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<document id="D7AE84EB0A4B9AA31E646EC8B7437B22" ID-CLB-Dataset="51460" ID-DOI="10.5281/zenodo.4619710" ID-GBIF-Dataset="3d6e0742-833f-461a-b5c4-800796980bda" ID-ISSN="1175­5326" ID-Zenodo-Dep="4619710" IM.metadata_requiresApprovalFor="plazi" IM.taxonomicNames_requiresApprovalFor="plazi" checkinTime="1459432144357" checkinUser="plazi" docAuthor="Ratcliffe, Brett C." docDate="2002" docId="039D87EFFF8A1B00FE81F9E672D411BC" docLanguage="en" docName="zt00032.pdf" docOrigin="Zootaxa 32" docStyle="DocumentStyle:83E4DCE73C651764D97D3683740715A6.1:zzz_test.0000.journal_article" docStyleId="83E4DCE73C651764D97D3683740715A6" docStyleName="zzz_test.0000.journal_article" docStyleVersion="1" docTitle="Pentodontini Mulsant 1842" docType="treatment" docVersion="12" lastPageNumber="33" masterDocId="FFA4FF97FF941B20FF89FF8D71661042" masterDocTitle="A checklist of the Scarabaeoidea (Coleoptera) of Panama" masterLastPageNumber="48" masterPageNumber="1" pageNumber="31" updateTime="1698192167068" updateUser="ExternalLinkService">
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<mods:title id="B9A422FCB45769E77F0C11CE8AC2F227">A checklist of the Scarabaeoidea (Coleoptera) of Panama</mods:title>
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<mods:date id="894B519FD0135F086EB6BBC700720E19">2002</mods:date>
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<treatment id="039D87EFFF8A1B00FE81F9E672D411BC" ID-DOI="http://doi.org/10.5281/zenodo.4619801" ID-GBIF-Taxon="119329580" ID-Zenodo-Dep="4619801" LSID="urn:lsid:plazi:treatment:039D87EFFF8A1B00FE81F9E672D411BC" httpUri="http://treatment.plazi.org/id/039D87EFFF8A1B00FE81F9E672D411BC" lastPageId="32" lastPageNumber="33" pageId="30" pageNumber="31">
<subSubSection id="C32E6572FF8A1B3EFE81F9E6708B16C7" box="[264,493,1643,1669]" pageId="30" pageNumber="31" type="nomenclature">
<paragraph id="8B8B36F9FF8A1B3EFE81F9E6708B16C7" blockId="30.[264,954,1643,1750]" box="[264,493,1643,1669]" pageId="30" pageNumber="31">
<heading id="D0C38195FF8A1B3EFE81F9E6708B16C7" bold="true" box="[264,493,1643,1669]" fontSize="11" level="1" pageId="30" pageNumber="31" reason="1">
<taxonomicName id="4C344D7AFF8A1B3EFE81F9E6708B16C7" ID-CoL="8RYTT" authorityName="Mulsant" authorityYear="1842" box="[264,493,1643,1669]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="tribe" tribe="Pentodontini">
<emphasis id="B940EAEBFF8A1B3EFE81F9E6708B16C7" bold="true" box="[264,493,1643,1669]" pageId="30" pageNumber="31">PENTODONTINI</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32E6572FF8A1B00FE81F91E72D411BC" lastPageId="32" lastPageNumber="33" pageId="30" pageNumber="31" type="multiple">
<paragraph id="8B8B36F9FF8A1B3EFE81F91E723116EC" blockId="30.[264,954,1643,1750]" box="[264,855,1683,1710]" pageId="30" pageNumber="31">
<taxonomicName id="4C344D7AFF8A1B3EFE81F91E732116EC" authority="Ratcliffe 1981" authorityName="Ratcliffe" authorityYear="1981" box="[264,583,1683,1710]" class="Insecta" family="Dynastidae" genus="Barutus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="B940EAEBFF8A1B3EFE81F91E70F216EF" bold="true" box="[264,404,1683,1709]" pageId="30" pageNumber="31">BARUTUS</emphasis>
Ratcliffe 1981
</taxonomicName>
. Coleop. Bull. 35: 463.
</paragraph>
<paragraph id="8B8B36F9FF8A1B3EFE81F93172DC1694" blockId="30.[264,954,1643,1750]" box="[264,954,1724,1750]" pageId="30" pageNumber="31">
<taxonomicName id="4C344D7AFF8A1B3EFE81F93173571694" authority="Ratcliffe 1981" authorityName="Ratcliffe" authorityYear="1981" box="[264,561,1724,1750]" class="Insecta" family="Scarabaeidae" genus="Canthomoechus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="hartmanni">hartmanni Ratcliffe 1981</taxonomicName>
. Coleop. Bull. 35: 468. Chiriqui.
