taxonomicName id="4C514D2CC5130E4DFE9DF8285DD2F87C" authority="(A. Milne-Edwards, 1867)" authorityName="A. Milne-Edwards" authorityYear="1867" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1867" box="[295,760,1928,1950]" class="Malacostraca" family="Portunidae" genus="Charybdis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="hellerii">
<emphasis id="B925EABDC5130E4DFE9DF8285ECAF87C" box="[295,480,1928,1950]" italics="true" pageId="16" pageNumber="217">Charybdis hellerii</emphasis>
(A. Milne-Edwards, 1867)
</taxonomicName>
, male, 47.0 x 68.0 mm, New Caledonia, RBINS, INV. 129463. A. Right orbital region (ventral view). B. Right basal antennal article.
</paragraph>
</caption>
<caption id="DF2E6627C5120E4CFF2DF8285BB4F85F" httpUri="https://zenodo.org/record/2624811/files/figure.png" pageId="17" pageNumber="218" startId="17.[151,250,1928,1950]" targetBox="[150,1436,193,1907]" targetPageId="17">
<paragraph id="8BEE36AFC5120E4CFF2DF8285BB4F85F" blockId="17.[151,1436,1928,1981]" pageId="17" pageNumber="218">
<emphasis id="B925EABDC5120E4CFF2DF8285E0AF87F" bold="true" box="[151,288,1928,1950]" pageId="17" pageNumber="218">FIGURE 13.</emphasis>
<taxonomicName id="4C514D2CC5120E4CFE9DF8285DD2F87C" authority="(A. Milne-Edwards, 1867)" authorityName="A. Milne-Edwards" authorityYear="1867" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1867" box="[295,760,1928,1950]" class="Malacostraca" family="Portunidae" genus="Charybdis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="218" phylum="Arthropoda" rank="species" species="hellerii">
<emphasis id="B925EABDC5120E4CFE9DF8285ECAF87C" box="[295,480,1928,1950]" italics="true" pageId="17" pageNumber="218">Charybdis hellerii</emphasis>
(A. Milne-Edwards, 1867)
</taxonomicName>
, male, 47.0 x 68.0 mm, New Caledonia, RBINS, INV. 129463. A. Right gonopod 1. B. Tip of right gonopod 1. The colour have been enhanced to increase the contrasts.
</paragraph>
</caption>
<caption id="DF2E6627C5110E4FFF2DF8285EC1F85F" httpUri="https://zenodo.org/record/2624813/files/figure.png" pageId="18" pageNumber="219" startId="18.[151,250,1928,1950]" targetBox="[150,1436,193,1907]" targetPageId="18">
<paragraph id="8BEE36AFC5110E4FFF2DF8285EC1F85F" blockId="18.[151,1436,1928,1981]" pageId="18" pageNumber="219">
<emphasis id="B925EABDC5110E4FFF2DF8285E08F87F" bold="true" box="[151,290,1928,1950]" pageId="18" pageNumber="219">FIGURE 14.</emphasis>
<taxonomicName id="4C514D2CC5110E4FFE96F8285C2FF87C" authority="(A. Milne-Edwards, 1867)" authorityName="A. Milne-Edwards" authorityYear="1867" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1867" box="[300,773,1928,1950]" class="Malacostraca" family="Portunidae" genus="Charybdis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="219" phylum="Arthropoda" rank="species" species="hellerii">
<emphasis id="B925EABDC5110E4FFE96F8285ECDF87C" box="[300,487,1928,1950]" italics="true" pageId="18" pageNumber="219">Charybdis hellerii</emphasis>
(A. Milne-Edwards, 1867)
</taxonomicName>
, male, 37.0 x 56.0 mm, Queensland, RBINS, INV. 129466. A. Dorsal habitus. B. Frontal region.
</paragraph>
</caption>
<caption id="DF2E6627C5100E4EFF2DF8285C59F83E" httpUri="https://zenodo.org/record/2624817/files/figure.png" pageId="19" pageNumber="220" startId="19.[151,250,1928,1950]" targetBox="[150,1436,193,1907]" targetPageId="19">
<paragraph id="8BEE36AFC5100E4EFF2DF8285C59F83E" blockId="19.[151,1436,1928,2012]" pageId="19" pageNumber="220">
<emphasis id="B925EABDC5100E4EFF2DF8285E08F87F" bold="true" box="[151,290,1928,1950]" pageId="19" pageNumber="220">FIGURE 15.</emphasis>
<taxonomicName id="4C514D2CC5100E4EFE96F8285C2FF87C" authority="(A. Milne-Edwards, 1867)" authorityName="A. Milne-Edwards" authorityYear="1867" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1867" box="[300,773,1928,1950]" class="Malacostraca" family="Portunidae" genus="Charybdis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="220" phylum="Arthropoda" rank="species" species="hellerii">
<emphasis id="B925EABDC5100E4EFE96F8285ECDF87C" box="[300,487,1928,1950]" italics="true" pageId="19" pageNumber="220">Charybdis hellerii</emphasis>
(A. Milne-Edwards, 1867)
</taxonomicName>
, male, 37.0 x 56.0 mm, Queensland, RBINS, INV. 129466. A. Lateral view of the chelipeds. B. Ventral habitus. C.
<taxonomicName id="4C514D2CC5100E4EFD08F8075DD3F85F" authorityName="Pearse" authorityYear="1952" box="[690,761,1959,1981]" class="Maxillopoda" family="Telsidae" genus="Telson" higherTaxonomySource="GBIF" kingdom="Animalia" order="Poecilostomatoida" pageId="19" pageNumber="223" phylum="Arthropoda" rank="genus">Telson</taxonomicName>
. D. Carpus and extremity of adjacent articles of right pereiopod 5 (ventral view). The colour have been enhanced to increase the contrasts.
</paragraph>
</caption>
<caption id="DF2E6627C5170E49FF2DF8285A55F85F" httpUri="https://zenodo.org/record/2624819/files/figure.png" pageId="20" pageNumber="221" startId="20.[151,250,1928,1950]" targetBox="[150,1436,193,1907]" targetPageId="20">
<paragraph id="8BEE36AFC5170E49FF2DF8285A55F85F" blockId="20.[151,1436,1928,1981]" pageId="20" pageNumber="221">
<emphasis id="B925EABDC5170E49FF2DF8285E08F87F" bold="true" box="[151,290,1928,1950]" pageId="20" pageNumber="221">FIGURE 16.</emphasis>
<taxonomicName id="4C514D2CC5170E49FE96F8285C2FF87C" authority="(A. Milne-Edwards, 1867)" authorityName="A. Milne-Edwards" authorityYear="1867" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1867" box="[300,773,1928,1950]" class="Malacostraca" family="Portunidae" genus="Charybdis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="221" phylum="Arthropoda" rank="species" species="hellerii">
<emphasis id="B925EABDC5170E49FE96F8285ECDF87C" box="[300,487,1928,1950]" italics="true" pageId="20" pageNumber="221">Charybdis hellerii</emphasis>
(A. Milne-Edwards, 1867)
</taxonomicName>
, male, 37.0 x 56.0 mm, Queensland, RBINS, INV. 129466. A. Right orbital region (ventral view). B. Right basal antennal article. The colour have been enhanced to increase the contrasts.
