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<document id="9C8694B5D9AFBE9ADF52AA810DFDAAAE" ID-DOI="10.5281/zenodo.280763" ID-GBIF-Dataset="3b9585ce-995d-4c11-99d5-eea5dcec4417" ID-ISSN="1175-5326" ID-Zenodo-Dep="280763" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460471218047" checkinUser="plazi" docAuthor="Komai, Tomoyuki, Reshmi, Rema &amp; Kumar, Appukuttan Nair Biju" docDate="2012" docId="03DDA531FFD3FF9BD19ECE1FFA88FD44" docLanguage="en" docName="zt03266p061.pdf" docOrigin="Zootaxa 3266" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Ciliopagurus grandis Komai, Reshmi &amp; Kumar, 2012, n. sp." docType="treatment" docVersion="7" lastPageNumber="61" masterDocId="FFE4DD49FFD3FF93D109C92AFFF6FFAB" masterDocTitle="A new species of the hermit crab genus Ciliopagurus Forest (Crustacea: Decapoda: Anomura: Diogenidae) from southern India" masterLastPageNumber="61" masterPageNumber="53" pageNumber="53" updateTime="1698305775244" updateUser="plazi">
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<mods:title id="9A5C9A0E4D71ED75F16F488D8047B1D6">A new species of the hermit crab genus Ciliopagurus Forest (Crustacea: Decapoda: Anomura: Diogenidae) from southern India</mods:title>
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<mods:namePart id="AEB315CC6226C7C3EFCB8E36753459C7">Komai, Tomoyuki</mods:namePart>
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<mods:namePart id="341368A9DB22972A5CDF020D2DF920B2">Reshmi, Rema</mods:namePart>
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<mods:namePart id="8F7CAEF7EA05230ED1EAD0381B7E371F">Kumar, Appukuttan Nair Biju</mods:namePart>
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<mods:title id="DD8B9C668E315FBD87470775A69630AE">Zootaxa</mods:title>
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<mods:date id="5E7540ADCD0811C46F4DEE46FA77CEA0">2012</mods:date>
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<treatment id="03DDA531FFD3FF9BD19ECE1FFA88FD44" ID-DOI="http://doi.org/10.5281/zenodo.6176578" ID-GBIF-Taxon="119515278" ID-Zenodo-Dep="6176578" LSID="urn:lsid:plazi:treatment:03DDA531FFD3FF9BD19ECE1FFA88FD44" httpUri="http://treatment.plazi.org/id/03DDA531FFD3FF9BD19ECE1FFA88FD44" lastPageId="8" lastPageNumber="61" pageId="0" pageNumber="53">
<subSubSection id="C36E47ACFFD3FF93D19ECE1FFEFDF8C4" pageId="0" pageNumber="53" type="nomenclature">
<paragraph id="8BCB1427FFD3FF93D19ECE1FFE17F8E5" blockId="0.[151,481,1844,1903]" box="[151,481,1844,1871]" pageId="0" pageNumber="53">
<heading id="D083A34BFFD3FF93D19ECE1FFE17F8E5" bold="true" box="[151,481,1844,1871]" fontSize="11" level="1" pageId="0" pageNumber="53" reason="1">
<emphasis id="B900C835FFD3FF93D19ECE1FFE17F8E5" bold="true" box="[151,481,1844,1871]" pageId="0" pageNumber="53">
<taxonomicName id="4C746FA4FFD3FF93D19ECE1FFE61F8E4" ID-CoL="8a1447e9-ff35-40a6-9757-b485c4b50480" box="[151,407,1845,1871]" class="Malacostraca" family="Diogenidae" genus="Ciliopagurus" kingdom="Animalia" order="Decapoda" pageId="0" pageNumber="53" phylum="Arthropoda" rank="species" species="grandis" status="sp. nov.">
<emphasis id="B900C835FFD3FF93D19ECE1FFE61F8E4" bold="true" box="[151,407,1845,1871]" italics="true" pageId="0" pageNumber="53">Ciliopagurus grandis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A233754EFFD3FF93D097CE1EFE17F8E5" box="[414,481,1844,1870]" pageId="0" pageNumber="53" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BCB1427FFD3FF93D19ECE7DFEFDF8C4" blockId="0.[151,481,1844,1903]" box="[151,267,1879,1903]" pageId="0" pageNumber="53">
(
<figureCitation id="134F08A2FFD3FF93D196CE7DFEF7F8C4" box="[159,257,1879,1903]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStartId-0="1.[151,250,1981,2004]" captionStartId-1="3.[151,250,1875,1898]" captionStartId-2="4.[151,250,906,929]" captionStartId-3="5.[151,250,1920,1943]" captionStartId-4="6.[151,250,1950,1973]" captionTargetBox-0="[173,1430,151,1937]" captionTargetBox-1="[176,1400,184,1874]" captionTargetBox-2="[175,1404,187,854]" captionTargetBox-3="[194,1399,185,1907]" captionTargetBox-4="[153,1427,184,1949]" captionTargetId-0="figure@1.[173,1437,141,1964]" captionTargetId-1="figure@3.[151,1415,157,1884]" captionTargetId-2="figure@4.[151,1436,173,885]" captionTargetId-3="figure@5.[168,1428,159,1908]" captionTargetId-4="figure@6.[151,1436,163,1963]" captionTargetPageId-0="1" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionTargetPageId-4="6" captionText-0="FIGURE 1. Ciliopagurus grandis n. sp., holotype, male (sl 13.7 mm), CBM-ZC 10663, showing coloration preserved in formalin. A, entire animal in dorsal view; B, same, lateral view." captionText-1="FIGURE 2. Ciliopagurus grandis n. sp., holotype, male (sl 13.7 mm), CBM-ZC 10663. A, shield and cephalic appendages, dorsal view (setae omitted from left side); B, left antennal acicle, dorsal view (setae omitted); C, endopod of left maxillule, lateral view; D, left third maxilliped, lateral view (setae omitted); E, same, ischium, ventral view; F, same, exopod, dorsal view (setae omitted); G, right fourth pereopod, lateral view; H, anterior lobe of sixth thoracic sternite, ventral view; I, sixth pleonal tergite, dorsal view; J, telson, dorsal view. Scale bars: 5 mm for A; 2 mm for H; 1 mm for B, D, F, G, I, J; 0.5 mm for C, E." captionText-2="FIGURE 3. Ciliopagurus grandis n. sp., holotype, male (sl 13.7 mm), CBM-ZC 10663. A, left maxilla, outer view (setae on scaphognathite omitted) l B, left first maxilliped, outer view; C, left second maxilliped, outer view. Scale bars: 2 mm." captionText-3="FIGURE 4. Ciliopagurus grandis n. sp., holotype, male (sl 13.7 mm), CBM-ZC 10663. A, left cheliped, lateral view; B, left chela and carpus, dorsal view; C, left cheliped, mesial view; D, close up of transverse ridges or striae on dorsal surface near base of fixed finger (setae omitted); E, close up of distal stridulating ridges on mesial surface of palm, setae partially shown. Scale bars: 5 mm for A C; 1 mm for D, E." captionText-4="FIGURE 5. Ciliopagurus grandis n. sp., holotype, male (sl 13.7 mm), CBM-ZC 10663. A, right second pereopod, lateral view (only setae on dactylus shown); B, same, mesial view (setae omitted); C, same, dactylus, ventral view; D, right third pereopod, lateral view (setae omitted); E, same, mesial view. Scale bars: 5 mm." httpUri-0="https://zenodo.org/record/280764/files/figure.png" httpUri-1="https://zenodo.org/record/280765/files/figure.png" httpUri-2="https://zenodo.org/record/280766/files/figure.png" httpUri-3="https://zenodo.org/record/280767/files/figure.png" httpUri-4="https://zenodo.org/record/280768/files/figure.png" pageId="0" pageNumber="53">Figs 15</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36E47ACFFD3FF91D19ECEB5FACEFF5C" lastPageId="2" lastPageNumber="55" pageId="0" pageNumber="53" type="materials_examined">
<paragraph id="8BCB1427FFD3FF93D19ECEB5FDB1F870" blockId="0.[151,1436,1951,2012]" pageId="0" pageNumber="53">
<emphasis id="B900C835FFD3FF93D19ECEB5FE88F813" bold="true" box="[151,382,1951,1976]" pageId="0" pageNumber="53">Material examined</emphasis>
.
<emphasis id="B900C835FFD3FF93D099CEB5FE00F81C" box="[400,502,1951,1975]" italics="true" pageId="0" pageNumber="53">
<typeStatus id="54CFAA85FFD3FF93D099CEB5FE00F81C" box="[400,502,1951,1975]" pageId="0" pageNumber="53" type="holotype">Holotype</typeStatus>
</emphasis>
: male (sl
<quantity id="4C8CB9C2FFD3FF93D37DCEB5FD16F81C" box="[628,736,1951,1975]" metricMagnitude="-2" metricUnit="m" metricValue="1.3699999999999999" pageId="0" pageNumber="53" unit="mm" value="13.7">13.7 mm</quantity>
), off Sakthikulangara, Kerala State,
<collectingCountry id="F36354B7FFD3FF93D584CEB5FB3FF81C" box="[1165,1225,1951,1975]" name="India" pageId="0" pageNumber="53">India</collectingCountry>
, depths
<quantity id="4C8CB9C2FFD3FF93D424CEB5FA6EF81C" box="[1325,1432,1951,1975]" metricMagnitude="1" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="0" pageNumber="53" unit="m" value="12.5" valueMax="15.0" valueMin="10.0">1015 m</quantity>
,
<date id="FFCA32E7FFD3FF93D19ECEE9FE8AF870" box="[151,380,1987,2012]" pageId="0" pageNumber="53" value="2011-01-24" valueMax="2011-01-29" valueMin="2011-01-24">2429 January 2011</date>
, CBM-ZC 10663.
</paragraph>
<caption id="DF0B44AFFFD2FF92D19ECE97FCEAF859" httpUri="https://zenodo.org/record/280764/files/figure.png" pageId="1" pageNumber="54" targetBox="[173,1430,151,1937]" targetPageId="1">
<paragraph id="8BCB1427FFD2FF92D19ECE97FCEAF859" blockId="1.[151,1436,1981,2034]" pageId="1" pageNumber="54">
<emphasis id="B900C835FFD2FF92D19ECE97FEF9F879" bold="true" box="[151,271,1981,2004]" pageId="1" pageNumber="54">FIGURE 1</emphasis>
.
<taxonomicName id="4C746FA4FFD2FF92D029CE97FE0BF878" box="[288,509,1981,2003]" class="Malacostraca" family="Diogenidae" genus="Ciliopagurus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="54" phylum="Arthropoda" rank="species" species="grandis" status="sp. nov.">
<emphasis id="B900C835FFD2FF92D029CE97FE0BF878" box="[288,509,1981,2003]" italics="true" pageId="1" pageNumber="54">Ciliopagurus grandis</emphasis>
</taxonomicName>
<emphasis id="B900C835FFD2FF92D30ECE94FDB2F878" bold="true" box="[519,580,1982,2003]" pageId="1" pageNumber="54">
<taxonomicNameLabel id="A233754EFFD2FF92D30ECE94FDB2F878" box="[519,580,1982,2003]" pageId="1" pageNumber="54" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype, male (sl 13.7 mm), CBM-ZC 10663, showing coloration preserved in formalin. A, entire animal in dorsal view; B, same, lateral view.
