treatments-xml/data/03/9D/87/039D87F55532FFCAFF4AF907FACBBA85.xml

<document ID-DOI="http://dx.doi.org/10.11646/zootaxa.4038.1.1" ID-GBIF-Dataset="066e84f1-206b-40cb-a615-2126696b813d" ID-ISSN="1175-5326" ID-Zenodo-Dep="289876" ID-ZooBank="1B62B78C-AA59-4417-A4FC-1CC9CED745E0" checkinTime="1461316934850" checkinUser="plazi" docAuthor="Janák, Jiří &amp; Bordoni, Arnaldo" docDate="2015" docId="039D87F55532FFCAFF4AF907FACBBA85" docLanguage="en" docName="zt04038p094.pdf" docOrigin="Zootaxa 4038 (1)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@509564af" docStyleName="zootaxa.2013.monograph" docTitle="Afrus spegazzinii Bernhauer 1915, comb. nov." docType="treatment" docVersion="6" lastPageNumber="12" masterDocId="FFA4FF8D553BFFC1FFDDFFA6FFFFBE25" masterDocTitle="Revision of the genus Thyreocephalus and description of Afrus gen. nov. of Africa south of the Sahara (Coleoptera: Staphylinidae: Staphylininae)" masterLastPageNumber="94" masterPageNumber="1" pageNumber="10" updateTime="1661641859159" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Revision of the genus Thyreocephalus and description of Afrus gen. nov. of Africa south of the Sahara (Coleoptera: Staphylinidae: Staphylininae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Janák, Jiří</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Bordoni, Arnaldo</mods:namePart>
</mods:name>
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<mods:titleInfo>
<mods:title>Zootaxa</mods:title>
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<mods:date>2015</mods:date>
<mods:detail type="volume">
<mods:number>4038</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
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<mods:start>1</mods:start>
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<mods:identifier type="DOI">http://dx.doi.org/10.11646/zootaxa.4038.1.1</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.6101601" ID-GBIF-Taxon="127693939" ID-Zenodo-Dep="6101601" LSID="urn:lsid:plazi:treatment:039D87F55532FFCAFF4AF907FACBBA85" httpUri="http://treatment.plazi.org/id/039D87F55532FFCAFF4AF907FACBBA85" lastPageId="11" lastPageNumber="12" pageId="9" pageNumber="10">
<subSubSection pageId="9" pageNumber="10" type="nomenclature">
<paragraph blockId="9.[151,740,1697,1756]" box="[151,740,1697,1723]" pageId="9" pageNumber="10">
<heading bold="true" box="[151,740,1697,1723]" fontSize="11" level="1" pageId="9" pageNumber="10" reason="1">
<emphasis bold="true" box="[151,740,1697,1723]" pageId="9" pageNumber="10">
<taxonomicName authority="Bernhauer, 1915" authorityName="Bernhauer" authorityYear="1915" box="[151,592,1697,1723]" class="Insecta" family="Staphylinidae" genus="Afrus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="spegazzinii" status="comb. nov.">
<emphasis bold="true" box="[151,359,1697,1723]" italics="true" pageId="9" pageNumber="10">Afrus spegazzinii</emphasis>
(Bernhauer, 
<number box="[523,584,1697,1723]" pageId="9" pageNumber="10" value="1915.0">1915</number>
)
</taxonomicName>
, 
<taxonomicNameLabel box="[606,740,1697,1723]" pageId="9" pageNumber="10">comb. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph blockId="9.[151,740,1697,1756]" box="[151,315,1731,1756]" pageId="9" pageNumber="10">
(
<figureCitation box="[159,238,1731,1756]" captionStart="FIGURES 1 – 2. 1" captionStartId="7.[151,264,1873,1895]" captionTargetBox="[197,1414,210,1817]" captionTargetId="figure@7.[164,1439,195,1851]" captionTargetPageId="7" captionText="FIGURES 1 – 2. 1, Afrus collarti (Cameron); 2, Afrus spegazzinii (Bernhauer). Scale 1 mm." httpUri="https://zenodo.org/record/289877/files/figure.png" pageId="9" pageNumber="10">Figs. 2</figureCitation>
, 
<figureCitation box="[250,307,1731,1756]" captionStart-0="FIGURES 3 – 7. 3, 4" captionStart-1="FIGURE 8" captionStart-2="FIGURES 9 – 15" captionStartId-0="8.[151,264,1877,1899]" captionStartId-1="9.[151,250,1599,1621]" captionStartId-2="10.[151,264,1914,1936]" captionTargetBox-0="[169,1423,213,1814]" captionTargetBox-1="[158,1436,196,1576]" captionTargetBox-2="[160,1413,355,1887]" captionTargetId-0="figure@8.[150,1436,193,1828]" captionTargetId-1="figure@9.[151,1436,193,1578]" captionTargetId-2="figure@10.[151,1436,337,1893]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionTargetPageId-2="10" captionText-0="FIGURES 3 – 7. 3, 4, Afrus collarti (Cameron); 5 – 7, Afrus spegazzinii (Bernhauer). 