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<document id="EB425578139F82645EDCF185A623CCBF" ID-DOI="10.11646/phytotaxa.361.2.2" ID-ISSN="1179-3163" ID-Zenodo-Dep="13705385" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725585223051" checkinUser="felipe" docAuthor="Ranjbar, Massoud, Askari, Ali &amp; Sw, Someh Mohammadi" docDate="2018" docId="03AB87D74272FFF2A3FBFA2288518559" docLanguage="en" docName="phytotaxa.361.2.2.pdf" docOrigin="Phytotaxa 361 (2)" docSource="http://dx.doi.org/10.11646/phytotaxa.361.2.2" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Centaurea palanganensis Ranjbar &amp; Askari 2018, sp. nov." docType="treatment" docVersion="2" lastPageNumber="166" masterDocId="FF92FFAF427BFFFDA373FFA489048644" masterDocTitle="A contribution to Centaurea sect. Microlophus (Asteraceae, Cardueae) in Iran" masterLastPageNumber="167" masterPageNumber="151" pageNumber="160" updateTime="1725597091092" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="264C6006C8AE8DA4B9369ECC6AEB7022" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="AD5C691712D02819CDA8D6D6C265973C">A contribution to Centaurea sect. Microlophus (Asteraceae, Cardueae) in Iran</mods:title>
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<mods:name id="923CFAD4E1E916E2497BF5AF67FD8DDF" type="personal">
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<mods:namePart id="58A30EEC45F95869A1820F011F5241C5">Ranjbar, Massoud</mods:namePart>
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<mods:namePart id="4224E2EDC0467A610144B603210261C0">Askari, Ali</mods:namePart>
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<mods:namePart id="D7CD0BC9137CEE01640A33EAFFC0529D">Sw, Someh Mohammadi</mods:namePart>
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<mods:title id="E9E3433D11236BA166D62E952111878A">Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part id="8ACB060743F773BCF14622B978F5776B">
<mods:date id="07E735E54760240D0CE1F8607521A07B">2018</mods:date>
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<mods:number id="3E86706A5ED3D6DFD6372FB25F296302">2018-07-19</mods:number>
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<mods:url id="4882C8AF79BEBD691B4B0CA4D1E38C40">http://dx.doi.org/10.11646/phytotaxa.361.2.2</mods:url>
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<mods:classification id="AA145C245B3A25443248D4D30CACB81D">journal article</mods:classification>
<mods:identifier id="B2AE176D5851B610E96F4386CDDEEC56" type="DOI">10.11646/phytotaxa.361.2.2</mods:identifier>
<mods:identifier id="864BE01E13D710907CDB3CF99ABCD529" type="ISSN">1179-3163</mods:identifier>
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<treatment id="03AB87D74272FFF2A3FBFA2288518559" ID-DOI="http://doi.org/10.5281/zenodo.13705359" ID-Zenodo-Dep="13705359" LSID="urn:lsid:plazi:treatment:03AB87D74272FFF2A3FBFA2288518559" httpUri="http://treatment.plazi.org/id/03AB87D74272FFF2A3FBFA2288518559" lastPageId="15" lastPageNumber="166" pageId="9" pageNumber="160">
<subSubSection id="C318654A4272FFF4A3FBFA228A6083E5" box="[136,868,1413,1441]" pageId="9" pageNumber="160" type="nomenclature">
<paragraph id="8BBD36C14272FFF4A3FBFA228A6083E5" blockId="9.[136,868,1413,1441]" box="[136,868,1413,1441]" pageId="9" pageNumber="160">
<heading id="D0F581AD4272FFF4A3FBFA228A6083E5" box="[136,868,1413,1441]" fontSize="11" level="2" pageId="9" pageNumber="160" reason="3">
<taxonomicName id="4C024D424272FFF4A3FBFA228B7883E5" ID-CoL="69GBL" authority="Ranjbar &amp; Askari" authorityName="Ranjbar &amp; Askari" authorityYear="2018" box="[136,636,1413,1441]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="9" pageNumber="160" phylum="Tracheophyta" rank="species" species="palanganensis" status="sp. nov.">
<emphasis id="B976EAD34272FFF4A3FBFA2288AA83E4" bold="true" box="[136,430,1413,1440]" italics="true" pageId="9" pageNumber="160">Centaurea palanganensis</emphasis>
Ranjbar &amp; Askari
</taxonomicName>
,
<emphasis id="B976EAD34272FFF4A1FAFA228BE483E4" bold="true" box="[649,736,1414,1440]" italics="true" pageId="9" pageNumber="160">
<taxonomicNameLabel id="A24557A84272FFF4A1FAFA228BE483E4" box="[649,736,1414,1440]" pageId="9" pageNumber="160" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
<figureCitation id="13392A444272FFF4A194FA2C8A5B83E5" box="[743,863,1416,1441]" captionStart-0="FIGURE 8" captionStart-1="FIGURE 9" captionStart-2="FIGURE 10" captionStart-3="FIGURE 11" captionStartId-0="11.[136,229,2029,2050]" captionStartId-1="12.[136,229,1879,1900]" captionStartId-2="13.[136,229,1980,2001]" captionStartId-3="14.[136,229,1994,2015]" captionTargetBox-0="[171,1405,205,2004]" captionTargetBox-1="[168,1431,239,1809]" captionTargetBox-2="[252,1382,200,1930]" captionTargetBox-3="[197,1345,237,1928]" captionTargetId-0="figure-14@11.[154,1434,190,2004]" captionTargetId-1="figure-17@12.[151,1436,190,1854]" captionTargetId-2="figure-14@13.[151,1436,190,1954]" captionTargetId-3="figure-17@14.[151,1436,190,1970]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionTargetPageId-2="13" captionTargetPageId-3="14" captionText-0="FIGURE 8. Centaurea palanganensis (Ranjbar &amp; Askari 39100, BASU). A. habit. Scale A = 5 cm." captionText-1="FIGURE 9. Centaurea palanganensis (Ranjbar &amp; Askari 39100, BASU). A. habit. Scale A = 5 cm." captionText-2="FIGURE 10. Centaurea palanganensis (Ranjbar &amp; Askari 39100, BASU). A. close up of capitule. B1–B3: outer phyllaries. C1–C3: median phyllaries. D1–D3: inner phyllaries. E: leaf stomata. F: leaf indumentum, Scale A = 1 cm." captionText-3="FIGURE 11. Morphological characters measured in Centaurea palanganensis (Ranjbar &amp; Askari 39100, BASU). A. middle stem leaf. B. peripheral floret. C. involucre. D. involucral bract. For character explanations see Table 1." figureDoi-0="http://doi.org/10.5281/zenodo.13705405" figureDoi-1="http://doi.org/10.5281/zenodo.13705407" figureDoi-2="http://doi.org/10.5281/zenodo.13705409" figureDoi-3="http://doi.org/10.5281/zenodo.13705411" httpUri-0="https://zenodo.org/record/13705405/files/figure.png" httpUri-1="https://zenodo.org/record/13705407/files/figure.png" httpUri-2="https://zenodo.org/record/13705409/files/figure.png" httpUri-3="https://zenodo.org/record/13705411/files/figure.png" pageId="9" pageNumber="160">Figs. 8–11</figureCitation>
.