</paragraph>
</caption>
<paragraph id="8BCB1427FFD1FF91D1CEC9B3FACEFF5C" blockId="2.[151,1437,151,2012]" pageId="2" pageNumber="55">
<emphasis id="B900C835FFD1FF91D1CEC9B3FECFFF1B" box="[199,313,153,176]" italics="true" pageId="2" pageNumber="55">
<typeStatus id="54CFAA85FFD1FF91D1CEC9B3FECFFF1B" box="[199,313,153,176]" pageId="2" pageNumber="55" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<specimenCount id="9D72DFAEFFD1FF91D04EC9BDFE6BFF04" box="[327,413,151,175]" pageId="2" pageNumber="55" type="male">2 males</specimenCount>
(sl 9.6,
<quantity id="4C8CB9C2FFD1FF91D0F0C9BDFD97FF04" box="[505,609,151,175]" metricMagnitude="-2" metricUnit="m" metricValue="1.28" pageId="2" pageNumber="55" unit="mm" value="12.8">12.8 mm</quantity>
),
<specimenCount id="9D72DFAEFFD1FF91D37DC9BDFD23FF04" box="[628,725,151,175]" pageId="2" pageNumber="55" type="female">1 female</specimenCount>
(sl 10.0 mm), same data as
<typeStatus id="54CFAA85FFD1FF91D506C9BDFB82FF04" box="[1039,1140,151,175]" pageId="2" pageNumber="55" type="holotype">holotype</typeStatus>
, CBM-ZC 10664;
<specimenCount id="9D72DFAEFFD1FF91D44CC9BDFA6DFF04" box="[1349,1435,151,175]" pageId="2" pageNumber="55" type="male">2 males</specimenCount>
(sl 17.1, 14.0 mm),
<specimenCount id="9D72DFAEFFD1FF91D07DC996FE23FF7F" box="[372,469,188,212]" pageId="2" pageNumber="55" type="female">1 female</specimenCount>
(sl
<quantity id="4C8CB9C2FFD1FF91D0F6C996FDACFF7F" box="[511,602,188,212]" metricMagnitude="-3" metricUnit="m" metricValue="9.8" pageId="2" pageNumber="55" unit="mm" value="9.8">9.8 mm</quantity>
), off Sakthikulangara, Kerala State,
<collectingCountry id="F36354B7FFD1FF91D2E6C996FBDAFF7F" box="[1007,1068,188,212]" name="India" pageId="2" pageNumber="55">India</collectingCountry>
, depths
<quantity id="4C8CB9C2FFD1FF91D58EC996FB1AFF7F" box="[1159,1260,188,212]" metricMagnitude="1" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="2" pageNumber="55" unit="m" value="12.5" valueMax="15.0" valueMin="10.0">1015 m</quantity>
,
<date id="FFCA32E7FFD1FF91D5F1C996FF27FF5C" pageId="2" pageNumber="55" value="2011-01-24" valueMax="2011-01-29" valueMin="2011-01-24">2429 January 2011</date>
, Museum of the Department of Aquatic Biology and Fisheries, University of Kerala, AR AN 1416.
</paragraph>
</subSubSection>
<subSubSection id="C36E47ACFFD1FF94D1CEC829FBE5FBBB" lastPageId="7" lastPageNumber="60" pageId="2" pageNumber="55" type="description">
<paragraph id="8BCB1427FFD1FF91D1CEC829FBE6FE07" blockId="2.[151,1437,151,2012]" pageId="2" pageNumber="55">
<emphasis id="B900C835FFD1FF91D1CEC829FEA7FEB7" bold="true" box="[199,337,259,284]" pageId="2" pageNumber="55">Description</emphasis>
. Cephalothorax (
<figureCitation id="134F08A2FFD1FF91D312C82EFDA8FEB7" box="[539,606,260,284]" captionStart="FIGURE 1" captionStartId="1.[151,250,1981,2004]" captionTargetBox="[173,1430,151,1937]" captionTargetId="figure@1.[173,1437,141,1964]" captionTargetPageId="1" captionText="FIGURE 1. Ciliopagurus grandis n. sp., holotype, male (sl 13.7 mm), CBM-ZC 10663, showing coloration preserved in formalin. A, entire animal in dorsal view; B, same, lateral view." httpUri="https://zenodo.org/record/280764/files/figure.png" pageId="2" pageNumber="55">Fig. 1</figureCitation>
A, B) fairly depressed dorsoventrally. Shield (
<figureCitation id="134F08A2FFD1FF91D573C82EFB36FEB7" box="[1146,1216,260,284]" captionStart="FIGURE 2" captionStartId="3.[151,250,1875,1898]" captionTargetBox="[176,1400,184,1874]" captionTargetId="figure@3.[151,1415,157,1884]" captionTargetPageId="3" captionText="FIGURE 2. Ciliopagurus grandis n. sp., holotype, male (sl 13.7 mm), CBM-ZC 10663. A, shield and cephalic appendages, dorsal view (setae omitted from left side); B, left antennal acicle, dorsal view (setae omitted); C, endopod of left maxillule, lateral view; D, left third maxilliped, lateral view (setae omitted); E, same, ischium, ventral view; F, same, exopod, dorsal view (setae omitted); G, right fourth pereopod, lateral view; H, anterior lobe of sixth thoracic sternite, ventral view; I, sixth pleonal tergite, dorsal view; J, telson, dorsal view. Scale bars: 5 mm for A; 2 mm for H; 1 mm for B, D, F, G, I, J; 0.5 mm for C, E." httpUri="https://zenodo.org/record/280765/files/figure.png" pageId="2" pageNumber="55">Fig. 2</figureCitation>
A) about 1.1 times longer than broad; anterolateral margins sloping; anterior margin between rostrum and lateral projections very slightly concave; posterior margin roundly truncate; dorsal surface flat, with scattered shallow pits anterolaterally and few tufts of stiff setae laterally; Y-shaped decalcified suture present posteriorly. Rostrum obtuse, rounded, slightly exceeding lateral projections. Lateral projections also obtuse, unarmed.
</paragraph>
<paragraph id="8BCB1427FFD1FF91D1CEC89DFE95FDF4" blockId="2.[151,1437,151,2012]" pageId="2" pageNumber="55">
Ocular peduncles (
<figureCitation id="134F08A2FFD1FF91D096C89DFE15FE64" box="[415,483,439,463]" captionStart="FIGURE 2" captionStartId="3.[151,250,1875,1898]" captionTargetBox="[176,1400,184,1874]" captionTargetId="figure@3.[151,1415,157,1884]" captionTargetPageId="3" captionText="FIGURE 2. Ciliopagurus grandis n. sp., holotype, male (sl 13.7 mm), CBM-ZC 10663. A, shield and cephalic appendages, dorsal view (setae omitted from left side); B, left antennal acicle, dorsal view (setae omitted); C, endopod of left maxillule, lateral view; D, left third maxilliped, lateral view (setae omitted); E, same, ischium, ventral view; F, same, exopod, dorsal view (setae omitted); G, right fourth pereopod, lateral view; H, anterior lobe of sixth thoracic sternite, ventral view; I, sixth pleonal tergite, dorsal view; J, telson, dorsal view. Scale bars: 5 mm for A; 2 mm for H; 1 mm for B, D, F, G, I, J; 0.5 mm for C, E." httpUri="https://zenodo.org/record/280765/files/figure.png" pageId="2" pageNumber="55">Fig. 2</figureCitation>
A) equal in length, long and slender, 0.8 times as long as shield and 7.07.5 times longer than corneal width, cylindrical, slightly inflated basally and not dilated in corneal region; dorsal surfaces with scattered very short setae. Ocular acicles narrowly triangular, mesial margins nearly straight, lateral margins sloping, terminating in small marginal spine; narrowly separated basally (distance between acicles less than half width of 1 acicle).
</paragraph>
<paragraph id="8BCB1427FFD1FF91D1CECB46FAEDFD44" blockId="2.[151,1437,151,2012]" pageId="2" pageNumber="55">
Antennal peduncle (
<figureCitation id="134F08A2FFD1FF91D0A0CB46FE1DFD2F" box="[425,491,620,644]" captionStart="FIGURE 2" captionStartId="3.[151,250,1875,1898]" captionTargetBox="[176,1400,184,1874]" captionTargetId="figure@3.[151,1415,157,1884]" captionTargetPageId="3" captionText="FIGURE 2. Ciliopagurus grandis n. sp., holotype, male (sl 13.7 mm), CBM-ZC 10663. A, shield and cephalic appendages, dorsal view (setae omitted from left side); B, left antennal acicle, dorsal view (setae omitted); C, endopod of left maxillule, lateral view; D, left third maxilliped, lateral view (setae omitted); E, same, ischium, ventral view; F, same, exopod, dorsal view (setae omitted); G, right fourth pereopod, lateral view; H, anterior lobe of sixth thoracic sternite, ventral view; I, sixth pleonal tergite, dorsal view; J, telson, dorsal view. Scale bars: 5 mm for A; 2 mm for H; 1 mm for B, D, F, G, I, J; 0.5 mm for C, E." httpUri="https://zenodo.org/record/280765/files/figure.png" pageId="2" pageNumber="55">Fig. 2</figureCitation>
A) moderately long. Ultimate segment reaching corneal bases, 0.40.5 times as long as shield and about twice length of penultimate segment, dorsally with scattered minute setae. Penultimate segment with several short to moderately short setae. Basal segment short; ventrodistal margin produced in triangular process; laterodistal margin denticulate; lateral surface unarmed or armed with minute spinule dorsally.
</paragraph>
<paragraph id="8BCB1427FFD1FF91D19ECBD6FEC7FBFC" blockId="2.[151,1437,151,2012]" pageId="2" pageNumber="55">
Antennal peduncles (
<figureCitation id="134F08A2FFD1FF91D080CBD6FE38FCBF" box="[393,462,764,788]" captionStart="FIGURE 2" captionStartId="3.[151,250,1875,1898]" captionTargetBox="[176,1400,184,1874]" captionTargetId="figure@3.[151,1415,157,1884]" captionTargetPageId="3" captionText="FIGURE 2. Ciliopagurus grandis n. sp., holotype, male (sl 13.7 mm), CBM-ZC 10663. A, shield and cephalic appendages, dorsal view (setae omitted from left side); B, left antennal acicle, dorsal view (setae omitted); C, endopod of left maxillule, lateral view; D, left third maxilliped, lateral view (setae omitted); E, same, ischium, ventral view; F, same, exopod, dorsal view (setae omitted); G, right fourth pereopod, lateral view; H, anterior lobe of sixth thoracic sternite, ventral view; I, sixth pleonal tergite, dorsal view; J, telson, dorsal view. Scale bars: 5 mm for A; 2 mm for H; 1 mm for B, D, F, G, I, J; 0.5 mm for C, E." httpUri="https://zenodo.org/record/280765/files/figure.png" pageId="2" pageNumber="55">Fig. 2</figureCitation>
A) reaching distal one-fourth to one-fifth of ocular peduncles. Fifth segment slightly flattened dorsoventrally, with scattered tufts of very short setae. Fourth segment unarmed or armed with minute spine dorsodistally, with short setae. Third segment with small spine at ventrodistal angle and some tufts of setae. Second segment with dorsolateral distal angle produced, terminating in simple or bifid spine, lateral margin unarmed; dorsomesial distal angle with minute spinule, mesial margin with stiff setae. First segment almost concealed by anterolateral margin of branchiostegite; ventrodistal margin with 1 minute spine lateral to antennal gland opening. Antennal acicle (
<figureCitation id="134F08A2FFD1FF91D304CAFEFDA5FC47" box="[525,595,980,1004]" captionStart="FIGURE 2" captionStartId="3.[151,250,1875,1898]" captionTargetBox="[176,1400,184,1874]" captionTargetId="figure@3.[151,1415,157,1884]" captionTargetPageId="3" captionText="FIGURE 2. Ciliopagurus grandis n. sp., holotype, male (sl 13.7 mm), CBM-ZC 10663. A, shield and cephalic appendages, dorsal view (setae omitted from left side); B, left antennal acicle, dorsal view (setae omitted); C, endopod of left maxillule, lateral view; D, left third maxilliped, lateral view (setae omitted); E, same, ischium, ventral view; F, same, exopod, dorsal view (setae omitted); G, right fourth pereopod, lateral view; H, anterior lobe of sixth thoracic sternite, ventral view; I, sixth pleonal tergite, dorsal view; J, telson, dorsal view. Scale bars: 5 mm for A; 2 mm for H; 1 mm for B, D, F, G, I, J; 0.5 mm for C, E." httpUri="https://zenodo.org/record/280765/files/figure.png" pageId="2" pageNumber="55">Fig. 2</figureCitation>
B) reaching proximal two-fifths of ocular peduncle, terminating in acute, simple or bifid spine; mesial margin with 1 or 2 tiny spines or spinules, lateral margin unarmed or 1 tiny spine. Flagellum slightly overreaching chelipeds; each article with some minute setae (&lt;half length of one article) on distal margin.
