treatments-xml/data/03/B6/3A/03B63A508D680165FF60142404AD4269.xml

<document id="A86AECAB09195A53BB859D8B9E36E386" ID-CLB-Dataset="28158" ID-DOI="10.11646/zootaxa.4563.1.1" ID-GBIF-Dataset="3e3068c1-c158-4d97-9aec-31fc80551edb" ID-ISSN="1175-5326" ID-Zenodo-Dep="2600858" ID-ZooBank="A44D4C11-ADA1-48A8-9F00-B46BDF7B2A49" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553170725873" checkinUser="plazi" docAuthor="Seeman, Owen D." docDate="2019" docId="03B63A508D680165FF60142404AD4269" docLanguage="en" docName="zootaxa.4563.1.1.pdf" docOrigin="Zootaxa 4563 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Megisthanus southcotti Seeman 2019, sp. nov." docType="treatment" docVersion="8" lastPageNumber="27" masterDocId="FF8F42288D7D017FFFF7166307564009" masterDocTitle="The megisthanid mites (Mesostigmata: Megisthanidae) of Australia" masterLastPageNumber="40" masterPageNumber="1" pageNumber="22" updateTime="1698702915705" updateUser="ExternalLinkService">
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<mods:title id="1E8574896C19C8F0544D66760C5BC214">The megisthanid mites (Mesostigmata: Megisthanidae) of Australia</mods:title>
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<mods:namePart id="2CF85C704A9E8E0A3FAB0A6551E54685">Seeman, Owen D.</mods:namePart>
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<mods:date id="17D393017D468C357870E6A1BBCE3898">2019</mods:date>
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<treatment id="03B63A508D680165FF60142404AD4269" ID-DOI="http://doi.org/10.5281/zenodo.5924269" ID-GBIF-Taxon="154474402" ID-Zenodo-Dep="5924269" LSID="urn:lsid:plazi:treatment:03B63A508D680165FF60142404AD4269" httpUri="http://treatment.plazi.org/id/03B63A508D680165FF60142404AD4269" lastPageId="26" lastPageNumber="27" pageId="21" pageNumber="22">
<subSubSection id="C305D8CD8D68016AFF6014240541426B" box="[151,535,583,610]" pageId="21" pageNumber="22" type="nomenclature">
<paragraph id="8BA08B468D68016AFF6014240541426B" blockId="21.[151,535,583,643]" box="[151,535,583,610]" pageId="21" pageNumber="22">
<heading id="D0E83C2A8D68016AFF6014240541426B" bold="true" box="[151,535,583,610]" fontSize="11" level="1" pageId="21" pageNumber="22" reason="1">
<emphasis id="B96B57548D68016AFF6014240541426B" bold="true" box="[151,535,583,610]" pageId="21" pageNumber="22">
<taxonomicName id="4C1FF0C58D68016AFF60142406E44268" authority="Seeman, 2019" authorityName="Seeman" authorityYear="2019" box="[151,434,583,609]" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="southcotti" status="sp. nov.">
<emphasis id="B96B57548D68016AFF60142406E44268" bold="true" box="[151,434,583,609]" italics="true" pageId="21" pageNumber="22">Megisthanus southcotti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A258EA2F8D68016AFE4E142B0541426B" box="[441,535,584,610]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C305D8CD8D68016AFF601409066E428A" box="[151,312,618,643]" pageId="21" pageNumber="22" type="description">
<paragraph id="8BA08B468D68016AFF601409066E428A" blockId="21.[151,535,583,643]" box="[151,312,618,643]" pageId="21" pageNumber="22">
<figureCitation id="132497C38D68016AFF601409066E428A" box="[151,312,618,643]" captionStart-0="FIGURE 12" captionStart-1="FIGURE 13" captionStart-2="FIGURE 14" captionStart-3="FIGURE 15" captionStartId-0="22.[151,250,1822,1844]" captionStartId-1="23.[151,250,1069,1091]" captionStartId-2="24.[151,250,1705,1727]" captionStartId-3="25.[151,250,1807,1829]" captionTargetBox-0="[157,1428,207,1794]" captionTargetBox-1="[162,1421,196,1043]" captionTargetBox-2="[180,1394,196,1617]" captionTargetBox-3="[202,1400,1080,1747]" captionTargetId-0="figure@22.[151,1436,193,1801]" captionTargetId-1="figure@23.[151,1436,193,1047]" captionTargetId-2="figure@24.[151,1436,193,1684]" captionTargetId-3="figure@25.[151,1436,1057,1786]" captionTargetPageId-0="22" captionTargetPageId-1="23" captionTargetPageId-2="24" captionTargetPageId-3="25" captionText-0="FIGURE 12. Megisthanus southcotti sp. nov. photomicrographs. (A) female, dorsal shield; (B) female, venter; (C) major male, venter; (D) major male, sternoventral shield. Scale bars: 200 µm (A–C), 100 µm (D)." captionText-1="FIGURE 13. Megisthanus southcotti sp. nov. female, photomicrographs. (A) ventral shield; (B) posterior dorsal shield; (C) ventrianal shield. Scale bars = 100 µm." captionText-2="FIGURE 14. Megisthanus southcotti sp. nov. (A) female, sternogenital region; (B) internal genitalia; (C) female, femur IV; (D) major male, sternogenital region. Abbreviations: g1 = anterior pair of genital setae; lg = latigynal remnant; mg = mesogynal shield; va = vaginal arms; vh = vaginal heads." captionText-3="FIGURE 15. Megisthanus southcotti sp. nov. (A) female subcapitulum and palp trochanter–genu (shown detached); (B) major male subcapitulum and palp trochanter–genu. Arrows indicate spurs." httpUri-0="https://zenodo.org/record/2600882/files/figure.png" httpUri-1="https://zenodo.org/record/2600884/files/figure.png" httpUri-2="https://zenodo.org/record/2600886/files/figure.png" httpUri-3="https://zenodo.org/record/2600888/files/figure.png" pageId="21" pageNumber="22">Figures 12–15</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C305D8CD8D68016AFF6014D106404526" pageId="21" pageNumber="22" type="materials_examined">
<paragraph id="8BA08B468D68016AFF6014D105E844CA" blockId="21.[151,1437,690,2011]" pageId="21" pageNumber="22">
<emphasis id="B96B57548D68016AFF6014D105FE42C2" bold="true" box="[151,680,690,715]" pageId="21" pageNumber="22">
Material examined. 
<materialsCitation id="3B77811B8D68016AFE6114D105FE42C2" ID-GBIF-Occurrence="2013950921" box="[406,680,690,715]" collectingDate="1996-01-31" collectionCode="QMS" collectorName="O. Seeman" country="Australia" location="Mossman Gorge" pageId="21" pageNumber="22" specimenCode="QMS 109486" specimenCount="1" stateProvince="Queensland" typeStatus="holotype">
<collectingCountry id="F308CBD68D68016AFE6114D1055142C2" box="[406,519,690,715]" name="Australia" pageId="21" pageNumber="22">Australia</collectingCountry>
, 
<collectingRegion id="49DB45A48D68016AFDEC14D105FE42C2" box="[539,680,690,715]" country="Australia" name="Queensland" pageId="21" pageNumber="22">Queensland</collectingRegion>
</materialsCitation>
</emphasis>
, ex 
<taxonomicName id="4C1FF0C58D68016AFD1114D0039642C2" authority="Kuwert, 1891" authorityName="Kuwert" authorityYear="1891" box="[742,1216,691,715]" class="Insecta" family="Passallidae" genus="Aulacocyclus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="fracticornis">
<emphasis id="B96B57548D68016AFD1114D0035B42C2" box="[742,1037,691,715]" italics="true" pageId="21" pageNumber="22">Aulacocyclus fracticornis</emphasis>
<bibRefCitation id="EF8EF6B78D68016AFBEC14D0039642C2" author="Kuwert, A. F." box="[1051,1216,691,715]" pageId="21" pageNumber="22" pagination="161 - 192" refId="ref22234" refString="Kuwert, A. F. (1891) Systematische Ubersicht der Passaliden-Arten und Gattungen. Deutsche Entomologische Zeitschrift, 1891, 161 - 192." type="journal article" year="1891">Kuwert, 1891</bibRefCitation>
</taxonomicName>
: 
<typeStatus id="54A435E48D68016AFB2514D1026C42C2" box="[1234,1338,690,715]" pageId="21" pageNumber="22" type="holotype">Holotype</typeStatus>
female, 
<location id="8EC0DD9D8D68016AFF6014B4060542E6" LSID="urn:lsid:plazi:treatment:03B63A508D680165FF60142404AD4269:8EC0DD9D8D68016AFF6014B4060542E6" box="[151,339,727,751]" country="Australia" name="Mossman Gorge" pageId="21" pageNumber="22" stateProvince="Queensland">Mossman Gorge</location>
, 
<date id="FFA1AD868D68016AFEA814B5069D42E6" box="[351,459,726,751]" pageId="21" pageNumber="22" value="1996-01-31">
<collectingDate id="EFE5546E8D68016AFEA814B5069D42E6" box="[351,459,726,751]" pageId="21" pageNumber="22" value="1996-01-31">31.i.1996</collectingDate>
</date>
, 
<collectorName id="26EAEE908D68016AFE2F14B4050042E6" box="[472,598,727,751]" pageId="21" pageNumber="22">O. Seeman</collectorName>
, ex beetle #66 (log 1, passalid 1, pr A) (
<specimenCode id="DBB9233D8D68016AFBEC14B403EC42E6" box="[1051,1210,726,751]" collectionCode="QMS" pageId="21" pageNumber="22">QMS 109486</specimenCode>
). 
<materialsCitation id="3B77811B8D68016AFB3A14B40335431A" ID-GBIF-Occurrence="2013950907" collectingDate="1996-01-31" collectionCode="QM" collectorName="O. Seeman" country="Australia" location="In" pageId="21" pageNumber="22" specimenCode="QMS 109488" specimenCount="15" specimenCount-female="15" stateProvince="Queensland" typeStatus="paratype">
<location id="8EC0DD9D8D68016AFB3A14B403B242E7" LSID="urn:lsid:plazi:treatment:03B63A508D680165FF60142404AD4269:8EC0DD9D8D68016AFB3A14B403B242E7" box="[1229,1252,727,750]" country="Australia" name="In" pageId="21" pageNumber="22" stateProvince="Queensland">In</location>
<collectionCode id="ED0E13838D68016AFB1B14B4024B42E6" box="[1260,1309,727,751]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="21" pageNumber="22" type="Museum">QM</collectionCode>
. 
<typeStatus id="54A435E48D68016AFADF14B402CF42E6" box="[1320,1433,727,751]" pageId="21" pageNumber="22" type="paratype">Paratypes</typeStatus>
: 
<specimenCount id="9D1940CF8D68016AFF6014980642431A" box="[151,276,762,787]" pageId="21" pageNumber="22" type="female">14 females</specimenCount>
, 7 major males, as follows. 