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C318654A4272FFF4A3FBFA0A88D58017" pageId="9" pageNumber="160" type="diagnosis">
<paragraph id="8BBD36C14272FFF4A3FBFA0A88D58017" blockId="9.[136,1452,1454,1691]" pageId="9" pageNumber="160">
<emphasis id="B976EAD34272FFF4A3FBFA0A89EF8387" bold="true" box="[136,235,1454,1475]" pageId="9" pageNumber="160">Diagnosis</emphasis>
:—
<taxonomicName id="4C024D424272FFF4A27AFA0A88FB8387" authorityName="Ranjbar &amp; Askari" authorityYear="2018" box="[265,511,1454,1475]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="9" pageNumber="160" phylum="Tracheophyta" rank="species" species="palanganensis">
<emphasis id="B976EAD34272FFF4A27AFA0A88FB8387" box="[265,511,1454,1475]" italics="true" pageId="9" pageNumber="160">Centaurea palanganensis</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="4C024D424272FFF4A10FFA0A8D5F8387" authority="(Janka 1872: 178) Gugler (1908: 201)" authorityName="Gugler" authorityPageNumber="201" authorityYear="1908" baseAuthorityName="Janka" baseAuthorityPageNumber="178" baseAuthorityYear="1872" box="[636,1115,1454,1475]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="9" pageNumber="160" phylum="Tracheophyta" rank="species" species="thracica">
<emphasis id="B976EAD34272FFF4A10FFA0A8BEC8387" box="[636,744,1454,1475]" italics="true" pageId="9" pageNumber="160">C. thracica</emphasis>
(
<bibRefCitation id="EF934B304272FFF4A186FA0A8A918387" author="Janka, V." box="[757,917,1454,1475]" pageId="9" pageNumber="160" pagination="174 - 181" refId="ref5974" refString="Janka, V. (1872) Plantarum novarum turcicarum breviarium. Oesterreichische Botanische Zeitschrift 22: 174 - 181. https: // doi. org / 10.1007 / BF 01644772" type="journal article" year="1872">Janka 1872: 178</bibRefCitation>
)
<bibRefCitation id="EF934B304272FFF4A0D1FA0A8D5F8387" author="Gugler, W." box="[930,1115,1454,1475]" pageId="9" pageNumber="160" pagination="15 - 297" refId="ref5950" refString="Gugler, W. (1908) Die Centaureen des Ungarischen National-Museums. Annales Historico-Naturales Musei Nationalis Hungarici 6: 15 - 297." type="journal article" year="1908">Gugler (1908: 201)</bibRefCitation>
</taxonomicName>
in cauline branching and colour of inner phyllaries, but differs from the latter by having median cauline leaves decurrent for
<quantity id="4CFA9B244272FFF4A75AFA768D9583A3" box="[1065,1169,1490,1511]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="1.8" metricValueMin="1.2" pageId="9" pageNumber="160" unit="mm" value="15.0" valueMax="18.0" valueMin="12.0">12–18 mm</quantity>
(vs.
<quantity id="4CFA9B244272FFF4A7B0FA768C1383A3" box="[1219,1303,1490,1511]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="9" pageNumber="160" unit="mm" value="6.0" valueMax="7.0" valueMin="5.0">5–7 mm</quantity>
), upper cauline leaves entire (vs. crenate), upper cauline leaves ovate (vs. oblong to lanceolate), involucres 7–20 ×
<quantity id="4CFA9B244272FFF4A7FAFA528DE1804F" box="[1161,1253,1526,1547]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" metricValueMax="1.5" metricValueMin="0.6" pageId="9" pageNumber="160" unit="mm" value="10.5" valueMax="15.0" valueMin="6.0">6–15 mm</quantity>
(vs. 30–40 ×
<quantity id="4CFA9B244272FFF4A603FA5289E2806B" metricMagnitude="-2" metricUnit="m" metricValue="3.25" metricValueMax="3.5" metricValueMin="3.0" pageId="9" pageNumber="160" unit="mm" value="32.5" valueMax="35.0" valueMin="30.0">30–35 mm</quantity>
), median phyllaries
<quantity id="4CFA9B244272FFF4A2DBF9BE8B0B806B" box="[424,527,1562,1583]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.3" metricValueMin="1.0" pageId="9" pageNumber="160" unit="mm" value="11.5" valueMax="13.0" valueMin="10.0">10–13 mm</quantity>
(vs.