3, 5, head and anterior part of pronotum (iop—internal ocular puncture, pop—posterior ocular punctures); 4, 7, lateral part of head (SG—subelliptical groove); 6, lateral part of ventral side of pronotum (SL—superior line, IL—inferior line). Scale 0.5 mm." captionText-1="FIGURE 8. Distribution of Afrus gen. nov. (▲ = A. collarti (Cameron), ● = A. spegazzinii (Bernhauer ))." captionText-2="FIGURES 9 – 15. Afrus spegazzinii (Bernhauer); 9, head and pronotum; 10, labrum; 11, male tergite 10; 12, male sternite 9; 13, aedeagus; 14, female genital segment (fp—fused pleurites, s—sternite, ss—supplementary sclerites); 15, metatibia (ac—apical ctenidium)." httpUri-0="https://zenodo.org/record/289878/files/figure.png" httpUri-1="https://zenodo.org/record/289879/files/figure.png" httpUri-2="https://zenodo.org/record/289880/files/figure.png" pageId="9" pageNumber="10">
5– 
<number box="[280,307,1731,1756]" pageId="9" pageNumber="10" value="15.0">15</number>
</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection box="[151,1361,1803,1825]" pageId="9" pageNumber="10" type="reference_group">
<paragraph blockId="9.[151,1361,1803,1825]" box="[151,1361,1803,1825]" pageId="9" pageNumber="10">
<treatmentCitationGroup box="[151,1361,1803,1825]" pageId="9" pageNumber="10">
<taxonomicName authority="Bernhauer, 1915a: 299" authorityName="Bernhauer" authorityPageNumber="299" authorityYear="1915" box="[151,670,1803,1825]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="spegazzinii">
<emphasis box="[151,431,1803,1825]" italics="true" pageId="9" pageNumber="10">Thyreocephalus spegazzinii</emphasis>
<treatmentCitation author="Bernhauer" box="[437,670,1803,1825]" page="299" pageId="9" pageNumber="10" year="1915">
<bibRefCitation author="Bernhauer" box="[437,619,1803,1825]" pageId="9" pageNumber="10" refString="Bernhauer, M. (1915 a) Neue Staphyliniden des tropischen Afrika. Verhandlungen der k. k. zoologisch-botanischen Gesellschaft in Wien, 65, 287 - 321." type="journal article" year="1915" yearSuffix="a">
Bernhauer, 
<number box="[554,608,1803,1825]" pageId="9" pageNumber="10" value="1915.0">1915</number>
a
</bibRefCitation>
: 
<number box="[629,670,1803,1825]" pageId="9" pageNumber="10" value="299.0">299</number>
</treatmentCitation>
</taxonomicName>
; 
<treatmentCitation author="Scheerpeltz" box="[680,929,1803,1825]" page="1317" pageId="9" pageNumber="10" year="1933">
<bibRefCitation author="Scheerpeltz" box="[680,866,1803,1825]" pageId="9" pageNumber="10" refString="Scheerpeltz, O. (1933) Staphylinidae VII. In: Schenkling, S. (Ed.), Coleopterorum Catalogus, 6 (129). Junk, Berlin, pp. 989 - 1500." type="journal article" year="1933">
Scheerpeltz, 
<number box="[811,866,1803,1825]" pageId="9" pageNumber="10" value="1933.0">1933</number>
</bibRefCitation>
: 
<number box="[875,929,1803,1825]" pageId="9" pageNumber="10" value="1317.0">1317</number>
</treatmentCitation>
; 
<treatmentCitation author="Coiffait" box="[939,1135,1803,1825]" page="156" pageId="9" pageNumber="10" year="1968">
<bibRefCitation author="Coiffait" box="[939,1084,1803,1825]" pageId="9" pageNumber="10" refString="Coiffait, H. (1968) Contribution a la connaissance des Xantholininae (Coleopteres Staphylinidae) du Gabon. Biologia Gabonica, 4 (2), 127 - 161." type="journal article" year="1968">
Coiffait, 
<number box="[1030,1084,1803,1825]" pageId="9" pageNumber="10" value="1968.0">1968</number>
</bibRefCitation>
: 
<number box="[1094,1135,1803,1825]" pageId="9" pageNumber="10" value="156.0">156</number>
</treatmentCitation>
; 
<treatmentCitation author="Herman" box="[1145,1357,1803,1825]" page="3767" pageId="9" pageNumber="10" year="2001">
<bibRefCitation author="Herman" box="[1145,1293,1803,1825]" pageId="9" pageNumber="10" refString="Herman, L. H. (2001) Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of the second millennium. VI. Staphylinine group (part 3). Bulletin of the American Museum of Natural History, 265, 3021 - 3840." type="journal article" year="2001">
Herman, 
<number box="[1239,1293,1803,1825]" pageId="9" pageNumber="10" value="2001.0">2001</number>
</bibRefCitation>
: 
<number box="[1303,1357,1803,1825]" pageId="9" pageNumber="10" value="3767.0">3767</number>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection lastPageId="11" lastPageNumber="12" pageId="9" pageNumber="10" type="materials_examined">
<paragraph blockId="9.[151,1436,1870,2003]" box="[151,493,1870,1895]" pageId="9" pageNumber="10">
<emphasis bold="true" box="[151,309,1870,1895]" pageId="9" pageNumber="10">
<typeStatus box="[151,210,1870,1895]" pageId="9" pageNumber="10">Type</typeStatus>
locality.