</paragraph>
</caption>
<caption id="DF2E6627C5160E48FF2DF8285EF6F85F" httpUri="https://zenodo.org/record/2624821/files/figure.png" pageId="21" pageNumber="222" startId="21.[151,250,1928,1950]" targetBox="[150,1436,193,1907]" targetPageId="21">
<paragraph id="8BEE36AFC5160E48FF2DF8285EF6F85F" blockId="21.[151,1435,1928,1981]" pageId="21" pageNumber="222">
<emphasis id="B925EABDC5160E48FF2DF8285E35F87F" bold="true" box="[151,287,1928,1950]" pageId="21" pageNumber="222">FIGURE 17.</emphasis>
<taxonomicName id="4C514D2CC5160E48FE9FF8285DD9F87C" authority="(A. Milne-Edwards, 1867)" authorityName="A. Milne-Edwards" authorityYear="1867" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1867" box="[293,755,1928,1950]" class="Malacostraca" family="Portunidae" genus="Charybdis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="222" phylum="Arthropoda" rank="species" species="hellerii">
<emphasis id="B925EABDC5160E48FE9FF8285EF4F87C" box="[293,478,1928,1950]" italics="true" pageId="21" pageNumber="222">Charybdis hellerii</emphasis>
(A. Milne-Edwards, 1867)
</taxonomicName>
, male, 37.0 x 56.0 mm, Queensland, RBINS, INV. 129466. A. Left gonopod 1 B. Tip of gonopod 1.
</paragraph>
</caption>
<caption id="DF2E6627C5150E4BFF2DFBEF5C1BFB87" box="[151,817,1103,1125]" httpUri="https://zenodo.org/record/2624823/files/figure.png" pageId="22" pageNumber="223" startId="22.[151,250,1103,1125]" targetBox="[170,1414,191,1065]" targetPageId="22">
<paragraph id="8BEE36AFC5150E4BFF2DFBEF5C1BFB87" blockId="22.[151,817,1103,1125]" box="[151,817,1103,1125]" pageId="22" pageNumber="223">
<emphasis id="B925EABDC5150E4BFF2DFBEF5FD0FB87" bold="true" box="[151,250,1103,1125]" pageId="22" pageNumber="223">FIGURE</emphasis>
18. Location of the study area, showing fishing sectors.
</paragraph>
</caption>
<paragraph id="8BEE36AFC5150E4BFF7DFB2F5E25FB2E" blockId="22.[151,1437,1167,2020]" pageId="22" pageNumber="223">
<emphasis id="B925EABDC5150E4BFF7DFB2F5EE2FB4A" box="[199,456,1167,1192]" italics="true" pageId="22" pageNumber="223">
<taxonomicName id="4C514D2CC5150E4BFF7DFB2F5E25FB4A" authorityName="Pearse" authorityYear="1952" box="[199,271,1167,1192]" class="Maxillopoda" family="Telsidae" genus="Telson" higherTaxonomySource="GBIF" kingdom="Animalia" order="Poecilostomatoida" pageId="22" pageNumber="223" phylum="Arthropoda" rank="genus">Telson</taxonomicName>
of mature males
</emphasis>
: segment 6 about as long as wide, with borders convex on their distal 0.2. Gonopods as illustrated.
</paragraph>
<paragraph id="8BEE36AFC5150E4BFF7DFB775DB3FAF6" blockId="22.[151,1437,1167,2020]" pageId="22" pageNumber="223">
<emphasis id="B925EABDC5150E4BFF7DFB775FD4FB12" bold="true" box="[199,254,1239,1264]" pageId="22" pageNumber="223">Size.</emphasis>
Up to
<date id="FFEF106FC5150E4BFEECFB775EA1FB0D" box="[342,395,1239,1264]" pageId="22" pageNumber="223" value="1985-10">85 x</date>
<quantity id="4CA99B4AC5150E4BFE2CFB775EC3FB0D" box="[406,489,1239,1264]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="22" pageNumber="223" unit="mm" value="60.0">60 mm</quantity>
(
<bibRefCitation id="EFC04B5EC5150E4BFE46FB785C35FB12" author="Milne-Edwards, A." box="[508,799,1239,1264]" pageId="22" pageNumber="223" pagination="263 - 288" refId="ref16786" refString="Milne-Edwards, A. (1867) Descriptions de quelques especes nouvelles de Crustaces Brachyures. Annales de la Societe entomologique de France, Series 4, 7, 263 - 288." type="journal article" year="1867">A. Milne-Edwards, 1867</bibRefCitation>
). The
<typeStatus id="54EA880DC5150E4BFCD6FB775CF9FB12" box="[876,979,1239,1264]" pageId="22" pageNumber="223" type="lectotype">lectotype</typeStatus>
of
<taxonomicName id="4C514D2CC5150E4BFBBAFB775BFAFB12" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1867" box="[1024,1232,1239,1264]" class="Malacostraca" family="Portunidae" genus="Charybdis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="hellerii">
<emphasis id="B925EABDC5150E4BFBBAFB775BFAFB12" box="[1024,1232,1239,1264]" italics="true" pageId="22" pageNumber="223">Charybdis hellerii</emphasis>
</taxonomicName>
is apparently the largest specimen of the species ever recorded.