<quantity id="4CFA9B244272FFF4A14DF9BE8BA3806B" box="[574,679,1562,1583]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" metricValueMax="1.4" metricValueMin="1.3" pageId="9" pageNumber="160" unit="mm" value="13.5" valueMax="14.0" valueMin="13.0">13–14 mm</quantity>
) long, median cauline leaves oblong or elliptic (vs. ovate to oblong), spine up to
<quantity id="4CFA9B244272FFF4A3CEF99A89F08017" box="[189,244,1598,1619]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="160" unit="mm" value="2.0">2 mm</quantity>
(vs. up to
<quantity id="4CFA9B244272FFF4A228F99A88908017" box="[347,404,1598,1619]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="9" pageNumber="160" unit="mm" value="5.0">5 mm</quantity>
) long.
</paragraph>
</subSubSection>
<subSubSection id="C318654A4272FFF2A3FBF9C68D158761" lastPageId="15" lastPageNumber="166" pageId="9" pageNumber="160" type="materials_examined">
<paragraph id="8BBD36C14272FFF4A3FBF9C6884980DF" blockId="9.[136,1452,1454,1691]" pageId="9" pageNumber="160">
<emphasis id="B976EAD34272FFF4A3FBF9C689DE8033" bold="true" box="[136,218,1634,1655]" pageId="9" pageNumber="160">Type:—</emphasis>
<collectingCountry id="F31576514272FFF4A3A9F9C6881C8033" box="[218,280,1634,1655]" name="Iran" pageId="9" pageNumber="160">IRAN</collectingCountry>
. Kurdestan, Tefin before Palangan village,
<geoCoordinate id="EE3650064272FFF4A1DDF9C68A378033" box="[686,819,1634,1655]" degrees="35" direction="north" minutes="05" orientation="latitude" pageId="9" pageNumber="160" precision="15" seconds="02" value="35.08389">35° 05’ 02’’ N</geoCoordinate>
,
<geoCoordinate id="EE3650064272FFF4A04FF9C68ABB8033" box="[828,959,1634,1655]" degrees="46" direction="east" minutes="36" orientation="longitude" pageId="9" pageNumber="160" precision="15" seconds="58" value="46.616108">46° 36’ 58’’ E</geoCoordinate>
,
<quantity id="4CFA9B244272FFF4A0BBF9C68D158033" box="[968,1041,1634,1655]" metricMagnitude="3" metricUnit="m" metricValue="1.097" pageId="9" pageNumber="160" unit="m" value="1097.0">1097 m</quantity>
,
<date id="FFBC10014272FFF4A769F9C68D898033" box="[1050,1165,1634,1655]" pageId="9" pageNumber="160" value="2016-05-05">5 May 2016</date>
,
<emphasis id="B976EAD34272FFF4A7E5F9C68C3B8033" box="[1174,1343,1634,1655]" italics="true" pageId="9" pageNumber="160">Ranjbar &amp; Askari</emphasis>
(
<typeStatus id="54B988634272FFF4A639F9C68CAC8033" box="[1354,1448,1634,1655]" pageId="9" pageNumber="160" type="holotype">Holotype</typeStatus>
:
<collectionCode id="ED13AE044272FFF4A3CEF92289F980DF" box="[189,253,1670,1691]" country="Iran" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15764" lsid="urn:lsid:biocol.org:col:15764" name="Bu-Ali Sina University" pageId="9" pageNumber="160" type="Herbarium">BASU</collectionCode>
<emphasis id="B976EAD34272FFF4A270F922883B80DF" box="[259,319,1670,1691]" italics="true" pageId="9" pageNumber="160">39100</emphasis>
).