<specimenCount id="9D1940CF8D68016AFDBD149805F8431A" box="[586,686,762,787]" pageId="21" pageNumber="22" type="female">1 female</specimenCount>
, same data as holotype (
<specimenCode id="DBB9233D8D68016AFC491498030A431A" box="[958,1116,763,787]" collectionCode="QMS" pageId="21" pageNumber="22">QMS 109488</specimenCode>
)
</materialsCitation>
; 
<materialsCitation id="3B77811B8D68016AFB98149805F9433E" ID-GBIF-Occurrence="2013950915" collectingDate="1996-01-31" collectionCode="QMS" collectorName="O. Seeman" country="Australia" location="In" pageId="21" pageNumber="22" specimenCode="QMS 109487" specimenCount="1" specimenCount-female="1" stateProvince="Queensland" typeStatus="paratype">
<specimenCount id="9D1940CF8D68016AFB9814980384431A" box="[1135,1234,762,787]" pageId="21" pageNumber="22" type="female">1 female</specimenCount>
, same data except ex beetle #71 (log 2 passalid 1) (
<specimenCode id="DBB9233D8D68016AFDFE157C05F1433E" box="[521,679,799,823]" collectionCode="QMS" pageId="21" pageNumber="22">QMS 109487</specimenCode>
)
</materialsCitation>
; 
<materialsCitation id="3B77811B8D68016AFD4C157C06DE4352" ID-GBIF-Occurrence="2013950900" collectingDate="1996-01-31" collectionCode="QMS" collectorName="O. Seeman" country="Australia" location="In" pageId="21" pageNumber="22" specimenCode="QMS109489" specimenCount="1" stateProvince="Queensland" typeStatus="paratype">
1 major male, same data except ex beetle #220 (log 1, passalid 2, pr A) (
<specimenCode id="DBB9233D8D68016AFF1E152006D64352" box="[233,384,835,859]" collectionCode="QMS" pageId="21" pageNumber="22">QMS109489</specimenCode>
)
</materialsCitation>
; 
<materialsCitation id="3B77811B8D68016AFE60152006684376" ID-GBIF-Occurrence="2013950902" collectingDate="1996-01-28" collectionCode="QMS" collectorName="O. Seeman" country="Australia" location="Atherton" municipality="Wongabel" pageId="21" pageNumber="22" specimenCode="QMS109490" specimenCount="1" specimenCount-female="1" stateProvince="Queensland" typeStatus="paratype">
<specimenCount id="9D1940CF8D68016AFE60152006A94352" box="[407,511,834,859]" pageId="21" pageNumber="22" type="female">1 female</specimenCount>
, 
<collectingMunicipality id="6BC4113C8D68016AFDF9152105D44352" box="[526,642,834,859]" pageId="21" pageNumber="22">Wongabel</collectingMunicipality>
, 
<location id="8EC0DD9D8D68016AFD66152105AE4352" LSID="urn:lsid:plazi:treatment:03B63A508D680165FF60142404AD4269:8EC0DD9D8D68016AFD66152105AE4352" box="[657,760,834,859]" country="Australia" municipality="Wongabel" name="Atherton" pageId="21" pageNumber="22" stateProvince="Queensland">Atherton</location>
, 
<date id="FFA1AD868D68016AFCF0152104254352" box="[775,883,834,859]" pageId="21" pageNumber="22" value="1996-01-28">
<collectingDate id="EFE5546E8D68016AFCF0152104254352" box="[775,883,834,859]" pageId="21" pageNumber="22" value="1996-01-28">28.i.1996</collectingDate>
</date>
, 
<collectorName id="26EAEE908D68016AFC75152003554352" box="[898,1027,835,859]" pageId="21" pageNumber="22">O. Seeman</collectorName>
, ex beetle #119 (log 1, passalid 3) (
<specimenCode id="DBB9233D8D68016AFF68150406604376" box="[159,310,871,895]" collectionCode="QMS" pageId="21" pageNumber="22">QMS109490</specimenCode>
)
</materialsCitation>
; 
<materialsCitation id="3B77811B8D68016AFEBE1504039C4376" ID-GBIF-Occurrence="2013950924" box="[329,1226,870,895]" collectingDate="1996-01-28" collectionCode="QMS" collectorName="O. Seeman" country="Australia" location="Atherton" municipality="Wongabel" pageId="21" pageNumber="22" specimenCode="QMS109491" specimenCount="1" stateProvince="Queensland" typeStatus="paratype">
1 major male, same data except ex beetle #86 (log 1, passalid 1) (
<specimenCode id="DBB9233D8D68016AFBDC150403944376" box="[1067,1218,871,895]" collectionCode="QMS" pageId="21" pageNumber="22">QMS109491</specimenCode>
)
</materialsCitation>
; 
<materialsCitation id="3B77811B8D68016AFB22150403A143AA" ID-GBIF-Occurrence="2013950906" collectingDate="1996-01-27" collectionCode="QMS" collectorName="O. Seeman" country="Australia" location="Atherton" municipality="Gadgarra" pageId="21" pageNumber="22" specimenCode="QMS 109492-3" specimenCount="1" specimenCount-female="1" stateProvince="Queensland" typeStatus="paratype">
<specimenCount id="9D1940CF8D68016AFB221504026C4376" box="[1237,1338,870,895]" pageId="21" pageNumber="22" type="female">1 female</specimenCount>
, 1 major male, 
<collectingMunicipality id="6BC4113C8D68016AFF2C15E9061043AA" box="[219,326,906,931]" pageId="21" pageNumber="22">Gadgarra</collectingMunicipality>
, 
<location id="8EC0DD9D8D68016AFEA615E906EE43AA" LSID="urn:lsid:plazi:treatment:03B63A508D680165FF60142404AD4269:8EC0DD9D8D68016AFEA615E906EE43AA" box="[337,440,906,931]" country="Australia" municipality="Gadgarra" name="Atherton" pageId="21" pageNumber="22" stateProvince="Queensland">Atherton</location>
, 
<date id="FFA1AD868D68016AFE3515E9057843AA" box="[450,558,906,931]" pageId="21" pageNumber="22" value="1996-01-27">
<collectingDate id="EFE5546E8D68016AFE3515E9057843AA" box="[450,558,906,931]" pageId="21" pageNumber="22" value="1996-01-27">27.i.1996</collectingDate>
</date>
, 
<collectorName id="26EAEE908D68016AFDCD15E805EE43AA" box="[570,696,907,931]" pageId="21" pageNumber="22">O. Seeman</collectorName>
, ex beetle #90 (log 2, passalid 7) (
<specimenCode id="DBB9233D8D68016AFBC215E803B943AA" box="[1077,1263,907,931]" collectionCode="QMS" pageId="21" pageNumber="22">QMS 109492–3</specimenCode>
)
</materialsCitation>
; 
<materialsCitation id="3B77811B8D68016AFAF515E804ED43CE" ID-GBIF-Occurrence="2013950916" collectingDate="1996-01-27" collectionCode="QMS" collectorName="O. Seeman" country="Australia" location="Atherton" municipality="Gadgarra" pageId="21" pageNumber="22" specimenCode="QMS 109494" specimenCount="1" stateProvince="Queensland" typeStatus="paratype">
1 major male, same data except ex beetle #92 (log 2, passalid 8) (
<specimenCode id="DBB9233D8D68016AFCF815CC04E543CE" box="[783,947,943,967]" collectionCode="QMS" pageId="21" pageNumber="22">QMS 109494</specimenCode>
)
</materialsCitation>
; 
<materialsCitation id="3B77811B8D68016AFC3B15CC04EA43E2" ID-GBIF-Occurrence="2013950920" collectingDate="1996-01-30" collectionCode="QMS" collectorName="O. Seeman" country="Australia" location="Cape Tribulation" municipality="Oliver Ck" pageId="21" pageNumber="22" specimenCode="QMS 109495" specimenCount="1" specimenCount-female="1" stateProvince="Queensland" typeStatus="paratype">
<specimenCount id="9D1940CF8D68016AFC3B15CC036143CE" box="[972,1079,942,967]" pageId="21" pageNumber="22" type="female">1 female</specimenCount>
, 
<collectingMunicipality id="6BC4113C8D68016AFBBF15CD039643CE" box="[1096,1216,942,967]" pageId="21" pageNumber="22">Oliver Ck</collectingMunicipality>
, 
<location id="8EC0DD9D8D68016AFB2615CC02CE43CE" LSID="urn:lsid:plazi:treatment:03B63A508D680165FF60142404AD4269:8EC0DD9D8D68016AFB2615CC02CE43CE" box="[1233,1432,942,967]" country="Australia" municipality="Oliver Ck" name="Cape Tribulation" pageId="21" pageNumber="22" stateProvince="Queensland">Cape Tribulation</location>
, 
<date id="FFA1AD868D68016AFF6015B1065543E2" box="[151,259,978,1003]" pageId="21" pageNumber="22" value="1996-01-30">
<collectingDate id="EFE5546E8D68016AFF6015B1065543E2" box="[151,259,978,1003]" pageId="21" pageNumber="22" value="1996-01-30">30.i.1996</collectingDate>
</date>
, 
<collectorName id="26EAEE908D68016AFEE715B006D943E2" box="[272,399,979,1003]" pageId="21" pageNumber="22">O. Seeman</collectorName>
, ex beetle #17 (log 3, passalid 2) (
<specimenCode id="DBB9233D8D68016AFCE215B004E343E2" box="[789,949,978,1003]" collectionCode="QMS" pageId="21" pageNumber="22">QMS 109495</specimenCode>
)
</materialsCitation>
; 
<materialsCitation id="3B77811B8D68016AFC3E15B0051F4406" ID-GBIF-Occurrence="2013950908" collectingDate="1996-01-30" collectionCode="QMS" collectorName="O. Seeman" country="Australia" location="Cape Tribulation" municipality="Oliver Ck" pageId="21" pageNumber="22" specimenCode="QMS 109496" specimenCount="1" stateProvince="Queensland" typeStatus="paratype">
1 major male, same data except ex beetle #14 (log 5, passalid 1) (
<specimenCode id="DBB9233D8D68016AFE55159405174406" box="[418,577,1014,1039]" collectionCode="QMS" pageId="21" pageNumber="22">QMS 109496</specimenCode>
)
</materialsCitation>
; 
<materialsCitation id="3B77811B8D68016AFDA3159407AE443A" ID-GBIF-Occurrence="2013950919" collectingDate="1996-01-30" collectionCode="ANIC" collectorName="O. Seeman" country="Australia" location="Cape Tribulation" municipality="Oliver Ck" pageId="21" pageNumber="22" specimenCode="ANIC 51- 006453" specimenCount="1" stateProvince="Queensland" typeStatus="paratype">
1 major male, same data except ex beetle #8 (log 2, passalid 1) (
<specimenCode id="DBB9233D8D68016AFADE159407A6443A" collectionCode="ANIC" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="21" pageNumber="22">ANIC 51- 006453</specimenCode>
)
</materialsCitation>
; 
<materialsCitation id="3B77811B8D68016AFEF112780622445E" ID-GBIF-Occurrence="2013950923" collectingDate="1996-01-30" collectionCode="ANIC" collectorName="O. Seeman" country="Australia" location="Cape Tribulation" municipality="Earthwatch Site" pageId="21" pageNumber="22" specimenCode="ANIC 51-006452" specimenCount="1" specimenCount-female="1" stateProvince="Queensland" typeStatus="paratype">