</emphasis>
<collectingCountry box="[316,407,1870,1895]" name="Senegal" pageId="9" pageNumber="10">Senegal</collectingCountry>
, Dakar.
</paragraph>
<paragraph blockId="9.[151,1436,1870,2003]" lastBlockId="10.[151,1436,151,285]" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
<emphasis bold="true" box="[199,635,1906,1931]" pageId="9" pageNumber="10">
<typeStatus box="[199,258,1906,1931]" pageId="9" pageNumber="10">Type</typeStatus>
material examined. 
<typeStatus box="[517,635,1906,1931]" pageId="9" pageNumber="10">Lectotype</typeStatus>
</emphasis>
(by present designation) ♂: 
<collectingCountry box="[981,1115,1906,1931]" name="Senegal" pageId="9" pageNumber="10">
<emphasis bold="true" box="[981,1115,1906,1931]" pageId="9" pageNumber="10">SENEGAL</emphasis>
</collectingCountry>
: “ 
<collectingCountry box="[1145,1236,1906,1931]" name="Senegal" pageId="9" pageNumber="10">Senegal</collectingCountry>
Dakkar, XII. 
<number box="[1393,1436,1907,1931]" pageId="9" pageNumber="10" value="913.0">913</number>
Spegazzini”, “ 
<taxonomicName box="[313,440,1943,1966]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="14" phylum="Arthropoda" rank="species" species="spegazzinii">
<emphasis box="[313,440,1943,1966]" italics="true" pageId="9" pageNumber="10">Spegazzinii</emphasis>
</taxonomicName>
, Brnh. 
<typeStatus box="[524,595,1943,1967]" pageId="9" pageNumber="10">Typus</typeStatus>
” (
<collectionCode LSID="urn:lsid:biocol.org:col:34795" box="[620,707,1943,1967]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="9" pageNumber="10">FMNH</collectionCode>
). The specimen is chosen as 
<typeStatus box="[1031,1134,1942,1967]" pageId="9" pageNumber="10">lectotype</typeStatus>
and the label: “ 
<typeStatus box="[1309,1436,1943,1967]" pageId="9" pageNumber="10">Lectotypus</typeStatus>
<taxonomicName authority="Bh., Bordoni &amp; Janak" authorityName="Bh., Bordoni &amp; Janak" box="[151,727,1978,2003]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="spegazzinii">
<emphasis box="[151,464,1978,2003]" italics="true" pageId="9" pageNumber="10">Thyreocephalus spegazzinii</emphasis>
Bh., Bordoni &amp; Janák
</taxonomicName>
des. 
<number box="[791,849,1979,2003]" pageId="9" pageNumber="10" value="2012.0">2012</number>
” was attached to this specimen. 
<typeStatus box="[1232,1397,1978,2003]" pageId="9" pageNumber="10">
<emphasis bold="true" box="[1232,1397,1978,2003]" pageId="9" pageNumber="10">Paralectotype</emphasis>
</typeStatus>
♂: 
<collectingCountry box="[151,285,151,176]" name="Senegal" pageId="10" pageNumber="11">
<emphasis bold="true" box="[151,285,151,176]" pageId="10" pageNumber="11">SENEGAL</emphasis>
</collectingCountry>
: “Dakkar, 
<collectingCountry box="[419,510,151,176]" name="Senegal" pageId="10" pageNumber="11">Senegal</collectingCountry>
, XII. 
<number box="[572,615,152,176]" pageId="10" pageNumber="11" value="913.0">913</number>
Spegazzini”, “ 
<taxonomicName box="[795,922,152,175]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="14" phylum="Arthropoda" rank="species" species="spegazzinii">
<emphasis box="[795,922,152,175]" italics="true" pageId="10" pageNumber="11">Spegazzinii</emphasis>
</taxonomicName>
, Brnh. Cotypus”(
<collectionCode LSID="urn:lsid:biocol.org:col:34795" box="[1132,1218,152,176]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="10" pageNumber="11">FMNH</collectionCode>
). The specimen is designated as 
<typeStatus box="[316,471,188,213]" pageId="10" pageNumber="11">paralectotype</typeStatus>
. The label: “ 
<typeStatus box="[622,789,188,213]" pageId="10" pageNumber="11">Paralectotypus</typeStatus>
<taxonomicName authority="Bh., Bordoni &amp; Janak" authorityName="Bh., Bordoni &amp; Janak" box="[800,1380,188,213]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="spegazzinii">
<emphasis box="[800,1113,188,213]" italics="true" pageId="10" pageNumber="11">Thyreocephalus spegazzinii</emphasis>
Bh., Bordoni &amp; Janák
</taxonomicName>
des. 
<number box="[151,209,224,248]" pageId="10" pageNumber="11" value="2012.0">2012</number>
” was attached to this specimen. Both specimens have the determination label “ 
<taxonomicName authority="Bh." authorityName="Bh." box="[1100,1360,223,248]" class="Insecta" family="Staphylinidae" genus="Afrus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="spegazzinii">
<emphasis box="[1100,1293,224,248]" italics="true" pageId="10" pageNumber="11">Afrus spegazzinii</emphasis>
(Bh.)
</taxonomicName>
, Janák &amp; Bordoni det. 
<number box="[326,387,260,284]" pageId="10" pageNumber="11" value="2014.0">2014</number>
”.