</paragraph>
<paragraph id="8BBD36C14272FFF2A3FBF9688D158761" blockId="9.[136,1452,1740,2089]" lastBlockId="15.[136,1452,160,797]" lastPageId="15" lastPageNumber="166" pageId="9" pageNumber="160">
Perennial plant,
<quantity id="4CFA9B244272FFF4A24EF96888A880A1" box="[317,428,1740,1765]" metricMagnitude="-1" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="9" pageNumber="160" unit="cm" value="70.0" valueMax="80.0" valueMin="60.0">60–80 cm</quantity>
tall, usually pale green throughout. Stem erect, densely leaved, simple or with 2–4 branches in upper part, mostly branched in lower to upper parts, predominately striate striped, thickened below capitula, densely covered with hirsute-articulate hairs. Leaves papyraceous, ± densely covered with hirsute-articulate hairs, margins scabrous. Basal and lower cauline leaves petiolate,
<quantity id="4CFA9B244272FFF4A1B2F89C8A348115" box="[705,816,1848,1873]" metricMagnitude="-1" metricUnit="m" metricValue="1.15" metricValueMax="1.3" metricValueMin="1.0" pageId="9" pageNumber="160" unit="cm" value="11.5" valueMax="13.0" valueMin="10.0">10–13 cm</quantity>
long, lyrate; terminal segment large, oblong to triangular or ovate, 7–13 ×
<quantity id="4CFA9B244272FFF4A223F8F888B38131" box="[336,439,1884,1909]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" metricValueMax="10.0" metricValueMin="4.0" pageId="9" pageNumber="160" unit="cm" value="7.0" valueMax="10.0" valueMin="4.0">4–10 cm</quantity>
; lateral segments smaller, in 2–3 pairs, oblong-lanceolate or lanceolate-triangular, 4–5 ×
<quantity id="4CFA9B244272FFF4A3FBF824880881DD" box="[136,268,1920,1945]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" metricValueMax="2.5" metricValueMin="1.5" pageId="9" pageNumber="160" unit="cm" value="2.0" valueMax="2.5" valueMin="1.5">1.5–2.5 cm</quantity>
. Median cauline leaves simple, oblong or elliptic, 6–8 ×
<quantity id="4CFA9B244272FFF4A0D4F8248D1181DD" box="[935,1045,1920,1945]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" metricValueMax="3.0" metricValueMin="1.2" pageId="9" pageNumber="160" unit="cm" value="2.1" valueMax="3.0" valueMin="1.2">1.2–3 cm</quantity>
, decurrent for
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, acute or acuminate at apex, margin crenate to sinuate, venation reticulate. Upper cauline leaves ovate 1.5–3 ×
<quantity id="4CFA9B244272FFF4A657F8008CA381F9" box="[1316,1447,1956,1981]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="9.0" metricValueMin="2.0" pageId="9" pageNumber="160" unit="cm" value="0.55" valueMax="0.9" valueMin="0.2">0.2–0.9 cm</quantity>
, acuminate to mucronate at apex, margin entire, uppermost ones slightly enveloping involucres. Involucres ovoidcupuliform or broadly ovoid, truncate at base, 0.7–2 ×
<quantity id="4CFA9B244272FFF4A199F8488A638E41" box="[746,871,2028,2053]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" metricValueMax="1.5" metricValueMin="0.6" pageId="9" pageNumber="160" unit="cm" value="1.05" valueMax="1.5" valueMin="0.6">0.6–1.5 cm</quantity>
long. Phyllaries multiseriate, imbricate, coriaceous, yellowish green, glabrous; outer phyllaries ovate,
<quantity id="4CFA9B244272FFF4A1C6F7B48A158E6D" box="[693,785,2064,2089]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="9" pageNumber="160" unit="mm" value="6.0" valueMax="7.0" valueMin="5.0">5–7 mm</quantity>
long, yellowish green. Median phyllaries oblong to elliptic,
<quantity id="4CFA9B244274FFF2A3FBFF04880186FD" box="[136,261,160,185]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.3" metricValueMin="1.0" pageId="15" pageNumber="166" unit="mm" value="11.5" valueMax="13.0" valueMin="10.0">10–13 mm</quantity>
long, yellowish green. Inner phyllaries oblong,
<quantity id="4CFA9B244274FFF2A044FF048AB786FD" box="[823,947,160,185]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="1.6" metricValueMin="1.4" pageId="15" pageNumber="166" unit="mm" value="15.0" valueMax="16.0" valueMin="14.0">14–16 mm</quantity>
long, pale purple in middle. Appendages a simple spine,
<quantity id="4CFA9B244274FFF2A259FF60888E8699" box="[298,394,196,221]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="15" pageNumber="166" unit="mm" value="1.5" valueMax="2.0" valueMin="1.0">1–2 mm</quantity>
long in median phyllaries, deciduous. Marginal flowers sterile, not radiant, central flowers hermaphrodite, up to
<quantity id="4CFA9B244274FFF2A20CFF4C88D48745" box="[383,464,232,257]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="15" pageNumber="166" unit="mm" value="12.0">12 mm</quantity>
long; peripheral sterile, not radiant, up to
<quantity id="4CFA9B244274FFF2A0C4FF4C8D0E8745" box="[951,1034,232,257]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" pageId="15" pageNumber="166" unit="mm" value="22.0">22 mm</quantity>
, limb lobes linear. Achenes
<quantity id="4CFA9B244274FFF2A63EFF4C8CA88745" box="[1357,1452,232,257]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="15" pageNumber="166" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">4–5 mm</quantity>
long. Pappus
<quantity id="4CFA9B244274FFF2A253FEA888798761" box="[288,381,268,293]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="15" pageNumber="166" unit="mm" value="5.5" valueMax="6.0" valueMin="5.0">5–6 mm</quantity>
long, inner row very short, multiseriate, whitish, scabrous.
</paragraph>
</subSubSection>
<caption id="DF7D66494271FFF7A3FBF8EB88C98194" ID-DOI="http://doi.org/10.5281/zenodo.13705403" ID-Zenodo-Dep="13705403" httpUri="https://zenodo.org/record/13705403/files/figure.png" pageId="10" pageNumber="161" startId="10.[136,229,1871,1892]" targetBox="[221,1365,209,1846]" targetPageId="10" targetType="figure">
<paragraph id="8BBD36C14271FFF7A3FBF8EB88C98194" blockId="10.[136,1452,1871,2000]" pageId="10" pageNumber="161">
<emphasis id="B976EAD34271FFF7A3FBF8EB89F98120" bold="true" box="[136,253,1871,1892]" pageId="10" pageNumber="161">FIGURE 7.</emphasis>
Representative meiotic cells of
<taxonomicName id="4C024D424271FFF7A15DF8EB8A628120" authorityName="Ranjbar &amp; Askari" authorityYear="2018" box="[558,870,1871,1892]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="161" phylum="Tracheophyta" rank="species" species="masjedsoleymanensis">
<emphasis id="B976EAD34271FFF7A15DF8EB8A628120" box="[558,870,1871,1892]" italics="true" pageId="10" pageNumber="161">Centaurea masjedsoleymanensis</emphasis>
</taxonomicName>
<emphasis id="B976EAD34271FFF7A018F8EB8A9F8120" bold="true" box="[875,923,1871,1892]" pageId="10" pageNumber="161">A–G</emphasis>
and
<taxonomicName id="4C024D424271FFF7A0BBF8EB8D6B8120" authorityName="Ranjbar &amp; Askari" authorityYear="2018" box="[968,1135,1871,1892]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="161" phylum="Tracheophyta" rank="species" species="palanganensis">
<emphasis id="B976EAD34271FFF7A0BBF8EB8D6B8120" box="[968,1135,1871,1892]" italics="true" pageId="10" pageNumber="161">C. palanganensis</emphasis>
</taxonomicName>
<emphasis id="B976EAD34271FFF7A707F8EB8DA18120" bold="true" box="[1140,1189,1871,1892]" pageId="10" pageNumber="161">H–O</emphasis>
.