</paragraph>
<paragraph id="8BEE36AFC5150E4BFF7DFABF5D43F806" blockId="22.[151,1437,1167,2020]" pageId="22" pageNumber="223">
<emphasis id="B925EABDC5150E4BFF7DFABF5E90FADA" bold="true" box="[199,442,1311,1336]" pageId="22" pageNumber="223">Habitat and biology.</emphasis>
Intertidal to
<quantity id="4CA99B4AC5150E4BFDF6FA805DADFAD5" box="[588,647,1312,1336]" metricMagnitude="1" metricUnit="m" metricValue="4.7" pageId="22" pageNumber="223" unit="m" value="47.0">47 m</quantity>
, substrates rocky/cobble, sandy, muddy bottoms, mangroves and coral reefs (
<bibRefCitation id="EFC04B5EC5150E4BFF61FAE35EF4FABE" author="Stephenson, W. &amp; Hudson, J. J. &amp; Campbell, B." box="[219,478,1347,1372]" pageId="22" pageNumber="223" pagination="491 - 507" refId="ref19047" refString="Stephenson, W., Hudson, J. J. &amp; Campbell, B. (1957) The Australian Portunids (Crustacea; Portunidae). II. The genus Charybdis. Australian Journal of Marine and Freshwater Research, 8 (4), 491 - 507. https: // doi. org / 10.1071 / MF 9570491" type="journal article" year="1957">
Stephenson
<emphasis id="B925EABDC5150E4BFEDEFAE55EB1FABE" box="[356,411,1347,1372]" italics="true" pageId="22" pageNumber="223">et al.</emphasis>
1957
</bibRefCitation>
;
<bibRefCitation id="EFC04B5EC5150E4BFE52FAE35DA1FABE" author="Crosnier, A." box="[488,651,1347,1372]" pageId="22" pageNumber="223" pagination="1 - 154" refId="ref12372" refString="Crosnier, A. (1962) Crustaces Decapodes Portunidae. In: Faune de Madagascar. Vol. 16. ORSTOM, CNRS, Paris, pp. 1 - 154." type="book chapter" year="1962">Crosnier 1962</bibRefCitation>
;
<bibRefCitation id="EFC04B5EC5150E4BFD2FFAE45C67FABE" author="Wee, D. P. C. &amp; Ng, P. K. L." box="[661,845,1347,1372]" pageId="22" pageNumber="223" pagination="1 - 128" refId="ref19791" refString="Wee, D. P. C. &amp; Ng, P. K. L. (1995) Swimming crabs of the genera Charybdis de Haan, 1833, and Thalamita Latreille, 1829 (Crustacea: Decapoda: Brachyura: Portunidae) from Peninsular Malaysia and Singapore. Raffles Bulletin of Zoology, 1 (Supplement), 1 - 128." type="journal article" year="1995">Wee &amp; Ng 1995</bibRefCitation>
;
<bibRefCitation id="EFC04B5EC5150E4BFCE2FAE35B45FABE" author="Apel, M. &amp; Spiridonov, V. A." box="[856,1135,1347,1372]" pageId="22" pageNumber="223" pagination="159 - 331" refId="ref11122" refString="Apel, M. &amp; Spiridonov, V. A. (1998) Taxonomy and zoogeography of the portunid crabs (Crustacea: Decapoda: Brachyura: Portunidae) of the Arabian Gulf and adjacent waters. Fauna of Arabia, 17, 159 - 331." type="journal article" year="1998">Apel &amp; Spiridonov 1998</bibRefCitation>
;
<bibRefCitation id="EFC04B5EC5150E4BFBC1FAE35A6AFABE" author="Ozcan, T. &amp; Katagan, T. &amp; Irmak, E." box="[1147,1344,1347,1372]" pageId="22" pageNumber="223" pagination="1 - 5" refId="ref17981" refString="Ozcan, T., Katagan, T. &amp; Irmak, E. (2010) An exotic crab, Charybdis hellerii (A. Milne-Edwards, 1867) along the Turkish Coasts. Biharean Biologist, 4 (1), 1 - 5." type="journal article" year="2010">Özcan et al. 2010</bibRefCitation>
;
<bibRefCitation id="EFC04B5EC5150E4BFAF1FAE35E38FA62" author="Dineen, J. F. &amp; Clark, P. F. &amp; Hines, A. H. &amp; Reed, S. A. &amp; Walton, H. P." pageId="22" pageNumber="223" pagination="774 - 805" refId="ref13561" refString="Dineen, J. F., Clark, P. F., Hines, A. H., Reed, S. A. &amp; Walton, H. P. (2001) Life history, larval description, and natural history of Charybdis hellerii (Decapoda, Brachyura, Portunidae), an invasive crab in the Western Atlantic. Journal of Crustacean Biology, 21 (3), 774 - 805. https: // doi. org / 10.1163 / 20021975 - 99990173" type="journal article" year="2001">
Dineen
<emphasis id="B925EABDC5150E4BFF2DFAC95FFAFA62" box="[151,208,1383,1408]" italics="true" pageId="22" pageNumber="223">et al.</emphasis>
2001
</bibRefCitation>
;
<bibRefCitation id="EFC04B5EC5150E4BFEA4FAC75EE0FA62" author="Naderloo, R." box="[286,458,1383,1408]" pageId="22" pageNumber="223" refId="ref17022" refString="Naderloo, R. (2017) Atlas of the crabs of the Persian Gulf. Springer International Publishing AG, Basel, 444 pp. https: // doi. org / 10.1007 / 978 - 3 - 319 - 49374 - 9" type="book" year="2017">Naderloo 2017</bibRefCitation>
). Exceptionally found as deep as
<quantity id="4CA99B4AC5150E4BFCF9FAC75C56FA9D" box="[835,892,1383,1408]" metricMagnitude="1" metricUnit="m" metricValue="8.6" pageId="22" pageNumber="223" unit="m" value="86.0">86 m</quantity>
(
<bibRefCitation id="EFC04B5EC5150E4BFC31FAC75B77FA62" author="Stephenson, W." box="[907,1117,1383,1408]" pageId="22" pageNumber="223" pagination="127 - 156" refId="ref18919" refString="Stephenson, W. (1972 a) Portunid crabs from the Indo-West-Pacific and Western America in the Zoological Museum, Copenhagen (Decapoda, Brachyura, Portunidae). Steenstrupia, 2 (9), 127 - 156." type="journal article" year="1972">Stephenson 1972a</bibRefCitation>
).