</paragraph>
<paragraph id="8B8B36F9FF8A1B3EFE81F88672621764" blockId="30.[264,1026,1803,1870]" box="[264,772,1803,1830]" pageId="30" pageNumber="31">
<taxonomicName id="4C344D7AFF8A1B3EFE81F886735F1764" authority="Hope 1837" authorityName="Hope" authorityYear="1837" box="[264,569,1803,1830]" class="Insecta" family="Dynastidae" genus="Bothynus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="B940EAEBFF8A1B3EFE81F88670CA1767" bold="true" box="[264,428,1803,1829]" pageId="30" pageNumber="31">BOTHYNUS</emphasis>
Hope 1837
</taxonomicName>
. Col.
<collectingCountry id="F3237669FF8A1B3EFDF4F88173D11764" box="[637,695,1804,1830]" name="Isle of Man" pageId="30" pageNumber="31">Man</collectingCountry>
. 1: 95.
</paragraph>
<paragraph id="8B8B36F9FF8A1B3EFEB1F8B97564170C" blockId="30.[264,1026,1803,1870]" box="[312,1026,1844,1870]" pageId="30" pageNumber="31">
<taxonomicName id="4C344D7AFF8A1B3EFEB1F8B973F5170C" authority="Blanchard 1846" authorityName="Blanchard" authorityYear="1846" box="[312,659,1844,1870]" class="Insecta" family="Scarabaeidae" genus="Cratocnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="B940EAEBFF8A1B3EFEB1F8B970A8170F" box="[312,462,1844,1869]" italics="true" pageId="30" pageNumber="31">Cratocnemis</emphasis>
Blanchard 1846
</taxonomicName>
. Insectes Amer. Merid., p. 191.
</paragraph>
<paragraph id="8B8B36F9FF8B1B3FFEB1FE9172F61174" blockId="31.[264,1322,284,550]" box="[312,912,284,310]" pageId="31" pageNumber="32">
<taxonomicName id="4C344D7AFF8B1B3FFEB1FE9173F81174" authority="Burmeister 1847" authorityName="Burmeister" authorityYear="1847" box="[312,670,284,310]" class="Insecta" family="Scarabaeidae" genus="Scaptophilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="B940EAEBFF8B1B3FFEB1FE9170A91177" box="[312,463,284,309]" italics="true" pageId="31" pageNumber="32">Scaptophilus</emphasis>
Burmeister 1847
</taxonomicName>
. Handb. Ent. 5: 122.
</paragraph>
<paragraph id="8B8B36F9FF8B1B3FFEB1FEC972F4111C" blockId="31.[264,1322,284,550]" box="[312,914,324,350]" pageId="31" pageNumber="32">
<taxonomicName id="4C344D7AFF8B1B3FFEB1FEC973C6111C" authority="Burmeister 1847" authorityName="Burmeister" authorityYear="1847" box="[312,672,324,350]" class="Insecta" family="Scarabaeidae" genus="Corynoscelis" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="B940EAEBFF8B1B3FFEB1FEC970B7111F" box="[312,465,324,349]" italics="true" pageId="31" pageNumber="32">Corynoscelis</emphasis>
Burmeister 1847
</taxonomicName>
. Handb. Ent. 5: 126.
</paragraph>
<paragraph id="8B8B36F9FF8B1B3FFE81FEE175D011C4" blockId="31.[264,1322,284,550]" box="[264,1206,364,390]" pageId="31" pageNumber="32">
<taxonomicName id="4C344D7AFF8B1B3FFE81FEE1730D11C4" authority="(Burmeister 1847)" baseAuthorityName="Burmeister" baseAuthorityYear="1847" box="[264,619,364,390]" class="Insecta" family="Dynastidae" genus="Bothynus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="complanus">complanus (Burmeister 1847)</taxonomicName>
. Handb. Ent. 5: 123. Canal Zone, Colón,
<collectingCountry id="F3237669FF8B1B3FFBD8FEE175D411C4" box="[1105,1202,364,390]" name="Panama" pageId="31" pageNumber="32">Panama</collectingCountry>
.
</paragraph>
<paragraph id="8B8B36F9FF8B1B3FFEB1FE1972A511EC" blockId="31.[264,1322,284,550]" box="[312,963,404,430]" pageId="31" pageNumber="32">
<taxonomicName id="4C344D7AFF8B1B3FFEB1FE19730511EC" authority="Bates 1888" authorityName="Bates" authorityYear="1888" box="[312,611,404,430]" class="Insecta" family="Dynastidae" genus="Bothynus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="simplicitarsis">
<emphasis id="B940EAEBFF8B1B3FFEB1FE1970B011EF" box="[312,470,404,429]" italics="true" pageId="31" pageNumber="32">simplicitarsis</emphasis>
Bates 1888
</taxonomicName>
. Biol. Cent.­Amer. 2 (2): 319.