<emphasis id="B976EAD34271FFF7A7C3F8EB8DCC8120" bold="true" box="[1200,1224,1871,1892]" pageId="10" pageNumber="161">A.</emphasis>
diakinesis.
<emphasis id="B976EAD34271FFF7A64BF8EB8C4A8120" bold="true" box="[1336,1358,1871,1892]" pageId="10" pageNumber="161">B.</emphasis>
anaphase I with bridge.
<emphasis id="B976EAD34271FFF7A261F8D7882D81CC" bold="true" box="[274,297,1907,1928]" pageId="10" pageNumber="161">C.</emphasis>
telophase I.
<emphasis id="B976EAD34271FFF7A2D5F8D788B981CC" bold="true" box="[422,445,1907,1928]" pageId="10" pageNumber="161">D.</emphasis>
anaphase II.
<emphasis id="B976EAD34271FFF7A14CF8D78B5181CC" bold="true" box="[575,597,1907,1928]" pageId="10" pageNumber="161">E.</emphasis>
metaphase I with precocious chromosome.
<emphasis id="B976EAD34271FFF7A08CF8D78D1581CC" bold="true" box="[1023,1041,1907,1928]" pageId="10" pageNumber="161">F.</emphasis>
anaphase I.
<emphasis id="B976EAD34271FFF7A7F8F8D78DA081CC" bold="true" box="[1163,1188,1907,1928]" pageId="10" pageNumber="161">G.</emphasis>
anaphase I with bridge.
<emphasis id="B976EAD34271FFF7A6E0F8D78CA881CC" bold="true" box="[1427,1452,1907,1928]" pageId="10" pageNumber="161">H.</emphasis>
anaphase I with bridge.
<emphasis id="B976EAD34271FFF7A218F83388A181E8" bold="true" box="[363,421,1943,1964]" pageId="10" pageNumber="161">I &amp; J.</emphasis>
metaphase I with precocious chromosome.
<emphasis id="B976EAD34271FFF7A035F8338A5B81E8" bold="true" box="[838,863,1943,1964]" pageId="10" pageNumber="161">K.</emphasis>
diakinesis.
<emphasis id="B976EAD34271FFF7A0BCF8338AE181E8" bold="true" box="[975,997,1943,1964]" pageId="10" pageNumber="161">L.</emphasis>
metaphase I.
<emphasis id="B976EAD34271FFF7A71BF8338D8081E8" bold="true" box="[1128,1156,1943,1964]" pageId="10" pageNumber="161">M.</emphasis>
anaphase I.
<emphasis id="B976EAD34271FFF7A789F8338C1681E8" bold="true" box="[1274,1298,1943,1964]" pageId="10" pageNumber="161">N.</emphasis>
anaphase I with laggard.
<emphasis id="B976EAD34271FFF7A3AFF81F89F18194" bold="true" box="[220,245,1979,2000]" pageId="10" pageNumber="161">O.</emphasis>
bivalent at diakinesis.
</paragraph>
</caption>
<caption id="DF7D66494270FFF6A3FBF8498D438E46" ID-DOI="http://doi.org/10.5281/zenodo.13705405" ID-Zenodo-Dep="13705405" box="[136,1095,2029,2050]" httpUri="https://zenodo.org/record/13705405/files/figure.png" pageId="11" pageNumber="162" startId="11.[136,229,2029,2050]" targetBox="[171,1405,205,2004]" targetPageId="11" targetType="figure">
<paragraph id="8BBD36C14270FFF6A3FBF8498D438E46" blockId="11.[136,1095,2029,2050]" box="[136,1095,2029,2050]" pageId="11" pageNumber="162">
<emphasis id="B976EAD34270FFF6A3FBF84989F98E46" bold="true" box="[136,253,2029,2050]" pageId="11" pageNumber="162">FIGURE 8.</emphasis>
<taxonomicName id="4C024D424270FFF6A270F84988FD8E46" authorityName="Ranjbar &amp; Askari" authorityYear="2018" box="[259,505,2029,2050]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="11" pageNumber="162" phylum="Tracheophyta" rank="species" species="palanganensis">
<emphasis id="B976EAD34270FFF6A270F84988FD8E46" box="[259,505,2029,2050]" italics="true" pageId="11" pageNumber="162">Centaurea palanganensis</emphasis>
</taxonomicName>
(
<emphasis id="B976EAD34270FFF6A175F8498BF08E46" box="[518,756,2029,2050]" italics="true" pageId="11" pageNumber="162">Ranjbar &amp; Askari 39100</emphasis>
, BASU).
<emphasis id="B976EAD34270FFF6A027F8498A618E46" bold="true" box="[852,869,2029,2050]" pageId="11" pageNumber="162">A</emphasis>
. habit. Scale
<emphasis id="B976EAD34270FFF6A094F8498AFC8E46" bold="true" box="[999,1016,2029,2050]" pageId="11" pageNumber="162">A</emphasis>
= 5 cm.