<bibRefCitation id="EFC04B5EC5150E4BFBCBFAC75A79FA62" author="Dineen, J. F. &amp; Clark, P. F. &amp; Hines, A. H. &amp; Reed, S. A. &amp; Walton, H. P." box="[1137,1363,1383,1408]" pageId="22" pageNumber="223" pagination="774 - 805" refId="ref13561" refString="Dineen, J. F., Clark, P. F., Hines, A. H., Reed, S. A. &amp; Walton, H. P. (2001) Life history, larval description, and natural history of Charybdis hellerii (Decapoda, Brachyura, Portunidae), an invasive crab in the Western Atlantic. Journal of Crustacean Biology, 21 (3), 774 - 805. https: // doi. org / 10.1163 / 20021975 - 99990173" type="journal article" year="2001">
Dineen
<emphasis id="B925EABDC5150E4BFB73FAC95A28FA62" box="[1225,1282,1383,1408]" italics="true" pageId="22" pageNumber="223">et al.</emphasis>
(2001)
</bibRefCitation>
stated that
<taxonomicName id="4C514D2CC5150E4BFF70FA2B5EB3FA46" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1867" box="[202,409,1419,1444]" class="Malacostraca" family="Portunidae" genus="Charybdis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="hellerii">
<emphasis id="B925EABDC5150E4BFF70FA2B5EB3FA46" box="[202,409,1419,1444]" italics="true" pageId="22" pageNumber="223">Charybdis hellerii</emphasis>
</taxonomicName>
appeared to be closely associated with habitat structure of both hard substrates and some plant structures and presents a secretive and cryptic behaviour during daytime. The same authors reported that they collected it only at salinities&gt; 28 ppt and
<bibRefCitation id="EFC04B5EC5150E4BFDCFFA745C76FA0E" author="Trento Occhi, T. V." box="[629,860,1491,1516]" pageId="22" pageNumber="223" refId="ref19458" refString="Trento Occhi, T. V. (2016) Tolerancia a variacao de salinidade e plasticidade osmorregulatoria do siri invasor Charybdis hellerii (Crustacea, Portunidae) em comparacao com o caranguejo nativo Menippe nodifrons (Crustacea, Xanthidae). Dissertacao apresentada como requisito parcial para a obtencao do titulo de mestre em Ecologia, Area de concentracao em Ecologia e Conservacao, Setor de Ciencias Biologicas da Universidade Federal do Parana, Curitiba, 49 pp." type="book" year="2016">Trento Occhi (2016)</bibRefCitation>
demonstrated that it cannot survive in low salinity brackish water, with 100% mortality after two days at 20 ppt. This crab reaches sexual maturity within about one year (
<bibRefCitation id="EFC04B5EC5150E4BFF62F9BB5E9CF9D6" author="Dineen, J. F. &amp; Clark, P. F. &amp; Hines, A. H. &amp; Reed, S. A. &amp; Walton, H. P." box="[216,438,1563,1588]" pageId="22" pageNumber="223" pagination="774 - 805" refId="ref13561" refString="Dineen, J. F., Clark, P. F., Hines, A. H., Reed, S. A. &amp; Walton, H. P. (2001) Life history, larval description, and natural history of Charybdis hellerii (Decapoda, Brachyura, Portunidae), an invasive crab in the Western Atlantic. Journal of Crustacean Biology, 21 (3), 774 - 805. https: // doi. org / 10.1163 / 20021975 - 99990173" type="journal article" year="2001">
Dineen
<emphasis id="B925EABDC5150E4BFE8FF9BD5E5BF9D6" box="[309,369,1563,1588]" italics="true" pageId="22" pageNumber="223">et al.</emphasis>
2001
</bibRefCitation>
). Its lifespan is estimated to 23 years and it reproduces continuously during the year (
<bibRefCitation id="EFC04B5EC5150E4BFF24F99F5EA9F9BA" author="Bolanos, J. A. &amp; Baeza, J. A. &amp; Hernandez, J. E. &amp; Lira, C. &amp; Lopez, R." box="[158,387,1599,1624]" pageId="22" pageNumber="223" pagination="469 - 474" refId="ref11373" refString="Bolanos, J. A., Baeza, J. A., Hernandez, J. E., Lira, C. &amp; Lopez, R. (2012) Population dynamics and reproductive output of the non-indigenous crab Charybdis hellerii in the south-eastern Caribbean Sea. Journal of the Marine Biological Association of the United Kingdom, 92 (3), 469 - 474. https: // doi. org / 10.1017 / S 002531541100052 X" type="journal article" year="2012">
Bolaños
<emphasis id="B925EABDC5150E4BFEBEF9E15E15F9BA" box="[260,319,1599,1624]" italics="true" pageId="22" pageNumber="223">et al.</emphasis>
2012
</bibRefCitation>
), but ovigerous females are less abundant during the warmest months (
<bibRefCitation id="EFC04B5EC5150E4BFB15F99F5FF9F99E" author="Mantelatto, F. L. M. &amp; Garcia, R. B." pageId="22" pageNumber="223" pagination="469 - 477" refId="ref16231" refString="Mantelatto, F. L. M. &amp; Garcia, R. B. (2001) Biological aspects of the nonindigenous portunid crab Charybdis hellerii in the western tropical South Atlantic. Bulletin of Marine Science, 68 (3), 469 - 477." type="journal article" year="2001">Mantelatto &amp; Garcia 2001</bibRefCitation>
). The smallest ovigerous female recorded was
<quantity id="4CA99B4AC5150E4BFD5DF9C35C66F999" box="[743,844,1635,1660]" metricMagnitude="-2" metricUnit="m" metricValue="3.46" pageId="22" pageNumber="223" unit="mm" value="34.6">34.6 mm</quantity>
carapace width (
<bibRefCitation id="EFC04B5EC5150E4BFBB6F9C35A13F99E" author="Mantelatto, F. L. M. &amp; Garcia, R. B." box="[1036,1337,1635,1660]" pageId="22" pageNumber="223" pagination="469 - 477" refId="ref16231" refString="Mantelatto, F. L. M. &amp; Garcia, R. B. (2001) Biological aspects of the nonindigenous portunid crab Charybdis hellerii in the western tropical South Atlantic. Bulletin of Marine Science, 68 (3), 469 - 477." type="journal article" year="2001">Mantelatto &amp; Garcia 2001</bibRefCitation>
) and the largest still immature females recorded by Oshiro &amp; da
<bibRefCitation id="EFC04B5EC5150E4BFCB4F9275CFDF942" author="Oshiro, L. M. Y. &amp; da Conceicao, L." box="[782,983,1671,1696]" pageId="22" pageNumber="223" pagination="213 - 216" refId="ref17919" refString="Oshiro, L. M. Y. &amp; da Conceicao, L. (2001) Maturidade sexual em femeas de Charybdis hellerii (A. Milne-Edwards, 1867) (Crustacea, Decapoda, Portunidade) da Baia de Sepetiba-RJ. Anais da XI Jornada de Iniciacao Cientifica da UFRRJ, 11 (2), 213 - 216." type="journal article" year="2001">Conceição (2001)</bibRefCitation>
were about
<quantity id="4CA99B4AC5150E4BFBD9F9275B99F97D" box="[1123,1203,1671,1696]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" pageId="22" pageNumber="223" unit="mm" value="45.0">45 mm</quantity>
cw. The diameter of the eggs ranges from
<quantity id="4CA99B4AC5150E4BFE3CF90C5D70F921" box="[390,602,1707,1732]" metricMagnitude="-4" metricUnit="m" metricValue="2.45" metricValueMax="2.66" metricValueMin="2.24" pageId="22" pageNumber="223" unit="mm" value="0.245" valueMax="0.266" valueMin="0.224">0.224 to 0.266 mm</quantity>
in Turkish populations (