</paragraph>
<paragraph id="8BCB1427FFD1FF91D1CECD4EFDD7FADC" blockId="2.[151,1437,151,2012]" pageId="2" pageNumber="55">
Mandible (not illustrated) without distinguishing characters. Maxillule with endopod bearing 7 bristles on moderately well-developed internal lobe, external lobe well-developed, slender, recurved (
<figureCitation id="134F08A2FFD1FF91D591CDADFB2AFB34" box="[1176,1244,1159,1183]" captionStart="FIGURE 2" captionStartId="3.[151,250,1875,1898]" captionTargetBox="[176,1400,184,1874]" captionTargetId="figure@3.[151,1415,157,1884]" captionTargetPageId="3" captionText="FIGURE 2. Ciliopagurus grandis n. sp., holotype, male (sl 13.7 mm), CBM-ZC 10663. A, shield and cephalic appendages, dorsal view (setae omitted from left side); B, left antennal acicle, dorsal view (setae omitted); C, endopod of left maxillule, lateral view; D, left third maxilliped, lateral view (setae omitted); E, same, ischium, ventral view; F, same, exopod, dorsal view (setae omitted); G, right fourth pereopod, lateral view; H, anterior lobe of sixth thoracic sternite, ventral view; I, sixth pleonal tergite, dorsal view; J, telson, dorsal view. Scale bars: 5 mm for A; 2 mm for H; 1 mm for B, D, F, G, I, J; 0.5 mm for C, E." httpUri="https://zenodo.org/record/280765/files/figure.png" pageId="2" pageNumber="55">Fig. 2</figureCitation>
C). Maxilla (
<figureCitation id="134F08A2FFD1FF91D467CDADFF52FB6F" captionStart="FIGURE 3" captionStartId="4.[151,250,906,929]" captionTargetBox="[175,1404,187,854]" captionTargetId="figure@4.[151,1436,173,885]" captionTargetPageId="4" captionText="FIGURE 3. Ciliopagurus grandis n. sp., holotype, male (sl 13.7 mm), CBM-ZC 10663. A, left maxilla, outer view (setae on scaphognathite omitted) l B, left first maxilliped, outer view; C, left second maxilliped, outer view. Scale bars: 2 mm." httpUri="https://zenodo.org/record/280766/files/figure.png" pageId="2" pageNumber="55">Fig. 3</figureCitation>
A) with basally broad, tapering endopod reaching scaphognathite in distal extension. First maxilliped (
<figureCitation id="134F08A2FFD1FF91D435CD86FA89FB6F" box="[1340,1407,1196,1220]" captionStart="FIGURE 3" captionStartId="4.[151,250,906,929]" captionTargetBox="[175,1404,187,854]" captionTargetId="figure@4.[151,1436,173,885]" captionTargetPageId="4" captionText="FIGURE 3. Ciliopagurus grandis n. sp., holotype, male (sl 13.7 mm), CBM-ZC 10663. A, left maxilla, outer view (setae on scaphognathite omitted) l B, left first maxilliped, outer view; C, left second maxilliped, outer view. Scale bars: 2 mm." httpUri="https://zenodo.org/record/280766/files/figure.png" pageId="2" pageNumber="55">Fig. 3</figureCitation>
B) with basal segment of exopod broad. Second maxilliped (
<figureCitation id="134F08A2FFD1FF91D221CDE5FC9DFB4C" box="[808,875,1231,1255]" captionStart="FIGURE 3" captionStartId="4.[151,250,906,929]" captionTargetBox="[175,1404,187,854]" captionTargetId="figure@4.[151,1436,173,885]" captionTargetPageId="4" captionText="FIGURE 3. Ciliopagurus grandis n. sp., holotype, male (sl 13.7 mm), CBM-ZC 10663. A, left maxilla, outer view (setae on scaphognathite omitted) l B, left first maxilliped, outer view; C, left second maxilliped, outer view. Scale bars: 2 mm." httpUri="https://zenodo.org/record/280766/files/figure.png" pageId="2" pageNumber="55">Fig. 3</figureCitation>
C) with basis-ischium fusion incomplete; exopod about twice of endopod in length, moderately broad. Third maxilliped (
<figureCitation id="134F08A2FFD1FF91D2B3CDDEFC0AFAA7" box="[954,1020,1268,1292]" captionStart="FIGURE 2" captionStartId="3.[151,250,1875,1898]" captionTargetBox="[176,1400,184,1874]" captionTargetId="figure@3.[151,1415,157,1884]" captionTargetPageId="3" captionText="FIGURE 2. Ciliopagurus grandis n. sp., holotype, male (sl 13.7 mm), CBM-ZC 10663. A, shield and cephalic appendages, dorsal view (setae omitted from left side); B, left antennal acicle, dorsal view (setae omitted); C, endopod of left maxillule, lateral view; D, left third maxilliped, lateral view (setae omitted); E, same, ischium, ventral view; F, same, exopod, dorsal view (setae omitted); G, right fourth pereopod, lateral view; H, anterior lobe of sixth thoracic sternite, ventral view; I, sixth pleonal tergite, dorsal view; J, telson, dorsal view. Scale bars: 5 mm for A; 2 mm for H; 1 mm for B, D, F, G, I, J; 0.5 mm for C, E." httpUri="https://zenodo.org/record/280765/files/figure.png" pageId="2" pageNumber="55">Fig. 2</figureCitation>
D) with basis and ischium separated; basis with 2 tiny corneous-tipped denticles; ischium with crista dentata well-developed, without accessory tooth (
<figureCitation id="134F08A2FFD1FF91D196CC16FF11FAFF" box="[159,231,1340,1364]" captionStart="FIGURE 2" captionStartId="3.[151,250,1875,1898]" captionTargetBox="[176,1400,184,1874]" captionTargetId="figure@3.[151,1415,157,1884]" captionTargetPageId="3" captionText="FIGURE 2. Ciliopagurus grandis n. sp., holotype, male (sl 13.7 mm), CBM-ZC 10663. A, shield and cephalic appendages, dorsal view (setae omitted from left side); B, left antennal acicle, dorsal view (setae omitted); C, endopod of left maxillule, lateral view; D, left third maxilliped, lateral view (setae omitted); E, same, ischium, ventral view; F, same, exopod, dorsal view (setae omitted); G, right fourth pereopod, lateral view; H, anterior lobe of sixth thoracic sternite, ventral view; I, sixth pleonal tergite, dorsal view; J, telson, dorsal view. Scale bars: 5 mm for A; 2 mm for H; 1 mm for B, D, F, G, I, J; 0.5 mm for C, E." httpUri="https://zenodo.org/record/280765/files/figure.png" pageId="2" pageNumber="55">Fig. 2</figureCitation>
E); merus and carpus unarmed on dorsodistal margins; exopod broad (broadest at proximal one-third), uncalcified in lateral half (
<figureCitation id="134F08A2FFD1FF91D0B6CC75FDF4FADC" box="[447,514,1375,1399]" captionStart="FIGURE 2" captionStartId="3.[151,250,1875,1898]" captionTargetBox="[176,1400,184,1874]" captionTargetId="figure@3.[151,1415,157,1884]" captionTargetPageId="3" captionText="FIGURE 2. Ciliopagurus grandis n. sp., holotype, male (sl 13.7 mm), CBM-ZC 10663. A, shield and cephalic appendages, dorsal view (setae omitted from left side); B, left antennal acicle, dorsal view (setae omitted); C, endopod of left maxillule, lateral view; D, left third maxilliped, lateral view (setae omitted); E, same, ischium, ventral view; F, same, exopod, dorsal view (setae omitted); G, right fourth pereopod, lateral view; H, anterior lobe of sixth thoracic sternite, ventral view; I, sixth pleonal tergite, dorsal view; J, telson, dorsal view. Scale bars: 5 mm for A; 2 mm for H; 1 mm for B, D, F, G, I, J; 0.5 mm for C, E." httpUri="https://zenodo.org/record/280765/files/figure.png" pageId="2" pageNumber="55">Fig. 2</figureCitation>
F).
</paragraph>
<paragraph id="8BCB1427FFD1FF97D1CECCAEFD1EFB13" blockId="2.[151,1437,151,2012]" lastBlockId="4.[151,1437,1004,1924]" lastPageId="4" lastPageNumber="57" pageId="2" pageNumber="55">
Chelipeds slightly unequal, short, left larger (
<figureCitation id="134F08A2FFD1FF91D3D9CCAEFCE7FA37" box="[720,785,1412,1436]" captionStart="FIGURE 1" captionStartId="1.[151,250,1981,2004]" captionTargetBox="[173,1430,151,1937]" captionTargetId="figure@1.[173,1437,141,1964]" captionTargetPageId="1" captionText="FIGURE 1. Ciliopagurus grandis n. sp., holotype, male (sl 13.7 mm), CBM-ZC 10663, showing coloration preserved in formalin. A, entire animal in dorsal view; B, same, lateral view." httpUri="https://zenodo.org/record/280764/files/figure.png" pageId="2" pageNumber="55">Fig. 1</figureCitation>
A). Left cheliped (
<figureCitation id="134F08A2FFD1FF91D2E2CCAEFBD9FA37" box="[1003,1071,1412,1436]" captionStart="FIGURE 4" captionStartId="5.[151,250,1920,1943]" captionTargetBox="[194,1399,185,1907]" captionTargetId="figure@5.[168,1428,159,1908]" captionTargetPageId="5" captionText="FIGURE 4. Ciliopagurus grandis n. sp., holotype, male (sl 13.7 mm), CBM-ZC 10663. A, left cheliped, lateral view; B, left chela and carpus, dorsal view; C, left cheliped, mesial view; D, close up of transverse ridges or striae on dorsal surface near base of fixed finger (setae omitted); E, close up of distal stridulating ridges on mesial surface of palm, setae partially shown. Scale bars: 5 mm for A C; 1 mm for D, E." httpUri="https://zenodo.org/record/280767/files/figure.png" pageId="2" pageNumber="55">Fig. 4</figureCitation>
AC) with dactylus subequal in length to palm; cutting edge with 13 prominent calcareous teeth proximally, terminating in very strong, black corneous claw; dorsomesial margin not markedly delimited, dorsal and mesial surface with 5 irregular, shallow, transverse furrows and irregular rows of small, corneous-tipped spines or transverse rows of corneous-tipped rods forming stridulating apparatus (stridulating rods restricted to mesial surface), these armature partially obscured by stiff setae; ventral surface with several tufts of stiff setae. Palm slightly longer than carpus; dorsolateral or lateral margin not delimited, dorsal, lateral and mesial surfaces each marked by shallow transverse furrows, each furrow on dorsal and lateral surfaces bounded proximally by low, setose, microscopically denticulate ridge (
<figureCitation id="134F08A2FFD1FF91D5FCCF55FAC1F93C" box="[1269,1335,1663,1687]" captionStart="FIGURE 4" captionStartId="5.[151,250,1920,1943]" captionTargetBox="[194,1399,185,1907]" captionTargetId="figure@5.[168,1428,159,1908]" captionTargetPageId="5" captionText="FIGURE 4. Ciliopagurus grandis n. sp., holotype, male (sl 13.7 mm), CBM-ZC 10663. A, left cheliped, lateral view; B, left chela and carpus, dorsal view; C, left cheliped, mesial view; D, close up of transverse ridges or striae on dorsal surface near base of fixed finger (setae omitted); E, close up of distal stridulating ridges on mesial surface of palm, setae partially shown. Scale bars: 5 mm for A C; 1 mm for D, E." httpUri="https://zenodo.org/record/280767/files/figure.png" pageId="2" pageNumber="55">Fig. 4</figureCitation>
D) (main ridges
<quantity id="4C8CB9C2FFD1FF91D1EDCF8EFEDAF917" box="[228,300,1700,1724]" metricMagnitude="-1" metricUnit="m" metricValue="1.905" metricValueMax="2.286" metricValueMin="1.524" pageId="2" pageNumber="55" unit="in" value="7.5" valueMax="9.0" valueMin="6.0">69 in</quantity>
number on dorsal surfaces, some of them interrupted; some interspersing striae also present); fixed finger distally with scattered corneous-tipped spines interspersed with stiff setae and proximally with short transverse ridges bearing corneous spinules; furrows of mesial surface bounded proximally by transverse rows of corneous-tipped rods forming stridulating apparatus (for details, see below); ventral surface laterally with 4 or 5 transverse ridges consisting of small corneous tubercles and mesially with 4 transverse striae, and with short to long setae on ridges and striae. Carpus 0.60.7 times as long as merus, subtriangular; dorsal, lateral and mesial surfaces each shallow transverse furrows, and each furrow bounded proximally by low, microscopically denticulate or smooth ridges (5 or 6 main ridges present; corneous-tipped spines conspicuous on dorsal surface); distal margin microscopically denticulate dorsally and laterally; dorsodistal margin with small corneous-tipped spine. Merus subtriangular; dorsal margins, lateral and to lesser extent mesial surfaces each with shallow transverse furrows, bounded proximally by low, microscopically denticulate or smooth ridges, each ridge dense short setae marginally; dorsal margin broadly arcuate, without conspicuous spines, dorsodistal margin with 24 small spines; mesial surface also with 1 small tubercle proximoventrally, ventromesial margin with 04 small tubercles; ventral surface with 1 distinct subdistal transverse ridge and some short transverse striae, lacking conspicuous spine. Ischium with ventromesial margin smooth to minutely denticulate.