</paragraph>
</caption>
<caption id="DF7D66494277FFF1A3FBF8F38D4D8128" ID-DOI="http://doi.org/10.5281/zenodo.13705407" ID-Zenodo-Dep="13705407" box="[136,1097,1879,1900]" httpUri="https://zenodo.org/record/13705407/files/figure.png" pageId="12" pageNumber="163" startId="12.[136,229,1879,1900]" targetBox="[168,1431,239,1809]" targetPageId="12" targetType="figure">
<paragraph id="8BBD36C14277FFF1A3FBF8F38D4D8128" blockId="12.[136,1097,1879,1900]" box="[136,1097,1879,1900]" pageId="12" pageNumber="163">
<emphasis id="B976EAD34277FFF1A3FBF8F389F98128" bold="true" box="[136,253,1879,1900]" pageId="12" pageNumber="163">FIGURE 9.</emphasis>
<taxonomicName id="4C024D424277FFF1A270F8F388FD8128" authorityName="Ranjbar &amp; Askari" authorityYear="2018" box="[259,505,1879,1900]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="12" pageNumber="163" phylum="Tracheophyta" rank="species" species="palanganensis">
<emphasis id="B976EAD34277FFF1A270F8F388FD8128" box="[259,505,1879,1900]" italics="true" pageId="12" pageNumber="163">Centaurea palanganensis</emphasis>
</taxonomicName>
(
<emphasis id="B976EAD34277FFF1A175F8F38BF08128" box="[518,756,1879,1900]" italics="true" pageId="12" pageNumber="163">Ranjbar &amp; Askari 39100</emphasis>
, BASU).
<emphasis id="B976EAD34277FFF1A027F8F38A618128" bold="true" box="[852,869,1879,1900]" pageId="12" pageNumber="163">A</emphasis>
. habit. Scale
<emphasis id="B976EAD34277FFF1A094F8F38AFC8128" bold="true" box="[999,1016,1879,1900]" pageId="12" pageNumber="163">A</emphasis>
= 5 cm.
</paragraph>
</caption>
<caption id="DF7D66494276FFF0A3FBF8188D3C81B1" ID-DOI="http://doi.org/10.5281/zenodo.13705409" ID-Zenodo-Dep="13705409" httpUri="https://zenodo.org/record/13705409/files/figure.png" pageId="13" pageNumber="164" startId="13.[136,229,1980,2001]" targetBox="[252,1382,200,1930]" targetPageId="13" targetType="figure">
<paragraph id="8BBD36C14276FFF0A3FBF8188D3C81B1" blockId="13.[136,1452,1980,2037]" pageId="13" pageNumber="164">
<emphasis id="B976EAD34276FFF0A3FBF81888088195" bold="true" box="[136,268,1980,2001]" pageId="13" pageNumber="164">FIGURE 10.</emphasis>
<taxonomicName id="4C024D424276FFF0A267F8188B088195" authorityName="Ranjbar &amp; Askari" authorityYear="2018" box="[276,524,1980,2001]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="13" pageNumber="164" phylum="Tracheophyta" rank="species" species="palanganensis">
<emphasis id="B976EAD34276FFF0A267F8188B088195" box="[276,524,1980,2001]" italics="true" pageId="13" pageNumber="164">Centaurea palanganensis</emphasis>
</taxonomicName>
(
<emphasis id="B976EAD34276FFF0A16FF8188A148195" box="[540,784,1980,2001]" italics="true" pageId="13" pageNumber="164">Ranjbar &amp; Askari 39100</emphasis>
, BASU).
<emphasis id="B976EAD34276FFF0A006F8188A828195" bold="true" box="[885,902,1980,2001]" pageId="13" pageNumber="164">A</emphasis>
. close up of capitule.
<emphasis id="B976EAD34276FFF0A716F8188DAC8195" bold="true" box="[1125,1192,1980,2001]" pageId="13" pageNumber="164">B1–B3</emphasis>
: outer phyllaries.
<emphasis id="B976EAD34276FFF0A62DF8188CA08195" bold="true" box="[1374,1444,1980,2001]" pageId="13" pageNumber="164">C1–C3</emphasis>
: median phyllaries.
<emphasis id="B976EAD34276FFF0A24CF844888281B1" bold="true" box="[319,390,2016,2037]" pageId="13" pageNumber="164">D1–D3</emphasis>
: inner phyllaries.
<emphasis id="B976EAD34276FFF0A146F8448B4181B1" bold="true" box="[565,581,2016,2037]" pageId="13" pageNumber="164">E</emphasis>
: leaf stomata.
<emphasis id="B976EAD34276FFF0A1A1F8448BE581B1" bold="true" box="[722,737,2016,2037]" pageId="13" pageNumber="164">F</emphasis>
: leaf indumentum, Scale
<emphasis id="B976EAD34276FFF0A0A4F8448AEC81B1" bold="true" box="[983,1000,2016,2037]" pageId="13" pageNumber="164">A</emphasis>
= 1 cm.