</paragraph>
<caption httpUri="https://zenodo.org/record/289880/files/figure.png" pageId="10" pageNumber="11" targetBox="[160,1413,355,1887]" targetPageId="10">
<paragraph blockId="10.[151,1436,1914,2002]" pageId="10" pageNumber="11">
<emphasis bold="true" box="[151,320,1914,1936]" pageId="10" pageNumber="11">FIGURES 9–15</emphasis>
. 
<taxonomicName authority="Bernhauer" authorityName="Bernhauer" box="[332,638,1914,1936]" class="Insecta" family="Staphylinidae" genus="Afrus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="spegazzinii">
<emphasis box="[332,506,1914,1936]" italics="true" pageId="10" pageNumber="11">Afrus spegazzinii</emphasis>
(Bernhauer)
</taxonomicName>
; 9, head and pronotum; 10, labrum; 11, male tergite 10; 12, male sternite 9; 13, aedeagus; 14, female genital segment (fp—fused pleurites, s—sternite, ss—supplementary sclerites); 15, metatibia (ac—apical ctenidium).
</paragraph>
</caption>
<paragraph blockId="11.[151,1437,151,1184]" pageId="11" pageNumber="12">
<emphasis bold="true" box="[199,697,151,176]" pageId="11" pageNumber="12">
Additional material examined. 
<collectingCountry box="[574,697,151,176]" name="Angola" pageId="11" pageNumber="12">ANGOLA</collectingCountry>
</emphasis>
: 
<number box="[713,727,152,176]" pageId="11" pageNumber="12" value="1.0">1</number>
spec.: Unterer Umba, Methner (
<collectionCode box="[1097,1183,152,176]" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="11" pageNumber="12">MFNB</collectionCode>
). 
<collectingCountry box="[1202,1333,151,176]" name="Burundi" pageId="11" pageNumber="12">
<emphasis bold="true" box="[1202,1333,151,176]" pageId="11" pageNumber="12">BURUNDI</emphasis>
</collectingCountry>
: 
<number box="[1349,1363,152,176]" pageId="11" pageNumber="12" value="2.0">2</number>
spec.: plaine de la Ruzizi, 
<date box="[382,498,188,213]" pageId="11" pageNumber="12" value="1966-02-17">
<number box="[382,410,188,213]" pageId="11" pageNumber="12" value="17.0">17</number>
.ii. 
<number box="[435,498,188,213]" pageId="11" pageNumber="12" value="1966.0">1966</number>
</date>
, N'Dani S. (
<collectionCode box="[639,728,188,212]" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="11" pageNumber="12">MRAC</collectionCode>
). 
<collectingCountry box="[748,917,188,213]" name="Cameroon" pageId="11" pageNumber="12">
<emphasis bold="true" box="[748,917,188,213]" pageId="11" pageNumber="12">CAMEROON</emphasis>
</collectingCountry>
: 
<number box="[934,948,188,212]" pageId="11" pageNumber="12" value="1.0">1</number>
spec.: Metet, A. I. Good C. M. Acc. 
<number box="[1380,1436,188,213]" pageId="11" pageNumber="12" value="6641.0">6641</number>
(
<collectionCode box="[159,245,224,248]" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="11" pageNumber="12">NMPC</collectionCode>
); 
<number box="[264,278,224,248]" pageId="11" pageNumber="12" value="1.0">1</number>
spec.: Joko, Heyne (
<collectionCode box="[513,599,224,248]" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="11" pageNumber="12">MFNB</collectionCode>
). 
<collectingCountry box="[618,1125,223,248]" name="Democratic Republic of the Congo" pageId="11" pageNumber="12">
<emphasis bold="true" box="[618,1125,223,248]" pageId="11" pageNumber="12">DEMOCRATIC REPUBLIC OF CONGO</emphasis>
</collectingCountry>
: 
<specimenCount box="[1140,1183,224,248]" pageId="11" pageNumber="12" type="generic">
<number box="[1140,1154,224,248]" pageId="11" pageNumber="12" value="1.0">1</number>
♂
</specimenCount>
: Libenge, 
<date box="[1304,1398,223,248]" pageId="11" pageNumber="12" value="1948-01-08">
<number box="[1304,1318,223,248]" pageId="11" pageNumber="12" value="8.0">8</number>
.i. 
<number box="[1334,1398,223,248]" pageId="11" pageNumber="12" value="1948.0">1948</number>
</date>
, R. Cremer &amp; M. Neuman (
<collectionCode box="[432,521,260,284]" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="11" pageNumber="12">IRSNB</collectionCode>
). 
<emphasis bold="true" box="[542,670,260,285]" pageId="11" pageNumber="12">
<collectingCountry box="[542,664,260,285]" name="Gambia" pageId="11" pageNumber="12">GAMBIA</collectingCountry>
:
</emphasis>
<number box="[680,708,260,284]" pageId="11" pageNumber="12" value="10.0">10</number>
spec.: Bathurst [= Banjul], 
<date box="[1035,1079,260,285]" pageId="11" pageNumber="12" value="1968-01">
i. 