<specimenCount id="9D1940CF8D68016AFEF112780638443A" box="[262,366,1050,1075]" pageId="21" pageNumber="22" type="female">1 female</specimenCount>
, 
<collectingMunicipality id="6BC4113C8D68016AFE8A12790563443A" box="[381,565,1050,1075]" pageId="21" pageNumber="22">Earthwatch Site</collectingMunicipality>
, 
<location id="8EC0DD9D8D68016AFDB31278045E443A" LSID="urn:lsid:plazi:treatment:03B63A508D680165FF60142404AD4269:8EC0DD9D8D68016AFDB31278045E443A" box="[580,776,1050,1075]" country="Australia" municipality="Earthwatch Site" name="Cape Tribulation" pageId="21" pageNumber="22" stateProvince="Queensland">Cape Tribulation</location>
, 
<collectingDate id="EFE5546E8D68016AFCE1127904D5443A" box="[790,899,1050,1075]" pageId="21" pageNumber="22" value="1996-01-30">
<date id="FFA1AD868D68016AFCE1127904D5443A" box="[790,899,1050,1075]" pageId="21" pageNumber="22" value="1996-01-30">30.i.1996</date>
</collectingDate>
, 
<collectorName id="26EAEE908D68016AFC6512780345443A" box="[914,1043,1051,1075]" pageId="21" pageNumber="22">O. Seeman</collectorName>
, ex beetle #51 (log 2, passalid 3) (
<specimenCode id="DBB9233D8D68016AFF69125C063A445E" box="[158,364,1086,1111]" collectionCode="ANIC" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="21" pageNumber="22">ANIC 51-006452</specimenCode>
)
</materialsCitation>
; 
<materialsCitation id="3B77811B8D68016AFE76125C05EC44CA" ID-GBIF-Occurrence="2013950922" collectingDate="1943-10-24" collectingDateMax="1950-01-10" collectingDateMin="1943-10-24" collectionCode="SAM, QM, ANIC" collectorName="J. C. Brooks &amp; D. L. Pollier &amp; Specimens" country="Australia" county="Longlands Gap Rd" location="Atherton Tablelands" municipality="Herberton" pageId="21" pageNumber="22" specimenCount="7" specimenCount-female="6" specimenCount-male="1" stateProvince="Queensland" typeStatus="paratype">
<specimenCount id="9D1940CF8D68016AFE76125C06A4445E" box="[385,498,1086,1111]" pageId="21" pageNumber="22" type="female">3 females</specimenCount>
, 
<specimenCount id="9D1940CF8D68016AFE08125C0519445E" box="[511,591,1086,1111]" pageId="21" pageNumber="22" type="male">1 male</specimenCount>
, 
<locationDeviation id="9A1292648D68016AFDAA125C0437445E" box="[605,865,1086,1111]" location="east Wondecla" pageId="21" pageNumber="22">
<quantity id="4CE726A38D68016AFDAA125C05E7445E" box="[605,689,1086,1111]" metricMagnitude="4" metricUnit="m" metricValue="1.2874752" pageId="21" pageNumber="22" unit="mi" value="8.0">8 miles</quantity>
east Wondecla
</locationDeviation>
, 
<date id="FFA1AD868D68016AFC99125C04B6445E" box="[878,992,1087,1111]" pageId="21" pageNumber="22" value="1943-10-24">
<collectingDate id="EFE5546E8D68016AFC99125C04B6445E" box="[878,992,1087,1111]" pageId="21" pageNumber="22" value="1943-10-24">24.x.1943</collectingDate>
</date>
, R.V. coll., ex passalid (
<accessionNumber id="944C16A58D68016AFB0E125C0218445E" box="[1273,1358,1087,1111]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J17309" pageId="21" pageNumber="22" type="EnaNcbi">J17309</accessionNumber>
–12). 
<specimenCount id="9D1940CF8D68016AFA79125C07A74472" pageId="21" pageNumber="22" type="female">2 females</specimenCount>
, 
<collectingCounty id="62C1F3CA8D68016AFEF6120106B44472" box="[257,482,1122,1147]" pageId="21" pageNumber="22">Longlands Gap Rd</collectingCounty>
, near 
<collectingMunicipality id="6BC4113C8D68016AFDD8120105F24472" box="[559,676,1122,1147]" pageId="21" pageNumber="22">Herberton</collectingMunicipality>
, 
<date id="FFA1AD868D68016AFD43120104764472" box="[692,800,1122,1147]" pageId="21" pageNumber="22" value="1950-01-10">
<collectingDate id="EFE5546E8D68016AFD43120104764472" box="[692,800,1122,1147]" pageId="21" pageNumber="22" value="1950-01-10">10.i.1950</collectingDate>
</date>
, 
<collectorName id="26EAEE908D68016AFCC6120004EB4472" box="[817,957,1122,1147]" pageId="21" pageNumber="22">J.C. Brooks</collectorName>
, ex small carabid (
<accessionNumber id="944C16A58D68016AFB53120003AC4472" box="[1188,1274,1123,1147]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J17307" pageId="21" pageNumber="22" type="EnaNcbi">J17307</accessionNumber>
–8). 
<specimenCount id="9D1940CF8D68016AFAD8120002CE4472" box="[1327,1432,1122,1147]" pageId="21" pageNumber="22" type="female">1 female</specimenCount>
, 
<location id="8EC0DD9D8D68016AFF6012E506284496" LSID="urn:lsid:plazi:treatment:03B63A508D680165FF60142404AD4269:8EC0DD9D8D68016AFF6012E506284496" box="[151,382,1158,1183]" country="Australia" county="Longlands Gap Rd" municipality="Herberton" name="Atherton Tablelands" pageId="21" pageNumber="22" stateProvince="Queensland">Atherton Tablelands</location>
, 
<date id="FFA1AD868D68016AFE7D12E505504496" box="[394,518,1158,1183]" pageId="21" pageNumber="22" value="1945-03-28">
<collectingDate id="EFE5546E8D68016AFE7D12E505504496" box="[394,518,1158,1183]" pageId="21" pageNumber="22" value="1945-03-28">28.iii.1945</collectingDate>
</date>
, 
<collectorName id="26EAEE908D68016AFDE512E405CF4496" box="[530,665,1158,1183]" pageId="21" pageNumber="22">D.L. Pollier</collectorName>
(
<accessionNumber id="944C16A58D68016AFD5E12E405A94496" box="[681,767,1159,1183]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J17328" pageId="21" pageNumber="22" type="EnaNcbi">J17328</accessionNumber>
). 
<collectorName id="26EAEE908D68016AFCE412E504DA4496" box="[787,908,1158,1183]" pageId="21" pageNumber="22">Specimens</collectorName>
with J-numbers in 
<collectionCode id="ED0E13838D68016AFB9F12E403FE4496" box="[1128,1192,1159,1183]" country="South Africa" httpUri="http://grbio.org/cool/mhew-9czz" name="South African Museum" pageId="21" pageNumber="22">SAM</collectionCode>
, others in 
<collectionCode id="ED0E13838D68016AFAE912E4021D4496" box="[1310,1355,1159,1183]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="21" pageNumber="22" type="Museum">QM</collectionCode>
except one 
<typeStatus id="54A435E48D68016AFF3112C8067144CA" box="[198,295,1195,1219]" pageId="21" pageNumber="22" type="paratype">paratype</typeStatus>
female and 
<typeStatus id="54A435E48D68016AFE4712C8054644CA" box="[432,528,1195,1219]" pageId="21" pageNumber="22" type="paratype">paratype</typeStatus>
male in 
<collectionCode id="ED0E13838D68016AFD8512C805EC44CA" box="[626,698,1195,1219]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="21" pageNumber="22">ANIC</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BA08B468D68016AFF3012AD06404526" blockId="21.[151,1437,690,2011]" pageId="21" pageNumber="22">
<emphasis id="B96B57548D68016AFF3012AD05C644EE" bold="true" box="[199,656,1230,1255]" pageId="21" pageNumber="22">
Other material examined, 
<materialsCitation id="3B77811B8D68016AFDF412AD05C644EE" ID-GBIF-Occurrence="2013950909" box="[515,656,1230,1255]" collectingDate="1996-01-30" collectorName="O. Seeman" location="Mt Lewis" municipality="Mt Lewis Rd" pageId="21" pageNumber="22" specimenCount="2" specimenCount-female="2" stateProvince="Queensland">
<collectingRegion id="49DB45A48D68016AFDF412AD05C644EE" box="[515,656,1230,1255]" country="Australia" name="Queensland" pageId="21" pageNumber="22">Queensland</collectingRegion>
</materialsCitation>
</emphasis>
, ex 
<taxonomicName id="4C1FF0C58D68016AFD4812AD03F644EE" authority="Mjoberg, 1917: 1" authorityName="Mjoberg" authorityPageNumber="1" authorityYear="1917" box="[703,1184,1230,1255]" class="Insecta" family="Passallidae" genus="Austropassalus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="hultgreni">
<emphasis id="B96B57548D68016AFD4812AD048044EF" box="[703,982,1230,1254]" italics="true" pageId="21" pageNumber="22">Austropassalus hultgreni</emphasis>
<bibRefCitation id="EF8EF6B78D68016AFC2A12AD03D144EE" author="Mjoberg, E. G." box="[989,1159,1230,1255]" pageId="21" pageNumber="22" pagination="1 - 19" refId="ref22407" refString="Mjoberg, E. G. (1917) Results of Dr. E. Mjoberg's Swedish scientific expeditions to Australia 1910 - 1913. 14. Cetonidae, Rutelidae, Passalidae, Chrysomelidae: subfam. Sagrinae, Cassidinae, Hispinae. Arkiv for Zoologi, 11, 1 - 19." type="journal article" year="1917">Mjöberg, 1917</bibRefCitation>
: 
<specimenCount id="9D1940CF8D68016AFB6512AC03F644EE" box="[1170,1184,1231,1255]" pageId="21" pageNumber="22" type="female">1</specimenCount>
</taxonomicName>
female, 
<collectingMunicipality id="6BC4113C8D68016AFAF612AC02C144EE" box="[1281,1431,1230,1255]" pageId="21" pageNumber="22">Mt Lewis Rd</collectingMunicipality>
, 
<location id="8EC0DD9D8D68016AFF601290065E4502" LSID="urn:lsid:plazi:treatment:03B63A508D680165FF60142404AD4269:8EC0DD9D8D68016AFF601290065E4502" box="[151,264,1266,1291]" municipality="Mt Lewis Rd" name="Mt Lewis" pageId="21" pageNumber="22" stateProvince="Queensland">Mt Lewis</location>
, 
<date id="FFA1AD868D68016AFEE1129106D44502" box="[278,386,1266,1291]" pageId="21" pageNumber="22" value="1996-01-30">
<collectingDate id="EFE5546E8D68016AFEE1129106D44502" box="[278,386,1266,1291]" pageId="21" pageNumber="22" value="1996-01-30">30.i.1996</collectingDate>
</date>
, 
<collectorName id="26EAEE908D68016AFE67129005464502" box="[400,528,1267,1291]" pageId="21" pageNumber="22">O. Seeman</collectorName>
, (beetle #57, log 1, passalid 2). 
<specimenCount id="9D1940CF8D68016AFC70129004B84502" box="[903,1006,1266,1291]" pageId="21" pageNumber="22" type="female">1 female</specimenCount>
, same data, except beetle #68 (log 1, passalid 5).