</paragraph>
</caption>
<caption id="DF7D66494275FFF3A3FBF86E8AEE8E47" ID-DOI="http://doi.org/10.5281/zenodo.13705411" ID-Zenodo-Dep="13705411" httpUri="https://zenodo.org/record/13705411/files/figure.png" pageId="14" pageNumber="165" startId="14.[136,229,1994,2015]" targetBox="[197,1345,237,1928]" targetPageId="14" targetType="figure">
<paragraph id="8BBD36C14275FFF3A3FBF86E8AEE8E47" blockId="14.[136,1452,1994,2051]" pageId="14" pageNumber="165">
<emphasis id="B976EAD34275FFF3A3FBF86E880C819B" bold="true" box="[136,264,1994,2015]" pageId="14" pageNumber="165">FIGURE 11.</emphasis>
Morphological characters measured in
<taxonomicName id="4C024D424275FFF3A1F2F86E8A72819B" authorityName="Ranjbar &amp; Askari" authorityYear="2018" box="[641,886,1994,2015]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="14" pageNumber="165" phylum="Tracheophyta" rank="species" species="palanganensis">
<emphasis id="B976EAD34275FFF3A1F2F86E8A72819B" box="[641,886,1994,2015]" italics="true" pageId="14" pageNumber="165">Centaurea palanganensis</emphasis>
</taxonomicName>
(
<emphasis id="B976EAD34275FFF3A0F0F86E8D6B819B" box="[899,1135,1994,2015]" italics="true" pageId="14" pageNumber="165">Ranjbar &amp; Askari 39100</emphasis>
, BASU).
<emphasis id="B976EAD34275FFF3A7BDF86E8DE2819B" bold="true" box="[1230,1254,1994,2015]" pageId="14" pageNumber="165">A.</emphasis>
middle stem leaf.
<emphasis id="B976EAD34275FFF3A6E6F86E8CA8819B" bold="true" box="[1429,1452,1994,2015]" pageId="14" pageNumber="165">B.</emphasis>
peripheral floret.
<emphasis id="B976EAD34275FFF3A25CF84A88438E47" bold="true" box="[303,327,2030,2051]" pageId="14" pageNumber="165">C.</emphasis>
involucre.
<emphasis id="B976EAD34275FFF3A2C0F84A88CF8E47" bold="true" box="[435,459,2030,2051]" pageId="14" pageNumber="165">D.</emphasis>
involucral bract. For character explanations see Table 1.
</paragraph>
</caption>
<subSubSection id="C318654A4274FFF2A3CEFE948B748729" pageId="15" pageNumber="166" type="etymology">
<paragraph id="8BBD36C14274FFF2A3CEFE948DD9870D" blockId="15.[136,1452,160,797]" box="[189,1245,304,329]" pageId="15" pageNumber="166">
<emphasis id="B976EAD34274FFF2A3CEFE94883A870D" bold="true" box="[189,318,304,329]" pageId="15" pageNumber="166">Etymology</emphasis>
:—The specific epithet refers to Palangan village, where the new taxon was found.
</paragraph>
<paragraph id="8BBD36C14274FFF2A3CEFEF08B748729" blockId="15.[136,1452,160,797]" box="[189,624,340,365]" pageId="15" pageNumber="166">
<emphasis id="B976EAD34274FFF2A3CEFEF0883D8729" bold="true" box="[189,313,340,365]" pageId="15" pageNumber="166">Phenology</emphasis>
:—Flowering April to June.
</paragraph>
</subSubSection>
<subSubSection id="C318654A4274FFF2A3CEFEDC88518559" pageId="15" pageNumber="166" type="distribution">
<paragraph id="8BBD36C14274FFF2A3CEFEDC8C96879D" blockId="15.[136,1452,160,797]" pageId="15" pageNumber="166">
<emphasis id="B976EAD34274FFF2A3CEFEDC88E187D5" bold="true" box="[189,485,376,401]" pageId="15" pageNumber="166">Distribution and habitat:</emphasis>
—
<taxonomicName id="4C024D424274FFF2A172FEDC8A1987D5" authorityName="Ranjbar &amp; Askari" authorityYear="2018" box="[513,797,376,401]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="15" pageNumber="166" phylum="Tracheophyta" rank="species" species="palanganensis">
<emphasis id="B976EAD34274FFF2A172FEDC8A1987D5" box="[513,797,376,401]" italics="true" pageId="15" pageNumber="166">Centaurea palanganensis</emphasis>
</taxonomicName>
occurs in Palangan area, a mountainous region and a village in Zhavehrud Rural District, in Central District of Kamyaran County, Kurdestan Province,
<collectingCountry id="F31576514274FFF2A707FE388DA187F1" box="[1140,1189,412,437]" name="Iran" pageId="15" pageNumber="166">Iran</collectingCountry>
. Phytogeographically it is an Irano-Turanian element and grows in dry clay, stony slopes and screes, at elevations of
<quantity id="4CFA9B244274FFF2A7E5FE648C35879D" box="[1174,1329,448,473]" metricMagnitude="3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="15" pageNumber="166" unit="m" value="1500.0" valueMax="2000.0" valueMin="1000.0">1000–2000 m</quantity>
(
<figureCitation id="13392A444274FFF2A633FE648C80879D" box="[1344,1412,448,473]" captionStart="FIGURE 6" captionStartId="7.[136,229,2056,2077]" captionTargetBox="[229,1358,1003,2029]" captionTargetId="figure-372@7.[228,1359,1001,2031]" captionTargetPageId="7" captionText="FIGURE 6. Distribution of 1. Centaurea masjedsoleymanensis. 2. C. palanganensis, in Iran." figureDoi="http://doi.org/10.5281/zenodo.13705401" httpUri="https://zenodo.org/record/13705401/files/figure.png" pageId="15" pageNumber="166">Fig. 6</figureCitation>
).