</paragraph>
<paragraph id="8B8B36F9FF8B1B3FFEB1FE3172CC1194" blockId="31.[264,1322,284,550]" box="[312,938,444,470]" pageId="31" pageNumber="32">
<taxonomicName id="4C344D7AFF8B1B3FFEB1FE31732C1194" authority="Bates 1888" authorityName="Bates" authorityYear="1888" box="[312,586,444,470]" class="Insecta" family="Scarabaeidae" genus="Brachycopris" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="monstrosus">
<emphasis id="B940EAEBFF8B1B3FFEB1FE3170DA1197" box="[312,444,444,469]" italics="true" pageId="31" pageNumber="32">monstrosus</emphasis>
Bates 1888
</taxonomicName>
. Biol. Cent.­Amer. 2 (2): 320.
</paragraph>
<paragraph id="8B8B36F9FF8B1B3FFE81FE6970C11264" blockId="31.[264,1322,284,550]" pageId="31" pageNumber="32">
<taxonomicName id="4C344D7AFF8B1B3FFE81FE6973E711BC" authority="(Taschenberg 1870)" baseAuthorityName="Taschenberg" baseAuthorityYear="1870" box="[264,641,484,510]" class="Insecta" family="Scarabaeidae" genus="Canthidium" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="quadridens">quadridens (Taschenberg 1870)</taxonomicName>
. Zeit. Ges. Natw. 1: 185. Canal Zone, Chiriqui,
<collectingCountry id="F3237669FF8B1B3FFB4FFE69744011BC" box="[1222,1318,484,510]" name="Panama" pageId="31" pageNumber="32">Panama</collectingCountry>
, Veraguas.
</paragraph>
<paragraph id="8B8B36F9FF8B1B3FFE81FDD672C91234" blockId="31.[264,1262,603,910]" box="[264,943,603,630]" pageId="31" pageNumber="32">
<taxonomicName id="4C344D7AFF8B1B3FFE81FDD673291234" authority="Bates 1888" authorityName="Bates" authorityYear="1888" box="[264,591,603,630]" class="Insecta" family="Dynastidae" genus="Euetheola" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="B940EAEBFF8B1B3FFE81FDD670A71237" bold="true" box="[264,449,603,629]" pageId="31" pageNumber="32">EUETHEOLA</emphasis>
Bates 1888
</taxonomicName>
. Biol. Cent.­Amer. 2 (2): 314.
</paragraph>
<paragraph id="8B8B36F9FF8B1B3FFE81FD0972F112DC" blockId="31.[264,1262,603,910]" box="[264,919,644,670]" pageId="31" pageNumber="32">
<taxonomicName id="4C344D7AFF8B1B3FFE81FD09733012DC" authority="(Burmeister 1847)" baseAuthorityName="Burmeister" baseAuthorityYear="1847" box="[264,598,644,670]" class="Insecta" family="Rutelidae" genus="Chlorota" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="bidentata">bidentata (Burmeister 1847)</taxonomicName>
. Handb. Ent. 5: 81. Darien.
</paragraph>
<paragraph id="8B8B36F9FF8B1B3FFEB1FD2172781284" blockId="31.[264,1262,603,910]" box="[312,798,684,710]" pageId="31" pageNumber="32">
?
<taxonomicName id="4C344D7AFF8B1B3FFECCFD21734C1284" authority="(Perty 1830)" authorityName="Perty" authorityYear="1830" baseAuthorityName="Perty" baseAuthorityYear="1830" box="[325,554,684,710]" class="Insecta" family="Rutelidae" genus="Chrysina" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="brevis">
<emphasis id="B940EAEBFF8B1B3FFECCFD2170EA1287" box="[325,396,684,709]" italics="true" pageId="31" pageNumber="32">brevis</emphasis>
(Perty 1830)
</taxonomicName>
. Del. Animal., p. 46.
</paragraph>
<paragraph id="8B8B36F9FF8B1B3FFEB1FD59721712AC" blockId="31.[264,1262,603,910]" box="[312,881,724,750]" pageId="31" pageNumber="32">
?
<taxonomicName id="4C344D7AFF8B1B3FFECCFD5973F612AC" authority="(Burmeister 1847)" baseAuthorityName="Burmeister" baseAuthorityYear="1847" box="[325,656,724,750]" class="Insecta" family="Scarabaeidae" genus="Anomiopus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="globosus">
<emphasis id="B940EAEBFF8B1B3FFECCFD5970CB12AF" box="[325,429,724,749]" italics="true" pageId="31" pageNumber="32">globosus</emphasis>
(Burmeister 1847)
</taxonomicName>
. Handb. Ent. 5: 91.