<bibRefCitation id="EFC04B5EC5150E4BFCD0F90B5B1EF926" author="Ozcan, T. &amp; Katagan, T. &amp; Irmak, E." box="[874,1076,1707,1732]" pageId="22" pageNumber="223" pagination="1 - 5" refId="ref17981" refString="Ozcan, T., Katagan, T. &amp; Irmak, E. (2010) An exotic crab, Charybdis hellerii (A. Milne-Edwards, 1867) along the Turkish Coasts. Biharean Biologist, 4 (1), 1 - 5." type="journal article" year="2010">
Özcan
<emphasis id="B925EABDC5150E4BFC03F90D5CC3F926" box="[953,1001,1707,1732]" italics="true" pageId="22" pageNumber="223">et al</emphasis>
. 2010
</bibRefCitation>
). Fecundity in
<taxonomicName id="4C514D2CC5150E4BFB63F90C5A60F926" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1867" box="[1241,1354,1707,1732]" class="Malacostraca" family="Portunidae" genus="Charybdis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="hellerii">
<emphasis id="B925EABDC5150E4BFB63F90C5A60F926" box="[1241,1354,1707,1732]" italics="true" pageId="22" pageNumber="223">C. hellerii</emphasis>
</taxonomicName>
is high and ranges from 22,550 eggs to 3,200,000 eggs per brood depending on size of the female (for review see
<bibRefCitation id="EFC04B5EC5150E4BFAF1F96F5E3CF8EE" author="Dineen, J. F. &amp; Clark, P. F. &amp; Hines, A. H. &amp; Reed, S. A. &amp; Walton, H. P." pageId="22" pageNumber="223" pagination="774 - 805" refId="ref13561" refString="Dineen, J. F., Clark, P. F., Hines, A. H., Reed, S. A. &amp; Walton, H. P. (2001) Life history, larval description, and natural history of Charybdis hellerii (Decapoda, Brachyura, Portunidae), an invasive crab in the Western Atlantic. Journal of Crustacean Biology, 21 (3), 774 - 805. https: // doi. org / 10.1163 / 20021975 - 99990173" type="journal article" year="2001">
Dineen
<emphasis id="B925EABDC5150E4BFF2DF9555FE0F8EE" box="[151,202,1779,1804]" italics="true" pageId="22" pageNumber="223">et al</emphasis>
. 2001
</bibRefCitation>
). It can store sperm to produce multiple broods (up to 6 broods per year in captive condition) and its larval development (through 6 zoeas and one megalopa stages) requires about 44 days at 24°C and 32-ppt salinity (
<bibRefCitation id="EFC04B5EC5150E4BFF24F89B5E59F8B6" author="Dineen, J. F. &amp; Clark, P. F. &amp; Hines, A. H. &amp; Reed, S. A. &amp; Walton, H. P." box="[158,371,1851,1876]" pageId="22" pageNumber="223" pagination="774 - 805" refId="ref13561" refString="Dineen, J. F., Clark, P. F., Hines, A. H., Reed, S. A. &amp; Walton, H. P. (2001) Life history, larval description, and natural history of Charybdis hellerii (Decapoda, Brachyura, Portunidae), an invasive crab in the Western Atlantic. Journal of Crustacean Biology, 21 (3), 774 - 805. https: // doi. org / 10.1163 / 20021975 - 99990173" type="journal article" year="2001">
Dineen
<emphasis id="B925EABDC5150E4BFF42F89D5E03F8B6" box="[248,297,1851,1876]" italics="true" pageId="22" pageNumber="223">et al</emphasis>
. 2001
</bibRefCitation>
).
<taxonomicName id="4C514D2CC5150E4BFE3CF89C5ED2F8B6" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1867" box="[390,504,1851,1876]" class="Malacostraca" family="Portunidae" genus="Charybdis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="hellerii">
<emphasis id="B925EABDC5150E4BFE3CF89C5ED2F8B6" box="[390,504,1851,1876]" italics="true" pageId="22" pageNumber="223">C. hellerii</emphasis>
</taxonomicName>
is an opportunistic feeder. Excluding organic matter, the most abundant food items found in its stomach by Sant'Anna
<emphasis id="B925EABDC5150E4BFD98F8C15D71F89A" box="[546,603,1887,1912]" italics="true" pageId="22" pageNumber="223">et al.</emphasis>
(2015) were crustaceans (25%), molluscs (21%) and macrophytes (13%). It is documented that
<taxonomicName id="4C514D2CC5150E4BFE31F8245ED7F87E" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1867" box="[395,509,1923,1948]" class="Malacostraca" family="Portunidae" genus="Charybdis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="hellerii">
<emphasis id="B925EABDC5150E4BFE31F8245ED7F87E" box="[395,509,1923,1948]" italics="true" pageId="22" pageNumber="223">C. hellerii</emphasis>
</taxonomicName>
is itself consumed by octopuses (
<bibRefCitation id="EFC04B5EC5150E4BFCCDF8235B51F87E" author="Sampaio, C. &amp; LS &amp; Rosa, I. L." box="[887,1147,1923,1948]" pageId="22" pageNumber="223" pagination="58" refId="ref18397" refString="Sampaio, C. LS. &amp; Rosa, I. L. (2005) Predation of an alien species of crab (Charybdis hellerii Milne Edwards) by a native Octopus species on NE Brazilian reefs. Coral Reefs, 25 (1), 58. https: // doi. org / 10.1007 / s 00338 - 005 - 0050 - 0" type="journal article" year="2005">Sampaio &amp; Rosa 2005</bibRefCitation>
). Parasitism by
<taxonomicName id="4C514D2CC5150E4BFA97F8235AB1F87E" box="[1325,1435,1923,1948]" class="Maxillopoda" family="Sacculinidae" genus="Sacculina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Kentrogonida" pageId="22" pageNumber="223" phylum="Arthropoda" rank="genus">
<emphasis id="B925EABDC5150E4BFA97F8235AB1F87E" box="[1325,1435,1923,1948]" italics="true" pageId="22" pageNumber="223">Sacculina</emphasis>
</taxonomicName>
has been reported in Australian populations [1.3% of infection] (
<bibRefCitation id="EFC04B5EC5150E4BFCD6F8075B44F822" author="Stephenson, W. &amp; Hudson, J. J. &amp; Campbell, B." box="[876,1134,1959,1984]" pageId="22" pageNumber="223" pagination="491 - 507" refId="ref19047" refString="Stephenson, W., Hudson, J. J. &amp; Campbell, B. (1957) The Australian Portunids (Crustacea; Portunidae). II. The genus Charybdis. Australian Journal of Marine and Freshwater Research, 8 (4), 491 - 507. https: // doi. org / 10.1071 / MF 9570491" type="journal article" year="1957">
Stephenson
<emphasis id="B925EABDC5150E4BFC4FF8095B04F822" box="[1013,1070,1959,1984]" italics="true" pageId="22" pageNumber="223">et al.</emphasis>
1957
</bibRefCitation>
) and
<collectingCountry id="F346763FC5150E4BFB17F8075BCDF822" box="[1197,1255,1959,1984]" name="India" pageId="22" pageNumber="223">India</collectingCountry>
(
<bibRefCitation id="EFC04B5EC5150E4BFB4CF8075FFBF806" author="Elumalai, V. &amp; Viswanathan, C. &amp; Prakinkumar, M. &amp; Raffi, S. M." pageId="22" pageNumber="223" pagination="337 - 339" refId="ref13753" refString="Elumalai, V., Viswanathan, C., Prakinkumar, M. &amp; Raffi, S. M. (2014) Infestation of parasitic barnacle Sacculina spp. in commercial marine crabs. Journal of Parasitic Diseases, 38 (3), 337 - 339. https: // doi. org / 10.1007 / s 12639 - 013 - 0247 - z" type="journal article" year="2014">
Elumalai
<emphasis id="B925EABDC5150E4BFADEF8095AB6F822" box="[1380,1436,1959,1984]" italics="true" pageId="22" pageNumber="223">et al.</emphasis>
2014
</bibRefCitation>
) but not yet in invasive populations.