</paragraph>
<paragraph id="8BBD36C14274FFF2A3CEFE408D9C8401" blockId="15.[136,1452,160,797]" pageId="15" pageNumber="166">
<emphasis id="B976EAD34274FFF2A3CEFE40885B87B9" bold="true" box="[189,351,484,509]" pageId="15" pageNumber="166">Relationships</emphasis>
:—
<taxonomicName id="4C024D424274FFF2A2F0FE408BA687B9" authorityName="Ranjbar &amp; Askari" authorityYear="2018" box="[387,674,484,509]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="15" pageNumber="166" phylum="Tracheophyta" rank="species" species="palanganensis">
<emphasis id="B976EAD34274FFF2A2F0FE408BA687B9" box="[387,674,484,509]" italics="true" pageId="15" pageNumber="166">Centaurea palanganensis</emphasis>
</taxonomicName>
is characterized with median cauline leaves decurrent, upper cauline leaves ovate, margin entire, and inner phyllaries pale purple. It is similar to
<taxonomicName id="4C024D424274FFF2A094FDAC8D628465" authorityName="Gugler" authorityYear="1908" baseAuthorityName="Janka" baseAuthorityYear="1872" box="[999,1126,520,545]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="15" pageNumber="166" phylum="Tracheophyta" rank="species" species="thracica">
<emphasis id="B976EAD34274FFF2A094FDAC8D628465" box="[999,1126,520,545]" italics="true" pageId="15" pageNumber="166">C. thracica</emphasis>
</taxonomicName>
in the colour of flowers and size of basal and lower cauline leaves, but differs in some important characters (see
<tableCitation id="C680037A4274FFF2A746FD888D838401" box="[1077,1159,556,581]" captionStart="TABLE 1" captionStartId="8.[136,224,160,183]" captionText="TABLE 1. Diagnostic morphological characters of Centaurea masjedsoleymanensis, C. pabotii, C. palanganensis and C. thracica." pageId="15" pageNumber="166">Table 1</tableCitation>
).
</paragraph>
<paragraph id="8BBD36C14274FFF2A3CEFDF48AB68491" blockId="15.[136,1452,160,797]" pageId="15" pageNumber="166">
<emphasis id="B976EAD34274FFF2A3CEFDF488E7842D" bold="true" box="[189,483,592,617]" pageId="15" pageNumber="166">IUCN Red List category</emphasis>
:—
<taxonomicName id="4C024D424274FFF2A175FDF48A21842D" authorityName="Ranjbar &amp; Askari" authorityYear="2018" box="[518,805,592,617]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="15" pageNumber="166" phylum="Tracheophyta" rank="species" species="palanganensis">
<emphasis id="B976EAD34274FFF2A175FDF48A21842D" box="[518,805,592,617]" italics="true" pageId="15" pageNumber="166">Centaurea palanganensis</emphasis>
</taxonomicName>
is known from a few small populations, consisting of no more than 15 plants, observed in a mountain between Palangan and Tefin villages (M. Ranjbar, pers. obs.). According to the IUCN Red List criteria (
<bibRefCitation id="EF934B304274FFF2A299FD3C8B7184F5" author="IUCN &amp; Species Survival Commission, IUCN" box="[490,629,664,689]" pageId="15" pageNumber="166" refId="ref6006" refString="IUCN (2001) IUCN Red List Categories and Criteria. Version 3.1. Species Survival Commission, IUCN, Gland and Cambridge, 33 pp." type="book" year="2001">IUCN 2001</bibRefCitation>
), this species could be classified as Critically Endangered (CR D). This category includes taxa for which fewer than 50 mature individuals exist.
</paragraph>
<paragraph id="8BBD36C14274FFF2A3CEFD4488518559" blockId="15.[136,1452,160,797]" pageId="15" pageNumber="166">
<emphasis id="B976EAD34274FFF2A3CEFD4488C784BD" bold="true" box="[189,451,736,761]" pageId="15" pageNumber="166">Chromosome number</emphasis>
:—
<taxonomicName id="4C024D424274FFF2A295FD448A0084BD" authorityName="Ranjbar &amp; Askari" authorityYear="2018" box="[486,772,736,761]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="15" pageNumber="166" phylum="Tracheophyta" rank="species" species="palanganensis">
<emphasis id="B976EAD34274FFF2A295FD448A0084BD" box="[486,772,736,761]" italics="true" pageId="15" pageNumber="166">Centaurea palanganensis</emphasis>
</taxonomicName>
is a diploid species with the base chromosome number of 2
<emphasis id="B976EAD34274FFF2A6EEFD448CAF84BD" box="[1437,1451,736,761]" italics="true" pageId="15" pageNumber="166">n</emphasis>
= 2
<emphasis id="B976EAD34274FFF2A3DEFCA089BD8559" box="[173,185,772,797]" italics="true" pageId="15" pageNumber="166">x</emphasis>
= 16 (
<figureCitation id="13392A444274FFF2A271FCA088438559" box="[258,327,772,797]" captionStart="FIGURE 7" captionStartId="10.[136,229,1871,1892]" captionTargetBox="[221,1365,209,1846]" captionTargetId="figure-17@10.[221,1366,209,1846]" captionTargetPageId="10" captionText="FIGURE 7. Representative meiotic cells of Centaurea masjedsoleymanensis A–G and C. palanganensis H–O. A. diakinesis. B. anaphase I with bridge. C. telophase I. D. anaphase II. E. metaphase I with precocious chromosome. F. anaphase I. G. anaphase I with bridge. H. anaphase I with bridge. I &amp; J. metaphase I with precocious chromosome. K. diakinesis. L. metaphase I. M. anaphase I. N. anaphase I with laggard. O. bivalent at diakinesis." figureDoi="http://doi.org/10.5281/zenodo.13705403" httpUri="https://zenodo.org/record/13705403/files/figure.png" pageId="15" pageNumber="166">Fig. 7</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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