</paragraph>
<paragraph id="8B8B36F9FF8B1B3FFE81FD7175881354" blockId="31.[264,1262,603,910]" box="[264,1262,764,790]" pageId="31" pageNumber="32">
<taxonomicName id="4C344D7AFF8B1B3FFE81FD7173251354" authority="(Burmeister 1847)" baseAuthorityName="Burmeister" baseAuthorityYear="1847" box="[264,579,764,790]" class="Insecta" family="Cetoniidae" genus="Rhipiphorina" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="humilis">humilis (Burmeister 1847)</taxonomicName>
. Handb. Ent. 5: 93. Canal Zone, Chiriqui, Colón,
<collectingCountry id="F3237669FF8B1B3FFB00FD71758C1354" box="[1161,1258,764,790]" name="Panama" pageId="31" pageNumber="32">Panama</collectingCountry>
.
</paragraph>
<paragraph id="8B8B36F9FF8B1B3FFEB1FCA972BE137C" blockId="31.[264,1262,603,910]" box="[312,984,804,830]" pageId="31" pageNumber="32">
<taxonomicName id="4C344D7AFF8B1B3FFEB1FCA97328137C" authority="LeConte 1856" authorityName="LeConte" authorityYear="1856" box="[312,590,804,830]" class="Insecta" family="Scarabaeidae" genus="Ligyrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="rugiceps">
<emphasis id="B940EAEBFF8B1B3FFEB1FCA970FB137F" box="[312,413,804,829]" italics="true" pageId="31" pageNumber="32">rugiceps</emphasis>
LeConte 1856
</taxonomicName>
. Proc. Acad. Nat. Sci. Phil. 8: 21.
</paragraph>
<paragraph id="8B8B36F9FF8B1B3FFEB1FCC1724E1324" blockId="31.[264,1262,603,910]" box="[312,808,844,870]" pageId="31" pageNumber="32">
<taxonomicName id="4C344D7AFF8B1B3FFEB1FCC1737A1324" authority="Casey 1915" authorityName="Casey" authorityYear="1915" box="[312,540,844,870]" class="Adenophorea" family="Plectidae" genus="Plectus" kingdom="Animalia" order="Araeolaimida" pageId="31" pageNumber="32" phylum="Nematoda" rank="species" species="parvus">
<emphasis id="B940EAEBFF8B1B3FFEB1FCC170EE1327" box="[312,392,844,869]" italics="true" pageId="31" pageNumber="32">parvus</emphasis>
Casey 1915
</taxonomicName>
. Mem. Coleop. 6: 172.
</paragraph>
<paragraph id="8B8B36F9FF8B1B3FFEB1FCF9723F13CC" blockId="31.[264,1262,603,910]" box="[312,857,884,910]" pageId="31" pageNumber="32">
<taxonomicName id="4C344D7AFF8B1B3FFEB1FCF9732A13CC" authority="Casey 1915" authorityName="Casey" authorityYear="1915" box="[312,588,884,910]" class="Insecta" family="Dynastidae" genus="Euetheola" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="hondurana">
<emphasis id="B940EAEBFF8B1B3FFEB1FCF970DF13CF" box="[312,441,884,909]" italics="true" pageId="31" pageNumber="32">hondurana</emphasis>
Casey 1915
</taxonomicName>
. Mem. Coleop. 6: 188.
</paragraph>
<paragraph id="8B8B36F9FF8B1B3FFE81FC33722E139B" blockId="31.[264,1322,958,1904]" box="[264,840,958,985]" pageId="31" pageNumber="32">
<taxonomicName id="4C344D7AFF8B1B3FFE81FC337334139B" authority="Erichson 1847" authorityName="Erichson" authorityYear="1847" box="[264,594,958,985]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="B940EAEBFF8B1B3FFE81FC3370FB139A" bold="true" box="[264,413,958,984]" pageId="31" pageNumber="32">TOMARUS</emphasis>
Erichson 1847
</taxonomicName>
. Arch. Natur. 13: 95.
</paragraph>
<paragraph id="8B8B36F9FF8B1B3FFEB1FC6A72371443" blockId="31.[264,1322,958,1904]" box="[312,849,999,1025]" pageId="31" pageNumber="32">
<taxonomicName id="4C344D7AFF8B1B3FFEB1FC6A73061443" authority="Burmeister 1847" authorityName="Burmeister" authorityYear="1847" box="[312,608,999,1025]" class="Insecta" family="Scarabaeidae" genus="Ligyrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="B940EAEBFF8B1B3FFEB1FC6A70F71442" box="[312,401,999,1024]" italics="true" pageId="31" pageNumber="32">Ligyrus</emphasis>
Burmeister 1847
</taxonomicName>
. Handb. Ent. 5: 542.