</paragraph>
<caption id="DF0B44AFFFD0FF90D19ECE79FA87F84F" httpUri="https://zenodo.org/record/280765/files/figure.png" pageId="3" pageNumber="56" targetBox="[176,1400,184,1874]" targetPageId="3">
<paragraph id="8BCB1427FFD0FF90D19ECE79FA87F84F" blockId="3.[151,1436,1875,2020]" pageId="3" pageNumber="56">
<emphasis id="B900C835FFD0FF90D19ECE79FEFBF8C2" bold="true" box="[151,269,1875,1898]" pageId="3" pageNumber="56">FIGURE 2</emphasis>
.
<taxonomicName id="4C746FA4FFD0FF90D015CE7EFE01F8C1" box="[284,503,1876,1898]" class="Malacostraca" family="Diogenidae" genus="Ciliopagurus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="56" phylum="Arthropoda" rank="species" species="grandis" status="sp. nov.">
<emphasis id="B900C835FFD0FF90D015CE7EFE01F8C1" box="[284,503,1876,1898]" italics="true" pageId="3" pageNumber="56">Ciliopagurus grandis</emphasis>
</taxonomicName>
<emphasis id="B900C835FFD0FF90D0F6CE7EFDCFF8C2" bold="true" box="[511,569,1876,1897]" pageId="3" pageNumber="56">
<taxonomicNameLabel id="A233754EFFD0FF90D0F6CE7EFDCFF8C2" box="[511,569,1876,1897]" pageId="3" pageNumber="56" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype, male (sl 13.7 mm), CBM-ZC 10663. A, shield and cephalic appendages, dorsal view (setae omitted from left side); B, left antennal acicle, dorsal view (setae omitted); C, endopod of left maxillule, lateral view; D, left third maxilliped, lateral view (setae omitted); E, same, ischium, ventral view; F, same, exopod, dorsal view (setae omitted); G, right fourth pereopod, lateral view; H, anterior lobe of sixth thoracic sternite, ventral view; I, sixth pleonal tergite, dorsal view; J, telson, dorsal view. Scale bars: 5 mm for A; 2 mm for H; 1 mm for B, D, F, G, I, J; 0.5 mm for C, E.
</paragraph>
</caption>
<caption id="DF0B44AFFFD7FF97D19ECAA0FAC1FC14" httpUri="https://zenodo.org/record/280766/files/figure.png" pageId="4" pageNumber="57" targetBox="[175,1404,187,854]" targetPageId="4">
<paragraph id="8BCB1427FFD7FF97D19ECAA0FAC1FC14" blockId="4.[151,1436,906,959]" pageId="4" pageNumber="57">
<emphasis id="B900C835FFD7FF97D19ECAA0FEFBFC0B" bold="true" box="[151,269,906,929]" pageId="4" pageNumber="57">FIGURE 3</emphasis>
.
<taxonomicName id="4C746FA4FFD7FF97D012CAA1FE03FC0A" box="[283,501,907,929]" class="Malacostraca" family="Diogenidae" genus="Ciliopagurus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="57" phylum="Arthropoda" rank="species" species="grandis" status="sp. nov.">
<emphasis id="B900C835FFD7FF97D012CAA1FE03FC0A" box="[283,501,907,929]" italics="true" pageId="4" pageNumber="57">Ciliopagurus grandis</emphasis>
</taxonomicName>
<emphasis id="B900C835FFD7FF97D0F5CAA1FDC1FC0B" bold="true" box="[508,567,907,928]" pageId="4" pageNumber="57">
<taxonomicNameLabel id="A233754EFFD7FF97D0F5CAA1FDC1FC0B" box="[508,567,907,928]" pageId="4" pageNumber="57" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype, male (sl 13.7 mm), CBM-ZC 10663. A, left maxilla, outer view (setae on scaphognathite omitted)l B, left first maxilliped, outer view; C, left second maxilliped, outer view. Scale bars: 2 mm.
</paragraph>
</caption>
<paragraph id="8BCB1427FFD7FF97D1CECDEEFC20FB77" blockId="4.[151,1437,1004,1924]" box="[199,982,1220,1244]" pageId="4" pageNumber="57">Right cheliped similar to left cheliped in armature and ornamentation.</paragraph>
<paragraph id="8BCB1427FFD7FF97D1CECDC2FC58FAC7" blockId="4.[151,1437,1004,1924]" pageId="4" pageNumber="57">
Stridulating apparatus consisting of 4 main ridges extending to ventral half of mesial surface (occasionally interrupted) and interspersing 4 or 5 short dorsal ridges or elevations bearing longitudinal corneous rods (most rods each terminating distally in small spine) (
<figureCitation id="134F08A2FFD7FF97D365CC1AFD47FAE3" box="[620,689,1328,1352]" captionStart="FIGURE 4" captionStartId="5.[151,250,1920,1943]" captionTargetBox="[194,1399,185,1907]" captionTargetId="figure@5.[168,1428,159,1908]" captionTargetPageId="5" captionText="FIGURE 4. Ciliopagurus grandis n. sp., holotype, male (sl 13.7 mm), CBM-ZC 10663. A, left cheliped, lateral view; B, left chela and carpus, dorsal view; C, left cheliped, mesial view; D, close up of transverse ridges or striae on dorsal surface near base of fixed finger (setae omitted); E, close up of distal stridulating ridges on mesial surface of palm, setae partially shown. Scale bars: 5 mm for A C; 1 mm for D, E." httpUri="https://zenodo.org/record/280767/files/figure.png" pageId="4" pageNumber="57">Fig. 4</figureCitation>
C). Distalmost ridge bearing about 20 corneous rods decreasing in length ventrally (
<figureCitation id="134F08A2FFD7FF97D05FCC7EFE6CFAC7" box="[342,410,1364,1388]" captionStart="FIGURE 4" captionStartId="5.[151,250,1920,1943]" captionTargetBox="[194,1399,185,1907]" captionTargetId="figure@5.[168,1428,159,1908]" captionTargetPageId="5" captionText="FIGURE 4. Ciliopagurus grandis n. sp., holotype, male (sl 13.7 mm), CBM-ZC 10663. A, left cheliped, lateral view; B, left chela and carpus, dorsal view; C, left cheliped, mesial view; D, close up of transverse ridges or striae on dorsal surface near base of fixed finger (setae omitted); E, close up of distal stridulating ridges on mesial surface of palm, setae partially shown. Scale bars: 5 mm for A C; 1 mm for D, E." httpUri="https://zenodo.org/record/280767/files/figure.png" pageId="4" pageNumber="57">Fig. 4</figureCitation>
E). In total, about 5060 corneous rods present.
</paragraph>
<paragraph id="8BCB1427FFD7FF97D1CECC52FD7AF82F" blockId="4.[151,1437,1004,1924]" pageId="4" pageNumber="57">
Ambulatory legs (
<figureCitation id="134F08A2FFD7FF97D09BCC52FE23FA3B" box="[402,469,1400,1424]" captionStart="FIGURE 5" captionStartId="6.[151,250,1950,1973]" captionTargetBox="[153,1427,184,1949]" captionTargetId="figure@6.[151,1436,163,1963]" captionTargetPageId="6" captionText="FIGURE 5. Ciliopagurus grandis n. sp., holotype, male (sl 13.7 mm), CBM-ZC 10663. A, right second pereopod, lateral view (only setae on dactylus shown); B, same, mesial view (setae omitted); C, same, dactylus, ventral view; D, right third pereopod, lateral view (setae omitted); E, same, mesial view. Scale bars: 5 mm." httpUri="https://zenodo.org/record/280768/files/figure.png" pageId="4" pageNumber="57">Fig. 5</figureCitation>
A, B, D, F) moderately long, second overreaching tips of chelipeds by 0.50.7 lengths of dactyli; generally similar in armature and ornamentation. Dactyli 1.11.3 times longer than propodi, terminating in strong terminal claws, in lateral view gently curved ventrally, in dorsal view nearly straight; dorsal surfaces each with low transverse ridges, showing as arcuate or squamiform ridges on dorsal halves of lateral and mesial surfaces, and stiff short to long setae arising from these ridges; lateral and mesial surfaces each with row of arcuate or squamiform ridges on ventral half and tufts of stiff setae; ventral surfaces with row of 1520 slender corneous spines increasing in size distally (cf.
<figureCitation id="134F08A2FFD7FF97D33ECF6AFD8CF9F3" box="[567,634,1600,1624]" captionStart="FIGURE 5" captionStartId="6.[151,250,1950,1973]" captionTargetBox="[153,1427,184,1949]" captionTargetId="figure@6.[151,1436,163,1963]" captionTargetPageId="6" captionText="FIGURE 5. Ciliopagurus grandis n. sp., holotype, male (sl 13.7 mm), CBM-ZC 10663. A, right second pereopod, lateral view (only setae on dactylus shown); B, same, mesial view (setae omitted); C, same, dactylus, ventral view; D, right third pereopod, lateral view (setae omitted); E, same, mesial view. Scale bars: 5 mm." httpUri="https://zenodo.org/record/280768/files/figure.png" pageId="4" pageNumber="57">Fig. 5</figureCitation>
B). Propodi each with series of prominent ridges and rows of short fine stiff setae circumscribing segment (6 main ridges and a few interspersed, short, occasionally interrupted ridges); dorsal and ventral surfaces each with additional tufts of long stiff setae. Carpi each with corneous-tipped dorsodistal spine; dorsal and lateral, and to lesser extent, mesial surfaces each with series of transverse ridges and rows of short fine setae; dorsal ridges with 13 minute to tiny corneous-tipped spines in second, unarmed in third. Meri with dorsal surfaces each having row of low transverse ridges extending laterally and mesially, all ridges interrupted, bearing fine short setae; dorsal surface also with sparse long setae proximally; ventral surfaces each with spinulose ridges (second) or almost smooth ridges (third) and additional tufts of long stiff setae, ventrolateral distal margin with small distal spine in second, unarmed in third. Ischium with ventral margin bearing row of minute spinules (second) or unarmed (third).
</paragraph>
<caption id="DF0B44AFFFD6FF96D19ECEAAFDB1F858" httpUri="https://zenodo.org/record/280767/files/figure.png" pageId="5" pageNumber="58" targetBox="[194,1399,185,1907]" targetPageId="5">
<paragraph id="8BCB1427FFD6FF96D19ECEAAFDB1F858" blockId="5.[151,1437,1920,2035]" pageId="5" pageNumber="58">
<emphasis id="B900C835FFD6FF96D19ECEAAFEFBF83D" bold="true" box="[151,269,1920,1943]" pageId="5" pageNumber="58">FIGURE 4</emphasis>
.
<taxonomicName id="4C746FA4FFD6FF96D012CEABFE02F83C" box="[283,500,1921,1943]" class="Malacostraca" family="Diogenidae" genus="Ciliopagurus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="58" phylum="Arthropoda" rank="species" species="grandis" status="sp. nov.">
<emphasis id="B900C835FFD6FF96D012CEABFE02F83C" box="[283,500,1921,1943]" italics="true" pageId="5" pageNumber="58">Ciliopagurus grandis</emphasis>
</taxonomicName>
<emphasis id="B900C835FFD6FF96D0F5CEABFDC0F83D" bold="true" box="[508,566,1921,1942]" pageId="5" pageNumber="58">
<taxonomicNameLabel id="A233754EFFD6FF96D0F5CEABFDC0F83D" box="[508,566,1921,1942]" pageId="5" pageNumber="58" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype, male (sl 13.7 mm), CBM-ZC 10663. A, left cheliped, lateral view; B, left chela and carpus, dorsal view; C, left cheliped, mesial view; D, close up of transverse ridges or striae on dorsal surface near base of fixed finger (setae omitted); E, close up of distal stridulating ridges on mesial surface of palm, setae partially shown. Scale bars: 5 mm for AC; 1 mm for D, E.
</paragraph>
</caption>
<caption id="DF0B44AFFFD5FF95D19ECEB4FCA6F859" httpUri="https://zenodo.org/record/280768/files/figure.png" pageId="6" pageNumber="59" targetBox="[153,1427,184,1949]" targetPageId="6">
<paragraph id="8BCB1427FFD5FF95D19ECEB4FCA6F859" blockId="6.[151,1437,1950,2034]" pageId="6" pageNumber="59">
<emphasis id="B900C835FFD5FF95D19ECEB4FEFDF81F" bold="true" box="[151,267,1950,1973]" pageId="6" pageNumber="59">FIGURE 5</emphasis>
.