</paragraph>
</subSubSection>
<subSubSection id="C305D8CD8D68016AFF30135906AA467A" pageId="21" pageNumber="22" type="diagnosis">
<paragraph id="8BA08B468D68016AFF30135906AA467A" blockId="21.[151,1437,690,2011]" pageId="21" pageNumber="22">
<emphasis id="B96B57548D68016AFF3013590617455A" bold="true" box="[199,321,1338,1363]" pageId="21" pageNumber="22">Diagnosis.</emphasis>
<emphasis id="B96B57548D68016AFEBF1358055C455A" box="[328,522,1339,1363]" italics="true" pageId="21" pageNumber="22">Female and male</emphasis>
. Sternal setae st4 absent. Seta pd2 of femur IV on rudimentary projection. Dorsal shield finely punctate. Ventrianal shield widest at posterior level of anus. 
<emphasis id="B96B57548D68016AFC3E133D034B457F" box="[969,1053,1374,1398]" italics="true" pageId="21" pageNumber="22">Female</emphasis>
. Dorsal shield ovate, length 
<quantity id="4CE726A38D68016AFAAA133D07AB4593" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.05" metricValueMin="1.95" pageId="21" pageNumber="22" unit="mm" value="2.0" valueMax="2.05" valueMin="1.95">1.95– 2.05 mm</quantity>
, maximum width 
<quantity id="4CE726A38D68016AFE3E13E105384593" box="[457,622,1410,1435]" metricMagnitude="-4" metricUnit="m" metricValue="9.5" metricValueMax="10.5" metricValueMin="8.5" pageId="21" pageNumber="22" unit="mm" value="0.95" valueMax="1.05" valueMin="0.85">0.85–1.05 mm</quantity>
, not including secondary sclerotisation. Most dorsal shield setae 
<quantity id="4CE726A38D68016AFABF13E007FB45B6" metricMagnitude="0" metricUnit="m" metricValue="2.667" metricValueMax="3.048" metricValueMin="2.286" pageId="21" pageNumber="22" unit="in" value="105.0" valueMax="120.0" valueMin="90.0">90–120 in</quantity>
length, with fine but conspicuous barbs; some marginal setae thicker and longer; podonotum and opisthonotum densely setose. Sternogynal shields weakly fused posteriorly, each bearing 4–7 setae; internal genitalia with rudimentary latigynal element. Ventrianal shield a rounded trapezoid. 
<emphasis id="B96B57548D68016AFC35138C031F460F" box="[962,1097,1518,1542]" italics="true" pageId="21" pageNumber="22">Major male</emphasis>
. Broader than female, dorsal shield length 
<quantity id="4CE726A38D68016AFEC6107106814623" box="[305,471,1554,1579]" metricMagnitude="-3" metricUnit="m" metricValue="2.075" metricValueMax="2.2" metricValueMin="1.95" pageId="21" pageNumber="22" unit="mm" value="2.075" valueMax="2.2" valueMin="1.95">1.95–2.20 mm</quantity>
, width 
<quantity id="4CE726A38D68016AFDDA107105824623" box="[557,724,1554,1579]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="1.6" metricValueMin="1.4" pageId="21" pageNumber="22" unit="mm" value="1.5" valueMax="1.6" valueMin="1.4">1.40–1.60 mm</quantity>
; setae on soft cuticle laterad dorsal shield short, length 15–30; 130–160 setae around genital opening; corniculi lacking medial inner tooth; setae h1-h2 smooth, h3 and palp coxal setae smooth or with 1–2 barbs.
</paragraph>
</subSubSection>
<subSubSection id="C305D8CD8D680166FF30101D043A43E4" lastPageId="25" lastPageNumber="26" pageId="21" pageNumber="22" type="description">
<paragraph id="8BA08B468D68016AFF30101D047547D2" blockId="21.[151,1437,690,2011]" pageId="21" pageNumber="22">
<emphasis id="B96B57548D68016AFF30101D06EE469E" bold="true" box="[199,440,1662,1687]" pageId="21" pageNumber="22">
Description. 
<emphasis id="B96B57548D68016AFE96101D06EE469E" bold="true" box="[353,440,1662,1687]" italics="true" pageId="21" pageNumber="22">Female</emphasis>
</emphasis>
. 
<emphasis id="B96B57548D68016AFE30101C0577469F" box="[455,545,1663,1686]" italics="true" pageId="21" pageNumber="22">Dorsum</emphasis>
(
<figureCitation id="132497C38D68016AFDC5101D05C9469E" box="[562,671,1662,1687]" captionStart="FIGURE 12" captionStartId="22.[151,250,1822,1844]" captionTargetBox="[157,1428,207,1794]" captionTargetId="figure@22.[151,1436,193,1801]" captionTargetPageId="22" captionText="FIGURE 12. Megisthanus southcotti sp. nov. photomicrographs. (A) female, dorsal shield; (B) female, venter; (C) major male, venter; (D) major male, sternoventral shield. Scale bars: 200 µm (A–C), 100 µm (D)." httpUri="https://zenodo.org/record/2600882/files/figure.png" pageId="21" pageNumber="22">Figs 12A</figureCitation>
, 
<figureCitation id="132497C38D68016AFD5B101C0588469E" box="[684,734,1663,1687]" captionStart="FIGURE 13" captionStartId="23.[151,250,1069,1091]" captionTargetBox="[162,1421,196,1043]" captionTargetId="figure@23.[151,1436,193,1047]" captionTargetPageId="23" captionText="FIGURE 13. Megisthanus southcotti sp. nov. female, photomicrographs. (A) ventral shield; (B) posterior dorsal shield; (C) ventrianal shield. Scale bars = 100 µm." httpUri="https://zenodo.org/record/2600884/files/figure.png" pageId="21" pageNumber="22">13B</figureCitation>
). Idiosomal length 
<quantity id="4CE726A38D68016AFC4C101C0376469F" box="[955,1056,1663,1687]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="21" pageNumber="22" unit="mm" value="2.3">2.30 mm</quantity>
(
<quantity id="4CE726A38D68016AFBC7101C038C469E" box="[1072,1242,1663,1687]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" metricValueMax="2.3" metricValueMin="2.2" pageId="21" pageNumber="22" unit="mm" value="2.25" valueMax="2.3" valueMin="2.2">2.20–2.30 mm</quantity>
), width 
<quantity id="4CE726A38D68016AFAC0101D02CB469F" box="[1335,1437,1662,1687]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="21" pageNumber="22" unit="mm" value="1.5">1.50 mm</quantity>
(
<quantity id="4CE726A38D68016AFF6810C1061C46B2" box="[159,330,1698,1723]" metricMagnitude="-3" metricUnit="m" metricValue="1.375" metricValueMax="1.5" metricValueMin="1.25" pageId="21" pageNumber="22" unit="mm" value="1.375" valueMax="1.5" valueMin="1.25">1.25–1.50 mm</quantity>
), body length (including 
<taxonomicName id="4C1FF0C58D68016AFD9910C105AE46B2" authorityName="Krefft" authorityYear="1871" box="[622,760,1698,1723]" class="Chromadorea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="21" pageNumber="37" phylum="Nematoda" rank="genus">gnathosoma</taxonomicName>
, to tip of gnathotectum) 
<quantity id="4CE726A38D68016AFBEE10C003D646B3" box="[1049,1152,1699,1723]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="21" pageNumber="22" unit="mm" value="2.8">2.80 mm</quantity>
(
<quantity id="4CE726A38D68016AFB6710C1026B46B2" box="[1168,1341,1698,1723]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" metricValueMax="2.8" metricValueMin="2.6" pageId="21" pageNumber="22" unit="mm" value="2.7" valueMax="2.8" valueMin="2.6">2.60–2.80 mm</quantity>
). Dorsal shield ovate, length 
<quantity id="4CE726A38D68016AFE8D10A5068846D7" box="[378,478,1734,1759]" metricMagnitude="-3" metricUnit="m" metricValue="1.95" pageId="21" pageNumber="22" unit="mm" value="1.95">1.95 mm</quantity>
(
<quantity id="4CE726A38D68016AFE1A10A505C046D6" box="[493,662,1734,1759]" metricMagnitude="-3" metricUnit="m" metricValue="1.9249999999999998" metricValueMax="2.05" metricValueMin="1.8" pageId="21" pageNumber="22" unit="mm" value="1.9249999999999998" valueMax="2.05" valueMin="1.8">1.80–2.05 mm</quantity>
), width 
<quantity id="4CE726A38D68016AFD0710A5040346D7" box="[752,853,1734,1759]" metricMagnitude="-4" metricUnit="m" metricValue="9.5" pageId="21" pageNumber="22" unit="mm" value="0.95">0.95 mm</quantity>
(
<quantity id="4CE726A38D68016AFC9410A5035A46D6" box="[867,1036,1734,1759]" metricMagnitude="-4" metricUnit="m" metricValue="9.5" metricValueMax="10.5" metricValueMin="8.5" pageId="21" pageNumber="22" unit="mm" value="0.95" valueMax="1.05" valueMin="0.85">0.85–1.05 mm</quantity>
); secondary sclerotisation absent or present (including 
<typeStatus id="54A435E48D68016AFE9B10890687470A" box="[364,465,1770,1795]" pageId="21" pageNumber="22" type="holotype">holotype</typeStatus>
, 
<figureCitation id="132497C38D68016AFE2910890511470A" box="[478,583,1770,1795]" captionStart="FIGURE 12" captionStartId="22.[151,250,1822,1844]" captionTargetBox="[157,1428,207,1794]" captionTargetId="figure@22.[151,1436,193,1801]" captionTargetPageId="22" captionText="FIGURE 12. Megisthanus southcotti sp. nov. photomicrographs. (A) female, dorsal shield; (B) female, venter; (C) major male, venter; (D) major male, sternoventral shield. Scale bars: 200 µm (A–C), 100 µm (D)." httpUri="https://zenodo.org/record/2600882/files/figure.png" pageId="21" pageNumber="22">Fig. 12A</figureCitation>
), mostly posterolaterally and posteriorly; maximum length of dorsal shield including secondary sclerotisation 
<quantity id="4CE726A38D68016AFDE9116C05D3472F" box="[542,645,1807,1831]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="21" pageNumber="22" unit="mm" value="2.2">2.20 mm</quantity>
, width 
<quantity id="4CE726A38D68016AFD2F116C0416472F" box="[728,832,1807,1831]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="21" pageNumber="22" unit="mm" value="1.1">1.10 mm</quantity>
. Dorsal shield hypertrichous, especially mid-dorsally, setation sparser at margins and off dorsal shield. Most dorsal shield setae with several conspicuous but fine barbs, length 80–100; anterior and lateral setae similar but longer, and several marginal setae thicker and much longer (up to 360); posterior dorsal shield setae slightly longer, length 90–120 (
<figureCitation id="132497C38D68016AFC6911190353479A" box="[926,1029,1914,1939]" captionStart="FIGURE 13" captionStartId="23.[151,250,1069,1091]" captionTargetBox="[162,1421,196,1043]" captionTargetId="figure@23.[151,1436,193,1047]" captionTargetPageId="23" captionText="FIGURE 13. Megisthanus southcotti sp. nov. female, photomicrographs. (A) ventral shield; (B) posterior dorsal shield; (C) ventrianal shield. Scale bars = 100 µm." httpUri="https://zenodo.org/record/2600884/files/figure.png" pageId="21" pageNumber="22">Fig. 13B</figureCitation>
). Dorsal shield cuticle with obvious but fine punctation. Setae in soft cuticle with small barbs, on small platelets; mediolateral setae close to dorsal shield long (170–320), other setae much shorter (50–115).