</paragraph>
<paragraph id="8B8B36F9FF8B1B3FFEB1FB82722B146B" blockId="31.[264,1322,958,1904]" box="[312,845,1039,1065]" pageId="31" pageNumber="32">
<taxonomicName id="4C344D7AFF8B1B3FFEB1FB827326146B" authority="Casey 1915" authorityName="Casey" authorityYear="1915" box="[312,576,1039,1065]" class="Insecta" family="Scarabaeidae" genus="Ligyrodes" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="B940EAEBFF8B1B3FFEB1FB8270CA146A" box="[312,428,1039,1064]" italics="true" pageId="31" pageNumber="32">Ligyrodes</emphasis>
Casey 1915
</taxonomicName>
. Mem. Coleop. 6: 178.
</paragraph>
<paragraph id="8B8B36F9FF8B1B3FFEB1FBBA722D1413" blockId="31.[264,1322,958,1904]" box="[312,843,1079,1105]" pageId="31" pageNumber="32">
<taxonomicName id="4C344D7AFF8B1B3FFEB1FBBA73581413" authority="Casey 1915" authorityName="Casey" authorityYear="1915" box="[312,574,1079,1105]" class="Insecta" family="Scarabaeidae" genus="Euligyrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="B940EAEBFF8B1B3FFEB1FBBA70CC1412" box="[312,426,1079,1104]" italics="true" pageId="31" pageNumber="32">Euligyrus</emphasis>
Casey 1915
</taxonomicName>
. Mem. Coleop. 6: 185.
</paragraph>
<paragraph id="8B8B36F9FF8B1B3FFEB1FBD27257143B" blockId="31.[264,1322,958,1904]" box="[312,817,1119,1145]" pageId="31" pageNumber="32">
<taxonomicName id="4C344D7AFF8B1B3FFEB1FBD27342143B" authority="Casey 1915" authorityName="Casey" authorityYear="1915" box="[312,548,1119,1145]" class="Insecta" genus="Grylius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="B940EAEBFF8B1B3FFEB1FBD270F6143A" box="[312,400,1119,1144]" italics="true" pageId="31" pageNumber="32">Grylius</emphasis>
Casey 1915
</taxonomicName>
. Mem. Coleop. 6: 189.
</paragraph>
<paragraph id="8B8B36F9FF8B1B3FFEB1FB0A723C14E3" blockId="31.[264,1322,958,1904]" box="[312,858,1159,1185]" pageId="31" pageNumber="32">
<taxonomicName id="4C344D7AFF8B1B3FFEB1FB0A732A14E3" authority="Casey 1915" authorityName="Casey" authorityYear="1915" box="[312,588,1159,1185]" class="Insecta" family="Scarabaeidae" genus="Anagrylius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="B940EAEBFF8B1B3FFEB1FB0A70DF14E2" box="[312,441,1159,1184]" italics="true" pageId="31" pageNumber="32">Anagrylius</emphasis>
Casey 1915
</taxonomicName>
. Mem. Coleop. 6: 204.
</paragraph>
<paragraph id="8B8B36F9FF8B1B3FFEB1FB227229148B" blockId="31.[264,1322,958,1904]" box="[312,847,1199,1225]" pageId="31" pageNumber="32">
<taxonomicName id="4C344D7AFF8B1B3FFEB1FB227324148B" authority="Casey 1915" authorityName="Casey" authorityYear="1915" box="[312,578,1199,1225]" class="Insecta" family="Dynastidae" genus="Ligyrellus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="B940EAEBFF8B1B3FFEB1FB2270CB148A" box="[312,429,1199,1224]" italics="true" pageId="31" pageNumber="32">Ligyrellus</emphasis>
Casey 1915
</taxonomicName>
. Mem. Coleop. 6: 206.
</paragraph>
<paragraph id="8B8B36F9FF8B1B3FFE81FB5A73A4155B" blockId="31.[264,1322,958,1904]" pageId="31" pageNumber="32">
<taxonomicName id="4C344D7AFF8B1B3FFE81FB5A726A14B3" authority="(Palisot de Beauvois 1805)" baseAuthorityName="Palisot de Beauvois" baseAuthorityYear="1805" box="[264,780,1239,1265]" class="Insecta" family="Dynastidae" genus="Ligyrellus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="bituberculatus">bituberculatus (Palisot de Beauvois 1805)</taxonomicName>
. Insect. Afr. et Amer., p. 103. Canal Zone, Chiriqui, Colón, Darien,
<collectingCountry id="F3237669FF8B1B3FFDD4FB7273D8155B" box="[605,702,1279,1305]" name="Panama" pageId="31" pageNumber="32">Panama</collectingCountry>
.