<number box="[1047,1079,260,285]" pageId="11" pageNumber="12" value="68.0">68</number>
</date>
, Palm (
<collectionCode box="[1170,1233,260,284]" httpUri="http://grbio.org/cool/nhq2-n6y1" name="St Petersburg State University" pageId="11" pageNumber="12">ZML</collectionCode>
, 
<collectionCode box="[1248,1312,260,284]" pageId="11" pageNumber="12">JJRC</collectionCode>
); 
<number box="[1334,1361,260,284]" pageId="11" pageNumber="12" value="11.0">11</number>
spec.: Bakau, 
<date box="[238,389,295,320]" pageId="11" pageNumber="12" value="1984-11-06" valueMax="1984-11-26" valueMin="1984-11-06">
<number box="[238,298,295,320]" pageId="11" pageNumber="12" value="16.0" valueMax="26.0" valueMin="6.0">6–26</number>
.xi. 
<number box="[329,389,295,320]" pageId="11" pageNumber="12" value="1984.0">1984</number>
</date>
, T. Palm (
<collectionCode box="[506,569,296,320]" httpUri="http://grbio.org/cool/nhq2-n6y1" name="St Petersburg State University" pageId="11" pageNumber="12">ZML</collectionCode>
, 
<collectionCode box="[583,647,296,320]" pageId="11" pageNumber="12">JJRC</collectionCode>
); 
<number box="[668,682,296,320]" pageId="11" pageNumber="12" value="1.0">1</number>
spec.: oil palm and mangrove veg. close to the beach about 
<quantity box="[1378,1436,295,320]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="11" pageNumber="12" unit="km" value="5.0">5 km</quantity>
SSW Gunjur, 
<geoCoordinate box="[329,415,332,357]" direction="north" orientation="latitude" pageId="11" pageNumber="12" precision="555" value="18.45">
<number box="[329,393,332,357]" pageId="11" pageNumber="12" value="18.45">18.45</number>
N
</geoCoordinate>
, 
<geoCoordinate box="[437,527,332,356]" direction="west" orientation="longitude" pageId="11" pageNumber="12" precision="555" value="-20.3">
<number box="[437,500,332,356]" pageId="11" pageNumber="12" value="20.3">20.30</number>
W
</geoCoordinate>
, Lund Univ. Syst. Dept. Sweden-Gambia/ 
<collectingCountry box="[1055,1143,332,357]" name="Senegal" pageId="11" pageNumber="12">Senegal</collectingCountry>
, Nov. 1977, 
<date box="[1316,1436,332,357]" pageId="11" pageNumber="12" value="1977-11-13">
13.xi. 
<number box="[1375,1436,332,357]" pageId="11" pageNumber="12" value="1977.0">1977</number>
</date>
Cederholm-Danielsson-Hammarstedt-Hedquist-Samuelsson (
<collectionCode box="[844,909,368,392]" httpUri="http://grbio.org/cool/nhq2-n6y1" name="St Petersburg State University" pageId="11" pageNumber="12">ZML</collectionCode>
). 
<collectingCountry box="[933,1037,367,392]" name="Ghana" pageId="11" pageNumber="12">
<emphasis bold="true" box="[933,1037,367,392]" pageId="11" pageNumber="12">GHANA</emphasis>
</collectingCountry>
: 
<number box="[1058,1072,368,392]" pageId="11" pageNumber="12" value="1.0">1</number>
spec.: Northern Prov., Banda- Nkwanta, 
<quantity box="[268,343,404,429]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="11" pageNumber="12" unit="m" value="150.0">150 m</quantity>
, 
<geoCoordinate box="[356,428,404,428]" direction="north" orientation="latitude" pageId="11" pageNumber="12" precision="555" value="8.22">
<number box="[356,407,404,428]" pageId="11" pageNumber="12" value="8.22">8.22</number>
N
</geoCoordinate>
, 
<geoCoordinate box="[441,517,404,428]" direction="west" orientation="longitude" pageId="11" pageNumber="12" precision="555" value="-2.08">
<number box="[441,490,404,428]" pageId="11" pageNumber="12" value="2.08">2.08</number>
W
</geoCoordinate>
, 
<date box="[530,668,404,429]" pageId="11" pageNumber="12" value="1965-08-10">
<number box="[530,559,404,429]" pageId="11" pageNumber="12" value="10.0">10</number>
.viii. 
<number box="[608,668,404,429]" pageId="11" pageNumber="12" value="1965.0">1965</number>
</date>
, S. Endrödy-Younga (
<collectionCode LSID="urn:lsid:biocol.org:col:34991" box="[921,1013,404,428]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="11" pageNumber="12">HNHM</collectionCode>
). 
<collectingCountry box="[1033,1132,404,429]" name="Kenya" pageId="11" pageNumber="12">
<emphasis bold="true" box="[1033,1132,404,429]" pageId="11" pageNumber="12">KENYA</emphasis>
</collectingCountry>
: 
<number box="[1148,1162,404,428]" pageId="11" pageNumber="12" value="3.0">3</number>
spec.: Victoria Nyanza, Arcipelago di Sesse, Bogala, 
<number box="[506,565,440,464]" pageId="11" pageNumber="12" value="1908.0">1908</number>
, E. Bayon (
<collectionCode box="[714,815,440,464]" pageId="11" pageNumber="12">MCSNG</collectionCode>
, 
<collectionCode box="[833,902,440,464]" pageId="11" pageNumber="12">ABFI</collectionCode>
); 
<number box="[927,941,440,464]" pageId="11" pageNumber="12" value="1.0">1</number>
spec.: 
<collectingCountry box="[1033,1125,439,464]" name="Kenya" pageId="11" pageNumber="12">Malindi</collectingCountry>
, 
<date box="[1142,1264,439,464]" pageId="11" pageNumber="12" value="1992-09-30">
<number box="[1142,1172,439,464]" pageId="11" pageNumber="12" value="30.0">30</number>
.ix. 