</paragraph>
<caption id="DF60DBCE8D6B0169FF60117D04A5475A" httpUri="https://zenodo.org/record/2600882/files/figure.png" pageId="22" pageNumber="23" startId="22.[151,250,1822,1844]" targetBox="[157,1428,207,1794]" targetPageId="22">
<paragraph id="8BA08B468D6B0169FF60117D04A5475A" blockId="22.[151,1436,1822,1875]" pageId="22" pageNumber="23">
<emphasis id="B96B57548D6B0169FF60117D0649473A" bold="true" box="[151,287,1822,1844]" pageId="22" pageNumber="23">FIGURE 12.</emphasis>
<taxonomicName id="4C1FF0C58D6B0169FED3117D0546473D" authority="Seeman, 2019" authorityName="Seeman" authorityYear="2019" box="[292,528,1822,1844]" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="southcotti" status="sp. nov.">
<emphasis id="B96B57548D6B0169FED3117D0546473D" box="[292,528,1822,1844]" italics="true" pageId="22" pageNumber="23">Megisthanus southcotti</emphasis>
</taxonomicName>
<emphasis id="B96B57548D6B0169FDE2117C0530473D" bold="true" box="[533,614,1823,1844]" pageId="22" pageNumber="23">
<taxonomicNameLabel id="A258EA2F8D6B0169FDE2117C0530473D" box="[533,614,1823,1844]" pageId="22" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
photomicrographs. (A) female, dorsal shield; (B) female, venter; (C) major male, venter; (D) major male, sternoventral shield. Scale bars: 200 µm (A–C), 100 µm (D).
</paragraph>
</caption>
<caption id="DF60DBCE8D6A0168FF60124E0574446B" httpUri="https://zenodo.org/record/2600884/files/figure.png" pageId="23" pageNumber="24" startId="23.[151,250,1069,1091]" targetBox="[162,1421,196,1043]" targetPageId="23">
<paragraph id="8BA08B468D6A0168FF60124E0574446B" blockId="23.[151,1436,1069,1122]" pageId="23" pageNumber="24">
<emphasis id="B96B57548D6A0168FF60124E0677444B" bold="true" box="[151,289,1069,1091]" pageId="23" pageNumber="24">FIGURE 13.</emphasis>
<taxonomicName id="4C1FF0C58D6A0168FEDD124E054E444A" authority="Seeman, 2019" authorityName="Seeman" authorityYear="2019" box="[298,536,1069,1091]" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="southcotti" status="sp. nov.">
<emphasis id="B96B57548D6A0168FEDD124E054E444A" box="[298,536,1069,1091]" italics="true" pageId="23" pageNumber="24">Megisthanus southcotti</emphasis>
</taxonomicName>
<emphasis id="B96B57548D6A0168FDD7124E0524444B" bold="true" box="[544,626,1069,1090]" pageId="23" pageNumber="24">
<taxonomicNameLabel id="A258EA2F8D6A0168FDD7124E0524444B" box="[544,626,1069,1090]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
female, photomicrographs. (A) ventral shield; (B) posterior dorsal shield; (C) ventrianal shield. Scale bars = 100 µm.
</paragraph>
</caption>
<paragraph id="8BA08B468D6A0168FF3012E902CB44E3" blockId="23.[151,1437,1161,2014]" pageId="23" pageNumber="24">
<emphasis id="B96B57548D6A0168FF3012E9065A44A8" box="[199,268,1162,1185]" italics="true" pageId="23" pageNumber="24">Venter</emphasis>
(
<figureCitation id="132497C38D6A0168FED712EA06DB44A8" box="[288,397,1161,1186]" captionStart="FIGURE 12" captionStartId="22.[151,250,1822,1844]" captionTargetBox="[157,1428,207,1794]" captionTargetId="figure@22.[151,1436,193,1801]" captionTargetPageId="22" captionText="FIGURE 12. Megisthanus southcotti sp. nov. photomicrographs. (A) female, dorsal shield; (B) female, venter; (C) major male, venter; (D) major male, sternoventral shield. Scale bars: 200 µm (A–C), 100 µm (D)." httpUri="https://zenodo.org/record/2600882/files/figure.png" pageId="23" pageNumber="24">Figs 12B</figureCitation>
, 
<figureCitation id="132497C38D6A0168FE6C12EA055B44A8" box="[411,525,1161,1185]" captionStart="FIGURE 13" captionStartId="23.[151,250,1069,1091]" captionTargetBox="[162,1421,196,1043]" captionTargetId="figure@23.[151,1436,193,1047]" captionTargetPageId="23" captionText="FIGURE 13. Megisthanus southcotti sp. nov. female, photomicrographs. (A) ventral shield; (B) posterior dorsal shield; (C) ventrianal shield. Scale bars = 100 µm." httpUri="https://zenodo.org/record/2600884/files/figure.png" pageId="23" pageNumber="24">13A, 13C</figureCitation>
, 
<figureCitation id="132497C38D6A0168FDEC12EA056144A8" box="[539,567,1161,1185]" captionStart="FIGURE 14" captionStartId="24.[151,250,1705,1727]" captionTargetBox="[180,1394,196,1617]" captionTargetId="figure@24.[151,1436,193,1684]" captionTargetPageId="24" captionText="FIGURE 14. Megisthanus southcotti sp. nov. (A) female, sternogenital region; (B) internal genitalia; (C) female, femur IV; (D) major male, sternogenital region. Abbreviations: g1 = anterior pair of genital setae; lg = latigynal remnant; mg = mesogynal shield; va = vaginal arms; vh = vaginal heads." httpUri="https://zenodo.org/record/2600886/files/figure.png" pageId="23" pageNumber="24">14</figureCitation>
A–B). Setae st1 sparsely barbed, length 140 (135–155). Sternal shield (
<figureCitation id="132497C38D6A0168FA9812EA079E44CC" captionStart="FIGURE 14" captionStartId="24.[151,250,1705,1727]" captionTargetBox="[180,1394,196,1617]" captionTargetId="figure@24.[151,1436,193,1684]" captionTargetPageId="24" captionText="FIGURE 14. Megisthanus southcotti sp. nov. (A) female, sternogenital region; (B) internal genitalia; (C) female, femur IV; (D) major male, sternogenital region. Abbreviations: g1 = anterior pair of genital setae; lg = latigynal remnant; mg = mesogynal shield; va = vaginal arms; vh = vaginal heads." httpUri="https://zenodo.org/record/2600886/files/figure.png" pageId="23" pageNumber="24">Fig. 14A</figureCitation>
) reticulate, with 2 pairs of setae, 5–6 pairs of pores opening ventrally, without pores opening laterally; setae st2–3 sparsely barbed, st2 length 110 (100–110), st3 length 90 (75–95); st4 absent; shield medial length 60 (55–70).
</paragraph>
<paragraph id="8BA08B468D6A0168FF301296044445CB" blockId="23.[151,1437,1161,2014]" pageId="23" pageNumber="24">
Sternogynal shields (
<figureCitation id="132497C38D6A0168FE4F129605734504" box="[440,549,1269,1294]" captionStart="FIGURE 14" captionStartId="24.[151,250,1705,1727]" captionTargetBox="[180,1394,196,1617]" captionTargetId="figure@24.[151,1436,193,1684]" captionTargetPageId="24" captionText="FIGURE 14. Megisthanus southcotti sp. nov. (A) female, sternogenital region; (B) internal genitalia; (C) female, femur IV; (D) major male, sternogenital region. Abbreviations: g1 = anterior pair of genital setae; lg = latigynal remnant; mg = mesogynal shield; va = vaginal arms; vh = vaginal heads." httpUri="https://zenodo.org/record/2600886/files/figure.png" pageId="23" pageNumber="24">Figs 14A</figureCitation>
) weakly fused posteriorly, genital opening length 255 (255–265), width 285 (280–290), each sternogynal shield length 220 (220–240), weakly lineate-reticulate, each bearing 4 (4–7, asymmetry present) setae and one lyrifissure; anterior seta thicker, longer 90 (85–110) than posterior setae (50–65). Internal genitalia comprises three obvious elements (
<emphasis id="B96B57548D6A0168FCF7130104744570" box="[768,802,1378,1401]" italics="true" pageId="23" pageNumber="24">mg</emphasis>
, 
<emphasis id="B96B57548D6A0168FCC3130104184570" box="[820,846,1378,1401]" italics="true" pageId="23" pageNumber="24">va</emphasis>
, 
<emphasis id="B96B57548D6A0168FC961302042A4570" box="[865,892,1377,1401]" italics="true" pageId="23" pageNumber="24">vh</emphasis>
) and rudimentary latigynal (
<emphasis id="B96B57548D6A0168FB3A130203B54570" box="[1229,1251,1377,1401]" italics="true" pageId="23" pageNumber="24">lg</emphasis>
) elements (
<figureCitation id="132497C38D6A0168FA991302079F4594" captionStart="FIGURE 14" captionStartId="24.[151,250,1705,1727]" captionTargetBox="[180,1394,196,1617]" captionTargetId="figure@24.[151,1436,193,1684]" captionTargetPageId="24" captionText="FIGURE 14. Megisthanus southcotti sp. nov. (A) female, sternogenital region; (B) internal genitalia; (C) female, femur IV; (D) major male, sternogenital region. Abbreviations: g1 = anterior pair of genital setae; lg = latigynal remnant; mg = mesogynal shield; va = vaginal arms; vh = vaginal heads." httpUri="https://zenodo.org/record/2600886/files/figure.png" pageId="23" pageNumber="24">Fig. 14B</figureCitation>
); rudimentary latigynal elements fused with thickened margin of sternogynal shield, with small porose area near attachment of vaginal arms with sternogynal shield.
</paragraph>
<paragraph id="8BA08B468D6A0168FF3013AE03FC465B" blockId="23.[151,1437,1161,2014]" pageId="23" pageNumber="24">
Ventral shield (
<figureCitation id="132497C38D6A0168FE8313AE06B645EC" box="[372,480,1485,1510]" captionStart="FIGURE 12" captionStartId="22.[151,250,1822,1844]" captionTargetBox="[157,1428,207,1794]" captionTargetId="figure@22.[151,1436,193,1801]" captionTargetPageId="22" captionText="FIGURE 12. Megisthanus southcotti sp. nov. photomicrographs. (A) female, dorsal shield; (B) female, venter; (C) major male, venter; (D) major male, sternoventral shield. Scale bars: 200 µm (A–C), 100 µm (D)." httpUri="https://zenodo.org/record/2600882/files/figure.png" pageId="23" pageNumber="24">Figs 12B</figureCitation>
, 
<figureCitation id="132497C38D6A0168FE1913AE054945EC" box="[494,543,1485,1509]" captionStart="FIGURE 13" captionStartId="23.[151,250,1069,1091]" captionTargetBox="[162,1421,196,1043]" captionTargetId="figure@23.[151,1436,193,1047]" captionTargetPageId="23" captionText="FIGURE 13. Megisthanus southcotti sp. nov. female, photomicrographs. (A) ventral shield; (B) posterior dorsal shield; (C) ventrianal shield. Scale bars = 100 µm." httpUri="https://zenodo.org/record/2600884/files/figure.png" pageId="23" pageNumber="24">13A</figureCitation>
) extends 750 (725–820) posteriorly from posterior margin of genital opening; maximum width behind coxa IV 340 (340–370); shield with 9–12 pairs of setae around shield margin and 5–11 setae within shield and close to sternogynal shield; setae often expressed asymmetrically; setae smooth, length 40– 90; shield reticulate posteriorly and posterior to genital opening, remainder lineate-reticulate.