</paragraph>
<paragraph id="8B8B36F9FF8B1B3FFEB1FAAA72AB1503" blockId="31.[264,1322,958,1904]" box="[312,973,1319,1345]" pageId="31" pageNumber="32">
<taxonomicName id="4C344D7AFF8B1B3FFEB1FAAA734A1503" authority="Bates 1888" authorityName="Bates" authorityYear="1888" box="[312,556,1319,1345]" class="Insecta" family="Scarabaeidae" genus="Ligyrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="latifovea">
<emphasis id="B940EAEBFF8B1B3FFEB1FAAA70F81502" box="[312,414,1319,1344]" italics="true" pageId="31" pageNumber="32">latifovea</emphasis>
Bates 1888
</taxonomicName>
. Biol. Cent.­Amer., Coleop. 2: 315.
</paragraph>
<paragraph id="8B8B36F9FF8B1B3FFEB1FAC2729C152A" blockId="31.[264,1322,958,1904]" box="[312,1018,1358,1384]" pageId="31" pageNumber="32">
<taxonomicName id="4C344D7AFF8B1B3FFEB1FAC2735F152A" authority="Arrow 1913" authorityName="Arrow" authorityYear="1913" box="[312,569,1358,1384]" class="Insecta" family="Scarabaeidae" genus="Ligyrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="maximus">
<emphasis id="B940EAEBFF8B1B3FFEB1FAC270C6152A" box="[312,416,1359,1384]" italics="true" pageId="31" pageNumber="32">maximus</emphasis>
Arrow 1913
</taxonomicName>
. Ann. Mag. Nat. Hist. (ser. 8) 11: 466.
</paragraph>
<paragraph id="8B8B36F9FF8B1B3FFEB1FAFA72AA15D2" blockId="31.[264,1322,958,1904]" box="[312,972,1398,1424]" pageId="31" pageNumber="32">
<taxonomicName id="4C344D7AFF8B1B3FFEB1FAFA736C15D2" authority="Arrow 1914" authorityName="Arrow" authorityYear="1914" box="[312,522,1398,1424]" class="Insecta" family="Scarabaeidae" genus="Ligyrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="latus">
<emphasis id="B940EAEBFF8B1B3FFEB1FAFA701715D2" box="[312,369,1399,1424]" italics="true" pageId="31" pageNumber="32">latus</emphasis>
Arrow 1914
</taxonomicName>
. Ann. Mag. Nat. Hist. (ser. 8) 14: 273.
</paragraph>
<paragraph id="8B8B36F9FF8B1B3FFE81FA1375D915FA" blockId="31.[264,1322,958,1904]" box="[264,1215,1438,1464]" pageId="31" pageNumber="32">
<taxonomicName id="4C344D7AFF8B1B3FFE81FA13737315FA" authority="(DeGeer 1774)" baseAuthorityName="DeGeer" baseAuthorityYear="1774" box="[264,533,1438,1464]" class="Insecta" family="Scarabaeidae" genus="Canthomoechus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="ebenus">ebenus (DeGeer 1774)</taxonomicName>
. Mem. Ins. 4: 317. Canal Zone, Colón,
<collectingCountry id="F3237669FF8B1B3FFC6DFA13752215FA" box="[996,1092,1438,1464]" name="Panama" pageId="31" pageNumber="32">Panama</collectingCountry>
, San Blas.
</paragraph>
<paragraph id="8B8B36F9FF8B1B3FFEB1FA4A727B15A2" blockId="31.[264,1322,958,1904]" box="[312,797,1478,1504]" pageId="31" pageNumber="32">
<emphasis id="B940EAEBFF8B1B3FFEB1FA4A70FB15A2" box="[312,413,1479,1504]" italics="true" pageId="31" pageNumber="32">cordatus</emphasis>
Fabricius 1792. Ent. Syst. 1: 31.
</paragraph>
<paragraph id="8B8B36F9FF8B1B3FFEB1FA627235164A" blockId="31.[264,1322,958,1904]" box="[312,851,1518,1544]" pageId="31" pageNumber="32">
<emphasis id="B940EAEBFF8B1B3FFEB1FA6270B2164A" box="[312,468,1519,1544]" italics="true" pageId="31" pageNumber="32">scarabaeinus</emphasis>
Perty 1830. Del. Animal., p. 46.
</paragraph>
<paragraph id="8B8B36F9FF8B1B3FFE81F99B75C81672" blockId="31.[264,1322,958,1904]" box="[264,1198,1558,1584]" pageId="31" pageNumber="32">
<taxonomicName id="4C344D7AFF8B1B3FFE81F99B73751672" authority="(Latreille 1813)" baseAuthorityName="Latreille" baseAuthorityYear="1813" box="[264,531,1558,1584]" class="Insecta" family="Dynastidae" genus="Ligyrellus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="fossor">fossor (Latreille 1813)</taxonomicName>
. Voyage Humboldt and Bonpland 2: 11. Coclé,
<collectingCountry id="F3237669FF8B1B3FFBC3F99B75CD1672" box="[1098,1195,1558,1584]" name="Panama" pageId="31" pageNumber="32">Panama</collectingCountry>
.