<number box="[1203,1264,439,464]" pageId="11" pageNumber="12" value="1992.0">1992</number>
</date>
, L. Bartolozzi (
<collectionCode box="[159,243,476,500]" httpUri="http://grbio.org/cool/mvk4-k796" name="Museo Zoologico La Specola, Universita di Firenze" pageId="11" pageNumber="12">MZUF</collectionCode>
); 
<number box="[267,281,476,500]" pageId="11" pageNumber="12" value="3.0">3</number>
spec.: Lamu, Manda 
<collectingCountry box="[542,613,476,501]" name="Iceland" pageId="11" pageNumber="12">Island</collectingCountry>
, 
<date box="[630,800,476,501]" pageId="11" pageNumber="12" value="1976-09-20" valueMax="1976-10-02" valueMin="1976-09-20">
<number box="[630,661,476,501]" pageId="11" pageNumber="12" value="20.0">20</number>
.ix– 
<number box="[704,718,476,501]" pageId="11" pageNumber="12" value="2.0">2</number>
.x. 
<number box="[740,800,476,501]" pageId="11" pageNumber="12" value="1976.0">1976</number>
</date>
, N. Sanfilippo (
<collectionCode box="[992,1093,476,500]" pageId="11" pageNumber="12">MCSNG</collectionCode>
, 
<collectionCode box="[1109,1178,476,500]" pageId="11" pageNumber="12">ABFI</collectionCode>
); 
<number box="[1201,1215,476,500]" pageId="11" pageNumber="12" value="1.0">1</number>
spec.: Kitale env., Kapenguria, 
<date box="[298,429,511,536]" pageId="11" pageNumber="12" value="1995-12-16">
<number box="[298,327,511,536]" pageId="11" pageNumber="12" value="16.0">16</number>
.xii. 
<number box="[368,429,511,536]" pageId="11" pageNumber="12" value="1995.0">1995</number>
</date>
, Smrz (
<collectionCode box="[520,618,512,536]" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="11" pageNumber="12">NHMW</collectionCode>
); 
<number box="[640,654,512,536]" pageId="11" pageNumber="12" value="1.0">1</number>
spec.: Muhoroni, Welks (
<collectionCode box="[954,1038,512,536]" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="11" pageNumber="12">TMSA</collectionCode>
). 
<collectingCountry box="[1059,1206,511,536]" name="Tanzania" pageId="11" pageNumber="12">
<emphasis bold="true" box="[1059,1206,511,536]" pageId="11" pageNumber="12">TANZANIA</emphasis>
</collectingCountry>
: 
<number box="[1224,1238,512,536]" pageId="11" pageNumber="12" value="1.0">1</number>
spec.: Morogoro Prov., Doma, 
<geoCoordinate box="[305,433,548,573]" direction="south" orientation="latitude" pageId="11" pageNumber="12" precision="15" value="-6.9430556">
06° 
<number box="[345,376,548,573]" pageId="11" pageNumber="12" value="56.0">56</number>
' 
<number box="[379,408,548,573]" pageId="11" pageNumber="12" value="35.0">35</number>
''S
</geoCoordinate>
, 
<geoCoordinate box="[444,574,548,573]" direction="east" orientation="longitude" pageId="11" pageNumber="12" precision="15" value="37.268055">
<number box="[444,474,548,573]" pageId="11" pageNumber="12" value="37.0">37</number>
° 
<number box="[484,512,548,573]" pageId="11" pageNumber="12" value="16.0">16</number>
'05''E
</geoCoordinate>
, 
<date box="[585,716,548,573]" pageId="11" pageNumber="12" value="2004-07-21">
<number box="[585,613,548,573]" pageId="11" pageNumber="12" value="21.0">21</number>
.vii. 
<number box="[653,716,548,573]" pageId="11" pageNumber="12" value="2004.0">2004</number>
</date>
, L. Bartolozzi &amp; A. Sforzi (
<collectionCode box="[1026,1111,548,572]" httpUri="http://grbio.org/cool/mvk4-k796" name="Museo Zoologico La Specola, Universita di Firenze" pageId="11" pageNumber="12">MZUF</collectionCode>
). 
<collectingCountry box="[1128,1225,548,573]" name="Sudan" pageId="11" pageNumber="12">
<emphasis bold="true" box="[1128,1225,548,573]" pageId="11" pageNumber="12">SUDAN</emphasis>
</collectingCountry>
: 
<number box="[1239,1253,548,572]" pageId="11" pageNumber="12" value="1.0">1</number>
spec.: Shambat, 
<date box="[151,231,583,608]" pageId="11" pageNumber="12" value="1966-10">
x. 
<number box="[170,231,583,608]" pageId="11" pageNumber="12" value="1966.0">1966</number>
</date>
, P. Štys (
<collectionCode box="[332,397,584,608]" pageId="11" pageNumber="12">JJRC</collectionCode>
).