</paragraph>
<paragraph id="8BA08B468D6A0168FF30103E03904693" blockId="23.[151,1437,1161,2014]" pageId="23" pageNumber="24">
Ventrianal shield a rounded trapezoid (
<figureCitation id="132497C38D6A0168FD73103E05BD467C" box="[644,747,1629,1654]" captionStart="FIGURE 13" captionStartId="23.[151,250,1069,1091]" captionTargetBox="[162,1421,196,1043]" captionTargetId="figure@23.[151,1436,193,1047]" captionTargetPageId="23" captionText="FIGURE 13. Megisthanus southcotti sp. nov. female, photomicrographs. (A) ventral shield; (B) posterior dorsal shield; (C) ventrianal shield. Scale bars = 100 µm." httpUri="https://zenodo.org/record/2600884/files/figure.png" pageId="23" pageNumber="24">Fig. 13C</figureCitation>
), anterior width 350 (350–420), maximum width 530 (530– 570), length 320 (315–370); shield reticulate; 12–16 loosely paired smooth setae, length 55–90.
</paragraph>
<paragraph id="8BA08B468D6A0168FF3010C605EC470F" blockId="23.[151,1437,1161,2014]" pageId="23" pageNumber="24">
Peritrematal shield hypertrichous, post–peritrematal region extends 450 (450–520) behind coxa IV into metapodal region (
<figureCitation id="132497C38D6A0168FE9A10AA068346E8" box="[365,469,1737,1762]" captionStart="FIGURE 12" captionStartId="22.[151,250,1822,1844]" captionTargetBox="[157,1428,207,1794]" captionTargetId="figure@22.[151,1436,193,1801]" captionTargetPageId="22" captionText="FIGURE 12. Megisthanus southcotti sp. nov. photomicrographs. (A) female, dorsal shield; (B) female, venter; (C) major male, venter; (D) major male, sternoventral shield. Scale bars: 200 µm (A–C), 100 µm (D)." httpUri="https://zenodo.org/record/2600882/files/figure.png" pageId="23" pageNumber="24">Fig. 12B</figureCitation>
); setae laterad legs barbed, length 25–50; anterior margin of peritrematal shield with three pairs of long barbed setae, length 150–260.
</paragraph>
<paragraph id="8BA08B468D6A0168FF30117203AA47D7" blockId="23.[151,1437,1161,2014]" pageId="23" pageNumber="24">
<taxonomicName id="4C1FF0C58D6A0168FF30117206004720" authorityName="Krefft" authorityYear="1871" box="[199,342,1809,1833]" class="Chromadorea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="23" pageNumber="24" phylum="Nematoda" rank="genus">
<emphasis id="B96B57548D6A0168FF30117206004720" box="[199,342,1809,1833]" italics="true" pageId="23" pageNumber="24">Gnathosoma</emphasis>
</taxonomicName>
(
<figureCitation id="132497C38D6A0168FE90117206874723" box="[359,465,1809,1834]" captionStart="FIGURE 15" captionStartId="25.[151,250,1807,1829]" captionTargetBox="[202,1400,1080,1747]" captionTargetId="figure@25.[151,1436,1057,1786]" captionTargetPageId="25" captionText="FIGURE 15. Megisthanus southcotti sp. nov. (A) female subcapitulum and palp trochanter–genu (shown detached); (B) major male subcapitulum and palp trochanter–genu. Arrows indicate spurs." httpUri="https://zenodo.org/record/2600888/files/figure.png" pageId="23" pageNumber="24">Fig. 15A</figureCitation>
). Gnathotectum reticulate, medial length 680 (660–690). Subcapitulum with setae h1 240 (230–250), h2 155 (150–160), h3 245 (235–250), pc 135 (135–150), all setae barbed; medial subcapitulum with some fine striae, hypostome smooth. Corniculi length 230 (230–255) with medial tooth. Palp trochanter with ventro-distal spur, length 15 (12–15), palp femur with tiny distal spur, length 5 (5–8). Chelicerae with entire fixed digit length 610 (610–660), movable digit length 355 (350–370). Morphology as per genus description. Fixed digit with 12 teeth; cheliceral seta length 35–45 (not visible in 
<typeStatus id="54A435E48D6A0168FCEC11A604D447D7" box="[795,898,1989,2014]" pageId="23" pageNumber="24" type="holotype">holotype</typeStatus>
). Movable digit with 10–11 teeth.
</paragraph>
<caption id="DF60DBCE8D650167FF6010CA053F46F5" httpUri="https://zenodo.org/record/2600886/files/figure.png" pageId="24" pageNumber="25" startId="24.[151,250,1705,1727]" targetBox="[180,1394,196,1617]" targetPageId="24">
<paragraph id="8BA08B468D650167FF6010CA053F46F5" blockId="24.[151,1436,1705,1788]" pageId="24" pageNumber="25">
<emphasis id="B96B57548D650167FF6010CA064946B7" bold="true" box="[151,287,1705,1727]" pageId="24" pageNumber="25">FIGURE 14.</emphasis>
<taxonomicName id="4C1FF0C58D650167FED310CA055946B6" authority="Seeman, 2019" authorityName="Seeman" authorityYear="2019" box="[292,527,1705,1727]" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="southcotti" status="sp. nov.">
<emphasis id="B96B57548D650167FED310CA055946B6" box="[292,527,1705,1727]" italics="true" pageId="24" pageNumber="25">Megisthanus southcotti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A258EA2F8D650167FDE210C9053246B6" box="[533,612,1706,1727]" pageId="24" pageNumber="25" rank="species">
<emphasis id="B96B57548D650167FDE210C9050846B6" bold="true" box="[533,606,1706,1727]" pageId="24" pageNumber="25">sp. nov</emphasis>
.
</taxonomicNameLabel>
(A) female, sternogenital region; (B) internal genitalia; (C) female, femur IV; (D) 
<taxonomicName id="4C1FF0C58D650167FF6010AB078446D7" box="[151,210,1736,1758]" pageId="24" pageNumber="25" rank="species" species="major">major</taxonomicName>
male, sternogenital region. Abbreviations: g1 = anterior pair of genital setae; lg = latigynal remnant; mg = mesogynal shield; va = vaginal arms; vh = vaginal heads.
</paragraph>
</caption>
<paragraph id="8BA08B468D650167FF301146023647C4" blockId="24.[151,1436,1828,2033]" pageId="24" pageNumber="25">
<emphasis id="B96B57548D650167FF30114607AA4735" box="[199,252,1829,1852]" italics="true" pageId="24" pageNumber="25">Legs</emphasis>
(
<figureCitation id="132497C38D650167FEE61147062B4734" box="[273,381,1828,1853]" captionStart="FIGURE 14" captionStartId="24.[151,250,1705,1727]" captionTargetBox="[180,1394,196,1617]" captionTargetId="figure@24.[151,1436,193,1684]" captionTargetPageId="24" captionText="FIGURE 14. Megisthanus southcotti sp. nov. (A) female, sternogenital region; (B) internal genitalia; (C) female, femur IV; (D) major male, sternogenital region. Abbreviations: g1 = anterior pair of genital setae; lg = latigynal remnant; mg = mesogynal shield; va = vaginal arms; vh = vaginal heads." httpUri="https://zenodo.org/record/2600886/files/figure.png" pageId="24" pageNumber="25">Fig. 14C</figureCitation>
). Setation as per genus description. Selected setal measurements: genu II setae ad1, pd1 subequal, 85–100; seta pd1 on genu III very long, 220–300, much longer than ad1, 100–110, tarsus III ad3 moderately long, length 210–250, femur IV pd1 moderately long, 200–250, genu IV ad1 long, 375–410, pd1 long, length 435–510, tibia IV pd1 long, length 410–460 and tarsus IV ad4 long, length 320–340, pd3 long, length 315– 325. Femur IV with small projections bearing setae pd2, ad2, and slightly larger projection bearing setae ad3.
</paragraph>
<paragraph id="8BA08B468D650166FF3011B406674149" blockId="24.[151,1436,1828,2033]" lastBlockId="25.[151,1437,151,1005]" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25">
<emphasis id="B96B57548D650167FF3011B4065547F9" bold="true" box="[199,259,2007,2032]" italics="true" pageId="24" pageNumber="25">Male</emphasis>
(major). 
<emphasis id="B96B57548D650167FE8C11BA068347F9" box="[379,469,2009,2032]" italics="true" pageId="24" pageNumber="25">Dorsum</emphasis>
. Idiosomal length 
<quantity id="4CE726A38D650167FD4911BB043D47F9" box="[702,875,2008,2032]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" metricValueMax="2.4" metricValueMin="2.2" pageId="24" pageNumber="25" unit="mm" value="2.3" valueMax="2.4" valueMin="2.2">2.20–2.40 mm</quantity>
, width 2.00– 
<quantity id="4CE726A38D650167FBFB11BB032C47F9" box="[1036,1146,2008,2032]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="24" pageNumber="25" unit="mm" value="2.3">2.30 mm</quantity>
, body length (including 
<taxonomicName id="4C1FF0C58D640166FF6016F4067740B9" authorityName="Krefft" authorityYear="1871" box="[151,289,151,176]" class="Chromadorea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="25" pageNumber="37" phylum="Nematoda" rank="genus">gnathosoma</taxonomicName>
, to tip of gnathotectum) 
<quantity id="4CE726A38D640166FDC816F405FF40A6" box="[575,681,151,176]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="25" pageNumber="26" unit="mm" value="2.6">2.60 mm</quantity>
. Dorsal shield ovate, length 
<quantity id="4CE726A38D640166FC0516F403CF40A6" box="[1010,1177,151,176]" metricMagnitude="-3" metricUnit="m" metricValue="2.075" metricValueMax="2.2" metricValueMin="1.95" pageId="25" pageNumber="26" unit="mm" value="2.075" valueMax="2.2" valueMin="1.95">1.95–2.20 mm</quantity>
, width 
<quantity id="4CE726A38D640166FB0716F402CE40A6" box="[1264,1432,151,176]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="1.6" metricValueMin="1.4" pageId="25" pageNumber="26" unit="mm" value="1.5" valueMax="1.6" valueMin="1.4">1.40–1.60 mm</quantity>
; secondary sclerotisation absent or present, expanding laterally and posteriorly, length of dorsal shield including secondary sclerotisation 
<quantity id="4CE726A38D640166FE5816BC050340FE" box="[431,597,223,248]" metricMagnitude="-3" metricUnit="m" metricValue="2.175" metricValueMax="2.3" metricValueMin="2.05" pageId="25" pageNumber="26" unit="mm" value="2.175" valueMax="2.3" valueMin="2.05">2.05–2.30 mm</quantity>
, width 
<quantity id="4CE726A38D640166FD5D16BC040440FE" box="[682,850,223,248]" metricMagnitude="-3" metricUnit="m" metricValue="1.625" metricValueMax="1.75" metricValueMin="1.5" pageId="25" pageNumber="26" unit="mm" value="1.625" valueMax="1.75" valueMin="1.5">1.50–1.75 mm</quantity>
. Dorsum hypertrichous. Setae on dorsal shield tiny, smooth, length &lt;5 excepting few anterior and posterior setae up to 
<quantity id="4CE726A38D640166FC70176704964114" box="[903,960,260,285]" metricMagnitude="-1" metricUnit="m" metricValue="7.62" pageId="25" pageNumber="26" unit="in" value="30.0">30 in</quantity>
length. Soft membrane with setae smooth, length 15–30.