</paragraph>
</subSubSection>
<subSubSection id="C34B6524C5140E4AFF7DFF375BA5FD82" pageId="23" pageNumber="224" type="distribution">
<paragraph id="8BEE36AFC5140E4AFF7DFF375BA5FD82" blockId="23.[151,1436,151,608]" pageId="23" pageNumber="224">
<emphasis id="B925EABDC5140E4AFF7DFF375E4AFF52" bold="true" box="[199,352,151,176]" pageId="23" pageNumber="224">Distribution.</emphasis>
Native:
<collectingCountry id="F346763FC5140E4AFE05FF375D78FF52" box="[447,594,151,176]" name="South Africa" pageId="23" pageNumber="224">South Africa</collectingCountry>
, East Africa,
<collectingCountry id="F346763FC5140E4AFD52FF375C5BFF52" box="[744,881,151,176]" name="Madagascar" pageId="23" pageNumber="224">Madagascar</collectingCountry>
, Red Sea, Gulf of
<collectingRegion id="4995F84DC5140E4AFB87FF375B56FF52" box="[1085,1148,151,176]" country="Yemen" name="'Adan" pageId="23" pageNumber="224">Aden</collectingRegion>
,
<collectingCountry id="F346763FC5140E4AFB3DFF385BF5FF52" box="[1159,1247,152,176]" name="Yemen" pageId="23" pageNumber="224">Socotra</collectingCountry>
, southern
<collectingCountry id="F346763FC5140E4AFAEAFF385ABDFF52" box="[1360,1431,152,176]" name="Oman" pageId="23" pageNumber="224">Oman</collectingCountry>
, Persian Gulf, Gulf of
<collectingCountry id="F346763FC5140E4AFE2AFF1C5EFDFF36" box="[400,471,188,212]" name="Oman" pageId="23" pageNumber="224">Oman</collectingCountry>
,
<collectingCountry id="F346763FC5140E4AFE5EFF1C5D6CFF37" box="[484,582,188,213]" name="Pakistan" pageId="23" pageNumber="224">Pakistan</collectingCountry>
,
<collectingCountry id="F346763FC5140E4AFDE8FF1C5DA4FF37" box="[594,654,188,213]" name="India" pageId="23" pageNumber="224">India</collectingCountry>
,
<collectingCountry id="F346763FC5140E4AFD21FF1C5C27FF37" box="[667,781,188,213]" name="Sri Lanka" pageId="23" pageNumber="224">Sri Lanka</collectingCountry>
, Mergui-Archipelago, Andaman Sea,
<collectingCountry id="F346763FC5140E4AFB01FF1C5A0BFF37" box="[1211,1313,188,213]" name="Thailand" pageId="23" pageNumber="224">Thailand</collectingCountry>
,
<collectingCountry id="F346763FC5140E4AFA95FF1C5AB2FF37" box="[1327,1432,188,213]" name="Malaysia" pageId="23" pageNumber="224">Malaysia</collectingCountry>
,
<collectingCountry id="F346763FC5140E4AFF2DFF7F5E32FF1A" box="[151,280,223,248]" name="Philippines" pageId="23" pageNumber="224">Philippines</collectingCountry>
,
<collectingCountry id="F346763FC5140E4AFE9FFF7F5EBFFF1A" box="[293,405,223,248]" name="Indonesia" pageId="23" pageNumber="224">Indonesia</collectingCountry>
,
<collectingCountry id="F346763FC5140E4AFE18FF7F5D3DFF1A" box="[418,535,223,248]" name="Singapore" pageId="23" pageNumber="224">Singapore</collectingCountry>
,
<collectingCountry id="F346763FC5140E4AFD9EFF7F5DA4FF1A" box="[548,654,223,248]" name="Australia" pageId="23" pageNumber="224">Australia</collectingCountry>
,
<collectingCountry id="F346763FC5140E4AFD21FF405C61FF1A" box="[667,843,223,248]" name="New Caledonia" pageId="23" pageNumber="224">New Caledonia</collectingCountry>
(
<bibRefCitation id="EFC04B5EC5140E4AFCE6FF7F5B27FF1A" author="Naderloo, R." box="[860,1037,223,248]" pageId="23" pageNumber="224" refId="ref17022" refString="Naderloo, R. (2017) Atlas of the crabs of the Persian Gulf. Springer International Publishing AG, Basel, 444 pp. https: // doi. org / 10.1007 / 978 - 3 - 319 - 49374 - 9" type="book" year="2017">Naderloo 2017</bibRefCitation>
). Invasive: Mediterranean (
<bibRefCitation id="EFC04B5EC5140E4AFAFFFF7F5FF9FEFE" author="Steinitz, W." pageId="23" pageNumber="224" pagination="1 - 90" refId="ref18875" refString="Steinitz, W. (1929) Die Wanderung indopazifischer Arten ins Mittelmeer seit Beginn der Quartarperiode. International Revue der Gesamten Hydrobiologie und Hydrographie, 22, 1 - 90. https: // doi. org / 10.1002 / iroh. 19290220102" type="journal article" year="1929">Steinitz 1929</bibRefCitation>
;
<bibRefCitation id="EFC04B5EC5140E4AFF64FEA45E5BFEFE" author="Monod, T." box="[222,369,260,285]" pageId="23" pageNumber="224" pagination="135 - 141" refId="ref16912" refString="Monod, T. (1930) Uber einige indo-pazifische Dekapoden der Meeresfauna Syriens. Zoologischer Anzeiger, 92 (5 / 6), 135 - 141." type="journal article" year="1930">Monod 1930</bibRefCitation>
;
<bibRefCitation id="EFC04B5EC5140E4AFEC7FEA45D1CFEFE" author="Galil, B. &amp; Froglia, C. &amp; Noel, P." box="[381,566,260,285]" pageId="23" pageNumber="224" refId="ref14584" refString="Galil, B., Froglia, C. &amp; Noel, P. (2002) CIESM Atlas of Exotic Species in the Mediterranean Vol. 2. Crustaceans: Decapods and Stomatopods. CIESM Publishers, Monte Carlo, 192 pp." type="book" year="2002">
Galil
<emphasis id="B925EABDC5140E4AFE06FEA55EC7FEFF" box="[444,493,260,285]" italics="true" pageId="23" pageNumber="224">et al</emphasis>