</paragraph>
<paragraph blockId="11.[151,1437,151,1184]" box="[199,1213,620,645]" pageId="11" pageNumber="12">
<emphasis bold="true" box="[199,676,620,645]" pageId="11" pageNumber="12">
Cited material (not examined). 
<collectingCountry box="[573,676,620,645]" name="Gabon" pageId="11" pageNumber="12">GABON</collectingCountry>
</emphasis>
: Makokou; 
<collectingCountry box="[812,946,620,645]" name="Senegal" pageId="11" pageNumber="12">
<emphasis bold="true" box="[812,946,620,645]" pageId="11" pageNumber="12">SENEGAL</emphasis>
</collectingCountry>
: Thiès (
<bibRefCitation author="Coiffait" box="[1037,1200,620,645]" pageId="11" pageNumber="12" refString="Coiffait, H. (1968) Contribution a la connaissance des Xantholininae (Coleopteres Staphylinidae) du Gabon. Biologia Gabonica, 4 (2), 127 - 161." type="journal article" year="1968">
Coiffait, 
<number box="[1140,1200,620,645]" pageId="11" pageNumber="12" value="1968.0">1968</number>
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="12" type="description">
<paragraph blockId="11.[151,1437,151,1184]" pageId="11" pageNumber="12">
<emphasis bold="true" box="[199,372,655,680]" pageId="11" pageNumber="12">Redescription.</emphasis>
Body length 
<quantity box="[524,636,655,680]" metricMagnitude="-3" metricUnit="m" metricValue="6.75" metricValueMax="7.0" metricValueMin="6.5" pageId="11" pageNumber="12" unit="mm" value="6.75" valueMax="7.0" valueMin="6.5">6.5-7 mm</quantity>
; length from anterior margin of head to posterior margin of elytra: 
<quantity metricMagnitude="-3" metricUnit="m" metricValue="3.6" metricValueMax="3.7" metricValueMin="3.5" pageId="11" pageNumber="12" unit="mm" value="3.6" valueMax="3.7" valueMin="3.5">3.5– 3.7 mm</quantity>
. Head black, pronotum and abdomen reddish brown, elytra light yellow, light brown around scutellum, scutellum, antennae and legs light brown (
<figureCitation box="[635,706,727,752]" captionStart="FIGURES 1 – 2. 1" captionStartId="7.[151,264,1873,1895]" captionTargetBox="[197,1414,210,1817]" captionTargetId="figure@7.[164,1439,195,1851]" captionTargetPageId="7" captionText="FIGURES 1 – 2. 1, Afrus collarti (Cameron); 2, Afrus spegazzinii (Bernhauer). Scale 1 mm." httpUri="https://zenodo.org/record/289877/files/figure.png" pageId="11" pageNumber="12">
Fig. 
<number box="[689,706,728,752]" pageId="11" pageNumber="12" value="2.0">2</number>
</figureCitation>
). Labrum lobate, as in 
<figureCitation box="[972,1056,727,752]" captionStart="FIGURES 9 – 15" captionStartId="10.[151,264,1914,1936]" captionTargetBox="[160,1413,355,1887]" captionTargetId="figure@10.[151,1436,337,1893]" captionTargetPageId="10" captionText="FIGURES 9 – 15. Afrus spegazzinii (Bernhauer); 9, head and pronotum; 10, labrum; 11, male tergite 10; 12, male sternite 9; 13, aedeagus; 14, female genital segment (fp—fused pleurites, s—sternite, ss—supplementary sclerites); 15, metatibia (ac—apical ctenidium)." httpUri="https://zenodo.org/record/289880/files/figure.png" pageId="11" pageNumber="12">
Fig. 
<number box="[1025,1056,728,752]" pageId="11" pageNumber="12" value="10.0">10</number>
</figureCitation>
. Head and pronotum with micropunctation. Head and pronotum and related punctation as in 
<figureCitation box="[854,967,764,789]" captionStart="FIGURES 3 – 7. 3, 4" captionStartId="8.[151,264,1877,1899]" captionTargetBox="[169,1423,213,1814]" captionTargetId="figure@8.[150,1436,193,1828]" captionTargetPageId="8" captionText="FIGURES 3 – 7. 3, 4, Afrus collarti (Cameron); 5 – 7, Afrus spegazzinii (Bernhauer). 3, 5, head and anterior part of pronotum (iop—internal ocular puncture, pop—posterior ocular punctures); 4, 7, lateral part of head (SG—subelliptical groove); 6, lateral part of ventral side of pronotum (SL—superior line, IL—inferior line). Scale 0.5 mm." httpUri="https://zenodo.org/record/289878/files/figure.png" pageId="11" pageNumber="12">Figs. 5, 7</figureCitation>
, 
<figureCitation box="[981,998,764,788]" captionStart="FIGURES 9 – 15" captionStartId="10.[151,264,1914,1936]" captionTargetBox="[160,1413,355,1887]" captionTargetId="figure@10.[151,1436,337,1893]" captionTargetPageId="10" captionText="FIGURES 9 – 15. Afrus spegazzinii (Bernhauer); 9, head and pronotum; 10, labrum; 11, male tergite 10; 12, male sternite 9; 13, aedeagus; 14, female genital segment (fp—fused pleurites, s—sternite, ss—supplementary sclerites); 15, metatibia (ac—apical ctenidium)." httpUri="https://zenodo.org/record/289880/files/figure.png" pageId="11" pageNumber="12">
<number box="[981,998,764,788]" pageId="11" pageNumber="12" value="9.0">9</number>
</figureCitation>
. Two postocular punctures situated in slightly elongate fossa. Elytra semimembranous, longer and wider than pronotum, with marked humeral angles. Surface with fine and dense punctation, arranged in numerous series, among which are setiferous series near suture, and median and lateral series. Abdomen shiny with traces of transverse micro-striation and fine, dense punctation, provided by long pubescence.