</paragraph>
<paragraph id="8BA08B468D640166FF30172E06674269" blockId="25.[151,1437,151,1005]" pageId="25" pageNumber="26">
<emphasis id="B96B57548D640166FF30172E065A416D" box="[199,268,333,356]" italics="true" pageId="25" pageNumber="26">Venter</emphasis>
(
<figureCitation id="132497C38D640166FED4172F0621416D" box="[291,375,332,357]" captionStart="FIGURE 12" captionStartId="22.[151,250,1822,1844]" captionTargetBox="[157,1428,207,1794]" captionTargetId="figure@22.[151,1436,193,1801]" captionTargetPageId="22" captionText="FIGURE 12. Megisthanus southcotti sp. nov. photomicrographs. (A) female, dorsal shield; (B) female, venter; (C) major male, venter; (D) major male, sternoventral shield. Scale bars: 200 µm (A–C), 100 µm (D)." httpUri="https://zenodo.org/record/2600882/files/figure.png" pageId="25" pageNumber="26">Fig. 12</figureCitation>
C–D, 14D). Setae st1 barbed, length 125–140. Sternoventral shield reticulate, length from posterior margin of genital opening 860–940. Sternal setae st2 and st3 sparsely barbed, length 115–130 and 115– 135, respectively; sternoventral shield (
<figureCitation id="132497C38D640166FDA117F7059741A5" box="[598,705,404,429]" captionStart="FIGURE 14" captionStartId="24.[151,250,1705,1727]" captionTargetBox="[180,1394,196,1617]" captionTargetId="figure@24.[151,1436,193,1684]" captionTargetPageId="24" captionText="FIGURE 14. Megisthanus southcotti sp. nov. (A) female, sternogenital region; (B) internal genitalia; (C) female, femur IV; (D) major male, sternogenital region. Abbreviations: g1 = anterior pair of genital setae; lg = latigynal remnant; mg = mesogynal shield; va = vaginal arms; vh = vaginal heads." httpUri="https://zenodo.org/record/2600886/files/figure.png" pageId="25" pageNumber="26">Figs 14D</figureCitation>
) highly hypertrichous posterior to setae st3, with 130–150 setae around genital opening; posterior sternoventral shield with setae along margin; these setae smooth or sparsely barbed, shortest setae 70–80 (posterolateral setae) and longest setae 115–135 (between, and just behind, coxa IV); posterior sternoventral shield (
<figureCitation id="132497C38D640166FE00179C05374211" box="[503,609,511,536]" captionStart="FIGURE 12" captionStartId="22.[151,250,1822,1844]" captionTargetBox="[157,1428,207,1794]" captionTargetId="figure@22.[151,1436,193,1801]" captionTargetPageId="22" captionText="FIGURE 12. Megisthanus southcotti sp. nov. photomicrographs. (A) female, dorsal shield; (B) female, venter; (C) major male, venter; (D) major male, sternoventral shield. Scale bars: 200 µm (A–C), 100 µm (D)." httpUri="https://zenodo.org/record/2600882/files/figure.png" pageId="25" pageNumber="26">Fig. 12D</figureCitation>
) with two discs of porose cuticle (diameter 130–145), each bearing an anterior and posterior pair of unmodified pair of smooth setae 20–35, and a highly modified pair of setae centrally, length 40–50.
</paragraph>
<paragraph id="8BA08B468D640166FF30140F030042A1" blockId="25.[151,1437,151,1005]" pageId="25" pageNumber="26">
Genital opening (
<figureCitation id="132497C38D640166FE7D140F06A6428D" box="[394,496,620,645]" captionStart="FIGURE 14" captionStartId="24.[151,250,1705,1727]" captionTargetBox="[180,1394,196,1617]" captionTargetId="figure@24.[151,1436,193,1684]" captionTargetPageId="24" captionText="FIGURE 14. Megisthanus southcotti sp. nov. (A) female, sternogenital region; (B) internal genitalia; (C) female, femur IV; (D) major male, sternogenital region. Abbreviations: g1 = anterior pair of genital setae; lg = latigynal remnant; mg = mesogynal shield; va = vaginal arms; vh = vaginal heads." httpUri="https://zenodo.org/record/2600886/files/figure.png" pageId="25" pageNumber="26">Fig. 14D</figureCitation>
) length 115–120, width 90–110, surrounded by a ring of heavily sclerotised cuticle; this cuticle with 6–10 loosely paired pores laterad and posterolaterad genital opening.
</paragraph>
<paragraph id="8BA08B468D640166FF3014D706F0431C" blockId="25.[151,1437,151,1005]" pageId="25" pageNumber="26">
Ventrianal shield (
<figureCitation id="132497C38D640166FE6D14D7055442C5" box="[410,514,692,717]" captionStart="FIGURE 12" captionStartId="22.[151,250,1822,1844]" captionTargetBox="[157,1428,207,1794]" captionTargetId="figure@22.[151,1436,193,1801]" captionTargetPageId="22" captionText="FIGURE 12. Megisthanus southcotti sp. nov. photomicrographs. (A) female, dorsal shield; (B) female, venter; (C) major male, venter; (D) major male, sternoventral shield. Scale bars: 200 µm (A–C), 100 µm (D)." httpUri="https://zenodo.org/record/2600882/files/figure.png" pageId="25" pageNumber="26">Fig. 12C</figureCitation>
) a curved trapezoid, length 300–365, anterior width 420–450, maximum width 585–660, widest just behind anus; shield reticulate, with 14–16 loosely paired setae. Peritreme and peritrematal shield similar to female.
</paragraph>
<paragraph id="8BA08B468D640166FF30154305A343C1" blockId="25.[151,1437,151,1005]" pageId="25" pageNumber="26">
<taxonomicName id="4C1FF0C58D640166FF30154306004331" authorityName="Krefft" authorityYear="1871" box="[199,342,800,824]" class="Chromadorea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="25" pageNumber="26" phylum="Nematoda" rank="genus">
<emphasis id="B96B57548D640166FF30154306004331" box="[199,342,800,824]" italics="true" pageId="25" pageNumber="26">Gnathosoma</emphasis>
</taxonomicName>
(
<figureCitation id="132497C38D640166FE9C157C068D4331" box="[363,475,799,824]" captionStart="FIGURE 15" captionStartId="25.[151,250,1807,1829]" captionTargetBox="[202,1400,1080,1747]" captionTargetId="figure@25.[151,1436,1057,1786]" captionTargetPageId="25" captionText="FIGURE 15. Megisthanus southcotti sp. nov. (A) female subcapitulum and palp trochanter–genu (shown detached); (B) major male subcapitulum and palp trochanter–genu. Arrows indicate spurs." httpUri="https://zenodo.org/record/2600888/files/figure.png" pageId="25" pageNumber="26">Figs 15B</figureCitation>
). Gnathotectum similar to female, medial length 780–810; subcapitulum similar, except with submedial projections (inner length 20-40). Setae h1 smooth, 200–215; setae h2 smooth, 125–155, setae h3 smooth or with 1–2 barbs, 210–215, palp coxal seta smooth or with few small barbs, 150–160. Corniculi straight, lacking tooth-like processes, length 365–385. Palp with distoventral cusps on trochanter (length 25–30) and femur (length 40–45). Chelicerae same as female.
</paragraph>
<paragraph id="8BA08B468D640166FF3015B6043A43E4" blockId="25.[151,1437,151,1005]" box="[199,876,980,1005]" pageId="25" pageNumber="26">
<emphasis id="B96B57548D640166FF3015B607AA43E5" box="[199,252,981,1004]" italics="true" pageId="25" pageNumber="26">Legs</emphasis>
. Similar to female except distal spurs and spines larger.
</paragraph>
</subSubSection>
<caption id="DF60DBCE8D640166FF60116C0406474D" httpUri="https://zenodo.org/record/2600888/files/figure.png" pageId="25" pageNumber="26" startId="25.[151,250,1807,1829]" targetBox="[202,1400,1080,1747]" targetPageId="25">
<paragraph id="8BA08B468D640166FF60116C0406474D" blockId="25.[151,1436,1807,1860]" pageId="25" pageNumber="26">
<emphasis id="B96B57548D640166FF60116C0649472D" bold="true" box="[151,287,1807,1829]" pageId="25" pageNumber="26">FIGURE 15.</emphasis>
<taxonomicName id="4C1FF0C58D640166FED2116C0547472C" authority="Seeman, 2019" authorityName="Seeman" authorityYear="2019" box="[293,529,1807,1829]" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="southcotti" status="sp. nov.">
<emphasis id="B96B57548D640166FED2116C0547472C" box="[293,529,1807,1829]" italics="true" pageId="25" pageNumber="26">Megisthanus southcotti</emphasis>
</taxonomicName>
<emphasis id="B96B57548D640166FDE0116C0531472D" bold="true" box="[535,615,1807,1828]" pageId="25" pageNumber="26">
<taxonomicNameLabel id="A258EA2F8D640166FDE0116C0531472D" box="[535,615,1807,1828]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(A) female subcapitulum and palp trochanter–genu (shown detached); (B) major male subcapitulum and palp trochanter–genu. Arrows indicate spurs.
</paragraph>
</caption>
<subSubSection id="C305D8CD8D640165FF30111305434115" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26" type="diagnosis">
<paragraph id="8BA08B468D640165FF30111305434115" blockId="25.[151,1436,1904,2037]" lastBlockId="26.[151,1437,151,608]" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26">
<emphasis id="B96B57548D640166FF30111306874780" bold="true" box="[199,465,1904,1929]" pageId="25" pageNumber="26">Differential diagnosis.</emphasis>
This species is most similar to 
<taxonomicName id="4C1FF0C58D640166FCBC1112048A4781" authorityName="Seeman" authorityYear="2019" box="[843,988,1904,1928]" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="zachariei">
<emphasis id="B96B57548D640166FCBC111204344781" box="[843,866,1905,1928]" italics="true" pageId="25" pageNumber="26">M</emphasis>
. 
<emphasis id="B96B57548D640166FC831113048A4781" box="[884,988,1904,1928]" italics="true" pageId="25" pageNumber="26">zachariei</emphasis>
</taxonomicName>
in having females with both setae st4 absent and their dorsal setae of all one form, being thin and sparsely barbed. Both species are also the only 
<taxonomicName id="4C1FF0C58D640166FF6011DA067047D8" authorityName="Thorell" authorityYear="1882" box="[151,294,1977,2001]" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="25" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis id="B96B57548D640166FF6011DA067047D8" box="[151,294,1977,2001]" italics="true" pageId="25" pageNumber="26">Megisthanus</emphasis>
</taxonomicName>
found on the genus 
<taxonomicName id="4C1FF0C58D640166FDDF11DA05EB47D8" authorityName="Kaup" authorityYear="1868" box="[552,701,1977,2001]" class="Insecta" family="Passallidae" genus="Aulacocyclus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis id="B96B57548D640166FDDF11DA05EB47D8" box="[552,701,1977,2001]" italics="true" pageId="25" pageNumber="26">Aulacocyclus</emphasis>
</taxonomicName>
. Female 
<taxonomicName id="4C1FF0C58D640166FCC711DA049F47D8" authorityName="Seeman" authorityYear="2019" box="[816,969,1977,2001]" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="southcotti">
<emphasis id="B96B57548D640166FCC711DA041147D9" box="[816,839,1977,2000]" italics="true" pageId="25" pageNumber="26">M</emphasis>
. 