. 2002
</bibRefCitation>
;
<bibRefCitation id="EFC04B5EC5140E4AFDFBFEA45C0EFEFF" author="Yokes, B. &amp; Galil, B. S." box="[577,804,260,285]" pageId="23" pageNumber="224" pagination="747 - 755" refId="ref19856" refString="Yokes, B. &amp; Galil, B. S. (2006) New records of alien decapods (Crustacea) from the Mediterranean coast of Turkey, with a description of a new palaemonid species. Zoosystema, 28 (3), 747 - 755." type="journal article" year="2006">Yokes &amp; Galil 2006</bibRefCitation>
;
<bibRefCitation id="EFC04B5EC5140E4AFC8AFEA45CD2FEFE" author="Yokes, M. B. &amp; Karhan, S. U. &amp; Okus, E. &amp; Yusek, A. &amp; Aslan-Yilmaz, A. &amp; Yilmaz, I. N. &amp; Demirel, N. &amp; Demir, V. &amp; Galil, B. S." box="[816,1016,260,285]" pageId="23" pageNumber="224" pagination="162 - 168" refId="ref19905" refString="Yokes, M. B., Karhan, S. U., Okus, E., Yusek, A., Aslan-Yilmaz, A., Yilmaz, I. N., Demirel, N., Demir, V. &amp; Galil, B. S. (2007) Alien Crustacean Decapods from the Aegean Coast of Turkey. Aquatic Invasions, 2 (3), 162 - 168. https: // doi. org / 10.3391 / ai. 2007.2.3.2" type="journal article" year="2007">
Yokes
<emphasis id="B925EABDC5140E4AFCC6FEA55C9EFEFF" box="[892,948,260,285]" italics="true" pageId="23" pageNumber="224">et al.</emphasis>
2007
</bibRefCitation>
); Hawai'i (
<bibRefCitation id="EFC04B5EC5140E4AFBD5FEA45A10FEFF" author="Edmondson, C. H." box="[1135,1338,260,285]" pageId="23" pageNumber="224" pagination="217 - 274" refId="ref13723" refString="Edmondson, C. H. (1954) Hawaiian Portunidae. Occasional Papers of Bernice P. Bishop Museum, 21 (12), 217 - 274." type="journal article" year="1954">Edmondson 1954</bibRefCitation>
: isolated record; Evans
<emphasis id="B925EABDC5140E4AFE86FE895E5CFEA2" box="[316,374,295,320]" italics="true" pageId="23" pageNumber="224">et al.</emphasis>
2018: population); south Pacific: sea-chest of a fishing vessel in
<collectingCountry id="F346763FC5140E4AFBDBFE885BD3FEA2" box="[1121,1273,295,320]" name="New Zealand" pageId="23" pageNumber="224">New Zealand</collectingCountry>
(
<bibRefCitation id="EFC04B5EC5140E4AFAB3FE875E0DFE86" author="Dodgshun, T. &amp; Coutts, A." pageId="23" pageNumber="224" pagination="35" refId="ref13683" refString="Dodgshun, T. &amp; Coutts, A. (2003) Opening the lid on sea chests. Seafood New Zealand, 11, 35. [Not seen, cited by Brockerhoff &amp; McLay (2011)]" type="journal article" year="2003">Dodgshun &amp; Coutts 2003</bibRefCitation>
); western Atlantic from the Atlantic coast of the
<collectingCountry id="F346763FC5140E4AFCD8FEEC5CB0FE86" box="[866,922,332,356]" name="United States of America" pageId="23" pageNumber="224">USA</collectingCountry>
to the south of
<collectingCountry id="F346763FC5140E4AFBEDFEEC5BB1FE87" box="[1111,1179,332,357]" name="Brazil" pageId="23" pageNumber="224">Brazil</collectingCountry>
(for review see
<bibRefCitation id="EFC04B5EC5140E4AFAE6FEEC5E1AFE6A" author="Negri Pereira, M." pageId="23" pageNumber="224" refId="ref17215" refString="Negri Pereira, M. (2016) Registro de Charybdis hellerii (Milne Edwards, 1867) em aguas do litoral brasileiro (Decapoda: Portunidae). Tese apresentada a Faculdade de Filosofia, Ciencias e Letras de Ribeirao Preto da USP, como parte das exigencias para a obtencao do titulo de Doutor em Ciencias Area: Biologia Comparada, viii + 164 pp." type="book" year="2016">Negri Pereira 2016</bibRefCitation>
;
<bibRefCitation id="EFC04B5EC5140E4AFEFEFECF5D37FE6A" author="Negri, M. &amp; Schubart, C. D. &amp; Mantelatto, F. L." box="[324,541,367,392]" pageId="23" pageNumber="224" pagination="1771 - 1798" refId="ref17133" refString="Negri, M., Schubart, C. D. &amp; Mantelatto, F. L. (2018) Tracing the introduction history of the invasive swimming crab Charybdis hellerii (A. Milne-Edwards, 1867) in the Western Atlantic: evidences of high genetic diversity and multiple introductions. Biological Invasions, 20 (7), 1771 - 1798. https: // doi. org / 10.1007 / s 10530 - 018 - 1660 - 0" type="journal article" year="2018">
Negri
<emphasis id="B925EABDC5140E4AFE28FED15EE0FE6A" box="[402,458,367,392]" italics="true" pageId="23" pageNumber="224">et al</emphasis>
. 2018
</bibRefCitation>
); tropical eastern Atlantic:
<collectingCountry id="F346763FC5140E4AFCD7FECF5B38FE6A" box="[877,1042,367,392]" name="Guinea-Bissau" pageId="23" pageNumber="224">Guinea-Bissau</collectingCountry>
[
<geoCoordinate id="EE655068C5140E4AFB9DFECF5B8DFE6A" box="[1063,1191,367,392]" direction="north" orientation="latitude" pageId="23" pageNumber="224" precision="5" value="11.5342">11.5342°N</geoCoordinate>
,