<taxonomicName id="4C746FA4FFD5FF95D011CEB5FE07F81E" box="[280,497,1951,1973]" class="Malacostraca" family="Diogenidae" genus="Ciliopagurus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="59" phylum="Arthropoda" rank="species" species="grandis" status="sp. nov.">
<emphasis id="B900C835FFD5FF95D011CEB5FE07F81E" box="[280,497,1951,1973]" italics="true" pageId="6" pageNumber="59">Ciliopagurus grandis</emphasis>
</taxonomicName>
<emphasis id="B900C835FFD5FF95D0FECEB5FDC6F81F" bold="true" box="[503,560,1951,1972]" pageId="6" pageNumber="59">
<taxonomicNameLabel id="A233754EFFD5FF95D0FECEB5FDC6F81F" box="[503,560,1951,1972]" pageId="6" pageNumber="59" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype, male (sl 13.7 mm), CBM-ZC 10663. A, right second pereopod, lateral view (only setae on dactylus shown); B, same, mesial view (setae omitted); C, same, dactylus, ventral view; D, right third pereopod, lateral view (setae omitted); E, same, mesial view. Scale bars: 5 mm.
</paragraph>
</caption>
<paragraph id="8BCB1427FFD4FF94D1CEC9BDFF15FECF" blockId="7.[151,1436,151,2013]" pageId="7" pageNumber="60">
Fourth pereopods (
<figureCitation id="134F08A2FFD4FF94D0AFC9BDFE18FF04" box="[422,494,151,175]" captionStart="FIGURE 2" captionStartId="3.[151,250,1875,1898]" captionTargetBox="[176,1400,184,1874]" captionTargetId="figure@3.[151,1415,157,1884]" captionTargetPageId="3" captionText="FIGURE 2. Ciliopagurus grandis n. sp., holotype, male (sl 13.7 mm), CBM-ZC 10663. A, shield and cephalic appendages, dorsal view (setae omitted from left side); B, left antennal acicle, dorsal view (setae omitted); C, endopod of left maxillule, lateral view; D, left third maxilliped, lateral view (setae omitted); E, same, ischium, ventral view; F, same, exopod, dorsal view (setae omitted); G, right fourth pereopod, lateral view; H, anterior lobe of sixth thoracic sternite, ventral view; I, sixth pleonal tergite, dorsal view; J, telson, dorsal view. Scale bars: 5 mm for A; 2 mm for H; 1 mm for B, D, F, G, I, J; 0.5 mm for C, E." httpUri="https://zenodo.org/record/280765/files/figure.png" pageId="7" pageNumber="60">Fig. 2</figureCitation>
G) semichelate; dactylus distinctly longer than palm, somewhat curved distally, overreaching tip of fixed finger by 0.2 length, terminating in small corneous claw, lateral surface with row of corneous spines ventrally; preungual process absent. Propodal rasp very well developed, encompassing half to twothirds of lateral surface, consisting of small to elongate scales increasing in size distally. Carpus with or without dorsodistal spine. Tufts of stiff long setae present on dorsal surfaces of dactylus to merus and ventral surfaces of merus.
</paragraph>
<paragraph id="8BCB1427FFD4FF94D1CEC845FE23FE2C" blockId="7.[151,1436,151,2013]" box="[199,469,367,391]" pageId="7" pageNumber="60">Fifth pereopods chelate.</paragraph>
<paragraph id="8BCB1427FFD4FF94D1CEC8BEFCC7FE64" blockId="7.[151,1436,151,2013]" pageId="7" pageNumber="60">
Anterior lobe of sixth thoracic sternite (of third pereopods) (
<figureCitation id="134F08A2FFD4FF94D264C8BEFC46FE07" box="[877,944,404,428]" captionStart="FIGURE 2" captionStartId="3.[151,250,1875,1898]" captionTargetBox="[176,1400,184,1874]" captionTargetId="figure@3.[151,1415,157,1884]" captionTargetPageId="3" captionText="FIGURE 2. Ciliopagurus grandis n. sp., holotype, male (sl 13.7 mm), CBM-ZC 10663. A, shield and cephalic appendages, dorsal view (setae omitted from left side); B, left antennal acicle, dorsal view (setae omitted); C, endopod of left maxillule, lateral view; D, left third maxilliped, lateral view (setae omitted); E, same, ischium, ventral view; F, same, exopod, dorsal view (setae omitted); G, right fourth pereopod, lateral view; H, anterior lobe of sixth thoracic sternite, ventral view; I, sixth pleonal tergite, dorsal view; J, telson, dorsal view. Scale bars: 5 mm for A; 2 mm for H; 1 mm for B, D, F, G, I, J; 0.5 mm for C, E." httpUri="https://zenodo.org/record/280765/files/figure.png" pageId="7" pageNumber="60">Fig. 2</figureCitation>
H) broadly subsemicircular. Eighth thoracic sternite (of fifth pereopods) with shallow groove anteriorly.
</paragraph>
<paragraph id="8BCB1427FFD4FF94D1CEC8F6FC35FD97" blockId="7.[151,1436,151,2013]" pageId="7" pageNumber="60">Males with 4 unpaired left pleopods (second to fifth pleopods; length: fifth = fourth&gt; third&gt; second); each exopod well developed, while endopods greatly reduced, 0.20.3 length of exopods. Female with 4 biramous unpaired pleopods, with both rami multi-segmented and well developed.</paragraph>
<paragraph id="8BCB1427FFD4FF94D1CECB6DFD1AFD44" blockId="7.[151,1436,151,2013]" pageId="7" pageNumber="60">
Uropods asymmetrical. Sixth pleonal tergite (
<figureCitation id="134F08A2FFD4FF94D3CCCB6DFCFEFDF4" box="[709,776,583,607]" captionStart="FIGURE 2" captionStartId="3.[151,250,1875,1898]" captionTargetBox="[176,1400,184,1874]" captionTargetId="figure@3.[151,1415,157,1884]" captionTargetPageId="3" captionText="FIGURE 2. Ciliopagurus grandis n. sp., holotype, male (sl 13.7 mm), CBM-ZC 10663. A, shield and cephalic appendages, dorsal view (setae omitted from left side); B, left antennal acicle, dorsal view (setae omitted); C, endopod of left maxillule, lateral view; D, left third maxilliped, lateral view (setae omitted); E, same, ischium, ventral view; F, same, exopod, dorsal view (setae omitted); G, right fourth pereopod, lateral view; H, anterior lobe of sixth thoracic sternite, ventral view; I, sixth pleonal tergite, dorsal view; J, telson, dorsal view. Scale bars: 5 mm for A; 2 mm for H; 1 mm for B, D, F, G, I, J; 0.5 mm for C, E." httpUri="https://zenodo.org/record/280765/files/figure.png" pageId="7" pageNumber="60">
Fig.
<date id="FFCA32E7FFD4FF94D3F0CB6DFCFEFDF4" box="[761,776,583,607]" pageId="7" pageNumber="60">2</date>
</figureCitation>
I) about 1.5 times longer than broad, divided into two parts by distinct transverse groove across at posterior two-thirds; lateral indentations deep, narrow; posterior part with shallow median groove, posterolateral margins unarmed. Telson (
<figureCitation id="134F08A2FFD4FF94D27CCBA5FC41FD0C" box="[885,951,655,679]" captionStart="FIGURE 2" captionStartId="3.[151,250,1875,1898]" captionTargetBox="[176,1400,184,1874]" captionTargetId="figure@3.[151,1415,157,1884]" captionTargetPageId="3" captionText="FIGURE 2. Ciliopagurus grandis n. sp., holotype, male (sl 13.7 mm), CBM-ZC 10663. A, shield and cephalic appendages, dorsal view (setae omitted from left side); B, left antennal acicle, dorsal view (setae omitted); C, endopod of left maxillule, lateral view; D, left third maxilliped, lateral view (setae omitted); E, same, ischium, ventral view; F, same, exopod, dorsal view (setae omitted); G, right fourth pereopod, lateral view; H, anterior lobe of sixth thoracic sternite, ventral view; I, sixth pleonal tergite, dorsal view; J, telson, dorsal view. Scale bars: 5 mm for A; 2 mm for H; 1 mm for B, D, F, G, I, J; 0.5 mm for C, E." httpUri="https://zenodo.org/record/280765/files/figure.png" pageId="7" pageNumber="60">Fig. 2</figureCitation>
J) with posterior lobes broadly and roundly subtriangular, greatly unequal (left much larger than right), with narrow, deep median cleft; terminal margins unarmed, but with row of moderately long stiff setae.
</paragraph>
<paragraph id="8BCB1427FFD4FF94D1CECBD1FCFBFC63" blockId="7.[151,1436,151,2013]" pageId="7" pageNumber="60">
<emphasis id="B900C835FFD4FF94D1CECBD1FE20FCBF" bold="true" box="[199,470,763,788]" pageId="7" pageNumber="60">Coloration in formalin</emphasis>
. (
<figureCitation id="134F08A2FFD4FF94D0E4CBD6FDDAFCBF" box="[493,556,764,788]" captionStart="FIGURE 1" captionStartId="1.[151,250,1981,2004]" captionTargetBox="[173,1430,151,1937]" captionTargetId="figure@1.[173,1437,141,1964]" captionTargetPageId="1" captionText="FIGURE 1. Ciliopagurus grandis n. sp., holotype, male (sl 13.7 mm), CBM-ZC 10663, showing coloration preserved in formalin. A, entire animal in dorsal view; B, same, lateral view." httpUri="https://zenodo.org/record/280764/files/figure.png" pageId="7" pageNumber="60">Fig. 1</figureCitation>
A, B). Shield orange, anterior margin whitish. Posterior carapace pale orange; branchiostegite and ventrolateral portion of posterior carapace crimson. Ocular peduncle white on dorsal face, otherwise red. Antennular and antennal peduncles whitish, antennal flagellum uniformly pale orange. Chelipeds and ambulatory legs generally orange, transverse ridges lined by red. Pleon generally white, without markings; sixth somite, telson and uropods orange. Claws of chelipeds and ambulatory legs and corneous spinules on ambulatory dactyli black or dark brown. See
<figureCitation id="134F08A2FFD4FF94D399CA85FD38FC6C" box="[656,718,943,967]" captionStart="FIGURE 1" captionStartId="1.[151,250,1981,2004]" captionTargetBox="[173,1430,151,1937]" captionTargetId="figure@1.[173,1437,141,1964]" captionTargetPageId="1" captionText="FIGURE 1. Ciliopagurus grandis n. sp., holotype, male (sl 13.7 mm), CBM-ZC 10663, showing coloration preserved in formalin. A, entire animal in dorsal view; B, same, lateral view." httpUri="https://zenodo.org/record/280764/files/figure.png" pageId="7" pageNumber="60">Fig. 1</figureCitation>
A, B.