</paragraph>
<paragraph id="8B8B36F9FF8B1B3FFEB1F9B27533161A" blockId="31.[264,1322,958,1904]" box="[312,1109,1598,1624]" pageId="31" pageNumber="32">
<taxonomicName id="4C344D7AFF8B1B3FFEB1F9B2739D161A" authority="(Palisot de Beauvois 1805)" baseAuthorityName="Palisot de Beauvois" baseAuthorityYear="1805" box="[312,763,1598,1624]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="antillarum">
<emphasis id="B940EAEBFF8B1B3FFEB1F9B270D2161A" box="[312,436,1599,1624]" italics="true" pageId="31" pageNumber="32">antillarum</emphasis>
(Palisot de Beauvois 1805)
</taxonomicName>
. Insect. Afr. et Amer., p. 104.
</paragraph>
<paragraph id="8B8B36F9FF8B1B3FFEB1F9EA756A16C2" blockId="31.[264,1322,958,1904]" box="[312,1036,1638,1664]" pageId="31" pageNumber="32">
<taxonomicName id="4C344D7AFF8B1B3FFEB1F9EA73FD16C2" authority="Apolinar 1927" authorityName="Apolinar" authorityYear="1927" box="[312,667,1638,1664]" class="Insecta" family="Scarabaeidae" genus="Aprosterna" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="castaneipennis">
<emphasis id="B940EAEBFF8B1B3FFEB1F9EA708016C2" box="[312,486,1639,1664]" italics="true" pageId="31" pageNumber="32">castaneipennis</emphasis>
Apolinar 1927
</taxonomicName>
. Bol. Soc. Colom. Sci. 16: 120.
</paragraph>
<paragraph id="8B8B36F9FF8B1B3FFE81F90370FB1692" blockId="31.[264,1322,958,1904]" pageId="31" pageNumber="32">
<taxonomicName id="4C344D7AFF8B1B3FFE81F903736D16EA" authority="(Erichson 1848)" baseAuthorityName="Erichson" baseAuthorityYear="1848" box="[264,523,1678,1704]" class="Insecta" family="Dynastidae" genus="Ligyrellus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="gyas">gyas (Erichson 1848)</taxonomicName>
. Schomb. Reise Brit.
<collectingCountry id="F3237669FF8B1B3FFC9AF903720116EA" box="[787,871,1678,1704]" name="Guyana" pageId="31" pageNumber="32">Guiana</collectingCountry>
3: 561. Canal Zone, Chiriqui, Colón,
<collectingCountry id="F3237669FF8B1B3FFEB1F93B70FF1692" box="[312,409,1718,1744]" name="Panama" pageId="31" pageNumber="32">Panama</collectingCountry>
.
</paragraph>
<paragraph id="8B8B36F9FF8B1B3FFEB1F952757816BA" blockId="31.[264,1322,958,1904]" box="[312,1054,1758,1784]" pageId="31" pageNumber="32">
<taxonomicName id="4C344D7AFF8B1B3FFEB1F952733D16BA" authority="Arrow 1914" authorityName="Arrow" authorityYear="1914" box="[312,603,1758,1784]" class="Insecta" family="Scarabaeidae" genus="Ligyrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="amazonicus">
<emphasis id="B940EAEBFF8B1B3FFEB1F95270A416BA" box="[312,450,1759,1784]" italics="true" pageId="31" pageNumber="32">amazonicus</emphasis>
Arrow 1914
</taxonomicName>
. Ann. Mag. Nat. Hist. (ser. 8) 14: 273.
</paragraph>
<paragraph id="8B8B36F9FF8B1B3FFE81F88B755D1762" blockId="31.[264,1322,958,1904]" box="[264,1083,1798,1824]" pageId="31" pageNumber="32">
<taxonomicName id="4C344D7AFF8B1B3FFE81F88B73251762" authority="(Burmeister 1847)" baseAuthorityName="Burmeister" baseAuthorityYear="1847" box="[264,579,1798,1824]" class="Insecta" family="Dynastidae" genus="Ligyrellus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="nasutus">nasutus (Burmeister 1847)</taxonomicName>
. Handb. Ent. 5: 120. Canal Zone, Chiriqui.