</paragraph>
<paragraph blockId="11.[151,1437,151,1184]" pageId="11" pageNumber="12">
<collectingCountry box="[199,260,943,968]" name="Malta" pageId="11" pageNumber="12">Male</collectingCountry>
. Tergite 
<number box="[357,385,944,968]" pageId="11" pageNumber="12" value="10.0">10</number>
and sternite 
<number box="[530,544,944,968]" pageId="11" pageNumber="12" value="9.0">9</number>
of male genital segment as in 
<figureCitation box="[889,1025,943,968]" captionStart="FIGURES 9 – 15" captionStartId="10.[151,264,1914,1936]" captionTargetBox="[160,1413,355,1887]" captionTargetId="figure@10.[151,1436,337,1893]" captionTargetPageId="10" captionText="FIGURES 9 – 15. Afrus spegazzinii (Bernhauer); 9, head and pronotum; 10, labrum; 11, male tergite 10; 12, male sternite 9; 13, aedeagus; 14, female genital segment (fp—fused pleurites, s—sternite, ss—supplementary sclerites); 15, metatibia (ac—apical ctenidium)." httpUri="https://zenodo.org/record/289880/files/figure.png" pageId="11" pageNumber="12">Figs. 11, 12</figureCitation>
. Aedeagus (
<figureCitation box="[1163,1244,943,968]" captionStart="FIGURES 9 – 15" captionStartId="10.[151,264,1914,1936]" captionTargetBox="[160,1413,355,1887]" captionTargetId="figure@10.[151,1436,337,1893]" captionTargetPageId="10" captionText="FIGURES 9 – 15. Afrus spegazzinii (Bernhauer); 9, head and pronotum; 10, labrum; 11, male tergite 10; 12, male sternite 9; 13, aedeagus; 14, female genital segment (fp—fused pleurites, s—sternite, ss—supplementary sclerites); 15, metatibia (ac—apical ctenidium)." httpUri="https://zenodo.org/record/289880/files/figure.png" pageId="11" pageNumber="12">
Fig. 
<number box="[1216,1244,944,968]" pageId="11" pageNumber="12" value="13.0">13</number>
</figureCitation>
) relatively large, 
<quantity box="[151,255,980,1005]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" pageId="11" pageNumber="12" unit="mm" value="1.25">1.25 mm</quantity>
long; median lobe protruding, with rounded apex; internal sac narrow, folded on itself, covered with evident scales; parameres short.
</paragraph>
<paragraph blockId="11.[151,1437,151,1184]" pageId="11" pageNumber="12">
<emphasis bold="true" box="[199,465,1052,1077]" pageId="11" pageNumber="12">Differential diagnosis.</emphasis>
<taxonomicName box="[476,671,1052,1076]" class="Insecta" family="Staphylinidae" genus="Afrus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="spegazzinii">
<emphasis box="[476,671,1052,1076]" italics="true" pageId="11" pageNumber="12">Afrus spegazzinii</emphasis>
</taxonomicName>
differs from 
<taxonomicName box="[831,948,1052,1077]" class="Insecta" family="Staphylinidae" genus="Afrus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="collarti">
<emphasis box="[831,948,1052,1077]" italics="true" pageId="11" pageNumber="12">A. collarti</emphasis>
</taxonomicName>
in the shorter impression with postocular setiferous punctures behind the eyes, yellowish elytra, smaller body size and apically narrowed median lobe of the aedeagus.
</paragraph>
</subSubSection>
<subSubSection box="[199,1332,1159,1184]" pageId="11" pageNumber="12" type="distribution">
<paragraph blockId="11.[151,1437,151,1184]" box="[199,1332,1159,1184]" pageId="11" pageNumber="12">
<emphasis bold="true" box="[199,352,1159,1184]" pageId="11" pageNumber="12">Distribution.</emphasis>
The species is distributed in West, Central, East and Southern (
<collectingCountry box="[1063,1149,1159,1184]" name="Angola" pageId="11" pageNumber="12">Angola</collectingCountry>
) Africa (
<figureCitation box="[1250,1317,1159,1184]" captionStart="FIGURE 8" captionStartId="9.[151,250,1599,1621]" captionTargetBox="[158,1436,196,1576]" captionTargetId="figure@9.[151,1436,193,1578]" captionTargetPageId="9" captionText="FIGURE 8. Distribution of Afrus gen. nov. (▲ = A. collarti (Cameron), ● = A. spegazzinii (Bernhauer ))." httpUri="https://zenodo.org/record/289879/files/figure.png" pageId="11" pageNumber="12">
Fig. 
<number box="[1302,1317,1160,1184]" pageId="11" pageNumber="12" value="8.0">8</number>
</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>