</paragraph>
<paragraph id="8BCB1427FFD4FF94D1CECAF9FBE5FBBB" blockId="7.[151,1436,151,2013]" pageId="7" pageNumber="60">
<emphasis id="B900C835FFD4FF94D1CECAF9FEDDFC47" bold="true" box="[199,299,979,1004]" pageId="7" pageNumber="60">Housing</emphasis>
. Using gastropod shells, e.g.,
<taxonomicName id="4C746FA4FFD4FF94D375CAFEFBE7FC47" authority="Linnaeus, 1767" authorityName="Linnaeus" authorityYear="1767" box="[636,1041,980,1004]" class="Gastropoda" family="Xenophoridae" genus="Xenophora" kingdom="Animalia" order="Littorinimorpha" pageId="7" pageNumber="60" phylum="Mollusca" rank="species" species="solaris">
<emphasis id="B900C835FFD4FF94D375CAFEFCBDFC47" box="[636,843,980,1004]" italics="true" pageId="7" pageNumber="60">Xenophora solaris</emphasis>
(Linnaeus, 1767)
</taxonomicName>
(Discopoda:
<taxonomicName id="4C746FA4FFD4FF94D5A1CAFEFABFFC47" box="[1192,1353,980,1004]" class="Gastropoda" family="Xenophoridae" kingdom="Animalia" order="Littorinimorpha" pageId="7" pageNumber="60" phylum="Mollusca" rank="family">Xenophoridae</taxonomicName>
),
<taxonomicName id="4C746FA4FFD4FF94D457CAFFFE58FBA4" authority="Sowerby, 1825" authorityName="Sowerby" authorityYear="1825" class="Gastropoda" family="Ficidae" genus="Ficus" kingdom="Animalia" order="Littorinimorpha" pageId="7" pageNumber="60" phylum="Mollusca" rank="species" species="gracilis">
<emphasis id="B900C835FFD4FF94D457CAFFFF1BFBBB" italics="true" pageId="7" pageNumber="60">Ficus gracilis</emphasis>
(Sowerby, 1825)
</taxonomicName>
and
<taxonomicName id="4C746FA4FFD4FF94D0EDCAD3FCECFBA4" authority="Lamarck, 1822" authorityName="Lamarck" authorityYear="1822" box="[484,794,1015,1040]" class="Gastropoda" family="Ficidae" genus="Ficus" kingdom="Animalia" order="Littorinimorpha" pageId="7" pageNumber="60" phylum="Mollusca" rank="species" species="ficoides">
<emphasis id="B900C835FFD4FF94D0EDCAD3FDA1FBBB" box="[484,599,1016,1040]" italics="true" pageId="7" pageNumber="60">F. ficoides</emphasis>
(Lamarck, 1822)
</taxonomicName>
(Discopoda:
<taxonomicName id="4C746FA4FFD4FF94D2B8CADDFBF3FBA4" box="[945,1029,1015,1039]" class="Gastropoda" family="Ficidae" kingdom="Animalia" order="Littorinimorpha" pageId="7" pageNumber="60" phylum="Mollusca" rank="family">Ficidae</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C36E47ACFFD4FF94D1CECD31FEF7FBFC" pageId="7" pageNumber="60" type="distribution">
<paragraph id="8BCB1427FFD4FF94D1CECD31FEF7FBFC" blockId="7.[151,1436,151,2013]" pageId="7" pageNumber="60">
<emphasis id="B900C835FFD4FF94D1CECD31FEAFFB9F" bold="true" box="[199,345,1051,1076]" pageId="7" pageNumber="60">Distribution</emphasis>
. So far known only from the
<typeStatus id="54CFAA85FFD4FF94D3A4CD37FD2BFB9E" box="[685,733,1053,1077]" pageId="7" pageNumber="60">type</typeStatus>
locality, off Sakthikulangara, Kerala State,
<collectingCountry id="F36354B7FFD4FF94D5C7CD36FAFCFB9F" box="[1230,1290,1052,1076]" name="India" pageId="7" pageNumber="60">India</collectingCountry>
, at depths of
<quantity id="4C8CB9C2FFD4FF94D19ECD15FF0BFBFC" box="[151,253,1087,1111]" metricMagnitude="1" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="7" pageNumber="60" unit="m" value="12.5" valueMax="15.0" valueMin="10.0">
10
<specimenCount id="9D72DFAEFFD4FF94D1CBCD15FF0BFBFC" box="[194,253,1087,1111]" pageId="7" pageNumber="60" type="generic">15 m</specimenCount>
</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C36E47ACFFD4FF9BD1CECD4EFAEBFD67" lastPageId="8" lastPageNumber="61" pageId="7" pageNumber="60" type="discussion">
<paragraph id="8BCB1427FFD4FF94D1CECD4EFB77F83E" blockId="7.[151,1436,151,2013]" pageId="7" pageNumber="60">
<emphasis id="B900C835FFD4FF94D1CECD4EFEC2FBD7" bold="true" box="[199,308,1124,1148]" pageId="7" pageNumber="60">Remarks</emphasis>
.
<taxonomicName id="4C746FA4FFD4FF94D04CCD4EFDCFFBD7" box="[325,569,1124,1148]" class="Malacostraca" family="Diogenidae" genus="Ciliopagurus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="60" phylum="Arthropoda" rank="species" species="grandis" status="sp. nov.">
<emphasis id="B900C835FFD4FF94D04CCD4EFDCFFBD7" box="[325,569,1124,1148]" italics="true" pageId="7" pageNumber="60">Ciliopagurus grandis</emphasis>
</taxonomicName>
<emphasis id="B900C835FFD4FF94D34ACD4EFD73FBD7" bold="true" box="[579,645,1124,1148]" pageId="7" pageNumber="60">
<taxonomicNameLabel id="A233754EFFD4FF94D34ACD4EFD73FBD7" box="[579,645,1124,1148]" pageId="7" pageNumber="60" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is similar to
<taxonomicName id="4C746FA4FFD4FF94D22DCD4FFBB0FBD7" authority="Forest, 1952" authorityName="Forest" authorityYear="1952" box="[804,1094,1124,1148]" class="Malacostraca" family="Diogenidae" genus="Ciliopagurus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="60" phylum="Arthropoda" rank="species" species="caparti">
<emphasis id="B900C835FFD4FF94D22DCD4FFC6CFBD7" box="[804,922,1125,1148]" italics="true" pageId="7" pageNumber="60">C. caparti</emphasis>
(Forest, 1952)
</taxonomicName>
from the eastern Atlantic,
<taxonomicName id="4C746FA4FFD4FF94D48BCD4FFE56FB34" authority="Forest, 1995" authorityName="Forest" authorityYear="1995" class="Malacostraca" family="Diogenidae" genus="Ciliopagurus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="60" phylum="Arthropoda" rank="species" species="albatrossi">
<emphasis id="B900C835FFD4FF94D48BCD4FFEF0FB0B" italics="true" pageId="7" pageNumber="60">C. albatrossi</emphasis>
Forest, 1995
</taxonomicName>
from Hawaii,
<taxonomicName id="4C746FA4FFD4FF94D346CDA2FC53FB34" authority="Lewinsohn, 1969" authorityName="Lewinsohn" authorityYear="1969" box="[591,933,1159,1184]" class="Malacostraca" family="Diogenidae" genus="Ciliopagurus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="60" phylum="Arthropoda" rank="species" species="shebae">
<emphasis id="B900C835FFD4FF94D346CDA2FD34FB0B" box="[591,706,1160,1184]" italics="true" pageId="7" pageNumber="60">C. shebae</emphasis>
(Lewinsohn, 1969)
</taxonomicName>
from the Indo-West Pacific,
<taxonomicName id="4C746FA4FFD4FF94D5F5CDA2FED2FB6F" authority="Forest, 1995" authorityName="Forest" authorityYear="1995" class="Malacostraca" family="Diogenidae" genus="Ciliopagurus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="60" phylum="Arthropoda" rank="species" species="macrolepis">
<emphasis id="B900C835FFD4FF94D5F5CDA2FA6DFB0B" box="[1276,1435,1160,1184]" italics="true" pageId="7" pageNumber="60">C. macrolepis</emphasis>
Forest, 1995
</taxonomicName>
from
<collectingCountry id="F36354B7FFD4FF94D063CD86FE2CFB6F" box="[362,474,1196,1220]" name="Indonesia" pageId="7" pageNumber="60">Indonesia</collectingCountry>
, and
<taxonomicName id="4C746FA4FFD4FF94D31FCD87FD04FB6F" authority="Forest, 1995" authorityName="Forest" authorityYear="1995" box="[534,754,1196,1220]" class="Malacostraca" family="Diogenidae" genus="Ciliopagurus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="60" phylum="Arthropoda" rank="species" species="liui">
<emphasis id="B900C835FFD4FF94D31FCD87FDA8FB6F" box="[534,606,1196,1220]" italics="true" pageId="7" pageNumber="60">C. liui</emphasis>
Forest, 1995
</taxonomicName>
from
<collectingCountry id="F36354B7FFD4FF94D231CD87FC8EFB6F" box="[824,888,1197,1220]" name="Japan" pageId="7" pageNumber="60">Japan</collectingCountry>
and
<collectingCountry id="F36354B7FFD4FF94D2B9CD86FC00FB6F" box="[944,1014,1196,1220]" name="China" pageId="7" pageNumber="60">China</collectingCountry>
. Shared characters are (Forest 1995): the ultimate segment of the antennular peduncle is more than 0.3 times as long as the shield; transverse ridges on the palms of the chelae are mostly minutely denticulate, lacking distinct tubercles or spines; the palms of the chelipeds have four or more main transverse ridges, interspersed by shorter striae; and the meri of the chelipeds are devoid of prominent tubercles or spines on the ventral surface. However,
<taxonomicName id="4C746FA4FFD4FF94D2E6CC17FB9FFAFF" box="[1007,1129,1340,1364]" class="Malacostraca" family="Diogenidae" genus="Ciliopagurus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="60" phylum="Arthropoda" rank="species" species="grandis" status="sp. nov.">
<emphasis id="B900C835FFD4FF94D2E6CC17FB9FFAFF" box="[1007,1129,1340,1364]" italics="true" pageId="7" pageNumber="60">C. grandis</emphasis>
</taxonomicName>
<emphasis id="B900C835FFD4FF94D57BCC16FB42FAFF" bold="true" box="[1138,1204,1340,1364]" pageId="7" pageNumber="60">
<taxonomicNameLabel id="A233754EFFD4FF94D57BCC16FB42FAFF" box="[1138,1204,1340,1364]" pageId="7" pageNumber="60" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is distinctive in the weakly armed antennal acicles, which bear at most one lateral and one or two mesial spines. In the other species, the antennal acicle is normally armed with five or more spines on the mesial margin and with two or more spines on the lateral margin. The simple ocular acicle distinguishes
<taxonomicName id="4C746FA4FFD4FF94D234CC82FC42FA6B" box="[829,948,1448,1472]" class="Malacostraca" family="Diogenidae" genus="Ciliopagurus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="60" phylum="Arthropoda" rank="species" species="grandis" status="sp. nov.">
<emphasis id="B900C835FFD4FF94D234CC82FC42FA6B" box="[829,948,1448,1472]" italics="true" pageId="7" pageNumber="60">C. grandis</emphasis>
</taxonomicName>
<emphasis id="B900C835FFD4FF94D2B3CC82FC0EFA6B" bold="true" box="[954,1016,1448,1472]" pageId="7" pageNumber="60">
<taxonomicNameLabel id="A233754EFFD4FF94D2B3CC82FC0EFA6B" box="[954,1016,1448,1472]" pageId="7" pageNumber="60" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
from
<taxonomicName id="4C746FA4FFD4FF94D534CC82FB3AFA6B" box="[1085,1228,1448,1472]" class="Malacostraca" family="Diogenidae" genus="Ciliopagurus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="60" phylum="Arthropoda" rank="species" species="albatrossi">
<emphasis id="B900C835FFD4FF94D534CC82FB3AFA6B" box="[1085,1228,1448,1472]" italics="true" pageId="7" pageNumber="60">C. albatrossi</emphasis>
</taxonomicName>
,
<taxonomicName id="4C746FA4FFD4FF94D5D3CC82FABEFA6B" box="[1242,1352,1448,1472]" class="Malacostraca" family="Diogenidae" genus="Ciliopagurus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="60" phylum="Arthropoda" rank="species" species="shebae">
<emphasis id="B900C835FFD4FF94D5D3CC82FABEFA6B" box="[1242,1352,1448,1472]" italics="true" pageId="7" pageNumber="60">C. shebae</emphasis>
</taxonomicName>
,
<taxonomicName id="4C746FA4FFD4FF94D45FCC82FA6AFA6B" box="[1366,1436,1448,1472]" class="Malacostraca" family="Diogenidae" genus="Ciliopagurus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="60" phylum="Arthropoda" rank="species" species="liui">
<emphasis id="B900C835FFD4FF94D45FCC82FA6AFA6B" box="[1366,1436,1448,1472]" italics="true" pageId="7" pageNumber="60">C. liui</emphasis>
</taxonomicName>
and
<taxonomicName id="4C746FA4FFD4FF94D1CFCCE7FE94FA4F" box="[198,354,1484,1508]" class="Malacostraca" family="Diogenidae" genus="Ciliopagurus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="60" phylum="Arthropoda" rank="species" species="macrolepis">
<emphasis id="B900C835FFD4FF94D1CFCCE7FE94FA4F" box="[198,354,1484,1508]" italics="true" pageId="7" pageNumber="60">C. macrolepis</emphasis>
</taxonomicName>
, as the ocular acicle is bifid in the latter four species. The relatively short antennal acicle, which only slightly overreaches the fourth segment of the antennal peduncle, differentiates
<taxonomicName id="4C746FA4FFD4FF94D579CCDAFB1CF9A3" box="[1136,1258,1520,1544]" class="Malacostraca" family="Diogenidae" genus="Ciliopagurus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="60" phylum="Arthropoda" rank="species" species="grandis" status="sp. nov.">
<emphasis id="B900C835FFD4FF94D579CCDAFB1CF9A3" box="[1136,1258,1520,1544]" italics="true" pageId="7" pageNumber="60">C. grandis</emphasis>
</taxonomicName>
<emphasis id="B900C835FFD4FF94D5FDCCDAFAC1F9A3" bold="true" box="[1268,1335,1520,1544]" pageId="7" pageNumber="60">
<taxonomicNameLabel id="A233754EFFD4FF94D5FDCCDAFAC1F9A3" box="[1268,1335,1520,1544]" pageId="7" pageNumber="60" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
from
<taxonomicName id="4C746FA4FFD4FF94D48BCCDAFF1EF987" class="Malacostraca" family="Diogenidae" genus="Ciliopagurus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="60" phylum="Arthropoda" rank="species" species="caparti">
<emphasis id="B900C835FFD4FF94D48BCCDAFF1EF987" italics="true" pageId="7" pageNumber="60">C. caparti</emphasis>