<geoCoordinate id="EE655068C5140E4AFB00FED05A17FE6A" box="[1210,1341,368,392]" direction="west" orientation="longitude" pageId="23" pageNumber="224" precision="5" value="-17.077">17.0770°W</geoCoordinate>
: (http:// www.discoverlife.org/mp/20l?id=GBIF773871744 accessed
<date id="FFEF106FC5140E4AFCE3FE345CFCFE4F" box="[857,982,404,429]" pageId="23" pageNumber="224" value="2018-03-28">28.iii.2018</date>
)] and
<collectingCountry id="F346763FC5140E4AFB8FFE345B52FE4F" box="[1077,1144,404,429]" name="Benin" pageId="23" pageNumber="224">Benin</collectingCountry>
(present material). An accidental isolated record on the hull of a ship in northwest
<collectingCountry id="F346763FC5140E4AFCF9FE175CA9FE32" box="[835,899,439,464]" name="Spain" pageId="23" pageNumber="224">Spain</collectingCountry>
has been reported by
<bibRefCitation id="EFC04B5EC5140E4AFB3AFE185A4EFE32" author="Cuesta, J. A. &amp; Almon, B. &amp; Perez-Dieste, J. &amp; Trigo, J. E. &amp; Banon, R." box="[1152,1380,439,464]" pageId="23" pageNumber="224" pagination="619 - 630" refId="ref12454" refString="Cuesta, J. A., Almon, B., Perez-Dieste, J., Trigo, J. E. &amp; Banon, R. (2016) Role of ships' hull fouling and tropicalization process on European carcinofauna: new records in Galician waters (NW Spain). Biological Invasions, 19, 619 - 630. https: // doi. org / 10.1007 / s 10530 - 015 - 1034 - 9" type="journal article" year="2016">
Cuesta
<emphasis id="B925EABDC5140E4AFB6EFE195A2CFE32" box="[1236,1286,439,464]" italics="true" pageId="23" pageNumber="224">et al</emphasis>
. (2016)
</bibRefCitation>
. The Mediterranean populations arrived from the Red Sea via the Suez Canal (
<bibRefCitation id="EFC04B5EC5140E4AFC5AFE7C5B8AFE16" author="Galil, B. &amp; Froglia, C. &amp; Noel, P." box="[992,1184,476,501]" pageId="23" pageNumber="224" refId="ref14584" refString="Galil, B., Froglia, C. &amp; Noel, P. (2002) CIESM Atlas of Exotic Species in the Mediterranean Vol. 2. Crustaceans: Decapods and Stomatopods. CIESM Publishers, Monte Carlo, 192 pp." type="book" year="2002">
Galil
<emphasis id="B925EABDC5140E4AFB99FE7D5B77FE17" box="[1059,1117,476,501]" italics="true" pageId="23" pageNumber="224">et al.</emphasis>
2002
</bibRefCitation>
;
<bibRefCitation id="EFC04B5EC5140E4AFB14FE7C5A5DFE16" author="Negri, M. &amp; Schubart, C. D. &amp; Mantelatto, F. L." box="[1198,1399,476,501]" pageId="23" pageNumber="224" pagination="1771 - 1798" refId="ref17133" refString="Negri, M., Schubart, C. D. &amp; Mantelatto, F. L. (2018) Tracing the introduction history of the invasive swimming crab Charybdis hellerii (A. Milne-Edwards, 1867) in the Western Atlantic: evidences of high genetic diversity and multiple introductions. Biological Invasions, 20 (7), 1771 - 1798. https: // doi. org / 10.1007 / s 10530 - 018 - 1660 - 0" type="journal article" year="2018">
Negri
<emphasis id="B925EABDC5140E4AFB4DFE7D5A1BFE17" box="[1271,1329,476,501]" italics="true" pageId="23" pageNumber="224">et al.</emphasis>
2018
</bibRefCitation>
). It seems that the sources of invasive Western Atlantic populations are manyfold, consisting both of specimens from the western Indian Ocean and Mediterranean Sea group (major source) and from the Eastern Indian + western Pacific group (minor source, southern
<collectingCountry id="F346763FC5140E4AFDFFFDE75DA3FD82" box="[581,649,583,608]" name="Brazil" pageId="23" pageNumber="224">Brazil</collectingCountry>
only) (
<bibRefCitation id="EFC04B5EC5140E4AFD61FDE75C9FFD82" author="Negri Pereira, M." box="[731,949,583,608]" pageId="23" pageNumber="224" refId="ref17215" refString="Negri Pereira, M. (2016) Registro de Charybdis hellerii (Milne Edwards, 1867) em aguas do litoral brasileiro (Decapoda: Portunidae). Tese apresentada a Faculdade de Filosofia, Ciencias e Letras de Ribeirao Preto da USP, como parte das exigencias para a obtencao do titulo de Doutor em Ciencias Area: Biologia Comparada, viii + 164 pp." type="book" year="2016">Negri Pereira 2016</bibRefCitation>
;
<bibRefCitation id="EFC04B5EC5140E4AFC7AFDE75BA8FD82" author="Negri, M. &amp; Schubart, C. D. &amp; Mantelatto, F. L." box="[960,1154,583,608]" pageId="23" pageNumber="224" pagination="1771 - 1798" refId="ref17133" refString="Negri, M., Schubart, C. D. &amp; Mantelatto, F. L. (2018) Tracing the introduction history of the invasive swimming crab Charybdis hellerii (A. Milne-Edwards, 1867) in the Western Atlantic: evidences of high genetic diversity and multiple introductions. Biological Invasions, 20 (7), 1771 - 1798. https: // doi. org / 10.1007 / s 10530 - 018 - 1660 - 0" type="journal article" year="2018">
Negri
<emphasis id="B925EABDC5140E4AFBBDFDE95B15FD82" box="[1031,1087,583,608]" italics="true" pageId="23" pageNumber="224">et al.</emphasis>
2018
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>