<emphasis id="B96B57548D640166FCAC11DA049F47D8" box="[859,969,1977,2001]" italics="true" pageId="25" pageNumber="26">southcotti</emphasis>
</taxonomicName>
are most easily distinguished from 
<taxonomicName id="4C1FF0C58D640166FA8A11DA07A947FC" authorityName="Seeman" authorityYear="2019" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="zachariei">
<emphasis id="B96B57548D640166FA8A11DA02C247D9" box="[1405,1428,1977,2000]" italics="true" pageId="25" pageNumber="26">M</emphasis>
. 
<emphasis id="B96B57548D640166FF6011BE07A947FC" box="[151,255,2013,2037]" italics="true" pageId="25" pageNumber="26">zachariei</emphasis>
</taxonomicName>
by the length of the sternogynal setae (60–110 versus 
<quantity id="4CE726A38D640166FC9711BF049547FC" box="[864,963,2012,2037]" metricMagnitude="-1" metricUnit="m" metricValue="6.984999999999999" metricValueMax="8.89" metricValueMin="5.08" pageId="25" pageNumber="26" unit="in" value="27.5" valueMax="35.0" valueMin="20.0">20–35 in</quantity>
<taxonomicName id="4C1FF0C58D640166FC3E11BE030147FC" authorityName="Seeman" authorityYear="2019" box="[969,1111,2013,2037]" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="zachariei">
<emphasis id="B96B57548D640166FC3E11BE04B647FD" box="[969,992,2013,2036]" italics="true" pageId="25" pageNumber="26">M</emphasis>
. 
<emphasis id="B96B57548D640166FC1811BE030147FC" box="[1007,1111,2013,2037]" italics="true" pageId="25" pageNumber="26">zachariei</emphasis>
</taxonomicName>
), and by the number of setae behind the genital shields within the ventral shield margin (5–11 setae 
<emphasis id="B96B57548D670165FC4416FA049C40B9" box="[947,970,153,176]" italics="true" pageId="26" pageNumber="27">vs</emphasis>
1–2 setae). This reduction in hypertrichy in 
<taxonomicName id="4C1FF0C58D670165FF4216DE061240DD" authorityName="Seeman" authorityYear="2019" box="[181,324,188,212]" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="zachariei">
<emphasis id="B96B57548D670165FF4216DE079A40DD" box="[181,204,189,212]" italics="true" pageId="26" pageNumber="27">M</emphasis>
. 
<emphasis id="B96B57548D670165FF2B16DF061240DD" box="[220,324,188,212]" italics="true" pageId="26" pageNumber="27">zachariei</emphasis>
</taxonomicName>
is also apparent in males (
<figureCitation id="132497C38D670165FD8116DF058B40DD" box="[630,733,188,213]" captionStart="FIGURE 22" captionStartId="34.[151,250,1933,1955]" captionTargetBox="[199,1380,210,1889]" captionTargetId="figure@34.[186,1410,193,1915]" captionTargetPageId="34" captionText="FIGURE 22. Megisthanus zachariei sp. nov. (A) female, posterior dorsal shield; (B) female, sternogenital region; (C) internal genitalia; (D) female, femur IV; (E) major male, sternogenital region. Abbreviations: g1 = anterior pair of genital setae; lg = latigynal remnant; mg = mesogynal shield; va = vaginal arms; vh = vaginal heads." httpUri="https://zenodo.org/record/2600902/files/figure.png" pageId="26" pageNumber="27">Fig. 22E</figureCitation>
). Female 
<taxonomicName id="4C1FF0C58D670165FCBB16DE04B440DD" authorityName="Seeman" authorityYear="2019" box="[844,994,188,212]" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="southcotti">
<emphasis id="B96B57548D670165FCBB16DE043540DD" box="[844,867,189,212]" italics="true" pageId="26" pageNumber="27">M</emphasis>
. 
<emphasis id="B96B57548D670165FC8416DF04B440DD" box="[883,994,188,212]" italics="true" pageId="26" pageNumber="27">southcotti</emphasis>
</taxonomicName>
also have short dorsal setae with large barbs (longer dorsal setae with small barbs in 
<taxonomicName id="4C1FF0C58D670165FD6D1682047140F1" authorityName="Seeman" authorityYear="2019" box="[666,807,224,248]" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="zachariei">
<emphasis id="B96B57548D670165FD6D168205E740F1" box="[666,689,225,248]" italics="true" pageId="26" pageNumber="27">M</emphasis>
. 
<emphasis id="B96B57548D670165FD481683047140F1" box="[703,807,224,248]" italics="true" pageId="26" pageNumber="27">zachariei</emphasis>
</taxonomicName>
) and a denser setation in the mid-opisthonotal region of the dorsal shield (
<figureCitation id="132497C38D670165FEAA1767069F4115" box="[349,457,260,285]" captionStart="FIGURE 12" captionStartId="22.[151,250,1822,1844]" captionTargetBox="[157,1428,207,1794]" captionTargetId="figure@22.[151,1436,193,1801]" captionTargetPageId="22" captionText="FIGURE 12. Megisthanus southcotti sp. nov. photomicrographs. (A) female, dorsal shield; (B) female, venter; (C) major male, venter; (D) major male, sternoventral shield. Scale bars: 200 µm (A–C), 100 µm (D)." httpUri="https://zenodo.org/record/2600882/files/figure.png" pageId="26" pageNumber="27">Figs 12A</figureCitation>
, 
<figureCitation id="132497C38D670165FE231767055E4115" box="[468,520,260,284]" captionStart="FIGURE 21" captionStartId="33.[151,250,1957,1979]" captionTargetBox="[170,1410,633,1925]" captionTargetId="figure@33.[161,1425,625,1936]" captionTargetPageId="33" captionText="FIGURE 21. Megisthanus zachariei sp. nov. photomicrographs. (A) female dorsal shield; (B) female venter; (C) female posterior dorsal shield; (D) major male ventrianal shield. Scale bars: 200 µm (A–B); 100 µm (C–D)." httpUri="https://zenodo.org/record/2600900/files/figure.png" pageId="26" pageNumber="27">21A</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C305D8CD8D670165FF30174405834211" pageId="26" pageNumber="27" type="discussion">
<paragraph id="8BA08B468D670165FF30174403A6418E" blockId="26.[151,1437,151,608]" pageId="26" pageNumber="27">
<emphasis id="B96B57548D670165FF301744066D4149" bold="true" box="[199,315,295,320]" pageId="26" pageNumber="27">Remarks.</emphasis>
This species is usually found on 
<taxonomicName id="4C1FF0C58D670165FD43174B04014149" authorityName="Kuwert" authorityYear="1891" box="[692,855,296,320]" class="Insecta" family="Passallidae" genus="Aulacocyclus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="fracticornis">
<emphasis id="B96B57548D670165FD43174B05934136" box="[692,709,296,319]" italics="true" pageId="26" pageNumber="27">A</emphasis>
. 
<emphasis id="B96B57548D670165FD23174B04014149" box="[724,855,296,320]" italics="true" pageId="26" pageNumber="27">fracticornis</emphasis>
</taxonomicName>
in north-eastern 
<collectingRegion id="49DB45A48D670165FBE1174403C84149" box="[1046,1182,295,320]" country="Australia" name="Queensland" pageId="26" pageNumber="27">Queensland</collectingRegion>
. Each beetle tended to carry just one mite each (9 of 11 
<taxonomicName id="4C1FF0C58D670165FDFC172F05FB416D" authorityName="Kuwert" authorityYear="1891" box="[523,685,332,356]" class="Insecta" family="Passallidae" genus="Aulacocyclus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="fracticornis">
<emphasis id="B96B57548D670165FDFC172F054A416A" box="[523,540,332,355]" italics="true" pageId="26" pageNumber="27">A</emphasis>
. 
<emphasis id="B96B57548D670165FDDD172F05FB416D" box="[554,685,332,356]" italics="true" pageId="26" pageNumber="27">fracticornis</emphasis>
</taxonomicName>
). Two specimens were collected from the unusual passalid species 
<taxonomicName id="4C1FF0C58D670165FF60171306FB4181" authorityName="Mjoberg" authorityYear="1917" box="[151,429,368,392]" class="Insecta" family="Passallidae" genus="Austropassalus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="hultgreni">
<emphasis id="B96B57548D670165FF60171306FB4181" box="[151,429,368,392]" italics="true" pageId="26" pageNumber="27">Austropassalus hultgreni</emphasis>
</taxonomicName>
. Despite the atypical host, the specimens agree closely with 
<taxonomicName id="4C1FF0C58D670165FBA2171203BE4181" authorityName="Seeman" authorityYear="2019" box="[1109,1256,368,392]" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="southcotti">
<emphasis id="B96B57548D670165FBA21712033A4181" box="[1109,1132,369,392]" italics="true" pageId="26" pageNumber="27">M</emphasis>
. 
<emphasis id="B96B57548D670165FB8D171303BE4181" box="[1146,1256,368,392]" italics="true" pageId="26" pageNumber="27">southcotti</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BA08B468D670165FF3017F705834211" blockId="26.[151,1437,151,608]" pageId="26" pageNumber="27">
The specimens from the South Australian Museum were all marked by Dr Herbert Womersley as “ 
<taxonomicName id="4C1FF0C58D670165FF5417DB06FB41D9" authority="Seeman, 2019" authorityName="Seeman" authorityYear="2019" box="[163,429,440,464]" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="southcotti" status="sp. nov.">
<emphasis id="B96B57548D670165FF5417DB06FB41D9" box="[163,429,440,464]" italics="true" pageId="26" pageNumber="27">Megisthanus southcotti</emphasis>
</taxonomicName>
<emphasis id="B96B57548D670165FE4C17D406A941D9" bold="true" box="[443,511,439,464]" pageId="26" pageNumber="27">
<taxonomicNameLabel id="A258EA2F8D670165FE4C17D406A941D9" box="[443,511,439,464]" pageId="26" pageNumber="27" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
”, a name that was never published. This material from the South Australian Museum includes an unusual record of two females from a carabid beetle. Considering the strong preference for passalid hosts, this record is treated with suspicion.
</paragraph>
</subSubSection>
<subSubSection id="C305D8CD8D670165FF30144704AD4269" pageId="26" pageNumber="27" type="etymology">
<paragraph id="8BA08B468D670165FF30144704AD4269" blockId="26.[151,1437,151,608]" pageId="26" pageNumber="27">
<emphasis id="B96B57548D670165FF30144706184234" bold="true" box="[199,334,548,573]" pageId="26" pageNumber="27">Etymology.</emphasis>
This species is named for Dr Ronald Southcott, thus fulfilling Dr Herbert Womersley’s intentions to honour one of Australia’s most significant acarologists with this patronym.
</paragraph>
</subSubSection>
</treatment>
</document>