</taxonomicName>
,
<taxonomicName id="4C746FA4FFD4FF94D1FFCF3FFE7EF987" box="[246,392,1556,1580]" class="Malacostraca" family="Diogenidae" genus="Ciliopagurus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="60" phylum="Arthropoda" rank="species" species="macrolepi">
<emphasis id="B900C835FFD4FF94D1FFCF3FFE7EF987" box="[246,392,1556,1580]" italics="true" pageId="7" pageNumber="60">C. macrolepi</emphasis>
</taxonomicName>
s and
<taxonomicName id="4C746FA4FFD4FF94D0C0CF3FFDE6F987" box="[457,528,1556,1580]" class="Malacostraca" family="Diogenidae" genus="Ciliopagurus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="60" phylum="Arthropoda" rank="species" species="liui">
<emphasis id="B900C835FFD4FF94D0C0CF3FFDE6F987" box="[457,528,1556,1580]" italics="true" pageId="7" pageNumber="60">C. liui</emphasis>
</taxonomicName>
. In the latter three species, the antennal acicle reaches nearly to the midlength of the fourth segment of the antennal peduncle. The greatly unequal posterior lobes of the telson, of which the left lobe is roundly subtriangular, separates
<taxonomicName id="4C746FA4FFD4FF94D35ACF77FD3CF9DF" box="[595,714,1628,1652]" class="Malacostraca" family="Diogenidae" genus="Ciliopagurus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="60" phylum="Arthropoda" rank="species" species="grandis" status="sp. nov.">
<emphasis id="B900C835FFD4FF94D35ACF77FD3CF9DF" box="[595,714,1628,1652]" italics="true" pageId="7" pageNumber="60">C. grandis</emphasis>
</taxonomicName>
<emphasis id="B900C835FFD4FF94D3D9CF76FCF9F9DF" bold="true" box="[720,783,1628,1652]" pageId="7" pageNumber="60">
<taxonomicNameLabel id="A233754EFFD4FF94D3D9CF76FCF9F9DF" box="[720,783,1628,1652]" pageId="7" pageNumber="60" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
from
<taxonomicName id="4C746FA4FFD4FF94D25ACF77FC33F9DF" box="[851,965,1629,1652]" class="Malacostraca" family="Diogenidae" genus="Ciliopagurus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="60" phylum="Arthropoda" rank="species" species="caparti">
<emphasis id="B900C835FFD4FF94D25ACF77FC33F9DF" box="[851,965,1629,1652]" italics="true" pageId="7" pageNumber="60">C. caparti</emphasis>
</taxonomicName>
,
<taxonomicName id="4C746FA4FFD4FF94D2DBCF77FBB7F9DF" box="[978,1089,1628,1652]" class="Malacostraca" family="Diogenidae" genus="Ciliopagurus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="60" phylum="Arthropoda" rank="species" species="shebae">
<emphasis id="B900C835FFD4FF94D2DBCF77FBB7F9DF" box="[978,1089,1628,1652]" italics="true" pageId="7" pageNumber="60">C. shebae</emphasis>
</taxonomicName>
and
<taxonomicName id="4C746FA4FFD4FF94D57FCF77FB4BF9DF" box="[1142,1213,1628,1652]" class="Malacostraca" family="Diogenidae" genus="Ciliopagurus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="60" phylum="Arthropoda" rank="species" species="liui">
<emphasis id="B900C835FFD4FF94D57FCF77FB4BF9DF" box="[1142,1213,1628,1652]" italics="true" pageId="7" pageNumber="60">C. liui</emphasis>
</taxonomicName>
. Information on the coloration in life or in well-preserved condition is available for
<taxonomicName id="4C746FA4FFD4FF94D267CFAAFC17F933" box="[878,993,1664,1688]" class="Malacostraca" family="Diogenidae" genus="Ciliopagurus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="60" phylum="Arthropoda" rank="species" species="caparti">
<emphasis id="B900C835FFD4FF94D267CFAAFC17F933" box="[878,993,1664,1688]" italics="true" pageId="7" pageNumber="60">C. caparti</emphasis>
</taxonomicName>
,
<taxonomicName id="4C746FA4FFD4FF94D2F8CFAAFB97F933" box="[1009,1121,1664,1688]" class="Malacostraca" family="Diogenidae" genus="Ciliopagurus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="60" phylum="Arthropoda" rank="species" species="shebae">
<emphasis id="B900C835FFD4FF94D2F8CFAAFB97F933" box="[1009,1121,1664,1688]" italics="true" pageId="7" pageNumber="60">C. shebae</emphasis>
</taxonomicName>
and
<taxonomicName id="4C746FA4FFD4FF94D593CFAAFB17F933" box="[1178,1249,1664,1688]" class="Malacostraca" family="Diogenidae" genus="Ciliopagurus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="60" phylum="Arthropoda" rank="species" species="liui">
<emphasis id="B900C835FFD4FF94D593CFAAFB17F933" box="[1178,1249,1664,1688]" italics="true" pageId="7" pageNumber="60">C. liui</emphasis>
</taxonomicName>
, among the five species considered.
<taxonomicName id="4C746FA4FFD4FF94D073CF8EFD91F917" box="[378,615,1700,1724]" class="Malacostraca" family="Diogenidae" genus="Ciliopagurus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="60" phylum="Arthropoda" rank="species" species="caparti">
<emphasis id="B900C835FFD4FF94D073CF8EFD91F917" box="[378,615,1700,1724]" italics="true" pageId="7" pageNumber="60">Ciliopagurus caparti</emphasis>
</taxonomicName>
is distinctive in having longitudinal stripes on the shield (Forest 1995).
<taxonomicName id="4C746FA4FFD4FF94D19ECFE2FE77F94B" box="[151,385,1736,1760]" class="Malacostraca" family="Diogenidae" genus="Ciliopagurus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="60" phylum="Arthropoda" rank="species" species="shebae">
<emphasis id="B900C835FFD4FF94D19ECFE2FE77F94B" box="[151,385,1736,1760]" italics="true" pageId="7" pageNumber="60">Ciliopagurus shebae</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C746FA4FFD4FF94D310CFE2FD67F94B" box="[537,657,1736,1760]" class="Malacostraca" family="Diogenidae" genus="Ciliopagurus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="60" phylum="Arthropoda" rank="species" species="grandis" status="sp. nov.">
<emphasis id="B900C835FFD4FF94D310CFE2FD67F94B" box="[537,657,1736,1760]" italics="true" pageId="7" pageNumber="60">C. grandis</emphasis>
</taxonomicName>
<emphasis id="B900C835FFD4FF94D390CFE2FD2EF94B" bold="true" box="[665,728,1736,1760]" pageId="7" pageNumber="60">
<taxonomicNameLabel id="A233754EFFD4FF94D390CFE2FD2EF94B" box="[665,728,1736,1760]" pageId="7" pageNumber="60" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
in the dorsally white carapace (versus generally orange in the shield and pale orange in the posterior carapace), uniformly orange ocular peduncles (versus dorsally white and remainder red), uniformly whitish ambulatory dactyli (versus generally orange with red transverse ridges) and the dark reddish brown abdomen ornamented by white transverse bands (versus generally white).
<taxonomicName id="4C746FA4FFD4FF94D5D0CE1EFA6AF8E7" box="[1241,1436,1844,1868]" class="Malacostraca" family="Diogenidae" genus="Ciliopagurus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="60" phylum="Arthropoda" rank="species" species="liui">
<emphasis id="B900C835FFD4FF94D5D0CE1EFA6AF8E7" box="[1241,1436,1844,1868]" italics="true" pageId="7" pageNumber="60">Ciliopagurus liui</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C746FA4FFD4FF94D02ACE72FE6FF8DB" box="[291,409,1880,1904]" class="Malacostraca" family="Diogenidae" genus="Ciliopagurus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="60" phylum="Arthropoda" rank="species" species="grandis" status="sp. nov.">
<emphasis id="B900C835FFD4FF94D02ACE72FE6FF8DB" box="[291,409,1880,1904]" italics="true" pageId="7" pageNumber="60">C. grandis</emphasis>
</taxonomicName>
<emphasis id="B900C835FFD4FF94D0A9CE72FE28F8DB" bold="true" box="[416,478,1880,1904]" pageId="7" pageNumber="60">
<taxonomicNameLabel id="A233754EFFD4FF94D0A9CE72FE28F8DB" box="[416,478,1880,1904]" pageId="7" pageNumber="60" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
in the uniformly orange ocular peduncles and the reddish brown abdomen marked by numerous transverse to longitudinal white lines (
<bibRefCitation id="EFE569D6FFD4FF94D3B0CE57FB87F83E" author="Miyake" box="[697,1137,1916,1941]" pageId="7" pageNumber="60" refString="Miyake, S. (1982) Japanese Crustacean Decapods and Stomatopods in Color. Vol. 1. Macrura, Anomura and Stomatopoda. Hoikusha, Osaka, 261 pp., 56 pls." type="book" year="1982">
Miyake 1982;
<emphasis id="B900C835FFD4FF94D254CE54FB87F83E" box="[861,1137,1916,1941]" italics="true" pageId="7" pageNumber="60">
Tas rizopagurus
<taxonomicName id="4C746FA4FFD4FF94D511CE56FB87F83E" box="[1048,1137,1916,1941]" class="Malacostraca" family="Diogenidae" genus="Ciliopagurus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="60" phylum="Arthropoda" rank="species" species="krempfi">krempfi</taxonomicName>
</emphasis>
</bibRefCitation>
).
</paragraph>
<paragraph id="8BCB1427FFD4FF9BD1CECE88FAACFECF" blockId="7.[151,1436,151,2013]" lastBlockId="8.[151,1436,151,752]" lastPageId="8" lastPageNumber="61" pageId="7" pageNumber="60">
Among the species characterized by the smooth or microscopically denticulate transverse ridges on the chelae,
<taxonomicName id="4C746FA4FFD4FF94D19ECEEFFDD8F876" authority="Forest, 1952" authorityName="Forest" authorityYear="1952" box="[151,558,1989,2013]" class="Malacostraca" family="Diogenidae" genus="Ciliopagurus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="60" phylum="Arthropoda" rank="species" species="krempfi">
<emphasis id="B900C835FFD4FF94D19ECEEFFE70F876" box="[151,390,1989,2013]" italics="true" pageId="7" pageNumber="60">Ciliopagurus krempfi</emphasis>
(Forest, 1952)
</taxonomicName>
from the Indo-West Pacific also somewhat resembles the present new species in having simple ocular acicles and the short antennal acicles. Nevertheless,
<taxonomicName id="4C746FA4FFDBFF9BD519C9B2FB7FFF1B" box="[1040,1161,152,176]" class="Malacostraca" family="Diogenidae" genus="Ciliopagurus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="61" phylum="Arthropoda" rank="species" species="krempfi">
<emphasis id="B900C835FFDBFF9BD519C9B2FB7FFF1B" box="[1040,1161,152,176]" italics="true" pageId="8" pageNumber="61">C. krempfi</emphasis>
</taxonomicName>
is readily distinguished from
<taxonomicName id="4C746FA4FFDBFF9BD1DDC997FEBCFF7F" box="[212,330,188,212]" class="Malacostraca" family="Diogenidae" genus="Ciliopagurus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="61" phylum="Arthropoda" rank="species" species="grandis" status="sp. nov.">
<emphasis id="B900C835FFDBFF9BD1DDC997FEBCFF7F" box="[212,330,188,212]" italics="true" pageId="8" pageNumber="61">C. grandis</emphasis>
</taxonomicName>
<emphasis id="B900C835FFDBFF9BD059C996FE79FF7F" bold="true" box="[336,399,188,212]" pageId="8" pageNumber="61">
<taxonomicNameLabel id="A233754EFFDBFF9BD059C996FE79FF7F" box="[336,399,188,212]" pageId="8" pageNumber="61" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
by the fewer main transverse ridges on the palms of the chelipeds and the subequal posterior lobes of the telson. In
<taxonomicName id="4C746FA4FFDBFF9BD096C9CAFDE1FF53" box="[415,535,224,248]" class="Malacostraca" family="Diogenidae" genus="Ciliopagurus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="61" phylum="Arthropoda" rank="species" species="krempfi">
<emphasis id="B900C835FFDBFF9BD096C9CAFDE1FF53" box="[415,535,224,248]" italics="true" pageId="8" pageNumber="61">C. krempfi</emphasis>
</taxonomicName>
, there are four main, complete transverse ridges on the palm of the cheliped, whereas, there are seven or eight main transverse ridges, of which three or four are interrupted, in
<taxonomicName id="4C746FA4FFDBFF9BD5E9C82FFAA0FEB7" box="[1248,1366,260,284]" class="Malacostraca" family="Diogenidae" genus="Ciliopagurus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="61" phylum="Arthropoda" rank="species" species="grandis" status="sp. nov.">
<emphasis id="B900C835FFDBFF9BD5E9C82FFAA0FEB7" box="[1248,1366,260,284]" italics="true" pageId="8" pageNumber="61">C. grandis</emphasis>
</taxonomicName>
<emphasis id="B900C835FFDBFF9BD454C82EFA6DFEB7" bold="true" box="[1373,1435,260,284]" pageId="8" pageNumber="61">
<taxonomicNameLabel id="A233754EFFDBFF9BD454C82EFA6DFEB7" box="[1373,1435,260,284]" pageId="8" pageNumber="61" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
The coloration is also very different between the two: in
<taxonomicName id="4C746FA4FFDBFF9BD225C802FC55FEEB" box="[812,931,296,320]" class="Malacostraca" family="Diogenidae" genus="Ciliopagurus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="61" phylum="Arthropoda" rank="species" species="krempfi">
<emphasis id="B900C835FFDBFF9BD225C802FC55FEEB" box="[812,931,296,320]" italics="true" pageId="8" pageNumber="61">C. krempfi</emphasis>
</taxonomicName>
, the carapace is entirely whitish; the ocular peduncles are uniformly yellow; and the ambulatory dactyli are uniformly yellow, without distinct markings.