</paragraph>
<paragraph id="8B8B36F9FF8B1B3FFEB1F8A272BC170A" blockId="31.[264,1322,958,1904]" box="[312,986,1838,1864]" pageId="31" pageNumber="32">
<taxonomicName id="4C344D7AFF8B1B3FFEB1F8A2735F170A" authority="Bates 1888" authorityName="Bates" authorityYear="1888" box="[312,569,1838,1864]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="pygidialis">
<emphasis id="B940EAEBFF8B1B3FFEB1F8A270CD170A" box="[312,427,1839,1864]" italics="true" pageId="31" pageNumber="32">pygidialis</emphasis>
Bates 1888
</taxonomicName>
. Biol. Cent.­Amer., Coleop. 2: 317.
</paragraph>
<paragraph id="8B8B36F9FF8B1B3FFE81F8DB72B21732" blockId="31.[264,1322,958,1904]" box="[264,980,1878,1904]" pageId="31" pageNumber="32">
<taxonomicName id="4C344D7AFF8B1B3FFE81F8DB73731732" authority="(Endrodi 1968)" authorityName="Endrodi" authorityYear="1968" baseAuthorityName="Endrodi" baseAuthorityYear="1968" box="[264,533,1878,1904]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="similis">similis (Endrödi 1968)</taxonomicName>
. Fol. Ent. Hung. 21: 166. Canal Zone.
</paragraph>
<paragraph id="8B8B36F9FFB41B00FE81FE9672541174" blockId="32.[264,1050,283,350]" box="[264,818,283,310]" pageId="32" pageNumber="33">
<taxonomicName id="4C344D7AFFB41B00FE81FE9673371174" authority="Prell 1934" authorityName="Prell" authorityYear="1934" box="[264,593,283,310]" class="Insecta" family="Dynastidae" genus="Parapucaya" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="B940EAEBFFB41B00FE81FE9670AB1177" bold="true" box="[264,461,283,309]" pageId="32" pageNumber="33">PARAPUCAYA</emphasis>
Prell 1934
</taxonomicName>
. Ent. Zeit. 47: 162.
</paragraph>
<paragraph id="8B8B36F9FFB41B00FE81FEC9757C111C" blockId="32.[264,1050,283,350]" box="[264,1050,324,350]" pageId="32" pageNumber="33">
<taxonomicName id="4C344D7AFFB41B00FE81FEC9736F111C" authority="Prell 1934" authorityName="Prell" authorityYear="1934" box="[264,521,324,350]" class="Insecta" family="Scarabaeidae" genus="Pseudoserica" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="amazonica">amazonica Prell 1934</taxonomicName>
. Ent. Zeit. 47: 162. Bocas del Toro, Chiriqui.
</paragraph>
<paragraph id="8B8B36F9FFB41B00FE81FE1E72EF11EC" blockId="32.[264,1189,403,510]" box="[264,905,403,430]" pageId="32" pageNumber="33">
<taxonomicName id="4C344D7AFFB41B00FE81FE1E737011EC" authority="Ohaus 1910" authorityName="Ohaus" authorityYear="1910" box="[264,534,403,430]" class="Insecta" family="Dynastidae" genus="Pucaya" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="B940EAEBFFB41B00FE81FE1E701811EF" bold="true" box="[264,382,403,429]" pageId="32" pageNumber="33">PUCAYA</emphasis>
Ohaus 1910
</taxonomicName>
. Deutsche Ent. Zeit. 1910: 675.
</paragraph>
<paragraph id="8B8B36F9FFB41B00FE81FE3175C31194" blockId="32.[264,1189,403,510]" box="[264,1189,444,470]" pageId="32" pageNumber="33">
<taxonomicName id="4C344D7AFFB41B00FE81FE3173641194" authority="Ohaus 1910" authorityName="Ohaus" authorityYear="1910" box="[264,514,444,470]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="castanea">castanea Ohaus 1910</taxonomicName>
. Deutsche Ent. Zeit. 1910: 676. Bocas del Toro, Chiriqui.
</paragraph>
<paragraph id="8B8B36F9FFB41B00FEB1FE6972D411BC" blockId="32.[264,1189,403,510]" box="[312,946,484,510]" pageId="32" pageNumber="33">
<taxonomicName id="4C344D7AFFB41B00FEB1FE69732D11BC" authority="Beck 1942" authorityName="Beck" authorityYear="1942" box="[312,587,484,510]" class="Insecta" family="Melolonthidae" genus="Astaena" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="columbiana">
<emphasis id="B940EAEBFFB41B00FEB1FE6970A411BF" box="[312,450,484,509]" italics="true" pageId="32" pageNumber="33">columbiana</emphasis>
Beck 1942
</taxonomicName>
. Bull. Soc. Ent.
<collectingCountry id="F3237669FFB41B00FC83FE69723C11BC" box="[778,858,484,510]" name="France" pageId="32" pageNumber="33">France</collectingCountry>
47: 47.
</paragraph>
</subSubSection>
</treatment>
</document>
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