</paragraph>
<paragraph id="8BCB1427FFDBFF9BD1CEC85AFAEBFD67" blockId="8.[151,1436,151,752]" pageId="8" pageNumber="61">
From Indian waters, two other congeneric species are known,
<taxonomicName id="4C746FA4FFDBFF9BD285C85AFB48FE2C" authority="Herbst, 1804" authorityName="Herbst" authorityYear="1804" box="[908,1214,367,392]" class="Malacostraca" family="Diogenidae" genus="Ciliopagurus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="61" phylum="Arthropoda" rank="species" species="strigatus">
<emphasis id="B900C835FFDBFF9BD285C85AFBE6FE23" box="[908,1040,368,392]" italics="true" pageId="8" pageNumber="61">C. strigatus</emphasis>
(Herbst, 1804)
</taxonomicName>
and
<taxonomicName id="4C746FA4FFDBFF9BD5FEC85AFE15FE07" authority="(Alcock, 1905) (Forest 1995)" authorityName="(Alcock, 1905) (Forest" authorityYear="1995" class="Malacostraca" family="Diogenidae" genus="Ciliopagurus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="61" phylum="Arthropoda" rank="species" species="tenebrarum">
<emphasis id="B900C835FFDBFF9BD5FEC85AFA6AFE23" box="[1271,1436,368,392]" italics="true" pageId="8" pageNumber="61">C. tenebrarum</emphasis>
(
<bibRefCitation id="EFE569D6FFDBFF9BD197C8BEFECEFE07" author="Alcock" box="[158,312,404,428]" pageId="8" pageNumber="61" refString="Alcock, A. (1905) Anomura. Fasc. I. Pagurides. Catalogue of the Indian decapod Crustacea in the collections of the Indian Museum, 2, 1 - 197. Indian Museum, Calcutta." type="journal article" year="1905">Alcock, 1905</bibRefCitation>
) (Forest 1995)
</taxonomicName>
. This new species is the third known from
<collectingCountry id="F36354B7FFDBFF9BD2CEC8BEFBF2FE07" box="[967,1028,404,428]" name="India" pageId="8" pageNumber="61">India</collectingCountry>
. Differentiation between the present new species and these two species is easy:
<taxonomicName id="4C746FA4FFDBFF9BD381C892FCFBFE7B" box="[648,781,440,464]" class="Malacostraca" family="Diogenidae" genus="Ciliopagurus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="61" phylum="Arthropoda" rank="species" species="strigatus">
<emphasis id="B900C835FFDBFF9BD381C892FCFBFE7B" box="[648,781,440,464]" italics="true" pageId="8" pageNumber="61">C. strigatus</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C746FA4FFDBFF9BD2A1C892FBD7FE7B" box="[936,1057,440,464]" class="Malacostraca" family="Diogenidae" genus="Ciliopagurus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="61" phylum="Arthropoda" rank="species" species="grandis" status="sp. nov.">
<emphasis id="B900C835FFDBFF9BD2A1C892FBD7FE7B" box="[936,1057,440,464]" italics="true" pageId="8" pageNumber="61">C. grandis</emphasis>
</taxonomicName>
<emphasis id="B900C835FFDBFF9BD520C892FB9CFE7B" bold="true" box="[1065,1130,440,464]" pageId="8" pageNumber="61">
<taxonomicNameLabel id="A233754EFFDBFF9BD520C892FB9CFE7B" box="[1065,1130,440,464]" pageId="8" pageNumber="61" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
in the multispinose ocular acicle;
<taxonomicName id="4C746FA4FFDBFF9BD1E2C8F7FE64FE5F" box="[235,402,476,500]" class="Malacostraca" family="Diogenidae" genus="Ciliopagurus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="61" phylum="Arthropoda" rank="species" species="tenebrarum">
<emphasis id="B900C835FFDBFF9BD1E2C8F7FE64FE5F" box="[235,402,476,500]" italics="true" pageId="8" pageNumber="61">C. tenebrarum</emphasis>
</taxonomicName>
differs from the new species in the presence of small prominent tubercles on transverse ridges on the chelae and the short antennal peduncle only slightly reaching beyond the midlength of the ocular peduncle. The coloration is also very different among the three species. For example, the ocular peduncle is entirely red in
<taxonomicName id="4C746FA4FFDBFF9BD033CB62FE4AFDCB" box="[314,444,584,608]" class="Malacostraca" family="Diogenidae" genus="Ciliopagurus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="61" phylum="Arthropoda" rank="species" species="strigatus">
<emphasis id="B900C835FFDBFF9BD033CB62FE4AFDCB" box="[314,444,584,608]" italics="true" pageId="8" pageNumber="61">C. strigatus</emphasis>
</taxonomicName>
(cf.
<bibRefCitation id="EFE569D6FFDBFF9BD0E6CB6DFD1CFDF4" author="Poupin" box="[495,746,583,607]" pageId="8" pageNumber="61" refString="Poupin, J. &amp; Malay, M. C. (2009) Identification of a Ciliopagurus strigatus (Herbst, 1804) species-complex, with description of a new species from French Polynesia (Crustacea, Decapoda, Anomura, Diogenidae). Zoosystema, 31, 209 - 232." type="journal article" year="2009">Poupin &amp; Malay 2009</bibRefCitation>
) and
<taxonomicName id="4C746FA4FFDBFF9BD22FCB62FC3CFDCB" box="[806,970,584,608]" class="Malacostraca" family="Diogenidae" genus="Ciliopagurus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="61" phylum="Arthropoda" rank="species" species="tenebrarum">
<emphasis id="B900C835FFDBFF9BD22FCB62FC3CFDCB" box="[806,970,584,608]" italics="true" pageId="8" pageNumber="61">C. tenebrarum</emphasis>
</taxonomicName>
(cf.
<bibRefCitation id="EFE569D6FFDBFF9BD2F5CB6DFB79FDF4" author="Alcock" box="[1020,1167,583,607]" pageId="8" pageNumber="61" refString="Alcock, A. (1905) Anomura. Fasc. I. Pagurides. Catalogue of the Indian decapod Crustacea in the collections of the Indian Museum, 2, 1 - 197. Indian Museum, Calcutta." type="journal article" year="1905">Alcock 1905</bibRefCitation>
), but it is dorsally white in
<taxonomicName id="4C746FA4FFDBFF9BD1BACB47FEDFFD2F" box="[179,297,620,644]" class="Malacostraca" family="Diogenidae" genus="Ciliopagurus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="61" phylum="Arthropoda" rank="species" species="grandis" status="sp. nov.">
<emphasis id="B900C835FFDBFF9BD1BACB47FEDFFD2F" box="[179,297,620,644]" italics="true" pageId="8" pageNumber="61">C. grandis</emphasis>
</taxonomicName>
<emphasis id="B900C835FFDBFF9BD026CB46FE9BFD2F" bold="true" box="[303,365,620,644]" pageId="8" pageNumber="61">
<taxonomicNameLabel id="A233754EFFDBFF9BD026CB46FE9BFD2F" box="[303,365,620,644]" pageId="8" pageNumber="61" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
The chelipeds and ambulatory legs are banded with white or light yellow and red in
<taxonomicName id="4C746FA4FFDBFF9BD413CB47FA6AFD2F" box="[1306,1436,621,644]" class="Malacostraca" family="Diogenidae" genus="Ciliopagurus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="61" phylum="Arthropoda" rank="species" species="strigatus">
<emphasis id="B900C835FFDBFF9BD413CB47FA6AFD2F" box="[1306,1436,621,644]" italics="true" pageId="8" pageNumber="61">C. strigatus</emphasis>
</taxonomicName>
(cf.
<bibRefCitation id="EFE569D6FFDBFF9BD1CDCBA5FE34FD0C" author="Poupin" box="[196,450,655,679]" pageId="8" pageNumber="61" refString="Poupin, J. &amp; Malay, M. C. (2009) Identification of a Ciliopagurus strigatus (Herbst, 1804) species-complex, with description of a new species from French Polynesia (Crustacea, Decapoda, Anomura, Diogenidae). Zoosystema, 31, 209 - 232." type="journal article" year="2009">Poupin &amp; Malay 2009</bibRefCitation>
); banded with light red and dark red, with tips of the distal four segments being white in
<taxonomicName id="4C746FA4FFDBFF9BD19ECB9FFECCFD67" box="[151,314,692,716]" class="Malacostraca" family="Diogenidae" genus="Ciliopagurus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="61" phylum="Arthropoda" rank="species" species="tenebrarum">
<emphasis id="B900C835FFDBFF9BD19ECB9FFECCFD67" box="[151,314,692,716]" italics="true" pageId="8" pageNumber="61">C. tenebrarum</emphasis>
</taxonomicName>
(cf.
<bibRefCitation id="EFE569D6FFDBFF9BD067CB9EFDF4FD67" author="Alcock" box="[366,514,692,716]" pageId="8" pageNumber="61" refString="Alcock, A. (1905) Anomura. Fasc. I. Pagurides. Catalogue of the Indian decapod Crustacea in the collections of the Indian Museum, 2, 1 - 197. Indian Museum, Calcutta." type="journal article" year="1905">Alcock 1905</bibRefCitation>
); and generally orange with slender dark red bands in
<taxonomicName id="4C746FA4FFDBFF9BD568CB9FFB21FD67" box="[1121,1239,692,716]" class="Malacostraca" family="Diogenidae" genus="Ciliopagurus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="61" phylum="Arthropoda" rank="species" species="grandis" status="sp. nov.">
<emphasis id="B900C835FFDBFF9BD568CB9FFB21FD67" box="[1121,1239,692,716]" italics="true" pageId="8" pageNumber="61">C. grandis</emphasis>
</taxonomicName>
<emphasis id="B900C835FFDBFF9BD5D7CB9EFAEBFD67" bold="true" box="[1246,1309,692,716]" pageId="8" pageNumber="61">
<taxonomicNameLabel id="A233754EFFDBFF9BD5D7CB9EFAEBFD67" box="[1246,1309,692,716]" pageId="8" pageNumber="61" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C36E47ACFFDBFF9BD1CECBF2FA88FD44" box="[199,1406,727,752]" pageId="8" pageNumber="61" type="etymology">
<paragraph id="8BCB1427FFDBFF9BD1CECBF2FA88FD44" blockId="8.[151,1436,151,752]" box="[199,1406,727,752]" pageId="8" pageNumber="61">
<emphasis id="B900C835FFDBFF9BD1CECBF2FEB0FD5B" bold="true" box="[199,326,728,752]" pageId="8" pageNumber="61">Etymology</emphasis>
. From the Latin
<taxonomicName id="4C746FA4FFDBFF9BD30ACBF2FDAFFD5B" box="[515,601,728,752]" class="Malacostraca" family="Diogenidae" genus="Ciliopagurus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="61" phylum="Arthropoda" rank="species" species="grandis">
<emphasis id="B900C835FFDBFF9BD30ACBF2FDAFFD5B" box="[515,601,728,752]" italics="true" pageId="8" pageNumber="61">grandis</emphasis>
</taxonomicName>
(= large), in reference to the large body size of the present new species.
</paragraph>
</subSubSection>
</treatment>
</document>